Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia) - Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII

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Title: Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia) - Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Author: Haeckel, Ernst, 1834-1919
Language: English
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BY H.M.S. CHALLENGER DURING THE YEARS 1873-1876, FIRST PART: PORULOSA
(SPUMELLARIA AND ACANTHARIA)***

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https://archive.org/details/reportonradiolar01haecrich

Second Part: Subclass Osculosa; Index
Plates

Transcriber's note:

Text enclosed by underscores is in italics (_Actissa_).

Text enclosed by hash marks is in "gesperrt"
(wide-spaced) type (#Larcoidea#).

Page numbers enclosed by curly braces (example: {25}) have
been incorporated to facilitate the use of the Index in
Part II.

Some typographical errors in the printed work have been
corrected. They are listed at the end of the text.

The Addenda & Errata (Second Part, pp. 1763-4) have been
applied without further comment.

REPORT ON THE SCIENTIFIC RESULTS OF THE VOYAGE OF H.M.S. CHALLENGER
DURING THE YEARS 1873-76

Under the Command of Captain George S. Nares, R.N., F.R.S.
and the Late Captain Frank Tourle Thomson, R.N.

Prepared Under the Superintendence of
the Late Sir C. Wyville Thomson, KNT., F.R.S., &c.
Regius Professor of Natural History in the University of Edinburgh
Director of the Civilian Scientific Staff on Board
and Now of
John Murray
One of the Naturalists of the Expedition

ZOOLOGY--VOL. XVIII.

FIRST PART

Published by Order of Her Majesty's Government

Printed for Her Majesty's Stationary Office
and Sold by
London:--Eyre & Spottiswoode, East Harding Street, Fetter Lane
Edinburgh:--Adam & Charles Black
Dublin:--Hodges, Figgis, & Co.
1887

Price (in Two Parts, with a Volume of Plates) £5, 10s.

CONTENTS.

REPORT on the RADIOLARIA collected by H.M.S. CHALLENGER during the years
1873-1876.

By ERNST HAECKEL, M.D., Ph.D., Professor of Zoology in the University of
Jena.

FIRST PART.--PORULOSA.

(#SPUMELLARIA AND ACANTHARIA.#)

EDITORIAL NOTES.

The Report on the RADIOLARIA by Professor Ernst Haeckel of Jena occupies
the whole of the present Volume, the text being bound up in Two Separate
Parts and the Plates in a Third Part. The Report forms Part XL. of the
Zoological Series of Reports on the Scientific Results of the Expedition,
and is the largest single Report of the series which has up to this time
been published.

The Manuscript of the Systematic Part was written by Professor Haeckel in
the English language, and was received by me in instalments on the 12th
August 1884, 13th July and 4th December 1885, and 3rd June 1886. The
Introduction was written in German and was translated into the English
language by Mr. W. E. Hoyle of the Challenger Editorial Staff; the German
text being received in instalments between the 15th July 1886, and the 25th
January 1887.

The Challenger Naturalists found the representatives of this group of
animals to be universally distributed throughout ocean waters, and their
dead remains to be nearly equally widely distributed over the floor of the
ocean, the relative abundance and the species differing, however, with
change of locality, and their abundance or variety being intimately
connected with some of the most interesting and intricate problems of
general oceanography.

It was a fortunate circumstance that so distinguished a Naturalist, with
such an intimate knowledge of the RADIOLARIA, should have been willing to
undertake the laborious examination and description of the extensive
collections made during the Expedition. Professor Haeckel has devoted ten
years of his life to this work, and this Report sets forth the results of
his labours, on the conclusion of which he will be congratulated by all
Naturalists. The entire literature of the RADIOLARIA (from 1834 to 1884) is
completely recorded, and the older species (both living and fossil)
redescribed, so that the Report is a complete Monograph, which will be an
invaluable aid to all future Investigators.

CHALLENGER OFFICE, 32 QUEEN STREET,
EDINBURGH, _1st February 1887._

THE

VOYAGE OF H.M.S. CHALLENGER.

#ZOOLOGY.#

Report on the RADIOLARIA collected by H.M.S. Challenger during the Years
1873-76. By ERNST HAECKEL, M.D., Ph.D., Professor of Zoology in the
University of Jena.

PREFACE.

The significance of the Radiolaria in regard to the relations of life in
the ocean has been increased in a most unexpected manner by the discoveries
of the Challenger. Large swarms of these delicate Rhizopoda were found not
only at the surface of the open ocean but also in its different
bathymetrical zones. Thousands of new species make up the wonderful
Radiolarian ooze, which covers large areas of the deep-sea bed, and was
brought up from abysses of from 2000 to 4000 fathoms by the sounding
machine of the Challenger. They open a new world to morphological
investigation.

When ten years ago (in the autumn of 1876) I accepted the enticing
invitation of Sir Wyville Thomson to undertake the investigation of these
microscopic creatures, I hoped to be able to accomplish the task with some
degree of completeness within a period of from three to five years, but the
further my investigations proceeded the more immeasurable seemed the range
of forms, like the boundless firmament of stars. I soon found myself
compelled to decide between making a detailed study of a selection of
special forms or giving as complete a survey as possible of the varied
forms of the whole class; and I decided upon the latter course, having
regard both to the general plan of the Challenger Reports, and to the
interests of our acquaintance with the class as a whole. I must, however,
confess at the close of my work that my original intention is far from
having been fulfilled. The extraordinary extent and varied difficulties of
the undertaking must excuse the many deficiencies.

The special examination of the Challenger collection was for the most part
completed in the summer of 1881; I collected its results in my Entwurf
eines Radiolarien-Systems auf Grund von Studien der Challenger-Radiolarien
(Jenaische Zeitschr. f. Naturw., Bd. xv., 1881). Since the manuscript of
this preliminary communication was completed only a few days before my
departure for Ceylon, and since I was unable to correct the proofs myself,
several errors have crept into the Prodromus Systematis Radiolarium
included in it. These have been corrected in the following more extensive
working out of it. Even at that time I had distinguished 630 genera and
more than 2000 species; but on the revision of these, which I undertook
immediately on my return from India, this number was considerably
increased. The total number of forms here described amounts to 739 genera
and 4318 species; of these 3508 are new, as against 810 previously
described. In spite of this large number, however, and in spite of the
astonishing variety of the new and marvellous forms, the riches of the
Challenger collection are by no means exhausted. A careful and patient
worker who would devote a second decade to the work, would probably
increase the number of new forms (especially of the smaller ones) by more
than a thousand; but for a really complete examination, the lifetime of one
man would not suffice.

The richest source of the Challenger material is the Radiolarian ooze of
the central Pacific Ocean (Stations 265 to 274). This remarkable deep-sea
mud consists for the greater part of well-preserved siliceous shells of
Polycystina (SPUMELLARIA and NASSELLARIA). Not less important, however,
especially for the study of the ACANTHARIA and PHÆODARIA, are the wonderful
preparations stained with carmine and mounted in Canada balsam on the spot
by Dr. John Murray. One such preparation (_e.g._, from Station 271) often
contains twenty or thirty, sometimes even fifty new species. In many of
these preparations the individual parts of the unicellular organism are so
well preserved that they show clearly the characteristic peculiarities of
the legions and orders. Since the material for these preparations was taken
with the tow-net, not only from the surface of the sea but also from
different bathymetrical zones, it furnishes valuable conclusions regarding
the chorology, as well as the physiology and morphology of the group. For
many new discoveries I am indebted to the study of such preparations, of
which I have examined about a thousand from 168 different Stations (compare
§ 240). In addition to these about 100 bottles were handed to me,
containing partly bottom-deposits, partly tow-net gatherings.

Sir Wyville Thomson, who directed the investigations of the Challenger with
so much devotion, and only partly saw its results, has laid me under a deep
debt of obligation; not less is this the case, however, with his successor,
Dr. John Murray. I am especially indebted to both gentlemen for the freedom
they have allowed me in the carrying out of my work, and especially for the
permission to include a description of all known Radiolaria in the
Challenger Report, which has thus become a second edition many times
enlarged of my Monograph published in 1862. Since all previous literature
of the subject has been consulted and critically revised, it is hoped that
this Report will form a useful foundation for future investigations. All
names of sufficiently described Radiolaria published during the first half
century of our knowledge of the class (from 1834 to 1884), are inserted in
alphabetical order in the index at the end of this work.

In addition to the treasures of the Challenger, my own collection of
Radiolaria has yielded many new forms whose description is here included.
On my journeys to the Mediterranean (an account of which is given in the
introduction to my Monograph of the Medusæ), I have given special attention
to these delicate microscopic organisms for more than thirty years. Besides
the various points on the Mediterranean, the Atlantic Ocean at the Canaries
(in the winter of 1866-67) yielded many interesting new forms; whilst my
voyage across the Indian Ocean, from Aden to Bombay, in November 1881,
thence to Ceylon and back by Socotra in March 1882, was still more
productive. In particular, some extended excursions which I had the
opportunity of making from Belligemma and Matura (at the southern extremity
of Ceylon) gave me an insight into the rich treasures of the Indian Ocean.

Most important, however, as regards the knowledge of the Indian Radiolaria,
are the collections which Captain Heinrich Rabbe of Bremen has so
beautifully preserved during his many voyages through that region. In the
neighbourhood of Madagascar and the Cocos Islands more especially, and also
in the Sunda Archipelago, he met with large swarms of Radiolaria, among
which were many new and remarkable forms. These were of special value for
completing the chorology, and the more so since the course of the
Challenger in the Indian Ocean lay very far to the southwards. I will
therefore take this opportunity of repeating my best thanks to Captain
Rabbe for the friendly donation of his valuable collection.

The Radiolarian fauna of the North Atlantic Ocean, which was previously but
little known and only slightly increased by the investigations of the
Challenger, received a valuable increase from the interesting collections
made by Dr. John Murray on various expeditions to the Færöe Islands (on the
"Knight Errant" in 1880 and on the "Triton" in 1882). A large number of new
Radiolaria were captured in the Færöe Channel, partly at the surface of the
Gulf Stream, partly at various depths, and the proof was thus furnished
that at certain points in the North Atlantic Ocean Radiolaria are very
richly developed. I am further indebted to Dr. John Murray for the free use
of this important material as well as for much other assistance in the
carrying out of my work. Another rich source of Radiolaria I found in the
alimentary canal of pelagic animals from all seas. Medusæ, Siphonophoræ,
Salpæ, Pteropoda, Heteropoda, Crustacea, &c., which live partly at the
surface of the sea and partly at various depths, and swallow large masses
of Radiolaria, often contain numbers of their shells well-preserved in
their intestine. The alimentary canal of Fishes and Cephalopods too, which
live upon these pelagic animal frequently contains considerable quantities
of siliceous shells; and another newly discovered source has been found in
the coprolites of the Jurassic period, which consist largely of Radiolarian
skeletons.

In the investigation of this complicated system of organisms, I have
endeavoured on the one hand to give accurately the forms and dimensions of
the species observed, and on the other hand to present a survey of the
relationships of the different genera and families; and in this I have
striven especially to combine the phylogenetic aims of the natural system
with the essentially artificial divisions of a practical classification.
Being, however, a conscientious supporter of the theory of descent, I can
of course lay no stress upon the value of the categories, which are here
distinguished as Legions, Orders, Families, Genera, &c. All these
artificial systematic grades I regard as of merely relative value; and from
the same cause I attach no importance to the distinction of all the species
here described; many of them are probably only developmental stages, and
like my predecessors I have determined their boundaries on subjective
grounds. In the systematic working out of so much material one always runs
the risk of doing either too much or too little in the way of creating
species; but in the light of the theory of descent this danger is of no
consequence.

In the carrying out of this extensive task the friendly aid of Dr. Reinhold
Teuscher of Jena was of the greatest benefit to me; at my request he was at
the trouble of making a large number of accurate drawings with the camera
lucida, and he also undertook a long series, amounting to some 8000,
accurate micrometric measurements, which were of the greatest value in the
attempt to settle the important question of the constancy of the various
species; I have alluded to this in a note at the conclusion of the Report
(p. 1760). My best thanks are due to Dr. Teuscher for the patient and
careful manner in which he discharged these tedious tasks.

The figures of new species of Radiolaria (about 1600 in number) which
appear in the atlas of one hundred and forty plates accompanying this
Report, were nearly all drawn with the camera lucida, partly by Mr. Adolph
Giltsch and partly by myself. The names of the genera which appear at the
bottom of the plates have in many cases been changed since they were
printed off, as may be seen from the explanations which accompany them. Had
it been possible to complete the examination of the material before the
plates were commenced this might have been avoided, and in many cases a
better selection of figures might have been made. All the drawings have
been made upon the stone by the practised hand of Mr. Adolph Giltsch, in
his usual masterly manner, and his lithographic work, which has lasted
fully ten years, is the more valuable since he has himself microscopically
studied the greater part of the species figured. The fact that the atlas
presents so full a picture of the marvellous wealth of form of the
Radiolaria is especially due to his lively interest in the work, to his
unwearying care, and to his morphological acuteness. May it be the means of
inducing many naturalists to study more deeply this inexhaustible kingdom
of microscopic life, whose endless variety of wonderful forms justifies the
saying--_Natura in minimis maxima_.

CONTENTS.

FIRST PART.

GENERAL INTRODUCTION-- PAGE

I. Anatomical Section (§§ 1-140), i
Chapter I. The Unicellular Organism, i
" II. The Central Capsule, xxiv
" III. The Extracapsulum, li
" IV. The Skeleton, lxviii

II. Biogenetical Section (§§ 141-200), xciii
Chapter V. Ontogeny (Individual Development), xciii
" VI. Phylogeny (Genealogical Development), ci

III. Physiological Section (§§ 201-225), cxxviii
Chapter VII. Vegetative Functions, cxxviii
" VIII. Animal Functions, cxl

IV. Chorological Section (§§ 226-250), cxlvi
Chapter IX. Geographical Distribution, cxlvi
" X. Geological Distribution, clxiv

V. Bibliographical Section (§§ 251-254), clxxvi

SYSTEMATIC PART, 1

I. Subclass PORULOSA, 6
Legion I. SPUMELLARIA vel PERIPYLEA, 6
Order 1. Colloidea, 10
" 2. Beloidea, 28
" 3. Sphæroidea, 50
" 4. Prunoidea, 284
" 5. Discoidea, 402
" 6. Larcoidea, 599

Legion II. ACANTHARIA vel ACTIPYLEA, 716
Order 7. Actinelida, 728
" 8. Acanthonida, 740
" 9. Sphærophracta, 795
" 10. Prunophracta, 859

SECOND PART.

II. Subclass OSCULOSA, 889
Legion III. NASSELLARIA vel MONOPYLEA, 889
Order 11. Nassoidea, 895
" 12. Plectoidea, 898
" 13. Stephoidea, 931
" 14. Spyroidea, 1015
" 15. Botryodea, 1103
" 16. Cyrtoidea, 1126

Legion IV. PHÆODARIA vel CANNOPYLEA, 1521
Order 17. Phæocystina, 1542
" 18. Phæosphæria, 1590
" 19. Phæogromia, 1642
" 20. Phæoconchia, 1710

Note on the Dimensions and Measurements, 1760

ADDENDA, 1761

ERRATA, 1763

INDEX, 1765

GENERAL INTRODUCTION.

----

ANATOMICAL SECTION.

A SKETCH OF OUR KNOWLEDGE OF THE ORGANISATION OF THE RADIOLARIA IN THE YEAR
1884.

----

CHAPTER I.--THE UNICELLULAR ORGANISM.

(§§ 1-50.)

1. _Definition of the Radiolaria._--RADIOLARIA are marine Rhizopoda, whose
unicellular body always consists of two main portions, separated by a
membrane; an inner _Central capsule_ (with one or more nuclei) and an
_Extracapsulum_ (the external calymma, which has no nucleus, and the
pseudopodia); the endoplasm of the former and the exoplasm of the latter
are connected by openings in the capsule-membrane. The central capsule is
partly the general central organ of the Radiolarian cell, partly the
special organ of reproduction, since its intracapsular protoplasm, along
with the nuclei embedded in it, serves for the formation of flagellate
spores. The extracapsulum is partly the general organ for intercourse with
the outer world (by means of the pseudopodia), partly the special organ of
protection (calymma) and nutrition (sarcomatrix). The majority of
Radiolaria develop also a skeleton for support and protection, which
presents the utmost variety of form, and is generally composed of silica,
sometimes of an organic substance (acanthin). The Radiolarian cell usually
leads an isolated existence (Monozoa _vel_ Monocyttaria); only in a small
minority (of one legion) are the unicellular organisms united in colonies
or coenobia (Polyzoa _vel_ Polycyttaria).

The extent of the Radiolaria, as limited by the above definition, which I
have made as compact as possible, differs in several important respects
from that allowed to the group by all previous diagnoses. The shortest
expression of its scope might perhaps be:--Rhizopoda with central capsule
and calymma; for the most important character of the Radiolaria, and that
by which they are distinguished from all other Rhizopoda, is the
differentiation of the unicellular body into two principal parts of equal
importance and their separation by a constant capsule-membrane.

2. _The Two Subclasses of the Radiolaria._--The systematic catalogue of the
Radiolaria, which forms the second part of this Report, and is brought up
to the year {ii}1884, contains 20 orders, 85 families, 739 genera, and 4318
species. The consideration that but a small proportion of the ocean his yet
been investigated renders it likely, however, that even this large number
does not include the half of the recent species. The great progress which
our knowledge of the organisation of the Radiolaria has made, by means of
comparative study, renders it possible to arrange this enormous mass of
forms in four main divisions or legions, and these are again related in
pairs, so that two divisions of the highest rank or subclasses are
constituted, the _Porulosa_ (or _Holotrypasta_) and _Osculosa_ (or
_Merotrypasta_).

The division of the Radiolaria into two subclasses and four legions (or
principal orders), I sought to establish in 1883 in a communication on
the Orders of the Radiolaria (Sitzb. Jena Gesellsch. Med. u. Naturwiss.,
February 16, 1883). As a believer in the theory of descent, I regard all
the systematic arrangements of specialists as artificial, and all their
divisions as subjective abstractions, and hence I shall be guided in the
establishment of such groups as subclasses, legions, orders, &c., by
purely practical considerations, especially by the desire to give as
ready a survey as possible of the complex multitude of forms (compare §§
154 to 156).

3. _Porulosa or Holotrypasta._--The subclass Porulosa or Holotrypasta
includes the two legions, PERIPYLEA or SPUMELLARIA, and ACTIPYLEA or
ACANTHARIA, which agree in the following constant and important
characters:--(1) The _Central Capsule_ is primitively a sphere, and retains
this homaxon form in the majority of the species. (2) The _Membrane_ of the
central capsule is everywhere perforated by very numerous minute pores, but
possesses no larger principal aperture (osculum). (3) The _Pseudopodia_
radiate in all directions and in great numbers from the central capsule,
passing through its pores. (4) The _Equilibrium_ of the floating
unicellular body is in most Porulosa pantostatic (indifferent) or
polystatic (plural-stable), since a vertical axis is either absent, or, if
present, has its two poles similarly constituted. (5) The _Ground-forms_ of
the skeleton are therefore almost always either spherotypic or
isopolar-monaxon, very rarely zygotypic. The two legions of the Porulosa
are distinguished mainly by the skeleton of the SPUMELLARIA (or PERIPYLEA)
being siliceous, never centrogenous, nor composed of acanthin, whilst in
the ACANTHARIA (or ACTIPYLEA) it is always centrogenous and made up of
acanthin; hence in the former the nucleus is always central, in the latter
always excentric.

4. _Osculosa or Merotrypasta._--The subclass Osculosa or Merotrypasta
includes the two legions MONOPYLEA or NASSELLARIA, and CANNOPYLEA or
PHÆODARIA, which agree in the following constant and important
characters:--(1) The Central Capsule is originally monaxon (ovoid or
spheroidal) and retains this ground-form in most of the species. (2) The
_Membrane_ of the central capsule possesses a single large principal
aperture (osculum) at the basal pole of the vertical main axis. (3) The
_Pseudopodia_ radiate from a stream of sarcode which passes out from the
central capsule only on one side, namely, through the principal aperture.
(4) The _Equilibrium_ of the floating body is {iii}monostatic or unistable,
since the two poles of the principal axis are always more or less different
from each other. (5) The _Ground-forms_ of the skeleton are, therefore, for
the most part grammotypic (centraxon) or zygotypic (centroplan), rarely
spherotypic. The two legions of the Osculosa are distinguished chiefly by
the principal opening (osculum) being closed by a porous plate (porochora
with its podoconus) in the NASSELLARIA (or MONOPYLEA), and by a radiate
cover (operculum with its astropyle) in the PHÆODARIA (or CANNOPYLEA).

5. _The four Legions of Radiolaria._--The four principal groups of
Radiolaria, to which we have given the name "legions," are natural units,
since the most important peculiarities in the structure of the central
capsule are quite constant within the limits of the same legion, and since
all the forms in the same legion may be traced without violence to the same
phylogenetic stem. The four legions are, however, related to each other, in
so far as they all exhibit those characters which distinguish the
Radiolaria from other Protista. The two which compose the Porulosa (§ 3)
seem somewhat more nearly related to each other than to the two which make
up the Osculosa (§ 4). When, however, the attempt is made to bring them all
into a phylogenetic relationship, it undoubtedly appears that the
SPUMELLARIA (or PERIPYLEA) are the primitive stem, out of which the other
three have been developed as independent branches. All three have been
derived, probably independently, from the most ancient stem-form of the
SPUMELLARIA, the spherical _Actissa_.

6. _Peripylea or Spumellaria._--Those Radiolaria which we call "PERIPYLEA"
on account of the constitution of their central capsule, or "SPUMELLARIA"
on account of the nature of their skeleton, are separated from the other
three legions of the class by the combination of the following constant
characters:--(1) The _Membrane_ of the central capsule is single and evenly
perforated all over by innumerable fine pore-canals, but without any larger
principal opening (osculum). (2) The _Nucleus_ always lies centrally in the
SPUMELLARIA monozoa and is serotinous, for it divides only at a later
period into the nuclei of the spores; in the SPUMELLARIA polyzoa it is
precocious, and divides early into many small nuclei. (3) The _Pseudopodia_
are exceedingly numerous and distributed evenly over the whole surface of
the central capsule. (4) The _Calymma_ contains no phæodium. (5) The
_Skeleton_ is seldom wanting, is never centrogenous, and is always
siliceous. (6) The _Ground-form_ of the central capsule is originally
spherical (often modified); that of the skeleton is also spherical or, in
the majority of cases, derived in different ways from the sphere.

7. _Actipylea or Acantharia._--These Radiolaria which we call "ACTIPYLEA"
on account of the constitution of their central capsule, or "ACANTHARIA"
from the formation of their skeleton, are separated from the other three
legions by the combination of the following constant characters:--(1) The
_Membrane_ of the central capsule is single and {iv}perforated by numerous
fine pore-canals, which are regularly distributed in series or groups, but
without a larger principal opening (osculum). (2) The _Nucleus_ is always
excentric and generally precocious, since it divides early by a peculiar
process of budding into numerous small nuclei. (3) The _Pseudopodia_ are
very numerous and distributed regularly in groups (or series united into a
network). (4) The _Calymma_ contains no phæodium. (5) The _Skeleton_ is
generally present, always centrogenous, and composed of acanthin. (6) The
_Ground-form_ of the central capsule is originally spherical (often
modified), that of the skeleton polyaxon (often modified).

8. _Monopylea or Nassellaria._--Those Radiolaria which we call "MONOPYLEA"
from the formation of their central capsule, or "NASSELLARIA" from the
nature of their skeleton, are distinguished from the other three legions of
the class by the combination of the following constant characters:--(1) The
_Membrane_ of the central capsule is single, and has only one large
principal opening (osculum) at the basal pole of the vertical main axis;
this osculum is closed by a perforated lid (porochora or operculum porosum)
from which there arises within the central capsule a peculiar cone of
threads or pseudopodia (podoconus). (2) The _Nucleus_ is usually excentric
and is always serotinous, since it only divides at a comparatively late
period into spore-nuclei. (3) The _Pseudopodia_ are not very numerous and
arise by division of a single stem or bundle of threads of sarcode, which
issues from the porochora. (4) The _Calymma_ contains no phæodium. (5) The
_Skeleton_ (very rarely absent) is never centrogenous, but always
extracapsular and siliceous. (6) The _Ground-form_ of the central capsule
is always monaxon (with a vertical allopolar main axis), originally ovoid,
often modified; that of the skeleton is also generally monaxon, often
modified (triradial or bilateral).

9. _Cannopylea or Phæodaria._--Those Radiolaria which we call "CANNOPYLEA"
from the constitution of their central capsule, or "PHÆODARIA" on account
of their peculiar phæodium, are distinguished from the other three legions
by the combination of the following characters:--(1) The _Membrane_ of the
central capsule is double, consisting of a strong outer and delicate inner
capsule, and has only one principal opening (osculum) at the basal pole of
the vertical main axis; this osculum is closed by a radiate cover
(astropyle or operculum radiatum), from the centre of which arises an
external tubular spout (proboscis). Occasionally a few small accessory
openings (parapylæ) are present besides the principal opening. (2) The
_Nucleus_ lies centrally or subcentrally in the capsule (in the vertical
main axis), and is serotinous, inasmuch as it only divides at a late period
into spore-nuclei. (3) The _Pseudopodia_ are usually very numerous and
arise from a thick sarcomatrix, formed by the spreading out of a thick stem
of sarcode, which issues from the astropyle. (4) The _Calymma_ always
contains a phæodium or peculiar voluminous excentric mass of pigment. (5)
The _Skeleton_ (very rarely absent) is never centrogenous, always
extracapsular and formed of a silicate of carbon. (6) The {v}_Ground-form_
of the central capsule is always monaxon (with a vertical allopolar main
axis) and generally spheroidal; that of the skeleton is very varied.

10. _Synopsis of the Subclasses and Legions:--_

11. _Individuality of the Radiolaria._--Like other Protozoa the Radiolaria
are unicellular organisms, the whole fully developed organisation of which
falls under the category of a single cell, both morphologically and
physiologically. Since this view is based upon the composition of the
individual body out of two different morphological elements, nucleus and
protoplasm, it is at once justified in the case of the majority of
Radiolaria, in which the plasmatic body encloses only a single nucleus (the
so-called "Binnen-Bläschen"); such is the case in all the SPUMELLARIA
monozoa, NASSELLARIA and PHÆODARIA. This aspect of the case might appear
doubtful in those Radiolaria in which the simple primary cell-nucleus
divides early into numerous small secondary nuclei, as is the case in the
SPUMELLARIA polyzoa and most ACANTHARIA. Strictly speaking, the
multinucleate central capsule should in such cases be regarded as a
syncytium; but since the individual unity of the unicellular organism is as
clearly defined in these precocious multinuclear Radiolaria as in the
ordinary serotinous forms, the former must be considered unicellular
Rhizopods just as are the latter. This mode of regarding {vi}the case is
the more necessary, inasmuch as the early division of the nucleus has no
further influence upon the organisation. Just as in many other classes of
the Protista there are monozootic (solitary) and polyzootic (social) forms,
so also in the Radiolaria there are in addition to the ordinary monozootic
or monobious forms certain families in which colonies or coenobia are
formed by the association of individuals; this distinction may be expressed
by the terms "Monocyttaria" and "Polycyttaria."

The unicellular nature of the Radiolaria was first established by Richard
Hertwig in 1879 (L. N. 33),[1] and brought into conformity with our
present histiological knowledge and the new reform of the cell-theory.
Huxley, however, who was in 1851 the first to examine living Radiolaria
accurately, declared _Thalassicolla nucleata_ to be a unicellular
Protozoon, and the individual central capsules of _Sphærozoum punctatum_
to be cells, but, owing to the then condition of the cell-theory, he was
unable to give a conclusive demonstration of this view. Later, when
Johannes Müller in 1858 and myself in 1862 recognised the peculiar
"yellow cells" which occur in large numbers in many Radiolaria as true
nucleated cells, it appeared impossible any longer to maintain the
unicellular nature of the Radiolaria; also the great complication which I
showed to exist in the structure of _Thalassicolla_ appeared to
contradict it. Only after Cienkowski (1871) and Brandt (1881) had shown
that the "yellow cells" do not belong to the Radiolarian organism, but
are symbiotic unicellular algæ, was it possible to revive and demonstrate
anew the unicellular nature of the Radiolaria.

12. _Morphological Individuality._--From the morphological standpoint the
individuality of the unicellular elementary organism is obvious in the
ordinary solitary Radiolaria (Monobia), and is to be so regarded that the
whole body with all its constituent parts, and not merely the central
capsule, is to be regarded as _a cell_. Naturally the xanthellæ or yellow
cells (§§ 76, 90), which as independent algæ live in symbiosis with many
Radiolaria, must be excluded. The unicellular organisation of the
Radiolaria is further to be distinguished from that of the other Protista,
inasmuch as an internal membrane (capsule-membrane) separates the central
(medullary) from the peripheral (cortical) portion. In the coenobia of the
social Radiolaria (or Polycyttaria), the morphological individuality
persists only as regards the medullary portions of the aggregated cells
(the individual central capsules), while the cortical portions fuse
completely to form a common extracapsulum. Hence in these SPUMELLARIA
polyzoa two different stages of morphological individuality must be
distinguished, the _Cell_ as a _Morphon of the first stage_, and the
_Coenobium_ as a _Morphon of the second stage_.

13. _Physiological Individuality._--From the physiological standpoint also
the individuality of the unicellular organism is immediately obvious in the
case of the ordinary solitary Radiolaria (Monobia); as in other Protista it
fulfils all the functions of life by itself alone. This physiological
individuality of the monobious Radiolarian cell is furthermore not
influenced by the xanthellæ, which live as independent algæ in symbiosis
with many Radiolaria; even though these often by the production of starch
assist in the {vii}nourishment of the Radiolaria, yet they are by no means
indispensable to them. On the other hand, the physiological individuality
offers more complicated relations in the social Radiolaria (Polycyttaria)
which live united in colonies or coenobia. Here the actual _Bion_ (or the
fully developed physiological individual) is not represented by the
individual cells, but by the whole multicellular coenobium, which in each
species has a definite form and size. In these coenobia, which are usually
spherical or cylindrical jelly-like masses, several millimeters in
diameter, numerous cells are so intimately united that only their medullary
portions (the central capsule with the endoplasm) remain independent; the
cortical portions (calymma and exoplasm) on the contrary uniting into a
common extracapsulum. This discharges, as a whole, the functions of
locomotion, sensation, and inception of nutriment, while the separate
central capsules act in the main only as reproductive organs (forming
spores) and partly also as the central organs of metastasis (digestion).
Each coenobium may also be regarded as a polycyttarium, _i.e._, a
"multicellular Radiolarian," whose numerous central capsules represent so
many sporangia or spore-capsules.

On this head compare the section in my monograph of 1862 (L. N. 16),
entitled Die Organisation der Radiolarien-Colonien; _Polyzoen_ oder
_Polycyttarien?_ (pp. 116 to 126); and also R. Hertwig, Zur Histologie
der Radiolarien, 1876 (L. N. 26, p. 23).

14. _Monocyttaria_ and _Polycyttaria_.--In the majority of the Radiolaria
each unicellular organism passes its individual life in an isolated
condition (as a Monocyttarium). Only in a part of the SPUMELLARIA numerous
unicellular individuals are united into societies which are regarded as
coenobia or colonies (Polycyttaria). This is the case in three different
families belonging to the PERIPYLEA, in the Collozoida (without a skeleton,
Pl. 3), the Sphærozoida (with a Beloid skeleton, Pl. 4), and the
Collosphærida (with a Sphæroid skeleton, Pls. 5-8). All three families of
Polycyttaria (or social Radiolaria), agree in their mode of forming
colonies, since the central capsules of the social individuals remain
separate and lie in a common jelly-like mass, which is formed by the fusion
of their extracapsulum. The chief part of the voluminous colonies, which
attain a diameter of several millimetres (sometimes more than 1 cm.), and
are generally spherical, ellipsoidal or cylindrical, consists therefore of
the jelly-like calymma, and this is penetrated by a sarcoplegma, to whose
meshes all the individual organisms contribute by means of the pseudopodia,
which radiate from their sarcomatrix. A further peculiarity in which the
social SPUMELLARIA differ from the solitary consists in the fact that the
former are precocious and the latter serotinous in the division of the
nucleus (§ 64). Whilst in the solitary or monozootic SPUMELLARIA the middle
of the central capsule is occupied by the simple nucleus, and this divides
only at a late period (immediately before the formation of spores) into the
numerous spore nuclei, in the colonial or polyzootic SPUMELLARIA this
division takes place very early, and the middle of each central capsule is
usually occupied by an oil-globule.

{viii}The colonial Radiolaria were described as early as the year 1834 by
Meyen, the first investigator of the class, under the name _Sphærozoum_,
and, as _Palmellaria_, compared with the gelatinous colonies of the
Nostochineæ. The first accurate observations upon their structure were,
however, made in 1851 by Huxley, who described examples of all three
families under the name _Thalassicolla punctata_. More extended,
however, were the investigations of Johannes Müller, who in his
fundamental work (1858) divided the whole class Radiolaria into
_Solitaria_ and _Polyzoa_. The _Radiolaria solitaria_ he divided into
Thalassicolla, Polycystina and Acanthometra, the _Radiolaria polyzoa_
into Sphærozoa (without a shell) and Collosphæra (with a shell). The
most accurate delineation of the Polycyttaria was given by Hertwig in his
beautiful memoir, Zur Histologie der Radiolarien (1876). Quite recently,
however (1886), since the completion of my manuscript upon the Challenger
Radiolaria, a very complete Monograph of the Polycyttaria has appeared by
Karl Brandt, Die colonie-bildenden Radiolarien (Sphærozoen) des Golfes
von Neapel und der angrenzenden Meeres-Abschnitte (276 pp., 8 pls.,
Berlin). It contains in particular most valuable contributions to the
physiology and histology.

15. _The Central Capsule and Extracapsulum._--The special peculiarity of
the unicellular Radiolarian organism, by which it is clearly distinguished
from all other Rhizopoda (and indeed from most other Protista), is its
differentiation into two separate chief constituents, the central capsule
and extracapsulum, and the formation of a special membrane which separates
them. This, the capsule-membrane, is not to be compared with an ordinary
cell-membrane, as an external layer, but rather to be regarded as an
internal differentiated product. The extracapsulum or external (cortical)
portion of the body is in most Radiolaria more voluminous than the central
capsule or inner (medullary) portion. The exoplasm of the former (the
cortical or extracapsular protoplasm) is emphatically different from the
endoplasm of the latter (the medullary or intracapsular protoplasm).
Besides the most important vital processes are distributed by division of
labour so completely between them that they appear most distinctly
co-ordinated. The central capsule is on the one hand the general central
organ of the "cell-soul" for the discharge of its sensory and motor
functions (comparable to a ganglion-cell), on the other hand the special
organ of reproduction (sporangium). The extracapsulum, also, is not less
significant, since on the one hand its calymma acts as a protecting
envelope to the central capsule, as a support to the pseudopodia, and a
foundation for the skeleton or a matrix for the development of the shell,
and on the other hand its pseudopodia are of the utmost importance as
peripheral organs of movement and sensation as well as of nutrition and
respiration. The _central capsule_ and the _extracapsulum_ are therefore to
be regarded both morphologically and physiologically as the two
_characteristic co-ordinated principal parts_ of the unicellular
Radiolarian organism.

In most of the more modern delineations of the Radiolaria the central
capsule is regarded as the "cell proper" and its membrane as the
"cell-wall." The following facts are opposed to the correctness of this
interpretation:--1. In most Radiolaria the exoplasm is clearly different
from {ix}the endoplasm, and the former is more voluminous than the
latter. 2. In all Radiolaria the division of labour is so carried out
between the central capsule and the extracapsulum, that the physiological
significance and independence of both principal parts of the cell is
almost equally great. 3. It is only in the ACANTHARIA that the formation
of the skeleton takes place within the central capsule; in all the other
three legions it is quite independent of it.

16. _The Malacoma and Skeleton._--Whilst the division of the unicellular
organism into central capsule and extracapsulum is undoubtedly the most
important character of the Radiolarian organism, the development of a
skeleton of peculiar and most varied form is of very striking significance.
This skeleton is _always a secondary product of the cell_, but is always
anatomically so independent, and so clearly marked off from the soft parts
or malacoma, that it seems advisable to regard both separately in a general
morphological survey. The skeleton stands in a different relation to each
of the two principal constituents of the malacoma. Only in the ACANTHARIA
is it centrogenous and developed from the central capsule outwards. In the
other three legions the skeleton never arises in the centre of the capsule;
in the NASSELLARIA and PHÆODARIA it is always extracapsular; in the
SPUMELLARIA it is also outside the central capsule originally, but
afterwards becomes often surrounded by it, and finally lies in most cases
partly within and partly without the central capsule. The chemical basis of
the skeleton in the ACANTHARIA is the curious acanthin (an organic
substance allied to chitin), in the PHÆODARIA a silicate of carbon, and in
the NASSELLARIA and SPUMELLARIA silica.

17. _Ground-Forms of the Radiolaria (Promorphology)._--The ground-forms of
the Radiolaria exhibit a greater variety than those of any other class in
the organic world, greater indeed than is to be found in all the remaining
groups together. For every conceivable ground-form which can be defined in
the system of promorphology is actually present in the Radiolaria; their
skeleton exhibits, as it were, in material existence, certain geometrical
ground-forms which are found in no other organisms. The cause of this
unexampled richness in different forms lies chiefly in the static relations
of the Radiolaria, which swim freely in the sea, partly also in the
peculiar plasticity of their protoplasm and the material of their
skeletons.

Regarding the general system of ground-forms compare my Generelle
Morphologie (1866, Bd. i. pp. 375-552; Bd. iv., Allgemeine
Grundformenlehre). The ground-forms there proposed and systematically
defined have, however, found but little acceptance (chiefly, no doubt,
owing to the difficult and complicated nomenclature); but having now,
twenty years after their publication, anew carefully revised and
critically studied them, I can find no sufficient reason for abandoning
the principles there adopted. On the contrary the study of the Challenger
Radiolaria during the last ten years, with its incomparable wealth of
forms, has only confirmed the accuracy of my system of ground-forms. The
customary treatment of these in zoological and botanical handbooks (such
as those of Claus and Sachs) is quite insufficient.

{x}18. _The Principal Groups of Geometrical Ground-Forms._--The great
variety of the geometrical ground-forms which are actually realised in the
variously shaped bodies of the Radiolaria, renders it desirable to classify
these in as small a number as possible of principal groups and a larger
number of subdivisions. As extensive principal groups four at least must be
distinguished; the _Centrostigma_ or Sphærotypic, the _Centraxonia_ or
Grammotypic, the _Centroplana_ or Zygotypic, and the _Acentrica_ or Atypic.
The natural centre of the body, about which all its parts are regularly
arranged, is in the first group a point (stigma), in the second a straight
line (principal axis), in the third a plane (sagittal plane), in the fourth
a centre is of course wanting.

19. _The Centrostigma or Sphærotypic Ground-Forms._--The first group of
geometrical ground-forms, here distinguished as sphærotypic or the
centrostigma, is undoubtedly the most important among the Radiolaria,
inasmuch as if these be considered monophyletic, it must be the original
one from which all the other ground-forms have been derived. The common
character of all these sphærotypic ground-forms is that their natural
centre is a point (stigma); thus there is no single principal axis (or
protaxon) such as is characteristic of the two following groups. The
sphærotypic ground-forms are subdivided into two important smaller groups,
the _spheres_ (Homaxonia) and the _endospherical polyhedra_ (Polyaxonia).
The spherical ground-forms, fully developed in the central capsule and
calymma of _Actissa_ and the #Sphæroidea# as well as in many ACANTHARIA,
present no different axes; all possible axes passing through the centre of
the body are equal (Homaxonia). In the endospherical polyhedra, on the
contrary, numerous axes (three at least) may be distinguished, which are
precisely equal to each other and different from all the remaining axes
(Polyaxonia). If the extremities of these axes, or the poles, which are all
equidistant from the common centre, be united by straight lines, a
polyhedral figure is produced whose angles all lie in the surface of the
sphere. According as the poles of the axes are at equal, subequal, or at
different distances from each other, we may divide the endospherical
polyhedra into regular, subregular and irregular. (See Gener. Morphol., Bd.
i. pp. 404-416.)

20. _The Centraxonia or Grammotypic Ground-Forms._--The second principal
group of organic ground-forms, here called grammotypic or centraxonia, is
characterised by the fact that a straight line (gramma) or a single
principal axis (protaxon) forms the natural centre of the body. This
important and extensive group is divided into two subgroups, those with one
axis (Monaxonia) and those with crossed axes (Stauraxonia); in the latter
different secondary transverse or cross-axes may be distinguished, but not
in the former. In the Monaxonia, therefore, every transverse section of the
body perpendicular to the principal axis is a circle, in the Stauraxonia,
on the contrary, a polygon. The Monaxonia are further subdivided into two
groups, in one of which the two poles of the principal axis {xi}are equal
and similar (Isopolar), in the other of which they are different
(Allopolar); in the former the two halves of the body, which are separated
by the equatorial plane (or the largest transverse plane, perpendicular to
the principal axis), are equal, in the latter unequal. Among the isopolar
uniaxial ground-forms (Monaxonia isopola) may be mentioned the ellipsoidal,
spheroidal, lenticular, &c.; to the allopolar uniaxial forms (Monaxonia
allopola) belong the conical, hemispherical, ovoid, &c. In the same way the
pyramidal ground-forms with crossed axes are divisible into two groups,
according as the two poles of the principal axis are equal or not. The
ground-form of the former is the double pyramid, that of the latter the
single pyramid. Both the double and the single pyramids may again be
subdivided, each into two important lesser groups, the regular and the
amphithect. In the first division the equatorial plane of the double and
the basal plane of the single pyramid is a regular polygon (square, &c.),
whilst in the other division it is an elongated or amphithect polygon
(rhombus, &c.); the crossed axes are equal in the former, unequal in the
latter. (See Gener. Morphol., Bd. i. pp. 416-494.)

21. _The Centroplana or Zygotypic Ground-Forms._--The third principal group
of ground-forms includes those which are bilaterally symmetrical in the
ordinary sense, or zeugitic or zygotypic; the natural centre of their body
is a plane. These forms are the only ones in which the distinction between
right and left is possible, since their body is divided by the median plane
(planum sagittale) into two symmetrical halves (right and left). In all
these zeugites the position of every part is determined by three axes
perpendicular to each other, and of these three dimensive axes two are
allopolar, one is isopolar. The two unlike poles of the principal (or
longitudinal) axis are the oral and aboral, the two unlike poles of the
sagittal (or vertical) axis are the dorsal and ventral; the two similar
poles of the frontal (or transverse) axis, however, are the right and left.
This important group of zeugitic or bilateral forms may also be divided
into two clearly distinct lesser groups, the _Amphipleura_ and the
_Zygopleura_. In the Amphipleura (or bilaterally radial ground-forms) the
"radial two-sided" body is produced by modification of a regular pyramid
(as _Spatangus_ from _Echinus_), and hence is composed of several (not less
than three) antimeres. In the Zygopleura (or bilaterally symmetrical
ground-forms) on the other hand, the bodies consist of two antimeres (as in
all the higher animals, Vertebrata, Arthropoda, &c.). (See Gener. Morphol.,
Bd. i. pp. 495-527.)

22. _The Acentrica or Atypic Ground-Forms._--Among the acentrica or
anaxonia are included all those ground-forms which are absolutely
irregular, and in which neither a definite centre nor constant axes can be
distinguished (_e.g._, most Sponges). These quite irregular ground-forms
are very rare among the Radiolaria, but nevertheless there may be referred
to them the amoeboid central capsule of some #Colloidea# (_Collodastrum_,
p. 27, Pl. 3, figs. 4, 5) among the SPUMELLARIA, the irregular shells of
many Collosphærida {xii}(Pl. 8, fig. 2), and the absolutely irregular
shells of the Phorticida and Soreumida among the #Larcoidea#. (See Gener.
Morphol., Bd. i. p. 400.)

23. _The Subsidiary Groups of Geometrical Ground-Forms._--The four natural
principal groups of ground-forms, which have just been defined according to
the nature of the centre of their bodies, may be divided again into
numerous subsidiary groups, defined by the relations of the constant axes
and the two poles of each axis, as well as by the number of the axes and
the differentiation of the secondary with respect to the principal axis.
The most important of these subsidiary groups into which the principal ones
are immediately divided are the following:--(1) The _Centrostigma_ (or
sphærotypic) are divided into spheres (Homaxonia) and endospherical
polyhedra (Polyaxonia). (2) The _Centraxonia_ (or grammotypic) into
uniaxial (Monaxonia) and those with crossed axes (Stauraxonia); among the
former of these may be distinguished the isopolar (phacotypic) and the
allopolar (conotypic); among the latter the double and single pyramids. (3)
The _Centroplana_ (or bilaterals) are divided into amphipleura (or
bilaterally radial) and zygopleura (or bilaterally symmetrical). (4) The
_Acentrica_ (or Anaxonia) or absolutely irregular ground-forms, present no
special subdivisions.

For a complete system of the geometrical ground-forms and their relation
to promorphological classification, see Gener. Morphol., Bd. i. pp.
555-558.

24. _The Spherical or Homaxon Ground-Form._--The spherical is the only
absolutely regular ground-form, since only in it are all axes which pass
through the centre equal; it is very often realised among the Radiolaria,
especially in the SPUMELLARIA and ACANTHARIA, where it furnishes the common
original ground-form, but it is often to be seen in the shells of many
PHÆODARIA (in most #Phæosphæria#); on the other hand, it is never found
among the NASSELLARIA. Geometrical spheres, in the strict sense of the
term, are only to be found among the SPUMELLARIA and ACANTHARIA, namely, in
the central capsule of many #Collodaria# (Pls. 1, 2) and all #Sphæroidea#
(Pls. 11-30) as well as many Acanthometra and Acanthophracta (Pls.
128-138). Nevertheless, speaking generally, one includes those central
capsules and skeletons which have been distinguished here as endospherical
polyhedra. (On these ground-forms see Gener. Morphol., Bd. i. pp. 404-406.)

25. _The Endospherical Polyhedral Ground-Form._--The endospherical
polyhedron or polyaxon ground-form naturally follows the spherical or
homaxon. Under it are included all polyhedra whose angles fall in the
surface of a sphere; this ground-form is especially common among the
SPUMELLARIA, especially in the shells of #Sphæroidea#, but is also found
among the ACANTHARIA (especially in the Astrolophida and #Sphærophracta#),
as well as among the #Phæosphæria# (in most genera of the Orosphærida,
Sagosphærida, and Aulosphærida). Strictly speaking, all those
lattice-shells which have {xiii}been incorrectly called "spherical" belong
to this category, for they are none of them true spheres in the geometrical
sense (like the central capsules of the #Sphæroidea#), but rather
endospherical polyhedra, whose angles are indicated by the nodal points of
the lattice shell, or the radial spines which spring from them. These
endospherical polyhedra may be divided into three groups, the regular,
subregular, and irregular. Of _regular polyhedra_, properly so-called, it
may be shown geometrically that only five can exist, namely, the regular
tetrahedron, cube, octahedron, dodecahedron, and icosahedron. All these are
actually manifested among the Radiolaria, although but seldom. Much more
common are the _subregular endospherical polyhedra_, _e.g._, spherical
lattice-shells with regular hexagonal meshes of equal size; they are never
exactly equal nor perfectly regular, but the divergences are so
insignificant that they escape superficial observation (Pl. 20, figs. 3, 4;
Pl. 26, figs. 1-3). On the contrary in the _irregular endospherical
polyhedra_ the meshes of the lattice-sphere are more or less different in
size and often in form also (Pl. 28, figs. 4, 8; Pl. 30, figs. 4, 6). The
five truly regular polyhedra require separate notice on account of their
importance. (See Gener. Morphol., Bd. i. p. 406.)

26. _The Regular Icosahedral Ground-Form._--The ground-form whose
geometrical type is the regular icosahedron (bounded by twenty equilateral
triangles) is rarely exemplified, but it occurs among the PHÆODARIA in the
Circoporid genus _Circogonia_ (Pl. 117, fig. 1), and also in certain
Aulosphærida, but, apparently, only as an accidental variation (_e.g._,
_Aulosphæra icosahedra_). Furthermore, this ground-form may also be assumed
to occur in those #Sphæroidea# whose spherical lattice-shells bear twelve
equidistant radial spines (_e.g._, many species of _Acanthosphæra_,
_Heliosphæra_, and other Astrosphærida); the basal points of these spines
indicate the twelve angles of the regular icosahedron. (See on this head
Gener. Morphol., Bd. i. p. 411.)

27. _The Regular Dodecahedral Ground-Form._--The ground-form whose
geometrical type is the regular dodecahedron (or pentagonal dodecahedron),
bounded by twelve equilateral and equiangular pentagons, is very rarely
found perfectly developed, as in _Circorrhegma dodecahedra_ (Pl. 117, fig.
2). This form is by no means so common among the Radiolaria as in the
pollen grains of plants (_e.g._, _Buchholzia maritima_, _Fumaria spicata_,
_Polygonum amphibium_, &c.). It can, however, be regarded as present in all
those #Sphæroidea# whose spherical lattice-shells bear twenty equal and
equidistant radial spines (_e.g._, many species of _Acanthosphæra_,
_Heliosphæra_, and other Astrosphærida); the basal points of these spines
mark out the twenty angles of the regular pentagonal dodecahedron. (See
Gener. Morphol., Bd. i. p. 412.)

28. _The Regular Octahedral Ground-Form._--The ground-form whose
geometrical type is the regular octahedron (bounded by eight equilateral
triangles), commonly appears among the SPUMELLARIA in the family
Cubosphærida (p. 169, Pls. 21-25). In {xiv}these #Sphæroidea# the typical
ground-form is usually indicated by six equal radial spines, which are
opposed to each other in pairs, and lie in three similar axes perpendicular
to each other; these are the three axes of the tesseral crystallographic
system; one of them is vertical, whilst the other two cross each other at
right angles in its centre. Occasionally, too, the spherical form of the
lattice-shell passes over into that of the regular octahedron (Pl. 22,
figs. 8, 10). The same form recurs in _Circoporus_ (Pl. 117, fig. 6) among
the PHÆODARIA. In the vegetable kingdom it is exhibited by the antheridia
of _Chara_. It is not found in the NASSELLARIA and ACANTHARIA. (See Gener.
Morphol., Bd. i. p. 412.)

29. _The Regular Cubic Ground-Form._--The ground-form whose geometrical
type is that of a die or cube, is actually presented in a very striking
manner by various Radiolaria. Among the SPUMELLARIA it occurs in certain
#Sphæroidea#, _e.g._, in the Astrosphærid genera _Centrocubus_ and
_Octodendron_ (Pl. 18, figs. 1-3); in these the central medullary shell is
a complete cube, bounded by six equal squares, from the eight angles of
which eight equal radial spines project. This form can also be regarded as
present in those #Sphæroidea# whose spherical lattice-shell bears eight
equal and equidistant radial spines (many Astrosphærida). Besides these the
cubic ground-form is to be seen in certain NASSELLARIA of the family
Tympanida, especially in _Lithocubus_ (Pl. 82, fig. 12; Pl. 94, fig. 13),
in many species of _Acrocubus_, _Microcubus_, &c.; the twelve bars of its
lattice-skeleton correspond often exactly to the edges of the cube. (See
Gener. Morphol., Bd. i. p. 413.)

30. _The Regular Tetrahedral Ground-Form._--The ground-form whose
geometrical type is the regular tetrahedron, bounded by four equilateral
triangles, occurs less frequently in the Radiolaria than the other four
regular polyhedra. Among the SPUMELLARIA it is found in the #Beloidea#, and
especially in those members of the Thalassosphærida and Sphærozoida whose
spicules bear four equal branches, diverging at equal angles from a common
centre. Precisely the same structure is seen also among the NASSELLARIA in
some #Plectoidea#, as in _Tetraplagia_ among the Plagonida, and
_Tetraplecta_ among the Plectanida. The skeleton of both these genera
consists of four equal rods, which radiate at equal angles from a common
centre, just as do the axes of the regular tetrahedron. The tetrahedral
form of these #Plectoidea# is the more important and interesting since on
the one hand it is related to the similar spicular form of the #Beloidea#,
and on the other perhaps furnishes the starting point from which _Cortina_
among the NASSELLARIA may be derived (_Plagoniscus_, _Plectaniscus_). (See
Gener. Morphol., Bd. i. p. 415.)

31. _The Isopolar-Monaxon or Phacotypic Ground-Form._--The isopolar
uniaxial or phacotypic ground-form is characterised by the possession of a
vertical main axis with {xv}equal poles, whilst no transverse axes are
differentiated. All horizontal planes which cut the axis at right angles
are circles, and increase in size from the poles towards the equator. The
most important ground-forms of this group are the _phacoid_ (the lens or
oblate spheroid) and the _ellipsoid_ (or prolate spheroid). Phacoids (or
geometrical lenses with blunt margins) are very often presented by the
central capsules of the #Discoidea# and of many ACANTHARIA (Quadrilonchida
and Hexalaspida), but the lattice-shells of many SPUMELLARIA and ACANTHARIA
exhibit the same form, as also do a few PHÆODARIA (_e.g._, _Aulophacus_).
True geometrical ellipsoids are seen in the central capsules of many
#Prunoidea# among the SPUMELLARIA, and of many Amphilonchida and
Belonaspida among the ACANTHARIA. Furthermore, the lattice shells of many
species of these groups retain the same essential form, _e.g._, many
Ellipsida, Druppulida, and Spongurida (Pls. 13-17, and 39), as well as most
Belonaspida. (See Gener. Morphol., Bd. i. p. 422.)

32. _Allopolar-Monaxon or Conotypic Ground-Form._--The allopolar uniaxial
or conotypic ground-form is characterised by the possession of a vertical
main axis whose two poles are unlike, while no transverse axes are
differentiated. All horizontal planes cutting the main axis at right
angles are circles, and decrease more rapidly from the largest plane
towards the basal than towards the apical pole. The most important
ground-forms of this group are the ovoid, the cone, and the hemisphere.
They often occur (and in geometrical perfection) in the egg-shaped central
capsule and podoconus of the NASSELLARIA, as well as in the shells of
several groups of this legion, particularly in the Cyrtocalpida or
Monocyrtida eradiata (Pl. 51, figs. 10-13), and in many Stichocyrtida
eradiata; furthermore, they are also seen among the PHÆODARIA, _e.g._,
certain Challengerida (Pl. 99, figs. 19-22). (See Gener. Morphol., Bd. i.
p. 426.)

33. _The Regular Dipyramidal or Quadrilonchial Ground-Form._--The
ground-forms whose geometrical type is the regular double pyramid are
characterised by a vertical main axis which possesses equal poles, and
which is crossed at its centre by several equal transverse axes. The
horizontal equatorial plane is therefore a regular polygon, and divides the
body into two equal regular pyramids. The simplest and commonest form of
this group is the quadratic octahedron, the ground-form of the quadratic
crystallographic system; its equatorial plane is a square. This regular
dipyramidal ground-form occurs among the SPUMELLARIA in the shells of the
Staurosphærida as well as of many #Discoidea#, in which several equidistant
radial spines or arms lie in the quadratic equatorial plane of the body,
and project from the margin of the lenticular disc (_e.g._, _Sethostaurus_,
Pl. 31; _Histiastrum_, Pl. 46, &c.). It is, however, among the ACANTHARIA
that the most important part is played by this ground-form (and especially
by the quadratic octahedron); it forms the basis of all those
#Acanthometra# and #Acanthophracta# in which twenty radial spines are
disposed according to the Müllerian Law, and in which {xvi}the four
equatorial spines are of equal dimensions (Icosacantha). (See Gener.
Morphol., Bd. i. p. 436-446.)

34. _The Amphithect Dipyramidal or Lentelliptical Ground-Forms._--The
ground-forms whose geometrical type is the lenticular or "triaxial"
ellipsoid, may also be designated amphithect double pyramids; they are
characterised by the possession of a vertical main axis which has similar
poles, and is crossed at its middle by two transverse axes, unequal but
isopolar. The horizontal equatorial plane of the body is therefore an
amphithect or elongated polygon (a rhombus in the simplest case possible),
and divides the whole body into two equal amphithect pyramids. The simplest
and commonest form of this group is the rhombic octahedron, which is also
the ground-form of the rhombic crystallographic system. It plays an
important part in those ACANTHARIA in which twenty radial spines are
disposed according to the Müllerian Law, but in which the two pairs of
equatorial spines are unequal (different geotomical and hydrotomical axes,
see p. 719); to this category belong the Amphilonchida (Pl. 132),
Belonaspida (Pl. 136), Hexalaspida (Pl. 139), and Diploconida (Pl. 140). A
form essentially identical obtains also among the SPUMELLARIA in the
majority of the #Larcoidea#, both in their triaxial lattice-shells, and in
their lentelliptical central capsules, which present geometrically accurate
triaxial ellipsoids, with three unequal isopolar axes at right angles to
each other. (See Gener. Morphol., Bd. i. p. 446-452.)

35. _The Regular Pyramidal Ground-Forms._--The ground-forms whose
geometrical type is the regular pyramid, and which are the most conspicuous
in the Medusæ, Polyps, Corals, and regular Echinoderms (the Radiata of
earlier authors), are almost confined among the Radiolaria to the legion
NASSELLARIA; they occur, however, in the great majority of these, and
especially in those families which may be classed together as "#Cyrtoidea#
triradiata et multiradiata." Strictly speaking, however, almost all these
NASSELLARIA, at all events in their origin, are bilateral or dipleuric,
since the primary sagittal ring with its characteristic apophyses marks out
the sagittal median plane, and further, since the three feet of the basal
tripod are usually divided into an unpaired dorsal (pes caudalis) and two
paired ventral or lateral (pedes pectorales, dexter et sinister). On the
other hand, it is noteworthy, firstly, that among the primitive
#Plectoidea# there are perfectly regular radial forms, without any
indication of an original bilateral symmetry, and secondly, that similar
forms are also very common among the #Cyrtoidea#, probably as secondary
radial forms, developed from primitive bilateral ones. Similar cases also
occur in certain PHÆODARIA (_e.g._, the Medusettida and Tuscarorida, Pls.
100, 120), but they are entirely wanting among the ACANTHARIA and
SPUMELLARIA. The multiradial NASSELLARIA have arisen from the triradial by
the interpolation of three, six, nine, or more interradial and adradial
secondary apophyses between the three primary perradial ones. (See Gener.
Morphol., Bd. i. pp. 459-874.)

{xvii}36. _The Amphithect Pyramidal Ground-Forms._--The ground-forms whose
geometrical type is the amphithect pyramid, are distinguished from the
regular pyramidal forms, just discussed, chiefly by the form of the basal
plane, which is not a regular, but an amphithect or elongated polygon (in
the simplest case a rhombus). Hence in this case the allopolar main axis of
the body is crossed by two transverse axes which are isopolar and at right
angles, but are unequal; they cannot, however be distinguished as sagittal
and frontal axes as is the case in the zeugites. In the animal as well as
in the vegetable kingdom, an important part is played by this ground-form,
_e.g._, in the Ctenophora, where it is the rhombic pyramid. Among the
Radiolaria it is not common, though it is clearly expressed among the
NASSELLARIA in a number of #Stephoidea# (Stephanida and Tympanida), as well
as in many #Spyroidea# (_e.g._, the bipedal Zygospirida). It is very
accurately developed among the PHÆODARIA in the bivalved #Phæoconchia#
(Pls. 121-128), where the two valves of the shell (dorsal and ventral) are
generally exactly alike, their median keels corresponding to the poles of
the sagittal axis. In the slit between the two valves lie the two secondary
openings (right and left) of the tripylean central capsule, corresponding
to the two poles of the frontal axis, and the main axis stands
perpendicularly to both these, its oral pole being indicated by the
astropyle, or principal aperture. (See Gener. Morphol., Bd. i. pp.
479-494.)

37. _The Amphipleural Ground-Forms._--By the term amphipleural ground-forms
are to be understood those usually defined as "bilaterally radial"; their
geometrical type is a half amphithect pyramid. The best known examples of
this form in the animal kingdom are the bilateral five-rayed Echinoderms
(_Spatangus_, _Clypeaster_), in the vegetable kingdom the symmetrical
five-rayed flowers (_Viola_, _Trifolium_). The three dimensive axes have
the same relation as in the zygopleura, to be next discussed, and which
also resemble them in being divisible only by one plane (the sagittal
median plane) into two equal halves. They differ, however, the amphipleural
body not being made up of two antimeres, but of at least three pairs of
antimeres (or three parameres), being therefore primitively radial. Hence
each of the symmetrical halves of the body contains more than one antimere.
Among the Radiolaria this form does not occur in the SPUMELLARIA,
ACANTHARIA, or PHÆODARIA; it is very common, however, among the
NASSELLARIA; many #Cyrtoidea# multiradiata and #Spyroidea# multiradiata
show this bilaterally radial ground-form, inasmuch as the body consists of
two symmetrical halves, and is also composed of numerous (usually three,
six, nine, or more) radial parameres. In the multiradiate Dicyrtida and
Tricyrtida the cephalis (the first joint) is usually bilateral, whilst the
thorax (the second joint) is multiradial. (See Gener. Morphol., Bd. i. pp.
495-506.)

38. _The Zygopleural Ground-Forms._--As zygopleural or dipleural
ground-forms, as opposed to the amphipleural, are classed those zeugites or
centroplana which are known {xviii}as "bilaterally symmetrical" in the
strictest sense of the term. This is the most important ground-form in the
animal kingdom, inasmuch as it obtains almost exclusively among the higher
animals (Vertebrata, Articulata, Mollusca, Vermes). The body consists of
only two antimeres, which correspond to the two symmetrical halves of the
body. Of the three dimensive axes two are allopolar, one isopolar; the oral
pole of the longitudinal main axis is different from the aboral; the dorsal
pole of the sagittal axis is different from the ventral; but the right pole
of the frontal axis is equal to the left. The right antimere is usually
precisely similar to the left (Eudipleura), more rarely it is slightly
dissimilar or asymmetrical (Dysdipleura). Among the Radiolaria this
ground-form is entirely wanting in the Porulosa or Holotrypasta
(SPUMELLARIA and ACANTHARIA), but on the contrary it is very common in the
Osculosa or Merotrypasta (NASSELLARIA and PHÆODARIA). In the NASSELLARIA it
is of special importance, for the typical _Cortina_ (the combination of the
primary sagittal ring with the basal tripod) exhibits the zygopleural
ground-form clearly sketched out; indeed it is usually clearly seen even in
the sagittal ring itself, for its ventral segment is more strongly curved
than the dorsal; its basal (or oral) pole is always different from the
apical (or aboral). Of the three feet of the basal tripod the unpaired
(caudal) one is directed dorsally and backwards, the two paired (pectoral)
ones ventrally and forwards. The majority of the NASSELLARIA may be
regarded as modifications of this original ground-form. Its relation to the
primitively triradiate tripod presents a still unsolved problem, and the
numerous relations of the zygopleural to the multiradiate ground-forms in
the NASSELLARIA are exceedingly complicated. The zygopleural ground-form is
less widely distributed among the PHÆODARIA, though it is very
characteristically developed in the rich and varied group of Challengerida
(Pl. 99). (See Gener. Morphol., Bd. i. pp. 507-527.)

39. _Synopsis of the Geometrical Ground-Forms:--_

Principal Groups of Ground-Forms.
Subsidiary Groups of Ground-Forms.
Geometrical Type.
Examples.

_Tetraplagia_, _Tetraplecta_, &c.

II. CENTRAXONIA. {xix}
The geometrical centre of the body is a straight line (the vertical main
axis).

Constant transverse axes (perpendicular to the main axis) are wanting in
the Monaxonia (which have circular transverse sections); on the contrary
they are differentiated in the Stauraxonia (which have polygonal
transverse sections).
|
+-III. MONAXONIA.
| Uniaxial ground-forms or centraxonia without transverse axes. The
| transverse planes (perpendicular to the main axis) are circles.
| |
| +-8. _Monaxonia isopola._
| | (Spheroids and ellipsoids; both poles of the main axis similar.)
| |
| | Central capsule and lattice-shell of of many #Discoidea# (lenses) and
| | #Prunoidea# (ellipsoids), Belonaspida, &c.
| |
| +-9. _Monaxonia allopola._
| (Cone, ovoid and hemisphere; the two poles of the axis dissimilar.)
|
| Central capsule and lattice-shell of many NASSELLARIA, especially
| the #Cyrtoidea# eradiata (Cyrtocalpida, &c.).
|
+-IV. STAURAXONIA.
Pyramidal ground-forms or centraxonia with transverse axes. The
transverse planes (perpendicular to the main axis) are either regular
or amphithect polygons.
|
+-10. _Dipyramides regulares._
| (Quadratic octahedron, or quadrilonchial forms and regular double
| pyramids.)
|
| ACANTHARIA with twenty radial spines, the four equatorial being
| equal. Multiradial #Discoidea# and Staurosphærida.
|
+-11. _Dipyramides amphithectæ._
| (Rhombic octahedron, lentellipsoid, and amphithect double pyramids.)
|
| ACANTHARIA with twenty radial spines, whose four equatorial spines
| are unequal but paired. Many #Larcoidea#.
|
+-12. _Pyramides regulares._
| (Regular pyramids.)
|
| Many NASSELLARIA (triradial and multiradial). Medusettida and
| Tuscarorida.
|
+-13. _Pyramides amphithectæ._
(Rhombic pyramids.)

#Phæoconchia#. Bipedal #Spyroidea# and #Stephoidea#.

III. CENTROPLANA.
The geometrical centre of the body is a plane (the sagittal plane).
Constant transverse axes (perpendicular to the main axis) are wanting in
the Monaxonia (which have circular transverse sections); on the contrary
they are differentiated in the Stauraxonia (which have polygonal
transverse sections).
|
+-V. BILATERALIA (or ZEUGITA).
Bilateral forms in the general sense, with right and left halves.
|
+-14. _Amphipleura_
| (Bilaterally radial ground-form.)
|
| Many #Cyrtoidea# and #Spyroidea# multiradiata.
|
+-15. _Zygopleura._
(Bilaterally symmetrical ground-form.)

Most NASSELLARIA (primitively at least), many Challengerida.

IV. ACENTRA.
There is no geometrical centre.
|
+-VI. ANAXONIA.
No definite axes can be determined.
|
+-16. _Irregularia._
(Absolutely irregular ground-forms.)

_Collodastrum_, _Collosphæra_, Phorticida, Soreumida.

40. _Mechanical Causes of the Geometrical Ground-Forms._--The great variety
of ground-forms exhibited by the Radiolaria is of special interest, since
in most instances their causes admit of recognition, and since they are so
intimately related to each other that even in the remaining cases the
assumption that they have arisen by purely mechanical _causæ efficientes_
seems justified. In this respect the first rank is taken by statical
conditions, especially the indifferent or stable equilibrium of the whole
organism, which floats freely in the water. With regard to these
fundamental statical relations, three principal groups of ground-forms may
be distinguished, pantostatic, polystatic, and monostatic.

41. _Pantostatic Ground-Forms._--By pantostatic or indifferently stable
ground-forms are meant those in which the centre of gravity coincides with
the centre of the body, so that they are in equilibrium in any given
position. Strictly speaking, the only form which possesses perfectly
indifferent equilibrium is the sphere, that being the only truly homaxon
and perfectly regular form. Nevertheless, in a somewhat wider sense many
Polyaxonia, especially the endospherical polyhedra with very numerous
sides, may be {xx}included in this category. Such indifferently stable
bodies are found among the SPUMELLARIA in many #Collodaria# and
#Sphæroidea#, as well as in the Astrolophida among the ACANTHARIA. On the
contrary they are entirely wanting among the NASSELLARIA and PHÆODARIA,
since their central capsule constantly presents a main axis with a
differentiated basal pole, and determines the position of stable
equilibrium.

42. _Polystatic Ground-Forms._--Those ground forms are defined as
polystatic or multistable in which the body is in equilibrium in several
different positions (though not in an infinite number). The number of these
positions is usually twice as many as that of the constant equal isopolar
axes exhibited by the form. Hence the regular polyhedra have as many
positions of equilibrium as they have angles or sides, the icosahedron
twenty, dodecahedron twelve, octahedron eight, cube six, tetrahedron four.
The isopolar monaxon ground-forms (lens, ellipsoid, cylinder) and the
diplopyramidal ground forms (quadrilonchial and lentelliptical) have two
positions of stable equilibrium, since the two poles of the vertical axis
are equal and similar and the body is divided into equal halves by the
equatorial plane. This is the case in many SPUMELLARIA (especially
#Discoidea#, #Prunoidea#, and #Larcoidea#), as well as in the great
majority of ACANTHARIA. Perhaps the same holds good also in certain
NASSELLARIA (_e.g._, isopolar Tympanida) and PHÆODARIA (_e.g._, isopolar
#Phæosphæria#), though here unistable equilibrium appears to be
necessitated by the constant main axis of the central capsule and the
differentiated basal pole of the main axis.

43. _Monostatic Ground-Forms._--Those ground-forms are classed as
monostatic or unistable in which the body is in equilibrium only in one
position, since the centre of gravity of the body lies in a constant
vertical axis below its centre. This fixed position is only rarely and
exceptionally found among the SPUMELLARIA (_e.g._, in _Xiphostylus_,
_Sphærostylus_, _Lithomespilus_, _Lithapium_) and among the ACANTHARIA
(_e.g._, in _Zygostaurus_ and _Amphibelone_). On the contrary it is quite
usual among the NASSELLARIA and PHÆODARIA (with but few exceptions); for
here a vertical main axis, with a differentiated basal pole, is determined
even by the formation of the central capsule, and usually also by the
corresponding structure of the skeleton. Among the NASSELLARIA this basal
pole, with the porochora of the central capsule, appears always to be the
lower; as also in most #Phæogromia# among the PHÆODARIA. In the peculiar
bivalved #Phæoconchia#, on the other hand, the basal pole with the
cannopyle is directed upwards; as also in the Challengerida and
Tuscarorida. The #Phæosphæria# and #Phæocystina# are probably to a large
extent polystatic. In general unistable equilibrium may be assumed in the
following categories of ground-forms:--(1) Allopolar monaxon (conical and
ovoid); (2) pyramidal (regular and amphithect); (3) Centroplana
(amphipleura and zygopleura); (4) Anaxonia.

{xxi}44. _Principal Axes._--From the foregoing consideration of the
statical conditions and their direct causal connection with the geometrical
ground-forms of the Radiolaria, the great mechanical significance of the
differentiation of definite axes in these unicellular free-swimming
organisms will be manifest. The most important of these is the primary main
axis (axis principalis, or protaxon), which in all cases has a vertical
direction. It is wanting in the Centrostigma (spheres and endospherical
polyhedra), and in the Anaxonia (acentra). It is isopolar in the phacotypic
forms (Monaxonia isopola), and in the double pyramids (Stauraxonia
isopola). It is allopolar in all monastatic ground-forms, in the conotypic
forms (Monaxonia allopola), pyramids (Stauraxonia allopola), and the
Centroplana (or bilateral forms).

45. _Secondary or Transverse Axes._--In contrast to the vertical main axis
all the other constant axes differentiated in the body may be called
"secondary axes," or "transverse axes," since they cross the former at
definite points. All ground-forms whose vertical axis is crossed by a fixed
number of such axes at definite angles may be called "Stauraxonia." They
are divided into two groups, double pyramids and single pyramids; in the
former the two poles of the main axis (or the two halves of the body
separated by the equatorial plane) are similar (Stauraxonia homopola), in
the latter dissimilar (Stauraxonia heteropola). If all the secondary axes
be equal, the stauraxon ground-form is regularly radial. If some of them be
unequal they are arranged in certain relations towards two primary
transverse axes, perpendicular to each other, to which all the other
secondary axes are subsidiary; the ground-forms are then either amphithect
or bilateral. The two primary transverse axes, which may also be designated
"ideal transverse axes" (euthyni), divide the vertical main axis in its
centre; one of them is the sagittal, the other the frontal. These three
dimensive axes give the factors which accurately determine the ground-form
and the dimensions in most Radiolaria; the vertical main axis determines
the length (principal axis); one horizontal transverse axis determines the
thickness (sagittal axis), and the other the breadth (frontal axis). Those
ground-forms in which the transverse axes are isopolar are termed
"amphithect," and those in which the one (frontal or lateral) is isopolar
and the other (sagittal or dorso-ventral) is allopolar, are termed
"bilateral," or better "zeugitic."

46. _Primary and Secondary Ground-Forms._--The geometrical sphere must be
regarded as the original ground-form of the Radiolaria; it being understood
that its monophyletic derivation from a single stem-form, _Actissa_, is
correct. The simplest forms of _Actissa_ (_Procyttarium_, Pl. 1, fig. 1)
are in fact geometrically _perfect spheres_; indeed even the individual
parts which compose their unicellular bodies (nucleolus, nucleus, central
capsule and calymma) are concentric spheres. But in addition the central
capsules of most other SPUMELLARIA, especially the #Sphæroidea#, as well as
of many ACANTHARIA {xxii}are true spheres. Furthermore the simple or
concentrically composed lattice-spheres of #Sphæroidea#, #Sphærophracta#,
and #Phæosphæria# may be regarded as spheres, although strictly speaking
they are endospherical polyhedra. From the primary spherical form of the
Radiolaria all other secondary forms may be derived in the following
order:--1. By the development of a main axis the Monaxonia arise. 2. By the
development of transverse axes the Stauraxonia arise. 3. In both groups
(Monaxonia and Stauraxonia) the two poles (or upper and lower halves of the
body) are at first similar (Isopola). 4. By differentiation in the two
poles or halves of the body (distinction between the basal pole and the
apical) the forms with different poles (Allopola) arise. 5. The transverse
axes of the Stauraxonia are at first equal (regular pyramids and double
pyramids). 6. By differentiation in the transverse axes (distinction
between the sagittal and the frontal axis) the amphithect pyramids and
double pyramids arise. 7. From the amphithect pyramids the Amphipleura
arise by differentiation of both poles of the sagittal axis. 8. The
zygopleural ground-form appears last, as the simplest form of the
Amphipleura.

47. _The Ground-Forms of the Spumellaria._--The SPUMELLARIA, being the
oldest and most primitive Radiolaria, have for the most part either
indifferent or multistable equilibrium; _e.g._, all #Colloidea# and
#Beloidea# which have a spherical central capsule, and also most
#Sphæroidea#. Among these primitive Centrostigma true spheres and
endospherical polyhedra are represented in the utmost variety, and the
regular polyhedra in particular. By the development of a vertical main axis
these Centrostigma have also given rise to very numerous Centraxonia, which
are usually isopolar, very rarely allopolar. Sometimes they are Monaxonia
(circular in transverse section), sometimes Stauraxonia (polygonal in
transverse section). The vertical main axis is longer in the #Prunoidea#,
shorter in the #Discoidea# than any of the other axes. The #Larcoidea# are
distinguished by their lentelliptical or triaxial ellipsoid form; the three
different but isopolar axes corresponding with those of the rhombic
octahedron; but even among the #Sphæroidea#, #Prunoidea#, and #Discoidea#,
this form is sometimes produced by the differentiation of two different
transverse axes at right angles to each other. Whilst these ground-forms
(Centraxonia and Centrostigma) occur in the utmost variety among the
SPUMELLARIA, the centroplanar (or true bilateral) ground-form is entirely
wanting.

48. _The Ground-Forms of Acantharia._--In the small family Astrolophida,
which contains the most archaic forms of the legion (_Actinelius_,
_Astrolophus_), the ACANTHARIA show a direct relation to the most primitive
SPUMELLARIA (_Actissa_), and like these have indifferent equilibrium; their
central capsule is a sphere, their calymma an endospherical polyhedron,
whose angles are indicated by the distal ends of the numerous {xxiii}equal
radial spines. In the great majority of ACANTHARIA, however (all
#Acanthonida# and #Acanthophracta#), twenty radial spines are present,
regularly distributed, according to Müller's icosacanthan law, in five
parallel circles, each containing four crossed spines (p. 717). Usually the
twenty spines are equal, and the ground-form is the quadratic octahedron,
or a regular double pyramid with sixteen sides. But in some groups (the
Amphilonchida and Prunophracta) two opposite equatorial spines are much
more strongly developed than the other eighteen, and therefore the
hydrotomical axis in the equatorial plane is larger than the geotomical
axis (p. 719); the isopolar stauraxonian form passes over into the
allopolar, and the ground-form becomes the rhombic octahedron or the
amphithect double pyramid (compare §§ 33 and 34, and p. 720). The
centroplanar ground-form is entirely wanting in the ACANTHARIA.

49. _The Ground-Forms of the Nassellaria._--The NASSELLARIA all possess
monostatic ground-forms, inasmuch as by the very structure of their
monopylean central capsule a vertical main axis is necessitated, whose
basal pole occupies the porochora. The same arrangement is also for the
most part clearly recognisable in the corresponding structure of the
skeleton, which is generally either centraxon or centroplanar. Among their
manifold skeletal forms different larger groups of ground-forms may be
recognised according as the vertical allopolar main axis is crossed by
differentiated transverse axes or not (Stauraxonia or Monaxonia); the
former are either triradial or multiradial. The triradial, with three
lateral or terminal radial apophyses, constitute the greater part of the
NASSELLARIA, and have probably been derived originally from the triradial
#Plectoidea# (_Triplagia_, _Triplecta_); a more careful examination,
however (especially with reference to the structure of the cortinar
septum), reveals the fact that the ground-form is not strictly regularly
pyramidal (with three equal radii), but amphipleural (with two paired
ventral and one unpaired dorsal radius), and that it usually passes over
into a distinctly zygopleural form. The same holds true of the multiradial
NASSELLARIA, where for the most part three interradial or six adradial
(sometimes more) apophyses are intercalated between the three primary
perradial ones; sometimes here also the ground-form is a quite regular
hexagonal or nonagonal pyramid, but usually it is more or less amphithect
or amphipleural. Among the eradial NASSELLARIA, which have no radial
apophyses, the ground-form is sometimes allopolar monaxon (conical, ovoid,
hemispherical, &c.), sometimes amphithect pyramidal (even in the simplest
Stephanida, _Archicircus_, &c.), or sometimes distinctly zygopleural or
bilateral (many #Plectellaria#).

50. _The Ground-Forms of the Phæodaria._--The PHÆODARIA agree with the
NASSELLARIA in the possession of a primitively centraxon ground-form, and
like them are monostatic, since a vertical main axis whose basal pole
passes through the astropyle is present, owing to the characteristic
structure of their cannopylean central capsule. In {xxiv}the great majority
of PHÆODARIA the spheroidal central capsule also possesses a pair of
parapylæ near the opposite apical pole of the main axis (Tripylea), and
these determine (as the right and left secondary openings) an isopolar
frontal axis. Hence, strictly speaking, in most PHÆODARIA the central
capsule has the geometrical ground-form of the amphithect pyramid (as in
the Ctenophora), with an allopolar vertical main axis, and two unequal, but
isopolar, horizontal transverse axes. In many PHÆODARIA the skeleton also
has this amphithect pyramidal ground-form, _e.g._, the bivalved
#Phæoconchia# and part of the #Phæogromia#. On the contrary, in the rest of
the PHÆODARIA the skeleton exhibits very various geometrical ground-forms,
independent of that of the central capsule. In the #Phæosphæria# it forms
preferably spheres or endospherical polyhedra, as also in the Castanellida
and Circoporida among the #Phæogromia#; among the Circoporida there are
also seen with remarkable distinctness the regular polyhedra (especially
the dodecahedron and icosahedron). Isopolar monaxonia are found among the
Aulosphærida (_Aulatractus_) and Orosphærida; allopolar monaxonia among the
Challengerida (_Lithogromia_). The Medusettida and Tuscarorida show various
forms of regular pyramids (allopolar Stauraxonia); and finally, the
Challengerida are for the most part centroplanar or bilateral. Thus the
PHÆODARIA present a great wealth of different geometrical ground-forms in
the development of their skeleton, not in that of their central capsule.

CHAPTER II.--THE CENTRAL CAPSULE.

51. _Components of the Central Capsule._--In all Radiolaria without
exception, at some period of life or other, the central portion of the soft
body is separated from the peripheral portion by an independent,
anatomically recognisable membrane; this membrane with all its contents is
designated the central capsule, and is the peculiar central organ of the
unicellular body, which distinguishes the Radiolaria most clearly from the
other Rhizopoda. In the great majority of the Radiolaria the volume of the
central capsule is less than that of the surrounding peripheral soft body
which we place in opposition to it as "extracapsulum." The
"capsule-membrane," which separates these two constituents, arises very
early in most Radiolaria, and persists throughout their whole life. In some
species, however, the membrane only appears later, immediately before the
formation of the spores, and hence is absent for a considerable period.
Regarded as a whole, then, the capsule consists of the following
parts:--(1) the capsule-membrane; (2) the enclosed endoplasm, or
intracapsular protoplasm; (3) the nucleus. But in addition, many other
non-essential structures may be enclosed in the central capsule, especially
hyaline spheres (vacuoles), fatty spheres, pigment granules, crystals, &c.

The central capsule was first described in my Monograph in 1862 (pp.
69-82) as the most characteristic component of the Radiolarian organism,
and distinguished from the whole extracapsular {xxv}soft body. The fact
that it has recently been reported as absent by various authors is due to
their having observed young or unripe specimens, before the formation of
the spores. In some species of #Polycyttaria# and ACANTHARIA the membrane
persists only a very short time.

52. _The Primary Form of the Central Capsule._--The form of the central
capsule is originally a geometrical sphere; and if in accordance with our
monophyletic hypothesis all Radiolaria are to be derived from one common
stem-form (_Actissa_, see p. 12), then the central capsule of this common
stem-form must be regarded as perfectly spherical (_Procyttarium_, p. 13,
Pl. 1, fig. 1). Since, further, the enclosed nucleus and the surrounding
calymma of this primitive archaic form must also be spheres, and since the
nucleus lies in the centre of the body, and the protoplasm is evenly
distributed between it and the membrane, it follows that no axes or
excentrically differentiated parts are to be distinguished in this most
primitive Radiolarian. Rather in the primary central capsule all parts are
concentrically and evenly arranged round its centre. This primary spherical
form becomes modified in most Radiolaria into various secondary
ground-forms, which are correlated partly with the structure of the capsule
itself, and partly also with the development of openings in its membrane.
In general the ground-form of the central capsule is polyaxon in the
Porulosa (SPUMELLARIA and ACANTHARIA); but in the Osculosa centraxon forms
are more frequently observed; in the NASSELLARIA the ovoid (allopolar
monaxon) form is predominant, and in the PHÆODARIA the rhomboid or
amphithect pyramid. In these latter, the astropyle indicates the basal pole
of the vertical main axis, whilst the two parapylæ (right and left) mark
the poles of the frontal transverse axis. In the NASSELLARIA the centre of
the porochora corresponds with the basal pole of the main axis, whilst no
transverse axes are originally present.

53. _The Secondary Forms of the Central Capsule._--The original purely
spherical form of the central capsule persists only in the minority of the
Radiolaria, namely, the greater part of the SPUMELLARIA and ACANTHARIA; it
passes over into various other secondary forms in the majority of the
class, in the whole of the NASSELLARIA and PHÆODARIA, and in a considerable
portion of the SPUMELLARIA and ACANTHARIA. These secondary or derived forms
may be divided into two quite distinct groups, which may be designated
endometamorphic and exometamorphic; in the former the cause of the
divergence of the secondary form from the sphere lies in the internal
structure of the central capsule; in the latter it lies in the external
influence exerted by the growth of the skeleton. Obviously the former
series of modifications is more significant than the latter.

54. _The Endometamorphic Forms of the Central Capsule._--The secondary
forms of the central capsule, which are due to internal causes connected
with its growth, are as follows:--

{xxvi}A. _The Ellipsoidal Central Capsule_, with one axis elongated, so
that it becomes the vertical main axis of the body.

_a._ Among the SPUMELLARIA, _Actiprunum_ (p. 14), _Colloprunum_ (p. 25,
Pl. 3, fig. 9), most #Prunoidea# (p. 288).

_b._ Among the ACANTHARIA, many Amphilonchida (p. 782, Pl. 132, figs.
2, 6), and Belonaspida (p. 861).

_c._ Among the NASSELLARIA, many #Plectoidea# (p. 905, Pl. 91, figs. 5,
9), #Stephoidea# (p. 937, Pl. 81, fig. 16), Monocyrtida (Pl. 51, fig.
3), &c.

B. _The Cylindrical Central Capsule_, with considerable elongation of the
vertical main axis, which is several times as long as the horizontal
transverse axis.

_a._ Amongst the SPUMELLARIA, _Collophidium_ (p. 26, Pl. 3, figs. 1-3)
and many #Prunoidea# (_Spongurus_, &c.).

_b._ Among the ACANTHARIA, some Amphilonchida.

C. _The Discoidal, Spheroidal, or Lenticular Central Capsule_, with one
axis shorter than the others, which becomes the vertical main axis.

_a._ Among the SPUMELLARIA, _Actidiscus_ (p. 15), _Collodiscus_ (p.
27), and the large group #Discoidea# (p. 408).

_b._ Among the ACANTHARIA, many Quadrilonchida (p. 768, Pl. 131), and
most Hexalaspida (p. 874).

_c._ Among the NASSELLARIA, certain #Stephoidea# and #Cyrtoidea#.

_d._ Among the great legion PHÆODARIA the spheroidal central capsule is
almost always more or less flattened in the direction of the main axis
(p. 1525, Pls. 101-128).

D. _The Lentelliptical Central Capsule_ (or triaxial ellipsoid), with
three unequal but isopolar axes at right angles to each other, the
sections in all three dimensions of space being ellipses.

_a._ Among the SPUMELLARIA, _Actilarcus_ and the large group
#Larcoidea# (p. 604).

_b._ Among the ACANTHARIA, certain Amphilonchida and Belonaspida.

E. _The Polymorphic, Amoeboid or Irregular Central Capsule._

_a._ Among the SPUMELLARIA, _Collodastrum_ (p. 28, Pl. 3, figs. 4, 5),
and some #Larcoidea#.

55. _The Exometamorphic Forms of the Central Capsule._--The secondary forms
of the central capsule, which are brought about by external causes, chiefly
dependent on the formation of the skeleton, are very various and in many
cases devoid of special interest; in other instances, on the contrary, they
are of great importance, because of the clear relation of cause and effect
which can be traced between the development of the skeleton and of the
capsule. The most important phenomena to be recorded in this connection are
as follows:--

{xxvii}I. SPUMELLARIA.--(A) In many of the #Sphæroidea#, the central
capsule of which is originally enclosed by a simple lattice-sphere, it
puts out protrusions through the meshes of the shell, thus forming
club-shaped processes, corresponding in number with the meshes of the
lattice (Pl. 11, figs. 1, 5; Pl. 20, fig. 1_a_; Pl. 27, fig. 3, &c.). The
whole surface of the spherical capsule may thus be covered with numerous
independent radial clubs of equal size, but usually they unite again
outside the shell to form a simple sphere with smooth surface. (B) In
many #Prunoidea# whose originally ellipsoidal body has become cylindrical
by the marked prolongation of the main axis, the central capsule is
divided by a series of constrictions into segments, which correspond with
the annular constrictions of the skeleton (Pls. 39, 40). (C) In most
#Discoidea# whose lentiform or discoidal shell develops radial arms at
its margin, the central capsule sends out processes into these arms, and
adapts itself to the stellate form of the skeleton (p. 409, Pl. 43, fig.
15; Pl. 47, &c.) (D) In many #Larcoidea# whose growth is originally
lentelliptical, but later spiral or irregular, the central capsule
follows the mode of growth and develops irregular protuberances.

II. ACANTHARIA.--Whilst the central capsule of most ACANTHARIA retains
its primitive spherical form, in a minority of the group it passes over
into various secondary forms, which are directly determined by the growth
of the skeleton; especially common are lappet or club-shaped prominences
which follow the larger radial spines. Hence the central capsule may
assume the form of a violin, with two lobes corresponding to the two
poles of the elongated main axis, as in many Amphilonchida (p. 782, Pl.
132, fig. 10), and the Diploconida (p. 884, Pl. 140). On the other hand
the central capsule becomes cruciform, with four lobes disposed at right
angles, as in Lithoptera and other Quadrilonchida (p. 768, Pl. 131, fig.
10, &c.).

III. NASSELLARIA.--The primitive ellipsoid or ovoid form of the central
capsule persists only in a few NASSELLARIA, such as the simplest and most
archaic forms, the Nassellida, many #Plectoidea#, #Stephoidea#,
Monocyrtida, &c. In the great majority of the NASSELLARIA, on the
contrary, the ellipsoid or ovoid form passes over into a secondary form
which is usually characterised by the presence of lobes, and is obviously
dependent upon the previous development of the skeleton. In many
#Stephoidea# and #Spyroidea# (probably the majority), a bilobed central
capsule is formed (with symmetrically equal right and left lobes), since
the primary vertical sagittal ring interferes with the growth in the
median plane (Pl. 90, figs. 7-10). In other {xxviii}#Spyroidea#, on the
contrary, and the majority of the #Cyrtoidea#, the central capsule forms
at its basis rounded lobes, which protrude and hang down from the meshes
of the cortinar plate; and since this latter has usually three or four
large pores, the capsule similarly develops three or four processes (Pl.
53, fig. 19; Pl. 55, figs. 4-11; Pl. 59, figs. 4-13; Pl. 60, figs. 3-7;
Pl. 65, fig. 1).

56. _The Membrane of the Central Capsule._--The capsule-membrane or
envelope of the central capsule is both morphologically and physiologically
one of the most important parts of the Radiolarian body, for it separates
its two main constituents, the capsule with its nucleus and endoplasm and
the extracapsulum with the calymma and exoplasm. The capsule-membrane is
invariably present at some time or other during the life of the organism,
even though in a few species it may persist only for a short time. It is
characterised in general by its power of resistance to chemical and
physical reagents, and appears to be related to the elastic tissues or
perhaps even more to the chitinous substances. Its thickness is usually
less than 0.0001, though in certain groups it ranges between 0.001 and
0.002, and in many of the larger Radiolaria (such as Collida and PHÆODARIA)
it may attain a thickness of 0.003 to 0.006 or more. In the three legions
SPUMELLARIA, ACANTHARIA, and NASSELLARIA the capsule-membrane is single,
while in the PHÆODARIA it is always double, being composed of a firm outer
and a delicate inner membrane, which are in contact at only few points.
Usually it is quite structureless, except for its apertures; the thicker
membrane showing occasionally a fine concentric lamination. In certain
large #Colloidea# (_e.g._, _Thalassicolla_, Pl. 1, fig. 5_b_) the membrane
is covered on the inner surface by a network of polygonal ridges, and in
some large PHÆODARIA with remarkable small curved rods (Pl. 114, fig. 13).
In all Radiolaria the membrane is perforated by definite openings or pores,
through which the intracapsular and extracapsular protoplasm are in direct
communication. These openings (or "pylae") show very characteristic and
constant differences in the four legions, which have given rise to the
names--PERIPYLEA, ACTIPYLEA, MONOPYLEA, CANNOPYLEA.

The capsule-membrane was first indicated as the most important and
absolutely constant component of all Radiolaria, and as the differential
character of the class, in my Monograph (1862, pp. 69-71). The careful
investigations of R. Hertwig have confirmed this view and at the same
time have yielded the most important conclusions regarding the nature and
systematic significance of the openings in the capsule (_op. cit._, 1879,
pp. 105-107). On the contrary, Karl Brandt has recently propounded the
theory that the capsule-membrane is by no means a constant part of the
Radiolarian organism, but is lacking in certain species of _Collozoum_
and _Sphærozoum_ (1881, p. 392). This contradiction is explained by the
fact that in some #Collodaria# and #Acanthometra# the formation of the
central capsule takes place much later than in the other Radiolaria, in
some {xxix}species indeed only just prior to the development of the swarm
spores. I have recognised the presence of it in all species which I have
investigated (more than a thousand), and even in those in which Brandt
denies its existence. It is often very delicate and may easily be
overlooked, especially when the contents of the capsule are colourless,
but in all cases by the prudent application of staining fluids and other
reagents its presence may be demonstrated. Even in those cases in which
the contour of the capsule was not visible, and its contents appeared to
pass without definite boundary into the matrix of the extracapsulum, it
was possible by the use of appropriate stains or reagents, which would
not penetrate the capsule, or of those solvents which were capable of
dissolving its contents and of causing it to swell up like a distended
bladder, to recognise the existence of the membrane. Those Radiolaria in
which it is truly absent are young animals of species in which the
membrane is only formed immediately before sporification, and persists
but for a short time (_e.g._, species of _Collozoum_, _Sphærozoum_,
_Acanthometra_, _Acanthochiasma_, &c.).

57. _The Capsule-Openings of the Peripylea (or Spumellaria)._--The
capsule-membrane of the PERIPYLEA is generally perforated by extremely fine
and numerous pores, which are distributed at equal distances over the whole
surface, and are precisely alike in all parts of the capsule. Hence the
SPUMELLARIA may be called "Holotrypasta" or "Porulosa"; they agree with the
ACTIPYLEA in being devoid of an osculum or operculum; they are
distinguished from the latter group mainly in that their pores are equally
distributed over the whole surface of the capsule, whilst in the ACTIPYLEA
the pores are disposed in definite groups or lines, separated by large
imporous areas.

The central capsule of the SPUMELLARIA, with its innumerable fine and
evenly distributed pores, must be regarded as the primitive arrangement,
from which the different central capsules of the three other legions have
been developed. The central capsule of the ACTIPYLEA has been derived
from that of the PERIPYLEA by reduction in the number of the pores and
their distribution in definite, regularly disposed areas in the membrane.
The central capsule of the Osculosa is characterised by the formation of
a special main-aperture (osculum) at the basal pole, which is closed in
the MONOPYLEA by the porochora, and in the CANNOPYLEA by the astropyle;
the remaining pores, with the exception of the accessory openings of many
CANNOPYLEA, remain undeveloped in both these legions. In the same way
Hertwig regards the central capsule of the PERIPYLEA as the primitive
form (1879, L. N. 33, p. 107).

58. _The Capsule-Openings of the Actipylea (or Acantharia)._--The
capsule-membrane of the ACTIPYLEA is perforated by very numerous fine
pores, which are regularly distributed over the surface of the central
capsule, and separated by imporous intervals. Hence the ACANTHARIA belong
to the "Holotrypasta" or "Porulosa"; they have neither osculum nor
operculum, and agree in this particular with the PERIPYLEA; but they are
separated from these latter chiefly by the fact that their pores are much
less numerous, and marked off into regularly arranged groups or lines by
imporous intervals. In the PERIPYLEA, on the contrary, the pores are much
more numerous and are evenly distributed over the whole surface of the
capsule.

{xxx}The central capsule of the ACANTHARIA has hitherto been for the most
part confounded with that of the SPUMELLARIA, and no clear distinction
has been drawn in this respect between the two legions of the Porulosa.
Hertwig, who in 1879 first discovered the remarkably different structure
of the Osculosa (NASSELLARIA and PHÆODARIA), recognised no distinction
between the structure of the capsules in the PERIPYLEA and ACTIPYLEA (his
Acanthometrea), and supposed that in both these legions "very fine pores
were evenly distributed in large numbers over the capsule-membrane"
(_loc. cit._, p. 106). I have, however, during the last few years
convinced myself, by the careful comparative investigation of numerous
ACANTHARIA, that in this respect they are quite distinct from the
SPUMELLARIA (with perhaps the exception of the Astrolophida, which are
nearly related to the primitive _Actissa_). The number of pores in the
ACTIPYLEA is usually very much smaller than in the PERIPYLEA, and they
are regularly arranged in groups.

59. _The Capsule-Openings of the Monopylea (or Nassellaria.)_--The
capsule-membrane of the MONOPYLEA always possesses a single large
main-opening, an osculum, which lies at the basal pole of the main axis,
and is closed by a circular perforated lid (operculum porosum). When seen
from the surface this lid appears as a clearly defined porous area
(porochora or area porosa), and forms the horizontal base of a peculiar
cone, which stands vertically in the interior of the capsule and may be
designated the "thread-cone" (podoconus). The NASSELLARIA may hence be
termed "Merotrypasta" or "Osculosa," like the CANNOPYLEA; the structure and
significance of the circular lid (operculum), which closes the main-opening
(osculum) is, however, quite different in the two legions. Whilst the lid
of the CANNOPYLEA (astropyle) is solid, traversed by radial ribs, and only
perforated in its centre by a short tube (proboscis), in the MONOPYLEA the
operculum (porochora) is always perforated by numerous vertical fine pores,
and is in connection with the peculiar internal "pseudopodial cone"
(podoconus, Pl. 51, figs. 5, 13; Pl. 81, fig. 16; Pl. 91, fig. 5; Pl. 98,
fig. 13). The pores are separated by small vertical, highly refractive rods
(opercular rhabdillæ); these become intensely stained by carmine, and are
either evenly distributed over the surface of the porochora or arranged in
definite groups. The outer or distal end of each rod is rounded, sometimes
thickened like a club or split into lobes; the inner or proximal end is
usually pointed, and stands in connection with a myophane thread of the
podoconus (see § 79). The primary circular form of the porochora, in which
the opercular rhabdillæ are evenly distributed in a horizontal plane,
undergoes various secondary modifications in many NASSELLARIA. The
triradial structure of the skeleton, which characterises the majority of
the legion, causes a splitting of the base of the central capsule into
three or four lobes; this division also affects the porochora, which lies
in the centre of the base, so that the rhabdillæ become arranged in three
or four equal circles. If, however, the lobes of the central capsule become
larger and protrude through the three or four collar pores of the cortinar
septum, the central porochora may separate entirely into three or four
elongated tracts, which lie on the axial side of the magnified lobes; the
rhabdillæ are then arranged over the whole surface of {xxxi}these tracts,
on the outer aspect of which run the longitudinal myophane fibrillæ of the
podoconus (compare §§ 79 and 99).

The porous area of the MONOPYLEA was first described by Hertwig in 1879,
and shown to be the characteristic main-opening of the central capsule in
various families belonging to this legion (L. N. 33, pp. 71, 73, 83, 106,
Taf. vii., viii.). According to his view "the capsule-membrane in the
porous area becomes thickened around each pore into a rod, perforated by
a canal," and the intracapsular protoplasm passes outwards through these
fine canals (_loc. cit._, p. 106). I am not able to share this
interpretation, but think rather that I have convinced myself by the
examination of some living NASSELLARIA, and of many well-stained and
preserved preparations in the Challenger collection, that the rods are
_solid_, specially modified portions of the capsular wall, and that the
protoplasm does not pass through them but through pores which lie between
them.

60. _The Capsule-Openings of the Cannopylea (or Phæodaria)._--The
capsule-membrane of the CANNOPYLEA always possesses only a single large
main-opening or osculum, which lies at the basal pole of the vertical main
axis, and is closed by a circular radiated lid (operculum radiatum). This
operculum appears, when seen from the surface, as a sharply defined
stellate area (astropyle), from the middle of which arises a shorter or
longer cylindrical tube, the proboscis. Hence the PHÆODARIA, like the
MONOPYLEA, belong to the "Merotrypasta" or "Osculosa"; the structure and
significance of the circular operculum, which closes the main-opening
(osculum), are, however, quite different in the two legions. Whilst the
operculum of the MONOPYLEA (porochora) is perforated by numerous fine
vertical pores, and connected with the peculiar internal pseudopodial cone
(podoconus), this structure is entirely wanting in the CANNOPYLEA, and
instead of it there is a solid operculum, with radial ribs which originate
at the base of its central tubular mouth; this tube (proboscis) is
cylindrical, often conical at the base, of very variable length and with a
round aperture at either end. In spite of the great difference which the
various families of CANNOPYLEA exhibit in the formation of their skeleton
and its appendages, the constitution of this characteristic stellate
main-opening (astropyle) is always essentially the same; both the stellate
operculum itself, and the proboscis which rises from its centre, show only
slight differences in the various groups. In addition to this large
main-opening most PHÆODARIA possess several small accessory openings
(parapylæ); and usually two of these are present, placed symmetrically
right and left of the aboral pole of the main axis and in the frontal plane
(Pl. 101, figs. 2, 6, 10; Pl. 104, figs. 1, 2_a_). Sometimes there are more
numerous accessory openings (three to six or more) regularly arranged, as
in the two peculiar families, Circoporida and Tuscarorida; occasionally
also there is only a single parapyle, at the aboral pole of the main axis
(_e.g._, in _Tuscaridium_). The parapylæ seem to be quite absent in the
families Challengerida, Medusettida, Castanellida, and perhaps also in
other PHÆODARIA. The form and structure of the small accessory openings
appear to be always the same. The {xxxii}outer capsule-membrane is elevated
in the form of a short cylindrical tube or "apertural ring" (collare
paraboscidis), the external margin of which bends inwards, and at the base
of the ring passes over into the delicate internal capsule membrane. Upon
this apertural ring is situated a longer or shorter "apertural cone"
(paraboscis), which is a tubular, cylindrical or conical, prolongation of
the membrane, open externally.

The peculiar capsule-openings of the PHÆODARIA were first discovered and
carefully described by Hertwig in 1879 (L. N. 33, pp. 95, 107). He found
in all the six genera which he examined _three_ openings, a main-opening
at the basal pole of the main axis and two accessory openings, one on
either side of the apical pole; hence he named the whole group
"TRIPYLEA." This name, however, is not applicable to the numerous
PHÆODARIA mentioned above, which have only a main opening without any
accessory openings, nor to those genera in which the number of the latter
is variable. I have, therefore, replaced Hertwig's designation by the
term "CANNOPYLEA," which has reference to the peculiar tubular form of
the opening. This I find much more developed in many PHÆODARIA than
Hertwig has represented, and I must also, in certain particulars, dissent
from his delineation of the minute structure, although this is in the
main remarkably accurate.

61. _The Nucleus._--The nucleus, enclosed in the central capsule of all
Radiolaria, behaves in every respect like a true cell-nucleus, and thus
lies at the base of the now universal opinion, that the whole Radiolarian
organism, in spite of its varied development and remarkable variations, is
unicellular and remains throughout life a true individual cell. This
important theory is not invalidated by the fact that the nucleus undergoes
peculiar modifications in many groups, and in certain groups presents
appearances seldom or never seen elsewhere.

62. _Uninuclear and Multinuclear Radiolaria (Monocaryotic and
Polycaryotic)._--All Radiolaria present two different conditions in respect
of the behaviour of the nucleus, since in their young stages they are
uninuclear (_monocaryotic_), and in later stages multinuclear
(_polycaryotic_). This is readily explained by the fact that each
individual Radiolarian is developed from a simple unicellular swarm-spore,
and that afterwards, before the formation of swarm-spores, the single
nucleus divides into many small nuclei. Thus in the Radiolaria the nucleus
is pre-eminently the _organ of reproduction and inheritance_. The division
of the originally single nucleus into many small nuclei may take place,
however, at very different periods, so that the Radiolaria may be divided
in this respect into precocious and serotinous.

63. _Serotinous and Precocious Radiolaria._--In the great majority of the
Radiolaria the division of the nucleus takes place only at a late period, a
short time or even immediately before the process of spore formation; it
then breaks up rapidly into numerous small nuclei (always more than one
hundred, sometimes many thousands), and each of these {xxxiii}either
becomes itself the nucleus of a swarm-spore, or by repeated division gives
rise to a group of spore-nuclei. All those Radiolaria which are uninuclear
during the greater part of their existence, and in which the process of
division is late, and takes place rapidly, are called "serotinous" or
late-dividing forms. To this category belong all PHÆODARIA and NASSELLARIA,
as well as all the solitary or monozoic SPUMELLARIA and some ACANTHARIA. On
the other hand, the name "precocious," or early dividing, is applied to
those Radiolaria in which the division of the nucleus takes place very
early, and in which, therefore, the cell is multinuclear during the greater
part of its existence. This is the case in all the social or polyzootic
Radiolaria (Polycyttaria, Pls. 3-8), and also in the great majority of the
ACANTHARIA, both #Acanthometra# and #Acanthophracta#. In the last two
groups, however, there are numerous exceptions, and these are seen in
remarkably large species, characterised by the great size of the central
capsule. From a phylogenetic point of view, the conclusion is allowable
that the precocious forms are secondary, and have arisen by adaptive
modification from the primitive serotinous stem. In the Polycyttaria (or
social SPUMELLARIA, _i.e._, the three families Collozoida, Sphærozoida, and
Collosphærida), the cause of the adaptation lies most probably in the
formation of the colony itself, for all these three families are so closely
related to three corresponding families of serotinous, monozootic
Radiolaria (Thalassicollida, Thalassosphærida, Ethmosphærida), that certain
species of the latter are hardly to be distinguished from isolated
individuals of the former. Perhaps the remarkable formation of the large
central oil-globule, which particularly characterises the Polycyttaria, is
the prime cause of their early nuclear division. In the ACANTHARIA the
cause is most likely to be found in the characteristic _centrogenous
development_ of their acanthin skeleton, whose radial bars first of all
appear in the centre of the capsule. Hence arises directly the excentric
position of the nucleus, which in the archaic stem of ACANTHARIA
(_Actissa?_) was probably central. In any case, but little weight is to be
laid upon the precocious division of the nucleus in the ACANTHARIA in
general, inasmuch as in certain species (both #Acanthometra# and
#Acanthophracta#) the more usual serotinous division persists.

64. _Central and Excentric Nuclei._--The position of the nucleus in the
interior of the central capsule was no doubt primitively central, and this
situation in the geometrical centre of the original spherical central
capsule has been accurately retained in all monozootic SPUMELLARIA; in the
polyzootic families of this legion (Polycyttaria), on the contrary, it is
obscured by the precocious division of the nucleus. In the other three
legions, which may be phylogenetically derived from the SPUMELLARIA, the
position of the nucleus is rarely central, but usually excentric, or at
most subcentral. In the ACANTHARIA (both #Acanthometra# and
#Acanthophracta#) the central position of the nucleus is at once excluded
by the constantly centrogenous development of the skeleton; the nucleus is
therefore always excentric, and may lie at either side; it usually
{xxxiv}divides very early into numerous separate nuclei, which are usually
distributed in the peripheral portions of the central capsule. In the
NASSELLARIA the development of the porochora, and of the podoconus which
stands upon it, brings about the formation of a vertical axis, and in
consequence the central capsule assumes a monaxon form (usually ovoid or
conical); the nucleus then lies in the main axis, but excentrically between
the apex of the podoconus and the aboral pole. In many NASSELLARIA,
however, especially when the podoconus is so large that its apex approaches
the aboral pole of the central capsule, the nucleus is pressed to one side
and lies quite excentrically. The PHÆODARIA exhibit a different
arrangement; the large spheroidal nucleus is always subcentral, so that its
main axis corresponds with that of the concentric spheroidal central
capsule; but since the astropyle always occupies the oral pole of the
latter, and since the distance of the nucleus from this pole is always
somewhat different from its distance from the other, it follows that,
strictly speaking, the nucleus never lies accurately in the geometrical
centre.

65. _Homogeneous and Allogeneous Nuclei._--The nucleus of the Radiolaria
not only exhibits a similar structure and composition, and suffers similar
modifications to those which are found to occur in the case of other
cell-nuclei, but also to some extent shows very peculiar developmental
forms, which are seldom or never found in other cells. In the first place
the nuclei may be divided into homogeneous and allogeneous, the former are
structureless and consist of a uniform mass of nuclein, whilst the latter
are composed of different substances and show various structural relations.
_Homogeneous_ nuclei, whose whole mass is uniform and exhibits no
structural differentiation, are probably always to be found in the
swarm-spores; in the fully developed Radiolarian body they are found only
in the first legion, SPUMELLARIA, and that both in many Monozoa (especially
small #Sphæroidea# and #Prunoidea#) and in the Polyzoa (or Polycyttaria).
The whole mass of these homogeneous nuclei, which are usually spherical or
ellipsoidal, consists of uniform, perfectly clear and transparent nuclein,
and becomes evenly stained by carmine, hæmatoxyline, &c. They may be
readily distinguished by these means from the clear vacuoles or "hyaline
vesicles," which are evenly distributed in the endoplasm of many
Radiolaria, and may be confused with the former. _Allogeneous_ nuclei,
which are always composed of different parts and often show complicated
structural relations, are found developed in the great majority of
Radiolaria. The most important differentiation exhibited by these secondary
forms is the separation of the nuclear mass into a firm nuclear substance
(caryoplasm) and a fluid nuclear juice (caryolymph). In addition in each
nucleus a nucleolus is visible, and often several or many may be seen (see
§§ 67 to 70).

66. _The Form of the Nucleus._--The nucleus of the Radiolaria shows greater
variations in form and structure than are to be found in the majority of
cell-nuclei; {xxxv}exception must, however, be made in the case of many
animal ovicells, which, in their peculiar form and composition, often
recall large Radiolarian nuclei. With respect to the external shape two
main forms may be distinguished, as primary and secondary. The _primary
form_ of the Radiolarian nucleus is the sphere; it occurs not only in most
swarm-spores, but also in most adult forms belonging to the legion
SPUMELLARIA, and in individual instances in other groups; indeed the nuclei
of most SPUMELLARIA, as also the concentric central capsules in which they
lie, are true geometrical spheres. The _secondary forms_ of the nucleus are
found in the majority of adult Radiolaria, and arise from the primary
spherical forms in various ways, either by the elongation or contraction of
one axis, or by the formation of apophyses or processes. The most important
of these secondary forms are as follows:--

1. _Ellipsoidal nuclei_, arising by elongation of one principal axis;
very common among the NASSELLARIA, as well as in many #Prunoidea# and
#Larcoidea# among the SPUMELLARIA; also in several ACANTHARIA.

2. _Discoidal nuclei_, arising by contraction of one principal axis,
sometimes lenticular or spheroidal, biconvex, sometimes shaped like a
disc or coin; especially common in the #Discoidea# among the SPUMELLARIA,
also in some ACANTHARIA; the large nucleus of the PHÆODARIA is always
spheroidal or almost spherical, with a slightly shortened main axis.

3. _Stellate nuclei_, spherical, and armed with evenly distributed radial
club-shaped or conical processes; rare but very characteristic,
especially in the two large Thalassicollida _Thalassopila_ (Pl. 1, fig.
3), and _Thalassophysa_ (Monogr. d. Radiol., Taf. i.); also in some
#Sphærellaria# (Pl. 11, fig. 5).

4. _Amoeboid nuclei_, with unequal processes irregularly arranged, in
certain irregular forms of SPUMELLARIA and ACANTHARIA.

5. _Lobate nuclei_, with several (usually two or three) large ovoid or
pyriform lobes, which protrude into corresponding larger lobes of the
central capsule, in many NASSELLARIA, especially the multiarticulate
#Cyrtoidea# (Pl. 59, figs. 12, 13). The budding nucleus of the ACANTHARIA
is also lobate (Pl. 129, figs. 6-11).

67. _The Nucleus of the Peripylea._--The nucleus of the SPUMELLARIA or
PERIPYLEA shows in certain groups a very primitive arrangement, indeed the
archaic structure from which the various forms of nuclei of other
Radiolaria may be derived; but on the other hand, in other groups it
exhibits very peculiar and remarkable differentiations. In the first place
it may be noted that the monozootic or solitary SPUMELLARIA usually possess
a single serotinous nucleus, which only divides into numerous swarm-spores
at a late period; {xxxvi}whilst, on the contrary, the polyzootic colonial
SPUMELLARIA (or Polycyttaria) are uninuclear only in the young state (Pl.
3, fig. 12), and speedily present numerous small homogeneous nuclei, which
have arisen by precocious division of a single nucleus; these are usually
spherical and 0.008 to 0.012 mm. in diameter. The serotinous nucleus of the
monozootic SPUMELLARIA, in many divisions of this large legion, and
especially in the simply constituted #Sphæroidea#, is a homogeneous sphere
of nuclein, lying in the middle of the central capsule. In many other cases
it assumes the form of a spherical vesicle ("Binnen-Bläschen"), whose fluid
or semi-fluid contents are enclosed by a more or less firm membrane. This
vesicle often contains a single central spherical _nucleolus_ (Pl. 1, figs.
1_l_, 4_l_), but sometimes a variable number of small excentric nucleoli
(Pl. 1, figs. 1_a_, 2_a_). The nuclear membrane is often somewhat thick,
presenting a double contour, and in such cases may even exhibit a fine
radial striation, the expression of minute pores (Pl. 1, fig. 2_a_). In the
colossal nuclei (as much as 1 to 2 mm. in diameter) of certain large
Thalassicollida the nucleolus presents a very remarkable form, becoming
stellate by the protrusion of processes, which may again branch in a
dendritic fashion (as in the common _Thalassicolla nucleata_), or it may
develop into a very long cylindrical thread, which is disposed in
serpentine coils, and in _Thalassophysa pelagica_ passes into the different
cæcal processes of the stellate nucleus. In many #Sphæroidea#, whose
skeleton is composed of numerous concentric lattice spheres, the small
central spherical nucleus lies at first within the innermost of these (the
medullary shell); but afterwards it grows through the meshes of the
lattice-work, and the radiating club-shaped processes thus formed (Pl. 11,
fig. 5) unite with each other outside the medullary shell, and form an
external nuclear sphere which completely encloses the latter. In the
Polysphærida (with several concentric lattice-shells) and in the
Spongosphærida (with spongy lattice-spheres), this process may be several
times repeated, so that eventually the central spherical nucleus attains
considerable dimensions, and encloses two or more concentric lattice-shells
with their radial connecting rods. The nuclear membrane is in these cases
usually penetrated by radial bars, which connect the outermost of the
enclosed shells with the remaining cortical shells which surround the
central capsule. The same remarkable arrangement is also very common among
the #Discoidea#. The small spherical primary nucleus is in such instances
immediately surrounded by the innermost earliest developed lattice-shell,
around which the concentric rings are subsequently deposited; it then grows
out through the meshes, and the processes fuse outside the ring to form a
homogeneous lentiform nucleus (Pl. 43, fig. 15). The same process recurs in
certain #Prunoidea# and #Larcoidea#, whilst in other SPUMELLARIA of these
groups (_e.g._, Pylonida) the lobate processes of the nucleus remain free.

Both the simple serotinous nucleus of the monozootic SPUMELLARIA, and the
numerous precocious nuclei of the Polycyttaria, were first described in
my Monograph in 1862, the former as the "endocyst" ("Binnen-Bläschen"),
the latter as "spherical transparent vesicles" ("Kugelige
{xxxvii}wasserhelle Bläschen"). I was in error, however, in regarding the
latter as identical with the so-called "hyaline spherules" in the central
capsule of many Monozoa, which rather belong to the category of
intracapsular vacuoles (see § 72). The credit of recognising, by the aid
of the modern methods of staining, the distinctness of these two
structures, which may readily be mistaken for each other, and of
demonstrating the true nature both of the serotinous and precocious
nuclei, belongs to Richard Hertwig (1879, L. N. 33).

68. _The Nucleus of the Actipylea._--The nucleus of the ACANTHARIA or
ACTIPYLEA shows very peculiar relations in respect of structure and
division, particularly special forms of lobular budding, which belong to
the characteristic peculiarities of this singular legion, and are not found
among other Radiolaria. The position of the nucleus is _always excentric_,
even in the youngest ACANTHARIA, for the centrogeneous formation of the
skeleton, the constant development of the earliest radial portions of it in
the middle of the central capsule, forces the nucleus from its normal
central position. The majority of the ACANTHARIA, like most Polycyttaria,
are precocious, the primary nucleus early dividing into numerous small
nuclei (see note A below). Nevertheless there are many exceptions to this
rule in different families, _e.g._, _Stauracantha_, _Xiphacantha_,
_Phatnacantha_, and _Pristacantha_ among the #Acanthometra#, and
_Stauraspis_, _Echinaspis_, _Dodecaspis_, and _Phatnaspis_ among the
#Acanthophracta#. In these instances the primary nucleus remains for a long
time as a simple excentric ellipsoidal or irregularly round body, even in
the fully developed stage, and only at a very late period (sometimes just
before the formation of the spores) divides into many small nuclei. Since
this serotinous division of the nucleus takes place in different genera of
very various groups, it can only be decided by further investigations how
widely it is spread among the ACANTHARIA, and upon what circumstances it is
dependent (see note B). The division of the nucleus appears to be
precocious in the majority of this legion, and a number of small nuclei
appear to be early formed by a peculiar process of budding; in most fully
developed ACANTHARIA these are disposed in one or two layers under the
surface of the central capsule, but if their numbers increase to any
considerable extent, the whole space between the skeletal rods becomes
filled with small nuclei; sometimes these are homogeneous, sometimes
vesicular, 0.002 to 0.012 mm. in diameter; usually they are spherical and
have a small nucleolus (compare Pl. 129, figs. 6-11, and note C).

A. The numerous nuclei, which are to be found in the central capsule of
most mature ACANTHARIA, were first described in my Monograph (1862) as
"spherical, transparent vesicles, provided with a small dark granule" (p.
374, Taf. xv. figs. 2, 5; Taf. xvi. figs. 2, 4; Taf. xxi. fig. 7, &c.).
Their more minute constitution and peculiar origin were first accurately
delineated by R. Hertwig (1879, _loc. cit._, pp. 11-24, Taf. i-iii.).

B. The fact that in a number of ACANTHARIA the nucleus does not divide
early as in the majority of the legion, but only at a later period, was
first observed by R. Hertwig in a species of #Acanthometra# (_Xiphacantha
serrata_), and a species of #Acanthophracta# (_Phatnaspis
{xxxviii}mülleri_ = _Haliommatidium mülleri_) (_loc. cit._, pp. 11 and
27). This serotinous division of the nucleus seems, however, to be rather
widely spread in both sublegions of the ACANTHARIA; I have found, not
only in the forms above mentioned, but also in several others belonging
to different genera, a single large excentric nucleus, even in those
individuals in which the skeleton was fully developed.

C. The peculiar mode of nuclear budding, by which these small nuclei
arise, appears to proceed in the following manner (Pl. 129). The
vesicular primary nucleus, which, in consequence of the centrogeneous
development of the skeleton protrudes as it grows into irregular lobes
(Pl. 129, fig. 9), assumes a peculiar concavo-convex form, sometimes that
of a hood or dish, sometimes that of a kidney or sausage. The convex
surface is apposed to the capsule-membrane, while the concave is turned
towards the central star of the skeleton (fig. 6). There is now formed at
the centre of the convex surface of the strong, doubly-contoured, nuclear
membrane, a flask-shaped invagination with a narrow neck and expanded
base; the membrane now becomes disposed in peculiar folds, which at the
narrow aperture of invagination appear as folds, but on the expanded body
of the flask take the form of concentric rings, laid closely side by side
(Pl. 129, fig. 10). The convex bottom of the flask, which is directed
towards the concave proximal side of the nucleus, becomes again
invaginated by a central conical apophysis of the enlarged nucleolus,
which is situated between them. Usually the nucleolus has already become
flattened into a lentiform shape, and upon its distal face a conical
apophysis has been developed, which is divisible into a darker proximal
and clearer distal portion. The tip of the latter appears to be in direct
connection with the nuclear membrane at the centre of the base of the
flask-shaped invagination (figs. 6, 10). At this stage of development the
nucleus of the ACANTHARIA generally presents the characteristic form of a
hood-shaped, concavo-convex vesicle, whose radial axis is also the axis
of the flask-shaped distal invagination, and of the depressed conical
nucleolus, which lies between the latter and the concave side of the
nucleus. After this peculiar invagination has persisted for some time in
connection with the enlarged nucleolus, both disappear, and then a
remarkable growth of lobular processes takes place on the concave
proximal side of the hood or kidney-shaped nucleus; from four to eight
knobs of unequal size usually appear, and their thickened wall encloses a
variable number of small of nucleoli; these are at first few but
afterwards more numerous (fig. 7). Subsequently these knobs or lobes
become completely separated by constriction from the original central
mass of the nucleus, and appear as so many separate independent
"sausage-shaped bodies" in the hollow central capsule (fig. 8). Each of
the bodies now appears, and at first on its convex aspect, to form a
large number of small nucleoli, which either separate by constriction
from it or become free by its breaking up and lie in numbers in the
central capsule. Finally the buds or lobes of the nucleus break up
entirely into such nucleoli, which are evenly distributed in the central
capsule, and become the nuclei of the swarm-spores (fig. 11). Compare R.
Hertwig, L. N. 33, Taf. i.-iii. pp. 19-25.

69. _The Nucleus of the Monopylea._--The nucleus of the mature forms of the
NASSELLARIA or MONOPYLEA is generally simple or lobate, homogeneous or
vesicular and _excentric_, and appears only to divide into numerous small
nuclei just before the formation of the spores. Nevertheless I have
sometimes, though not often, seen in representatives of very various
families of the MONOPYLEA, the central capsule filled with many small
spherical homogeneous nuclei (Pl. 53, fig. 19). Hence all the families of
this legion appear to be serotinous, their simple primitive nucleus
persisting for a long period. It {xxxix}is commonly placed excentrically,
and most usually in the apical or aboral portion of the central capsule,
either between its apex and the podoconus, or quite excentrically on the
dorsal aspect. The simple nucleus of the NASSELLARIA usually appears to be
vesicular and to possess a somewhat firm membrane, clear contents, and a
rather large, dark coloured nucleolus. In many NASSELLARIA the nucleus is
spherical or ellipsoidal (Pl. 53, fig. 11); whilst in many #Stephoidea# and
#Spyroidea#, where the central capsule is constricted by the sagittal ring
and divided into two symmetrical lateral lobes, the nucleus partakes of the
same mode of growth and appears in the middle of the capsule as a
transversely placed ellipsoid or even as a short cylinder (Pl. 90, figs. 7,
9). The most remarkable modification in the form of the nucleus is to be
found in the multi-articulate #Cyrtoidea#. Here it is usually enclosed in
the cephalis and is spherical, ellipsoidal or spheroidal, often flattened
almost into a disc. If now the central capsule increase greatly in size and
put forth three or four clavate lobes which hang down through the pores of
the cortinar septum into the thorax (or even into the succeeding joints),
the nucleus usually undergoes similar modification, and three or four
finger-like apophyses are developed from its base, which project into the
corresponding lobes of the central capsule (Pl. 59, figs. 4, 12, 13).

The numerous small, spherical, homogeneous nuclei which are to be found
in the central capsules of those NASSELLARIA, which are ripe and about to
develop spores, were described in 1862 in my Monograph, as "numerous,
small, transparent, spherical cells" in the case of various #Cyrtoidea#
(_Arachnocorys_, _Lithomelissa_, _Eucecryphalus_, _Eucyrtidium_, &c.)
(_loc. cit._, pp. 302, 305, 309, 321, &c.), and I find them of the same
form and dimensions, but deeply stained with carmine in many preparations
in the Challenger collection. R. Hertwig has delineated them very
accurately in the case of _Tridictyopus_ (1879, _loc. cit._, p. 84, Taf.
vii. fig. 3). He was also the first to recognise the uninucleate
condition of the NASSELLARIA, which is much more frequently observed than
the serotinous multinucleate condition, and he described very clearly the
peculiar lobed nuclei which arise in #Cyrtoidea#, owing to the protrusion
of the nucleus through the cortinar septum (_loc. cit._, p. 85, Taf.
viii. figs. 3-8).

70. _The Nucleus of the Cannopylea._--The nucleus presents the same
remarkable structures in all species of the PHÆODARIA or CANNOPYLEA which
have been examined, and closely resembles the germinal vesicle of an
amphibian ovum, being a large spherical or spheroidal vesicle with numerous
nucleoli. Its diameter usually amounts to half or two-thirds, sometimes
even three-quarters, that of the central capsule. The vertical main axis of
the latter is also that of the nucleus, which usually lies somewhat nearer
to the aboral pole. The nucleus is generally rather more strongly
compressed in the direction of the main axis than the capsule itself. The
membrane of the vesicular nucleus is thin, but firm, and encloses a clear
or finely granular mass of nuclein. The number and size of the contained
nucleoli are variable even in one and the same species, and stand in
inverse ratio to each other, an obvious result of the gradual process of
division. Commonly {xl}from twenty to fifty roundish or spherical, strongly
refracting nucleoli, are present; more rarely there are several hundred
very small ones. Sometimes the nucleus is penetrated by fine trabeculæ, in
whose meshes lie the nucleoli (Pl. 101, fig. 2). In certain nuclei, which
contained a few large nucleoli, these were of irregular form, probably the
result of amoeboid movements (Pl. 101, fig. 1). In the formation of spores
in the CANNOPYLEA, the nucleus apparently becomes dissolved, and its
numerous nucleoli develop directly into the nuclei or mother-nuclei, which
produce the nuclei of the flagellate spores. Furthermore, many PHÆODARIA
seem to multiply by simple cell-division, since very commonly (especially
in the #Phæocystina# and #Phæoconchia#) two large nuclei (right and left),
may be met with in one central capsule; sometimes also a single large
nucleus, in which a sagittal constriction marks the commencing division of
the capsule (Pl. 101, figs. 2, 36; Pl. 104, fig. 3; Pl. 124, fig. 6, &c.).

The large nucleus of the PHÆODARIA was first described in my Monograph in
1862, in the case of _Aulacantha_ (p. 263), _Aulosphæra_ (p. 359), and
_Coelodendrum_ (p. 361), as a "large, spherical, thin-walled endocyst,"
from 0.1 to 0.2 mm. in diameter. More detailed descriptions, especially
with respect to the behaviour of the nucleoli were given by R. Hertwig in
1879 (L. N. 33, p. 97).

71. _The Endoplasm or Intracapsular Protoplasm._--In all Radiolaria the
intracapsular protoplasm, which, for the sake of brevity, may be termed
"endoplasm," constitutes originally, and especially in the earliest stages,
the only important content of the central capsule, except the nucleus. In
certain SPUMELLARIA and NASSELLARIA, of simple structure and of small
dimensions, this condition persists for a long period, and the endoplasm
then appears as a homogeneous, colourless, turbid or finely granular,
mucous, semi-solid mass, which cannot be distinguished from the ordinary
undifferentiated protoplasm of young cells; no definite structure, and in
particular, no fibrillar network, can be discovered in it even by the use
of the customary reagents. In the great majority of the Radiolaria,
however, this primitive homogeneous condition of the endoplasm is very
transient, and it soon undergoes definite modifications, becoming
differentiated into separate parts or producing new constituent contents.
Such products of the internal protoplasm are in particular hyaline spheres
(vacuoles and alveoles), oil-globules, pigment-bodies, crystals, &c. The
most important of the differentiations which take place in the endoplasm is
that into an internal, granular, _medullary_ substance and an external,
fibrillar, _cortical_ substance; although the various legions behave
somewhat differently in this respect (§§ 77-80).

72. _Intracapsular Hyaline Spheres._--The central capsule of very many
Radiolaria contains in its endoplasm numerous spherical bodies of varying
size, which consist of watery or albuminous fluid, and have previously been
regarded as nuclei, or described as products of the internal protoplasm,
under various names, such as "spherical transparent {xli}vesicles" (see
note A, below), "albumen spheres" (see B), "gelatinous spheres" (see C),
"alveolar cells" (see D), &c. Some of these spheres are perfectly
transparent, structureless and of varying refractive power, producing the
impression of drops of fluid; others contain various formed constituents,
such as oil-globules, fat-granules, pigment-granules, concretions,
crystals, &c. From a morphological point of view they may all be divided
into two categories, membraneless vacuoles and vesicular alveoles. The
_vacuoles_ are simple spherical drops of fluid or of gelatinous material,
devoid of a special envelope, but immediately surrounded by the endoplasm.
The _alveoles_, on the other hand, are true vesicles with a thin spherical
envelope, enclosing a drop of fluid or jelly. This envelope is commonly
very thin, homogeneous, and often scarcely discernible, so that in practice
a sharp line of demarcation cannot be drawn between alveoles and vacuoles;
the former are usually somewhat larger than the latter. The fact is,
nevertheless, certain that the hyaline spheres, which may be isolated on
rupturing the central capsule of many Radiolaria, in certain cases,
particularly in large species, possess a clear, anatomically demonstrable
membrane, whilst in others no such appearance is presented. It may be
assumed that the vesicular alveoles are developed from the drop-like
vacuoles by increase in size, and by the precipitation of a delicate
envelope from the endoplasm. The character common to all these hyaline
spheres, whether vacuoles or alveoles, is found in their aqueous, not
adipose, constitution, and in their clear transparent appearance, which
allows of no structure (the above-mentioned contained bodies excepted)
being recognised. Their refractive power and consistency vary somewhat, and
probably their chemical constitution still more. Sometimes they are
strongly refractive and shining, and sometimes feebly refractive and pale;
their consistency shows all intermediate stages between a thin fluid, which
readily disappears in water, and a firm, insoluble jelly. As regards their
chemical composition (which is probably very variable), the hyaline spheres
may be best divided into two groups, the organic and inorganic. The
_inorganic hyaline spheres_ are simple drops of saline solution without any
carbonaceous constituent; the _organic_, on the other hand, contain a small
quantity of organic matter dissolved in the watery fluid, and may be either
albuminous or gelatinous spheres. The formed contents which are commonly
present are of very various natures, usually small fat-granules, more
rarely larger fat-granules or pigment-granules, sometimes concretions or
crystals. In many groups, especially among the large PHÆODARIA and
#Collodaria#, the numerous hyaline spheres are remarkable for their equal
size and even distribution throughout the endoplasm (Pl. 1, figs. 1, 4; Pl.
104, fig. 2, &c.). In some genera belonging to the Thalassicollida the
alveoles are of enormous size (Pl. 1, figs. 2, 3); they then become
flattened by mutual pressure into polyhedra and distend the central capsule
to unusual dimensions (in _Physematium_ and _Thalassolampe_ 8 to 12 mm.).

A. The "_spherical hyaline vesicles_," which I described in my Monograph
(1862, p. 71) as among the most important and constant contents of the
central capsule, are partly vacuoles, {xlii}partly homogeneous nuclei.
Most recent investigators, Bütschli in particular (1882, L. N. 41), have
pointed out and rightly criticised this confusion. The criticism might,
however, have been more justly expressed by stating that, in the
preparation of my Monograph (1859-1862), I did not make use of modern
methods of demonstrating the nucleus by staining fluids, which were quite
unknown at the time, and only discovered a decade later. In fact, without
the aid of such reagents, it is quite impossible to distinguish between
the various "spherical transparent vesicles," of which those found in the
central capsule of the PHÆODARIA and many monozootic #Collodaria# are
simple vacuoles lying in the endoplasm, whilst, on the other hand, those
of the Polycyttaria and many other Radiolaria are true homogeneous
nuclei. For not only are the general appearance of the small clear
spheres, their refractive power, and regular distribution in the
endoplasm quite similar, but they are also of much the same size, for the
diameter ranges from 0.005 to 0.015 mm., being generally between 0.008
and 0.012 mm. In addition to this there is generally in each hyaline
sphere a dark brightly shining granule, which, in the case of the
vacuole, is simply a fat-granule, whilst in the case of the nucleus, it
is a true nucleolus. The small hyaline spheres in the young uninucleate
capsules of the Polycyttaria are simple vacuoles (Pl. 3, fig. 12), whilst
in the ripe multinucleate capsules they are true nuclei (Pl. 3, figs. 3,
8, 9), and it is quite impossible to discriminate between these two
conditions without the use of reagents. This has been expressly
recognised by R. Hertwig, who has the merit of having been the first to
clearly distinguish, by the aid of staining fluids, between these two
different constituents (1879, L. N. 33, p. 108).

B. The "_albumen spheres_," which were first observed by A. Schneider in
1858 in the common cosmopolitan _Thalassicolla nucleata_ (L. N. 13, p.
40), and which appear to occur in only a few other Thalassicollida, are
distinguished from the ordinary hyaline spheres of about the same size by
their higher refractive power and by certain albuminoid reactions,
especially the coagulation of a membranous envelope under the influence
of certain reagents (see my Monograph, p. 250, and Hertwig, L. N. 26,
1876, p. 46). They often enclose various formed contents, and require
further investigation.

C. The _gelatinous spheres_ of various sizes, found in the endoplasm of
the Radiolaria, agree in their reactions (especially in staining by
certain reagents) with the common extracapsular jelly of the calymma, and
are hence distinguishable both from the true (coagulable) "albumen
sphere," and from the ordinary watery vacuoles.

D. The _alveoles_, which are only accurately known in the case of certain
large monozootic #Collodaria#, but which also seem to occur in the
central capsule of other remarkably large Radiolaria, were described in
my Monograph in the case of _Thalassolampe margarodes_ and _Physematium
mülleri_, under the name "intracapsular alveolar cells" (1862, pp. 77,
254, 257). They are not, however, true nucleated cells, and the body
described as a nucleus is not such in reality. Nevertheless these large
hyaline spheres do possess a special envelope, as I have recently
convinced myself by the examination of ruptured central capsules of
_Thalassolampe maxima_, _Thalassopila cladococcus_, and _Physematium
atlanticum_ (Pl. 1, figs. 2, 3). The central capsule of these
#Collodaria# becomes distended to most unusual dimensions (2 to 12 mm. in
diameter) by the great development of these large hyaline vesicles, each
of which measure from 0.1 to 0.5 mm. in diameter.

73. _The Intracapsular Fat-Globules._--Fat is present in the central
capsule of all Radiolaria in larger or smaller quantities, and generally
appears in the form of very {xliii}numerous, small, spherical granules,
which are either distributed evenly in the endoplasm (as an emulsion) or
enclosed in the vacuoles; the latter, in particular, is the case in most
PHÆODARIA, perhaps generally. In this group each vacuole contains as a rule
a single dark, shining fat-granule, and sometimes also an irregular bunch
composed of from two to five or more granules. In addition to these small
fat-granules (_granula adiposa_) which are always present, the central
capsule of many Radiolaria contains also larger fat-globules (_globuli
adiposi_). These appear to be generally wanting in the PHÆODARIA, and are
on the whole rare in the ACANTHARIA; whilst, on the contrary, they are very
common in the NASSELLARIA and SPUMELLARIA. The Polycyttaria or social
Radiolaria are as a rule distinguished by the possession of a single large
central oil-globule, which lies in the centre of the central capsule, and
is on an average about one-third of it in diameter (Pl. 3, figs. 4, 5).
This is absent, however, in those young capsules of the Polycyttaria in
which the primary nucleus is centrally situated (Pl. 3, fig. 12). Those
species of Polycyttaria whose central capsule reaches a considerable size,
often enclose numerous oil-globules, and in _Collophidium_ (species of
_Collozoum_ with an elongated cylindrical capsule, Pl. 3, figs. 1, 3) the
axis of each capsule is occupied by a row of numerous oil-globules. In the
monozootic SPUMELLARIA, in which the nucleus is always centrally situated,
the large oil-globules are, of course, excentric, being in apposition to
the inner surface of the capsule-membrane (Pl. 1, fig. 3; Pl. 2, figs. 2,
5). In the #Discoidea# the oil-globules, which are often present in large
numbers, form elegant concentric rings around the central nucleus, and in
those species with segmented arms, there are one or more transverse rows in
each segment (Pl. 43, fig. 15). In the NASSELLARIA the number and
distribution of the oil-globules are dependent upon the form of the central
capsule. When this is simple, without lobes, and ovoid or conical, they
generally lie in its aboral half above the podoconus (Pl. 51, figs. 5, 13;
Pl. 97, fig. 1). When, on the contrary, the basal portion of the capsule
sends out three or four dependent processes (as in the majority of the
#Cyrtoidea#), a large globule may generally be seen in the swollen distal
part of each conical or ovoid lobe (Pl. 53, fig. 19; Pl. 60, figs. 4-7). In
many #Stephoidea# and #Spyroidea#, whose central capsule is separated into
two lateral portions by the constriction corresponding to the sagittal
ring, each of these contains either a single large globule or a group of
small ones (Pl. 90, figs. 7, 10). These oil-globules are usually colourless
and highly refractive; rarely they are yellow or brown, sometimes
rose-coloured, or an intense blood-red (_e.g._, in _Thalassophysa
sanguinolenta_) or even orange (in _Physematium mülleri_). In many
SPUMELLARIA, and particularly in the Polycyttaria, an albuminous substratum
may be recognised in them, which is sometimes disposed in layers, and after
extraction of the fat presents the appearance of a laminated sphere. The
physiological significance of the oil-globules is twofold; in the first
place they tend to diminish the specific gravity of the organism; in the
second they may be utilised as a reserve store {xliv}of nutriment. In the
latter respect they are of special importance in the process of
spore-formation, each flagellate spore usually containing a fat-granule.

74. _The Intracapsular Pigment-Bodies._--In the majority of Radiolaria when
observed alive, the central capsule is coloured, only in the minority is it
colourless. The colour is never diffuse, but always due to the formation
of definite pigment granules or vesicles, which are sometimes distributed
evenly throughout the endoplasm, sometimes aggregated in the central or
peripheral regions. Their form may be either spherical, irregularly
rounded, or polyhedral. They vary much in dimensions, but in most cases
are immeasurably small, and appear under a high magnifying power as fine
dust; occasionally, however, their diameter may amount to from 0.001 to
0.005 or more. The chemical constitution of the intracapsular pigment is
unknown in most Radiolaria, and is probably very various. In many instances
the pigment-granules consist of fat, in others not. The commonest colours
are yellow, red, and brown; violet and blue are rare, and green still
rarer. Sometimes a definite tone of colour prevails throughout a whole
group, and may then be attributed to inheritance, _e.g._, red is found in
most #Sphæroidea#, and blue in the Polycyttaria (see note A). One colour is
almost always constant in the members of the same species. True
pigment-cells, belonging to the Radiolarian organism, do not occur within
the central capsule. The peculiar yellow cells which are found in the
central capsule of many ACANTHARIA are symbiotic xanthellæ (see § 76).

A. The number of Radiolaria whose pigment has been examined in the living
state, is too small to allow of any general conclusions being drawn.
Regarding the different colours known, see my Monograph, L. N. 16, p. 76.

75. _The Intracapsular Crystals._--The crystals found in the central
capsule of many Radiolaria may be divided into two groups, of very
different significance; small crystals, which are very widely distributed,
and large crystals, which occur in only a few genera. The _small crystals_
may also be termed "spore-crystals," since each swarm-spore often contains
such a crystal. They are rod-like or spindle-shaped, and consist of an
organic substance which probably serves as a reserve of nutriment for the
developing spores. Such spore-crystals have been observed in numerous
SPUMELLARIA and ACANTHARIA belonging to various families, and are probably
present throughout the two legions which make up the Porulosa. On the other
hand, they have not been noticed in the Osculosa (NASSELLARIA and
PHÆODARIA), the few swarm-spores belonging to these groups which have been
observed not exhibiting any crystals. The _large crystals_, which occur in
small numbers in the endoplasm, have hitherto only been observed in a few
species of SPUMELLARIA, belonging to the Polycyttaria. They were first
noticed in the common _Collosphæra huxleyi_, and regarded as coelestin.
They are also found in the central capsule of many other Collosphærida,
_e.g._, _Buccinosphæra_ (Pl. 5, figs. 11, 12). Crystal-masses,
crystal-sheaves, or spherical masses of radiating acicular crystals are
enclosed in {xlv}the vacuoles or "albumen globules" of _Thalassicola
nucleata_ and other Thalassicollida, as well as in the central capsule of
_Coelographis_ and some other PHÆODARIA (Pl. 127, figs. 4-7). All these
large crystals are probably to be regarded as excretory products.

75A. _The Intracapsular Concrements._--Concretions, either mineral or
organic, of varying form and constitution, are to be found in the endoplasm
of Radiolaria belonging to very different families. They are most abundant
and multiform in _Thalassicolla nucleata_, being usually circular or
elliptical discs, which are concentrically laminated and highly refractive,
resembling starch-grains. Among them twin forms may frequently be observed,
as though the concrements were in process of division (see note A). Similar
amyloid concretions are to be seen in the central capsule of different
SPUMELLARIA and NASSELLARIA, _e.g._, in _Cephalospyris triangulata_ (Pl.
96, fig. 28). Violin-shaped, highly refractive concrements have been
observed in the central capsule of numerous SPUMELLARIA, NASSELLARIA, and
ACANTHARIA, _e.g._, _Thalassosphæra_, _Spongosphæra_, _Plegmosphæra_,
_Cyrtocalpis_, _Peripyramis_, _Botryocella_, &c. (see note B). The chemical
constitution of these concrements is insufficiently known.

A. The amyloid concretions of _Thalassicolla nucleata_ have been
described in detail in my Monograph (pp. 80, 250, Taf. iii. figs. 2, 3),
and by R. Hertwig in the Histologie der Radiolarien (1876, p. 47, Taf.
iii. figs. 9-13).

B. The violin-shaped concretions of _Thalassosphæra bifurca_ have been
figured in my Monograph (pp. 80, 261, Taf. xii. fig. 1).

76. _The Intracapsular Xanthellæ._--The xanthellæ, zooxanthellæ, or
symbiotic "yellow cells" are found within the central capsule only in the
ACANTHARIA, whilst in other Radiolaria they only occur in the
extracapsulum. They are most frequent in the #Acanthometra#, rarer in the
#Acanthophracta#, but even in the former they are often wanting. Their
number is very variable, but usually small, from ten to thirty in one
capsule. They lie for the most part immediately below the capsule membrane,
in the cortical layer of the endoplasm. The form of the yellow cells is
either spherical or ellipsoidal, often also spheroidal or even lentiform.
The diameter varies from 0.01 to 0.03 mm. They possess a distinct membrane
and an excentric nucleus, and contain numerous yellow pigment-granules in
the endoplasm. This yellow pigment dissolves in mineral acids to form a
green fluid, and in other respects also behaves somewhat differently from
the yellow pigment in the extracapsular yellow cells of the SPUMELLARIA and
NASSELLARIA. In both cases, however, the xanthellæ are not integral
portions of the organism, but unicellular algae, living as parasites or
symbiontes in the body.

A. The yellow cells in the central capsule of the ACANTHARIA were first
observed by Joh. Müller (L. N. 12, pp. 14, 47). In my Monograph I
described them at greater length, and indicated their differences from
the extracapsular yellow cells of other Radiolaria (L. N. 16, pp. 77,
86). Since then, R. Hertwig has demonstrated their cellular nature (L. N.
33, pp. 12, 113), and still more recently {xlvi}Brandt has given further
accurate information regarding their occurrence, constitution, and
physiological significance (L. N. 39, ii. Art., p. 235, figs. 62-73).

77. _The Endoplasm of the Peripylea._--The intracapsular protoplasm of the
SPUMELLARIA or PERIPYLEA is usually distinguished by a more or less
complete radial arrangement, which does not occur in the same form in other
Radiolaria; it may be regarded as characteristic of this legion, for it
probably occurs in all the species at some period of life or other, and
stands in a direct causal relationship with the typical structure of the
capsule-membrane in all the "PERIPYLEA" (see note A). For as this is
commonly perforated by very numerous pores distributed at equal intervals
over the whole surface of the capsule, and since a communication between
the intra- and extracapsular sarcode takes place through these, the radiate
structure of the endoplasm may be readily explained as due to the influence
of radial currents which take place continuously or intermittently in the
endoplasm. This radiate structure is most obvious when the endoplasm
contains no secondary products or only an insignificant amount of these,
and thus appears colourless and almost homogeneous, or only finely
granular. Under these circumstances, an optical section of the central
capsule usually reveals a distinct radial striation; numerous narrow,
straight, dark streaks alternating regularly with still narrower clear
ones; the latter consist of homogeneous, the former of more or less
granular protoplasm (Pl. 20, fig. 1_a_). Often there may be distinguished
in each darker streak a single straight row of strongly refracting (fat?)
granules, sometimes several such rows. Occasionally the whole endoplasm
becomes divided up into a number of large "radial wedges," club-shaped,
conical or pyramidal masses of granular protoplasm, separated by clear
divisions of hyaline plasma (_e.g._, in _Actissa radiata_, p. 14, where in
the optical section of the central capsule, between the membrane and the
nucleus, twenty-five dark radial wedges of equal size were separated by
thick clear partitions of hyaline protoplasm). In the majority of the
SPUMELLARIA this radial striation is partially or entirely concealed by the
formation of pigment or of other products. Very often it is only visible in
the cortical layer, which lies immediately below the capsule-membrane (Pl.
1, figs. 1, 3). The remarkable "centripetal cones" which characterise the
Thalassicollid genus _Physematium_, and were formerly described as
"centripetal cell-groups," are probably a special development of these
cortical radial wedges; they are conical cortical bodies, regularly
distributed on the inner surface of the membrane of the central capsule,
and disposed with the apex turned towards the centre (see note B). More
rarely than in the cortical layer, a similar radial structure is to be
found in the innermost medullary layer immediately surrounding the nucleus.
Here the endoplasm sometimes breaks up into fine radial threads, which are
anatomically separable and hang down from the free nucleus as thin
processes (see note C). In some cases it is also possible to isolate radial
rods from the cortical layer of teased out central capsules.

{xlvii}A. The radial structure of the endoplasm was first described in my
Monograph (1862, p. 74), though R. Hertwig (1879, p. 112) was the first
to indicate its typical significance in the case of the PERIPYLEA, and to
demonstrate its causal relation with the radial currents in the central
capsule of this legion. More recent investigations have led me to the
conviction that this phenomenon is more widespread, and often more
strongly developed, than was formerly imagined, and that it is probably
one of the typical characters of all SPUMELLARIA (at least of the
Monozoa).

B. The centripetal cones of _Physematium_, which have hitherto been known
only in these colossal Thalassosphærida, were fully described in my
Monograph under the name "conical centripetal cell-groups"; by their
first discoverer, A. Schneider (L. N. 13), they were termed "nests," and
compared with the "nests" (central capsules) of the Polycyttaria. In the
_Physematium mülleri_ of the Mediterranean (hitherto only observed by
Schneider and myself at Messina) it appeared as though each centripetal
cone were composed of a group of from three to nine (usually four or
five) slender wedge-shaped cells, whose common centripetal apex was
produced into a radial thread of sarcode (L. N. 16, p. 258, Taf. iii.
fig. 7). Since then (1866) I have observed at Lanzerote, in the Canary
Islands, a nearly related form, which I take to be _Physematium
atlanticum_, Meyen. In this, however, the "centripetal cell-groups" were
wanting, and the whole cortical layer of the endoplasm was cleft into
numerous radial portions, each enclosing a nucleus (probably the
mother-cells of flagellate spores, see p. 35).

C. The radial fibres of the medullary endoplasm which cling to an
extracted nucleus have been observed by Hertwig in certain #Sphæroidea#
(_Diplosphæra_, _Arachnosphæra_) (L. N. 33, p. 40).

78. _The Endoplasm of the Actipylea._--The intracapsular protoplasm of the
ACANTHARIA or ACTIPYLEA is often distinguished by a partial or complete
radial arrangement like that of the PERIPYLEA, but differing in the number,
size, form, and distribution of the radial portions into which the
endoplasm is differentiated. For since the pores of the capsule membrane
are distributed at equal distances all over the surface in the SPUMELLARIA,
whilst in the ACANTHARIA they are arranged in definite groups, and since
the number and arrangement of the pores has a direct influence upon the
internal currents of the endoplasm, it follows that the radial structure in
the latter legion must be very different from that in the former. In
addition to this there must not be forgotten the important influence which
the early centrogenous formation of the skeletal rods exercises upon the
disposition and growth of the intracapsular structures. Hence the endoplasm
of the ACANTHARIA does not separate into innumerable thin, closely packed
radial wedges or cortical radial rods, but into a small number of large
pyramidal portions between which run the radially disposed heterogeneous
portions of the contents of the capsule, viz., the radial bars of acanthin
and the peculiar intracapsular "axial threads." As a direct consequence of
the regular disposition of these heterogeneous radial portions, which is
often characteristic of the various families of the ACANTHARIA, a
corresponding differentiation of the endoplasm is brought about; it divides
into a number of conical or pyramidal portions (radial pyramids), whose
bases rest upon the capsule-membrane and whose apices are directed towards
the centre of {xlviii}the capsule (the central star of the skeleton).
These radial pyramids are, however, but rarely visible, being usually more
or less concealed by a dark pigment.

The differentiations of the endoplasm in the central capsule of the
ACTIPYLEA have been but little investigated, but they appear to vary
somewhat in the different groups of this legion. In all ACANTHARIA in
which the twenty radial bars are regularly arranged according to the
Müllerian law (see p. 717) and in which axial threads constant in number
and disposition run between them from the central star to the
capsule-membrane, it obviously follows that the endoplasm must be divided
into more or less distinct radial pyramids, and this must the case
whether these take the form of continuous tracts or of actually separable
portions. The regular polygonal figures, often seen on the surface of the
central capsule (with special distinctness in _Acanthometron elasticum_
and _Acanthometron pellucidum_) separated by a network of granular
threads, are the bases of such radial pyramids (see Hertwig, L. N. 43, p.
12, Taf. i. figs. 1-7).

79. _The Endoplasm of the Monopylea._--The intracapsular protoplasm of the
NASSELLARIA or MONOPYLEA is distinguished from that of any of the other
three legions by the development of a quite peculiar fibrillar structure,
the axial "pseudopodial cone," which may shortly be termed the "podoconus"
(foot-cone). Since this is in direct correlation with the peculiar
structure of the capsular opening, the large "porochora," which is situated
at the basal pole of the main axis, it is quite as characteristic of the
legion as the latter itself (see note A). The podoconus is primitively a
vertical regular cone whose circular base occupies the horizontal porochora
or "basal porous area" of the central capsule, while its vertical axis
coincides with that of the latter. The apex of the cone, usually somewhat
rounded off, is therefore directed towards the aboral or apical pole of the
central capsule and separated from it by a larger or smaller interval. In
this interval the nucleus originally lies (as in Pl. 51, fig. 13; Pl. 98,
fig. 13); but it is usually displaced subsequently and lies excentrically.
The cone is of very variable height; on an average its vertical height is
about equal to the diameter of its horizontal base; these dimensions are,
however, dependent upon the form of the central capsule; the height being
greater in slender ovoid or conical capsules, and less in depressed
sphæroidal or discoidal ones, than the diameter of the base. The podoconus
consists of differentiated endoplasm, which becomes more deeply stained by
carmine and offers greater resistance to solvents than the surrounding
finely granular protoplasm. The apex, especially, becomes very intensely
stained. It always exhibits a very characteristic fine but distinct
striation, numerous straight radial lines diverging from the apex of the
cone towards the base. The number of these striæ appears to correspond with
that of the vertical rods in the porochora, and each of these latter stands
apparently in direct communication with the basal end of an apical stria (§
59). These threads are probably differentiated constant contractile threads
of endoplasm, or even myophanes, comparable with the contractile cortical
threads of the CANNOPYLEA and the permanent axial threads of the ACTIPYLEA.
The numerous modifications, {xlix}undergone by the form and contents of
the central capsule in the different groups of MONOPYLEA, especially those
due to the formation of the skeleton, are not without influence upon the
podoconus. The most important divergencies from the above described primary
form are the following:--(1) The vertical axial cone becomes oblique, its
axis inclining in the sagittal plane and approaching either the dorsal or
the ventral wall of the capsule; the cause of this appears to be usually
the excentric development of the growing nucleus or the formation of a
large oil-globule. (2) The smooth mantle of the podoconus becomes divided
by three longitudinal furrows into three equal prominent ridges, which
correspond to three circular lobes in the porochora; the cause of this
basal triradial lobular formation lies probably in the triradial
development of the skeleton in many NASSELLARIA or in the cortinar
structure of the collar septum. (3) The simple podoconus splits into three
or four elongated lobes, which eventually become almost completely
separated and correspond to the lobes of the central capsule, in the axial
wall of which they lie as longitudinally striated bands. The behaviour of
these bands justifies the hypothesis that the podoconus is a muscular
differentiated portion of the endoplasm and is composed of myophane
fibrillæ, whose contraction determines the opening of the central capsule.

A. The podoconus of the MONOPYLEA was first described by R. Hertwig in
1879, and recognised as a characteristic component of the central capsule
in the most various groups of this legion (in #Plectoidea#, #Stephoidea#,
#Spyroidea#, and #Cyrtoidea#; see his figures, _loc. cit._, Taf. vii.,
viii., and the description, pp. 71, 73, 83, 106). Hertwig called it the
"pseudopodial cone," and regarded it as a conical process of the
capsule-membrane, which is developed from this latter and projects from
the porous area into the interior of the central capsule; "it is
penetrated by fine canals which arise at the apex of the cone, diverge
towards the base, and terminate there in the rods of the pseudopodial
area. The intracapsular protoplasm penetrates at the apex of the
pseudopodial cone into its fine canals, runs along them and emerges from
the rods of the porous area in the form of slender threads" (_loc. cit._,
p. 19). I cannot agree with this view of Hertwig, although I have been
able to confirm the accuracy of his description by my own observations
upon numerous excellently stained and preserved preparations in the
Challenger collection. As I have proved by numerous teased out
preparations, and as Hertwig himself correctly states, "the cone is more
readily detached from the membrane than from the protoplasm, when the
capsule is teased" (_loc. cit._, p. 73). Hence I regard the podoconus not
as a differentiated portion of the capsule-membrane but as endoplasm, and
believe that it is composed of myophanes or "contractile muscular
fibrils" in the same manner as the cortical layer of the CANNOPYLEA.
Probably the contraction of these fibrils serves to raise the opercular
rods and hence to allow the exit of the endoplasm through the pores which
lie between these opercular rhabdillae (compare § 59).

80. _The Endoplasm of the Cannopylea._--The intracapsular protoplasm of the
PHÆODARIA or CANNOPYLEA is distinguished from that of the other three
legions by several characteristic peculiarities, which are very important,
since they stand in causal relation to the typical structure of the
capsule-membrane and in particular of its {l}remarkable aperture. In the
case of many and perhaps of all PHÆODARIA the endoplasm is differentiated
into a granular medullary and a thin fibrillar cortical layer, the former
of which usually encloses numerous small vacuoles, while the latter
contains muscular fibrillæ. In the voluminous central capsule of large
PHÆODARIA the whole cortical layer of the endoplasm, which lies immediately
below the delicate inner capsule-membrane, sometimes appears delicately and
regularly striated, and most distinctly so under the apertures, towards the
centre of each of which the dark striæ are radially directed (see note A,
below). These striæ are probably contractile muscular fibrillæ; or
"myophanes," by whose contraction the openings are voluntarily widened. In
the Tripylea this fibrillar star is much more strongly developed under the
astropyle (the main opening) than under the parapylæ (or accessory
openings); and probably the peculiar radial structure of the operculum of
the former is due to the stronger development of these radial fibrils
(being their impression). In many PHÆODARIA, indeed, the fine myophane
fibrils are only visible under the apertures, whilst in others they form a
continuous fibrillar cortical layer on the whole inner surface of the inner
capsule-membrane; the fine fibrillæ run meridionally from one pole of the
main axis to the other; perhaps the whole central capsule may change its
form in consequence of their contractions. The medullary portion of the
endoplasm, which lies below this thin cortical layer, is usually finely
granular in the PHÆODARIA, and permeated by numerous spherical vacuoles,
which are noteworthy from their equal size and regular distribution. Each
clear vacuole usually contains a dark shining fat-granule, more rarely a
group of such granules (see note B). Compare § 60, and Pl. 101, figs. 1-3;
Pl. 104, figs. 1, 2; Pl. 111, fig. 2; Pl. 128, fig. 2, &c.

A. The fine fibrillæ in the cortical layer of the endoplasm were first
described by Hertwig in 1879 (L. N. 33, p. 98, Taf x. figs. 6-10). He
found them, however, only below the three openings in the capsule of the
Tripylea, where they form three stellate groups of fibrils. I find them
very clearly shown, and with especial distinctness, under the astropyle
in most PHÆODARIA of which I have had the opportunity of examining
well-stained and preserved central capsules. In many cases, also, the
striation is not confined to the apertures, but spreads over the whole
cortical layer. Perhaps this constitutes in all PHÆODARIA a thin
myophane-sheet, whose contractile fibrils run from one pole of the main
axis to the other and cause by their contraction changes in the form of
the spheroidal central capsule.

B. The granular medullary portion of the endoplasm of the PHÆODARIA, with
its numerous clear spherical vacuoles, was first described in my
Monograph (1862), in the case of _Aulacantha_ (p. 263), _Aulosphæra_ (p.
359), and _Coelodendrum_ (p. 361) as a "finely granular, mucous substance
(intracapsular sarcode), packed more or less closely with clear spherical
vesicles from 0.005 to 0.015 mm. in diameter, each of which contains one
or two, rarely three, dark shining granules." That these clear spheres
are true vacuoles was first clearly proved by Hertwig (L. N. 33, p. 98).
As a rule all the vacuoles of the same central capsule are of equal size
(generally from 0.008 to 0.012 mm. in diameter), and are distributed at
equal intervals throughout the finely granular endoplasm.

{li}CHAPTER III.--THE EXTRACAPSULUM.

(§§ 81-100).

81. _The Components of the Extracapsulum._--The extracapsulum or
extracapsular malacoma, under which name are included all those parts of
the soft body which lie outside the central capsule, consists of the
following constant, and important constituents:--(1) The _calymma_ or
extracapsular jelly-veil; (2) the _sarcomatrix_ or layer of exoplasm
immediately surrounding the membrane of the central capsule; (3) the
_sarcodictyum_ or network of exoplasm, covering the surface of the calymma;
(4) the _pseudopodia_ or radial fibres of exoplasm, which may again be
subdivided into intracalymmar pseudopodia, uniting the sarcomatrix and
sarcodictyum, and extracalymmar pseudopodia, radiating freely into the
water outside the calymma.

82. _The Calymma._--The calymma or extracapsular jelly-veil of the
Radiolaria is always the most voluminous portion of the extracapsulum, and
in spite of its simple structureless constitution is of great morphological
and physiological importance. In all Radiolaria this gelatinous mantle
completely surrounds the central capsule, but is separated from its outer
surface by a continuous, though thin, layer of exoplasm, the sarcomatrix.
The pseudopodia radiating from the latter pierce the calymma, form the
sarcodictyum at its surface, and radiate from its nodal points freely into
the surrounding water. The calymma is rarely visible in living freshly
captured Radiolaria, examined in sea-water, for its gelatinous substance is
perfectly hyaline, colourless and pellucid, and possesses the same
refractive index as sea-water; but when the object is removed from this
fluid and transferred to carmine solution or some other colouring matter,
the extent and figure of the calymma become apparent, for the staining
fluid does not at first penetrate into the gelatinous material. When this
has taken place, however (after a longer or shorter time), and the
gelatinous material has become coloured, its form and size may be observed
by the converse experiment; the object is transferred once more to water
and the outlines of the calymma become as clear as those of the central
capsule. The same is the case with dead specimens in which the sticky
surface of the calymma has become covered with dust.

The jelly-veil of the Radiolaria was recognised even by the earliest
observers of the group, Meyen (1834), and Huxley (1851), and compared
with that of the Palmellaria; the former noticed it in _Physematium_ and
_Sphærozoum_ (L. N. 1, p. 283), and the latter in _Thalassicolla_ and
_Collosphæra_ (L. N. 5, p. 433). In all these SPUMELLARIA, both in the
monozootic _Thalassicolla_ and in the polyzootic _Sphærozoum_ and
_Collosphæra_, the calymma is very voluminous and filled with large
alveoli. Meyen called them "muco-gelatinous masses, in the interior of
which are contained small equal-sized vesicles"; Huxley likewise found
clear vesicles in the jelly and compared them with Dujardin's vacuoles.
Johannes Müller observed the jelly-veil in many different Radiolaria, in
particular in the #Acanthometra#, first discovered by him, but
erroneously believed that it only originated {lii}after death by
liquefaction of the sarcode (L. N. 12, p. 6). This mistake is, however,
easy to understand, since in living Radiolaria the calymma is usually
invisible on account of its perfect transparency, whilst in dead
specimens it is usually quite distinct on account of the dust clinging to
its adhesive surface. I myself believed that the formation of the
voluminous hyaline jelly-veil was only partially due to liquefaction
after death, but that it was to some extent present in the living
organism and that it might vanish and subsequently reappear by means of
imbibition (L. N. 16, pp. 109, 110). R. Hertwig was the first to
demonstrate, in 1879, that the jelly-veil is constantly present in living
Radiolaria, that it forms the basis of the extracapsular malacoma and
surrounds the central capsule as a second protective sheath (L. N. 33, p.
114).

83. _The Structure of the Calymma._--The extracapsular jelly-veil appears
structureless in most Radiolaria, inasmuch as it represents a homogeneous
pellucid excretion of the exoplasm and contains neither fibres nor other
formed structures. In some groups, however, definite structural characters
become secondarily developed. The most common and striking of these is the
formation of alveoles, which takes place in the extracapsulum (see § 86).
In consequence of this the calymma assumes a remarkable frothy consistency
and appears to be composed of large, clear, thin-walled vesicles; this is
especially the case in the #Collodaria# (#Colloidea#, Pls. 1, 3, and
#Beloidea#, Pls. 2, 4), and in many large PHÆODARIA, especially among the
#Phæocystina# (Phæodinida and Cannorrhaphida, Pl. 101, and Aulacanthida,
Pls. 102-104). More rarely the calymma is not permeated by vacuoles, but
there appear in it fine striæ parallel to the surface as though it were
composed of thin concentric laminæ like an onion; perhaps these are the
expressions of a different quantity of water in the various layers. In the
calymma of many Radiolaria thin, straight, radial lines are to be seen,
which are probably pseudopodia, and not to be attributed to any structural
modification, or they may be slender canals which serve for the exit of the
pseudopodia. On the outer surface of the calymma of different Radiolaria,
and especially in the ACANTHARIA, a peculiar network of fibres is to be
found, composed of polygonal meshes, like elastic fibres, probably due to a
local thickening of the jelly. These polygonal meshes are often very
regularly distributed between the radial spines of the #Acanthometra#, and
stand in a definite relation to them. The fibres which form the meshes are
often rather strong, resembling elastic fibres, as above-mentioned, and
either simple or composed of bundles of very fine fibrillæ (L. N. 33, p.
15, Taf. i. fig. 1, Taf. ii. fig. 4).

84. _The Consistency of the Calymma._--The gelatinous material of which the
calymma of the Radiolaria consists is a pellucid mass, rich in water and
usually quite hyaline and structureless; its consistency is very variable.
In the majority of the Radiolaria it may perhaps be about equal to that of
the jelly which composes the umbrella of most Medusæ; but as in these
latter it may vary between very wide extremes, constituting on the one hand
a very soft jelly-mantle, offering but little {liii}resistance to
mechanical influences and almost disintegrating under the eyes of the
observer, and on the other hand forming a firm gelatinous shell, comparable
to cartilage in hardness, elasticity, and power of mechanical resistance.
In many Radiolaria of large dimensions with an alveolar calymma (especially
in numerous #Collodaria# and PHÆODARIA) this may be split by means of
dissecting needles and the central capsule extracted like the stone from a
cherry, and then it is easy to ascertain that the firmness and elasticity
of this jelly-veil are not less than those of a cherry. The different
degrees of consistency in the various Radiolaria may be dependent either
upon the relative amount of water which they contain, or upon qualitative
or quantitative variations in the organic substance of which the jelly
consists. Great importance is to be attached to the considerable
consistency of the calymma, because it furnishes the indispensable
groundwork for the deposition of many parts of the skeleton and
particularly of the lattice-shells.

85. _The Primary and Secondary Calymma._--In most Radiolaria the external
form and volume of the calymma are different at different stages of growth,
and this difference is mainly dependent upon the development of the
skeleton. Hence it is advisable to distinguish in general the primary from
the secondary calymma. The _primary calymma_ is in the great majority of
Radiolaria a perfect sphere, in the middle of which lies the concentric
central capsule; on the surface of this gelatinous plate the primary
spherical lattice-shell is secreted in most SPUMELLARIA and
#Acanthophracta#, as well as in those PHÆODARIA which possess a spherical
shell; in the remaining PHÆODARIA also and in the NASSELLARIA, where the
lattice-shell is not spherical but monaxon, it is secreted on the surface
of the primary calymma. This takes place at a definite time, very important
in the development of the Radiolarian, which for the sake of brevity we
shall term the "_lorication-period_." Since the firm surface of the primary
calymma furnishes the necessary foundation for the deposition of the
primary lattice-shell, it is of the greatest mechanical significance in all
shell-bearing Radiolaria. The _secondary calymma_ arises only after the
lorication-period by further growth of the primitive jelly-mantle and in
the fully developed Radiolarian usually encloses wholly or partially the
external parts of the skeleton, in consequence of which it assumes the most
various forms. Very often the secondary calymma is polyhedral, being
stretched between the radial spines of the skeleton, the distal ends of the
latter then forming the fixed points of the gelatinous polyhedron.

86. _The Extracapsular Vacuoles and Alveoles._--The calymma of the
Radiolaria usually appears completely homogeneous and hyaline without any
structure; sometimes it encloses numerous clear vesicles, vacuoles or
alveoles, and then assumes a frothy appearance, the expression of a more or
less distinct alveolar structure. {liv}The clear vesicles to which this is
due are either spherical, or polyhedral from mutual pressure, and like the
similar ones in the central capsule may be divided into membraneless
vacuoles and vesicular alveoles. The _vacuoles_ are simple drops of fluid,
without a special envelope, and immediately surrounded by the gelatinous
substance of the calymma, in which they appear as simple cavities. The
_alveoles_ on the contrary are true vesicles, with a thin envelope, which
encloses a drop of fluid or a globule of jelly; in the latter case its
contents are different in refracting power and amount of contained water
from the substance of the surrounding calymma. A sharp boundary between the
membraneless vacuoles and the vesicular alveoles cannot be drawn in the
case of the extracapsular hyaline spheres any more than in the
intracapsular; the envelope of the alveoles is sometimes very distinct and
even anatomically separable, whilst at other times it is very thin and
scarcely recognisable; it may occasionally arise and disappear within a
very short time (see note A). There is no doubt that in the calymma as in
the central capsule the vesicular alveoles are secondary products, which
have arisen from the vacuoles by the secretion of an enveloping membrane.
This membrane is either a delicate sheath of exoplasm, or a firmer and more
resistant skin, distinct from the exoplasm, and probably an excretion from
it (_e.g._, Pl. 4, figs. 2, 3). In many cases the outer surface even of the
vacuoles is covered by a network of pseudopodia, which form a sarcoplegma
similar to a fenestrated alveolar membrane. The colourless pellucid fluid
in the vacuoles and alveoles is usually simple sea-water, more rarely it
contains a small quantity of albumen ("albumen-spheres") or jelly
("gelatinous spheres"). The size of these spheres is very variable. Quite
small vacuoles may be found in the calymma of many Radiolaria. Large
vacuoles, on the other hand, producing the appearance of an alveolar
structure, are confined to but few groups, to a part of the SPUMELLARIA
(#Colloidea#, #Beloidea#, and a few #Sphæroidea#), and to the #Phæocystina#
(PHÆODARIA with incomplete skeleton); besides they occur only rarely in
individual genera, _e.g._, _Nassella_ among the skeletonless NASSELLARIA.
Since the volume of the calymma is much increased by the development of
vacuoles, and the power of mechanical resistance is at the same time much
increased, the fact is explained that the vacuoles occur mainly in
Radiolaria which have no skeleton or only an incomplete one (see note B).
Among the monozootic #Collodaria# the alveolar structure is especially well
developed in the following genera; _Thalassicolla_ (Pl. 1, figs. 4, 5),
_Thalassophysa_, _Thalassoplancta_, _Lampoxanthium_ (Pl. 2, figs. 1, 2);
among the PHÆODARIA in most genera of the Phæodinida, Cannorrhaphida and
Aulacanthida (Pls. 101-104), and probably also in other voluminous
PHÆODARIA (_e.g._, #Phæosphæria#). The alveoles or vacuoles in the calymma
of these large Radiolaria lie usually in several layers, one above another,
and increase in size from within outwards. The Polycyttaria or social
Radiolaria (the three families Collozoida, Sphærozoida and Collosphærida)
without exception have an alveolar structure, and the special form of
{lv}their colonies or coenobia is to a great extent determined by the
development, number, size and arrangement of the alveoles in their calymma
(compare Pls. 3-8). In these cases there is not unfrequently developed a
large central alveole (see note C) whose thickened wall encloses a globe of
jelly and serves as the central support of the whole colony (Pl. 5, fig.
1). Still more striking, however, is the arrangement of certain
Polycyttaria, where each individual of the colony (or each central capsule
with its calymma) is enclosed in a large alveole, whose firm wall often
attains considerable thickness (Pl. 4, figs. 2, 3). The whole colony then
appears as an aggregate of numerous cells, each of which possesses two
envelopes, the inner central capsule and the outer alveolar membrane;
between these lies in the Collosphærida the siliceous lattice-shell (Pl. 6,
fig. 2). These pericapsular alveoles may be regarded as an outer cell-wall
more correctly than the membrane of the central capsule itself, but the
arrangement may also be compared to the temporary encystation of other
Protista (see note D).

A. The extracapsular vacuoles in the calymma were first observed in 1851
by Huxley, in _Thalassicolla_ and _Sphærozoum_, and compared with
Dujardin's sarcode vacuoles (L. N. 5). Afterwards J. Müller noticed that
generally these "large clear vesicles are covered by a fine membrane,"
and hence he called them "alveoles" (L. N. 12, pp. 3, 7, &c.). In my
Monograph I have described them more in detail as "extracapsular
alveoles" (1862, p. 88, Tafs. i.-iii. xxxii.-xxxv.). Ever since then the
point has been debated whether these clear spaces are simple vacuoles in
the sense of Huxley or vesicular alveoles as stated by J. Müller. This
contention is unnecessary, for both varieties are present, and often no
sharp line can be drawn between them. R. Hertwig has recently come to the
conclusion that they are as a rule "membraneless vacuoles," but that they
"sometimes become surrounded by a special envelope" (L. N. 33, p. 31). He
even succeeded "in extracting from a _Collosphæra_ the large vesicle
which lies in the centre of many colonies and removing its covering of
central capsules and jelly."

B. The _mechanical importance_ of the alveolar structure, which certainly
increases the elasticity and mechanical resistance of the voluminous
calymma, has not yet been sufficiently realised; in the case of those
Radiolaria which have no skeleton, or at all events no lattice-shell, it
may take the place of this as a protective envelope. Furthermore, by
taking in and giving out water it may discharge a hydrostatic function,
causing the organism to rise or sink in the water.

C. The _large central alveole_ found in the colonies of many Polycyttaria
(especially Collosphærida) and first described in my Monograph (Taf.
xxxiv. fig. 1), has since then been observed by Hertwig, Bütschli, and
other investigators, and recognised as the "central support of the whole
colony, surrounded by a delicate membrane" (compare L. N. 33, p. 31, and
L. N. 41, p. 436). In a colony of _Trypanosphæra transformata_ (Pl. 5,
fig. 1), which I observed living while in Ceylon in 1881, the membrane of
the large central alveole was surrounded by a firm network of
sarcoplegma, and could be mechanically isolated from the central
jelly-sphere which it enclosed.

D. The _pericapsular alveoles_, figured in Pl. 4, figs. 2, 3, from a
_Sphærozoum_, and in Pl. 6, fig. 2, from a _Siphonosphæra_, were very
well preserved in some preparations in the Challenger collection; perhaps
their development coincides with the formation of spores, and may be
regarded as an encystation.

{lvi}87. _The Extracapsular Fat-Globules._--Fat is probably as widely
distributed in the exoplasm as in the endoplasm of the Radiolaria; a
considerable proportion of the small, dark, highly refractive granules
appear to consist of fat; most likely they are for the most part direct
products of metastasis. These widely-spread granules, which are sometimes
coloured, and which by their passive motion produce the phenomenon of
granular circulation in the exoplasm, are not the only fatty structures in
the extracapsulum; larger globules sometimes occur. In certain large
#Collodaria# (_e.g._, _Thalassicolla melacapsa_, Pl. 1, fig. 5;
_Thalassophysa sanguinolenta_, &c.) radial series of oil-globules are found
in the calymma, especially in its proximal portion; in others the central
capsule is surrounded by a layer of oil-globules (situated in the
sarcomatrix). In the PHÆODARIA a part of the phæodium appears to consist of
fat-globules.

88. _The Extracapsular Pigment._--The formation of colouring matters in the
extracapsulum is on the whole rare in the Radiolaria, apart from the
"yellow cells" (see § 91) and from the peculiar phæodium of the PHÆODARIA,
which will be separately treated of in the next paragraph. Considerable
masses of extracapsular pigment, usually black or blue, rarely brown or
red, are found only in a few Radiolaria belonging to the first three
legions; most often in the SPUMELLARIA. Some large #Collodaria#, _e.g._,
the common _Thalassicolla nucleata_ and a few other species of this genus
(Pl. 1, fig. 4), are characterised by a rich deposit of black or blue
pigment in the sarcomatrix and in the proximal portion of the calymma.
Brown pigment is deposited in the calymma of many #Sphæroidea# and
#Discoidea#, as well as of some NASSELLARIA (_Cystidium_, _Tridictyopus_,
&c.). In a part of the ACANTHARIA red pigment granules are thickly strewn
in the sarcoplegma and pass along the free pseudopodia, as for example in
_Actinelius purpureus_ and _Acanthostaurus purpurascens_. The composition
and significance of these extracapsular pigments are not completely known.

On the extracapsular pigment of _Thalassicolla nucleata_, compare my
Monograph, pp. 87, 251. On the red extracapsular pigment-granules of the
ACANTHARIA, see L. N. 19, pp. 345, 364, &c.

89. _The Phæodium of the Phæodaria._--The PHÆODARIA, which are
distinguished from the other three legions of Radiolaria by the double
membrane of the central capsule, and the peculiar structure of the
main-opening (astropyle), differ also in other points, the most important
of which is the constant presence of a voluminous mass of extracapsular
pigment. This possesses a peculiar constitution and special significance,
and is not to be confounded with the extracapsular pigment-granules of
other Radiolaria (_e.g._, _Thalassicolla_), and hence it has been
distinguished by the name "Phæodium," and the individual granules which
compose it as "Phæodella" (see note A). The phæodium is always excentric in
position relatively to the central capsule, of which it {lvii}surrounds the
oral half in the form of a voluminous concavo-convex cap, hiding the
astropyle at its basal pole so completely that the latter is rarely visible
until the phæodium has been removed (Pls. 99-104; Pl. 115, fig. 8; Pl. 123,
&c.). The central capsule is generally almost completely embedded in the
phæodium, so that only its aboral pole (with the two parapylæ in the
TRIPYLEA) projects. In the #Phæogromia#, in which the lattice-shell
possesses a special opening and the central capsule lies excentrically in
the aboral position of its interior, the phæodium occupies the oral aspect,
between the capsule and the aperture (Pls. 99, 100, 118-120, &c.). In the
peculiar family Coelographida (Pls. 126-128) a special receptacle (galea
with its rhinocanna) for the phæodium is developed outside the bivalve
shell, within which the central capsule lies. The proboscis, which in all
PHÆODARIA arises from the centre of the astropyle, lies in the vertical
axis of the phæodium and is entirely surrounded by it. The volume of the
phæodium in the majority of the PHÆODARIA may be said to be about as great
as that of the central capsule, although in some species it is considerably
larger. Its colour is always dark, usually between green and brown,
commonly olive-green or blackish-brown, rarely reddish-brown or black. The
phæodellæ or pigment-granules which make up the greater part of the
phæodium (see note B) are irregular in form and unequal in size and show no
definite structure; usually they are spherical or ellipsoidal, and exhibit
fine parallel striæ which run transversely or obliquely (Pl. 101, fig. 3,
6, 10; Pl. 103, fig. 1, &c.). Between the larger granules is usually found
a thick dust-like mass of innumerable very small grains. The physiological
significance of this peculiar phæodium is still unknown, but is probably
considerable, judging from its large size and especially from its constant
topographical relation to the astropyle; the latter consideration would
lead to the supposition that it plays an important part in the nutrition
and metastasis of the PHÆODARIA (see note C).

A. The phæodium of _Aulacantha_, _Thalassoplancta_, and _Coelodendrum_
was first described in 1862, in my Monograph, as an excentric
extracapsular mass of pigment of blackish-brown or olive-green colour
(pp. 87, 262, 264, 361, Taf. ii. iii. xxxii.). Since then John Murray,
who investigated many living PHÆODARIA during the Challenger expedition,
has shown its general distribution in this legion (Proc. Roy. Soc. Lond.,
vol. xxiv. p. 536, 1876). From the constancy of its presence I gave the
legion the name PHÆODARIA in 1879 (L. N. 34).

B. With regard to the special composition of the phæodium and the
constitution of the phæodellæ, see the general description of the
PHÆODARIA, pp. 1533-1537.

C. Perhaps the phæodellæ are to some extent symbiontes with the
PHÆODARIA; the xanthellæ present in most other Radiolaria are absent in
this legion.

90. _The Extracapsular Xanthellæ._--Xanthellæ or Zooxanthellæ, symbiotic
"yellow cells," are very commonly found in the extracapsulum of the
Radiolaria, especially in many SPUMELLARIA and NASSELLARIA; whilst in the
ACANTHARIA similar yellow cells usually only occur within the central
capsule, and in the PHÆODARIA their {lviii}presence has not been certainly
demonstrated. The extracapsular Xanthellæ are found most abundantly in the
#Collodaria#, both in the monozootic Thalassicollida and in the polyzootic
Sphærozoida. They occur in smaller numbers in the #Sphærellaria#, and in
many divisions of the latter they seem to be entirely absent. Also it
sometimes happens that, though present in large numbers in some
SPUMELLARIA, they are entirely absent in others nearly related to them;
indeed, this has also been observed in the case of different individuals of
the same species. This fact alone is sufficient to show that the Xanthellæ
are not an integral part of the Radiolarian organism (as was formerly
believed) but parasites or more correctly symbiontes, which live as
inhabitants of the calymma. More recent investigations have shown, that
besides the yellow pigment-grains they contain starch or an amyloid
substance, that is to say, vegetable reserve materials, that their thin
envelope contains cellulose, and that their yellow colouring-matter
resembles chlorophyll and is related to that of the Diatomaceæ
("Diatomin"). Hence they are now generally regarded as unicellular Algæ,
nearly related to those which occur as symbiontes in other marine animals
(_Exuviella_, &c.). The starch, which they develop with the formation of
oxygen, may serve as nutriment to the Radiolaria, while the carbonic acid
yielded by the latter is also beneficial to the Xanthellæ. The form of the
Xanthellæ is usually spherical and elliptical, often also sphæroidal or
discoidal. Their diameter is usually between 0.008 and 0.012 mm., rarely
more or less. The differences exhibited by Xanthellæ which live in
different groups of Radiolaria demand further investigation, which will
perhaps lead to the establishment of several species of the genus
_Zooxanthella_. At present _Zooxanthella extracapsularis_, in the calymma
of SPUMELLARIA and NASSELLARIA, may be clearly distinguished from
_Zooxanthella intracapsularis_, in the central capsule of the ACANTHARIA.

The "yellow cells" were first described in 1851 by Huxley, in the
#Collodaria#, and afterwards by J. Müller (1858) in many SPUMELLARIA and
NASSELLARIA. In my Monograph (1862, pp. 84-87) I gave a detailed account
of their structure and increase by division, and laid special emphasis on
the fact that they are the only elements in the Radiolarian organism
which "are _undoubtedly cells_ in the strict histological sense of the
word." Afterwards, in my Beiträge zur Plastiden-Theorie, I showed the
constant presence of "starch in the yellow cells of the Radiolaria"
(1870, L. N. 21). Shortly afterwards Cienkowski observed that the yellow
cells live independently and reproduce themselves after the death of the
Radiolaria, and in consequence first put forth the hypothesis that they
do not belong to the Radiolarian organism, but that they are unicellular
Algæ parasitic upon it (1871, L. N. 22). This view was ten years later
more fully established by Karl Brandt, and elucidated by comparison with
the symbiosis of the gonidia of Algæ, and the hyphæ of Fungi in the
formation of Lichens, which had in the meantime become known (1881, L. N.
38). Brandt gave this unicellular yellow Alga the name _Zooxanthella
nutricola_, and afterwards gave fuller details regarding its remarkable
vital relations (L. N. 39). Patrick Geddes, who named it _Philozoon_,
supplemented this account and showed experimentally that it gives off
oxygen under the influence of sun-light (1882, L. N. 42, 43). In
consequence {lix}of this there is no doubt that all Xanthellæ (the
_Zooxanthella extracapsularis_ of SPUMELLARIA and NASSELLARIA, and the
_Zooxanthella intracapsularis_ of the ACANTHARIA, and possibly also the
_Zooxanthella phæodaris_ of the PHÆODARIA) do not originally belong to
the Radiolarian organism, as was believed up to the time of Cienkowski,
but penetrate actively into it from without, or are taken in passively by
means of the pseudopodia. In any case their symbiosis, when they are
associated with the Radiolarian cell in large numbers, may be of great
advantage to both parties, since the metastasis of the Xanthella is
vegetable, that of the Radiolarian animal in character. In any case their
symbiosis is to a large extent accidental, by no means as necessary as in
the case of the Lichens. See on these points in addition to Brandt and
Geddes (_loc. cit._) also Geza Enz, Das Consortial-Verhältniss von Algen
und Thieren, Biol. Centralbl., Bd. ii. No. 15, 1883, Oskar Hertwig, Die
Symbiose oder das Genossenschaftsleben im Thierreich, Jena, 1883, and
Bütschli, Die Radiolarien, in Bronn's Klass. u. Ord. d. Thierreichs, 1882
(L. N. 41, pp. 456-462).

91. _The Exoplasm or Extracapsular Protoplasm._--The extracapsular
protoplasm, which may be shortly termed the "exoplasm" (or ectosarc), is
primitively in all Radiolaria (and especially in their earliest development
stages) the only important constituent of the extracapsulum, besides the
calymma. Although the extracapsular and intracapsular protoplasm of the
Radiolaria are everywhere in direct communication, and although the
openings in the membrane of the central capsule bring about an interchange
between them, still the two portions of sarcode show certain constant and
characteristic differences, which are due to the physiological division of
labour between the central and peripheral parts of the body and their
corresponding morphological differentiation. The extracapsular, like the
intracapsular, protoplasm is originally homogeneous, but may afterwards
become differentiated in various ways, producing the special constituents
of the extracapsulum. Such "external protoplasmic products" are vacuoles,
pigment-bodies, &c. More important, however, are the topographically
different sections into which the exoplasm may be divided according to its
relations to the central capsule and the calymma. In this respect the
following parts may be generally distinguished--(1) the _Sarcomatrix_, or
fundamental layer of the exoplasm, which surrounds the central capsule as a
continuous sheath of sarcode and separates it from the calymma; (2) the
_Sarcoplegma_, an irregular network of the exoplasm, which spreads
throughout the gelatinous material of the calymma; (3) the _Sarcodictyum_
or network of sarcode on the outer surface of the calymma; and (4) the
_Pseudopodia_, which project outwards from the latter and radiate into the
water.

92. _The Sarcomatrix._--The sarcomatrix, being "the fundamental layer of
the pseudopodia" (or "matrix of the exoplasm"), constitutes the proximal
innermost section of the extracapsular sarcode, and in all Radiolaria forms
a thin continuous mucous layer, which covers the whole outer surface of the
central capsule and separates it from the surrounding calymma (see note A,
below). The sarcomatrix communicates internally {lx}through the openings of
the central capsule with the endoplasm, whilst externally the pseudopodia
or mucous threads arise from it, which by their union form the sarcoplegma.
The sarcomatrix is only interrupted in the SPUMELLARIA and ACANTHARIA by
those parts of the skeleton which perforate the membrane of the central
capsule. In all NASSELLARIA and PHÆODARIA, as in the #Collodaria#, it
appears as a perfectly continuous sarcode-envelope of the central capsule.
Its thickness is variable; in general it is most strongly developed in the
SPUMELLARIA and PHÆODARIA, less so in the NASSELLARIA, and is thinnest in
the ACANTHARIA. The thickness seems, however, to vary even in one and the
same individual, the difference depending partly upon the different stages
of development and partly upon nutritional conditions. After abundant
inception of nutriment the thin protoplasmic layer of the matrix is
thickened and turbid, rich in granules and irregular masses, which are
probably due to enclosed but only half-digested food; xanthellæ also, as
well as foreign bodies taken up with the nutriment, such as frustules of
Diatoms and shells of smaller Radiolaria, and of pelagic infusoria, larvæ,
&c., are often, especially in large individuals, aggregated in considerable
quantities in the matrix. After long fasting, on the contrary, this is poor
in these enclosed bodies and in granules; it then forms a thin colourless
more or less hyaline mucous coating to the central capsule. From a
physiological standpoint the sarcomatrix is to be regarded as the _central
organ of the extracapsulum_, and as of pre-eminent significance. Probably
it is not only the most important organ for the nutrition of the Radiolaria
(especially for digestion and assimilation in particular), but perhaps is
also the central organ of perception. On the other hand the sarcomatrix
belongs to those components of the Radiolarian organism which take no part
in the formation of the skeleton.

A. The sarcomatrix was first described in my Monograph in 1862 (p. 110)
as the "Mutterboden der Pseudopodien," possessing a pre-eminent
physiological importance. Compare also my paper on the sarcode elements
of the Rhizopoda (Zeitschr. f. wiss. Zool., Bd. xv. p. 342, 1865).

93. _The Sarcoplegma._--By the name sarcoplegma, as distinguished from the
remaining extracapsular sarcode, is understood the intracalymmar web of
exoplasm or "ectosarcode network," which ramifies within the gelatinous
mass of the calymma. Internally it is in direct connection with the
continuous sheath (sarcomatrix), which encloses the central capsule, whilst
externally it is in contact with the superficial sarcode network
(sarcodictyum) which surrounds the calymma. The configuration of this
exoplasmic web, which penetrates the jelly-veil in all directions, is
exceedingly variable; in most Radiolaria it is extremely irregular in form,
like the protoplasmic network in the ground-substance of many kinds of
connective tissue. In some groups, however, it assumes a rather regular
shape which it appears to retain (_e.g._, in many ACANTHARIA). It must be
assumed also that in those instances where the consistency {lxi}of the
calymma approaches that of cartilage, the tracks of the exoplasmic threads
remain constant, but accurate observations are wanting as to how far the
configuration of the sarcoplegma is constant or variable in the different
groups, as well as regarding its peculiar behaviour in those Radiolaria
whose calymma is characterised by the formation of vacuoles or alveoles
(see § 86). Usually it envelops the larger alveoles in the form of a
reticulate veil. In many #Collodaria# the exoplasm is aggregated at certain
points of the intracalymmar web, so that large balls or amoeboid bodies
appear to be distributed between the alveoles, _e.g._, in _Thalassophysa
pelagica_ and _Thalassicolla melacapsa_ (Pl. 1, figs. 4, 5). The
sarcoplegma is metamorphosed directly into silex in the Radiolaria
spongiosa, or those genera which possess a spongy cortical skeleton, and
were formerly known as Spongurida; to this category belong the
Spongosphærida (Pl. 18) and Spongodiscida (Pl. 47) as well as certain
NASSELLARIA and PHÆODARIA. The single siliceous spicules, which are
irregularly interwoven to form the spongy web, are to be regarded as the
silicified threads of the intracalymmar sarcode network. From a
physiological point of view the sarcoplegma is of importance both for the
nutrition and motion of the Radiolaria, since it brings the sarcomatrix and
the sarcodictyum, with the pseudopodia which radiate from it, into direct
communication.

94. _The Sarcodictyum._--The sarcodictyum may be defined as the
extracalymmar network of exoplasm, and is a reticular covering which lies
upon the outer surface of the gelatinous calymma. Internally, the
sarcodictyum is in direct communication with the sarcoplegma, or the web of
exoplasmic threads which ramifies in the gelatinous substance of the
calymma; externally, on the other hand, the pseudopodia radiate freely from
it; thus its relation to these is similar to that which the sarcomatrix
bears to the roots of the sarcoplegma. Relations similar to those which
have led to the separation of the primary from the secondary calymma,
induce us to distinguish also a primary and secondary sarcodictyum. The
original or _primary sarcodictyum_ ramifies over the surface of the
original or primary calymma, and like this is of pre-eminent importance in
the formation of the primary lattice-shell; if we regard the surface of the
primary calymma as the indispensable foundation for the deposition of this
latter, then the primary sarcodictyum furnishes the material from which it
is developed: silex in the SPUMELLARIA and NASSELLARIA, a silicate of
carbon in the PHÆODARIA, and acanthin in the ACANTHARIA. It may indeed be
said that the primary lattice-shell of the Radiolaria arises by a direct
chemical metamorphosis of the primary sarcodictyum, by a chemical
precipitation of the dissolved skeletal material (silex, silicate, or
acanthin), which was stored up in the exoplasm of the sarcodictyum. Hence a
deduction from the special conformation of the former to that of the latter
is permissible. The particular form of the primary lattice-sphere with its
regular or irregular meshes is due to the corresponding form of the primary
sarcodictyum; both regular and irregular forms of this {lxii}commonly
occurring. The form of the _regular sarcodictyum_ with circular or regular
polygonal, usually hexagonal, meshes is constantly maintained during the
formation of the regular lattice-shells (_e.g._, Pl. 12, figs. 5-10; Pl.
52, figs. 8-20; Pl. 96, figs. 2-6; Pl. 113, figs. 1-6). The form of the
_irregular sarcodictyum_, on the other hand, with irregular polygonal or
roundish meshes, persists during the development of the irregular
lattice-shells (_e.g._, Pls. 29, 70, 97, 106). All this is true also of the
_secondary sarcodictyum_, or the exoplasmic network which ramifies over the
surface of the secondary calymma. The secondary lattice-shells, which are
deposited on the surface of the latter, retain the configuration of the
secondary sarcodictyum, by the chemical metamorphosis of which they have
originated; this is the case in many SPUMELLARIA which develop several
concentric lattice-shells (Pl. 29), in some NASSELLARIA (Pl. 54, fig. 5),
in the Phractopeltida among the ACANTHARIA (Pl. 133), and in the
double-shelled PHÆODARIA, Cannosphærida, and part of the Coelodendrida and
Coelographida (Pls. 112, 121, 128). In those Radiolaria which form no
lattice-shell whatever, the conformation of the sarcodictyum is usually
irregular, with meshes of irregular form and unequal size; sometimes,
however, they seem to be very regular, as in many #Acanthometra# (Pl. 129,
fig. 4).

95. _The Pseudopodia._--On the whole the pseudopodia or thread-like
processes of the exoplasm exhibit in the Radiolaria the same characteristic
peculiarities as in all true Rhizopoda; they are usually very numerous,
long and thin, flexible and sensitive filaments of sarcode, which show the
peculiar phenomena of granular movement. Their physiological significance
is in several respects very great, for they serve as active organs for the
inception of nutriment, for locomotion, sensation, and the formation of the
skeleton (see note A, below). The presence of a calymma, however, which
distinguishes the Radiolaria from the other Rhizopoda, brings about certain
modifications in the behaviour of the pseudopodia. If in general all the
threads, which arise from the sarcomatrix or fundamental layer and radiate
outwards, be called "pseudopodia," then that part of them which is included
in the gelatinous substance of the calymma and forms the sarcoplegma may be
termed the "collopodia" (or intracalymmar pseudopodia), and the remaining
portion, which passes outwards from the sarcodictyum freely into the water,
may be described as "astropodia" (or extracalymmar pseudopodia). In many
Radiolaria these two portions present some differences in morphological and
physiological respects, and certain distinctions are probably generally
present (see note B). Apart from this universal differentiation in the
different groups of the Radiolaria, specially modified forms of pseudopodia
may be recognised as the axopodia and myxopodia of the ACANTHARIA (see §
95, A), and the sarcode-flagellum of certain SPUMELLARIA (see note C).

A. The pseudopodia of the Radiolaria have been so fully described in my
Monograph, in 1862, both morphologically and physiologically, that I need
only refer to the account there given {lxiii}(pp. 89-127); for
supplementary observations see R. Hertwig (1879, L. N. 33, p. 117) and
Bütschli (1882, L. N. 41, pp. 437-445).

B. The _Astropodia_, or free radiating pseudopodia, are in many
Radiolaria more or less clearly distinguishable from the collopodia,
which form the sarcoplegma within the calymma; how far these distinctions
depend upon a permanent differentiation (especially in the ACANTHARIA and
PHÆODARIA) needs further investigation.

C. The _sarcode-flagellum_ (perhaps better termed _axoflagellum_) was
first described in my Monograph (1862, p. 115) in the case of various
#Discoidea# (Taf. xxviii. figs. 5, 8; Taf. xxx. fig. 1). Hertwig has
given a substantially similar account of the organ in some other
#Discoidea# (L. N. 33, p. 67, Taf. vi. figs. 10, 11); probably this
peculiar structure is confined to the order #Discoidea# among the
SPUMELLARIA, but is widely distributed within its limits. The
axoflagellum is a thick cylindrical thread of sarcode, finely striated
and pointed towards its free end. It always lies in the equatorial plane
of the discoidal body, and always unpaired in one of its axes; in the
triradiate #Discoidea# it is in the axis of the unpaired principal arm
and opposite to it (Pl. 43, fig. 15). In the Ommatodiscida (p. 500, Pl.
48, figs. 8, 19, 20) the axoflagellum probably passes out through the
peculiar marginal ostium of the shell. Perhaps it is always connected
with the central nucleus by intracapsular axial fibres, and is to be
regarded as a specially differentiated bundle of pseudopodia (or
axopodia?).

95A. _The Myxopodia and Axopodia._--The two forms of pseudopodia which we
distinguish as myxopodia and axopodia differ markedly from each other both
morphologically and physiologically. The _myxopodia_, or ordinary free
pseudopodia, which are found in large numbers in all Radiolaria, and
constitute their most important peripheral organs, are simple homogeneous
exoplasmic threads, which arise from the sarcodictyum or extracalymmar
sarcode network, and radiate freely into the water; here they may branch
and combine by anastomosis to form a changeable network, but they never
contain an axial thread. The _axopodia_, on the other hand, are
differentiated pseudopodia, which consist of a firm radial thread, and a
soft covering of exoplasm; they penetrate the whole calymma in a radial
direction and project freely from its surface, and generally (if not
always) they are produced inwards to the middle of the central capsule,
perforating its membrane; their proximal end is lost in a dark central heap
of granules. Such axopodia are at present known with certainty only in the
ACANTHARIA, where they are widely, and perhaps universally, distributed.
Their development in this legion probably stands in direct causal relation
to the peculiar structure of the central capsule and the centrogenous
formation of the skeleton. Since the radial skeletal rods of the
#Acanthometra# possess originally a thin coating of protoplasm, it may be
said that the centrogenous axopodia of this group became differentiated in
two ways, the firm axial threads of one section remaining very thin and
covered by protoplasm, whilst those of the other section became
metamorphosed into radial bars of acanthin. This hypothesis acquires more
probability from the regular distribution and arrangement of the axopodia
in the ACANTHARIA; they usually stand at fixed intervals {lxiv}between the
radial bars, singly or in groups; sometimes their number seems to be not
greater than that of the bars, whilst in other cases a circlet or group of
axopodia corresponds to each radial bar. Perhaps their fine axial thread
consists of acanthin. At all events the axopodia are constant organs
(probably sensory, like the "palpocils") and not retractile like the
movable myxopodia.

The axial threads in the pseudopodia of the #Acanthometra# were first
discovered by R. Hertwig, who accurately described their peculiar
structure and arrangement (L. N. 33, pp. 16, 117).

96. _The Myophriscs of the Acanthometra._--The #Acanthometra# are
characterised by a very peculiar differentiation of the exoplasm, namely,
by the formation of myophriscs or contractile threads from the
sarcodictyum. In most (and perhaps in all) ACANTHARIA of this order each
radial bar is surrounded by a circlet of such contractile threads, which
was first described as a "ciliary corona" (see note A, below). The number
of contractile threads in each circlet usually amounts to from ten to
twenty, rarely being more than thirty and less than eight; it often appears
to be constant in the individual species (see note B). In the living state
the myophriscs are long, thin filaments, the pointed distal end of which is
inserted into the radial bar, whilst the thicker proximal end is attached
to the surface of the calymma, which is elevated round the base of each rod
into the form of a gelatinous cone or skeletal sheath (see note C).
Probably the myophriscs lie on the outer surface of the apical portion of
this gelatinous cone, and are hence to be regarded as exoplasmic threads
differentiated from the sarcodictyum. Sometimes, however (as in
_Acanthochiasma_), they fuse into a contractile membrane and form the
envelope of a cone, whose interior is occupied by a gelatinous papilla of
the calymma. On mechanical irritation the myophriscs contract rapidly and
suddenly, like muscle-fibrillæ, becoming at the same time thicker, and
hence are very different from pseudopodia. Their distal point of insertion
being fixed to the firm acanthin rod, they raise by their contraction the
skeletal sheath, to which their bases are attached or in the surface of
which they lie. The result of their contraction is therefore a distention
and increase in volume of the calymma, with which is no doubt connected an
inception of water into the gelatinous mass, and hence a diminution in its
specific gravity. Probably the #Acanthometra# contract their myophriscs
voluntarily when they wish to rise in the water; when these relax the
calymma collapses owing to its elasticity, water is then expelled and the
specific gravity increases. From a physiological point of view, then, the
myophriscs are to be regarded as a hydrostatic apparatus, morphologically
as myophanes or muscular fibrillæ, such as also occur in the intracapsular
protoplasm (see §§ 77-80). On more violent irritation and after the death
of the #Acanthometra# the myophriscs separate from the radial bars and
remain attached to the distal ends of the conical gelatinous sheaths as
free "ciliary coronas." At the same time, {lxv}they melt into short, thick,
hyaline rods, the so-called "gelatinous cilia." The myophriscs are found
only in the order #Acanthometra#, and are wanting in the #Acanthophracta#,
as well as in the other three legions of Radiolaria.

A. The "_ciliary coronas_" on the skeletal rods of dead #Acanthometra#
were first described by the discoverer of this order, Johannes Müller,
and referred to as "the stumps of the contracted, thickened threads" (L.
N. 12, p. 11, Taf. xi.).

B. The "_number of the gelatinous cilia_" I found constant in certain
species of #Acanthometra#, and stated in my Monograph (L. N. 16, p. 115)
"that here is to be found the first differentiation of the diffuse
sarcode into definite organs of regular definite number, size, and
position, which deserve the name tentacles rather than pseudopodia."

C. The nature of the myophriscs as fibrillæ allied to muscles was first
discovered by R. Hertwig, who described them as "structures of peculiar
nature," under the name of "contractile threads," and pointed out in
detail their histological and physiological peculiarities (L. N. 33, pp.
16-19, Taf. i.).

97. _The Exoplasm of the Peripylea._--The extracapsular protoplasm of the
SPUMELLARIA or PERIPYLEA is in communication with the intracapsular sarcode
by the innumerable fine pores of the capsule-membrane, and like these pores
is evenly distributed over the whole surface. The sarcomatrix which
immediately surrounds the central capsule is moderately strong, and sends
out innumerable long, thin pseudopodia, which probably correspond to the
pores of the membrane. Their number is markedly greater in the SPUMELLARIA
than in the other three legions. The ramifications and communications which
the radiating fibres of the sarcomatrix undergo within the calymma,
apparently present the most manifold variations, so that the sarcoplegma or
intracalymmar network thus formed has very diverse forms. On the surface of
the calymma the exoplasmic threads constitute a variously disposed
sarcodictyum, a regular or irregular exoplasmic network, by the
silicification of which a primary lattice-shell arises in the majority of
the SPUMELLARIA. The free ends of the pseudopodia, which arise from this
extracalymmar network and radiate out into the water, appear in most
SPUMELLARIA to be relatively short, but exceedingly numerous. Specially
modified pseudopodia and axial threads in particular do not seem to occur
in this legion. Perhaps, however, among the latter may be reckoned the
remarkable pseudopodia which combine to form the sarcode flagellum in many
#Discoidea# (and perhaps in other SPUMELLARIA). This axoflagellum is a
particularly strong thread of sarcode, arising from a definite point in the
central capsule; it is cylindrical or slenderly conical in form, much
longer, stronger, and more contractile than the ordinary pseudopodia; it
contracts in a serpentine fashion on mechanical irritation and seems to
originate by the fusion of a bundle of pseudopodia (compare § 95, C).

98. _The Exoplasm of the Actipylea._--The extracapsular protoplasm of the
ACANTHARIA or ACTIPYLEA differs in several important respects from that of
other {lxvi}Radiolaria, and appears to undergo more significant
differentiations than that of the three other legions. Since the pores in
the wall of the central capsule are not distributed evenly and at equal
intervals over its whole surface (as in the PERIPYLEA), but rather exhibit
a regular disposition in groups at unequal intervals, the number of
projecting pseudopodia is much less and the law of their arrangement
different from that which obtains in the PERIPYLEA (§ 58). In many and
probably in all ACANTHARIA they are divided into two groups, those which
arise from the centre of the capsule and possess firm axial threads, and
those which have not these characters (compare § 95, A). The axopodia, or
stiff pseudopodia with axial threads, arise from the centre of the capsule,
are present in much smaller numbers than the soft and flexible myxopodia,
and are regularly disposed between the radial bars of acanthin, usually so
that they are as far removed from them as possible, _i.e._, in the centre
between each three or four bars; these latter may indeed be regarded as
strongly developed axial threads, which have become changed into acanthin
(§ 95, A). The soft myxopodia, or pseudopodia without axial threads, are
much more numerous than the others, and arise from the sarcodictyum or
exoplasmic network which ramifies over the surface of the calymma. Their
number and arrangement seem, however, in many (if not in all) ACANTHARIA to
be regular and not to possess the extraordinary variability seen in the
other three legions. In many #Acanthometra# the sarcodictyum exhibits a
symmetrical conformation, with regular or subregular, polygonal (mostly
hexagonal) meshes, and generally the stronger threads of the sarcodictyum
secrete a firm, homogeneous or fibrillar, striated substance, which forms a
network of ridges on the surface of the calymma. In the #Acanthophracta#
the place of this is taken by the acanthin network of the primary
lattice-shell. The axopodia of the #Acanthometra# are usually about as long
as the radial spines between which they stand; their stiff axial thread is
surrounded by a soft sheath of protoplasm, communicating with the thin
sarcomatrix which surrounds the central capsule. Numerous branches pass
into the calymma from the exoplasmic sheath of the axial threads, and form
by their interweaving a loose sarcoplegma. The most peculiar differentiated
products of the exoplasm of the ACANTHARIA, however, are the myophane
fibrillæ of the #Acanthometra#, which have already been described under the
name of myophriscs (§ 96).

99. _The Exoplasm of the Monopylea._--The extracapsular protoplasm of the
NASSELLARIA or MONOPYLEA arises only from the porochora, or the
intracapsular podoconus, the oral base of which is formed by this porous
area. The pseudopodia or protoplasmic threads which pass through the pores
of the latter, united into a bundle, are not very numerous (in most
NASSELLARIA probably between thirty and ninety), and unite just outside it
to form a thick discoid sarcomatrix; this covers the porochora completely
below, and spreads out in the form of a thin envelope of exoplasm over the
whole {lxvii}surface of the central capsule; at the apical portion of the
latter the sarcomatrix is often so thin that it can only be recognised by
the aid of reagents; it separates the membrane of the central capsule from
the surrounding calymma. The pseudopodia, which penetrate the latter and by
loose anastomoses from a wide-meshed sarcoplegma within it, are usually not
very numerous. The greater part of them radiate in a bunch downwards from
the basal disc of the sarcomatrix, and a smaller number arise from the
thinner envelope which covers the remainder of the central capsule (Pl. 51,
fig. 13; Pl. 65, fig. 1; Pl. 81, fig. 16). On the outer surface of the
calymma the collopodia, which have passed through it, unite to form the
sarcodictyum, and through the silicification of this the primary
lattice-shell arises in the great majority of the NASSELLARIA. From the
surface of the sarcodictyum arise the astropodia, or free pseudopodia which
radiate outwards into the water. Their number in most MONOPYLEA is
relatively small, but their length appears to be very great.

100. _The Exoplasm of the Cannopylea._--The extracapsular protoplasm of the
PHÆODARIA or CANNOPYLEA is much better developed as regards volume than in
the other three legions, and is connected with the intracapsular sarcode by
only a few apertures in the capsule-membrane. In most PHÆODARIA three of
these are present, the astropyle or main-opening at the oral pole of the
main axis, and the two lateral parapylæ or accessory openings on either
side of the aboral pole (§ 60). In several families the latter appear to be
wanting, whilst in others their number is increased; these families have
not yet, however, been observed during life. The protoplasm projects both
from the oral main-opening and from the two aboral accessory openings in
the form of a thick cylindrical rod; the tube into which each opening is
produced in many PHÆODARIA (longer in the case of the astropyle, shorter in
the parapylæ) being regarded as an excretion from this protoplasmic
cylinder. The sarcode threads within the tube appear like a bundle of
fibrils, either quite hyaline or finely striated. After issuing from the
mouth of the aperture they pass over into a thick sarcomatrix, which
surrounds the central capsule entirely and separates it from the enclosing
calymma. In the neighbourhood of the basal astropyle the sarcomatrix is
usually swollen into a thick lenticular disc, which is in direct contact
with the peculiar phæodium of this legion (§ 89). The pseudopodia, which
radiate from the sarcomatrix, and form by anastomosis a wide-meshed
sarcoplegma within the calymma, are usually not very numerous in the
PHÆODARIA, but are very strong. Sometimes two stronger bundles of
collopodia may be distinguished at the two poles of the main axis, an oral
bundle (in the direction of the proboscis of the astropyle) and an aboral
bundle (at the opposite pole between the parapylæ). The collopodia of the
sarcoplegma unite at the surface of the calymma into a regular or irregular
sarcodictyum, which, in most PHÆODARIA produces by the secretion of a
peculiar silicate the primary lattice-shell. {lxviii}The free astropodia,
which pass outwards from the sarcodictyum into the water, are in most
PHÆODARIA very numerous (Pl. 101, fig. 10). Since, however, only a few
species of this great legion have been observed in a living state, their
pseudopodia require further accurate examination.

CHAPTER IV.--THE SKELETON.

(§§ 101-140).

101. _The Significance of the Skeleton._--The skeleton of the Radiolaria is
developed in such exceedingly manifold and various shapes, and exhibits at
the same time such wonderful regularity and delicacy in its adjustments,
that in both these respects the present group of Protista excels all other
classes of the organic world. For, in spite of the fact that the
Radiolarian organism always remains merely a single cell, it shows the
potentiality of the highest complexity to which the process of skeleton
formation can be brought by a single cell. All that has been brought to
pass in this direction by single tissue-cells of animals and plants does
not attain the extremely high stage of development of the Radiolaria. Only
very few Rhizopoda of this very rich and varied class fail to exhibit the
power of forming this firm supporting and protecting organ--indeed, only
ten of the seven hundred and thirty-nine genera which are enrolled in the
list of the Challenger collection, namely, six genera of SPUMELLARIA (five
Thalassicollida, _Actissa_, _Thalassolampe_, _Thalassopila_,
_Thalassicolla_, _Thalassophysa_, Pl. 1, and one genus of Collozoida,
_Collozoum_, Pl. 3), and in addition two genera of NASSELLARIA (the
Nassellida, _Cystidium_ and _Nassella_, Pl. 91, fig. 1), and two genera of
PHÆODARIA (the Phæodinida, _Phæocolla_ and _Phæodina_, Pl. 101, figs. 1,
2). These skeletonless forms of Radiolaria are, however, of extreme
interest, since they include the original stem-forms of the whole class as
well as of its four legions. All Radiolaria which form skeletons have
originated from soft and skeletonless stem-forms by adaptation, and that
polyphyletically, for the skeletal types of the four legions have been
developed independently of each other (§ 108).

102. _The Chemical Peculiarities of the Skeleton._--The chemical
composition of the skeleton shows very marked variations in the different
legions of the Radiolaria. The two legions SPUMELLARIA and NASSELLARIA
(united formerly as "Polycystina") form their skeleton of pure silica (see
note A, below); the legion PHÆODARIA of a silicate of carbon (see note B),
and the ACANTHARIA of a peculiar organic substance--acanthin (see note C).
This explains the well-known fact that the deposits of fossil Radiolaria
(or Polycystine marls) are composed exclusively of the skeletons of
SPUMELLARIA and NASSELLARIA, those of the ACANTHARIA and PHÆODARIA being
entirely absent (in the case of the last group, however, exception must be
made in favour of the Dictyochida, or those PHÆODARIA {lxix}whose skeleton
is made up of isolated scattered tangential siliceous fragments). The
enormous deposits of Radiolarian skeletons in the deep sea of today, which
constitute the Radiolarian ooze, consist, like the fossil Polycystine
marls, almost exclusively of the shells of SPUMELLARIA and NASSELLARIA,
though here the acanthin skeletons of the ACANTHARIA may be present in very
small numbers, and the silicate skeletons of the PHÆODARIA, which offer
more resistance to the solvent action of sea-water, somewhat more
abundantly. Calcareous skeletons do not occur in the Radiolaria (see note
D).

A. The pure siliceous skeletons of the Polycystina were first recognised
in 1833 by Ehrenberg in chalky marls (L. N. 2, p. 117). Since the two
legions ACANTHARIA and PHÆODARIA were entirely unknown to Ehrenberg, his
name Polycystina has reference only to the SPUMELLARIA and NASSELLARIA.

B. The silicate skeleton of the PHÆODARIA was formerly taken by me for a
purely siliceous one. When I described the first PHÆODARIA in my
Monograph in 1862, I was only acquainted with five genera and seven
species, whilst the number of PHÆODARIA here described from the
Challenger amounts to eighty-four genera and four hundred and sixty-five
species. In the vast majority of these (though not in all) the skeleton
becomes more or less intensely stained by carmine, and is also more or
less charred at a red heat, in some even becoming of a blackish-brown. In
many PHÆODARIA, furthermore, the hollow skeletal tubes are destroyed by
the continued action of heat. They are also, for the most part, strongly
acted upon, or even destroyed by boiling caustic alkalis, whilst boiling
mineral acids have no effect upon them. The best method of cleaning the
skeletons of PHÆODARIA from their soft parts is to heat them in
concentrated sulphuric acid, and then add a drop of fuming nitric acid;
in this they are not dissolved even on prolonged heating. From these
facts it would appear that the skeletons of the PHÆODARIA consist of a
compound of organic substance and silica, or a "carbonic silicate." The
more intimate composition yet remains to be discovered, as also the
manifold differences which the various families of PHÆODARIA seem to show
in respect of its composition. The small skeletal fragments of the
Dictyochida (the only remains of PHÆODARIA which occur as fossils) appear
to consist of pure silica.

C. The acanthin skeleton of the ACANTHARIA was first described as such in
my Monograph (1862, pp. 30-32). Johannes Müller, the discoverer of this
legion, took them for siliceous skeletons and defined the #Acanthometra#
as "Radiolaria without lattice-shell, but with siliceous radial spines"
(L. N. 12, p. 46). I formerly supposed that the acanthin skeletons in
some of the ACANTHARIA were partially or wholly metamorphosed into
siliceous skeletons, but, according to the investigations of R. Hertwig,
this does not appear to be the case; he showed that the skeletons of the
most varied #Acanthometra# and #Acanthophracta# are completely dissolved
under the longer or shorter action of acids, and supposes that in all
ACANTHARIA, without exception, the skeleton is composed of acanthin
(1879, L. N. 33, p. 120). Quite recently Brandt has found that the
acanthin spines dissolve not only in acids, alkalis, and "liquor
conservativus" (as I had shown), but also in solutions of carbonate of
soda (1 per cent.), and even of common salt (10 to 20 per cent.); he
concludes from this that they consist of an albuminoid substance
(vitellin) (L. N. 38, p. 400). I am unable to share this view, for I have
never been able to see some of the most important reactions of albumen in
any of the skeletons which I have examined, such for example as the
xanthoproteic reaction, the red coloration with Millon's test, &c. They
do not become {lxx}yellow either with nitric acid or with iodine. In
dilute mineral acids they dissolve more rapidly than in concentrated. My
usual method of cleansing the skeleton of ACANTHARIA (which has been
practised with the same result on thousands of specimens) consists in
heating the preparation in a small volume of concentrated sulphuric acid
and then adding a drop of fuming nitric acid; all other constituents (the
whole central capsule and the calymma) are thus very rapidly destroyed;
the skeleton remains quite uninjured and withstands the combined action
of the mineral acids for a longer or shorter time, though on prolonged
heating it also is dissolved. I do not therefore regard acanthin as an
albuminous substance, but as one related to chitin.

D. Calcareous skeletons have not been certainly demonstrated in the
Radiolaria, and probably do not occur. Sir Wyville Thomson in his
Atlantic (1877, L. N. 31, vol. i. p. 233, fig. 51) described under the
name _Calcaromma calcarea_, a Radiolarian which contained scattered in
its calymma numerous calcareous corpuscles "resembling the rowels of
spurs." These are identical with the "toothed bodies, recalling crystal
balls," which Johannes Müller figured in the Mediterranean _Thalassicolla
morum_ so early as 1858, and compared with the "siliceous asterisks of
_Tethya_" (L. N. 12, p. 28, Taf. vii. figs. 1, 2). I formerly regarded
these peculiar calcareous corpuscles, whose solubility in mineral acids I
had observed, as spicules of a Thalassicollid, and hence described the
species in my Monograph as _Thalassosphæra morum_ (L. N. 16, p. 260). I
have, however, seen reason to change my view, and am now led to suppose
that those peculiar calcareous corpuscles, which may be named
"_Calcastrella_," are not formed by the Radiolarian itself, but are
foreign bodies which have been accidentally incorporated into the calymma
of a Thalassicollid (_Actissa_). These corpuscles occur, often in large
numbers, in many preparations in the Challenger collection, and in the
calymma of other Radiolaria, chiefly #Discoidea#, hence it would appear
that they are foreign bodies taken up by the pseudopodia and carried into
the calymma by the circulation of the sarcode. The Radiolaria which Sir
Wyville Thomson figured as _Calcaromma calcarea_, and Müller as
_Thalassicolla morum_, I regard as species of _Actissa_ (see p. 13),
perhaps _Actissa radiata_ of the Pacific, and _Actissa primordialis_ of
the Mediterranean (compare the description of the Thalassosphærida of the
Challenger collection, pp. 30, 31).

103. _The Physical Properties of the Skeleton._--The skeletons of all
Radiolaria are characterised pre-eminently by a high degree of _firmness_,
which fits them to serve as protective and supporting apparatus. This is
obvious in the case of the pure siliceous shells of the Polycystina; but
the acanthin framework of the ACANTHARIA also possesses a degree of
stiffness but little inferior, whilst the silicate skeletons of the
PHÆODARIA seem on the whole to be not so firm. The hollow skeletal tubes of
the last-named, which are filled with gelatinous material, are very brittle
on account of the delicacy of their walls. Their _elasticity_ also is very
small, whilst that of the acanthin spines is considerable. The thin long
needles of many ACANTHARIA are very elastic, as are also the bristle-like
siliceous spicules of many SPUMELLARIA. The _refractive power_ of the
skeleton in the various legions is very different, depending upon the
chemical constitution. The siliceous skeleton of the Polycystina
(SPUMELLARIA and NASSELLARIA) and the silicate skeleton of the PHÆODARIA
have the same refractive index as glycerine, and hence become invisible
when mounted in that fluid; they then become visible only on addition of
{lxxi}water, and are clearer in proportion to the quantity of water which
is added. The refractive index of acanthin is, however, very different from
that of glycerine, so that the skeletons of ACANTHARIA are readily visible
when mounted in this fluid. In water, the skeletons of all Radiolaria
appear about equally refractive, as also in Canada balsam. The substance of
the skeleton appears almost entirely hyaline, colourless, and transparent.
Very rarely it is faintly coloured (in some ACANTHARIA). A cloudy opaque
constitution is seen in some PHÆODARIA (especially in the "porcellanous
shells" of Tuscarorida and Circoporida, Pls. 100, 114-117); when dried,
these appear by reflected light milky-white or yellowish-white; the cause
of this opacity lies partly in the peculiar "cement-like structure" of
these porcellanous shells, partly in their fine porosity, and the minute
air-bubbles contained in their thick walls.

104. _The Elementary Structure of the Skeleton._--The general constitution
of the skeleton--or more accurately expressed, of the morphological
elements of which the skeleton consists--is of such a nature that it may be
termed structureless. Both the organic acanthin skeletons of the ACANTHARIA
and the silicate skeletons of the PHÆODARIA, as well as the inorganic
siliceous skeletons of the SPUMELLARIA and NASSELLARIA, appear under the
microscope perfectly homogeneous, transparent, colourless, and crystalline.
Only very rarely do they show traces of a concentric striation, which
arises from the deposition of the skeletal substance in layers; as, for
example, the thick spines of some PHÆODARIA (Pls. 105-107, &c.). Some of
the PHÆODARIA, however, form an exception to this rule, inasmuch as their
partially tubular skeletal elements possess a remarkable porcellanous
structure. In the tubular or Cannoid skeleton, which occurs in most
CANNOPYLEA, the lumen of the thin-walled flinty tube is filled with jelly,
and frequently a thin siliceous thread runs in its axis, and is connected
with the wall by transverse threads (§§ 127, 139). The elementary structure
of the opaque porcellanous shells, which distinguish the two families
Circoporida (Pls. 114-117) and Tuscarorida (Pl. 100), is quite peculiar.
Numerous fine siliceous spicules lie scattered irregularly in a finely
granular or porous matrix.

105. _Complete and Incomplete Lattice-Shells._--In the great majority of
Radiolaria (in all four legions) the skeleton has the form of a delicate
lattice-shell or a receptacle in which the central capsule is enclosed. In
a small minority, however, this is not the case. The skeleton then consists
only of isolated rigid pieces (radial or tangential spicules), or of a
simple ring (sagittal ring of the #Stephoidea#), or of a basal tripod with
or without a loose tissue of trabeculæ, &c. (#Plectoidea#); the central
capsule is then not surrounded by a special latticed receptacle, but only
rests upon the skeletal trabeculæ. According to these different
arrangements, two principal groups or sublegions may be distinguished in
each legion, of which one set (Cataphracta) are characterised by a complete
{lxxii}lattice-shell, whilst the others (Aphracta) are without it. The
RADIOLARIA APHRACTA, then, or Radiolaria without a complete skeleton, are
the #Collodaria# (p. 9), the #Acanthometra# (p. 725), the #Plectellaria#
(p. 895), and the #Phæocystina# (p. 1543). On the other hand, the
RADIOLARIA CATAPHRACTA, or Radiolaria with a complete skeleton, are the
#Sphærellaria# (p. 49), the #Acanthophracta# (p. 791), the #Cyrtellaria#
(p. 1015), and the #Phæocoscina# (p. 1590).

Upon this basis the first subdivision of the Radiolaria was made by
Johannes Müller, who recognised three groups:--"I. _Thalassicolla_,
without receptacle, naked or with spicules; II. _Polycystina_, with a
siliceous receptacle; III. _Acanthometra_, without receptacle, but with
siliceous radial spines" (L. N. 12, p. 16).

106. _The Ectolithia and Entolithia (Extracapsular and Intracapsular
Skeletons)._--The relation of the skeleton to the central capsule in the
Radiolaria is very various in many respects; in the first instance two
great groups, _Ectolithia_ and _Entolithia_ (see note A), may be
distinguished topographically by mere external observation; in the former
the skeleton lies entirely outside the central capsule; in the latter,
partially at all events, within it. The _Ectolithia_, with a completely
extracapsular skeleton, include all NASSELLARIA and PHÆODARIA, as well as a
great part of the SPUMELLARIA (all #Collodaria# and the most archaic forms
of #Sphærellaria#); the _Entolithia_, on the other hand, in which the
skeleton lies partly within, partly without the central capsule, include
all ACANTHARIA and the majority of the SPUMELLARIA (most #Sphærellaria#,
see note B).

A. The difference between Ectolithia and Entolithia was applied in my
Monograph in 1862 (p. 222) to separate the Monocyttaria into two main
groups. The arrangement was, however, quite artificial, being contrary to
the natural relations of the larger groups, as was shown seventeen years
later by the discovery of the different structural relations of the
central capsule.

B. Among the ACANTHARIA, which all possess primitively an intracapsular
and centrogenous skeleton, the remarkable _Cenocapsa_ (Pl. 133, fig. 11),
seems to furnish the single exception; in it the skeleton consists of a
simple spherical shell which encloses the concentric central capsule. The
exception is, however, only apparent; the twenty perspinal pores of the
shell show that they were originally in connection with twenty
centrogenous acanthin spines, and that those have disappeared by
retrograde metamorphosis.

107. _Perigenous and Centrogenous Skeletons._--Much more important than the
topographical relation of the skeleton to the central capsule, according to
which the Ectolithia and Entolithia are separated from each other (§ 106),
is the original development of the skeleton within or without the central
capsule, which gives rise to the distinction between perigenous and
centrogenous skeletons. _Centrogenous skeletons_ are found only in the
ACANTHARIA, which are further distinguished from all other Radiolaria by
their skeleton being formed of acanthin; in all ACANTHARIA the formation of
the skeleton begins in the middle of the central capsule, from which twenty
(the number is inconstant only in the {lxxiii}small group #Actinelida#)
radial spines are centrifugally developed. The three other legions, on the
contrary, possess on the whole a _perigenous_ skeleton, which _originally_
develops outside the central capsule and never in its middle. In the
NASSELLARIA and PHÆODARIA the skeleton retains this extracapsular position,
as also in the #Beloidea# and part of the #Sphærellaria# among the
SPUMELLARIA; in the great majority of the latter, however, the primary
perigenous skeleton is subsequently enveloped by the growing central
capsule, so that it lies partially within it (§ 109).

108. _Polyphyletic Origin of the Skeleton._--The skeleton of the Radiolaria
has undoubtedly originated polyphyletically, for it is impossible to derive
its manifold varieties from a single ground-form, or to regard them as
modifications of one type. It is much more probable that the different
skeletonless Radiolaria have entered upon different ways of skeleton
formation quite independently of each other. At the outset it is quite
clear that the skeletons of the _four legions have originated independently
of each other_. Further, it is certain that within the legion of the
SPUMELLARIA the Beloid skeletons of the #Collodaria# are not connected with
the Sphæroid skeletons of the #Sphærellaria# and the forms derived from
them (see § 109). In the same way the skeletons of the PHÆODARIA are
polyphyletic; probably in this legion the Beloid, Sphæroid, Cyrtoid, and
Conchoid skeletons have been developed quite independently (see § 112). In
the NASSELLARIA, on the other hand, it is possible that all the skeletal
forms are to be derived monophyletically from a single simple primitive
form (either the sagittal ring or basal tripod?) (see § 111). Still more
probable is it that the ACANTHARIA have arisen monophyletically, for all
the forms of their acanthin skeleton may be derived without violence from
_Actinelius_ (see § 110).

109. _The Skeleton of the Spumellaria._--The skeletons of the SPUMELLARIA
or PERIPYLEA consist of silica, and are very different and of independent
origin in the two orders of this legion. The first order, #Collodaria#,
have either no skeleton whatever (#Colloidea#, p. 10, Pls. 1, 3), or their
skeleton is _Beloid_, a loose extracapsular envelope of spicules,
consisting of numerous unconnected portions; the separate parts are usually
disposed tangentially, either as simple or compound siliceous spicules
(#Beloidea#, p. 28, Pls. 2, 4). The second order of SPUMELLARIA, on the
other hand (#Sphærellaria#, p. 49), develops a siliceous lattice-shell
which consists of a single piece, and is remarkable for the extraordinary
variety of its forms (pp. 50-715, Pls. 5-50). To this order belong not less
than three hundred genera and seventeen hundred species of the Challenger
Radiolaria (that is, about two-fifths of all the genera and species). In
spite of this extreme richness in different forms this large group must be
regarded as _monophyletic_, since all its forms may be quite naturally
derived from a common stem-form, a _simple lattice-sphere_ (_Cenosphæra_,
p. 61, Pl. 2). The twenty-eight families of #Sphærellaria# may be
distributed in four suborders, among which the #Sphæroidea# constitute the
{lxxiv}stem-forms, since they retain the original spherical shape (Pls.
5-8, 11-30). In the other three suborders a vertical main axis is
developed, which in #Prunoidea# is longer, in #Discoidea# shorter than the
other axes of the shell. Hence the shell of the #Prunoidea# (p. 284, Pls.
13, _bis_, 17, 39, 40) is ellipsoidal or cylindrical, that of the
#Discoidea#, on the other hand, lenticular or discoidal (p. 402, Pls.
31-38, 41-48). Finally, the shell of the fourth suborder, #Larcoidea#, is
lentelliptical; it has the ground-form of a triaxial ellipsoid, and is
characterised by the possession of three unequal dimensive axes, or three
isopolar axes of different lengths perpendicular to each other (p. 599,
Pls. 9, 10, 49, 50).

110. _The Skeleton of the Acantharia._--The skeletons of the ACANTHARIA or
ACTIPYLEA are distinguished from those of all other Radiolaria by two very
important peculiarities; in the first place, they consist not of silica but
of a peculiar organic substance, _Acanthin_, and secondly, their
development is centrogenous, numerous radial spines or acanthin spicules
being formed which are united in the middle of the central capsule. Hence
the ACANTHARIA are the only Radiolaria in which the skeleton originates
from the first in the middle of the central capsule. The number of radial
spines is primitively indefinite, variable, and often considerable (more
than a hundred), but in the great majority it is limited to twenty. In
accordance with this the legion may be divided into two orders, the more
archaic small group Adelacantha, with an indefinite number of spines, and
the more recent group, Icosacantha, which has been developed from them and
possesses twenty regularly disposed spines; of the three hundred and
seventy-two species of ACANTHARIA which have been hitherto described, about
five per cent. belong to the former, about ninety-five per cent. to the
latter division (see note A, below). The numerous genera of Icosacantha may
then be again divided into two suborders, of which the #Acanthonida# (p.
740, Pls. 130-132) produce no complete lattice-shell, and thus agree with
the #Actinelida#, with which they may be united as #Acanthometra# in the
broader sense (or ACANTHARIA without a lattice-shell). The
#Acanthophracta#, on the other hand (p. 791, Pls. 133-140), produce a
complete lattice-shell, usually by means of two opposite or four crossed
transverse processes, which arise from each radial spine and unite with
each other (see note B, below). In most #Acanthophracta# the lattice-shell
remains single; only in the Phractopeltida does it consist of two
concentric lattice-spheres (p. 847, Pl. 133, figs. 1-6). Furthermore, the
whole order #Acanthophracta# may be subdivided into two suborders according
to the different ground-form of the lattice-shell; this remains spherical
in the #Sphærophracta# (the three families Sphærocapsida, Dorataspida,
Phractopeltida, Pls. 133-138). On the other hand, it assumes another form
in the #Prunophracta#; it becomes ellipsoidal in the Belonaspida (Pl. 136,
figs. 6-9), discoidal or lentiform in the Hexalaspida (Pl. 139); and
finally takes the shape of a double cone in the Diploconida (Pl. 140).

{lxxv}A. The group Adelacantha consists only of the suborder
#Actinelida#, with the three families Astrolophida, Litholophida, and
Chiastolida (p. 728, Pl. 129, figs. 1-3); the number of the radial spines
is very different and variable, sometimes only from ten to sixteen, but
usually from thirty to fifty, and often more than one hundred; they are
generally irregularly distributed, and not as in the second main
division. This latter, the Icosacantha, always possesses _twenty_ radial
spines, which are regularly disposed according to a constant law, the
so-called "Müllerian" or "Icosacanthan" law; the twenty spines are always
so placed between the poles of a spineless axis that they form five zones
each of four spines; the four spines of each zone are equidistant from
each other, and also from the same pole, and alternate with those of the
neighbouring zones, so that the whole twenty lie in four meridian planes,
which cut out an angle of 45° (compare pp. 717-722, Pls. 130-140). In
spite of the manifold variations in form which are developed in the
Icosacantha, they may all be derived from a common stem-form,
_Acanthometron_ (p. 742), since the law of distribution of the twenty
spines is constantly inherited.

B. An exception is found in the peculiar family Sphærocapsida (p. 797,
Pl. 133, figs. 7-11; Pl. 135, figs. 6-10). Here the shell is composed of
innumerable small, perforated plates, which arise on the surface of the
calymma independently of the spines.

111. _The Skeleton of the Nassellaria._--The skeletons of the NASSELLARIA
or MONOPYLEA consist of silica, and are never composed of separate
portions, but constitute always a single continuous piece. The ground-form
is originally monaxon, corresponding to that of the central capsule, with a
constant difference between the two poles of the vertical main axis. The
ground-form is never spherical or polyaxon as in the lattice-shells of the
SPUMELLARIA, and the skeleton never consists of hollow tubes, as in the
PHÆODARIA. The legion NASSELLARIA may be divided into two orders; in the
#Plectellaria# (three suborders #Nassoidea#, #Plectoidea#, #Stephoidea#)
the skeleton does not form a complete lattice-shell; in the #Cyrtellaria#,
on the other hand, which are derived from these, the siliceous skeleton
forms a complete lattice-shell enclosing the central capsule. The number of
forms thus developed is astonishingly great, so that among the NASSELLARIA
no less than two hundred and seventy-four genera and sixteen hundred and
eighty-seven species may be distinguished, almost as many as in the
#Sphærellaria#. In spite of this great variety of forms the legion
MONOPYLEA is probably monophyletic; at least all the different skeletal
forms may be derived from three elements which are combined in the most
manifold fashion; (1) the _sagittal ring_, a simple siliceous ring, which
lies vertically in the sagittal plane of the body, encircles the central
capsule and comes into contact with it at the basal pole of the main axis
(§ 124); (2) the _basal or oral tripod_, composed of three diverging radial
spines, which meet in the middle of the basal pole of the central capsule
(or in the centre of the porochora) (§ 125); (3) the _cephalis_, or
lattice-head, a simple ovoid or subspherical lattice-shell, which encloses
the central capsule and stands in connection with it at the basal pole of
its main axis. Any one of these three important structural elements of the
NASSELLARIAN skeleton may possibly be the starting-point {lxxvi}for all the
remaining forms of the MONOPYLEA; the great difficulty in their
phylogenetic derivation lies in the facts that, on the one hand, any one of
the three elements may alone constitute the skeleton, and on the other
hand, in the great majority of the legion, two or three are united together
(compare §§ 182-185).

112. _The Skeleton of the Phæodaria._--The skeleton of the PHÆODARIA or
CANNOPYLEA is always extracapsular, usually consists of a silicate of
carbon (more rarely of pure silica), and in the majority of the legion is
composed of hollow cylindrical tubes, whose siliceous wall is very thin,
and whose lumen is filled with gelatinous material (§ 127). The manifold
and remarkable skeletal forms occurring in this legion are not
monophyletic, since they cannot be derived from a common stem-form; they
are, on the contrary, polyphyletic, various skeletonless PHÆODARIA
(Phæodinida) have independently acquired skeletons of different form and
composition. The legion PHÆODARIA can be subdivided into four orders, the
skeletons of which present the following important distinctions:--(1) The
#Phæocystina# possess only incomplete Beloid skeletons (§ 115), composed of
many separate pieces, sometimes tangentially (Cannorrhaphida, Pl. 101),
sometimes radially arranged (Aulacanthida, Pls. 102-105). (2) The
#Phæosphæria# form Sphæroid skeletons (§ 116), usually only a simple
lattice-shell without special aperture (Pls. 106-111); two concentric
shells united by radial bars occur only in the Cannosphærida (Pl. 112). (3)
The #Phæogromia# are distinguished by the formation of a simple Cyrtoid
skeleton (§ 123) resembling that of the Monocyrtida; the monothalamus
lattice-shell is usually ovoid or helmet-shaped, more rarely polyhedral or
almost spherical; a vertical main axis can always be distinguished, at the
basal pole of which is an aperture usually armed with teeth or spines (Pls.
99, 100, 113-120). (4) The #Phæoconchia# are distinguished from all other
Radiolaria by the possession of a bivalved shell like that of the
Conchifera; the two valves of this Conchoid skeleton must be distinguished
as dorsal and ventral, as in the Brachiopoda (Pls. 121-128). The fifteen
families of PHÆODARIA which are arranged in the four orders just mentioned,
present such great differences among themselves, that the skeleton must be
regarded as probably polyphyletic even within the limits of each order.

113. _Types of Skeletal Formation._--No less than twelve different
principal forms may be distinguished as morphological types of the
formation of the skeleton in the Radiolaria; some of these are peculiar to
a single legion or even to a smaller group; but sometimes the same form
occurs in several legions. Some types occur only in an isolated manner,
independently of the others, but most exist in various combinations with
other types. Of the twelve described below the Conchoid and Cannoid occur
only in the PHÆODARIA; the Plectoid and Circoid only in the NASSELLARIA;
the Astroid only in the ACANTHARIA; the remaining seven types are found in
several legions in the same form and hence are polyphyletic.

{lxxvii}114. _The Astroid Skeleton._--Under the name "Astroid" we place the
peculiar star-shaped skeletons of the ACANTHARIA in opposition to those of
all other Radiolaria, for they are separated from them not only
fundamentally by reason of the chemical nature of their substance
(Acanthin, § 102), but also by their centrogenous origin, and the resulting
stellate form (Pls. 129-140). The ACANTHARIA are the only Radiolaria in
which the skeleton arises within the central capsule by the formation of
numerous rays or radial spines of acanthin which project on all sides from
the centre. Originally these are united at this point, their conical or
pyramidal points meeting and being supported one upon another. In the great
majority of ACANTHARIA this loose apposition is constant, so that when the
soft parts are destroyed the skeleton falls to pieces. Only in a few forms
in this legion are the central ends of the spines fused so that the whole
skeleton forms a connected star (_Astrolithium_). The small group
Chiastolida (or Acanthochiasmida) is characterised by the fact that the two
rays which are opposite to one another in each axis unite and form a
diametral bar. The skeleton is almost always composed of twenty radial
spines, which are regularly disposed (Icosacantha), only in the small
primitive group #Actinelida# is the number variable (Adelacantha, § 110).

115. _The Beloid Skeleton._--As Beloid or spicular skeletons are grouped
together all those which consist of several disconnected portions; these
always lie outside the central capsule, either within the calymma or on its
surface. Such extracapsular Beloid skeletons are entirely wanting in the
ACANTHARIA and NASSELLARIA; they occur only in the #Beloidea# among the
SPUMELLARIA, and in the #Phæocystina# among the PHÆODARIA; the individual
Beloid portions of the former are solid, those of the latter hollow. In
both groups the simplest forms of the separate portions are simple
unbranched needles (_Thalassosphæra_, _Thalassoplancta_, _Physematium_,
_Belonozoum_, among the SPUMELLARIA; _Cannobelos_ and _Cannorrhaphis_ among
the PHÆODARIA); usually these spicules are disposed tangentially over the
surface of the calymma. Among the #Beloidea# branched spicules occur more
commonly than these simple ones; they are either stellate (with many rays
united in a centre) or twin-like, with a tangential bar, from each pole of
which two or three (seldom more) radial branches project (Pls. 2, 4). Among
the PHÆODARIA the subfamily Dictyochida is characterised by the annular
shape of its Beloid portions, either simple rings, or hat-shaped or
pyramidal bodies with a latticed cap over the ring (Pl. 101, figs. 3-14;
Pl. 114, figs. 7-13). The family Aulacanthida among the PHÆODARIA, alone
possesses hollow _radial tubes_, which penetrate the whole calymma, and
project distally over its surface, whilst their proximal ends rest upon the
surface of the central capsule. Although in these cases the enclosed
proximal end is always simple, the free distal end develops the most
various processes in adaptation to its prehensile functions (Pls. 102-105).

{lxxviii}116. _The Sphæroid Skeletons or Lattice-Spheres._--The
"lattice-spheres" or sphæroid skeletons are the simplest and most primitive
forms of lattice shells, and are widely distributed in the three legions
SPUMELLARIA, ACANTHARIA, and PHÆODARIA, whilst they are entirely wanting in
the NASSELLARIA. The round lattice-shell is either a true sphere in the
geometrical sense, or an endospherical polyhedron, _i.e._, a polyhedron,
all whose angles lie in the surface of a sphere (§ 25). In general,
_primary_ and _secondary_ lattice-spheres may be distinguished, of which
the former are secreted on the outer surface of the primary, the latter on
that of the secondary calymma (§ 85). Furthermore, _simple_ and _compound_
lattice-spheres may be distinguished, the latter of which consist of two or
more concentric lattice-spheres firmly united by radial bars; in such cases
the innermost lattice-sphere is always to be regarded as the oldest or
primary, all the succeeding ones as secondary, and the outermost as the
youngest (§ 129). The simple lattice-spheres are usually to be regarded as
primary; they may, however, occasionally be secondary, in which case the
primary shell, originally enclosed, has been lost by degeneration (as, for
example, in the case of the Aulosphærida and some #Sphærellaria#).

117. _The Lattice-Spheres of the Spumellaria._--The lattice-spheres or
Sphæroid skeletons of the SPUMELLARIA exhibit in spite of their simple type
of structure, an extraordinary variety in the formation of the lattice-work
and radial apophyses, so that in the systematic portion of this work no
less than one hundred and seven genera and six hundred and fifty species
are distinguished; these are united in one suborder, the #Sphæroidea# (pp.
50-284, Pls. 5-8, 11-30). It may be divided into two main divisions, the
_Monosphærida_ with a single primary lattice-sphere (Pls. 12-14, 21, 26,
27), and _Pliosphærida_ (or Sphæroidea concentrica) whose skeleton consists
of two or more concentric lattice-spheres united by radial bars. The latter
are subdivided into Dyosphærida with two concentric lattice-spheres (Pls.
16, 19, 20, 22, 28); Triosphærida, with three spheres (Pls. 17, 24, 29);
Tetrasphærida, with four (Pls. 23, 30); Polysphærida, with five or more
(Pls. 15, 23); and Spongosphærida, with spongy lattice-spheres (Pls. 18,
25). A special group is made up of the simple lattice-spheres of the social
Collosphærida (or Sphæroidea polyzoa) (Pls. 5-8); these are usually more or
less irregular, and characterised by the development of peculiar tubular
processes; the latter are generally wanting in the Sphæroidea monozoa,
whose lattice-shell is very regularly formed. This distinction is
interesting and important, inasmuch as the regular lattice-spheres are
explained by the independent development of the free-swimming Monozoa,
whilst the irregular spheres are due to the mutual dependence of the social
Polyzoa.

118. _The Lattice-Spheres of the Acantharia._--The lattice-shells or
Sphæroid skeletons of the ACANTHARIA are immediately distinguishable from
those of all other Radiolaria by their centrogenous development and the
central union of the radial spines by which they are supported; the only
exception is furnished by the remarkable genus _Cenocapsa_ {lxxix}(Pl. 133,
fig. 11), in which the radial spines are absent, not primitively, however,
but in consequence of degeneration; for the twenty cross-shaped perspinal
pores, originally due to the twenty radial spines, are still present. In
the most nearly allied genera, _Porocapsa_ (Pl. 133, fig. 7) and
_Cannocapsa_ (Pl. 133, fig. 8), the proximal part of the twenty radial
spines is still present, while their distal portion has degenerated; hence
in this case they do not stand in direct communication with the spherical
shell. On the other hand, this primitive connection persists in the genera
_Astrocapsa_ (Pl. 133, figs. 9, 10), and _Sphærocapsa_ (Pl. 135, figs.
6-10). The five genera just mentioned form the peculiar family
Sphærocapsida (pp. 795-802); the spherical shell is in these cases composed
of very numerous small plates disposed like a pavement, each plate or aglet
being perforated by a pore canal; in addition to which there are twenty
larger (perspinal) pores (or twenty cross-shaped groups each of four
aspinal pores) at those important points where primitively the twenty
radial spines penetrate the calymma. This peculiar porous "pavement shell"
has probably been developed (independently of the twenty radial spines)
upon the calymma of the #Acanthonida# (_Acanthonia_, p. 749) by the action
of the sarcodictyum; it has, therefore, quite a different morphological
significance from the spherical lattice-shell of the Dorataspida, which is
composed of tangential apophyses of the twenty Acanthonid spines (pp.
802-847, Pls. 134-138). Each radial spine here forms either two opposite or
four crossed transverse processes, and since their branches spread over the
surface of the spherical calymma and are united suturally at their
extremities, the peculiar lattice-sphere of the Dorataspida arises. This
extensive family is again divided into two subfamilies:--the Diporaspida
(Pls. 137, 138) possess always only two opposite apophyses, and form by the
union of their branches two opposite primary apertures or aspinal meshes.
The Tessaraspida, on the other hand (Pls. 135, 138), have always four
crossed transverse processes, and form by their union four primary aspinal
meshes. From the Diporaspida are probably to be derived the Phractopeltida
(p. 847, Pl. 133, figs. 1-6), the only ACANTHARIA which possess a double
lattice-sphere; their double concentric spherical shell may be compared
with that of the Dyosphærida.

119. _The Lattice-Spheres of the Phæodaria._--The lattice-spheres or
Sphæroid skeletons of the PHÆODARIA, which are generally developed quite
regularly, though occasionally in a modified form, fall in the order
#Phæosphæria# into two groups of very different structure, each of which
includes two families. The first group (_Phæosphæria inarticulata_)
contains the families Orosphærida (Pls. 106, 107) and Sagosphærida (Pl.
108); the lattice-work of the former consists of irregular polygonal meshes
and very coarse, partially hollow trabeculæ; in the latter, on the other
hand, it consists of triangular meshes and very slender filiform trabeculæ;
in both families the whole sphæroid skeleton forms a single unsegmented
piece as in most #Sphæroidea#. In the second group of {lxxx}#Phæosphæria#
(_Phæosphæria articulata_), on the other hand, the lattice-sphere is
segmented in quite a peculiar manner, and composed of hollow cylindrical
tangential tubes, which are separated by astral septa at the nodal points
of the network; this remarkable structure characterises the two families,
Aulosphærida (Pls. 109-111) and Cannosphærida (Pl. 112); the segmented
lattice-sphere of the former is simple and hollow; while that of the latter
is connected by centripetal radial tubes with a simple concentric inner
shell, which is sometimes solid, sometimes latticed, and provided with a
main-opening corresponding to the astropyle of the enclosed central
capsule. Since in the Aulosphærida also, hollow centripetal radial tubes
project from the segmented lattice-sphere, it is possible that they have
been derived from the Cannosphærida by the loss of the primitive internal
shell. A special peculiarity of many #Phæosphæria# (_Oroscena_,
_Sagoscena_, _Auloscena_, &c.) consists in the fact that the whole surface
of the lattice-sphere is regularly covered with pyramidal or tent-shaped
prominences (Pl. 106, fig. 4; Pl. 108, fig. 1; Pl. 110, fig. 1). A simple
lattice-sphere quite similar to that of most Monosphærida also constitutes
the skeleton of the Castanellida (Pl. 113), but since it possesses a
special main-opening, it must be referred promorphologically to the Cyrtoid
shells of the #Phæogromia#.

120. _The Prunoid Skeleton or Lattice-Ellipsoid._--The "lattice-ellipsoids"
or Prunoid skeletons have arisen from the lattice-spheres or Sphæroid
skeletons by more energetic growth and elongation of one axis; this is the
main axis of the body and is probably always vertical; its two poles are
commonly equal. The Prunoid skeleton is either a true ellipsoid in the
geometrical sense or an "endellipsoidal polyhedron" (_i.e._, a polyhedron,
all the angles of which lie in an ellipsoidal surface). By further
elongation of the main axis, the ellipsoidal form passes over into the
cylindrical, the polar surfaces of the cylinder being usually rounded,
rarely truncated. The rich order #Prunoidea# (pp. 284-402) contains
numerous modifications of this form of shell which arise on the one hand by
the formation of transverse constrictions, on the other by the apposition
of concentric secondary shells. In respect of the latter, simple and
compound Prunoid shells can be distinguished as in the case of the Sphæroid
shells. In the compound Prunoid shells either all the concentric
lattice-shells may be ellipsoidal or the inner may be spherical. More
important differences are found in the transverse annular constrictions,
which give the Prunoid skeleton a segmented appearance; in this respect,
three principal forms may be distinguished (p. 288):--(A) _Monoprunida_,
with unsegmented shell, having no transverse constriction (Pls. 15-17); (B)
_Dyoprunida_, having a shell with two segments and one (equatorial)
transverse constriction (Pl. 39); (C) _Polyprunida_, with three or more
parallel transverse constrictions, by means of which the shell is divided
into four or more segments (Pl. 40). In the same manner as the #Prunoidea#
have arisen from the #Sphæroidea# among the SPUMELLARIA by greater
{lxxxi}development of the vertical main axis, the ellipsoidal Belonaspida
have arisen from the spherical Dorataspida among the ACANTHARIA (p. 859;
Pl. 136, figs. 6-9; Pl. 139, figs. 8, 9). The main axis of the ellipsoid in
this case is always occupied by the opposite equatorial spines of the
hydrotomical axis (pp. 719, 860). In the legion PHÆODARIA a similar
prolongation of the main axis rarely occurs; it is found, however, in
_Aulatractus_ (Pl. 111, figs. 6, 7), the lattice-shell of this Aulosphærid
being sometimes truly fusiform, sometimes rather ellipsoidal or even
double-conical.

121. _The Discoid Skeletons or Lattice-Discs._--The "lattice-discs" or
Discoid skeletons are characteristic of the SPUMELLARIAN group #Discoidea#,
and have arisen from the lattice-spheres of the #Sphæroidea# by a less
development of one axis, which is the main axis of the body, and is
probably usually vertical; its two poles are always equal. The Discoid
lattice-shell is either a biconvex lens (with a thin margin), or a plane
disc (a shortened cylinder with thick margin), or some form intermediate
between the two. All Discoid shells show a horizontal median plane or
equatorial plane, by which they are divided into two equal halves, an upper
and lower; the margin of the lens itself is originally the equator. The
main axis, the shortest of all the axes of the shell, stands vertically in
the centre of the equatorial plane. Among the PHÆODARIA Discoid shells
rarely occur (_Aulophacus_), as also among the ACANTHARIA (Hexalaspida).

122. _The Larcoid Skeleton or Lentelliptical Lattice-Shell._--The
lentelliptical lattice-shells, which may be shortly designated "Larcoid,"
are especially characteristic of the #Larcoidea#, a large order of
SPUMELLARIA (pp. 599-715; Pls. 9, 10, 49, 50). In addition they recur among
the ACANTHARIA, in the small family Hexalaspida (p. 872, Pl. 139), and the
family Diploconida (p. 881, Pl. 140), which is derived from it. These
lentelliptical lattice-shells are all characterised by the clear
differentiation of three unequal, but isopolar dimensive axes, _i.e._, the
three geometrical axes, perpendicular to one another, which determine the
form of the shell, are of unequal length; the two poles of each are,
however, equal. The geometrical ground-form is, therefore, a triaxial
ellipsoid (§ 34). In the rich order #Larcoidea# the lentelliptical
lattice-shell shows many variations in its development.

123. _The Cyrtoid Skeleton._--Cyrtoid skeletons are those lattice-shells
which possess a vertical main axis with two different poles (Monaxonia
allopola); the upper pole is usually termed the apical, the lower the
basal. Such Cyrtoid shells are characteristic of the great majority of the
NASSELLARIA or MONOPYLEA (and especially of the #Cyrtellaria#); they are
also found in a large division of the PHÆODARIA (the #Phæogromia#), and in
some SPUMELLARIA. In general the manifold Cyrtoid shells may be divided
into two large groups, those with one and those with several chambers. The
_monothalamous_ Cyrtoid shells are usually ovoid, conical, cap- or
helmet-shaped; their {lxxxii}internal cavity is simple, without
constrictions or septa. Among the NASSELLARIA they occur in the Monocyrtida
(Pls. 51-54, 98), where they have received the name "Cephalis." A form of
shell, essentially the same, is found amongst the PHÆODARIA in the order
#Phæogromia#, more especially in the Challengerida (Pl. 99), Medusettida
(Pls. 118-120), and Tuscarorida (Pl. 100), many of these latter closely
resembling many Monocyrtida. Such monothalamous Cyrtoid shells occur much
more rarely among the SPUMELLARIA (_e.g._, among the #Prunoidea# in
_Lithapium_, _Lithomespilus_, _Druppatractus_, Pls. 13, 14, &c.).
Polythalamous Cyrtoid shells (Pls. 55-80) occur exclusively in the
NASSELLARIA, and exhibit in this legion an astonishing variety of
structure; they are distinguished from the monothalamous forms by the
development of internal septa, or of annular incomplete diaphragms, which
usually correspond to the external constrictions; their interior is thus
divided into two or more communicating compartments. Among the
polythalamous Cyrtoid shells may be distinguished three principal groups,
the Stichocyrtid, Zygocyrtid, and Polycyrtid. Zygocyrtid shells are
characteristic of the #Spyroidea# (Pls. 84-90), and are distinguished by a
bilobate cephalis (cephalis bilocularis); the median sagittal ring, or a
corresponding constriction, divides the shell into right and left
compartments. Polycyrtid shells (Pl. 96) are peculiar to the #Botryodea#,
and characterised by a multilobate cephalis (cephalis multilocularis).
Stichocyrtid shells are those in which the primary cephalis remains simple,
and new joints are successively added to its basal pole; such shells occur
in the majority of the #Cyrtoidea#. Secondary chambers are sometimes added
in the other two groups (#Botryodea# and #Spyroidea#). When, as often
happens in these polythalamous Cyrtoid shells, two or three distinct joints
follow each other, the first is called the "cephalis," the second the
"thorax," and the third the "abdomen" (Tricyrtida Pls. 64-75).

124. _The Circoid Skeleton._--This is a very important and remarkable type
of skeletal formation, which occurs exclusively in the legion NASSELLARIA,
where it plays a very prominent part; its characteristic element is the
"sagittal ring," a simple, vertical, siliceous ring, which surrounds the
central capsule in its sagittal plane, and is specially differentiated in
its basal portion. This "primary sagittal ring" whose vertical allopolar
main axis coincides with that of the Monopylean central capsule embraced by
it, is characteristic of all members of the order #Stephoidea# (p. 931,
Pls. 81-83, 92-94); here it forms by itself the skeleton of the Stephanida
(Pl. 81); in the Semantida (Pl. 92) it is combined with a horizontal basal
ring, in the Coronida (Pls. 82, 93) with a vertical frontal ring and in the
Tympanida (Pls. 83, 94) with two horizontal rings, an upper mitral and a
lower basal. In the great majority of these #Stephoidea# there often
develop in definite places characteristic processes or apophyses, whose
branches combine to form a loose tissue or an incomplete lattice-shell.
This becomes complete in the #Cyrtellaria#, the majority of which retain
more or less {lxxxiii}distinct traces of the sagittal ring. Hence the
skeletons of all NASSELLARIA may be derived monophyletically (Hypothesis A,
p. 893) from a simple sagittal ring (_Archicircus_ and _Lithocircus_, Pl.
81). This theory, however, encounters the great difficulty that in many
#Stephoidea# (_Cortina_, _Cortiniscus_, &c.) it is combined in a remarkable
manner with the basal tripod of the #Plectoidea#, whilst in these latter it
is entirely wanting (compare p. 894).

125. _The Plectoid Skeleton._--Those forms are distinguished as Plectoid in
which three, four, or more radial siliceous spines proceed from a common
point, which lies excentrically outside the central capsule and at the
basal pole of its vertical allopolar main axis. This peculiar type of
skeletal formation only occurs in the legion NASSELLARIA, and is specially
characteristic of the order #Plectoidea# (p. 898, Pl. 91). But since the
essential elements of this remarkable skeleton also occur in many other
NASSELLARIA, sometimes combined with the Circoid, sometimes with the
Cyrtoid skeleton, it perhaps has a fundamental significance in this legion;
at all events it is possible to derive monophyletically all the other forms
of this legion from it (Hypothesis B, p. 893). The simplest form of the
Plectoid skeleton is a tripod, the three feet of which either lie in a
horizontal plane (_Triplagia_, Pl. 91, fig. 2), or correspond to the three
edges of a low pyramid (_Plagiacantha_). A fourth ray is sometimes added,
which stands vertically upon the summit of the pyramid (_Plagoniscus_,
_Plagiocarpa_, Pl. 91, figs. 4, 5). In other #Plectoidea# three secondary
rays are intercalated between the three primary (Hexaplagida, &c.); seldom
the number is greatly increased (Polyplagida, &c.). The rays are rarely
simple, but usually branched; in the Plagonida (Pl. 91, figs. 2-6) the
branches remain free; in the Plectanida (Pl. 91, figs. 7-13) they are
united to form a loose wicker-work. From such a web a perfect Cyrtoid shell
may arise. Several forms of Plagonida may also be readily confounded with
the isolated triradiate or quadriradiate spicula of many Beloid skeletons
(_Sphærozoum_, _Lampoxanthium_, &c.).

126. _The Spongoid Skeleton._--From the simple lattice-skeleton which the
majority of Radiolaria possess, some of them develop a spongy shell; the
trabeculæ of the lattice-work, situated in one plane in the former, are
developed in the latter in different planes and cross irregularly in all
directions; thus arises a kind of wicker-work of more or less spongy
structure, usually with very thin trabeculæ and irregular meshes. Such
Spongoid shells are most common among the SPUMELLARIA, especially in the
#Sphæroidea# (Spongosphærida, Pl. 18) and #Discoidea# (Spongodiscida, Pls.
41-47), more rarely in the #Prunoidea# and #Larcoidea#. Lattice-work of
similar spongy structure occurs very seldom among the NASSELLARIA, _e.g._,
in some #Plectoidea# (Pl. 91) and #Cyrtoidea# (_Spongocyrtis_,
_Spongopyramis_, _Spongomelissa_, &c., Pl. 56, fig. 10; Pl. 64, figs. 5-10,
&c.). Among the PHÆODARIA spongy skeletons are very rare; they {lxxxiv}are
to be seen in some #Phæosphæria# (_Oroplegma_, Pl. 107, fig. 1;
_Sagoplegma_, Pl. 108, fig. 2; _Auloplegma_, Pl. 111, fig. 8). No Spongoid
skeletons are known among the ACANTHARIA.

127. _The Cannoid Skeleton._--Cannoid or tubular skeletons are those which
are composed of hollow tubes; they occur exclusively in the PHÆODARIA or
CANNOPYLEA. Tubular processes, nevertheless, occur in some other
Radiolaria, as, for example, among the SPUMELLARIA in a portion of the
Collosphærida (_Siphonosphæra_, _Caminosphæra_, Pls. 6, 7), and of the
#Prunoidea# (_Pipetta_, _Cannartus_, &c., Pl. 39, figs. 6-10, &c.), also
among the NASSELLARIA in _Theosyringium_ (Pl. 68, figs. 4-6), _Cannobotrys_
(Pl. 96, figs. 3, 4, 8-11, 20-22), &c. In all these cases, however, the
tubes are direct processes of the cavity of the shell, the trabeculæ of the
lattice-work being solid. Only in the CANNOPYLEA are the lattice-bars
themselves, the radial spines and appendicular organs, generally tubular
(hence the designation "Pansolenia"). The lumen of the thin-walled
siliceous tubes is filled with jelly, and hence the specific gravity of the
relatively large skeleton is considerably diminished. This peculiarity is
not found in all CANNOPYLEA; it is wanting in all Sagosphærida and
Concharida, as well as in a part of the Orosphærida and Castanellida; in
the latter there are found intermediate stages between hollow and solid
skeletal rods. Very often a fine siliceous thread runs in the axis of the
tubes, which is connected with its wall by lateral branches (Pl. 110, figs.
4, 6; Pl. 115, figs. 6, 7). More seldom the tubes are divided by horizontal
septa into a series of chambers (Medusettida, Pls. 118-120). The two
families Aulosphærida (Pls. 109-111) and Cannosphærida (Pl. 112) are
distinguished from all other PHÆODARIA by the fact that their tubes are
separated by astral septa in the nodal points of the lattice-shell (§§ 112,
134).

128. _The Conchoid Skeleton._--By the name "Conchoid skeletons" are
distinguished the bivalved lattice-shells which occur exclusively in the
legion PHÆODARIA; they are quite characteristic of the #Phæoconchia# or
_Phæodaria bivalvia_, which embrace three families:--Concharida (Pls.
123-125), Coelodendrida (Pls. 121, 122), and Coelographida (Pls. 126-128).
The two valves of the lattice-shell of the Concharida are simple,
hemispherical, or boat-shaped, whilst in the Coelodendrida and
Coelographida tubes grow out from them, which branch and usually give rise
by anastomosis to a second external bivalved shell. In all #Phæoconchia#
the two valves are so disposed about the central capsule that an open slit
remains between them, into which open the apertures of the central capsule;
and since all these _Phæodaria conchoidea_ are TRIPYLEA, with three typical
openings in the central capsule, and since the two lateral accessory
openings lie at either side of the aboral pole, and the unpaired
main-opening at the oral pole of the main axis, it follows that the two
valves are to be regarded as dorsal and ventral as in the Brachiopoda (not
right and left as in the Lamellibranchiata). The dorsal and ventral
{lxxxv}valves are usually equal, but in a portion of the Concharida they
present constant differences. In this family the two valves are attached to
each other by their free edges, just as in the bivalved Mollusca and
Diatoms; and these edges may either be smooth (Conchasmida, Pl. 123, figs.
1-6), or dentate (Conchopsida, Pls. 124, 125); the valvular connection of
the latter is sometimes strengthened by a special ligament which unites the
two valves at the aboral pole (Pl. 123, figs. 8, 9). The form of the valve
is sometimes hemispherical, sometimes boat-shaped, with a sagittal keel.

129. _Medullary and Conical Shells._--In all Radiolaria whose skeleton
consists of a double shell or of two concentric lattice-shells united by
radial bars, an inner medullary shell (testa medullaris) and an outer
cortical shell (testa corticalis) may be distinguished (see note A, below).
The medullary shell is usually to be regarded as a primary, the cortical as
a secondary structure. Such double shells occur among the SPUMELLARIA in
the Dyosphærida (Pls. 19, 20), as well as in many #Prunoidea# (Pls. 39,
40), #Discoidea# (Pls. 33, 34), and #Larcoidea# (Pls. 9, 10); among the
ACANTHARIA only in the family Phractopeltida (Pl. 133); among the
NASSELLARIA only in very few #Cyrtoidea# (_e.g._, _Periarachnium_, Pl. 55,
fig. 11), and finally among the PHÆODARIA in the Cannosphærida (Pl. 112) as
well as in part of the Coelodendrida (Pl. 121) and Coelographida (Pls. 127,
128). In most cases (if not always?) the cortical shell arises by the
growth of radial spines from the surface of the medullary shell; these
become united at equal distances from the centre by transverse apophyses,
the surface of the secondary calymma furnishing the basis for their
secretion (§ 85). Nevertheless, it seems that in many #Sphærellaria# the
formation of the whole cortical shell proceeds simultaneously (at a
definite dictyotic period) like that of the primary medullary shell (see
note B). Whilst in the PHÆODARIA, ACANTHARIA, and NASSELLARIA, at most two
concentric shells are formed, in many SPUMELLARIA their number increases
continuously with additional growth; in many #Sphærellaria# it rises to
four, eight, or even more, as well as in many #Discoidea# (if the
concentric, peripherally disposed rings of chambers be regarded as
incomplete flattened shells). In these cases either only the innermost
primary lattice-shell is to be styled "medullary shell," or at most the two
innermost (inner and outer medullary shells), all the others being
cortical.

A. The distinction between medullary and cortical shells was originally
based in my Monograph (1862, p. 50) upon the topographical relation of
the lattice-shells to the central capsule, inasmuch as I regarded all
intracapsular shells as medullary, all extracapsular as cortical.
Hertwig, however (1879, p. 122), rightly pointed out that this
distinction is unpractical, "because the same lattice-shell in the same
species may lie within or without the central capsule, according to the
size of the latter." He proposes, therefore, to restrict the term
medullary shell to the innermost, and to call all the others cortical; a
course which seems justified by the special significance of the primary
innermost lattice-shell ("as the point of origin of the radial spines").
But in most #Sphærellaria# which form three or more concentric shells,
the two innermost, which lie near together within the {lxxxvi}central
capsule, are very different in size and dictyosis from all the others
which lie outside, and are separated by wider interspaces (compare Pls.
17, 24, 29-32, 40, &c.). In these cases it appears better to regard the
two inner as inner and outer medullary shells, and all the others as
cortical shells. The character of the dictyosis in the intracapsular and
extracapsular shells is often so different that I have made it the basis
of separation of _Thecosphæra_ and _Rhodosphæra_ among the Liosphærida
(p. 60), of Elatommatida and Diplosphærida among the Astrosphærida (p.
208), &c.

B.--R. Hertwig (1879, L. N. 33, pp. 40, 123) separates the true
(simultaneously formed) "cortical shells" (_e.g._, of _Actinomma_,
_Cromyomma_) from the arachnoid "siliceous networks" (_e.g._, of
_Diplosphæra_ and _Arachnosphæra_) which are formed by the successive
union of tangential apophyses of the radial spines. Whether this
principle is right in the theory or not, it cannot be carried out
practically. Compare also Pl. 25, fig. 4.

130. _Dictyosis or Lattice Formation of the Skeleton._--In the great
majority of Radiolaria the dictyosis or formation of lattice-work, and
especially the formation of a variously-shaped "lattice-shell," plays such
an important part that the whole class has long been popularly known in
Germany by the name "lattice animalcules" ("Gitterthierchen" or
"Gitterlinge") (_Protista dictyota_). The old name Polycystina also (1838),
although referring only to the SPUMELLARIA and NASSELLARIA, is derived from
the lattice-work of the siliceous skeleton. The extremely various forms in
which this is manifested furnish the means of distinguishing species. The
specific conformation of the skeletal lattice-work is usually caused by the
special disposition of the sarcodictyum (§ 94), whose exoplasmatic threads
become silicified or (in the ACANTHARIA) converted into bars of acanthin.
In many cases, however, the form of the lattice is mainly dependent upon
the situation and form of the radial spines or of special processes from
them. With respect to their origin, two varieties of lattice may be
distinguished--simultaneous and successive. _Simultaneous dictyosis_ occurs
especially in the simple lattice-shells of the #Sphærellaria# and
PHÆODARIA, where, at a given moment ("dictyotic moment") the _whole_
lattice of the shell is excreted on the surface of the calymma. _Successive
dictyosis_, on the other hand, is found more particularly in the
lattice-shells of the ACANTHARIA (and in the concentric cortical shells of
many #Sphærellaria#), which develop from the separate lattice-plates formed
by the apophyses of the radial spines, and hence not at the same moment.
The lattice-shells of the #Cyrtellaria#, which gradually grow out from a
sagittal ring or a basal tripod, arise by successive dictyosis.

131. _Dictyosis of the Spumellaria._--Siliceous lattice-structures are
wanting in the first section of the SPUMELLARIA, the #Collodaria#, but in
the second section, #Sphærellaria#, they are developed in extraordinary
variety of details. In spite of this extreme richness in different forms,
the lattice-shells of the SPUMELLARIA may all be derived from one and the
same primitive ground-form, a simple lattice-sphere with regular hexagonal
meshes (_Phormosphæra_, p. 61, Pl. 12, figs. 9-11; _Heliosphæra_, &c.).
{lxxxvii}The siliceous bars which bound these regular and subregular meshes
are at first exceedingly then and filiform; afterwards they become thicker
or spread out laterally, whence the meshes often become round with a
hexagonal frame (Pl. 12, fig. 5; Pl. 28, fig. 1). If the latter vanish, a
lattice-shell with simple circular meshes is formed. Very commonly the
regular form of the meshes or pores becomes more or less irregular,
polygonal, or roundish. Hence, in general, four different principal forms
of dictyosis may be distinguished among the SPUMELLARIA; viz. (1) regular
or subregular _hexagonal_ meshes; (2) regular or subregular _circular_
meshes; (3) irregular _polygonal_ meshes; (4) irregular _roundish_ meshes.
The three latter forms are to be regarded as secondary, derived from the
primary first form. In those SPUMELLARIA which possess several concentric
lattice-shells enclosed one within another, either these have all the same
form of dictyosis, or the lattice-work of the innermost primary shell is
different from that of the outer secondary shells (Pls. 19, 20); sometimes
these latter also differ more or less among themselves (§ 129).

132. _Dictyosis of the Acantharia._--The lattice-structures of the
ACANTHARIA differ essentially from those of other Radiolaria in several
particulars. Firstly, they consist not of silica but of acanthin (§ 102);
secondly, they are always secondary formations, usually developed from
transverse processes of the primary centrogenous radial spines; thirdly,
their formation is not simultaneous (at the same time over the same shell),
but successive (proceeding from the individual radial spines tangentially
towards the middle of the intervals); fourthly, the configuration of the
network is due to the relative position of the spines and the mode of union
of their transverse apophyses. Since they are at right angles to the
spines, and since the branches of the apophyses are at right angles to
them, the original ground-form of their dictyosis is a lattice-work with
quadrangular meshes; these are often quite regular and square (Pl. 130,
figs. 5, 6; Pl. 136, figs. 2, 9, &c.); more commonly they are rectangular
or irregularly quadrangular (Pl. 131, fig. 10; Pl. 133, figs. 2, 3, &c.).
In the majority of the ACANTHARIA the quadrangular form of the meshes
passes over into an irregularly polygonal or roundish one (Pls. 137, 138).
Very often the primary meshes of the lattice-shells, which immediately
surround the radial spines, are larger and more regular ("aspinal pores"),
whilst the numerous secondary meshes between them are smaller and irregular
("coronal pores"; Pl. 135, figs. 1-4, &c.).

133. _Dictyosis of the Nassellaria._--The siliceous lattice-structures of
the NASSELLARIA are formed on the whole like those of the SPUMELLARIA, with
which they were formerly united under the name "Polycystina." In this group
also there may be distinguished as two main forms the regular and
irregular. In the NASSELLARIA the regular lattice-structures generally
exhibit hexagonal or circular meshes, whilst the irregular are either
polygonal or roundish; the irregular forms are, however, much more abundant
than the {lxxxviii}regular, and a further distinction from the SPUMELLARIA
consists in the fact that the primary skeletal elements, from which the
lattice is secondarily developed, exercise a predominant influence upon
their form. These primary elements in the majority of the NASSELLARIA are
to be seen in two morphologically most important structures:--first, the
_primary sagittal ring_, which embraces the central capsule in the median
plane (§ 124); and secondly, the _basal tripod_ (§ 125), whose three
diverging rays proceed from the base of the central capsule, whilst
commonly a fourth vertical ray supports the dorsal side of latter (compare
Pls. 81-91, p. 892). In the majority of the NASSELLARIA these two primary
elements appear in combination, whilst in others only one of them is
recognisable. In addition there occur numerous monaxon lattice-shells in
which neither of these elements can be recognised, but a simple ovoid
lattice-shell (cephalis) alone forms the whole skeleton or its primary part
(Pl. 51, fig. 13; Pl. 98, fig. 13). The great difficulty in the
morphological interpretation and phylogenetic derivation of the
NASSELLARIAN skeleton lies in the fact that each of these three
elements--the primary sagittal ring, the basal tripod, and the latticed
cephalis--may form the whole skeleton by itself or be combined with one or
both of the others (p. 893). Even nearly related or at all events very
similar forms may differ very greatly in this respect. With regard to the
manifold forms of their dictyosis it follows that it is partly dependent
upon one of the two first elements, partly independent. In the
#Plectellaria# (or those NASSELLARIA which do not possess a complete
lattice-shell) the lattice-work is usually irregular and arises by union of
the ramifications, which proceed either from the primary sagittal ring
(Pls. 81, 82, 92-94) or from the basal tripod (Pl. 91). In the
#Cyrtellaria# (or NASSELLARIA with a complete lattice-shell, Pls. 51-80),
on the other hand, the lattice-work is sometimes regular, sometimes
irregular, being often very different in the different joints of a
segmented shell (Pl. 72); a great part of it arises independently of the
two chief morphological elements, and develops according to laws similar to
those which regulate the dictyosis of the SPUMELLARIA.

134. _Dictyosis of the Phæodaria._--The lattice-structures of the
PHÆODARIA, which consist of a silicate of carbon (§ 102), are on the whole
not developed in such variety as those of the other Radiolaria, but exhibit
several essentially different types of structure, not reducible to a common
primitive type of lattice-work. In one portion of this legion there occurs
an ordinary simple lattice-work (as in SPUMELLARIA and NASSELLARIA), with
solid trabeculæ; of these the Castanellida (Pl. 113) and Concharida (Pls.
123-125) have usually regular or subregular, circular meshes, sometimes
hexagonally framed; the Orosphærida (Pls. 106, 107) large irregular
polygonal meshes with thick trabeculæ, the Sagosphærida (Pl. 108) large
triangular meshes with thin filiform trabeculæ. The Challengerida (Pl. 99)
are characterised by a very delicate regular lattice-work, with minute
hexagonal pores, like a Diatomaceous frustule. The Medusettida (Pls.
118-120) {lxxxix}show a peculiar alveolar structure, numerous small
compartments being enclosed between two parallel plates. In the Circoporida
(Pls. 114-117) and Tuscarorida (Pl. 100) the opaque porcellanous shell has
a peculiar cement structure (§ 104), and the lattice-structure is confined
for the most part to characteristic rings of pores at the base of the
hollow tubes, which arise from the shell. The most peculiar lattice-work,
however, appears in the segmented shell of the Aulosphærida (Pls. 109-111)
and Cannosphærida (Pl. 112). In the former the large meshes of the
lattice-work are usually subregular and triangular, in the latter
polygonal; the trabeculæ are hollow cylinders, filled with jelly, and
containing usually a central axial thread. In each nodal point of the
lattice, in which three or more tangential tubes meet, these are separated
by stellate or astral septa.

135. _Radial Spines of the Skeleton._--The skeleton in the great majority
of Radiolaria is armed with radial spines, which are of great importance in
the development of their general form and of their vital functions. From a
morphological point of view the number, arrangement, and disposition of the
spines is usually the determining factor as regards the general form of the
skeleton. Physiologically they discharge distinct functions, as organs of
protection and support; they act also, like the tentacles of the lower
animals, as prehensile organs, since their points, lateral branches, barbed
hooks, &c. serve to hold fast nutritive materials. In general main-spines
and accessory spines may be distinguished in most Radiolaria; the former
are of pre-eminent importance in determining the figure of the skeleton;
the latter are merely appendicular organs. The main-spines present such
characteristic and important differences in the various legions of
Radiolaria that they must be considered separately.

136. _Radial Spines of the Spumellaria._--The radial spines, which exhibit
most manifold variations in the large order #Sphærellaria#, present
characteristic differences in its four suborders. In the #Sphæroidea# their
number and disposition serve for the separation into families (p. 59); the
Cubosphærida (Pls. 21-25) always possess six radial main-spines, which
stand opposite to each other in pairs and lie in three diameters of the
shell, which are at right angles to each other and correspond to the axes
of the regular crystallographic system. The Staurosphærida (Pl. 15) have
four spines, which form a regular cross and stand opposite to each other in
pairs, in two axes at right angles. The Stylosphærida (Pls. 13-17) show
only two main-spines, which are opposed to each other in the vertical main
axis of the body. Finally, the Astrosphærida (Pls. 18-20, 26-30) are
characterised by a larger and variable number of radial spines (eight,
twelve, twenty or more), sometimes regularly, sometimes irregularly
arranged. Among the other #Sphærellaria# the #Prunoidea# (Pls. 13-17, 39,
40) are most allied to the Stylosphærida with two opposite main-spines; the
#Discoidea# (Pls. 31-47), on the other hand, to the Staurosphærida with
four crossed spines; there exist, however, #Discoidea# with two opposite,
three marginal, or numerous radial main-spines; it is {xc}characteristic of
this suborder that they all usually lie in the horizontal median plane of
the lenticular shell, arising from its equatorial margin. The #Larcoidea#
(Pls. 9, 10, 49, 50) show a great variety in the number and arrangement of
their radial main-spines, which in the different families of this suborder
stand in direct causal relation to the various forms of growth of the
shell; usually the primary main-spines lie either in the three different
dimensive axes, at right angles to each other, whose differentiation is
characteristic of the lentelliptical Larcoid shell (§§ 34, 122) or in
definite diagonal axes, which cut the former obliquely. The radial spines
of the SPUMELLARIA are _never_ united in the centre of the body, but arise
separately from the surface of the primary central lattice-shell (medullary
shell), more rarely from one of the secondary (cortical) shells, which
enclose it. Their form is originally three-edged (sometimes pyramidal,
sometimes prismatic); the cause of this is to be found in their origin from
the nodal points of the lattice-shell, whose meshes are primitively
hexagonal; hence three trabeculæ unite in each nodal point, and are
produced into three edges of the spine. Very commonly, however, the spines
are round (conical or cylindrical), more rarely polygonal. The three edges
are often delicately toothed, not unfrequently spirally twisted around the
axis of the spine (Pl. 21, figs. 1, 12).

137. _Radial Spines of the Acantharia_.--The radial spines of this legion
have a much greater significance than in the other three classes of
Radiolaria, since here alone they are the primary determining factors in
the skeletal structure, and grow outwards from the middle of the central
capsule. This centrogenous origin of the radial spines is as characteristic
of the ACANTHARIA as their chemical constitution, which is not siliceous
but acanthinic (§ 102). Furthermore, their form is in most cases so
peculiar that even an isolated ACANTHARIAN spine can be generally
distinguished from one belonging to either of the other three legions. In
the great majority of the ACANTHARIA (all #Acanthonida# and
#Acanthophracta#) twenty radial spines are constantly present, which,
disposed according to a definite geometrical law, make up the skeleton
(compare § 110 above and p. 717). The twenty spines are generally simply
apposed to each other in the centre (either by the surfaces or the edges of
their pyramidal base); more rarely they are completely united and form a
single star-like piece of acanthin (_Astrolithium_). Very rarely
(_Acanthochiasma_) each two opposite spines are united so that ten
diametric bars cross in the middle of the central capsule. Whilst in the
great majority of ACANTHARIA these twenty radial spines are present, the
small group #Actinelida# is characterised by the possession of an
inconstant, often very large number, sometimes over one hundred. Among
these #Actinelida# are probably to be found the stem-forms of the whole
legion. The variously modified spines of the ACANTHARIA may be grouped in
three main categories: (1) round (cylindrical or conical); (2) four-edged
(prismatic or pyramidal); (3) two-edged (leaf- or sword-shaped). The latter
very commonly bear two {xci}opposite transverse processes, the former four
crossed ones. By ramification and union of these apophyses arise the
lattice-shells of the #Acanthophracta# (excepting the Sphærocapsida).

138. _Radial Spines of the Nassellaria._--The radial spines in this legion
show as great a variety in their form as in the SPUMELLARIA, and, as in
them, are solid, siliceous bars, usually three-edged (prismatic or
pyramidal), or round (cylindrical or conical); more seldom they are
polygonal in section. The great majority of the NASSELLARIA are, however,
distinguished by a triradial structure, three primary radial bars diverging
from the base of the central capsule (usually from the centre of the
porochora); there is usually in addition a fourth apical spine, which
passes upwards vertically or obliquely on the dorsal aspect of the central
capsule. These three or four typical radial spines of the NASSELLARIA may
be derived with great probability from the basal tripod of the #Plectoidea#
(_Plagoniscus_, _Plectaniscus_, &c., Pl. 91); and since this tripod is very
characteristically combined in _Cortina_ and _Cortiniscus_ with the primary
sagittal ring of the #Stephoidea#, the three typical rays may be generally
designated "cortinar feet," in contradistinction to the other radial
processes of the NASSELLARIAN skeleton. One of the three descending basal
feet ("pes caudalis," Pls. 91-95, C) is always unpaired, and lies in the
vertical median plane (or sagittal plane), just as does the vertically
directed apical spine, which originally forms the dorsal bar of the
sagittal ring, and is produced upwards into the "apical horn," (marked _a_
on the plates). The other two basal feet are paired, and diverge right and
left, forwards and downwards ("pedes pectorales," _p.p._). Six-rayed
NASSELLARIA, in which three secondary (interradial) feet are intercalated
between the three primary (perradial) cortinar feet, are less common than
the three-rayed forms. In some groups the number rises still higher, nine,
twelve, or even more secondary feet being intercalated between the three
primary. Besides, accessory radial spines may be developed on different
parts of the shell, which have sometimes a definite relationship to the
typical radial spines, sometimes not. Their form and ramification are very
various (Pls. 51-98).

139. _Radial Spines of the Phæodaria._--The radial spines of the PHÆODARIA
are very clearly distinguished from those of other Radiolaria by the fact
that they are usually hollow tubes, rarely solid bars. As a rule, the tubes
are cylindrical, often slightly fusiform or conical, their siliceous wall
is very thin, and their lumen filled with jelly; a fine thread of silica
usually runs in the axis, and in several families is connected by fine
transverse threads with the wall of the tube (Pl. 110, figs. 4, 6; Pl. 115,
figs. 6, 7). The peculiar family Medusettida is characterised by a very
remarkable segmentation of the hollow spines (Pls. 118-120). Each tube is
divided by a series of septa into chambers, which communicate by a central
or excentric opening in each septum, an arrangement resembling the
siphuncle of the chambered Cephalopod shells. The number and
{xcii}arrangement of the radial tubes in most PHÆODARIA is indefinite and
very variable; only in a few families is the number constant in each
species and genus, and the disposition regular. The Medusettida (Pls.
118-120) resemble the NASSELLARIA, inasmuch as equal radial feet diverge
from the base of their shell, sometimes three in number (_Cortinetta_, Pl.
117, fig. 9), sometimes four (_Medusetta_, Pl. 120, figs. 1-4), sometimes
six (_Gazelletta_); _Gorgonetta_ is specially distinguished by the
possession of six ascending and six descending spines regularly alternating
(Pl. 119). The Tuscarorida (Pl. 100) usually have three or four equidistant
feet. The Circoporida (Pls. 115-117), on the other hand, rather approach
the #Sphæroidea#, their spherical or regular polyhedral shell having a
definite number of tubular radial spines, which arise at regular intervals
from their angles; _Circoporus_ has six, _Circospathis_ nine, _Circogonia_
twelve, and _Circorrhegma_ twenty radial tubes. Very rarely the tubes of
the PHÆODARIA are angular, usually they are round, more or less
cylindrical, though they are often bifurcated or even ramified, and exhibit
a great wealth of the most delicate appendages; siliceous hairs, bristles,
spines, barbed or anchor-like hooks, spathillæ, brushes, circlets, &c.
(compare Pls. 99-128).

140. _Main-Spines and Accessory Spines._--As accessory spines (Paracanthæ)
we oppose to the main-spines (Protacanthæ), just described, all those
processes which have no determining influence upon the formation of the
skeleton as a whole, but are to be regarded as secondary constituents of
the skeleton, or appendicular organs of inferior significance. They are
developed in the utmost variety, sometimes as hairs or bristles, sometimes
as thorns or clubs, either straight or curved (often zigzag), smooth or
barbed; sometimes standing vertically upon the shell, or directed towards
the centre, sometimes obliquely, or rising at a definite angle. In those
SPUMELLARIA whose lattice-shell consists of several concentric spheres, the
accessory spines generally arise from the outermost, the main-spines, on
the contrary, from the innermost. In the NASSELLARIA, multifarious forms of
accessory spines are especially developed in the order #Plectellaria#. In
the PHÆODARIA they are often furnished with delicate appendages, _e.g._,
anchor-hooks, spathillæ, coronets, &c. Among the ACANTHARIA the accessory
spines which arise from the surface of the shell in the #Acanthophracta#
are very characteristic. They are not radially disposed (like the similar
superficial spines of the SPUMELLARIA), but parallel to the radial
main-spines from whose transverse processes they arise. Since in all these
#Acanthophracta# the twenty radial main-spines are opposite to each other
in pairs, all the accessory spines (often several hundred) are parallel to
ten different regularly disposed axes of the lattice-shell (Pls. 134-138).

The skeletons of the Radiolaria, in addition to the general relations
which have been discussed above, present numerous and important special
differences in the various larger and smaller groups. These are indicated
in detail in the descriptions of the legions, orders, and families in the
systematic portion of this Report.

{xciii}BIOGENETICAL SECTION.

A SKETCH OF OUR KNOWLEDGE OF THE DEVELOPMENT OF THE RADIOLARIA IN THE YEAR
1884.

----

CHAPTER V.--ONTOGENY OR INDIVIDUAL DEVELOPMENT.

(§§ 141-152.)

141. _Individual Developmental Stages._--The germinal history of the
Radiolaria presents great obstacles to direct observation, and hence is
very incompletely known. The fragmentary observations, however (having been
made on Radiolaria of very various groups and supplemented by comparative
anatomical considerations), allow us to draw a general picture of the
essential developmental processes in this great class. It may probably be
assumed that in all Radiolaria, after maturation, the central capsule
discharges the function of a sporangium, and its contents are broken up
into numerous flagellate swarm-spores (zoospores). After these flagellate
swarm-spores (resembling _Astasia_) have emerged from the ruptured central
capsule, they probably pass over into a _Heliozoan_-stage (_Actinophrys_)
and then after the formation of a jelly-veil into the condition of
_Sphærastrum_. Afterwards, when a membrane is formed between the outer
jelly-veil and the inner nucleated cell-body, an _Actissa_-stage arises,
which exhibits in its simplest form the differentiation of the spherical
unicellular body into the central capsule and calymma. _Actissa_ thus
represents both ontogenetically and phylogenetically the primitive
condition of the Radiolarian organism, and may thus be regarded as the
point of departure of all other forms.

142. _The Astasia-Stage._--The formation of flagellate zoospores in the
mature central capsule is probably to be regarded as the common form of
individual development in all Radiolaria; since the whole contents are
utilised in the formation of these swarm-spores, and since the
extracapsulum takes no share in the process and perishes after they are
evacuated, the _central capsule_ may be regarded as a _sporangium_ (see
note A, below). The zoospores of the Radiolaria generally arise in the
following way:--the nucleus of the unicellular organism, sometimes early,
sometimes late (and in several different ways, §§ 63-70) breaks up into
numerous small nuclei, and each of these surrounds itself with a small
portion of the endoplasm. Very often, perhaps generally, this endoplasm
contains one or several fat-granules and sometimes also a small oblong
crystal; from the protoplasm {xciv}of the small roundish or ovoid cells
protrudes one or more vibratile flagella. The fully developed spores, which
commence their vibrations even within the central capsule, emerge when it
ruptures, and swim about freely in the surrounding water by means of the
flagellum. At this stage of its existence the young Radiolarian represents
essentially the simplest form of the Flagellata, such as _Astasia_ or
_Euglena_; the unicellular body is for the most part ovoid or
subcylindrical, sometimes fusiform or reniform, usually from 0.004 to 0.008
mm. in diameter (Pl. 1, fig. 1_c_; Pl. 129, fig. 11). In the anterior part
of the flagellate cell, immediately behind the base of the flagellum, lies
a homogeneous, spherical nucleus, whilst in the posterior part are usually
several small fat-granules and often also a small oblong crystal (hence the
name "crystal-spore," "Krystall-Schwärmer"). The number of vibrating
flagella, which are extremely long and fine, seems to be variable, usually
one, sometimes two, occasionally perhaps three, or even four or more (see
note B).

A. The formation of the motile spores in the central capsule was first
observed by J. Müller in _Acanthometra_ (1856, L. N. 10, p. 502), then by
A. Schneider in _Thalassicolla_ (1858, L. N. 13, p. 41), and finally by
myself in _Sphærozoum_ (1859, L. N. 16, p. 141). These older observations
were, however, incomplete, for the origin of the motile corpuscles from
the contents of the central capsule was not observed. The first complete
and detailed observations upon the formation of spores in the Radiolaria
were published in 1871 by Cienkowski (L. N. 22, p. 372, Taf. xxix.); they
relate to two different Polycyttaria, _Collosphæra_ and _Collozoum_.
These investigations were supplemented by R. Hertwig on _Collozoum_ and
_Thalassicolla_ (1876, L. N. 26, pp. 28, 43, &c.); on _Collozoum_ he made
the important discovery that the Polycyttaria form two kinds of spores,
one with and the other without crystals, and that the latter are divided
into macrospores and microspores (compare the chapter on "Reproduction,"
§§ 212-216). Quite recently Karl Brandt has confirmed these observations,
and has extended them to all the genera of Polycyttaria (1881, L. N. 38,
p. 393, and 1885, _loc. cit._).

B. The number of flagella, projecting from each spore, is very difficult
to determine, owing to their extraordinary length and slenderness. It
appeared to me that in the majority of those Radiolaria whose spores I
investigated only a single flagellum could be demonstrated with
certainty, although sometimes two, springing from a common base, seemed
to be present. Compare the chapter on "Reproduction," (§ 215) and the
recent work of Karl Brandt on Sphærozoea (1885, L. N. 52, pp. 145-174).

143. _The Actinophrys-Stage._--The fate of the flagellate zoospores which
emerge from the mature central capsule of the Radiolaria has not hitherto
been decided by actual observation; all attempts to rear the swarming
zoospores have been in vain, for they have soon died. From what we know,
however, of the comparative morphology of the Protista, the hypothesis is
fully justified, that between the _Astasia_-stage of the flagellate
swarm-spores, and the well-known _Actissa_-stage of the simplest
Radiolaria, there lies an intermediate developmental stage, which may be
regarded as being essentially the simplest Heliozoan form, _Actinophrys_ or
_Heterophrys_. The swarm-spore is very probably converted directly in to a
simple floating _Heliozoon_ by its elongated or ovoid body {xcv}becoming
spherical and by fine pseudopodia protruding all round instead of a single
flagellum; the nucleus at the same time assuming a central position.

144. _The Sphærastrum-Stage._--The _Actinophrys_-stage of the young
Radiolaria, which proceeds immediately from the flagellate zoospore, is
probably connected with the _Actissa_-stage by an intermediate form, which
may be regarded as a simple skeletonless _Heliozoon_ with a jelly-veil; a
well-known example of such a form is _Sphærastrum_ (in the solitary, not
the social condition) and _Heterophrys_. This important intermediate form
has arisen from the simple _Actinophrys_-stage by the excretion of an
external structureless jelly-veil, such as is formed in many other Protista
(_e.g._, in the encystation of many Infusoria). The young Radiolarian in
this second _Heliozoon_-stage becomes a simple cell with pseudopodia
radiating on all sides; its body consists of three concentric spheres, the
central nucleus, the protoplasmic body proper, and the surrounding calymma
or jelly-veil. When a firm membrane is developed between the last two
spheres this _Sphærastrum_-stage passes over into the _Actissa_.

The gap in our empirical knowledge which still exists between the
flagellate stage (§ 142) and the simplest Radiolarian stage (_Actissa_, §
145), can be filled hypothetically only by the assumption of several
_Heliozoon_-stages following one upon another. It is possible also that
the capsule-membrane is not formed between the endoplasm and exoplasm (as
here supposed), but that the membrane was formed first outside the cell
and the extracapsulum subsequently secreted around it.

145. _The Actissa-Stage._--The first SPUMELLARIAN genus, _Actissa_, is not
only the simplest form actually observed among the Radiolaria, and the true
prototype of the whole class, but also the simplest form under which the
Radiolarian organisation can be conceived. It is therefore extremely
probably that _Actissa_ not only forms the common stem-form of the whole
class in a phylogenetic sense, but is also its common ontogenetic or
germinal form. Probably in all Radiolaria the _Sphærastrum_-stage develops
immediately into the typical _Actissa_-stage, by the formation of a firm
membrane between the protoplasmic body of the spherical Heliozoan cell and
its jelly-veil. Thus arises the characteristic central capsule, which is
wanting in the nearly related Heliozoa. It is further probable that all
Radiolaria in their early stage will so far conform to the state of things
in _Actissa_ as to have the capsule-membrane of the spherical skeletonless
cell perforated everywhere by fine pores. This structure is retained in all
SPUMELLARIA, whilst in the other three legions those structural relations
of the capsule which are characteristic of each develop from the
_Actissa_-stage.

146. _The Ontogeny of the Spumellaria._--In the simplest case the
individual development in the SPUMELLARIA ceases with the _Actissa_-stage.
In all other genera of this legion diverging forms proceed from this, of
which the different growth of the three dimensive {xcvi}axes on the one
hand (§§ 44, 45), and the differentiation of the various parts of the
unicellular organism with the formation of the skeleton on the other, are
of pre-eminent significance. Even in the varying growth of the central
capsule in the different dimensions of space in the skeletonless
#Colloidea#, four different modes may be distinguished, which further, in
the corresponding development of the skeleton, furnish the basis for the
origin of the four orders of #Sphærellaria#. The most primitive and
simplest form of growth, equal extension in all directions, is found in the
spherical central capsule and the concentric spherical skeletons
(_Procyttarium_, #Sphæroidea#). When the growth of the central capsule
proceeds more rapidly in the direction of the vertical main axis than in
any other direction, the ellipsoidal or cylindrical central capsule
(_Actiprunum_) arises, and the vertically elongated skeleton of the
#Prunoidea#, which is derived from it. When, on the contrary, the growth of
the central capsule and lattice-shell is less in the direction of the
vertical main axis than in any other direction, the lenticular or discoid
central capsule (_Actidiscus_) arises, and the corresponding lenticular
shell of the #Discoidea#. Finally, even quite early in many SPUMELLARIA,
the growth of the central capsule and of the corresponding lattice-shell in
the three dimensive axes is different, and hence arise the lentelliptical
forms whose geometrical type is the triaxial ellipsoid or the rhombic
octahedron (_Actilarcus_, #Larcoidea#). Thus the origin of the four orders
of #Sphærellaria# is simply explained by a varying growth in the different
dimensive axes. The _primary_ (innermost) lattice-shell is in this legion
always _simultaneously_ developed (suddenly excreted at the moment of
lorication from the sarcodictyum). The _secondary_ lattice-shells, on the
other hand, which surround the former concentrically, and are united with
it by radial bars, arise _successively_ from within outwards.

147. _The Ontogeny of the Acantharia._--The individual development of the
ACANTHARIA in the simplest case (_Actinelius_) stops at a point which
differs from the _Actissa_-stage only in the change of radial axial threads
into acanthin spines. In the small group #Actinelida#, their number remains
variable and usually indeterminate (Adelacantha), whilst in the great
majority of the legion (#Acanthonida# and #Acanthophracta#) the number is
constantly twenty, and those spines are regularly arranged according to the
Müllerian law in five parallel circles, each containing four crossed spines
(Icosacantha). The simplest form among these latter is _Acanthometron_,
which may be regarded both ontogenetically and phylogenetically as the
common starting-point of all the Icosacantha. Within this extensive group
variations in the length of the dimensive axes appear, similar to those
observed in the SPUMELLARIA. In the Astrolonchida and #Sphærophracta# the
central capsule remains spherical, extending equally in all directions; and
correspondingly the lattice-shell, which is excreted on the surface of the
spherical calymma, remains spherical. In the Belonaspida (just as in the
#Prunoidea#) {xcvii}this form passes over into an ellipsoid by prolongation
of one axis; on the contrary, in the Hexalaspida (as in the #Discoidea#)
the discoidal or lenticular form arises by shortening of an axis. Finally,
in the Diploconida, and in some Hexalaspida in which the growth is
different in all three dimensive axes (as in the #Larcoidea#), both the
central capsule and the shell assume the lentelliptical form. The
lattice-shell of the #Acanthophracta# is usually successive in its
development, since from each of the twenty radial spines two or four
tangential apophyses proceed, whose branches subsequently unite and combine
to form the lattice-shell. Only in the peculiar Sphærocapsida can the
pavement-like shell arise simultaneously or in a moment of lorication.

148. _The Ontogeny of the Nassellaria._--The individual development of the
NASSELLARIA in the simplest instance remains stationary at the skeletonless
Nasselid stage (_Cystidium_, _Nassella_), which can be immediately derived
from the foregoing _Actissa_-stage by the disappearance of the pores in the
upper (apical) hemisphere of the central capsule, whilst in the lower
(basal) portion they are modified to form a porochora; the podoconus is
developed within the endoplasm upon this latter. Usually the spherical form
of the central capsule passes over into an ovoid or ellipsoidal one, the
vertical axis which passes through the centre of the porochora being
elongated. From the skeletonless Nassellida the other NASSELLARIA may be
derived both ontogenetically and phylogenetically by the excretion of an
extracapsular siliceous skeleton. Unfortunately, the earliest stages in the
formation of this skeleton are unknown, and hence no answer can at present
be given to the important question, in what order the three primary
skeletal elements of the NASSELLARIA (the basal tripod, sagittal ring, and
latticed cephalis) appear (compare §§ 111 and 182). If, for example, in
_Cortina_ and _Tripospyris_ the basal tripod were to appear first in the
ontogeny, and the sagittal ring were developed from this, then the
#Plectoidea# would be rightly considered to be the oldest forms in the
phylogeny of the skeleton-forming NASSELLARIA; and in the contrary case the
#Stephoidea# would be so regarded. The relations of growth in the three
dimensive axes are very variable in the NASSELLARIA; the three most
important factors in this respect (partly separately and partly in
combination) are; (1) the development of the basal tripod to a triradial
stauraxon form (the ground-form being a three-sided pyramid); (2) the
development of the sagittal ring in the median plane of the body (the
vertical axis having the poles different); (3) the development of the
latticed cephalis outside the central capsule (the poles of the vertical
axis being again different). Since the development both of the skeleton and
of the malacoma is characterised in most NASSELLARIA by the stronger growth
of the vertical axis and the differentiation of the two poles, the
allopolar monaxon ground-form acquires a predominant significance in this
legion (§ 32); the starting point of most of the further modifications is
the basal pole of the vertical main axis. Next to this the sagittal axis is
usually the most important determining factor (its dorsal and ventral poles
being {xcviii}usually different), more rarely the frontal axis (with equal
right and left poles). In the zygothalamous #Spyroidea# (as in the
#Stephoidea#) the formation of the shell proceeds from the sagittal ring,
whilst in the polythalamous #Cyrtoidea# the latticed cephalis is always the
starting point, from which a series of joints (thorax, abdomen, and in the
Stichocyrtida, the numerous post-abdominal joints) successively arise
(unipolar growth).

149. _The Ontogeny of the Phæodaria._--The individual development of the
PHÆODARIA in the simplest case stops with the skeletonless condition of the
Phæodinida (_Phæodina_, _Phæocolla_), which can be immediately derived from
the foregoing _Actissa_-stage by the disappearance of the pores in the
greater part of the central capsule, the characteristic astropyle being
developed at the basal pole (§ 60). Since this particular form and
structure of the spheroidal central capsule remains the same in all
PHÆODARIA, whilst the formation of their skeleton follows very different
directions, it follows that further common paths of development are
excluded both ontogenetically and phylogenetically. What will be laid down
in this respect as regards the phylogeny of the different groups of
PHÆODARIA (§§ 194-199) holds true also of their ontogeny. The relations of
growth in the three dimensive axes are hence very different in the
skeletons of the various groups of PHÆODARIA. This difference is best
marked in the #Phæoconchia#, whose bivalved lattice-shells have as their
ground-form the rhomboid pyramid of Ctenophora. In most #Phæogromia# the
monaxon lattice-shell may develop simultaneously by sudden excretion at a
particular moment of lorication; this is also the case with the polyaxon
lattice-shells of the #Phæosphæria#. In their future growth the development
of basal or radial apophyses is of special importance. In the majority of
the PHÆODARIA these apophyses are tubes of silicate filled with jelly
(often provided with an axial siliceous thread); thus their development is
distinguished by complications which are absent in the case of the other
three legions.

150. _Growth._--The growth of the body in the Radiolaria, as in all other
organisms, is the fundamental function of individual development (see note
A). All structural relations which this richest class of the Protista
exhibits may be referred to different forms of growth, either of the
unicellular malacoma or of the skeleton which it produces. In general the
special development of the skeleton is dependent upon that of the central
capsule, and of the sarcodictyum on the surface of the calymma; in the
further growth, however, the conditions are reversed, and the condition of
the skeleton already formed directly determines the further development of
the central capsule and of the calymma with its sarcodictyum. The four
legions of Radiolaria show, speaking generally, certain characteristic
differences in growth, which are due in great part to the different
structure and ground-form of their central capsule. In the two legions of
the Porulosa (SPUMELLARIA and ACANTHARIA), in which the central capsule is
originally spherical and {xcix}the ground-form of the skeleton either
polyaxon or isopolar monaxon, two fundamental and variously combined
directions of growth are recognisable; firstly, the _concentric_ growth
(equal increase of volume in all directions), and secondly, multipolar or
_diametral_ growth (hypertrophy of certain parts in the direction of
definite pairs of radii). A different state of things obtains, however, for
the most part, in the two legions of the Osculosa (NASSELLARIA and
PHÆODARIA), in which the central capsule possesses a vertical main axis
with different poles, and the structure of the skeleton is determined by
this allopolar monaxon ground-form. The two fundamental directions of
growth here combined in the most various ways are, firstly, _unipolar_
growth (starting from the basal pole of the vertical main axis), and
secondly, radial or _pyramidal_ growth (characterised by the different
development of separate parts in the direction of definite radii). Whilst
the growth of the _malacoma_ is dependent on intussusception (as in most
organic structures capable of imbibing), the growth of the _skeleton_ in
all Radiolaria takes place by apposition (see note B).

A. The earliest investigations into the modes of growth in the Radiolaria
are due to J. Müller (L. N. 12, pp. 21-33). More detailed communications
I gave myself in my Monograph (L. N. 16, pp. 150-159). The relations
there sketched have now, in consequence of the examination of the
Challenger collection, undergone many important additions, and in some
divisions, important modifications; these are for the most part treated
of in the general account of the separate families.

B. The view here maintained, that the skeleton of all Radiolaria grows
only by apposition, appeared formerly to have certain exceptions. I
thought I had shown that in _Coelodendrum_ the thin-walled tubes grew not
only in length but also in thickness, with continuous increase in the
lumen (L. N. 16, pp. 152, 360). Further K. Brandt concluded, from the
varying size of the median bars in the twin-spicules of _Sphærozoum_,
that these siliceous structures grow by intussusception (L. N. 38, p.
401). Both suppositions have been proved erroneous and I have come to the
opinion that in all Radiolaria the skeleton grows by apposition.

151. _Regeneration._--Whilst the general course of individual development
(perhaps without any exception in the Radiolaria), begins with the
formation of zoospores in the central capsule, there yet occurs in some
groups a different form of ontogeny, introduced by simple division of the
unicellular organism, and coming under the term "regeneration" in its wider
sense. This spontaneous division occurs quite commonly in the Polycyttaria
(or social SPUMELLARIA), and produces their colonies (compare the chapter
on Reproduction, § 213). On the contrary, it has not been observed in the
solitary SPUMELLARIA, nor in the ACANTHARIA and NASSELLARIA; possibly,
however, the peculiar ACANTHARIAN family, Litholophida, has arisen by the
division of Acanthonida (compare p. 734). Among the PHÆODARIA division is
commonly observed in the order #Phæocystina# (which have an incomplete
Beloid skeleton or none), and also in the #Phæoconchia#. In all these cases
the increase by division is nothing else than an ordinary case of
cell-division, in which bisection of the nucleus precedes that of the
central capsule. The regeneration by {c}which each of the two
daughter-cells develops to a complete mother-cell depends upon simple
growth. Another form of regeneration, different from this, has been
observed in _Thalassicolla_. If the central capsule be extracted
artificially from the large concentric calymma, the enucleated central
capsule produces a new extracapsulum, with sarcomatrix, pseudopodia, and
calymma. This experiment may be repeated several times with the same
result. (Compare A. Schneider, 1867, L. N. 20.)

152. _The Formation of Colonies._--The individual development of colonies
takes place in all three families of the Polycyttaria (Collozoida,
Sphærozoida, Collosphærida) in the same simple way, by the repeated
division of a single monozootic SPUMELLARIAN. Since these divisions only
affect the central capsule and not the extracapsulum, the sister-cells,
which arise by repeated division of the mother, remain enclosed in a common
rapidly growing calymma. Probably in all Polycyttaria the commencement of
the formation of colonies immediately follows the _Actissa_-stage of the
monozootic mother-cell (or takes place in the _Thalassicolla_-stage, which
arises from the former by the development of alveoles in the calymma). The
simple central nucleus separates (by direct nuclear division) into two
halves, and the central capsule follows this process of bisection, becoming
constricted in the middle between the two daughter nuclei (Pl. 3, fig. 12).
In the further growth of the colony the process of division proceeds in the
older, now multinucleate, central capsules, in which an oil-globule has
taken the place of the original nucleus; then the division of the
oil-globules precedes that of the central capsule (Pl. 5, fig. 1). Another
mode of growth of the colonies is the multiplication of the central
capsules by gemmulation, or the formation of the so-called "extracapsular
bodies" (Gemmulæ, § 214). The characteristic skeletal structure of the
different species appears at a later stage. Whether ripe central capsules
can emerge from the social bond of a coenobium, and, having become
isolated, establish the formation of a new colony, is very doubtful. The
various forms which the coenobium assumes in the different species of
Polycyttaria, are due partly to simple growth, partly to the development of
large vacuoles in the calymma.

The _form and size_ of the coenobia appear in many fully developed
Polycyttaria to exhibit specific differences, which require further
investigation; in the young stage, on the contrary, they are simple
spheres or ellipsoids, often cylindrical or sausage-shaped (Pl. 3, figs.
1, 4, 6, 11). In some species the cylindrical gelatinous bodies become
moniliform, and separated by transverse constrictions into many segments,
each of which encloses a large alveole (Pl. 3, fig. 10). The rare
ring-shape (Pl. 4, fig. 1) which I figured in 1862 in the case of
_Collozoum_ (L. N. 16, p. 522, Taf. xxxv. fig. 1), I have recently
observed in different species of Polycyttaria; it is capable of a very
simple mechanical explanation, both ends of a sausage-shaped colony
having been accidentally brought into contact by a wave and having united
by agglutination. Quite recently Brandt has given a very complete account
of the development, form, and growth of Polycyttarian colonies in his
work on the colonial Radiolaria of the Bay of Naples (1885, L. N. 52, pp.
71-85).

{ci}CHAPTER VI.--PHYLOGENY OR GENEALOGICAL DEVELOPMENT.

(§§ 153-200.)

153. _Sources of Phylogenetic Knowledge._--For the purpose of constructing
a hypothetical genealogical tree of the Radiolaria, as of all other
organisms, three sources of information are open to us, viz., palæontology,
comparative ontogeny, and comparative anatomy. In the present case,
however, these three sources are of very different value; the first two are
at present only very inadequately known and have only been partially
investigated, hence they can only be utilised to a very slight extent. The
comparative anatomy of the Radiolaria, on the other hand, is so completely
known, and affords such certain glimpses into the morphological relations
of the related groups, that by its aid we are in a position at all events
to lay down the general features of their phylogeny with some probability,
and to lay the foundation of a natural system.

154. _Natural and Artificial Systems._--Although in the classification of
the Radiolaria, as in the case of all other organisms, the natural system
must be regarded as the goal of systematic classification, our phylogenetic
knowledge of the Radiolaria is too fragmentary and inadequate to admit of
the systematic arrangement here adopted being regarded as a thoroughly
consistent natural system, that is, as representing the true genealogical
tree of the class. Owing, however, to the extraordinary variety of form of
the Radiolaria, and the complicated relationships of the larger and smaller
groups, a synoptical grouping of the different categories and the erection
of a complete, even if to some extent artificial, system, becomes a logical
necessity. Under these circumstances, and regard being had to both these
conditions, the following systematic treatment of the Radiolaria will
appear as a _compromise between the natural and artificial systems_, like
all other zoological and botanical classificatory attempts. On the one
hand, the attempt is made to arrange the larger and smaller groups as
nearly as possible according to their phylogenetic relationships, whilst,
on the other hand, the practice of circumscribing each by a definition as
clear and logical as possible has been carried out. Since these two efforts
naturally often come into contact, the insufficiency of many parts of the
arrangement is obvious, hence its hypothetical and provisional character is
emphatically stated.

155. _Systematic Categories._--The categories or different orders of
divisions have in the Radiolaria, as in all other organisms, no _absolute_
significance, but only a _relative_ value. In itself it is quite
unimportant whether the whole group be regarded, as at first, as a _family_
(Ehrenberg, 1847), or as an _order_ (J. Müller, 1858), or as a _class_
(Haeckel, {cii}1881). These different views are regulated, on the one hand,
by the known extent of the group and by the amount of our acquaintance with
it, and on the other, by comparison with related groups and by reference to
their conventional disposition. When, therefore, the whole class,
Radiolaria, is here divided into two subclasses, four legions, eight
orders, eighty-five families, &c., these artificial categories are drawn up
only in the conviction that by this means the easiest survey and most
thorough insight into the system as a whole may be attained; this latter
will indeed approach as far as possible the ideal of a natural system, but
must on numerous practical grounds always remain more or less artificial.
Since it is to be expected that with the progress of our systematic
knowledge the rank of the various categories will rise, it is possible that
in the future the arrangement of the group may be somewhat as
follows:--_Phylum_, RADIOLARIA; _Four Classes_, SPUMELLARIA, NASSELLARIA,
ACANTHARIA, PHÆODARIA; _Eight Legions_ (Nos. I.-VIII. in the following
Table); _Twenty Orders_ (Nos. 1-20 in the Table), &c.

Four Legions. Eight Sublegions. Twenty Orders. Typical Families.
{ {1. Colloidea, { 1a. Thalassicollida.
{I. COLLODARIA { { 1b. Collozoida.
{ (Spumellaria {
{ palliata) {2. Beloidea, { 2a. Thalassosphærida.
{ { { 2b. Sphærozoida.
I. Legion {
(or Subclass){ { { 3a. Ethmosphærida.
SPUMELLARIA { {3. Sphæroidea, { 3b. Collosphærida.
(PERIPYLEA) { {
{ {4. Prunoidea, { 4a. Ellipsida.
[Porulosa {II. SPHÆRELLARIA { { 4b. Zygartida.
peripylea.] { (Spumellaria {
{ loricata) {5. Discoidea, { 5a. Phacodiscida.
{ { { 5b. Porodiscida.
{ {
{ {6. Larcoidea, { 6a. Larnacida.
{ { { 6b. Pylonida.

{ {7. Actinelida, { 7a. Astrolophida.
{ { { 7b. Litholophida.
{III. ACANTHOMETRA { { 7c. Chiastolida.
{ (Acantharia {
II. Legion { palliata) { { 8a. Astrolonchida.
(or Subclass){ {8. Acanthonida, { 8b. Quadrilonchida.
ACANTHARIA { { { 8c. Amphilonchida.
(ACTIPYLEA) {
{ { { 9a. Sphærocapsida.
[Porulosa { {9. Sphærophracta, { 9b. Dorataspida.
actipylea.] {IV. ACANTHOPHRACTA { { 9c. Phractopeltida.
{ (Acantharia {
{ loricata) { {10a. Belonaspida.
{ {10. Prunophracta, {10b. Hexalaspida.
{ { {10c. Diploconida.
{ {11. Nassoidea, 11. Nassellida.
{ {
{V. PLECTELLARIA {12. Plectoidea, {12a. Plagonida.
{ (Nassellaria { {12b. Plectanida.
{ palliata) {
{ {13. Stephoidea, {13a. Stephanida.
{ { {13b. Tympanida.
{
III. Legion { {14. Spyroidea, {14a. Zygospyrida.
(or Subclass){ { {14b. Androspyrida.
Nassellaria { {
(MONOPYLEA) { { {15a. Cannobotryida.
{VI. CYRTELLARIA {15. Botryodea, {15b. Lithobotryida.
[Osculosa { (Nassellaria { {15c. Pylobotryida.
monopylea.] { loricata) {
{ { {16a. Monocyrtida.
{ {16. Cyrtoidea, {16b. Dicyrtida.
{ { {16c. Tricyrtida.
{ { {16d. Stichocyrtida.

{ { {17a. Phæodinida.
{ {17. Phæocystina, {17b. Cannorrhaphida.
{ { {17c. Aulacanthida.
{VII. PHÆOCYSTINA {
IV. Legion { (Phæodaria { {18a. Orosphærida.
(or Subclass){ palliata) {18. Phæosphæria, {18b. Aulosphærida.
Phæodaria { { {18c. Cannosphærida.
(CANNOPYLEA).{
{ { {19a. Challengerida.
[Osculosa { {19. Phæogromia, {19b. Castanellida.
cannopylea.]{VIII. PHÆOCOSCINA { {19c. Circoporida.
{ (Phæodaria {
{ loricata) { {20a. Concharida.
{ {20. Phæoconchia, {20b. Coelodendrida.
{ { {20c. Coelographida.

156. _Formation of Species._--The totality of similar forms, which we unite
in one species, and which in the earlier dogmatic systems was regarded as a
category of absolute value, possesses only a _relative value_ like all
other systematic categories (§ 155). According to the individual views of
the systematist and the general survey which he has attained of the smaller
and larger systematic groups, the conception of a species adopted in his
practical work will be wider or narrower. In the present systematic
arrangement a medium extent has been adopted. It is shown that in the
Radiolaria, as in all other extensive groups of organisms, the constancy of
the species is very variable in the different groups. Many families of
Radiolaria are very rich in "bad species," _i.e._, very _variable_ forms,
in which the process of the formation of species is seen in progress; such,
for example, are--among the SPUMELLARIA, the Sphærozoida, Stylosphærida,
Phacodiscida and Pylonida; among the ACANTHARIA, the Amphilonchida and
Phractopeltida; among the NASSELLARIA, the #Stephoidea# and #Botryodea#;
and among the PHÆODARIA, the Aulacanthida, Sagosphærida, Castanellida and
Concharida. On the {civ}other hand, in some families numerous "good
species" may be distinguished, since the intermediate connecting forms are
no longer present and the forms have become _relatively constant_. As
instances of such families may be mentioned, among the SPUMELLARIA, the
Astrosphærida, Cyphinida, Porodiscida and Tholonida; among the ACANTHARIA
the Quadrilonchida and Dorataspida; among the NASSELLARIA, the #Spyroidea#
and #Cyrtoidea#; among the PHÆODARIA, the Challengerida, Medusettida,
Circoporida and Coelographida. The more carefully the different groups are
studied, the more numerous the individuals of each species under
comparison, the greater becomes the number of "bad" species among the
Radiolaria, and the smaller the number of good ones. Originally, no doubt,
all "species bonæ" were "malæ." There may be observed in the manifold
skeletal forms of the Radiolaria, on the one hand, the utmost accuracy of
configuration, and on the other, the greatest variability, and hence a
careful comparative study of them leads to a firm conviction of the gradual
"Transformation of Species," and of the truth of the "Theory of Descent."

157. _Palæontological Development._--The palæontology of the Radiolaria
already offers very considerable material for study; but in consequence of
its incompleteness this is of little value for the study of the phylogeny
of the class. By far the larger portion of the fossil Radiolaria belong to
the Tertiary period; only quite recently have numerous well-preserved
fossil Radiolaria been described from the Mesozoic period, and especially
from the Jura. Of Palæozoic Radiolaria (from the coal measures) only slight
traces are known. Moreover, the fossil Radiolaria hitherto known have been
found only in very circumscribed and widely separated localities. The
majority of all the species belong to the small island of Barbados.
Although our palæontological acquaintance with the Radiolaria must
necessarily be incomplete for this reason, it is still more so since at
least thirty out of the eighty-five families (that is more than a third)
could not possibly leave any fossil remains, either because they possess no
skeleton, or because of its chemical composition.

Of the four legions of the Radiolaria, the ACANTHARIA (on account of the
solubility of their astroid acanthin skeletons) have entirely vanished
and have never been found fossil. Of the PHÆODARIA, whose silicate
skeleton is not as a rule capable of fossilisation, only one section
(Dictyochida) of a single family (Cannorrhaphida) has been observed
fossil. Hence the fossil remains of the Radiolaria belong almost
exclusively to the two legions, SPUMELLARIA and NASSELLARIA, which were
formerly united under the term "Polycystina." Among these, however, the
skeletonless Thalassicollida, Collozoida, and Nassellida could leave no
traces. Hence there only remain fifty-five families of which we might
expect to find fossil siliceous skeletons. Even of these, however,
scarcely the half are certainly known in the fossil condition, whilst of
the remainder nothing certain is known; for example, of the large order
#Larcoidea# (among the SPUMELLARIA) and of the #Stephoidea# (among the
NASSELLARIA) with a few isolated exceptions, no fossils are known. The
great majority of fossil Radiolaria belong to the two NASSELLARIAN orders
#Cyrtoidea# and #Spyroidea# (two relatively very highly developed
groups); next to these follow the orders {cv}#Discoidea# and #Sphæroidea#
among the SPUMELLARIA. From these palæontological facts it is obvious
that our present very incomplete acquaintance with the fossil Radiolaria
is quite insufficient to warrant us in drawing any conclusions from it
regarding the phylogenetic development or palæontological succession of
the individual groups.

158. _Origin of the Four Legions._--The agreement of all Radiolaria in
those constant and essential characters of the unicellular body, which
distinguish them from all other Protista (especially the differentiation of
the malacoma into a central capsule and extracapsulum), justifies the
conclusion that all members of this class have been developed from a common
undifferentiated stem-form. Only the simplest form of the SPUMELLARIA, a
skeletonless spherical cell with concentric spherical nucleus and calymma,
can be regarded as such. The simplest form of the Thalassicollida which is
now extant (_Actissa_, _Procyttarium_, p. 12), corresponds so exactly to
the morphological idea of that hypothetical stem-form that it may
unhesitatingly be regarded in a natural system as the common point of
origin of the whole class. On the other hand, _Actissa_ is so closely
related to the simple Heliozoa (_Actinophrys_, _Actinosphærium_,
_Heterophrys_, _Sphærastrum_, &c.) that its origin from this group of
Rhizopoda is exceedingly probable. The three legions ACANTHARIA,
NASSELLARIA, and PHÆODARIA are to be regarded as three main diverging
branches of the genealogical tree, which have been developed in different
directions and are only connected by their simplest stem-forms
(_Actinelius_, _Nassella_, _Phæodina_) with the stem-form of the
SPUMELLARIA, the primordial _Actissa_.

159. _Phylogeny of the Spumellaria._--The legion SPUMELLARIA or PERIPYLEA
is to be regarded as the common stem-group of the Radiolaria, and its
simplest form, _Actissa_, as the primitive genus or radical form of the
whole class; for it possesses in the simplest and most undifferentiated
form all those characters by which the Radiolaria are distinguished from
other Protista; all the other genera of the class may be derived from it by
successive modifications. Considered as a legion the whole group
SPUMELLARIA is undoubtedly monophyletic, for all its members possess those
essential characters by which it is distinctively marked off from the other
three legions, more especially a simple capsule-membrane, which is
everywhere evenly perforated by innumerable small pores; the nucleus lies
originally in the centre of the spherical central capsule. Furthermore, all
SPUMELLARIA lack those positive characters which distinguish the three
remaining legions--the centrogenous acanthin skeleton of the ACANTHARIA,
the basal porochora and the monaxon podoconus of the NASSELLARIA, the
astropyle and phæodium of the PHÆODARIA.

160. _Origin of the Spumellaria._--The genus _Actissa_ (p. 12, Pl. 1, fig.
1) presents the Radiolarian type in its simplest and most primitive form--a
spherical central capsule, which encloses in its middle a spherical
nucleus, and which is surrounded by a spherical calymma. The whole
unicellular body consists, therefore, of three concentric spheres, {cvi}and
possesses neither skeleton nor alveoles, nor other differentiated parts.
The innumerable fine pseudopodia, which issue from the central capsule
through the evenly distributed pores in its membrane, radiate in all
directions through the calymma and pass out over its surface. _Actissa_
can, therefore, be directly derived phylogenetically from the simplest
skeletonless Heliozoa (_Actinophrys_, _Heterophrys_, _Actinosphærium_,
_Sphærastrum_). The only essential difference between the two consists in
the development of the _central capsule_, which in _Actissa_ separates as a
distinct membrane the endoplasm from the exoplasm. This differentiation
which we regard is the most important distinguishing character of the
Radiolaria, has been transmitted by inheritance, along with the formation
of flagellate spores in the central capsule, from _Actissa_, the primitive
parent to all the other Radiolaria.

{cvii}161. _Hypothetical Genealogical Tree of the Spumellaria_:--

LARCOIDEA DISCOIDEA
~~~~~~~~~~~~~ ~~~~~~~~~~~~~~~~~~~~~~~
Streblonida PHACODISCARIA
| Coccodiscida
| THOLONIDA |
| | |
PRUNOIDEA | | |
~~~~~~~~~~~~~~~~~~ Soreumida| | |
| | | |
Zygartida | | | |
| | | | |
| | |Zonarida| |
| |Lithelida| | SPHÆROIDEA |CYCLODISCARIA
| | | | |~~~~~~~~~~~~~ | Spongodiscida
| | | | |Stylosphærida |Pylodiscida|
| Phorticida| | | | | | |
| | | | | | | | |
Panartida | | | | | | | |
| Artiscida| +----+---+ | | | |
| | | | | | | |
| | | | | Phacodiscida| |
|Spongodruppida | | | | | | |
| | | | | Staurosphærida | +-----+
| | | | Pylonida | | |
| | | | | | | |
Cyphinida | | +---------+ | | |
| | | | | | Porodiscida
| | | | Astrosphærida| | |
+-----+ | | | | | |
| | | | | Cenodiscida |
|Spongellipsida| Larnacida | | | |
| | | | | | | Archidiscida
| | | | |Cubosphærida | |
Druppulida | | | | | +--------+
| | | | | | |
| | | | | |Collosphærida |
| | | | | | | |
| | | LARNACILLA +-----+-----+ |
| | | (Trizonium) | |
| Spongurida | | | |
+------+-------+ | | |
| | | |
Ellipsida Larcarida Liosphærida Cenodiscida
(CENELLIPSIS) (CENOLARCUS) (CENOSPHÆRA) (CENODISCUS)
| | | |
[Actiprunum?] [Actilarcus?] [Procyttarium] [Actidiscus?]
| | | |
+---------------+------+-------+--------------+
|
CENOSPHÆRA (Common stem-form of all Sphærellaria?)
| |
| | POLYCYTTARIA
| ~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~~
|Collosphærida Collozoida Sphærozoida }
| | | | } BELOIDEA
+-----+ | Thalassosphærida }
| | |
Ethmosphærida | |
| | |
+----------------+----------+
|
COLLOIDEA
|
Thalassicollida
|
ACTISSA

{cviii}162. _Collodaria and Sphærellaria._--Whilst in all SPUMELLARIA the
malacoma agrees in possessing the characteristic features of the legion,
and thus justifies its derivation monophyletically from the common
stem-form _Actissa_, the different forms of skeleton, on the other hand,
cannot all be referred to the same fundamental form. More especially the
_spherical lattice-shell_, from which all the numerous skeletal forms of
the #Sphærellaria# may be derived, cannot have arisen from the incomplete
Beloid skeleton which characterises the #Beloidea# among the #Collodaria#.
It is probable rather that the formation of the skeleton has taken place
independently in those two groups of SPUMELLARIA. From the skeletonless
#Colloidea#, as the common stem-group of the SPUMELLARIA, two different
main groups have diverged, on the one hand the #Beloidea#, whose skeleton
consists of separate spicules scattered in the extracapsulum, and on the
other hand, the #Sphærellaria#, which have formed a simple lattice-sphere
around the central capsule; from this the manifold forms of the remaining
SPUMELLARIA may be derived.

163. _Descent of the Sphærellaria._--The extensive order #Sphærellaria#,
which includes all SPUMELLARIA with a complete lattice-shell, develops an
extraordinary variety of skeletal structures; these may, nevertheless, all
be derived without violence from a common stem-form, or simple spherical
lattice-shell, _Cenosphæra_. The main stem of the order, the extensive
suborder #Sphæroidea# (Pls. 5-30), is derived immediately from _Cenosphæra_
(p. 61, Pl. 12); three diverging branches of it being represented by the
other three suborders, the #Prunoidea# (Pls. 16, 17, 39, 40) being
developed by elongation, and the #Discoidea# (Pls. 31-48) by shortening of
the vertical main axis, whilst the #Larcoidea# (Pls. 9, 10, 49, 50) have
originated by the modification of the spherical lattice-shell into a
lentelliptical or triaxial ellipsoidal one. Although the monophyletic
derivation of all #Sphærellaria# from _Cenosphæra_ is exceedingly probable,
the possibility of a polyphyletic origin for the group is by no means
excluded. For even in the skeletonless primitive genus of all the
SPUMELLARIA, _Actissa_ (as well as in the social _Collozoum_), there are
found, in addition to the usual spherical types, other species (or
subgenera, p. 12) whose central capsule is not spherical but a modification
of the sphere; in _Actiprunum_ ellipsoidal; in _Actidiscus_ lenticular; in
_Actilarcus_ lentelliptical; if such modified forms of _Actissa_ were to
develop their lattice-shells independently, then their form would
correspond to that of the central capsule; and such simple ellipsoidal,
discoidal, and lentelliptical lattice-shells might have been the primitive
forms of the #Prunoidea#, #Discoidea# and #Larcoidea#.

164. _Genealogical Tree of the Sphæroidea._--_Cenosphæra_, the simplest
form of the spherical lattice-shell, may be unhesitatingly regarded as the
common stem-form of all the #Sphæroidea# (pp. 50-284, Pls. 5-30).
_Cenosphæra_ (p. 61, Pl. 12) arose directly from _Actissa_ simply by the
silicification of the spherical exoplasmatic network of the sarcodictyum
around the central capsule, on the surface of the concentric calymma. From
this simple siliceous extracapsular lattice-sphere all other forms of
#Sphæroidea# have arisen, in the main by the manifold combination of two
simple processes, first by the formation of radial spines on the surface of
the lattice-sphere, and second, the addition of concentric spherical
lattice-shells. Both processes may be utilised as the foundation for a
systematic treatment of the #Sphæroidea# (compare pp. 52-58).

If in the #Sphæroidea# the characteristic number and disposition of the
_radial spines_ be regarded as the most important heritable peculiarity
of the different families, then we have the following natural
arrangement:--(1) Liosphærida, without radial spines; (2) Cubosphærida,
with six radial spines (opposite in pairs in three axes perpendicular to
each other); (3) Staurosphærida, with four radial spines (in two axes
crossed at right angles); (4) Stylosphærida, with two opposite radial
spines (in the vertical main axis); and (5) Astrosphærida, with numerous
regularly or irregularly distributed radial spines (eight to twenty or
more). If, on the contrary, more stress be laid upon the number of the
concentric lattice-shells, then we have the following artificial
grouping:--(1) Monosphærida, with one simple lattice-sphere; (2)
Dyosphærida, with two concentric lattice-spheres; (3) Triosphærida, with
three; (4) Tetrasphærida, with four; (5) Polysphærida, with numerous
(five to twenty or more) concentric lattice-shells; (6) Spongosphærida,
with a spongy spherical shell. In general the former arrangement appears
more natural than the latter, since the number of primary radial spines,
which grow out from the primary lattice-sphere, determines their
ground-form from the outset, whatever may be the number of secondarily
added shells. Strictly speaking, according to the view adopted, these
Liosphærida which have several shells, on the outer surface of which
there are no radial spines, ought to be classified according to the
number and arrangement of their internal radial connecting beams and
distributed among the other families. The practical application of this
correct principle meets, however, with great difficulties. Also in many
cases the phylogenetic relations of the different #Sphæroidea# are more
complicated than would appear from both these classificatory principles.
In general their phylogeny will quite correspond with their ontogeny,
since from the innermost first formed {cix}lattice-shell (primary
medullary shell) a number of radial spines arises, and upon these the
secondary shells are formed from within outwards.

165. _Genealogical Tree of the Prunoidea._--The suborder #Prunoidea# is
very closely related to the #Sphæroidea#, and is distinguished from it by
the elongation of one axis; from the simple lattice-sphere (_Cenosphæra_)
is developed a latticed ellipsoid (_Cenellipsis_, Pl. 39, fig. 1). The
development of this vertical isopolar main axis is foreshadowed even among
the #Sphæroidea#, in that family in which two opposite radial spines grow
out of the primary lattice-sphere at the two poles of the vertical main
axis (Stylosphærida, Pls. 13, 14). These latter pass over without any sharp
boundary into those forms of #Prunoidea# whose ellipsoidal lattice-shell
bears two opposite main-spines (Stylatractida, Pls. 15, 16). Other very
intimate relationships between the #Sphæroidea# and #Prunoidea# are
indicated in certain of the latter by the fact that of the two concentric
lattice-shells the inner (medullary) shell is spherical, the outer
(cortical) shell ellipsoidal (Pl. 39, figs. 3, 7, 8, 14, 19); often three
concentric lattice-shells are present, of which the two inner are spherical
intracapsular medullary shells, whilst the outer is an extracapsular
cortical shell, ellipsoidal or cylindrical in form (Pl. 39, figs. 4, 12,
17, 18). Owing to the manifold nature of these phylogenetical relations and
the variety of their combinations, the derivation of the individual
#Prunoidea# from the #Sphæroidea# is rendered very difficult; in addition
to which it is possible that the simplest #Prunoidea# (_Cenellipsis_,
_Ellipsidium_) have been directly developed from the skeletonless
_Actiprunum_ (a form of _Actissa_ with ellipsoidal central capsule, p. 14)
by the excretion of a simple ellipsoidal lattice-shell on the surface of
their calymma.

The phylogeny of the #Prunoidea# is especially complicated by the
formation of peculiar transverse constrictions, perpendicular to the
longitudinal axis. They are wanting only in the Monoprunida (Ellipsida,
Druppulida, and Spongurida); the Dyoprunida (Artiscida and Cyphinida, Pl.
39, figs. 9-19) possess only one such constriction (in the equatorial
plane); the Polyprunida, on the other hand, have three, five, or more
parallel constrictions (Panartida and Zygartida, Pl. 40). The chambers,
which are separated off by these constrictions, may be regarded as polar
sections of incomplete cortical shells.

166. _Genealogical Tree of the Discoidea._--The suborder #Discoidea# is
closely related to the #Sphæroidea#, but separated from it by shortening of
one axis; from a simple lattice-sphere (_Cenosphæra_) a latticed lens or
flattened spheroid is developed, whose circular equatorial plane is larger
than any other section (_Cenodiscus_, Pl. 48, fig. 1). The formation of
this horizontal equatorial plane is perhaps indicated in that family of
#Sphæroidea# in which four crossed radial spines, lying in one plane, are
developed (Staurosphærida, Pls. 15, 31, 42). The morphological and
phylogenetical relations of the #Discoidea# to the #Sphæroidea# are
precisely the converse of those of the #Prunoidea#; in the latter the
vertical axis appears longer, in the former shorter than any {cx}other axis
of the body. The #Discoidea# are probably polyphyletic, having originated
from several different groups of #Sphæroidea#; at least two essentially
different main groups may be distinguished among them; of these the one is
characterised by the formation of a large extracapsular lenticular cortical
shell (Phacodiscaria), whilst in the other this typical "Phacoid shell" or
lattice-lens is wanting (Cyclodiscaria, compare pp. 403-409).

The Phacodiscida (Pls. 31-35) perhaps constitute the primitive group of
the Phacodiscaria, their lenticular or Phacoid cortical shell being
connected by radial bars with one or two concentric spherical medullary
shells; they may have originated directly from the Dyosphærida or
Triosphærida by flattening of the spheroidal cortical shell. From the
Phacodiscida the Cenodiscida (if indeed they be not the primitive
stem-form) have been developed by retrogression and loss of those
medullary shells. The Coccodiscida (Pls. 36-38), on the other hand, have
been developed from the Phacodiscida by the addition of concentric rings
of chambers, which may be regarded as incomplete cortical shells, only
the equatorial portion of which is developed. Perhaps the Porodiscida,
the primitive group of the Cyclodiscaria, have arisen in a similar way;
they lack, however, the typical Phacoid shell, the concentric rings of
chambers being directly applied to a small spherical medullary shell in
the equatorial plane (Pls. 41-46). If those rings from the commencement
be interrupted by three interradial gaps (gates) the family Pylodiscida
arises (Pl. 38, figs. 6-20). If, on the contrary, the concentric radially
divided chambers of the Porodiscida become quite irregular and spongy,
they pass over into the Spongodiscida (Pls. 46, 47). It is not, however,
impossible that part of the #Discoidea# (especially the Cenodiscida) have
originated directly from skeletonless #Collodaria# with a lenticular
central capsule, such as are found in a subgenus of _Actissa_
(_Actidiscus_, p. 15).

167. _Genealogical Tree of the Larcoidea._--The suborder #Larcoidea#
presents in the structure, composition, and development of its variously
formed lattice-shells much more complicated relations than the other
#Sphærellaria#; it is essentially distinguished from them by the
characteristic ground-form of its lattice-shells, which is a "lentellipsis"
or a triaxial ellipsoid (also the ground-form of the rhombic
crystallographic system, the rhombic octahedron). Hence all parts of the
body are regularly disposed with respect to three different dimensive axes;
all three axes, perpendicular one to another, are isopolar but of different
lengths; the longest is the vertical main axis, the mean the horizontal
frontal axis, the shortest the horizontal sagittal axis. In the great
majority of the #Larcoidea# the lentelliptical ground-form is indicated in
the central capsule, even when it is not at once obvious in the skeleton.
Since such lentelliptical central capsules are developed even in _Actissa_
(_Actilarcus_, p. 16), it is possible that the simplest #Larcoidea# may
have arisen directly from these by deposition of a simple lentelliptical
lattice-shell in the sarcodictyum, on the surface of the calymma
(_Cenolarcus_, Pl. 50, fig. 7). It is more probable, however, that these
simplest forms (_Cenolarcus_, _Larcarium_) have been developed from the
simplest #Sphæroidea# (_Cenosphæra_), by the spherical body growing
unequally in the three dimensions of space. It appears especially likely
{cxi}from a study of the concentrically disposed lattice-shells of some
#Larcoidea# (_Coccolarcus_, _Larcidium_, Pl. 50, fig. 8), in which the
inner medullary shell is spherical, the outer cortical shell more or less
elliptical. In the great majority of #Larcoidea# the latter arises in quite
a peculiar manner, three broad lattice-zones, which are developed in three
planes at right angles to each other, growing out from a small spherical or
lentelliptical medullary shell, _Trizonium_, _Larnacilla_ (compare pp. 600,
615, 628, &c.).

The trizonal _Larnacilla_-shell commences by the formation of a
transverse girdle, by the union of two lateral latticed processes, which
spring right and left in the equatorial plane from the poles of the
frontal axis of a lentelliptical medullary shell (_Monozonium_, p. 633,
Pl. 9, fig. 1). This is followed by a second lateral girdle, which lies
in the frontal plane and proceeds from its lateral poles (_Dizonium_, p.
634, Pl. 9, figs. 2, 3). Finally the sagittal girdle is formed, lying in
the sagittal plane and arising from the lateral girdle on the two poles
of the main axis (_Trizonium_, p. 637, Pl. 9, fig. 4). Whilst the gaps
between the three zones of this trizonal shell remain open in the
Pylonida, in _Larnacilla_, the important primitive form of the Larnacida,
they are closed by lattice-work (Pl. 50, figs. 3-8). From this trizonal
_Larnacilla_-shell the great majority of Larcoid shells may be derived.
Such a system of zones may be repeated (Diplozonaria) or even developed a
third time (Triplozonaria, p. 632). In most #Larcoidea# the zones are
secondarily connected by lattice-work. In the Tholonida (Pl. 10) each of
the two opposite latticed wings of a zone becomes a closed dome. In the
Zonarida (Pl. 50, figs. 9-12) these domes are partially or wholly
bisected by constrictions or latticed septa which are developed in the
three dimensive planes. The Lithelida (Pl. 49, figs. 1-7) are
characterised by the fact that one of each pair of opposite latticed
processes (or half zones) grows more strongly than the other, and that
the larger completely embraces the smaller so as to form a complicated
spiral. Whilst in this case the spiral lies in a plane, in the
Streblonida (Pl. 49, figs. 8, 9) it becomes turbinoid like a gastropod
shell and forms an ascending spiral. Finally, two small families of
#Larcoidea# are characterised by quite irregular growth (a very rare
occurrence among the Radiolaria); these are the simple-chambered
Phorticida (Pl. 49, figs. 10, 11) and the many chambered Soreumida (Pl.
49, figs. 12, 13). The phylogenetic relationship of these families of
#Larcoidea# is probably very complicated and demands closer investigation
(compare pp. 599-604).

168. _Descent of the Polycyttaria._--The polyzootic or colonial Radiolaria,
which we unite in the group Polycyttaria (sometimes known as "Sphærozoea"),
belong without doubt to the legion SPUMELLARIA, for they possess all the
peculiarities by which these PERIPYLEA are distinguished from the other
legions of the Radiolaria. Only the morphological position of the
Polycyttaria in that legion, and their phylogenetic relation to the
monozootic or solitary SPUMELLARIA, can be variously interpreted. The three
families which we distinguish among the Polycyttaria are so closely related
to three different families of the Monocyttaria, that they may be directly
derived from them by the formation of colonies. According to this
_triphyletic hypothesis_ the social skeletonless Collozoida (Pl. 3) would
be descended from the solitary Thalassicollida (Pl. 1), the polyzootic
Sphærozoida with a Beloid skeleton (Pl. 4) from the monozootic
{cxii}Thalassosphærida (Pl. 2), and the colonial Collosphærida with a
Sphæroid skeleton (Pls. 5-8) from the solitary Ethmosphærida (Pl. 12, &c.).
Many species of monozootic and polyzootic forms in all three groups are so
alike that they can only be distinguished by the fact that the one series
are colonial, the others solitary. On the other hand, there are some
reasons which would justify a monophyletic hypothesis for the Polycyttaria,
_e.g._, the precocious nuclear division; in this case it would be most
natural to hold that the Sphærozoida and Collosphærida have arisen as two
diverging branches from the Collozoida, whilst the latter are nothing else
than colonial Thalassicollida.

169. _Phylogeny of the Acantharia._--The legion ACANTHARIA or ACTIPYLEA is
distinguished by its peculiar acanthin skeleton, which develops
centrogenously, as well as by the disposition in groups of the pores in its
central capsule, and its excentric usually precocious nucleus; it is thus
so different from all other Radiolaria as undoubtedly to furnish,
phylogenetically considered, an independent stem (§ 7). This stem is only
connected at the root by _Actinelius_ with the primitive form of the
SPUMELLARIA, _Actissa_. The stem is monophyletic, since all the forms
belonging to it may be derived without violence from _Actinelius_ as a
common primitive form.

170. _Origin of the Acantharia._--The genus _Actinelius_ (p. 730, Pl. 129,
fig. 1), which may naturally be regarded as the common primitive form of
all ACANTHARIA, possesses a spherical central capsule, which in consequence
of the early division of the nucleus (§ 63), encloses numerous small
nuclei; from its centre arise many simple radial spines of equal size,
which penetrate the central capsule. A large number of radial pseudopodia
issue between the spines from the sarcomatrix which surrounds the capsule.
_Actinelius_ may have been directly derived from _Actissa_, the common
stem-form of all Radiolaria, by the division of the pseudopodia into two
groups, myxopodia, which remained soft, and axopodia, which became firm (§
95A). As the latter became changed into strong acanthin rods, and touched
each other in the centre, they forced the nucleus from its originally
central position and brought about its early division. _Actinelius_ is also
of all Radiolaria the form which, next to _Actissa_, most nearly approaches
the Heliozoa. If the stiff axial threads of _Actinosphærium_ be conceived
of as partially converted into acanthin spines, and its nucleated medullary
substance as separated from the alveolar cortical layer by a membrane
(central capsule), then _Actinelius_ would be produced.

{cxiii}171. _Hypothetical Genealogical Tree of the Acantharia_:--

{cxiv}172. _Adelacantha and Icosacantha._--The numerous forms of
ACANTHARIA, here disposed in twelve families and sixty-five genera, may be
divided phylogenetically into two main groups of very different
extent--_Adelacantha_ and _Icosacantha_. The more primitive group,
_Adelacantha_, have an indefinite and variable number of radial spines,
which are always quite simple in form and usually irregularly distributed;
this main division includes only the one order #Actinelida#, with six
genera, among which is _Actinelius_, the common stem-form of all the
ACANTHARIA. The more recent group, Icosacantha, includes all the other
ACANTHARIA (fifty-nine genera), and is very markedly distinguished from the
Adelacantha by the fact that the radial spines are always twenty in number,
and arranged according to Müller's law (compare pp. 717-725, and § 110).
Since this regular disposition (in five alternating zones each of four
spines) has been retained by inheritance in the whole of the Icosacantha,
it is probable that this large group has been developed monophyletically
from a twig of the Adelacantha; _Actinastrum_ (p. 732) and _Chiastolus_ (p.
738) still present connecting links between the former and the latter,
between _Actinelius_ and _Acanthometron_.

173. _Acanthonida and Acanthophracta._--The extensive main division
Icosacantha (§ 110), which embraces all ACANTHARIA with twenty radial
spines, disposed according to Müller's law, may be subdivided into two
large groups or orders:--the #Acanthonida# (p. 740, Pls. 130-132) and the
#Acanthophracta# (p. 791, Pls. 133-140). The latter possess a complete
extracapsular lattice-shell, which the former have not. The more recent
#Acanthophracta# may be derived phylogenetically from the more primitive
#Acanthonida# simply by the development of this lattice-shell, with which
process are usually (perhaps always) connected certain alterations in the
malacoma, _e.g._, degeneration of the myophriscs (§ 96). The most primitive
form of all Icosacantha is the genus _Acanthometron_ (p. 324), in which all
the twenty acanthin spines are of the simplest constitution and of equal
dimensions.

174. _Differentiation of the Acanthonida._--The order #Acanthonida#, which
embraces all Icosacantha which have no complete lattice-shell, divides
early into three main branches, the three families Astrolonchida,
Quadrilonchida, and Amphilonchida (p. 727, Pls. 130-132). The first of
these constitutes the common stem-group from which the other two as well as
the whole group #Acanthophracta# have been developed; the common stem-form
of all is _Acanthometron_ (§ 173). All the Astrolonchida (p. 740, Pl. 130)
have twenty radial spines of equal size and similar form. On the other
hand, in the Quadrilonchida (p. 766, Pl. 131) the four equatorial spines
differ from the others in size and sometimes also in form. In the
Amphilonchida (p. 781, Pl. 132) two opposite equatorial spines (lying in
the hydrotomical axis) are much larger than the other eighteen and of a
different shape. Of the three families of the #Acanthonida# the most
important is the primitive group Astrolonchida, for from this the various
stem-forms of the #Acanthophracta# arise. They are subdivided according to
the formation of the spines into three subfamilies: the Zygacanthida, with
simple spines without apophyses (or transverse processes); the
Phractacanthida, with two opposite apophyses on each radial {cxv}spine, and
the Stauracanthida, with four crossed apophyses on each radial spine. The
three genera of the Zygacanthida represent the stem-forms of the three
families, since the radial spines in _Acanthometron_ (the most primitive
form of #Acanthonida#) are cylindrical, in _Zygacantha_ two-edged, and in
_Acanthonia_ four-edged (p. 741).

175. _Capsophracta and Cladophracta._--The extensive order
#Acanthophracta#, which embraces all ACANTHARIA with a complete
lattice-shell, is polyphyletic, its main subdivisions have been developed
independently from different branches of the #Acanthonida#. The whole order
may be divided directly into two main groups, the #Capsophracta# and
#Cladophracta# (p. 793), which differ in the structure and the origin of
their lattice-shell. The group (or suborder) #Capsophracta# includes only
the single family Sphærocapsida (p. 795, Pl. 133, figs. 7-11; Pl. 135,
figs. 6-10); the lattice-shell arises independently of the twenty radial
spines, being made up like a pavement of innumerable small acanthin plates,
united by a kind of cement; each plate being perforated by a fine pore. In
addition twenty larger main pores (or groups of four pores each) are
present, corresponding to the twenty radial spines; these are always equal,
quadrangular prismatic, without transverse processes as in _Acanthonia_. In
the #Cladophracta#, which include the five remaining families of the
#Acanthophracta#, the structure and origin of the lattice-shell are quite
different; the lattice-shell is here made up of the branches of the
transverse processes, which radiate tangentially from the twenty radial
spines and are only united secondarily.

176. _Ascent of the Dorataspida._--The group #Cladophracta#, or those
ACANTHARIA whose lattice-shell arises by the union of transverse
processes of the twenty radial spines, includes five different
families, whose stem-group is the family Dorataspida, with a simple
spherical lattice-shell. This family itself is, however, diphyletic in
origin, being composed of two essentially and originally different
subfamilies--Diporaspida and Tessaraspida (p. 803). The Diporaspida (p.
808, Pls. 137, 138) have been developed from the Phractacanthida, and as
each radial spine of the latter bears two opposite apophyses, so the
lattice-shell of the former has forty primary aspinal pores (two on the
base of each spine). On the other hand, the Tessaraspida (p. 830, Pls. 135,
136) have been developed from the Stauracanthida, and as each radial spine
of the latter bears four crossed apophyses, so the lattice-shell of the
former has eighty primary aspinal pores (four at the base of each spine).

177. _Descent of the Diporaspida._--Whilst the Tessaraspida (§ 176) have
given rise to no new groups which could take rank as independent families,
no less than four separate families of ACANTHARIA have arisen from the
Diporaspida. The Phractopeltida (Pl. 133, figs. 1-6) are distinguished from
all other ACANTHARIA by the possession of two concentric spherical
lattice-shells, and have probably been developed from the {cxvi}Diporaspida
in the same way as the Dyosphærida from the Monosphærida among the
#Sphæroidea#; in that case the smaller inner lattice-sphere (medullary
shell) would be the primary, and the larger outer sphere (cortical shell)
the secondary; this latter shows forty primary aspinal pores like those of
the Diporaspida. The possibility is not excluded, however, that the small
inner lattice-sphere of the Phractopeltida is a secondary product. The
three remaining families, which must be regarded as descendants of the
Diporaspida, form together a single phylogenetic series, and are separated
from the primitive group mainly by the fact that the original spherical
form of the lattice-shell has been modified into one distinguished by an
elongated equatorial axis (the hydrotomical axis); hence the #Prunophracta#
(pp. 794-859). The ellipsoidal Belonaspida have arisen directly by
hypertrophy of the two opposite equatorial spines of this hydrotomical axis
(p. 859, Pl. 136, figs. 6-9; Pl. 139, figs. 8, 9; perhaps they have also
arisen directly from the Amphilonchida). In the lentelliptical Hexalaspida
(Pl. 139) all six spines which lie in the hydrotomical meridian plane (two
equatorial and four polar) are very strongly developed, the remaining
fourteen being rudimentary. Finally, in the Diploconida the two conical
sheaths of the two opposite hydrotomical equatorial spines are so
predominant that they take the chief part in the formation of the
hour-glass-shaped shell.

178. _Phylogeny of the Nassellaria._--The legion NASSELLARIA or MONOPYLEA
is so clearly characterised by the peculiar porochora, which closes the
osculum at the oral pole of the monaxon central capsule, and by the
podoconus connected with it, that there can be no doubt that
phylogenetically it represents an independent stem (§ 8). This stem is only
connected at its base by means of _Cystidium_ and _Nassella_ with _Actissa_
and _Thalassicolla_, the stem-forms of the SPUMELLARIA. This stem is
monophyletic, inasmuch as all its members may be derived without violence
from the skeletonless Nassellida (_Nassella_, _Cystidium_, p. 896, Pl. 91,
fig. 1).

179. _Origin of the Nassellaria._--The Nassellida (p. 896), which may
naturally be considered as the common stem-group of the NASSELLARIA, are
most nearly related among other Radiolaria to the Thalassicollida, and in
both these skeletonless families the simplest forms, _Cystidium_ and
_Actissa_ correspond; on the other hand, those which have arisen from them
by the formation of alveoles in the calymma (_Nassella_ and
_Thalassicolla_) also correspond. The origin of the simplest Nassellida
from these primitive Thalassicollida may be explained by supposing that the
numerous (formerly evenly distributed) pores of the capsule membrane became
obliterated in the upper (apical) half of the central capsule, whilst in
the lower (basal) half they became correspondingly more strongly developed;
hence the porochora was formed at the oral pole of the vertical main axis,
and a differentiation of the endoplasm proceeding from this gave rise to
the characteristic podoconus. Both these organs still at present exhibit
very various degrees of progressive development.

{cxvii}180. _Hypothetical Genealogical Tree of the Nassellaria._

CYRTOIDEA.
~~~~~~~~~~~~~~~~~~~~~~~~~
BOTRYODEA _Triradiata_
~~~~~~~~~~~~~
Pylobotryida Podocampida
| |
| _Eradiata_ | _Multiradiata_ SPYROIDEA
| | | ~~~~~~~~~~~~~~~~~
| Lithocampida | Phormocampida Androspyrida
| | | | |
Lithobotryida | Podocyrtida | |
| | | | |
| Theocyrtida | Phormocyrtida |
Tholospyrida
| | | | | |
Cannobotryida | Tripocyrtida | Phormospyrida |
| | | | | |
| Sethocyrtida | Anthocyrtida | |
| | | | | |
| | Tripocalpida | | |
| | | | | |
| Cyrtocalpida | Phænocalpida +---+----+
| | | | |
STEPHOIDEA | +---------+------------+ Zygospyrida
~~~~~~~~~~~~ | | (Spyroidea
triradiata)
Tympanida | Tripocalpida |
| | (Cyrtoidea triradiata monocyrtida) |
| | | |
Coronida | +------------------+-----------------------------+
| | |
| Semantida |
| | |
+--+---+ Cyrtellaria
| |
| Cortiniscus |
| | |
Stephanida | |
| | |
+-----+------+ |
| |
Cortina *--------- Cortinida (PLECTELLARIA)
(Cortina)
|
Plectaniscus ---- Plagoniscus ---- }
| }
Tetraplecta ---- Tetraplagia ---- } PLECTOIDEA
| } Plectanida
Plectophora ---- Plagiacantha ---- } |
| } |
Triplecta ---- Triplagia ---- } Plagonida
| }
Nassoidea
(Nassellida)
|
Nassella
(Cystidium)
|
Actissa

{cxviii}181. _Plectellaria and Cyrtellaria._--The extensive legion
NASSELLARIA far surpasses the other three legions in the endless variety of
its skeletal structures, and owing to the complicated relationships of its
numerous families presents no lack of difficult phylogenetic problems. All
NASSELLARIA may be divided first into two main groups or sublegions,
#Plectellaria# and #Cyrtellaria#; the latter having a complete
lattice-shell, the former not. Probably the #Cyrtellaria# have been
polyphyletically developed from several different groups of #Plectellaria#.
These groups are, however, connected in such manifold ways that a
monophyletic origin of all the NASSELLARIAN skeletons from one original
element is possible. Such a primitive element may have been furnished by
any one of three different skeletal parts, the sagittal ring, the basal
tripod, and the latticed cephalis (compare pp. 891-895, Bütschli, L. N. 40,
41).

182. _Phylogenetic Skeletal Elements of the Nassellaria._--The multiform
skeleton of the NASSELLARIA may be referred in different ways to one of the
three above-mentioned structural elements. Each of these (p. 891) may by
itself form the skeleton; the sagittal ring in the simplest #Stephoidea#
(_Archicircus_, _Lithocircus_), the basal tripod in the simplest
#Plectoidea# (_Triplagia_, _Plagiacantha_), the latticed cephalis in the
simplest #Cyrtoidea# (_Cyrtocalpis_, _Archicapsa_). In the great majority
of the NASSELLARIA, however, two of these elements, or even all three, are
found combined. In most #Cyrtellaria#, more especially, both the sagittal
ring and the basal tripod may be recognised in the lattice-shell, though
often only in slight rudiments or scarcely perceptible traces. In the
#Plectellaria# also (which possess no latticed cephalis) there are
individual genera with complete development both of the sagittal ring and
basal tripod; this important combination is especially well represented in
the Cortinida (_Cortina_, _Cortiniscus_, _Stephanium_, _Stephaniscus_,
_Tripocoronis_, &c.). The greatest difficulty as regards the phylogeny of
the NASSELLARIA lies in the fact that the most various combinations of the
three elements are presented by closely related or very similar forms. If,
in spite of this, a monophyletic hypothesis as to the origin of the
NASSELLARIA seems essential all sides of the three possible hypotheses must
receive full consideration and critical comparison (§§ 183-191).

183. _Ascent of the Nassellaria from the Plectoidea._--The monophyletic
hypothesis (No. 2, p. 893) which regards the basal tripod as the common
origin of the skeleton of all NASSELLARIA, starts from the simplest forms
of the #Plectoidea# (_Triplagia_, _Plagoniscus_, _Triplecta_,
_Plectaniscus_, &c., Pl. 91). All #Plectoidea# may be immediately derived
as diverging twigs of these, as well as all triradial and multiradial forms
of #Cyrtoidea# and #Spyroidea#; for in all these cases the distinctive
triradial (or the derived multiradial) form of skeleton appears directly
derivable from the simple basal tripod of the former. The same is perhaps
also true of many #Botryodea#. {cxix}Furthermore, certain important forms
of #Stephoidea# (_Cortina_, _Cortiniscus_, _Stephanium_, _Stephaniscus_,
&c.), which have a characteristic combination of the sagittal ring and
basal tripod, may be immediately derived from such forms of #Plectoidea# as
_Plagoniscus cortinaris_, _Plagiocarpa procortina_, _Plectaniscus
cortiniscus_, &c. On the contrary, those #Stephoidea# and #Cyrtoidea# in
which the basal tripod is wanting can only be derived from the #Plectoidea#
by the assumption that this structure has disappeared in consequence of
phylogenetic degeneration. The monophyletic derivation of the NASSELLARIA
from the #Plectoidea# has more internal probability than that from the
#Stephoidea#, since it is easier to suppose that the Cortinida (_Cortina_,
_Stephanium_, &c.) have been derived from the #Plectoidea# (_Plagoniscus_,
_Plagiocarpa_) than the converse. This view is the basis of the
hypothetical tree shown in § 180.

184. _Ascent of the Nassellaria from the Stephoidea._--The monophyletic
hypothesis (No. 1, p. 893) which regards the primary sagittal ring as the
common starting point of the skeleton in all NASSELLARIA, starts from the
simplest forms of #Stephoidea# (_Archicircus_, _Lithocircus_, &c., Pl. 81).
All #Stephoidea# and #Spyroidea# may be immediately derived from these, as
also the majority of the #Cyrtoidea# and probably of the #Botryodea#. Those
numerous forms of the last two groups, however, which possess no trace of a
sagittal ring, can only be derived from the former by the supposition that
the latter has completely disappeared in in consequence of gradual
phylogenetic degeneration. The same holds true also of the #Plectoidea#,
although certain forms (_e.g._, _Plagiocarpa procortina_, Pl. 91, fig. 5;
_Plectaniscus cortiniscus_, Pl. 91, fig. 9) appear to indicate the
commencing formation of the sagittal ring by the concrescence of two
branches, which approach each other from the upper part of the apical rod
and the ventral part of the basal rod. In any case, it is a fact of great
phylogenetic significance, that the primary sagittal ring in the cephalis
of the #Cyrtoidea# shows all conceivable stages of degeneration (compare
Bütschli, L. N. 40, 41, as well as the general account of and critical
comparison of the NASSELLARIA, pp. 889-895, &c.).

185. _Ascent of the Nassellaria from the Cyrtoidea._--The monophyletic
hypothesis (No. 3, p. 894) which regards the latticed cephalis as the
common point of origin of all the skeletons of the NASSELLARIA, starts from
the simplest forms of the #Cyrtoidea#, that is, from the Cyrtocalpida or
eradial Monocyrtida (_Archicorida_, _Archicapsida_, Pls. 51, 52, 98). All
#Cyrtoidea# and #Botryodea# may be regarded as divergent forms of these
monothalamous #Cyrtoidea#; the polythalamous simply by the addition of
fresh joints at the basal pole, the triradiate and multiradiate by the
development of three or more apophyses. The origin of the sagittal ring
(which presents every stage of development and degeneration in the
#Cyrtoidea#) may be regarded as a mechanical thickening of the latticed
plate in the sagittal circumference of the cephalis. By stronger
{cxx}development of this ring and coincident sagittal constriction of the
cephalis the order #Spyroidea# may be derived from the #Cyrtoidea#. On the
other hand, the #Plectellaria#, which possess no cephalis, and indeed no
complete lattice-shell whatever, may be derived from the Monocyrtida by the
assumption of a degeneration of this structure; the sagittal ring having
been preserved in the #Stephoidea#, and the tripod of the Tripocalpida in
the #Plectoidea#. Although such a monophyletic derivation of the
NASSELLARIA from the Cyrtocalpida is possible, and though here, too, the
Cortinida play an important part as connecting links, this hypothesis has
less internal probability than that of the derivation from the #Stephoidea#
(§ 184) or #Plectoidea# (§ 183).

186. _Genealogical Tree of the Plectoidea._--The order #Plectoidea#
includes those NASSELLARIA whose rudimentary skeleton does not contain the
characteristic sagittal ring of the #Stephoidea#, but consists of several
(at least three) radial spines, which proceed from a point in the centre of
the porochora. The branches of these radial spines remain free in the
Plagonida, whilst in the Plectanida they unite with each other to form a
loose meshwork (not, however, a complete lattice-shell). The number and
arrangement of the radial spines, which serve for generic distinctions, are
the same in both families, so that each genus of the Plectanida has arisen
from a corresponding genus of the Plagonida. The simplest Plagonida, which
possess a basal tripod (_Triplagia_ or _Plagiacantha_ with three rays,
_Tetraplagia_ with four rays) are probably to be regarded as forming the
common origin of the whole order. These agree with certain three- and
four-rayed skeletal pieces of the #Beloidea# (Thalassosphærida and
Sphærozoida); and also the four and six-rayed twinned pieces of the latter
(spicula bigemina and trigemina) repeat in the same fashion the skeleton of
the former (_Plagonidium_, _Plagonium_). This similarity, however, is a
mere analogy and possesses no phylogenetic significance. On the other hand,
certain Plagonida (_Plagoniscus_, _Plagiocarpa_), and the corresponding
genera of Plectanida (_Plectaniscus_, _Periplecta_) seem to have important
phylogenetic relations to certain #Stephoidea# (_Cortina_, _Cortiniscus_,
&c.); the sagittal ring of the latter having perhaps arisen by the vertical
apical spine of the former having been connected with their horizontal
basal rod by two ventral apophyses growing out opposite to each other
(compare pp. 902, 914, _Plagiocarpa procortina_, Pl. 91, fig. 5). In this
case the Plectanida would belong to the simplest stem-forms of the
NASSELLARIA.

187. _Genealogical Tree of the Stephoidea._--The order #Stephoidea#
includes all those NASSELLARIA whose skeleton does not form a complete
lattice-shell, but consists of one or more rings, and often of a loose
meshwork which arises by the union of branches of the rings. A _vertical
sagittal ring_ is constantly present, embracing the central capsule in the
median sagittal plane, and forming at its basal pole various processes, the
starting point for other skeletal forms. The most important of these is the
tripodal _Cortina_ {cxxi}(p. 950, § 182). The Stephanida are the most
archaic family among the #Stephoidea# (p. 937, Pl. 81), perhaps indeed
among all the NASSELLARIA (§ 184); in them the sagittal ring and its
processes alone constitute the skeleton; secondary rings and meshes are
wanting. Two diverging families, the Semantida and Coronida, have been
developed from the Stephanida, and from one of them the family Tympanida
has arisen.

The Semantida (p. 953, Pl. 92) develop a horizontal basal ring at the
oral side of the vertical sagittal ring; the basal meshes or lattice
gates, which remain between the former and the latter, are the important
cortinar pores (one pair jugular, one pair cardinal, p. 954); they
usually appear inherited in the cortinar septum of the #Cyrtellaria#. In
the Coronida (p. 967, Pls. 82, 94) a second vertical ring (the frontal
ring) appears in addition to the sagittal ring; it lies in the frontal
plane at right angles to the latter. Finally the Tympanida (p. 987, Pls.
93, 94) have probably arisen from the Semantida by the formation of a
second horizontal ring (mitral ring) parallel to the basal and attached
to the upper portion of the sagittal ring.

188. _Genealogical Tree of the Spyroidea._--The extensive order #Spyroidea#
is of especial interest in connection with the phylogeny of the
NASSELLARIA, since all its members show two well-developed skeletal
elements in combination, the sagittal ring of the #Stephoidea# and the
latticed cephalis of the #Cyrtoidea#; the majority possess also the basal
tripod of the #Plectoidea# (or a radial skeleton derived from it). Hence
there is a possibility of deriving the stem-forms of the #Spyroidea# from
each of these three groups. The four families of this order exhibit similar
relationships to those of the four families of #Cyrtoidea#; the common
stem-group is the family Zygospyrida; from this the Tholospyrida have
arisen by the development of a galea on the apical pole, the Phormospyrida
by the addition of a thorax on the basal pole. The Androspyrida may be
derived either from the Tholospyrida by the formation of a basal thorax, or
from the Phormospyrida by the development of an apical galea. Some groups,
however, such as the peculiar Nephrospyrida (Pl. 90) have probably been
developed directly from the #Stephoidea#.

189. _Genealogical Tree of the Botryodea._--The peculiar order #Botryodea#
(p. 1103), which is both difficult to investigate and insufficiently known,
presents great phylogenetic difficulties both as to its ascent and descent.
Probably the different genera of this order have been polyphyletically
developed from different groups of #Cyrtoidea# (perhaps also to some extent
of #Spyroidea#) by the formation of lobes in the cephalis. The three
families of #Botryodea# are related to each other in the same way as are
the three first families of the #Cyrtoidea#. From the single-jointed
Cannobotryida (corresponding to the Monocyrtida), the two-jointed
Lithobotryida (corresponding to the Dicyrtida), may be derived by the
development of a basal thorax, and from the latter the three-jointed
Pylobotryida (like the Tricyrtida) by the addition of an abdomen. In the
last two families the forms with an open basal mouth {cxxii}(Botryopylida
and Botryocyrtida) are to be regarded as primitive: the Botryocellida and
Botryocampida have arisen by the closure of this mouth with a basal
lattice-plate.

190. _Genealogical Tree of the Cyrtoidea._--The multiform and extensive
group #Cyrtoidea# presents the greatest difficulties to be found in the
phylogeny of the NASSELLARIA, because their morphological relations are
most complicated, and because similar forms very often appear to be of
quite different origin. The great majority of the #Cyrtoidea# show more or
less clearly a combination of the three structural elements: sagittal ring,
basal tripod, and latticed cephalis (p. 891). There are also, however,
numerous #Cyrtoidea#, whose skeleton no longer shows any trace of the
sagittal ring. Many of these show as the basis of the skeleton a strong
basal tripod with an apical spine, around which the cephalis has obviously
been secondarily developed, _e.g._, the remarkable Euscenida (p. 1146, Pls.
53, 97) and the interesting Callimitrida (p. 1217, Pls. 63, 64). These may
have been derived immediately from the #Plectoidea# without any relation to
the #Stephoidea#. There are also numerous true Monocyrtida, whose shell
consists of a simple latticed cephalis without a trace of the sagittal ring
or basal tripod (Cyrtocalpida, Pl. 51, figs. 9-13; Pl. 98, fig. 13); these
may have been developed directly from the skeletonless Nassellida by the
formation of a simple ovoid _Gromia_-like shell, and may have no relation
either to the #Stephoidea# or #Plectoidea#. On these grounds, as well as
from the complicated relationships of the many smaller groups of
#Cyrtoidea#, it is probable that the whole order has been developed
polyphyletically from different divisions of the #Plectellaria#.

191. _Systematic Arrangement of the Cyrtoidea._--Although for the reasons
just given no systematic arrangement of the #Cyrtoidea# can at present, or
for a long time in the future, be regarded as other than artificial, yet
some general principles of classification for this extensive group can be
laid down, which may serve as starting points for some future natural
disposition. This is especially true of the relations which in an
artificial system (p. 1129) were primarily utilised for the distinction of
twelve families and twenty-four subfamilies; the number of segments in the
shell, the number of radial apophyses (and parameres), and the constitution
of the basal aperture of the shell.

As regards the _number of segments_, separated by transverse
constrictions, of which the shell is composed, it is dependent upon the
secondary addition of new joints at the basal pole of the main axis.
Hence all many-jointed #Cyrtoidea# are to be derived from single-jointed
ones, and the four sections thus distinguished (Monocyrtida, Dicyrtida,
Tricyrtida, Stichocyrtida) form a phylogenetic series. Very often,
however, the primary cephalis disappears owing to retrograde
metamorphosis; and in such cases the single joint of the apparent
Monocyrtida is formed of the thorax (_e.g._, {cxxiii}Pls. 52, 54, &c.).
As regards the _number of radial apophyses_, three sections of
#Cyrtoidea# may be distinguished; the Pilocyrtida with three, the
Astrocyrtida with numerous apophyses, and the Corocyrtida with none (p.
1129). The last two may in general be regarded as two divergent branches
from the first, for the eradiate Corocyrtida have probably been formed
from the triradial Pilocyrtida by entire loss of the radial apophyses,
whilst on the other hand the multiradiate Astrocyrtida have arisen from
them by additions to the primary apophyses (interpolation of interradial
between the perradial ones). As regards the _constitution of the
shell-aperture_, the #Cyrtoidea# may be divided into Cyrtaperta and
Cyrtoclausa (p. 1129); in general the Cyrtoclausa (with latticed
shell-aperture) have arisen from the Cyrtaperta (with simple open mouth);
in many Monocyrtida the converse may be supposed, the simple basal mouth
having been formed by degeneration of a basal lattice.

192. _Phylogeny of the Phæodaria._--The legion PHÆODARIA or CANNOPYLEA is
so clearly marked off from other Radiolaria by the double membrane of the
central capsule and the astropyle at its oral pole, as well as by the
extracapsular phæodium, that it must be regarded phylogenetically as an
independent stem (§ 9). This stem is only connected at its root by
_Phæodina_ with the stem-form of the SPUMELLARIA, _Actissa_. The stem
itself is monophyletic, inasmuch it its members may be derived without
violence from the skeletonless Phæodinida (_Phæodina_, _Phæocolla_). On the
other hand, the formation of the skeleton of the PHÆODARIA is undoubtedly
polyphyletic, different Phæodinida having independently commenced the
formation of a skeleton and having carried it out in very different ways.

193. _Origin of the Phæodaria._--The Phæodinida (p. 1544, Pl. 101), which
may naturally be regarded as the common stem-group of the PHÆODARIA, have
their nearest relations among other Radiolaria in the Thalassicollida (p.
10); and since this family is to be regarded as the primitive group of all
Radiolaria, they may be directly derived from them phylogenetically. The
essential modifications by which the primitive Phæodinida have arisen from
the more archaic Thalassicollida are of three kinds; (1) the doubling of
the membrane of the central capsule; (2) the reduction of the numerous fine
pores in the membrane and the formation of an osculum, and of an astropyle
closing it, at the oral pole of the main axis; (3) the production of an
extracapsular phæodium. This last may, perhaps, be regarded as a
unilateral hypertrophy of the voluminous pigment masses which are deposited
in the sarcomatrix of certain Thalassicollida. Of the two genera of
Phæodinida hitherto known, probably _Phæodina_ (Pl. 101, fig. 2) approaches
the original stem of the PHÆODARIA more nearly than _Phæocolla_ (Pl. 101,
fig. 1), for the latter exhibits only the large main opening of the central
capsule (astropyle), whilst the former possesses also a pair of accessory
openings (parapylæ). The hypothetical stem-form (_Phæometra_) presumably
had a larger number of small parapylæ (like many Circoporida and
Tuscarorida), and the astropyle was probably but little differentiated from
them.

{cxxiv}194. _Hypothetical Genealogical tree of the Phæodaria:_--

{cxxv}195. _Phæocystina and Phæocoscina._--Whilst the malacoma of all
PHÆODARIA possesses the characteristics of the legion, and hence justifies
the assumption of a monophyletic origin, the skeleton, on the other hand,
shows in the different groups such manifold and fundamental variations that
a polyphyletic origin of the latter is indubitable. Different Phæodinida
have commenced the formation of the skeleton independently, and it has
progressed in different directions. In the #Phæocystina# it remained
incomplete and led to the formation of various Beloid skeletons, whilst the
#Phæocoscina# developed complete lattice-shells. Both of these divisions
too are to be regarded as polyphyletic, since the skeletal forms of the
different groups cannot be derived without violence from a common primitive
form.

196. _Phæocystina with a Beloid Skeleton._--The order #Phæocystina#
includes all PHÆODARIA which have no complete lattice-shell; it contains,
firstly, the skeletonless Phæodinida (the common stem-group of the legion),
and secondly, the Phæacanthida, or PHÆODARIA with a Beloid skeleton (§
115). The latter are divisible into several very different groups (at least
two or three) which are probably different in origin. The Aulacanthida
(Pls. 102-105) form radial tubes which perforate the calymma, their
proximal end resting upon the surface of the central capsule, whilst the
distal extremity projects freely outwards. The skeleton of the
Cannorrhaphida, on the other hand, is composed of many separate portions
which are never radially arranged but are either placed tangentially to the
surface of the calymma or scattered irregularly in its gelatinous mass.
Furthermore, in the three subfamilies of which this family is composed, the
individual skeletal portions are so different that they have probably
arisen independently of each other; in the Cannobelida they form
cylindrical tangential tubes (Pl. 101, figs. 3-5), in the Catinulida flat
basin or cap-like structures (Pl. 117, fig. 8), in the Dictyochida hollow
rings, from which small pyramids are developed by unilateral formation of
lattice-work (Pl. 101, figs. 9-14; Pl. 114, figs. 7-12).

197. _Phæosphæria with a Sphæroid Skeleton._--The order #Phæosphæria#
includes those PHÆODARIA which possess a spherical (sometimes slightly
modified) lattice-shell without the characteristic aperture of the
#Phæogromia#. They have probably arisen independently of these, though they
may have been derived from the Castanellida by loss of the shell-aperture,
which was present originally. The four families which we have distinguished
among the #Phæosphæria#, are so different in the structure of their
lattice-shell that their phylogenetic connection is doubtful. In the
Orosphærida (Pls. 106, 107) and the Sagosphærida (Pl. 108) the whole
lattice-shell consists of a single piece and is unjointed (without astral
septa); in the former it is very firm and massive, with thick laminated
trabeculæ and polygonal meshes; in the latter it is very delicate and
brittle, with filiform trabeculæ and large {cxxvi}triangular meshes. On
the other hand, the voluminous shell of the Aulosphærida (Pls. 109-111),
and of the Cannosphærida (Pl. 112), is characterised by a very peculiar
system of joints; it is composed of numerous separate cylindrical tubes,
which are placed tangentially and united at the nodes by stellate
partitions or astral septa. The Cannosphærida possess further a simple
central Cyrtoid shell, connected with the outer jointed shell by hollow
radial trabeculæ. Since many Aulosphærida possess rudiments of such
centripetal trabeculæ it is possible that these latter have been derived
from the former by the loss of the central Cyrtoid shell; the formation of
this monaxon shell perhaps indicates descent from the #Phæogromia#
(Castanellida).

198. _Phæogromia with a Cyrtoid Skeleton._--That order of the PHÆODARIA
which we designate #Phæogromia#, contains many very different forms, all
agreeing in the possession of a Cyrtoid skeleton, or a monaxon
lattice-shell, which has a large aperture at one pole of its vertical main
axis (§ 123). This Cyrtoid skeleton is sometimes ovoid or conical,
sometimes lentiform or helmet-shaped, sometimes polyhedral or almost
spherical. Although the principle of its structure is simple and often like
that of the Monocyrtida among the NASSELLARIA, yet the structure of the
wall and of the apophyses is so different in the various groups of the
#Phæogromia#, that the order is probably polyphyletic, and its Cyrtoid
shells have arisen independently of each other. Only in the Castanellida
(Pl. 113) does the shell-wall usually consist of simple lattice-work; in
the Challengerida, on the other hand (Pl. 99), it has an extremely fine
Diatom-like structure; in the Medusettida (Pls. 118-128) a peculiar
alveolar structure, and in the Circoporida (Pls. 114-117) and Tuscarorida
(Pl. 100) it possesses a characteristic porcellanous constitution (with
tangential spicules in a porous cement-mass); in the latter of these groups
the surface is smooth, in the former peculiarly tabulate; the two families
have also different stem-forms.

199. _Phæoconchia with a Conchoid Shell._--The order #Phæoconchia# (Pls.
121-128) is separated not only from all other PHÆODARIA, but also from all
other Radiolaria, by the possession of a bivalved shell resembling that of
a Lamellibranch; the two valves of this Conchoid skeleton are to be
interpreted as dorsal and ventral (§ 128). Probably these bivalved shells
are independent products, but possibly they may have been formed by the
bisection of a simple spherical lattice-shell; in the former case the
#Phæoconchia# would be directly descended from the Phæodinida, in the
latter from the Castanellida. The three families which we have
distinguished among the #Phæoconchia#, probably constitute a connected
stem, the most primitive group of which are the Concharida (Pls. 123-125).
From these the Coelodendrida (Pls. 121, 122) have next arisen by the
formation of a "galea" upon the apex of each valve, and the growth of
hollow tubes from this helmet-like structure. Finally, the Coelographida
{cxxvii}(Pls. 120-128) have been developed from the Coelodendrida by the
formation of a basal nasal tube (rhinocanna) from each galea, and the
formation of a median or paired frenulum, which connects the opening of the
nasal tube with the apex of the galea. In the Coelodendrida, as well as in
the Coelographida, there are two different subfamilies, of which the more
primitive (Coelodorida, Coelotholida) have free branches from the hollow
radial tubes, whilst the more recent (Coelodrymida, Coeloplegmida) form an
outer bivalved shell by anastomosis of the branches of the tubes.

200. _The Fundamental Biogenetic Law._--The causal connection between
ontogeny and phylogeny, which finds its most precise statement in the
fundamental biogenetic law, holds in general for the Radiolaria as for all
other organisms. In order to furnish direct proof of this, however, a
complete empirical knowledge both of individual and of palæontological
development would be necessary. In both these directions, as has been shown
in the foregoing chapters, our knowledge of the Radiolaria is very
incomplete and fragmentary, but still we are able to convince ourselves
indirectly of the validity of the law as applied to Radiolaria by the aid
of comparative anatomy. This is now so fully known to us (§§ 1-140) that we
are able not only to draw a complete and satisfactory picture of their
morphology, but also to arrive at most important conclusions regarding the
ontogeny and phylogeny of the individual groups. As regards the formation
of the multiform skeleton of the Radiolaria, most of the ontogenetic series
of forms, with which we have become acquainted by comparative anatomy, are
of _palingenetic_ nature; that is, they are primarily due to inheritance
and thus of direct phylogenetic significance. On the other hand, among the
ontogenetic phenomena of the Radiolaria, as far as they have yet been
investigated, only very few are _cenogenetic_, that is, brought about by
adaptive modification and without direct significance as regards phylogeny.

{cxxviii}PHYSIOLOGICAL SECTION.

----

CHAPTER VII.--VEGETATIVE FUNCTIONS.

(§§ 201-217.)

201. _Mechanism of the Functions._--The vital phenomena of the Radiolaria
are dependent upon the mechanical functions of their unicellular body, and
like those of all other organisms, are to be referred to physical and
chemical natural laws. All processes which appear in the life of the
Radiolaria are, therefore, ultimately to be explained by the attraction and
repulsion of the smallest particles, which compose the different portions
of their unicellular body; and the sensation of pleasure or the opposite is
in its turn the exciting cause of these elementary movements. Many adaptive
arrangements in the Radiolarian organism may produce the appearance of
being the premeditated result of causes working towards an end
("zweckthätig," _causæ finales_), but as opposed to this deceptive
appearance it must here be expressly stated that these may be recognised in
accordance with the developmental theory as the necessary consequence of
mechanical causes (_causæ efficientes_).

Our _physiological_ acquaintance with the Radiolaria has by no means
progressed so far as our _morphological_, so that the incomplete
communications which are placed here for the sake of completeness must be
regarded merely as preliminary fragments, not as fully elaborated
results. Since my recent investigations have been mainly in the direction
of morphology, I can add but little to the physiological conclusions,
which I stated at length in my monograph twenty-four years ago (L. N. 16,
pp. 127-165). Recently the vegetative physiology of the Radiolaria has
been much advanced by the recognition of the symbiosis with the Xanthellæ
(§ 205, L. N. 22, 39, 42). In addition Karl Brandt has recently (1885)
published several important contributions to the physiology of the
Polycyttaria or Sphaerozoea (L. N. 52).

202. _Distribution of Functions._--The distribution of the functions among
the various parts of the unicellular organism of the Radiolaria corresponds
directly to their anatomical composition, so that physiologically as well
as morphologically the central capsule and the extracapsulum appear as the
two coordinated main components. On the one hand the _central capsule_ with
its endoplasm and enclosed nucleus is the central organ of the "cell-soul"
(Zellseele), the unit regulating its animal and vegetative functions, and
the special organ of reproduction and inheritance. The _extracapsulum_
forms, on the other hand, by its calymma the protective envelope of the
central {cxxix}capsule, the support of the soft pseudopodia and the
substratum of the skeleton; the calymma acts also as a hydrostatic
apparatus, whilst the radiating pseudopodia are of the greatest importance
both as organs of nutrition and adaptation, as well as of motion and
sensation (§ 15). If, however, the vital functions as a whole be divided in
accordance with the usual convention into the two great groups of
_vegetative_ (nutrition and reproduction) and _animal_ (motion and
sensation), then the central capsule would be mainly the organ of
reproduction and sensation, and the extracapsulum the organ of nutrition
and motion.

The numerous separate vital phenomena, which by accurate physiological
investigation may be distinguished in the unicellular Radiolarian
organism, may be distributed in the above indicated conventional fashion
into a few larger and several smaller groups; it must always be borne in
mind, however, that these overlap in many respects, and that the division
of labour among the different organs in these Protista is somewhat
complicated, notwithstanding the apparent simplicity of their unicellular
organization. A general classification of the groups of functions is
difficult, because each individual organ discharges several different
functions. Thus the central capsule is pre-eminently the organ of
reproduction and inheritance, but not less (though less conspicuous) is
its importance as the psychical central organ, the unit regulating the
processes of sensation, motion, and also nutrition. In this last respect
it is comparable to the nerve-centres of the Metazoa, whilst the
peripheral nervous system of the latter (including the organs of sense
and the muscles) are in the present instance represented by the
pseudopodia, which are at the same time the most important organs of
nutrition and adaptation. In the calymma also in similar fashion several
different physiological functions are united.

203. _Metastasis._--The functions of metastasis and nutrition have in all
Radiolaria a purely animal character, so that these Rhizopoda from the
physiological standpoint are to be regarded as truly _unicellular animals_,
or Protozoa ("Urthiere"). Since they do not possess, like plants, the power
of forming synthetically the compounds (protoplasm, carbohydrates, &c.)
necessary for their sustenance, they are compelled to obtain them
ready-formed from other organisms. Like other true animals they evolve
carbon dioxide by the partial oxidation of those products, and hence they
successively take up the oxygen necessary to their existence from their
environment.

The question whether the Radiolaria are to be regarded as true animals I
discussed fully from various points of view in 1862, and finally answered
in the affirmative (L. N. 16, pp. 159-165). Afterwards, when in my
Generelle Morphologie (1866) I sought to establish the kingdom Protista,
I removed the Radiolaria along with the other Rhizopoda from the animal
kingdom proper and placed them in the kingdom Protista (Bd. i. pp.
215-220; Bd. ii. p. xxix). Compare also my Protistenreich (L. N. 32) and
my Natürliche Schöpfungsgeschichte (vii. Aufl., 1879, p. 364). Both these
steps appear fully justified when considered in the light of our present
increased knowledge. From the _physiological_ standpoint the Radiolaria
appear as unicellular _animals_, for in this respect the animal character
of their metastasis (that proper to an oxidising organism) furnishes the
sole {cxxx}criterion. On the other hand, from the morphological
standpoint, they are to be classed as neutral Protista, for in this
respect their unicellular character is the prominent feature, and
distinguishes them from all true multicellular animals (Metazoa). Compare
my Gastræa Theorie (1873, Jena. Zeitschr. für Naturwiss., Bd. viii. pp.
29, 53).

204. _Nutrition._--The nutritive materials which the Radiolaria require for
their sustenance, especially albuminates (plasma) and carbohydrates
(starch, &c.), they obtain partly from foreign organisms which they capture
and digest, and partly directly from the Xanthellæ or Philozoa, the
unicellular Algæ, with which they live in symbiosis (§ 205). _Zooxanthella
intracapsularis_, found in the ACANTHARIA (§ 76), is probably of the same
significance in this respect as _Zooxanthella extracapsularis_ of the
SPUMELLARIA and NASSELLARIA (§ 90); and perhaps the same is true also of
_Phæodella extracapsularis_ (or _Zoochlorella phæodaris_?) of the PHÆODARIA
(§ 89). The considerable quantity of starch or amyloid bodies, elaborated
by these inquiline symbiontes, as well as their protoplasm and nucleus, are
available, on their death, for the nutrition of the Radiolaria which
harbour them. Nutrition by means of other particles obtained by the
pseudopodia from the surrounding medium is by no means excluded; indeed it
may be regarded as certain that numerous Radiolaria (especially such as
contain no symbiotic Algoid cells) are nourished for the most part or
exclusively by this means. Diatoms, Infusoria, Thalamophora (Foraminifera)
as well as decaying particles of animal and vegetable tissues can be seized
directly by the pseudopodia and conveyed either to the sarcodictyum (on the
surface of the calymma) or to the sarcomatrix (on the surface of the
central capsule) in order to undergo digestion there. The indigestible
constituents (siliceous shells of Diatoms and Tintinnoidea, calcareous
shells of small Monothalamia and Polythalamia, &c.) are here collected
often in large numbers and removed by the streaming of the protoplasm.

The inception and digestion of nutriment, as it usually appears to take
place by the pseudopodia, has already been so fully treated in my
Monograph (L. N. 16, pp. 135-140), and since then in my paper on the
sarcode body of the Rhizopoda (L. N. 19, p. 342), that I have nothing of
importance to add. Quite recently Karl Brandt has expressed a doubt as to
whether the taking up of formed particles by the pseudopodia and their
aggregation in the calymma be really connected with the process of
nutrition. He is disposed rather to believe that these foreign bodies are
usually only accidentally and mechanically brought into the calymma, and
that the nourishment of the Radiolaria is derived exclusively or
pre-eminently from the symbiotic Xanthellæ (L. N. 52, pp. 88-93). I must,
however, maintain my former opinion, which I have only modified insomuch
that I now regard the sarcodictyum (on the outer surface of the calymma,
§ 94) rather than the sarcomatrix (on the outer surface of the central
capsule, § 92) as the principal seat of true digestion and assimilation.
From the sarcodictyum the dissolved and assimilated nutritive matters may
pass by the intracalymmar pseudopodia (or sarcoplegma, § 93) into the
sarcomatrix, and hence may reach the endoplasm through the openings in
the central capsule. To what an extent the Radiolaria are capable of
taking up even large formed bodies into the calymma, is shown by the
{cxxxi}striking instance of _Thalassicolla sanguinolenta_, which becomes
so deformed by the inception of numerous coccospheres and coccoliths,
that I described it as a special genus under the name _Myxobrachia_
(compare pp. 23, 30; also L. N. 21, p. 519, Taf. xviii., and L. N. 33, p.
37).

205. _Symbiosis._--Very many Radiolaria, but by no means all members of
this class, live in a definite commensal relation with yellow unicellular
Algæ of the group Xanthellæ. In the ACANTHARIA they live within the central
capsule (_Zooxanthella intracapsularis_, § 76), in the SPUMELLARIA and
NASSELLARIA, on the other hand, within the calymma but outside the central
capsule (_Zooxanthella extracapsularis_, § 90); in the PHÆODARIA a special
form of these symbiotic unicellular Algæ appears to inhabit the phæodium in
the extracapsulum, and to compose a considerable portion of the phæodellæ
(_Zooxanthella phæodaris_, § 90, or better perhaps _Zoochlorella
phæodaris_, § 89). Undoubtedly this commensal life is in very many cases of
the greatest physiological significance for both the symbiontes, for the
animal Radiolarian cells furnish the inquiline Xanthellæ not only with
shelter and protection, but also with carbon dioxide and other products of
decomposition for their nutriment; whilst on the other hand the vegetable
cells of the Xanthellæ yield the Radiolarian host its most important supply
of nutriment, protoplasm and starch, as well as oxygen for respiration.
Hence it is not only theoretically possible, but has been experimentally
proved, that Radiolaria which contain numerous Xanthellæ can exist without
extraneous nutriment for a long period in closed vessels of filtered
sea-water, kept exposed to the sunlight; the two symbiontes furnish each
other mutually with nourishment, and are physiologically supplementary to
each other by reason of the opposite nature of their metastasis. This
symbiosis is not necessary, however, for the existence of the Radiolaria;
for in many species the number of Xanthellæ is very variable and in many
others they are entirely wanting.

The symbiosis of the Radiolaria and Xanthellæ, or "yellow cells" (§§ 76,
90) was first discovered by Cienkowski in 1871 (L. N. 22). Ten years
later this important and often doubted fact was established by extended
observations and experiments almost simultaneously by Karl Brandt (L. N.
38, 39) and Patrick Geddes (L. N. 42, 43). This commensal life may be
compared with that of the lichens, in which an organism with vegetable
metastasis (the Algoid gonidia) and an organism with animal metastasis
(the Fungoid hyphæ) are intimately united for mutual benefit. But the
symbiosis of the Xanthellæ and Radiolaria is not as in the lichens a
phenomenon essential for their development, but has more or less the
character of an accidental association. The number of the inquiline
Xanthellæ is so variable even in one and the same species of Radiolaria,
that they do not appear to be exactly essential to its welfare; and in
many species they are entirely wanting. Their significance is
questionable in the case of those numerous deep-sea Radiolaria which live
in complete darkness, and in which, therefore, the Xanthellæ, even if
present, could excrete no oxygen on account of the want of light.
Nevertheless it is possible that the phæodellæ of the PHÆODARIA (usually
green, olive, or brown in colour), which are true cells, represent
vegetable symbiontes, {cxxxii}which in the absence of sunlight are able
to evolve oxygen by the aid of the phosphoresence of other abyssal
animals. Since the PHÆODARIA are, for the most part, dwellers in the
deep-sea, and since the voluminous phæodium must be of great
physiological importance, a positive solution of this hypothetical
question would be of no small interest (compare § 89).

206. _Respiration._--The respiration of the Radiolaria is animal in nature,
since all Protista of this class, like all other true Rhizopoda, take in
oxygen and give off carbon dioxide. Probably this process goes on
continuously and is tolerably active, as may be inferred from the fact that
Radiolaria cannot be kept for long in small vessels of sea-water unless
either they contain numerous Xanthellæ or the water is well aërated. The
oxygen is obtained from two sources, either from the surrounding water or
from the enclosed Xanthellæ, which in sunlight evolve considerable
quantities of this gas. Correspondingly, the carbon dioxide which is formed
during the process of oxidation of the Radiolaria is either given up to the
surrounding water or to the inquiline Xanthellæ, which utilise it for their
own sustenance (§§ 204, 205).

The significance of the symbiotic Xanthellæ for the respiration of the
enclosing Radiolaria may be shown experimentally in the following way. If
two Polycyttarian colonies of equal size, both of which contain numerous
Xanthellæ, be placed in equal quantities of filtered sea-water in sealed
glass tubes, and if one tube be placed in the dark the other in the
light, the colony in the former rapidly perishes, but not that in the
latter; the Xanthellæ excrete only under the influence of sunlight the
oxygen necessary for the life of the Radiolarian (compare Patrick Geddes,
L. N. 42, p. 304).

207. _Circulation._--In the protoplasm of all Radiolaria, both inside and
outside the central capsule, slow currents may be recognised which fall
under the general term circulation, and have already been compared to the
cyclosis in the interior of animal and vegetable cells, as well as to the
sarcode streams in the body of other Rhizopoda. These plasmatic currents or
"plasmorrheumata" probably continue throughout the whole life of the
Radiolaria, and are of fundamental importance for the performance of their
vital functions. They depend upon slow displacements of the molecules of
the plasma (plastidules or micellæ) and cause a uniform distribution of the
absorbed nutriment and a certain equalisation of the metastasis.
Furthermore they are of great importance also in the inception of
nutriment, the formation of the skeleton, locomotion, &c. Sometimes the
circulation is directly perceptible in the plasma itself; but usually it is
only visible owing to the presence of granules (sarcogranula), which are
suspended in the plasma in larger or smaller numbers. The movements of
these granules are usually regarded as passive, due to the active
displacement of the molecules of the plasma. Although the intracapsular
protoplasm is in communication with the extracapsular through the openings
in the capsule membrane, nevertheless the currents exhibit certain
differences {cxxxiii}in the two portions of the malacoma. It is sometimes
possible, however, to recognise the direct connection between them and to
observe how the granules pass through the openings in the capsule-membrane.

208. _Currents in the Endoplasm._--Intracapsular circulation or a certain
slow flowing of the plasma within the central capsule is probably just as
common in the Radiolaria as without it, but it is not so easy to observe in
the former case as in the latter. A more satisfactory proof of these
endoplasmatic currents is furnished by the arrangement of the protoplasm
within the central capsule, since this is (at all events in part) an effect
produced by them. In this respect the two main divisions of the class show
characteristic differences. In the Porulosa (the SPUMELLARIA, § 77, and the
ACANTHARIA, § 78) the endoplasm is in general distinguished by a more or
less distinct radial structure, which is to be regarded as the effect of
alternating centripetal and centrifugal radial streams. In the Osculosa, on
the other hand, this radial structure is absent and the intracapsular
plasmatic streams converge or diverge towards the osculum or main-opening
in the central capsule which lies at the basal pole of its main axis, and
through which the mass of the endoplasm issues into the calymma. The two
legions of the Osculosa, however, present differences in this respect. In
the NASSELLARIA (§ 79) the endoplasmatic currents appear to unite in an
axial main stream at the apex of the monaxon central capsule, and this
apical stream seems to split into a conical bundle, the individual threads
of which pass diverging between the myophane fibrillæ of the podoconus
towards the basis of the central capsule, and issue through the pores of
the porochora. In the PHÆODARIA (§ 80), on the other hand, meridional
currents of endoplasm are probably present on the inner surface of the
capsule, which flow from the aboral pole of the vertical main axis to its
basal pole, and return in the reverse direction.

209. _Currents in the Exoplasm._--Extracapsular circulation, or a distinct
flowing of the plasma outside the central capsule, may be readily observed
in all Radiolaria which are examined alive; this is most readily seen in
the astropodia, or those free pseudopodia which radiate from the
sarcodictyum on the surface of the calymma into the surrounding water. The
granular movement is often quite as clear in the sarcodictyum itself, and
may be recognised in the collopodia, which compose the irregular plasmatic
network within the calymma. More rarely it is possible to follow the
granular stream thence through the sarcomatrix, and further into the
interior of the central capsule. In general the direction of the
extracapsular protoplasmic streams is radial, and it is frequently
possible, even in a single free astropodium, to observe two streams
opposite in direction, the granules on one side of the radial sarcode
thread moving centripetally, those on the other side centrifugally. If the
threads branch, and neighbouring ones {cxxxiv}become united by connecting
threads, the circulation of the granules may proceed quite irregularly in
the network thus formed. The rapidity and character of the extracapsular
currents are subject to great variations.

The different forms of extracapsular sarcode currents have been already
very fully described in my Monograph (L. N. 16, pp. 89-126), and in my
critical essay on the sarcode body of the Rhizopoda (L. N. 19, p. 357,
Taf. XXVI.).

210. _Secretion._--Under the name _secretions_, in the strict sense, all
the skeletal formations of the Radiolaria may be included. They may be
divided according to their chemical composition into three different
groups: pure silica in the SPUMELLARIA and NASSELLARIA, a silicate of
carbon in the PHÆODARIA, and acanthin in the ACANTHARIA (compare § 102). It
may indeed be assumed that these skeletons arise directly by a chemical
metamorphosis (silicification, acanthinosis, &c.) of the pseudopodia and
protoplasmic network; and this view seems especially justified in the case
of the Astroid skeleton of the ACANTHARIA (§ 114), the Spongoid skeleton of
the SPUMELLARIA (§ 126), the Plectoid skeleton of the NASSELLARIA (§ 125),
the Cannoid skeleton of the PHÆODARIA (§ 127), and several other types. On
closer investigation, however, it appears yet more probable that the
skeleton does not arise by direct chemical metamorphosis of the protoplasm,
but by secretion from it; for when the dissolved skeletal material (silica,
acanthin) passes from the fluid into the solid state, it does not appear as
imbedded in the plasma, but as deposited from it. However, it must be borne
in mind that a hard line of demarcation can scarcely, if at all, be drawn
between these two processes. In the ACANTHARIA the intracapsular sarcode is
the original organ of secretion of the skeleton; in the other three
legions, on the other hand, the extracapsulum performs this function (§§
106, 107). In addition to the skeleton, we may regard as secretions (or
excretions) the intracapsular crystals (§ 75) and concretions (§ 75A), and
perhaps certain pigment-bodies (§§ 74, 88); and further, the calymma (§ 82)
may be considered to be a gelatinous secretion of the central capsule, and
perhaps also the capsule-membrane, in so far as it represents only a
secondary excretory product of the unicellular organism.

211. _Adaptation._--The innumerable and very various adaptive phenomena
which we meet with in the morphology of the Radiolaria, and especially in
that of their skeleton, are like other phenomena of the same kind, to be
ultimately referred to altered nutritional relations. These may be caused
directly either by the influence of external conditions of existence
(nutrition, light, temperature, &c.), or by the proper activity of the
unicellular organism (use or disuse of its organs, &c.), or, finally, by
the combined action of both causes in the struggle for existence. In very
many cases the cause to which the origin of a particular form of Radiolaria
is due may be directly perceived or at least guessed at with considerable
probability; thus, for example, the lattice-shells {cxxxv}may be explained
as protective coverings, the radial spines as defensive weapons, and the
anchor-hooks and spathillæ as organs of prehension, which are of advantage
to their possessors in the struggle for existence; the regular arrangement
of the radial spines in the Radiolaria may also be explained on hydrostatic
grounds, it being advantageous that the body should be maintained in a
definite position of equilibrium, &c. The well-known laws of _direct_ or
_actual adaptation_, which we designate cumulative, correlative, divergent
adaptation, &c., here explain a multitude of morphological phenomena. The
connection is less distinct in the case of the laws of _indirect_ or
_potential adaptation_, although this must play as important a part in the
formation of the Radiolaria as in that of other organisms (compare on this
head my Generelle Morphologie, Bd. ii. pp. 202-222).

212. _Reproduction._--The most common form of reproduction in the
Radiolaria is the formation of spores in the central capsule, which in this
respect is to be regarded as a sporangium (§ 215). In many Radiolaria
(Polycyttaria and PHÆODARIA), however, there occurs in addition an increase
of the unicellular organism by simple division (§ 213); upon this the
formation of colonies in the social Radiolaria is dependent (§ 14).
Reproduction by gemmation is much less common, and has hitherto been
observed only in the Polycyttaria (§ 214). In this group alone there also
occur at certain times two different forms of swarm-spores which copulate,
and thus indicate the commencement of sexual reproduction (Alternation of
Generations, § 216). The general organ of reproduction is in all cases the
central capsule, whilst the extracapsulum never takes an active part in the
process.

213. _Cell-Division._--Increase by cell-division among the Radiolaria in
the early stage, before the formation of the skeleton, is widely
distributed (perhaps even general?); in the adults of this class it is
rather rare and limited to certain groups. It is most readily observed in
the Polycyttaria; the growth of the colonies in this social group depends
mainly (and in many species exclusively) upon repeated spontaneous division
of the central capsule; all the individuals of each colony (in so far as
this has not arisen by the accidental fusion of two or more colonies) are
descendants of a single central capsule, which has arisen from an asexual
swarm-spore (§ 215) or from the copulation of two sexual swarm-spores (§
216). Whilst the central capsules of the colonies continually increase by
division, their calymma remains a common gelatinous sheath. Among the
SPUMELLARIA reproduction by simple cell-division probably occurs also in
many monozootic #Collodaria#. Among the ACANTHARIA the peculiar group
Litholophida has perhaps arisen by the spontaneous division of
#Acanthonida# (see p. 734). Among the PHÆODARIA increase by cell-division
seems to occur commonly in many groups, as in the #Phæocystina#, which have
no skeleton (Phæodinida, Pl. 101, {cxxxvi}fig. 2), or only an incomplete
Beloid skeleton (Cannorrhaphida, Pl. 101, figs. 3, 6, and Aulacanthida, Pl.
104, figs. 1-3). The #Phæosphæria# also (Aulosphærida, Coelacanthida) and
the #Phæogromia# (Tuscarorida, Challengerida) appear sometimes to divide;
at all events, their central capsule often contains two nuclei. Of special
interest is the spontaneous division of the #Phæoconchia#, especially the
Concharida (Pl. 124, fig. 6). In all monozootic Radiolaria, the nucleus
first divides by a median constriction into two equal halves (usually by
the mode of direct division); then the central capsule becomes constricted
in the middle (in the PHÆODARIA in the vertical main axis), and each
portion of the capsule retains its own nucleus. In the #Phæoconchia# each
half or daughter-cell corresponds to one valve of the shell, dorsal or
ventral, so that probably on subsequent separation each daughter-cell
retains one valve of the mother-cell, and forms a new one for itself by
regeneration (as in the Diatoms). In the polyzootic Radiolaria, which
already contain many small nuclei, but usually only a single central
oil-globule in each central capsule, the division of the latter is preceded
by that of the oil-globule. In many Polycyttaria the colony as a whole
multiplies by division.

The increase of the central capsule by division was first described in
1862 in my Monograph (L. N. 16, p. 146); since then R. Hertwig (L. N. 26,
p. 24) and K. Brandt (L. N. 52, p. 144) have confirmed my statement. In
the PHÆODARIA the division of the central capsule appears always to take
place in the main axis; in the bilateral sometimes in the sagittal,
sometimes in the frontal plane. In the Tripylea each daughter-cell seems
to retain one parapyle and half the astropyle (compare the general
description of the PHÆODARIA, Pl. 101, figs. 1-6, Pl. 104, figs. 1-3, and
also Hertwig, L. N. 33, p. 100, Taf. x. figs. 2, 11). Regarding the
spontaneous division of colonies of the Polycyttaria, see K. Brandt, L.
N. 52, p. 142.

214. _Cell-Gemmation._--Reproduction by gemmation has hitherto been
observed only in the social Radiolaria, but in them it appears to be widely
distributed, and in very young colonies is perhaps almost universally
present. The gemmules or capsular buds (hitherto described as
"extracapsular bodies") are developed on the surface of young central
capsules before these had secreted a membrane. They grow usually in
considerable numbers, from the surface of the central capsule, which is
sometimes quite covered with them. Each bud usually contains a
raspberry-like bunch of shining fatty globules, and by means of reagents a
few larger or a considerable number of smaller nuclei may be recognised in
them; the naked protoplasmic body of the bud is not enclosed by any
membrane. As soon as the buds have reached a certain size they are
constricted off from the central capsule and separated from it, being
distributed in the meshes of the sarcoplegma by the currents in the
exoplasm. Afterwards each bud becomes developed into a complete central
capsule by surrounding itself with a membrane when it has attained a
definite size. From the special relations of the process of nuclear
formation, which take place in the multiplication of the {cxxxvii}social
central capsules by gemmation and by cell-division, it would appear that
the capsules produced by the former method afterwards produce anisospores,
whilst those in the latter way yield isospores (§ 216).

The gemmules or capsular buds of the Polycyttaria were first accurately
described by Richard Hertwig (L. N. 26, pp. 37-39), under the name
"extracapsular bodies," and their significance rightly indicated; earlier
observers had incidentally mentioned and figured them, but had not seen
their origin from the central capsule. Quite recently Karl Brandt has
given a very painstaking account of them in the different Polycyttarian
genera (L. N. 52, pp. 179-198). In the Monocyttaria such a formation of
buds has not yet been observed. The basal lobes of the central capsule,
which occur in many NASSELLARIA, are not buds, but simple processes of
the capsule, due to its protrusion through the collar pores of the
cortinar septum (§ 55).

215. _Sporification._--Asexual reproduction by the formation of movable
flagellate spores has been hitherto observed only in a very small number of
genera; but since these belong to very different groups, and since the
comparative morphology of the capsule appears to be similar throughout as
regards the structure and development of its contents, it may be safely
assumed that this kind of reproduction occurs quite generally in the
Radiolaria. In all cases it is the contents of the central capsule, from
which the swarm-spores are formed, both nucleus and endoplasm taking an
equal share in the process; in all cases the spores produced are very
numerous, small, ovoid or reniform, and have one or two very long slender
flagella at one extremity (see §§ 141, 142). Since the whole contents of
the mature central capsule are used up in the formation of these flagellate
zoospores, it discharges the function of a sporangium. The division of the
simple primary nucleus into numerous small nuclei, which usually
(serotinous Radiolaria) takes place only shortly before sporification, but
sometimes (precocious Radiolaria, § 63) happens very early, is the
commencement of the often repeated process of nuclear division, which
terminates with the production of a very large number of small
spore-nuclei. The nucleolus often divides very peculiarly (§ 69, C). Each
spore nucleus becomes surrounded by a portion of endoplasm and usually
receives in addition one or more fatty granules, and sometimes also a small
crystal (hence the "crystal-spores"). The size of the flagellate zoospores
which emerge from the ruptured central capsule and swim freely in the water
by means of their flagellum, varies generally between 0.004 and 0.008 mm.
The extracapsulum is not directly concerned in the sporification, but
undergoes degeneration during the process and perishes at its conclusion.

The first complete and detailed observations on the formation of spores
in the Radiolaria were published by Cienkowski in 1871 and related to two
genera of Polycyttaria, the skeletonless _Collozoum_ and the
spherical-shelled _Collosphæra_ (L. N. 22, p. 372, Taf. xxix.). These
were subsequently continued and supplemented by R. Hertwig (1876, L. N.
26, pp. 26-42, and L. N. 33, p. 129), and a general summary of these
results has been given by Bütschli (L. N. 41, pp. 449-455).
{cxxxviii}Recently Karl Brandt has given a very detailed and fully
illustrated account of the sporification of the Polycyttaria (L. N. 52,
pp. 145-178). I have also had the opportunity during my sojourn in the
Canary Islands (1866), in the Mediterranean at Corfu (1877), and
Portofino (1880), as well as in Ceylon (1881), of observing the
development of flagellate zoospores from the central capsule of
individuals of all four legions: among the SPUMELLARIA in certain
#Colloidea#, #Beloidea#, #Sphæroidea#, and #Discoidea#, among the
ACANTHARIA in several #Acanthometra# and #Acanthophracta#, among the
NASSELLARIA in individuals belonging to the #Stephoidea#, #Plectoidea#,
and #Cyrtoidea#, and among the PHÆODARIA in one Castanellid. In most
zoospores I could distinctly observe only a single long flagellum;
sometimes, however, two or even three appeared to be present, but the
determination of their number is very difficult.

216. _Alternation of Generations._--A peculiar form of reproduction, which
may be designated "alternation of generations," appears to occur generally
in the Polycyttaria, but has not yet been observed in the Monocyttaria. All
#Collozoida#, #Sphærozoida#, and Collosphærida which have hitherto been
carefully and completely examined with respect to their development, are
distinguished by the production of two different kinds of swarm-spores,
isospores and anisospores. The _Isospores_ (or monogonous spores)
correspond to the ordinary asexual zoospores of the Monocyttaria (§ 215);
they possess a homogeneous, doubly refracting nucleus of uniform
constitution and develop asexually, without copulation. The _Anisospores_
(or amphigonous spores), on the other hand, are sexually differentiated and
possess a heterogeneous, singly refracting nucleus of twofold constitution;
they may therefore be distinguished as female macrospores and male
microspores. The _Macrospores_ (or gynospores, comparable with the female
macrogonidia of many Cryptogams) are larger, less numerous, and possess
larger nuclei, which are less easily stained, and have a fine filiform
trabecular network. On the other hand the _Microspores_ (or _androspores_,
comparable with the male microgonidia) are much smaller and more numerous,
and are distinguished by their smaller nuclei, which have thicker tuberculæ
and become stained more intensely. The gynospores and androspores are
developed in the #Collozoida# and #Sphærozoida# in the same individual, but
not in the Collosphærida. It is very probable that these two forms of
anisospores copulate with each other after their exit from the central
capsule and thus produce a new cell by the simplest mode of sexual
reproduction. But, since the same Polycyttaria, which produce these
anisospores, at other times give rise to ordinary or asexual isospores, it
is further possible that these two forms of reproduction alternate with
each other, and that the Polycyttaria thus pass through a true alternation
of generations. This has not yet been observed in the Monocyttaria, and
hence these latter seem to bear to the Polycyttaria a relation similar to
that in which the sexless solitary Flagellata (Astasiea) stand to the
sexual social Flagellata (Volvocinea). In the two analogous cases the
sexual differentiation may be regarded as a consequence of the social life
in the gelatinous colonies.

{cxxxix}The _sexual differentiation of the Polycyttaria_ was first
discovered in 1875 by R. Hertwig, and accurately described in the case of
_Collozoum inerme_ as occurring in addition to the formation of the
ordinary crystal-spores (L. N. 26, p. 36); compare also the general
discussion of Bütschli (L. N. 41, p. 52). Recently Karl Brandt has
demonstrated the formation of both homogeneous isospores (crystal-spores)
and heterogeneous anisospores (macro- and microspores) in seven different
species of Polycyttaria, and has come to the conclusion that in all
social Radiolaria there is a regular alternation between the former and
latter generations. Compare his elaborate account of the colonial
Radiolaria of the Gulf of Naples (L. N. 52, pp. 145-178).

217. _Inheritance._--Inheritance is to be regarded as the most important
accompaniment to the function of reproduction, and especially in the
present case, because the comparative morphology of the Radiolaria
furnishes abundant instances of the action of its different laws. The laws
of _conservative inheritance_ are illustrated by the comparative anatomy of
the larger groups; thus, in the four legions the characteristic
peculiarities of the central capsule are maintained unaltered in
consequence of continuous inheritance, although great varieties appear in
the skeleton in each legion. The individual parts of the skeleton furnish
by their development on the one hand and their degeneration on the other,
especially in the smaller groups, examples of _progressive inheritance_.
Thus in the SPUMELLARIA the constant formation of the primary lattice-shell
(a central medullary shell) and its ontogenetic relation to the secondary
one, which is developed concentrically round it, can only be explained
phylogenetically by conservative inheritance, whilst on the other hand the
characteristic differentiation of the axes in the various families of
SPUMELLARIA is to be explained by progressive inheritance. In the
ACANTHARIA the arrangement of the twenty radial spines (in accordance with
Müller's law, §§ 110, 172) was first acquired by a group of the most
archaic #Actinelida# (Adelacantha) through hydrostatic adaptation, and has
since been transmitted by inheritance to all the other families of the
legion (Icosacantha). The morphology of the NASSELLARIA is not less
interesting, because here several different heritable elements (the primary
sagittal ring and the basal tripod) combine in the most manifold ways in
the formation of the skeleton (compare §§ 123, 124, 182). The affinities of
the genera in the different families yield an astonishing variety of
interesting morphological phenomena, which can only be explained by
progressive inheritance. The same is true also of the PHÆODARIA. In this
legion the primary inheritance is especially manifested in the constant and
firm structure of the central capsule with its characteristic double wall
and astropyle, whilst the formation of the skeleton in this legion proceeds
in different directions by means of divergent adaptation. The morphology of
the Radiolaria thus proves itself a rich source of materials for the
physiological study of adaptation and inheritance.

{cxl}CHAPTER VIII.--ANIMAL FUNCTIONS.

(§§ 218-225.)

218. _Motion._--In addition to the internal movements which appear
generally in the unicellular Radiolaria and have already been mentioned as
plasmatic currents in treating of the circulation (§§ 207-209), two
different groups of external motor phenomena are to be observed in this
class: first, the _contraction_ of individual parts, which brings about
modifications of form (§ 220), and secondly, voluntary or reflex
_locomotion_ of the whole body (§ 220). These movements are partly due to
changes in form of undifferentiated plasmatic threads or sarcode filaments,
partly to the actual contraction of differentiated filaments which are
comparable to muscle fibrillæ, and must therefore be distinguished as
myophanes. In addition to this, endosmose and exosmose probably play an
important part in some of the locomotive phenomena, but nothing is yet
certainly known regarding these osmotic processes. We are at present
equally ignorant whether all the movements of the Radiolaria are simply
reflex (direct consequences of irritation) or whether they are in part
truly spontaneous.

219. _Suspension._--From direct observation of living Radiolaria, as well
as from deductive reasoning, based upon their morphology (and especially
their promorphology, §§ 17-50), the conclusion appears justified that all
Protista of this class in their normal condition float suspended in the
sea-water, either at the surface or at a definite depth. A necessary
condition of this hydrostatic suspension is that the specific gravity of
the Radiolarian organism must be equal to, or but slightly greater than
that of sea-water. The increase in specific gravity brought about by the
production of the siliceous skeleton, is compensated by the lighter fatty
globules, and partly perhaps by the calymma, especially when the latter
contains vacuoles or alveoles. The fluid or jelly contained in the latter
appears to be for the most part lighter than sea-water (containing no salt,
or only a very small quantity?). But if the specific gravity of the whole
body should be generally (or perhaps always) slightly greater than that of
sea-water, then the organism would be prevented from sinking, partly by the
increased friction, due to the radiating pseudopodia and the radial spines
usually present, and partly perhaps by active (if only feeble) movements of
the pseudopodia.

220. _Locomotion._--Active locomotion of the whole body, which is very
probably to be regarded as voluntary, occurs in the Radiolaria in three
different modes; (1) the vibratile movement of the flagellate swarm-spores;
(2) the swimming of the floating organisms; (3) the slow creeping of those
which rest accidentally upon the bottom. {cxli}The _vibratile_ movement of
the swarm-spores is the result of active sinuous oscillation of the single
or multiple flagellum, and is not essentially different from that of
ordinary flagellate Infusoria (see note A). Of the active swimming of
mature Radiolaria, only that form is known which is vertical in direction
and causes the sinking and rising in the sea-water. This is probably, for
the most part (perhaps exclusively), due to increase or diminution in the
specific gravity, which is perhaps brought about by the retraction or
protrusion of the pseudopodia; slow, oscillating, sinuous motions of these
organs have been directly observed to take place (though very slowly) in
suspended living Radiolaria. The most important hydrostatic organ is
probably the calymma, by the contraction of which the specific gravity is
increased, while it is diminished by its expansion; the contraction is
probably brought about by active contraction of the sarcodictyum, and is
connected with exosmosis, while the expansion is probably due to the
elasticity of the calymma and the inception of water by endosmosis. In the
#Acanthometra# (§ 96) the peculiar myophriscs appear to be charged with the
duty of distending the gelatinous envelope, and thus diminishing the
specific gravity; the latter increases again when the myophriscs are
relaxed, and the calymma contracts by virtue of its own elasticity (see
note B). The slow _creeping locomotion_ exhibited by Radiolaria on a glass
slide under the microscope, does not differ from that of the Thalamophora
(Monothalamia and Polythalamia), but can only occur normally when the
animal accidentally comes into contact with a solid surface or sinks to the
bottom of the sea. Whether this actually occurs periodically is not known
(see note C). The slow or gliding locomotion exhibited by creeping Monozoa
on a glass slide is due to muscle-like contractions of bundles of
pseudopodia, just as in the case of the social central capsules of Polyzoa,
which live together in the same coenobium and are able to move within their
common calymma sometimes centrifugally to its surface, sometimes towards
the centre where they aggregate into a roundish mass (see note D).

A. Regarding the movement of the flagella in mature swarm-spores compare
L. N. 22, p. 375; L. N. 26, pp. 31, 35; L. N. 41, p. 452, and L. N. 52,
p. 170.

B. On the active vertical swimming movements of mature Radiolaria,
especially the cause of sinking and rising, see L. N. 16, p. 134; L. N.
41, p. 443, and L. N. 52, pp. 97-102.

C. On the active horizontal creeping movements of mature Radiolaria on a
firm ground, compare L. N. 12, p. 10, and L. N. 16, pp. 132-134.

D. Regarding the motion of social central capsules within the same
coenobium and the changes thus brought about in the structure of the
calymma, see L. N. 16, pp. 119-127, and L. N. 52, pp. 75-82.

221. _Contraction._--Motions, which are due to the contraction of
individual portions and cause changes in volume or form, have been partly
already spoken of under the head of locomotion (§ 220) and are partly
connected with other functions. Examples may be seen in the contraction of
the central capsule and of the calymma. A certain {cxlii}contraction of the
central capsule is probably brought about by the myophanes, which arise by
differentiation of the endoplasm and hence may assume different forms in
the four legions. In the SPUMELLARIA, where numerous radial fibrillæ run
from the central nucleus to the capsule membrane (§ 77), the endoplasm is
probably driven out evenly through all the pores of the capsule membrane by
their simultaneous contraction, and hence the volume of the capsule is
diminished in all directions. The ACANTHARIA probably behave similarly,
but are different, inasmuch as the number of their contractile radial
fibrillæ is less, and special axial threads (§ 78) are already
differentiated. In the NASSELLARIA it is probable that owing to the
contraction of the divergent myophane fibrillæ in the podoconus the
vertical axis of the latter is shortened, the opercular rods of the
porochora are lifted, and the endoplasm driven out of its pores, so that
the volume of the monaxon central capsule is diminished (§ 79). In the
PHÆODARIA the same result is probably brought about by the contraction of
the cortical myophane fibrillæ, which run meridionally along the inside of
the capsule membrane from the apical to the basal pole of the vertical main
axis, where they are inserted into the periphery of the astropyle; since
the volume of the capsule is diminished by their contraction (their
spheroidal figure becoming more nearly spherical) the endoplasm will be
driven out through the proboscis of the astropyle. Whilst these
contractions of the central capsule are largely due to differentiated
muscle-like threads of endoplasm (myophanes), this appears to be but rarely
the case with the contractions of the extracapsulum (_e.g._, the myophriscs
of the #Acanthometra#, § 96). Most of the phenomena of contraction which
can be observed in the calymma and pseudopodia depend upon exoplasmatic
currents (§ 209).

222. _Protection._--Of the utmost importance, both for the physiology and
for the morphology of the Radiolaria are their manifold protective
functions, which we now consider under the heading "protection." From the
physiological point of view the consideration of the exposed situation in
which the delicate, free-swimming Radiolarian organism lives, and the
numerous dangers which beset it in the struggle for existence, would lead
_a priori_ to the expectation, that many special protective adaptations
would be developed by natural selection. On the other hand, morphological
experience shows us that this latter has been in action for immeasurable
periods, and has gradually produced an abundance of the most remarkable
protective modifications. Examples of these may be found in the formation
of the voluminous calymma, as a gelatinous protective covering for the
central capsule, and further, the formation of the capsule membrane itself,
which separates the generative contents of the central capsule from the
nutritive exoplasm. The phosphorescence of the central capsule, too (§
223), may be regarded as a useful protective arrangement; as also the
radiating of the numerous pseudopodia in all directions from the surface of
the calymma; for they are of great significance to the {cxliii}well-being
of the organism, both as sensory organs and as prehensile organs. By far
the most important and most varied means for the actual defence of the soft
body is to be seen in the endless modifications of the skeleton; first, in
the production of the enclosing lattice-shells and projecting radial
spines, but especially also in the very varied structure of the individual
parts of the skeleton, and in the special differentiation of the small
appendicular organs which grow out from it (hairs, thorns, spines, scales,
spathillæ, anchors, &c.). Finally "mimicry" possesses a considerable
significance among the different forms of adaptation which are to be
observed in this class.

223. _Phosphorescence._--Many Radiolarians shine in the dark, and their
phosphorescence presents the same phenomena as that of other luminous
marine organisms; it is increased by mechanical and chemical irritation, or
renewed if already extinguished. The light is sometimes greenish, sometimes
yellowish, and appears generally (if not always) to radiate from the
intracapsular fatty spheres (§ 73). Thus these latter unite several
functions, inasmuch as they serve, firstly, as reserve stores of nutriment,
secondly, as hydrostatic apparatus, and thirdly, as luminous organs for the
protection of the Radiolaria; probably the light acts by frightening other
animals, for the phosphorescent animals are provided with spines,
nettle-cells, poison glands or other defensive weapons. The production of
the light depends probably, as in other phosphorescent organisms, upon the
slow oxidation of the fat-globules, which combine with active oxygen in the
presence of alkalis. Phosphorescence is very likely widely distributed
among the Radiolaria.

The shining of the Radiolaria in the dark has been noticed by the
earliest observers of the class (see L. N. 1, p. 163, L. N. 16, p. 2, and
L. N. 52, pp. 136-139). In the winter of 1859 I observed the production
of light in the case of many monozootic and polyzootic Radiolaria, but
inadvertently omitted to record the fact in my Monograph. I made more
accurate observations in the winter of 1866 at Lanzerote in the Canary
Islands, and convinced myself the the light emanates from the central
capsule, and in particular from the fat-globules contained in it. In most
Polycyttaria (both #Collosphærida# and #Sphærozoida#), when each central
capsule contains a large central oil-globule the light radiates from it.
In _Collozoum serpentinum_ (Pl. 3, figs. 2, 3) each cylindrical central
capsule contains a row of luminous spherules like a string of beads. In
_Alacorys friderici_ (Pl. 65, fig. 1) the four-lobed central capsule
contains four shining points. Karl Brandt has recently made more detailed
communication on this point (L. N. 52, p. 137).

224. _Sensation._--The general irritability which we ascribe to all
organisms, and as the basis of which we regard the protoplasm, remains at
an inferior stage of development in the Radiolaria. For although they are
subject to various stimuli, and certainly possess a power of
discrimination, special sensory organs are not differentiated; the
peripheral portions of the protoplasm, and especially the pseudopodia,
rather act both as organs of the different kinds of sensation and various
modes of motion. That different Radiolaria have attained different degrees
of development in this respect may be seen {cxliv}partly by direct
observation of the reaction of the living organism towards various stimuli,
and partly by the comparison of the different conditions of existence under
which Radiolarians exist, both in the most various depths of the ocean and
in all climatic zones (see note A). In general the Radiolaria seem to be
sensitive to the following stimuli; (1) pressure (see note B); (2)
temperature (see note C); (3) light (see note D); (4) chemical composition
of the sea-water (see note E). The reaction towards these stimuli,
corresponding to the sensation of pleasure or dislike which they call
forth, is shown in various forms of motion of the protoplasm, changes in
the currents in it, contraction of the central capsule, changes in the
size, position, and form of the pseudopodia, changes in the volume of the
calymma (by the evocation of water), &c. Among the sensory functions of the
Radiolaria must be especially mentioned their remarkably developed
perception of hydrostatic equilibrium (see note F), as well as their
perception of distances, so clearly shown in the production of equal
lattice-meshes and other regularly formed skeletal structures (see note G).

A. I can add but little to the communication which I made twenty-four
years ago regarding sensation in the Radiolaria (L. N. 16, pp. 128-131).
The most important point would be the great difference in irritability
which must obtain between the pelagic, zonarial and abyssal Radiolaria,
which may be assumed from a consideration of their very different
conditions of existence as regards pressure, light, warmth, nutrition,
&c. It is natural to suppose that the numerous abyssal Radiolaria,
discovered by the Challenger, which live at great depths (2000 to 4500
fathoms) in complete darkness, in icy cold and under an enormous
pressure, must have quite different sensations of pleasure from their
pelagic relatives which live at the surface of the sea under an
equatorial sun. Karl Brandt has recently added much to our knowledge
regarding the special action of different vital conditions upon the
various Polycyttaria and the degrees of their irritability (L. N. 52, pp.
113-132).

B. Regarding the sensation of pressure or sensation of touch of the
Radiolaria and the various degrees of their mechanical irritability, see
L. N. 16, p. 129; L. N. 41, p. 464.

C. Regarding the sensation of warmth or temperature-sense and its
dependence upon different climatic relations, see L. N. 16, p. 129; L. N.
52, pp. 114-129.

D. Regarding the sensation of light, compare L. N. 16, p. 128; L. N. 42,
p. 304; L. N. 52, pp. 102-104, 114.

E. Regarding the sense of taste of the Radiolaria or their peculiar
sensitiveness towards the different chemical composition of the water,
change in its salinity, presence of organic impurities, &c., see L. N.
16, p. 130; L. N. 52, pp. 103, 113. This chemical irritability seems to
be the most highly developed sense in the Radiolaria, even more so than
their mechanical irritability.

F. The perception of hydrostatic equilibrium among the Radiolaria is
immediately visible from the position which their bodies, floating freely
in the water, assume spontaneously, and from the symmetrical development
of the skeleton, which by its gravitation necessitates a definite
position. It may be assumed that the development of the various
geometrical ground forms which correspond to a definite position of
equilibrium, is the result of this particular kind of perception (compare
§§ 40-45).

{cxlv}G. The plastic perception of distance of the pseudopodia is shown
by the symmetry with which the forms composing the regular skeletal
structures (_e.g._, the ordinary lattice-spheres with regular hexagonal
meshes, the radial spines with equidistant branches) are excreted from
the exoplasm. Both this form of sensation and the one first mentioned
(note F) have hitherto received scarcely any attention, but are deserving
of a thorough physiological investigation.

225. _The Cell-Soul (Zellseele)._--The common central vital principle,
commonly called the "soul," which is considered to be the regulator of all
vital functions, appears in the Radiolaria as in other Protista in its
simplest form, as the cell-soul. By the continual activity of this central
"psyche" all vital functions are maintained in unbroken action, and in
uniform correlation. It is also probable that by it the stimulations which
the peripheral portions of the cell receive from the outer world are first
transmitted into true sensation, and that, on the other hand, the volition,
which alone calls forth spontaneous movements, proceeds from it. The
central capsule is most likely the sole organ of this cell-soul or central
psychic organ, and the active portion may be either the endoplasm or the
nucleus, or both. The central capsule may thus (apart from its function as
a sporangium, § 215) be regarded as a simple ganglion cell, physiologically
comparable to the nervous centre of the higher animals, whilst the exoplasm
(sarcomatrix and pseudopodia) are to be compared to the peripheral nervous
system and sense organs of the latter. The great simplicity of the
functions of the cell-soul which appear in the Radiolaria, and the intimate
connection of their different psychic activities, give to these unicellular
Protista a special significance for the comprehension of the monistic
elements of a natural psychology.

Regarding the theory of the cell-soul as the only psychological theory
which is able to explain naturally the true nature of the life of the
soul in all organisms as well as in man, see my address on cell-souls and
soul-cells ("Zellseelen und Seelenzellen") in Gesammelte populäre
Vorträge aus dem Gebiete der Entwickelungslehre, Heft 1, p. 143; Bonn,
1878.

{cxlvi}CHOROLOGICAL SECTION.

----

CHAPTER IX.--GEOGRAPHICAL DISTRIBUTION.

(§§ 226-240.)

226. _Universal Marine Distribution._--Radiolaria occur in all the seas of
the world, in all climatic zones and at all depths. Probably under normal
conditions they always float freely in the water, whether their usual
position be at the surface (pelagic), or at a certain depth (zonarial), or
near to the bottom of the sea (abyssal). This appears both from numerous
direct observations, as well as from conclusions which may be drawn from
their organisation (and especially their promorphology) regarding their
floating life (compare §§ 40-50, 219, 220). Hitherto no observation has
been recorded, which justifies the assumption that Radiolaria live anywhere
upon the bottom of the sea (on stones, Algæ, or other firm substances),
either sessile or creeping. They perform the latter action, however, when
they fall accidentally upon a firm basis or are accidentally placed upon
it, but they seem normally always to float freely in the water with
pseudopodia radiating in all directions. Active free-swimming movements are
only met with in the case of the flagellate zoospores (§ 142). The
development of Radiolaria in large masses is very remarkable (see note A),
and in many parts of the ocean is so great that they play an important part
in the economy of marine life, especially as food for other pelagic and
abyssal animals (see note B). Medium salinity of the water seems to be most
favourable to their development in masses, although it is not unknown in
seas of high and low salinity (see note C). There are no Radiolaria in
fresh water (see note D).

A. The development of Radiolaria takes place in many parts of the ocean
in astonishingly large masses on the surface, in different strata, and
near the bottom. The #Collodaria# (and especially the Sphærozoida) often
cover the surface of the sea in millions, and form a shining layer,
phosphorescent in the dark like the _Noctilucæ_, as I observed in 1859 in
the Strait of Messina, in 1866 at the Canaries, and in 1881 in the Indian
Ocean. Similar masses of _Sphærozoum_ and _Acanthometron_ were seen by
Johannes Müller on the French and Ligurian coasts (L. N. 12), and John
Murray found another in the Gulf Stream, off the Færöe Islands, from the
surface to a depth of 600 fathoms; considerable masses of large PHÆODARIA
live there also.

B. The alimentary canal of Medusæ, Salpæ, Crustacea, Pteropoda, and many
other pelagic animals is a rich field for the discovery of Radiolaria,
and many of the species hereinafter described are from such sources.
Fossil coprolites too (_e.g._, those from the Jura) often contain many
Polycystina.

C. Some ACANTHARIA (#Acanthometra#) and PHÆODARIA (species of _Mesocena_
and _Dictyocha_) {cxlvii}live in the Baltic; I found their skeletons in
the alimentary canal of _Aurelia_, Ascidians and Copepods.

D. The so-called "fresh-water Radiolaria," which have been described by
Focke, Greeff, Grenacher and others, are all Heliozoa, without either
central capsule or calymma.

227. _Local distribution._--As regards their local distribution and its
boundaries the Radiolaria show in general the same relations as other
pelagic animals. Since they are only to a very slight extent, if at all,
capable of active horizontal locomotion, the dispersion of the different
species from their point of development (or "centre of creation") is
dependent upon oceanic currents, the play of winds and waves and all the
accidental causes which influence the transport of pelagic animals in
general. These passive migrations are here, however, as always, of the
greatest significance, and bring about the wide distribution of individual
species in a far higher degree than any active wanderings could do. Any one
who has ever followed a stream of pelagic animals for hours and seen how
millions of creatures closely packed together are in a short time carried
along for miles by such a current, will be in no danger of underestimating
the enormous importance of marine currents in the passive migration of the
fauna of the sea. Such constant currents may, however, be recognised both
near the bottom of the sea and at various depths, as well as at the
surface, and are therefore of just as much significance for the abyssal and
zonarial as for the pelagic Radiolaria. It is easy to explain by this means
how it is that so many animals of this class (probably indeed the great
majority) have a wide range of distribution. The number of _cosmopolitan_
species which live in the Pacific, Atlantic and Indian Oceans is already
relatively large. In each of these three great ocean basins, too, many
species show a wide distribution. On the other hand, there are very many
species which are hitherto known only from one locality, and probably many
small local faunas exist, characterised by the special development of
particular groups. The observations which we at present possess are too
incomplete, and the rich material of the Challenger is too incompletely
worked out, to enable any definite conclusions to be drawn regarding the
local distribution of Radiolaria.

The statements made in the systematic portion of this Report regarding
the distribution of the Challenger Radiolaria are very incomplete. In
most cases only one locality is mentioned, and that is the station (§
240) in the preparations or bottom deposit from which I first found the
species in question. Afterwards I often found the same species again in
one or more additional stations (not seldom in numerous preparations both
from the Pacific and Atlantic), without the possibility of adding them to
the habitat recorded under the description. The necessary accurate
determination and identification of the species (measuring the different
dimensions, counting the pores, &c.), would have occupied too much time,
and the writing of this extensive Report would have lasted not ten but
twenty or thirty years.

228. _Horizontal Distribution._--From the extensive collections of the
Challenger and from the other collections which have furnished a welcome
supplement to them, it appears {cxlviii}that Radiolaria are distributed
throughout all seas without distinction of zones and physical conditions,
even though these latter may be the cause of differences in their
qualitative and quantitative development. In the case of the Radiolaria as
well as of many other classes of animals, the law holds good that the
richest development of forms and the greatest number of species occurs
between the tropics, whilst the frigid zones (both Arctic and Antarctic)
exhibit great masses of individuals, but relatively few genera and species
(see note A). In the Challenger collection the greatest abundance of
species of Radiolaria is exhibited by those preparations which were
collected at low latitudes in the immediate neighbourhood of the equator;
this is true both of the Atlantic (Stations 346 to 349) and of the Pacific
(Stations 266 to 274); in the former the richest of all is Station 347
(lat. 0° 15' S.), in the latter Station 271 (lat. 0° 33' S.) (see note B).
From the tropics the abundance of species seems to diminish regularly
towards the poles, and more rapidly in the northern than in the southern
hemisphere; the latter also appears, considered as a whole, to possess more
species than the former. A limit to the life of the Radiolaria towards the
poles has not yet been found; the expeditions towards the North Pole (see
note C), like those towards the South (see note D), have obtained
bottom-deposits and ice enclosures which contained Radiolaria; in some of
the most northerly and most southerly positions which were reached the
number of Radiolaria enclosed in the ice was relatively great.

A. The greater abundance of Radiolaria in the tropical seas is probably
to be explained by the more favourable conditions of existence, and in
particular the larger quantity of nutritive material (especially of
decayed animals) and not by the higher temperature of the surface, for at
depths of from 2000 to 3000 fathoms where the abyssal Radiolaria live,
the temperature is but little above the freezing point or even below it
(compare the bottom temperatures in the list of Challenger Stations, §
240).

B. Station 271 of the Challenger Expedition, situated almost on the
equator in the Mid Pacific (lat. 0° 33' S.), exceeds all other parts of
the earth, hitherto known, in respect of its wealth in Radiolaria, and
this is true of the pelagic as well as of the zonarial and abyssal forms.
In the Station List the deposit at this point is stated to be
"Globigerina ooze"; but after the calcareous matter has been removed by
means of acid, the purest Radiolarian ooze remains, rich in varied and
remarkable species. More than one hundred new species have been described
from this Station alone.

C. Regarding the Arctic Radiolaria compare the contributions of Ehrenberg
(L. N. 24, pp. 138, 139, 195) and Brady on the English North Polar
Expedition, 1875-76 (Ann. and Mag. Nat. Hist., 1878, vol. i. pp. 425,
437).

D. Regarding the Antarctic Radiolaria, compare § 230, note A, and
Ehrenberg, Mikrogeologie (L. N. 6, Taf. xxxv., A.), also L. N. 24, pp.
136-139.

229. _Fauna of the Pacific Ocean._--From the splendid discoveries of the
Challenger, and the supplementary observations obtained from other sources,
the Pacific seems to be the ocean basin which is richest both
quantitatively and qualitatively in Radiolarian life, {cxlix}excelling both
the Indian and Atlantic Oceans in this respect. It may be assumed with
great probability that by far the largest portion of the Pacific has a
depth of between 2000 and 3000 fathoms, and that its bottom is covered
either with Radiolarian ooze (§ 237) or with a red clay (§ 239), which
contains many SPUMELLARIA and NASSELLARIA, and has probably been derived
for a great part from broken down and metamorphosed Radiolarian ooze (see
note A). Pure Radiolarian ooze was found by the Challenger eastwards in the
Central Pacific (over a wide area between lat. 12° N. and 12° S., Stations
265 to 274), and also westwards in the latitude of the Philippines, twenty
degrees to the east of them (between lat. 5° N. and 15° N.). The great
abundance of Radiolaria present in the neighbourhood of the Philippines and
in the Sunda Sea was already known from other investigations (note B). The
red clay also, which covers a great part of the bottom of the North
Pacific, and which was obtained of very constant composition by the
Challenger between lat. 35° N. and 38° N., from Japan to the meridian of
Honolulu (from long. 144° E. to 156° W.), is so pre-eminently rich in
Radiolaria that it often approaches in composition the Radiolarian ooze,
and has probably been derived from it. The track of the Challenger through
the tropical and northern parts of the Pacific describes nearly three sides
of a rectangle, which includes about half of the enormous Pacific basin,
and from this as well as from other supplementary observations it may with
great probability be concluded that by far the largest part of the bed of
the Pacific (at least three-fourths) is covered either with Radiolarian
ooze or with red clay, which contains a larger or smaller amount of the
remains of Radiolaria. With this agrees also the important fact that the
numerous preparations of pelagic materials and collections of pelagic
animals, which were collected by the Challenger in the Pacific, almost
always indicate a corresponding amount of Radiolarian life on the surface.
This is true in particular also of the South Pacific, between lat. 33° S.
and 40° S. (from long. 133° W. to 73° W., Stations 287 to 301); the surface
of this southern region and the different bathymetrical zones were rich in
new and peculiar species of Radiolaria.

A. Many specimens of bottom-deposits from the Pacific, which are entered
in the Challenger lists either as "red clay" or "Globigerina ooze,"
contain larger or smaller quantities of Radiolaria, and the number of
different species of SPUMELLARIA and NASSELLARIA which they contain is
often so great that the deposit might have been almost as appropriately
termed "Radiolarian ooze," _e.g._, Stations 241 to 245, and 270, 271
(compare §§ 236-239).

B. Pacific Radiolarian ooze was first obtained by Lieutenant Brooke (May
11, 1859) between the Philippines and Marianne Islands, from a depth of
3300 fathoms (lat. 18° 3' N., long. 129° 11' E.). Ehrenberg, who first
described it, found seventy-nine different species of Polycystina in it,
and reported "that their quantity and the number of different forms
increased with the depth" (Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, 1860, pp. 466, 588, 766).

230. _Fauna of the Indian Ocean._--As regards its Radiolarian fauna the
Indian Ocean is the least known of the three great basins. Still the few
limited spots, regarding which {cl}investigations are forthcoming, indicate
a very rich development of Radiolarian life. Probably it approaches more
nearly the fauna of the Pacific than that of the Atlantic, both as regards
the abundance and the morphological characters of its species. The
researches of the Challenger are very limited and incomplete as regards the
Indian Ocean, for the expedition only just touched upon this great ocean
basin (2000 to 3000 fathoms deep) at its two extremities (westwards at the
Cape of Good Hope and eastwards at Tasmania), its course lying for the most
part south of lat. 45° S. and extending beyond lat. 65° S. (from Station
149 to 158, south of lat. 50° S.). It is true that this portion of the
South Indian Ocean was shown to contain Radiolaria everywhere, but these
were more plentiful in individuals than in species. Only from Station 156
to Station 159 (between lat. 62° and 47° S., and long. 95° and 130° E.) was
the bottom, which consisted partly of Diatom ooze and partly of Globigerina
ooze, richer in species (see note A). The gaps left by the Challenger in
the investigation of the Indian Ocean, have, however, been to some extent
filled from other sources. As early as 1859 the English "Cyclops"
expedition had shown that the bottom of the Indian Ocean to the east of
Zanzibar (lat. 9° 37' S., long. 61° 33' W.) is covered with pure
Radiolarian ooze (see note B). Also since the Tertiary rocks of the Nicobar
Islands are for the most part of the same composition, and since a great
abundance of Radiolaria has been shown to be present both in the east part
of the ocean, between the Cocos Islands and the Sunda Archipelago (see note
C), and in the northern part or Arabian Sea between Socotra and Ceylon (see
note D); it may be assumed with great probability that the greater part of
the basin of the Indian Ocean, like that of the Pacific, is covered either
with Radiolarian ooze or with the characteristic red clay. With this agrees
the richness of the surface of the Indian Ocean in Radiolaria of the most
various groups, which has been more extensively demonstrated.

A. The Radiolarian fauna collected by the Challenger on the voyage from
the Cape to Melbourne, shows in part, namely, from Station 156 to Station
158, very peculiar and characteristic composition; in particular, the
Diatom ooze of Station 157 passes over in great part into a Radiolarian
ooze, mainly composed of #Sphærellaria#. This is worthy of a more
thorough investigation than I was able, owing to lack of material and
time, to give it.

B. The remarkably pure Radiolarian ooze of Zanzibar, discovered by
Ehrenberg in 1859, was the earliest known recent example of that deposit.
It was brought up by Captain Pullen of the English man-of-war "Cyclops,"
from a depth of 2200 fathoms, between Zanzibar and the Seychelles, and
"under a magnifying power of 300 diameters, showed at the first glance a
mass of almost pure Polycystina, such as no sample of a deep-sea deposit
has hitherto shown. It is very noticeable that in the whole of this mass
of living forms, no calcareous shells are to be seen" (Ehrenberg, L. N.
24, pp. 148, 149).

C. For the most important material from the Indian Ocean, I am indebted
to Captain Heinrich Rabbe of Bremen, who during many voyages in the
Indian Ocean, in his ship "Joseph Haydn," made numerous collections in
different localities with the tow-net and the trawl, and admirably
preserved the rich collections thus made. The greatest abundance of
Radiolaria was found in those {cli}obtained to the east of Madagascar,
and next in those from the neighbourhood of the Cocos Islands. I take
this opportunity of expressing my thanks to Captain Rabbe for the
liberality with which he placed all this valuable material at my
disposal.

D. On my voyage from Aden to Bombay, and thence to Ceylon (1881), and
especially on my return journey from Ceylon, between the Maldive Islands
and Socotra (1882), I carried on a number of experiments with a surface
net, which yielded a rich fauna of pelagic animals, and among them many
new species of Radiolaria, for observation. On several nights when the
smooth surface of the Indian Ocean, unrippled by any wind, shone with the
most lovely phosphorescent light, I drew up water from the surface with a
bucket, and obtained a rich booty. A number of other new species of
Radiolaria from very various parts of the Indian Ocean I obtained from
the alimentary canal of pelagic animals, such as Medusæ, Salpæ,
Crustacea, &c. Although the total number of Radiolaria known to me from
the Indian Ocean is much less than from the Atlantic and Pacific, there
are several new genera and numerous species among them, which show that a
careful study of this fauna will be of wide interest.

231. _Fauna of the Atlantic Ocean._--The Atlantic Ocean in all parts, of
which the pelagic fauna has been examined, has shown the same constant
presence of Radiolaria, and in certain parts of its abyssal deposits a
larger or smaller quantity of different types belonging to this class; on
the whole, however, its Radiolarian fauna is inferior to that of the
Pacific, and probably also to that of the Indian Ocean, both in quantity
and quality. Pure Radiolarian ooze, such as is so extensively found on the
floor of the Pacific, and in certain places in that of the Indian Ocean,
has not yet been found in the Atlantic (see § 237). The red clay, too, of
the deep Atlantic does not seem to be so rich in Radiolaria as that of the
Pacific; nevertheless, the number of species peculiar to the Atlantic is
very large, and at certain points the abundance of species as well as of
individuals seems to be scarcely less than in the Pacific. This is
especially true of the eastern equatorial zone not far from Sierra Leone,
Stations 347 to 352 (see note A); also of the South Atlantic between Buenos
Ayres and Tristan da Cunha, Stations 324, 325, 331 to 333 (see note B);
and, lastly, in the North Atlantic in the Gulf Stream and near the Canary
Islands (see note C). The fauna of the latter agrees for the most part with
that of the Mediterranean (see note D). In addition to the material
collected by the Challenger, other deep-sea investigations have furnished
bottom-deposits from different parts of the ocean, which have proved very
rich in Radiolaria (see note E). Furthermore, since the island of Barbados
consists for the most part of fossil Radiolarian ooze, it is very probable
that at certain parts of the tropical Atlantic true Radiolarian ooze, like
that of the Pacific and Indian Oceans, will eventually be found in depths
between 2000 and 3000 fathoms, perhaps over a considerable area.

A. The tropical zone of the eastern Atlantic seems to be especially rich
in peculiar Radiolaria of different species. This is shown by numerous
preparations from the surface, and from various depths (between lat. 3°
S. and 11° N., and long. 14° W. to 18° W.), which were made towards the
{clii}end of the cruise. Unfortunately no bottom-deposits were obtained
from the most important stations (except Nos. 346 and 347, depths 2350
and 2250 fathoms) in this region; at these the deposit was a Globigerina
ooze containing numerous different species of Radiolaria.

B. In the South Atlantic, between Buenos Ayres and Tristan da Cunha
(between lat. 35° S. and 43° S., long. 8° W. and 57° W.) there appears to
be a long stretch covered partly with Globigerina ooze (Stations 331 to
334), or red clay (Stations 329, 330), partly with blue mud (Stations 318
to 328), which contains not only large masses of individuals but numerous
peculiar species of SPUMELLARIA and NASSELLARIA. The preparations from
the surface-takings of this region are also rich in these, as well as in
peculiar PHÆODARIA.

C. The northern part of the Atlantic appears on the whole to be inferior
to the tropical and southern portions as regards its richness in
Radiolaria, and from the western half more especially, only few species
are known. From my researches at Lanzerote in 1866-67, it appears that
the pelagic fauna of the Canary Islands is very rich in them, as is also
the Gulf Stream in the neighbourhood of the Færöe Channel, according to
the investigations of John Murray (see his Report on the "Knight-Errant"
Expedition, Proc. Roy. Soc. Edin., vol. xi., 1882).

D. The Radiolaria of the Mediterranean are of special interest, because
almost all our knowledge of these organisms in the living conditions and
of their vital functions has been derived from investigations conducted
on its shores. Johannes Müller laid the foundation of this knowledge by
his investigations at Messina, and on the Ligurian and French coasts at
Nice, Cette, and St. Tropez (L. N. 10). The many new Radiolaria which I
described in my Monograph (L. N. 16, 1862), were for the most part taken
at Messina, the place which possesses a richer pelagic fauna than any
other, so far as is yet known, in the Mediterranean. Other new species I
found afterwards at Villafranca near Nice, in 1864 (L. N. 19), at
Portofino near Genoa (1880), at Corfu (1877), and at other points on the
coast. In Messina also, Richard Hertwig collected the material for his
valuable treatise on the Organisation of the Radiolaria (L. N. 33), after
he had previously made investigations into their histology at Ajaccio in
Corsica (L. N. 26). Lastly, at Naples, Cienkowski (L. N. 22) and Karl
Brandt (L. N. 38, 39, 52) carried out their important investigations into
the reproduction and symbiosis of the Radiolaria. With respect to the
character of its Radiolaria, the Mediterranean fauna is to be regarded as
a special province of the North Atlantic.

E. Among the smaller contributions which have been made towards our
knowledge of the Atlantic Radiolarian fauna, the communications of
Ehrenberg on the deposits obtained in sounding for the Atlantic cable,
and on the Mexican Gulf Stream near Florida, deserve special mention (L.
N. 24, pp. 138, 139-145).

232. _Vertical Distribution._--The most important general result of the
discoveries of the Challenger, as regards the vertical or bathymetrical
distribution of the Radiolaria, is the interesting fact that numerous
species of this class are found living at the most various depths of the
sea, and that certain species are limited to particular bathymetrical
zones, _i.e._, are adapted to the conditions which obtain there. In this
respect three different Radiolarian faunas may be distinguished, which may
be shortly termed "pelagic," "zonarial," and "abyssal." The _pelagic_
Radiolaria swim at the surface, and when they sink (_e.g._, in a stormy
sea), only descend to a small depth, probably not more than from {cliii}20
to 30 fathoms (§ 233). The complicated conditions of existence created by
the keen struggle for existence at the surface of the sea, give rise to the
formation of very numerous pelagic species, especially of Porulosa
(SPUMELLARIA and ACANTHARIA). The _abyssal_ Radiolaria are very different
from those just mentioned; they live at the bottom of the deep-sea, not
resting upon nor attached to it, but probably floating at a little distance
above it, and are adapted to the conditions of existence which obtain there
(§ 235). Here the Osculosa (NASSELLARIA and PHÆODARIA) seem to predominate.
The _zonarial_ Radiolaria live floating at various depths between the
pelagic and abyssal species (§ 234). In their morphological characters they
gradually approach the pelagic forms upwards and the abyssal downwards.

The views which have hitherto been held regarding the bathymetrical or
vertical distribution of the Radiolaria have been entirely altered by the
magnificent discoveries of the Challenger, and especially by the
important observations of Sir Wyville Thomson (L. N. 31) and John Murray
(L. N. 27). These two distinguished deep-sea explorers have, as a result
of their wide experience, been convinced that Radiolaria exist at all
depths of the ocean, and that there are large numbers of true deep-sea
species which are never found at the surface of the sea nor at slight
depths (L. N. 31, vol. i. pp. 236-238; L. N. 27, pp. 523, 525). The
result of my ten years' work upon the Challenger Radiolaria, and the
comparative study of more than a thousand mountings from all depths, has
only been to confirm this opinion, and I am further persuaded that it
will some day be possible by the aid of suitable nets (not yet invented)
to distinguish different faunistic zones in the various depths of the
sea. In this connection may be mentioned the specially interesting fact
that the species of Radiolaria of one and the same family present in the
different depths characteristic morphological distinctions, which
obviously correspond to their different physiological relations in the
struggle for existence. Owing to those extensive discoveries, the
representation which I gave in my Monograph (1862, L. N. 16, pp. 172-196)
of the vertical distribution of the Radiolaria, and of their life in the
greatest depths of the sea, has been entirely changed. Compare also
Bütschli (L. N. 41, p. 466).

233. _The Pelagic Fauna._--The surface of the open ocean seems everywhere,
at a certain distance from the coast at least, to be peopled by crowds of
living Radiolaria. In the tropical zone these pelagic crowds consist of
many different species, whilst in the frigid zones, on the other hand, they
are made up of many individuals belonging to but few species. Most of these
inhabitants of the surface may be regarded as truly pelagic species, which
either remain always at the surface or descend only very slightly below it.
Probably most Porulosa (both SPUMELLARIA and ACANTHARIA) belong to this
group; whilst but few Osculosa occur in it, and fewer PHÆODARIA than
NASSELLARIA. In general the pelagic Radiolaria are distinguished from the
abyssal by the more delicate and slender structure of their skeletons; the
pores of the lattice-shells are larger, the intervening trabeculæ thinner;
the armature of spines, spathillæ, anchors, &c., is more various and more
highly developed. Numerous forms are to be found among the pelagic
{cliv}Radiolaria which have either an incomplete skeleton or none at all.
When the pelagic forms leave the surface on account of unfavourable
weather, they appear only to sink to slight depths (probably not below 20
or 30 fathoms). Within the limits of the same family the size of the
pelagic species seems to be on an average greater than that of the related
abyssal forms.

234. _The Zonarial Fauna._--Between the pelagic fauna living at the surface
of the open sea and the abyssal, which floats immediately over the bottom,
there appears to be usually a middle fauna, which inhabits the different
bathymetrical zones of the intermediate water, and hence may be shortly
called the "zonarial" fauna. The different species of Radiolaria which
inhabit these different strata in the same vertical column of water present
differences corresponding to those of the plants composing the several
zones of vegetation, which succeed each other at different heights on a
mountain; they correspond to the different conditions of existence which
are presented by the different strata of water, and to which they have
become adapted in the struggle for existence. The existence of such
bathymetrical zones has been shown by those important, if not numerous,
observations of the Challenger, in which the tow-net was used at different
depths at one and the same Station. In several cases the character of the
Radiolarian fauna at different depths presented characteristic differences.

For the present, and until we are better acquainted with the characters
of the Radiolarian fauna at different depths, we may distinguish
provisionally the following _five bathymetrical zones_:--(1) The
_pelagic_ zone, extending from the surface to a depth of about 25
fathoms; (2) the _pellucid_ zone, extending from 25 to 150 fathoms, or as
far as the influence of the sunlight makes itself felt; (3) the _obscure_
zone, extending from 150 to 2000 fathoms, or from the depth at which
sunlight disappears to that at which the influence of the water
containing carbonic acid begins and the calcareous organisms vanish; (4)
the _siliceous_ zone, extending from 2000 or 2500 to about 3000 fathoms,
in which only siliceous not calcareous Rhizopoda are found, and in which
the peculiar conditions of the lowest regions have not yet appeared; (5)
the _abyssal_ zone, in which the accumulation of the oceanic deposits,
and the influence of the bottom currents, create new conditions of
existence. So far as our isolated and incomplete observations of the
zonarial Radiolarian fauna extend, it appears that the subclass Porulosa
(SPUMELLARIA and ACANTHARIA) predominates in the two upper zones, and as
the depth increases is gradually replaced by the subclass Osculosa
(NASSELLARIA and PHÆODARIA), so that the latter predominates in the two
lowest zones. The obscure zone which lies in the middle is probably the
poorest in species. In general, the morphological characters of the
zonarial fauna appear to change gradually upwards into the delicate form
of the pelagic and downwards into the robust constitution of the abyssal;
so also the average size of the individuals (within the limits of the
same family) appears to increase upwards and decrease downwards.

235. _The Abyssal Fauna._--The great majority of Radiolaria which have
hitherto been observed, and which are described in the systematic portion
of this Report, have been obtained from the bottom of the deep-sea, and
more than half of all the species have been {clv}derived from the pure
Radiolarian ooze, which forms the bed of the Central Pacific at depths of
from 2000 to 4000 fathoms (§ 237). Many of these abyssal forms were brought
up with the malacoma uninjured, and they show, both when mounted
immediately in balsam, and when preserved in alcohol, all the soft parts
almost as clearly as fresh preparations of pelagic Radiolaria. These
species are to be regarded as truly abyssal, _i.e._, as forms which live
floating only a little distance above the bottom of the deep-sea, having
become adapted to the peculiar conditions of life which obtain in the
lowest regions of the ocean. Probably the majority of the PHÆODARIA belong
to these abyssal Radiolaria, as well as a large number of NASSELLARIA, but
on the other hand, only a small number of ACANTHARIA and SPUMELLARIA are
found there. A character common to these abyssal forms, and rarely found in
those from the surface or from slight depths, is found in their small size
and their heavy massive skeletons, in which they strikingly resemble the
fossil Radiolaria of Barbados and the Nicobar Islands. The lattice-work of
the shell is coarser, its trabeculæ thicker and its pores smaller than in
pelagic species of the same group; also the apophyses (spines, spathillæ,
coronets, &c.), are much less developed than in the latter. From these true
abyssal Radiolaria must be carefully distinguished those species whose
empty skeletons, devoid of all soft parts, occur also in the Radiolarian
ooze of the deep-sea, but are clearly only the sunken remains of dead
forms, which have lived at the surface or in some of the upper zones.

236. _Deposits containing Radiolaria._--The richest collection of
Radiolaria is found in the deposits of ooze which form the bed of the
ocean. Although the pelagic material skimmed from the surface of the sea,
and the zonarial material taken by sinking the tow-net to various depths,
are always more or less rich in Radiolaria, still the number of species
thus obtained is, on the whole, much less than has hitherto been got merely
from deep-sea deposits. Of course the skeletons found in the mud of the
ocean-bed, may belong either to the abyssal species which live there (§
235), or to the zonarial (§ 234), or to the pelagic species (§ 233), for
the siliceous skeletons of these latter sink to the bottom after their
death. Almost all these remains found in the deposits belong to the
siliceous "Polycystina" (SPUMELLARIA and NASSELLARIA); PHÆODARIA occur but
sparingly, and ACANTHARIA are entirely wanting, for their acanthin skeleton
readily dissolves. The abundance of Radiolaria varies greatly according to
the composition and origin of the deposits. In general marine deposits may
be divided into two main divisions, terrigenous and abyssal, or, more
shortly, muds and oozes. The _terrigenous_ deposits (or muds) include all
those sediments which are made up for the most part of materials worn away
from the coasts of continents and islands, or brought down into the sea by
rivers. Their greatest extent from the coast is about 200 nautical miles.
They contain varying quantities of Radiolaria, but much fewer than those of
the next group. The _abyssal_ deposits (or oozes) usually commence at a
distance of from 100 to 200 nautical miles {clvi}from the coast. In general
they are characterised by great uniformity, corresponding to the constancy
of the conditions under which they are laid down; they may be divided into
three categories, the true Radiolarian ooze (§ 237), Globigerina ooze (§
238), and red clay (§ 239). Of these three most important deep-sea
formations the first is by far the richest in Radiolaria, although the
other two contain often very many siliceous shells.

The marvellous discoveries of the Challenger have thrown upon the nature
of marine deposits an entirely new light, which justifies most important
conclusions regarding the geographical distribution and geological
significance of the Radiolaria. Since Dr. John Murray and the Abbé Renard
will treat fully of these interesting relations in a forthcoming volume
of the Challenger series (Report on the Deep-Sea Deposits), it will be
sufficient here to refer to their preliminary publication already
published (Narrative of the Cruise of H.M.S. Challenger, 1885, vol. ii.
part ii. pp. 915-926); see also the earlier communications by John Murray
(1876, L. N. 27, pp. 518-537), and by Sir Wyville Thomson (The Atlantic,
L. N. 31, vol. i. pp. 206-246). In the Narrative (_loc. cit._, p. 916)
the following table of marine deposits is given:--

{Shore formations, } Found in inland
{Blue mud, } seas and along the
{Green mud and sand, } shores of
{Red mud, } continents.
Terrigenous deposits.{
{Volcanic mud and sand, } Found around oceanic
{Coral mud and sand, } islands and along the
{Coralline mud and sand, } shores of continents.

{Globigerina ooze, } Found in the abysmal
{Pteropod ooze, } regions of the
Abysmal deposits. {Diatom ooze, } ocean basins.
{Radiolarian ooze, }
{Red clay, }

237. _Radiolarian Ooze._--By Radiolarian ooze, in the strict sense of the
term, are understood those oceanic deposits, the greater part of which
(often more than three-quarters) is composed of the siliceous skeletons of
this class. Such _pure_ Radiolarian ooze has only been found in limited
areas of the Pacific and Indian Oceans. It is most conspicuous in the
Central Pacific, between lat. 12° N. and 8° S., long. 148° W. to 152° W.,
the depth being everywhere between 2000 and 3000 fathoms (Stations 266 to
268 and 272 to 274). In the deepest of the Challenger soundings (Station
225, 4475 fathoms) the bottom is composed of pure Radiolarian ooze, as well
as at the next Station in the Western Tropical Pacific (Station 226, 2300
fathoms), the latitude varying from 12° N. to 15° N., and the longitude
from 142° E. to 144° E. In the Indian Ocean also, pure Radiolarian ooze was
found in the year 1859 between Zanzibar and the Seychelles, this being the
first known example of it (§ 230). On the other hand, it has not yet been
found in the bed of the Atlantic; but the Tertiary formations of Barbados
(Antilles, § 231) like those of the Nicobar Islands (Further India), are to
be regarded as pure Radiolarian {clvii}ooze in the fossil condition.
_Mixed_ Radiolarian ooze is the name given to those deposits in which the
Radiolaria exceed any of the other organic constituents, although they do
not make up half the total mass. To this category belong a large number of
the Challenger soundings which are entered in the Station list either as
red clay or Globigerina ooze. Such mixed Radiolarian ooze has been
discovered (A) in the North Pacific in an elongated area of red clay
extending from Station 241 to Station 245 (perhaps even from Station 238 to
Station 253), that is, at least, from long. 157° E. to 175° E., between
lat. 35° N. and 37° N.; (B) in the tropical Central Pacific in the
Globigerina ooze of Stations 270 and 271. The ooze from the latter station,
situated almost on the equator (lat. 0° 33' S., long. 151° 34' W.), is
specially remarkable, for it has yielded more new species of SPUMELLARIA
and NASSELLARIA than any other Station, not excluding even the neighbouring
Stations 268, 269, and 272. Probably such mixed Radiolarian ooze is very
widely distributed in the depths of the ocean, as, for example, in the
South Pacific (Stations 288, 289, 300, and 302), and in the Southern Ocean
(Stations 156 to 159); also in the South Atlantic (Stations 324, 325, 331,
332) and in the tropical Atlantic (Stations 348 to 352). When carefully
purified and decalcified by acids, Radiolarian ooze appears as a fine
shining white powder; in the raw state it is yellowish or reddish,
sometimes reddish-brown or dark brown in colour, according to the quantity
of oxides of iron, manganese, &c., which it contains. Calcareous skeletons
(especially the tests of pelagic Foraminifera) do not occur at all or only
in very minute quantities in _pure_ Radiolarian ooze from more than 2000
fathoms, whilst specimens of _mixed_ ooze often contain considerable
quantities of them.

Pure Radiolarian ooze was first described by Dr. John Murray as regards
its peculiar nature and composition under the name "Radiolarian ooze"
(1876, L. N. 27, pp. 525, 526); compare also Sir Wyville Thomson (The
Atlantic, L. N. 31, vol. i. pp. 231-238), and John Murray (Narr. Chall.
Exp., L. N. 53, vol. i. pt. ii. pp. 920-926, pl. N. fig. 2). The
different specimens of pure Radiolarian ooze obtained by the Challenger
from the Pacific, and handed to me for investigation, are from depths of
from 2250 fathoms to 4475 fathoms, and may be divided according to their
composition into three different groups:--I. The Radiolarian ooze of the
Western Tropical Pacific, Stations 225 and 226, from depths of 4475 and
2300 fathoms (lat. 11° N. to 15° N., and long. 142° E. to 144° E.). II.
The Radiolarian ooze of the northern half of the Central Pacific,
Stations 265 to 269, from depths of 2550 to 2900 fathoms. III. The
Radiolarian ooze of the southern half of the Central Pacific, Stations
270 to 274, from depths of 2350 to 2925 fathoms. A fourth group would be
constituted by the Radiolarian ooze from the Philippines, which was
brought up by Brooke in 1860 near the Marianne Islands from 3300 fathoms,
and described by Ehrenberg (Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, 1860, p. 765). The Diatom ooze, too, found by the Challenger in
the Antarctic regions (Stations 152 to 157) is in some parts so rich in
Radiolaria that it passes over into true Radiolarian ooze. Regarding the
Radiolarian ooze from Zanzibar, obtained by Captain Pullen in 1859 from
2200 fathoms (§ 230), we have only the incomplete communications of
Ehrenberg (L. N. 24, p. 147). A more accurate knowledge of these deposits
from the Indian Ocean, and of {clviii}those which we may with probability
expect from the tropical eastern Atlantic, will be sure to increase very
widely our knowledge of the class.

238. _Globigerina Ooze._--Next to the Radiolarian ooze proper the
Globigerina ooze is the deposit which is richest in the remains of
Radiolaria. Often these are so abundant that it is doubtful to which
category the specimen should be referred (_e.g._, Stations 270 and 271, see
§ 237). In fact, the two pass without any sharp boundary into each other,
and both present transitions to the Diatom ooze. Next to red clay (§ 239),
Globigerina ooze is the most widely distributed of all sediments, and forms
a large part of the bed of the ocean at depths of 250 to 2900 fathoms
(especially between 1000 and 2000 fathoms). It covers extensive areas at
depths below 1800 fathoms, and in still deeper water is replaced by red
clay. It is a fine-grained white, grey, or yellowish powder, which
sometimes becomes coloured rose, red, or brown owing to the admixture of
oxides of iron and manganese. True Globigerina ooze consists for the most
part of the accumulated calcareous shells of pelagic Foraminifera,
principally _Globigerina_ and _Orbulina_, but also _Hastigerina_,
_Pulvinulina_, &c. It contains usually from 50 to 80 per cent. of calcium
carbonate, the extreme values being 40 and 95 per cent. After this has been
removed by acids, there remains a residue, which consists partly of the
siliceous shells of Radiolaria and Diatoms, and partly of mineral particles
identical with the volcanic elements of the red clay.

Regarding the composition and significance of the Globigerina ooze, see
John Murray (L. N. 27, pp. 523-525, and L. N. 53, vol. i. p. 919).
Recently this author has separated from the Globigerina ooze (_sensu
stricto_), the _Pteropod ooze_, distinguished from the former by the
greater abundance of Pteropod shells and calcareous shells of larger
pelagic organisms which it contains. It is found in moderate depths (at
most 1500 fathoms), and contains fewer Radiolaria.

239. _Red Clay._--This is quantitatively the most important of all deep-sea
deposits, covering by far the greatest extent of the three great ocean
basins at depths greater than 2200 fathoms. It thus far surpasses in area
the other deposits, both Radiolaria and Globigerina oozes, and commonly
forms a still deeper layer beneath them. Probably these three deep-sea
deposits together cover about three-eighths of the whole surface of the
earth, that is, about as much as all the continents together, whilst only
two-eighths are covered by the terrigenous deposits. Red clay is
principally composed of silicate of alumina, mixed in various proportions
with other finely granular substances; its usual red colour, which
sometimes passes over into grey or brown, is more especially due to
admixture of oxides of iron and manganese. Calcareous matter is usually
entirely wanting, or present only in traces, whilst free silica is found in
very variable, often considerable quantities. The chief mass of the red
clay consists of volcanic ashes, pumice, fragments of lava, &c., whilst a
large part of it is generally composed of shells of Radiolaria or fragments
of {clix}them; in many places the number of well-preserved skeletons
contained in the red clay is very considerable, so that it passes over
gradually into the Radiolarian ooze (_e.g._, in the North Pacific, Stations
238 to 253, see § 237). Hence it may be supposed that a large part of the
red clay consists of decomposed Radiolarian ooze.

The characteristic composition and fundamental significance of the red
clay in the formation of the deep-sea bed were first made known by the
discoveries of the Challenger (compare John Murray, 1876, L. N. 27, p.
527, and Narr. Chall. Exp., L. N. 53, vol. i. pt. ii. pp. 920-926, pl. N;
also Wyville Thomson, The Atlantic, L. N. 31, vol. i. pp. 226-229). The
mineral components of the red clay are for the most part of volcanic
origin, due to the decomposition of pumice, lava, &c. Among the organic
remains found in it, the siliceous skeletons of Radiolaria are by far the
most important, and their number is often considerable. A large portion
of the red clay appears to me to consist of broken down Radiolarian
shells, in which a peculiar metamorphism probably has taken place. Sir
Wyville Thomson was of opinion that a considerable proportion of it
consisted of the remains of Globigerina ooze, the calcareous constituents
of which had been removed by the carbon dioxide in the deep-sea water (L.
N. 31, _loc. cit._). Among these remains, however, the siliceous
skeletons of the Radiolaria play a significant and often the most
important part. Furthermore, John Murray has called attention to the fact
that in many deep-sea deposits yellow and red insoluble particles remain,
which unmistakably present the form of Radiolarian shells (L. N. 27, p.
513). At Station 303 he found "amorphous clayey matter, rounded yellow
minerals, many Radiolaria-shaped;" at Station 302 there was sediment
"consisting almost entirely of small rounded red mineral particles; many
of these had the form of both Foraminifera and Radiolaria; and it seemed
as if some substance had been deposited in and on these organisms."
Similar transitions from well-preserved Radiolarian shells into amorphous
mineral particles I have found in several other specimens of Challenger
soundings, and consider them a further argument for the supposition that
the Radiolaria often take an important share in the formation of the red
clay.

240. _List of Stations at which Radiolaria were observed on the Challenger
Expedition._--The 168 Stations recorded below, in soundings or surface
preparations from which I found Radiolaria, belong to the most various
parts of the sea which the Challenger traversed during her voyage round the
world; they constitute about half of the (364) observing Stations contained
in the official list published in the Narrative of the Cruise (Narr. Chall.
Exp., vol. i. part ii. Appendix ii.).

In addition to the particulars given in the list regarding the
geographical position of the Station, depth, temperature, and composition
of the bottom deposit, I have added the result of my investigations as
regards the relative abundance of the Radiolaria in each. The five
letters (A to E) denote the following degrees of frequency:--A, abundant
Radiolaria (AI, pure Radiolarian ooze; AII, mixed Radiolarian ooze); B,
very numerous Radiolaria (but not a predominating quantity); C, many
Radiolaria (medium quantity); D, few Radiolaria; E, very few Radiolaria
(as they occur almost always). In using these symbols regard has been had
to abundance of the abyssal as well as of the zonarial and pelagic forms
(§ 232); sometimes also the estimated number of Radiolaria has been
inserted, based upon information given by John Murray in his Preliminary
Report (L. N. 27), and in the Narrative of the Cruise (L. N. 53), as well
as by Henry B. Brady in his Report on the {clx}Foraminifera (Zool. Chall.
Exp., part xxii., 1884). From Stations 348 to 352 in the Eastern Tropical
Atlantic no specimens of the bottom were obtained, but a rich pelagic
Radiolarian fauna was demonstrated by numerous preparations from the
surface. The depths are given in fathoms and the temperature in degrees
Fahrenheit. In the column describing the nature of the bottom the
following abbreviations are used:--

rad. oz. = Radiolarian ooze (§ 237).
gl. oz. = Globigerina ooze (§ 238).
r. cl. = red clay (§ 239).
pt. oz. = Pteropod ooze (see p. clviii).
di. oz. = Diatom ooze (see p. clvii).
bl. m. = blue mud, } terrigenous deposits
gr. m. = green mud, } (see p. clvi).
volc. m. = volcanic mud, }
r. m. = red mud.

+----------+-----------+--------+------------+-----------+--------------+
| | | | Bottom | | Relative |
|Challenger| Locality. |Depth in|Temperature,| Nature of | Abundance of |
| Station. | |Fathoms.| ° F. | Bottom. | Radiolaria. |
+----------+-----------+--------+------------+-----------+--------------+
| | | | | | |
| 1. | N. Atl. | 1890 | 36.8 | gl. oz. | D few |
| 2. | " | 1945 | 36.8 | gl. oz. | E very few |
| 5. | " | 2740 | 37.0 | r. cl. | D few |
| 9. | " | 3150 | 36.8 | r. cl. | E very few |
| 24. | Tr. Atl. | 390 | ... | pt. oz. | D few |
| | | | | | |
| 32. | N. Atl. | 2250 | 36.7 | gl. oz. | E very few |
| 45. | " | 1240 | 37.2 | bl. m. | E " |
| 50. | " | 1250 | 38.0 | bl. m. | E " |
| 64. | " | 2700 | ... | r. cl. | D few |
| 76. | " | 900 | 40.0 | pt. oz. | D " |
| | | | | | |
| 98. | Tr. Atl. | 1750 | 36.7 | gl. oz. | C many |
| 106. | " | 1850 | 36.6 | gl. oz. | C " |
| 108. | " | 1900 | 36.8 | gl. oz. | C " |
| 111. | " | 2475 | 33.7 | gl. oz. | C " |
| 120. | " | 675 | ... | r. m. | D few |
| | | | | | |
| 132. | S. Atl. | 2050 | 35.0 | gl. oz. | C many |
| 134. | " | 2025 | 36.0 | gl. oz. | C " |
| 137. | " | 2550 | 34.5 | r. cl. | D few |
| 138. | " | 2650 | 35.1 | r. cl. | D " |
| 143. | S. Ind. | 1900 | 35.6 | gl. oz. | E very few |
| | | | | | |
| 144. | " | 1570 | 35.8 | gl. oz. | E " |
| 145. | " | 140 | ... | volc. s. | D few |
| 146. | " | 1375 | 35.6 | gl. oz. | C many |
| 147. | " | 1600 | 34.2 | di. oz. | C " |
| 148. | " | 210 | ... {| gravel, } | D few |
| | | | {| shells } | |
| | | | | | |
| 149H. | " | 127 | ... | volc. m. | D " |
| 150. | " | 150 | 35.2 | gravel | D " |
| 151. | " | 75 | ... | volc. m. | D " |
| 152. | " | 1260 | ... | di. oz. | C many |
| 153. | " | 1675 | ... | bl. m. | C " |
| | | | | | |
| 154. | " | 1800 | ... | bl. m. | C " |
| 155. | " | 1300 | ... | bl. m. | C " |
| 156. | " | 1975 | ... | di. oz. | B numerous |
| 157. | " | 1950 | 32.1 | di. oz. | B " |
| 158. | " | 1800 | 33.5 | gl. oz. | B " |
| | | | | | |
| 159. | " | 2150 | 34.5 | gl. oz. | B " |
| 160. | " | 2600 | 33.9 | r. cl. | C many |
| 162. | " | 38 | ... | sand | E very few |
| 163. | S. Pac. | 2200 | 34.5 | gr. m. | E " |
| 164A. | " | 1200 | ... | gr. m. | E " |
| | | | | | |
| 165. | S. Pac. | 2600 | 34.5 | r. cl. | D few |
| 166. | " | 275 | 50.8 | gl. oz. | D " |
| 169. | " | 700 | 40.0 | bl. m. | D " |
| 175. | Tr. Pac. | 1350 | 36.0 | gl. oz. | E very few |
| 181. | " | 2440 | 35.8 | r. cl. | E " |
| | | | | | |
| 193. | " | 2800 | 38.0 | bl. m. | D few |
| 195. | " | 1425 | 38.0 | bl. m. | C many |
| 197. | " | 1200 | 35.9 | bl. m. | D few |
| 198. | " | 2150 | 38.9 | bl. m. | C many |
| 200. | " | 250 | ... | gr. m. | B numerous |
| | | | | | |
| 201. | " | 82 | ... | st. & gra.| C many |
| 202. | " | 2550 | 50.5 | bl. m. | B numerous |
| 205. | " | 1050 | 37.0 | bl. m. | C many |
| 206. | " | 2100 | 36.5 | bl. m. | B numerous |
| 211. | " | 2225 | 50.5 | bl. m. | B " |
| | | | | | |
| 213. | " | 2050 | 38.8 | bl. m. | C many |
| 214. | " | 500 | 41.8 | bl. m. | C " |
| 215. | " | 2550 | 35.4 | r. cl. | C many |
| 216A | " | 2000 | 35.4 | gl. oz. | B numerous |
| 217. | " | 2000 | 35.2 | bl. m. | C many |
| | | | | | |
| 218. | " | 1070 | 36.4 | bl. m. | C " |
| 220. | " | 1100 | 36.2 | gl. oz. | C " |
| 221. | " | 2650 | 35.4 | r. cl. | B numerous |
| 222. | " | 2450 | 35.2 | r. cl. | B " |
| 223. | " | 2325 | 35.5 | gl. oz. | B " |
| | | | | | |
| 224. | " | 1850 | 35.4 | gl. oz. | B " |
| 225. | " | 4475 | 35.2 | rad. oz. | A very many |
| 226. | " | 2300 | 35.5 | rad. oz. | A " |
| 230. | N. Pac. | 2425 | 35.5 | r. cl. | C many |
| 231. | " | 2250 | 35.2 | bl. m. | C " |
| | | | | | |
| 232. | " | 345 | 41.1 | gr. m. | C " |
| 234. | " | 2675 | 35.8 | bl. m. | B numerous |
| 235. | " | 565 | 38.1 | gr. m. | D few |
| 236. | " | 775 | 37.6 | gr. m. | C many |
| 237. | " | 1875 | 35.3 | bl. m. | C " |
| | | | | | |
| 238. | " | 3950 | 35.0 | r. cl. | B numerous |
| 239. | " | 3625 | 35.1 | r. cl. | B " |
| 240. | " | 2900 | 34.9 | r. cl. | B " |
| 241. | " | 2300 | 35.1 | r. cl. | A very many |
| 242. | " | 2575 | 35.1 | r. cl. | AII " |
| | | | | | |
| 243. | " | 2800 | 35.0 | r. cl. | AII " |
| 244. | " | 2900 | 35.3 | r. cl. | AII " |
| 245. | " | 2775 | 34.9 | r. cl. | AII " |
| 246. | " | 2050 | 35.1 | gl. oz. | B numerous |
| 247. | " | 2530 | 35.2 | r. cl. | C many |
| | | | | | |
| 248. | " | 2900 | 35.1 | r. cl. | C " |
| 249. | " | 3000 | 35.2 | r. cl. | B numerous |
| 250. | " | 3050 | 35.0 | r. cl. | B " |
| 251. | " | 2950 | 35.1 | r. cl. | B " |
| 252. | " | 2740 | 35.3 | r. cl. | B " |
| | | | | | |
| 253. | " | 3125 | 35.1 | r. cl. | B " |
| 254. | " | 3025 | 35.0 | r. cl. | C many |
| 255. | " | 2850 | 35.0 | r. cl. | C " |
| 256. | " | 2950 | 35.2 | r. cl. | B numerous |
| 257. | " | 2875 | 34.9 | r. cl. | C many |
| | | | | | |
| 258. | " | 2775 | 35.2 | r. cl. | C " |
| 259. | Tr. Pac. | 2225 | 34.9 | r. cl. | C " |
| 261. | " | 2050 | 35.2 | volc. m. | C many |
| 262. | " | 2875 | 35.2 | r. cl. | C " |
| 263. | " | 2650 | 35.1 | r. cl. | B numerous |
| | | | | | |
| 264. | " | 3000 | 35.2 | r. cl. | C many |
| 265. | " | 2900 | 35.0 | r. cl. | A very many |
| 266. | " | 2750 | 35.1 | rad. oz. | A " |
| 267. | " | 2700 | 35.0 | rad. oz. | A " |
| 268. | " | 2900 | 34.8 | rad. oz. | A " |
| | | | | | |
| 269. | " | 2550 | 35.2 | rad. oz. | A " |
| 270. | " | 2925 | 34.6 | gl. oz. | A " |
| 271. | " | 2425 | 35.0 | gl. oz. | A " |
| 272. | " | 2600 | 35.1 | rad. oz. | A " |
| 273. | " | 2350 | 34.5 | rad. oz. | A " |
| | | | | | |
| 274. | " | 2750 | 35.1 | rad. oz. | A " |
| 275. | " | 2610 | 35.0 | r. cl. | B numerous |
| 276. | " | 2350 | 35.1 | r. cl. | C many |
| 280. | " | 1940 | 35.3 | gl. oz. | D few |
| 281. | " | 2385 | 34.9 | r. cl. | C many |
| | | | | | |
| 282. | S. Pac. | 2450 | 35.1 | r. cl. | C " |
| 283. | " | 2075 | 35.4 | gl. oz. | D few |
| 284. | " | 1985 | 35.1 | gl. oz. | C many |
| 285. | " | 2375 | 35.0 | r. cl. | D few |
| 286. | " | 2335 | 34.8 | r. cl. | D " |
| | | | | | |
| 287. | " | 2400 | 34.7 | r. cl. | D " |
| 288. | " | 2600 | 34.8 | r. cl. | B numerous |
| 289. | " | 2550 | 34.8 | r. cl. | B " |
| 290. | " | 2300 | 34.9 | r. cl. | C many |
| 291. | " | 2250 | 34.6 | r. cl. | C " |
| | | | | | |
| 292. | " | 1600 | 35.2 | gl. oz. | C " |
| 293. | " | 2025 | 34.4 | gl. oz. | C " |
| 294. | " | 2270 | 34.6 | r. cl. | D few |
| 295. | " | 1500 | 35.3 | gl. oz. | C many |
| 296. | " | 1825 | 35.3 | gl. oz. | D few |
| | | | | | |
| 297. | " | 1775 | 35.5 | gl. oz. | D " |
| 298. | " | 2225 | 35.6 | bl. m. | C many |
| 299. | " | 2160 | 35.2 | bl. m. | C " |
| 300. | " | 1375 | 35.5 | gl. oz. | B numerous |
| 302. | " | 1450 | 35.6 | gl. oz. | B " |
| | | | | | |
| 303. | " | 1325 | 36.0 | bl. m. | D few |
| 304. | " | 45 | ... | gr. m. | E very few |
| 318. | S. Atl. | 2040 | 33.7 | bl. m. | C few |
| 319. | " | 2425 | 32.7 | bl. m. | C " |
| 323. | " | 1900 | 33.1 | bl. m. | C " |
| | | | | | |
| 324. | " | 2800 | 32.6 | bl. m. | B numerous |
| 325. | " | 2650 | 32.7 | bl. m. | B " |
| 326. | " | 2775 | 32.7 | bl. m. | C many |
| 327. | " | 2900 | 32.8 | bl. m. | C " |
| 328. | " | 2900 | 32.9 | bl. m. | B numerous |
| | | | | | |
| 329. | " | 2675 | 32.3 | r. cl. | C many |
| 330. | " | 2440 | 32.7 | r. cl. | C " |
| 331. | " | 1715 | 35.4 | gl. oz. | B numerous |
| 332. | " | 2200 | 34.0 | gl. oz. | B " |
| 333. | " | 2025 | 35.3 | gl. oz. | B " |
| | | | | | |
| 334. | " | 1915 | 35.8 | gl. oz. | C many |
| 335. | " | 1425 | 37.0 | pt. oz. | D few |
| 338. | Tr. Atl. | 1990 | 36.3 | gl. oz. | D few |
| 340. | " | 1500 | 37.6 | pt. oz. | E very few |
| 341. | " | 1475 | 38.2 | pt. oz. | E " |
| | | | | | |
| 342. | " | 1445 | 37.5 | pt. oz. | D few |
| 343. | " | 425 | 40.3 | volc. s. | E very few |
| 344. | " | 420 | ... | volc. s. | E " |
| 345. | " | 2010 | 36.8 | gl. oz. | D few |
| 346. | " | 2350 | 34.0 | gl. oz. | C many |
| | | | | | |
| 347. | " | 2250 | 36.2 | gl. oz. | B numerous |
| 348. | " | (2450) | ... | (Pelag.) | B " |
| 349. | " | ... | ... | (Pelag.) | B " |
| 350. | " | ... | ... | (Pelag.) | B " |
| 351. | " | ... | ... | (Pelag.) | B " |
| | | | | | |
| 352. | " | ... | ... | (Pelag.) | B " |
| 353. | N. Atl. | 2965 | 37.6 | r. cl. | C many |
| 354. | " | 1675 | 37.8 | gl. oz. | D few |
+----------+-----------+--------+------------+-----------+--------------+

+----------+--------+-----------------------+---------------------------+
| | | | |
|Challenger| Date. |Latitude and Longitude.| Nearest Land. |
| Station. | | | |
+----------+--------+-----------------------+---------------------------+
| | 1873. | | |
| 1. |Feb. 15|27° 24' N., 16° 55' W.|S. of Tenerife. |
| 2. | " 17|25° 52' N., 19° 22' W.|S.W. of the Canary Islands.|
| 5. | " 21|24° 20' N., 24° 28' W.|S.W. of the Canary Islands.|
| 9. | " 26|23° 23' N., 35° 11' W.| (Ocean). |
| 24. |Mar. 25|18° 38' N., 65° 5' W.|Culebra (Antilles). |
| | | | |
| 32. |April 3|31° 49' N., 64° 55' W.|Bermuda. |
| 45. |May 3|38° 34' N., 72° 10' W.|S. of New York. |
| 50. | " 21|42° 8' N., 63° 39' W.|S. of Halifax. |
| 64. |June 20|35° 35' N., 50° 27' W.| (Ocean). |
| 76. |July 3|38° 11' N., 27° 9' W.|Azores. |
| | | | |
| 98. |Aug. 14| 9° 21' N., 18° 28' W.|W. of Sierra Leone. |
| 106. | " 25| 1° 47' N., 24° 26' W.| (Ocean). |
| 108. | " 27| 1° 10' N., 28° 23' W.| (Ocean). |
| 111. | " 31| 1° 45' S., 30° 58' W.| (Ocean). |
| 120. |Sept. 9| 8° 37' S., 34° 28' W.|Pernambuco. |
| | | | |
| 132. |Oct. 10|35° 25' S., 23° 40' W.|Tristan da Cunha. |
| 134. | " 14|36° 12' S., 12° 16' W.|Tristan da Cunha. |
| 137. | " 23|35° 59' S., 1° 34' E.| (Ocean). |
| 138. | " 25|36° 22' S., 8° 12' E.| (Ocean). |
| 143. |Dec. 19|36° 48' S., 19° 24' E.|Cape of Good Hope. |
| | | | |
| 144. | " 24|45° 57' S., 34° 39' E.| (Ocean). |
| 145. | " 27|46° 43' S., 38° 4' E.|Prince Edward Island. |
| 146. | " 29|46° 46' S., 45° 31' E.| (Ocean). |
| 147. | " 30|46° 16' S., 48° 27' E.|W. of the Crozet Islands. |
| | 1874. | | |
| 148. |Jan. 3|46° 47' S., 51° 37' E.|E. of the Crozet Islands. |
| | | | |
| 149H. | " 29|48° 45' S., 69° 14' E.|Kerguelen Island. |
| 150. |Feb. 2|52° 4' S., 71° 22' E.|N. of Heard Island. |
| 151. | " 7|52° 59' S., 73° 33' E.|Heard Island. |
| 152. | " 11|60° 52' S., 80° 20' E.|(Ocean). |
| 153. | " 14|65° 42' S., 79° 49' E.|Antarctic Ice. |
| | | | |
| 154. | " 19|64° 37' S., 85° 49' E.|Antarctic Ice. |
| 155. | " 23|64° 18' S., 94° 47' E.|Antarctic Ice. |
| 156. | " 26|62° 26' S., 95° 44' E.| (Ocean). |
| 157. |Mar. 3|53° 55' S., 108° 35' E.| (Ocean). |
| 158. | " 7|50° 1' S., 123° 4' E.| (Ocean). |
| | | | |
| 159. | " 10|47° 25' S., 130° 22' E.| (Ocean). |
| 160. | " 13|42° 42' S., 134° 10' E.| (Ocean). |
| 162. |April 2|39° 10' S., 146° 37' E.|Bass Strait. |
| 163. | " 4|36° 57' S., 150° 34' E.|Port Jackson. |
| 164A. |June 13|34° 9' S., 151° 55' E.|W. of Sydney. |
| | | | |
| 165. | " 17|34° 50' S., 155° 28' E.| (Ocean). |
| 166. | " 23|38° 50' S., 169° 20' E.|W. of New Zealand. |
| 169. |July 10|37° 34' S., 179° 22' E.|E. of New Zealand. |
| 175. |Aug. 12|19° 2' S., 177° 10' E.|Fiji Islands. |
| 181. | " 25|13° 50' S., 151° 49' E.|Louisiades. |
| | | | |
| 193. |Sept. 28| 5° 24' S., 130° 37' E.|Banda Sea. |
| 195. |Oct. 3| 4° 21' S., 129° 7' E.|Banda Sea. |
| 197. | " 14| 0° 41' N., 126° 37' E.|E. of Celebes. |
| 198. | " 20| 2° 55' N., 124° 58' E.|N. of Celebes. |
| 200. | " 23| 6° 47' N., 122° 28' E.|W. of Mindanao. |
| | | | |
| 201. | " 26| 7° 3' N., 121° 48' E.|W. of Mindanao. |
| 202. | " 27| 8° 32' N., 121° 55' E.|W. of Mindanao. |
| 205. |Nov. 13|16° 42' N., 119° 22' E.|W. of Luzon. |
| | 1875. | | |
| 206. |Jan. 8|17° 54' N., 117° 14' E.|W. of Luzon. |
| 211. | " 28| 8° 0' N., 121° 42' E.|W. of Mindanao. |
| | | | |
| 213. |Feb. 8| 5° 47' N., 124° 1' E.|S. of Mindanao. |
| 214. | " 10| 4° 33' N., 127° 6' E.|N. of Gilolo. |
| 215. | " 12| 4° 19' N., 130° 15' E.|N. of Gilolo. |
| 216A. | " 16| 2° 56' N., 134° 11' E.|S. of Pelew Islands. |
| 217. | " 22| 0° 39' S., 138° 55' E.|N. of New Guinea. |
| | | | |
| 218. |Mar. 1| 2° 33' S., 144° 4' E.|N. of New Guinea. |
| 220. | " 11| 0° 42' S., 147° 0' E.|N. of New Guinea. |
| 221. | " 13| 0° 40' N., 148° 41' E.| (Ocean). |
| 222. | " 16| 2° 15' N., 146° 16' E.| (Ocean). |
| 223. | " 19| 5° 31' N., 145° 13' E.|Carolines. |
| | | | |
| 224. | " 21| 7° 45' N., 144° 20' E.|Carolines. |
| 225. | " 23|11° 24' N., 143° 16' E.|Ocean } |
| 226. | " 25|14° 44' N., 142° 13' E.|Ocean } North-West Pacific,|
| 230. |April 5|26° 29' N., 137° 57' E.|Ocean } between Carolines |
| 231. | " 9|31° 8' N., 137° 8' E.|Ocean } and Japan. |
| | | | } |
| 232. |May 12|35° 11' N., 139° 28' E.|Ocean } |
| 234. |June 3|32° 31' N., 135° 39' E.|S. of Japan. |
| 235. | " 4|34° 7' N., 138° 0' E.|S. of Japan. |
| 236. | " 5|34° 58' N., 139° 29' E.|S. of Japan. |
| 237. | " 17|34° 37' N., 140° 32' E.|S. of Japan. |
| | | | |
| 238. | " 18|35° 18' N., 144° 8' E.|Ocean } |
| 239. | " 19|35° 18' N., 147° 9' E.|Ocean } |
| 240. | " 21|35° 20' N., 153° 39' E.|Ocean } |
| 241. | " 23|35° 41' N., 157° 42' E.|Ocean } |
| 242. | " 24|35° 29' N., 161° 52' E.|Ocean } |
| | | | } |
| 243. | " 26|35° 24' N., 166° 35' E.|Ocean } North Pacific, |
| 244. | " 28|35° 22' N., 169° 53' E.|Ocean } between Japan and |
| 245. | " 30|36° 23' N., 174° 31' E.|Ocean } San Francisco |
| 246. |July 2|36° 10' N., 178° 0' E.|Ocean } (35°-38° N. lat., |
| 247. | " 3|35° 49' N., 179° 57' W.|Ocean } 144°-156° W. long.)|
| | | | } |
| 248. | " 5|37° 41' N., 177° 4' W.|Ocean } |
| 249. | " 7|37° 59' N., 171° 48' W.|Ocean } |
| 250. | " 9|37° 49' N., 166° 47' W.|Ocean } |
| 251. | " 10|37° 37' N., 163° 26' W.|Ocean } |
| 252. | " 12|37° 52' N., 160° 17' W.|Ocean } |
| | | | } |
| 253. | " 14|38° 9' N., 156° 25' W.|Ocean |
| 254. | " 17|35° 13' N., 154° 43' W.|Ocean } |
| 255. | " 19|32° 28' N., 154° 33' W.|Ocean } North Pacific |
| 256. | " 21|30° 22' N., 154° 56' W.|Ocean } (35°-23° N. lat., |
| 257. | " 23|27° 33' N., 154° 55' W.|Ocean }154°-156° W. long.).|
| | | | } |
| 258. | " 24|26° 11' N., 155° 12' W.|Ocean } |
| 259. | " 26|23° 3' N., 156° 6' W.|Ocean } |
| 261. |Aug. 12|20° 18' N., 157° 14' W.|Sandwich Islands. |
| 262. | " 20|19° 12' N., 154° 14' W.|Sandwich Islands. |
| 263. | " 21|17° 33' N., 153° 36' W.|Ocean } |
| | | | } |
| 264. | " 23|14° 19' N., 152° 37' W.|Ocean } |
| 265. | " 25|12° 42' N., 152° 1' W.|Ocean } |
| 266. | " 26|11° 7' N., 152° 3' W.|Ocean } Tropical Central |
| 267. | " 28| 9° 28' N., 150° 49' W.|Ocean } Pacific, between |
| 268. | " 30| 7° 35' N., 149° 49' W.|Ocean } Sandwich and |
| | | | } |
| 269. |Sept. 2| 5° 54' N., 147° 2' W.|Ocean } Paumotu |
| 270. | " 4| 2° 34' N., 149° 9' W.|Ocean } (17° N. lat. to |
| 271. | " 6| 0° 33' S., 151° 34' W.|Ocean } 11° S. lat.). |
| 272. | " 8| 3° 48' S., 152° 56' W.|Ocean } |
| 273. | " 9| 5° 11' S., 152° 56' W.|Ocean } |
| | | | } |
| 274. | " 11| 7° 25' S., 152° 15' W.|Ocean } |
| 275. | " 14|11° 20' S., 150° 30' W.|Ocean } |
| 276. | " 16|13° 28' S., 149° 30' W.|Paumotu. |
| 280. |Oct. 4|18° 40' S., 149° 52' W.|S. of Tahiti. |
| 281. | " 6|22° 21' S., 150° 17' W.|Tubuai Islands. |
| | | | |
| 282. | " 7|23° 46' S., 149° 59' W.|Tubuai Islands. |
| 283. | " 9|26° 9' S., 145° 17' W.|N. of Oparo Island. |
| 284. | " 11|28° 22' S., 141° 22' W.|S. of Oparo Island. |
| 285. | " 14|32° 36' S., 137° 43' W.|Ocean } |
| 286. | " 16|33° 29' S., 133° 22' W.|Ocean } |
| | | | |
| 287. | " 10|36° 32' S., 132° 52' W.|Ocean } |
| 288. | " 21|40° 3' S., 132° 58' W.|Ocean } |
| 289. | " 23|39° 41' S., 131° 23' W.|Ocean } |
| 290. | " 25|39° 16' S., 124° 7' W.|Ocean } Open South Pacific |
| 291. | " 27|39° 13' S., 118° 49' W.|Ocean } Ocean, between New |
| | | | } Zealand and |
| 292. | " 29|38° 43' S., 112° 31' W.|Ocean } Valparaiso. |
| 293. |Nov. 1|39° 4' S., 105° 5' W.|Ocean } |
| 294. | " 3|39° 22' S., 98° 46' W.|Ocean } |
| 295. | " 5|38° 7' S., 94° 4' W.|Ocean } |
| 296. | " 9|38° 6' S., 88° 2' W.|Ocean } |
| | | | } |
| 297. | " 11|37° 29' S., 83° 7' W.|Ocean } |
| 298. | " 17|34° 7' S., 73° 56' W.|W. of Valparaiso. |
| 299. |Dec. 14|33° 31' S., 74° 43' W.|W. of Valparaiso. |
| 300. | " 17|33° 42' S., 78° 18' W.|N. of Juan Fernandez. |
| 302. | " 28|42° 43' S., 82° 11' W.| (Ocean). |
| | | | |
| 303. | " 30|45° 31' S., 78° 9' W.|W. of Patagonia. |
| 304. | " 31|46° 53' S., 75° 12' W.|W. of Patagonia. |
| | 1876. | | |
| 318. |Feb. 11|42° 32' S., 56° 29' W.| (Ocean). |
| 319. | " 12|41° 54' S., 54° 48' W.| (Ocean). |
| 323. | " 28|35° 39' S., 50° 47' W.|W. of Buenos Ayres. |
| | | | } |
| 324. | " 29|36° 9' S., 48° 22' W.|Ocean } |
| 325. |Mar. 2|36° 44' S., 46° 16' W.|Ocean } |
| 326. | " 3|37° 3' S., 44° 17' W.|Ocean } Open South Atlantic|
| 327. | " 4|36° 48' S., 42° 45' W.|Ocean } Ocean, between |
| 328. | " 6|37° 38' S., 39° 36' W.|Ocean } Buenos Ayres and |
| | | | } |
| 329. | " 7|37° 31' S., 36° 7' W.|Ocean } Tristan da Cunha |
| 330. | " 8|37° 45' S., 33° 0' W.|Ocean } (35°-37° S. lat., |
| 331. | " 9|37° 47' S., 30° 20' W.|Ocean } 21°-48° W. long.). |
| 332. | " 10|37° 29' S., 27° 31' W.|Ocean } |
| 333. | " 13|35° 36' S., 21° 12' W.|Ocean } |
| | | | |
| 334. | " 14|35° 45' S., 18° 31' W.|W. of Tristan da Cunha. |
| 335. | " 16|32° 24' S., 13° 5' W.|N. of Tristan da Cunha. |
| 338. | " 21|21° 15' S., 14° 2' W.|(Ocean). |
| 340. | " 24|14° 33' S., 13° 42' W.|Ocean } |
| 341. | " 25|12° 16' S., 13° 44' W.|Ocean } W. of St. Helena. |
| | | | |
| 342. | " 26| 9° 43' S., 13° 51' W.|Ocean } |
| 343. | " 27| 8° 3' S., 14° 27' W.|Ascension Island. |
| 344. |April 3| 7° 54' S., 14° 28' W.|Ascension Island. |
| 345. | " 4| 5° 45' S., 14° 25' W.|Ocean } |
| 346. | " 6| 2° 42' S., 14° 41' W.|Ocean } Tropical Atlantic, |
| | | | } between Ascension |
| 347. | " 7| 0° 15' S., 14° 25' W.|Ocean } and Sierra Leone. |
| 348. | " 9| 3° 10' N., 14° 51' W.|Ocean } |
| 349. | " 10| 5° 28' N., 14° 38' W.|Ocean } |
| 350. | " 11| 7° 33' N., 15° 16' W.|W. of Sierra Leone. |
| 351. | " 12| 9° 9' N., 16° 41' W.|W. of Sierra Leone. |
| | | | |
| 352. | " 13|10° 55' N., 17° 46' W.|W. of Sierra Leone. |
| 353. |May 3|26° 21' N., 33° 37' W.|W. of Canary Islands. |
| 354. | " 6|32° 41' N., 36° 6' W.|S. of Azores. |
+----------+--------+-----------------------+---------------------------+

{clxiv}CHAPTER X.--GEOGRAPHICAL DISTRIBUTION.

(§§ 241-250.)

241. _Historical Distribution._--Radiolaria are found fossil in all the
more important groups of the sedimentary rocks of the earth's crust. Whilst
a few years ago their well-preserved siliceous skeletons were only known in
considerable quantity from Cainozoic marls (§ 242), very many SPUMELLARIA
and NASSELLARIA have recently been found in Mesozoic and a few in Palæozoic
strata. By the aid of improved modern methods of investigation (especially
by the preparation of thin sections of very hard rocks) it has been shown
that many hard siliceous minerals, especially cryptocrystalline quartz,
contain numerous well-preserved Radiolaria, and sometimes are mainly
composed of closely compacted masses of such siliceous shells; of this kind
are many quartzites of the Jura (§ 243). These Jurassic quartzes
(Switzerland), as well as the Tertiary marls (Barbados) and clays (Nicobar
Islands), are to be regarded as "fossil Radiolarian ooze" (§ 237). Dense
masses of compressed SPUMELLARIA and NASSELLARIA form the principal part of
these rocks. Isolated or in smaller quantities, fossil Polycystina,
belonging to different families of SPUMELLARIA and NASSELLARIA, also occur
in in other rocks, and even in some of Palæozoic origin. Since specimens
have also been recently found both in Silurian and Cambrian strata, it may
be stated that as regards their historical distribution, Radiolaria occur
in all fossiliferous sedimentary deposits, from the oldest to those of the
present time.

242. _Cainozoic Radiolaria._--The great majority of fossil Radiolaria which
have hitherto been described, belong to the Cainozoic or Tertiary period,
and in fact, to its middle portion, the Miocene period. At this period the
richest and most important of all the Radiolarian formations were
deposited, such as the pure "Polycystine marl" of Barbados (see note A),
also that of Grotte in Sicily (see note B), and the clay of the Nicobar
Islands (see note C). Besides the above-mentioned deposits, which may be
designated "pure" fossil Radiolarian ooze, many deposits containing these
organisms have recently been discovered in widely separated parts of the
earth, partly of the nature of tripoli or marl, partly resembling clay.
Among these may be mentioned in the first place many coasts and islands of
the Mediterranean, both on the south coast of Europe (Sicily, Calabria,
Greece), and the north coast of Africa (from Oran to Tripoli). The
extensive layers of tripoli which are found in these Mediterranean Tertiary
mountains belong to the upper Miocene (Tortona stage), and consist partly
of marl rich in calcareous matter, and resembling chalk, partly passing
over into plastic clay or "Kieselguhr" (§ 246). The quantity of Radiolaria
contained varies, and is more conspicuous the fewer the calcareous shells
of Foraminifera present. Similar Tertiary Polycystine formations occur in
some parts of America (see note D); probably they have a very wide
distribution. In their general morphological characters, the Tertiary
SPUMELLARIA and NASSELLARIA {clxv}are related to those forms which are
found in the recent Radiolarian ooze of the depths of the Pacific,
especially to the species which are characteristic of the Challenger
Stations 225, 226, 265 and 268. Many living genera and families (_e.g._,
most #Larcoidea# and #Stephoidea#) have not yet been found in the Tertiary
formations.

A. The famous Polycystine marl of Barbados in the Antilles, which Robert
Schomburgk discovered forty years ago, belongs to the Miocene formation,
and is the richest and best known of all the important Radiolarian
deposits (see L. N. 16, pp. 5-8). After Ehrenberg had published in
December 1846 the first preliminary communication regarding its
composition out of masses of well-preserved Polycystina, he was able in
the following year to describe no less than 282 species from it; he
distributed these in 44 genera and 7 families (L. N. 4, 1847, p. 54). In
the year 1854 Ehrenberg published figures of 33 species in his
Mikrogeologie (L. N. 6, Taf. xxxvi.); but it was only in 1873 that he
published descriptions of 265 species (Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, Jan. 30, pp. 213-263). Finally there followed in 1875 his
Fortsetzung der Mikrogeologischen Studien, mit specieller Rücksicht auf
den Polycystinen-Mergel von Barbados (L. N. 25). On the thirty plates
which accompany this the last work of Ehrenberg, 282 species are figured
and named, of which 54 are SPUMELLARIA (13 #Sphæroidea#, 8 #Prunoidea#,
33 #Discoidea#), and 228 NASSELLARIA (2 #Stephoidea#, 38 #Spyroidea#, and
188 #Cyrtoidea#). The fourth section of this memoir contains a survey of
the Polycystine formation of Barbados (pp. 106-115), and the fifth
section the special description of a large specimen of rock from Mount
Hillaby in Barbados (see also L. N. 28, p. 117, and L. N. 41, pp.
476-478). The account given by Ehrenberg of the Polycystina of Barbados
is in many respects very incomplete, and very far from exhausting this
rich mine of remarkable forms. This may be readily seen from the
twenty-five plates of figures of Polycystins in the Barbados Chalk
Deposit published by Bury in 1862 (L. N. 17). The number of species here
figured (140 to 142) is about half of those given by Ehrenberg; and there
are among them numerous generic types, some of great interest, which were
entirely overlooked by the latter; _e.g._ _Saturnalis_ (#Sphæroidea#),
_Cannartidium_ (#Prunoidea#), _Tympanidium_ (#Stephoidea#),
_Cinclopyramis_ (#Cyrtoidea#), &c. Finally, Ehrenberg always (until 1875)
ignored Bury's atlas, which had been published thirteen years ago and was
quite accessible to him. How different were the contents of the two works
may easily be seen from the following abstract.

_Comparative View of the Species of Fossil Radiolaria from Barbados made
known by the figures of Bury in 1862 and of Ehrenberg in 1875._

+-------------------+--------------------+--------+-----------+-------+
| Legion. | Order. | Bury. | Ehrenberg.| Total.|
+-------------------+--------------------+--------+-----------+-------+
| I. Legion | { 1. Sphæroidea | 16 | 13 | 29 |
| SPUMELLARIA | { 2. Prunoidea | 10 | 8 | 18 |
| (PERIPYLEA). | { 3. Discoidea | 37 | 33 | 70 |
| | | | | |
| II. Legion | { 4. Stephoidea | 5 | 2 | 7 |
| NASSELLARIA | { 5. Spyroidea | 13 | 38 | 51 |
| (MONOPYLEA). | { 6. Cyrtoidea | 60 | 188 | 248 |
+-------------------+--------------------+--------+-----------+-------+
| Total, | 141 | 282 | 423 |
+----------------------------------------+--------+-----------+-------+

{clxvi}In 1882 Bütschli still further increased the number of known
Radiolaria from Barbados both by figures and descriptions (L. N. 40), and
gave in particular a very accurate morphological analysis of 12 new
NASSELLARIA (3 #Stephoidea#, 3 #Spyroidea#, and 6 #Cyrtoidea#; L. N. 40,
Taf. xxxii., xxxiii.). The number of the fossil species collected in the
Barbados marl is, however, greater than would appear from the
above-quoted communications. My respected friend, Dr. R. Teuscher, of
Jena, has, at my request, made a large number (about a thousand) of very
accurate drawings with the camera lucida of Polycystina from Barbados
(see p. 1760). From these it appears that the variations in the structure
of the shells, with respect to number, size, and form of the
lattice-pores, of the spines, &c., is much greater than would be supposed
from the figures of Ehrenberg and Bury. I have thus come to the
conviction that the number of species from Barbados (using the word
"species" in the sense understood by those authors) is not less than 400
and probably more than 500. Descriptions of some particularly interesting
new species from this series have been included in the systematic account
of the Challenger Radiolaria. A complete critical investigation of the
Radiolaria of Barbados, and especially an accurate comparison of these
Cainozoic species with the Mesozoic forms from the Jura, on the one hand,
and with recent types on the other, must be left to the future for its
accomplishment (see § 246).

B. The Cainozoic Polycystine tripoli or marl of the Mediterranean coast,
which is probably always of Miocene origin, forms very extensive mountain
ranges both in the south of Europe (Sicily, Calabria, Greece) and in the
north of Africa (from Oran to Tripoli) (§ 246). Hitherto, however, only
one locality has been thoroughly investigated, namely, Grotte in the
province of Girgenti in Sicily (L. N. 35). In the accurate account which
was given of it by Stöhr in 1880, 118 species were described, distributed
in 40 genera (L. N. 35; pp. 72-84); of these 118 species 78 are quite
new, 25 are identical with previously known fossils, and 29 identical
with living forms. Among them are 73 SPUMELLARIA (28 #Sphæroidea#, 8
#Prunoidea#, and 37 #Discoidea#), but only 40 NASSELLARIA (1
#Stephoidea#, 6 #Spyroidea#, and 33 #Cyrtoidea#), and 5 PHÆODARIA
(Dictyochida). The other parts of Sicily from which the same upper
Miocene tripoli has been investigated (belonging to the Tortona stage)
have proved less rich than Grotte. The best known of these places is
Caltanisetta, since upon three genera discovered here (_Haliomma_,
_Cornutella_, _Lithocampe_) the group Polycystina was founded by
Ehrenberg in 1838 (see L. N. 16, p. 3). Afterwards 31 species were
described from this locality, of which 23 were again found in Grotte. The
richest deposit on the Mediterranean coast, however, appears to be at
Oran. A small specimen of the Kieselguhr found there, which was recently
sent to me by Professor Steinman, proved to be pure Radiolarian ooze,
very similar to that now found in the Central Pacific, and contained many
hitherto undescribed species; it is deserving of careful investigation
and comparison.

C. Regarding the Tertiary Radiolarian clay of the Nicobar Islands, see §
247 and L. N. 25, pp. 116-120. Its fauna is incompletely known; probably
it is of Miocene or Oligocene origin.

D. Cainozoic tripoli, containing larger or smaller quantities of
Radiolaria, appears to be rather widely distributed in America. Ehrenberg
has described such from South America (polishing-slate from Morro di
Mijellones, on the coast between Chili and Bolivia), and from North
America (Richmond and Petersburg in Virginia, Piscataway in Maryland).
Similar deposits are also found in the Bermuda Islands (L. N. 4, 1855-56;
L. N. 6, Taf. 18; L. N. 16, pp. 3-9; L. N. 41, pp. 475-478, and L. N. 25,
pp. 2-6).

{clxvii}243. _Mesozoic Radiolaria._--From the Mesozoic or Secondary period
numerous well-preserved Radiolaria have recently been described. They
belong for the most part to the Jurassic formation (see notes A, B, C),
whilst the more recent Chalk (see note D) and the older Trias (see note E)
have hitherto yielded but few species. All the main divisions of the Jura,
both the upper (Malm) and the middle (Dogger), and especially the lower
(Lias) appear in certain localities to be very rich in well-preserved
shells of fossil Polycystina. Most of these are aggregated together in
coprolites and quartzites (jasper, chert, flint, &c., § 248). The majority
are #Cyrtoidea#, the minority #Sphæroidea# and #Discoidea# in almost equal
proportions; a few #Beloidea# (_Sphærozoum_) and #Phæocystina# (Dictyocha)
are also found among them. The general morphological character of these
Jurassic Radiolaria is very different from that of the nearly related
Tertiary and living forms. In general, their siliceous shells are firmer
and more massive, usually also somewhat larger, but of simpler structure.
The manifold delicate appendages (spines, bristles, feet, wings, &c.) which
are so richly developed in the living SPUMELLARIA and NASSELLARIA, and are
also well shown in the Tertiary species, are entirely wanting in the
majority of the Jurassic Polycystina. The #Sphæroidea# and #Prunoidea# are
all simple spherical or ellipsoidal lattice-shells (Monosphærida);
concentric lattice-shells (Polysphærida) are entirely wanting. The
#Cyrtoidea# are, for the most part, devoid of radial processes or basal
feet (Eradiata); triradiate and multiradiate forms, such as are found
abundantly in the recent and Tertiary formations, are very rare. The large
number of many-jointed forms (Stichocyrtida) and of #Cyrtoidea# with
latticed basal opening is very striking.

A. The most important work on the Jurassic Radiolaria, regarding which
but little was known prior to the year 1885, is the valuable and in some
respects very interesting Beiträge zur Kenntniss der fossilen Radiolarien
aus Gesteinen des Jura, by Dr. Rüst of Freiburg i. B. (1885,
Palæontographica, Bd. xxxi. 51 pp. with 12 plates). Unfortunately this
important work was issued only when about half of the present Report was
printed off, so that it was no longer possible to include the 234 species
there described in its systematic part. I have therefore elsewhere given
a list of the Jurassic Radiolaria, and at present only make the following
remarks:--Of the 234 species described, the larger half (130) belong to
the NASSELLARIA (#Cyrtoidea#), the smaller half (102) to the SPUMELLARIA
(38 #Sphæroidea#, 14 #Prunoidea#, and 50 #Discoidea#). In addition, there
are 2 PHÆODARIA depicted, and several spicules which are probably to be
referred to the #Beloidea#. Among the 130 #Cyrtoidea# (of which 2 are
described as #Botryodea#), there are 24 Monocyrtida, 14 Dicyrtida, 22
Tricyrtida, and 70 Stichocyrtida. Just as striking as the predominant
number of the last is the fact that there are only very few triradiate
(9) and multiradiate (4) species found among these 130 #Cyrtoidea#, as
also the large number of species with latticed basal opening;
#Stephoidea# appear to be entirely wanting. The rich material of jasper,
chert, flint, and coprolites in which Dr. Rüst found these Radiolaria, is
derived for the most part from the Jurassic rocks of Germany (Hanover,
South Bavaria), Tyrol, and Switzerland (compare § 248).

{clxviii}B. Jurassic Radiolaria from Italy, also found in jasper, which
are closely related to the forms from Germany and Switzerland described
by Dr. Rüst, were made known so long ago as 1880 by Dante Pantanelli in
his treatise I Diaspri della Toscana e i loro Fossili (Rome, 1880, 33 pp.
60 figs.). Pantanelli believes, however, that this jasper is for the most
part of Eocene origin; but from his description, and especially from the
morphological character of the forms which he figures, it appears very
probable "that these Tuscan jaspers from Galestro, like those of the
Swiss conglomerates, are found in a secondary locality and belong to the
Jurassic period" (Rüst, L. N. 51, p. 3). Unfortunately the figures of
Pantanelli are so small and incomplete that a reliable determination of
the species is hardly possible; for example, the lattice-work is only
given in ten of the sixty figures. Among the 32 recorded species 15 are
SPUMELLARIA (6 #Sphæroidea# and 9 #Discoidea#) and 17 NASSELLARIA (4
#Stephoidea# and 13 #Cyrtoidea#); many of which seem to be identical with
the forms more accurately described by Dr. Rüst (compare p. 1762).

C. From the Lias of the Alps and more particularly "from the lower
Liassic beds of the Schafberg near Salzburg," Dr. Emil von Dunikowski in
1882 described 18 species of fossil Radiolaria (L. N. 44, pp. 22-34, Taf.
iv.-vi.); most of these are #Sphæroidea# and #Discoidea# and appear to
have been more or less altered by petrological changes; their spongy
structure is probably secondary.

D. Cretaceous Radiolaria have been hitherto described only in very small
numbers; quite recently Dr. Rüst has found a larger number chiefly in
flints from the English chalk, but they have not yet been published. In
1876 Zittel described 6 very well-preserved species from the upper chalk
of North Germany (L. N. 29, pp. 76-96, Taf. ii.); among them were 1
#Sphæroidea#, 1 #Discoidea#, 1 Dictyocha, and 3 #Cyrtoidea#.

E. Triassic Radiolaria have recently been discovered by Dr. Rüst in
chert, but have not yet been described.

244. _Palæozoic Radiolaria._--The number of Radiolaria which are known from
the Palæozoic or Primary formations is much less than from either the
Mesozoic or Cainozoic periods. Here, however, the investigations of recent
times have yielded important information; a few species, at all events, of
Polycystina (mostly #Sphæroidea#) are now known from various Palæozoic
formations, and not only from the Permian ("Zechstein") and the
Coal-measures, but also from the older Devonian and Silurian systems. Even
in the still older Cambrian rocks a few fossil Radiolaria have been found.
All these Palæozoic Radiolaria are Polycystina of very simple form and
primitive structure, mostly simple SPUMELLARIA (latticed spheres,
ellipsoids, lenses, &c.), but partly also simple NASSELLARIA.

The important discoveries which have recently been made by Dr. Rüst
regarding the occurrence of Radiolaria in all the Palæozoic formations
have not yet been published. From conversations with this estimable
palæontologist I have learned, however, that he has pursued his fruitful
investigation of the Mesozoic quartzites (§ 243), and has met with no
less success in the case of similar Palæozoic structures. Although the
number of species hitherto discovered is relatively small, the important
conclusion appears to be warranted that they extend as far as the
Silurian and Cambrian systems. All these very ancient SPUMELLARIA
(#Sphæroidea#) and NASSELLARIA (#Cyrtoidea#) {clxix}exhibit very
primitive structural relations. The occurrence of fossil Polycystina in
the Carboniferous formation of England has been incidentally mentioned by
W. J. Sollas:--"In the carboniferous beds of North Wales pseudomorphs of
Radiolaria in calcite occur, along with minute quartz crystals" (Ann. and
Mag. Nat. Hist., 1880, ser. 5, vol. vi. p. 439); and in the siliceous
slate-beds of Saxony Rothpletz has shown the existence of a few
#Sphæroidea# (Zeitschr. d. Deutsch. Geol. Gesellsch., 1800, p. 447).

245. _Abundance of Radiolaria in the Various Rocks._--The relative quantity
of well-preserved or at all events recognisable Radiolaria in the different
rocks is very variable. In this respect three different degrees may be
distinguished, which may be called shortly "pure, mixed, and poor"
Radiolarian formations. The _pure_ Radiolarian rocks consist for the
greater part (usually much more than half, sometimes even more than
three-quarters) of closely compacted often calcined masses of siliceous
Polycystine shells. To this category belong the pure Miocene Polycystine
marls of Barbados (§ 246), the Tertiary Polycystine clay of the Nicobar
Islands (§ 247), and the Polycystine quartz of the Jura (§ 248). All these
pure Radiolarian rocks may be regarded as fossil Radiolarian ooze (§ 237),
and are certainly of deep-sea origin, having probably been deposited at
depths greater than 2000 fathoms. Their palæontological character also is
in favour of this view, for the abyssal Osculosa (§ 235) are more abundant
and richer in species than the pelagic Porulosa (§ 233). The elevation of
this deep-sea layer above the surface of the sea appears to have taken
place but seldom; it has only been observed on a large scale at Barbados
and in the Nicobar Islands. The _mixed_ Radiolarian rocks are much more
common; they were probably deposited at much less depths, or perhaps are
not true deep-sea formations at all. The siliceous shells of Polycystina
always constitute less than half (sometimes less than one-tenth) of their
mass, and are less prominent than other siliceous remains (Diatoms), or
calcareous remains (Foraminifera), or in some cases than the mineral
constituents (pumice, &c.). To this group belong many of the
above-mentioned Tertiary marls and clays (especially the Mediterranean
Tripoli), also many flints, cherts, and other quartzites from Mesozoic
strata (especially from the Jura), and probably also some palæozoic
quartzites. The marine ooze from which they have originated may have been
deposited at very various, even at slight, depths of the ocean. Formations
_poor_ in Radiolaria, which contain only a few species of SPUMELLARIA and
NASSELLARIA mingled with other fossil remains and mineral particles, occur
in all formations and are probably very widely distributed. Further careful
examination of thin sections (especially of coprolites) will yield here a
rich harvest of new forms. Both the mixed and the pure Radiolarian
formations may be divided according to their petrographic characters into
three groups, which, however, are connected by intermediate varieties--(1)
soft, chalky marl (§ 246), (2) plastic clay (§ 247), and (3) hard, flinty
quartz (§ 248).

{clxx}246. _Radiolarian Marl._--Those soft, friable rocks, which contain a
large quantity of calcareous matter, but consist for the most part of the
shells of SPUMELLARIA and NASSELLARIA, are called Radiolarian or
Polycystine marl, often more correctly Polycystine tripoli; the best known
example of them is the chalky marl of Barbados in the Antilles (§ 242). The
Tertiary mountain system of this island, which in Mount Hillaby rises to a
height of 1147 feet and includes about 15,800 acres, consists almost
exclusively of these remarkable masses of rock. Most of it appears as a
soft, earthy, often chalky marl, with a considerable but variable amount of
calcareous matter. Those specimens, the greater half of which is composed
of well-preserved siliceous shells of Polycystina, and which contain little
lime, approach the tripoli and "Kieselguhr." Those specimens, however,
which contain the largest amount of calcareous matter resemble common
writing chalk in consistency, and consist for the most part of shells of
Foraminifera and their fragments; of these there are only few species but
large numbers of individuals, generally in small fragments with a fine
calcareous powder between them. They may be regarded as fossil Globigerina
ooze (§ 238). In a third group of specimens from Barbados the quantity of
fragments of pumice and other volcanic matters predominates; the amount of
clay is also very considerable; these deposits pass over partly into actual
clay partly into volcanic tuff. A fourth group exhibits relations to a
coarser often ferruginous material, and although the shells of Polycystina
are less abundant in it, still it may be shown to be composed largely of
fragments and metamorphosed remains of them. The colour of this deposit,
which in some places passes over into sandstone, in others into clay, is
usually rather dark, grey, brown, sometimes red and occasionally black
(bituminous). The Radiolarian marls of the first two groups, which
sometimes approach the white chalk, sometimes the Kieselguhr, are grey, or
even pure white (see note A). The same constitution is exhibited by the
yellowish or white, very light and friable Polycystine marls of Sicily,
which in Caltanisetta approach the chalk, and in Grotte the Kieselguhr. In
Greece (Ægina, Zante, &c.), on the other hand, they pass over into plastic
clay, and the same occurs in the Baden marl of the Vienna basin. In North
Africa, however, on the Mediterranean shores of which the Radiolarian marl
seems to be very widely distributed (from Tripoli to Oran), it sometimes
becomes changed into actual firm polishing slate, sometimes into
pulverulent Kieselguhr or tripoli (Terra tripolitana, see note B). Most of
these Radiolarian marls appear to date from the middle Tertiary (Miocene)
period, and to be deep-sea formations.

A. The Polycystine marl of Barbados appears at different parts of the
island to present greater variations in its petrographical and
zoographical composition than would appear from Ehrenberg's description
(1875, L. N. 25, pp. 106-116). Through the kindness of one of my former
students, Dr. Dorner, to whom I take this opportunity of expressing my
thanks for the favour, I received a large number of specimens of Barbados
rock, taken from various parts of the island, and they exhibit very great
variations in their external appearance, their chemical composition, and
the {clxxi}Radiolaria which they contain. The white specimens resembling
Kieselguhr contained approximately 60 to 70 per cent. by volume of
Radiolarian shells, the yellowish marl 40 to 50 per cent., and the brown
and black (bituminous) marl 10 to 20 per cent. or less. Two analyses of
the first, which my friend Dr. W. Weber was good enough to carry out,
yielded different results from those which are given by Ehrenberg on the
basis of Rammelsberg's analyses (L. N. 25, p. 116). The results of both
are here given for comparison.

For further comparison I here add the three different analyses of Miocene
Tripoli-marls from Sicily, given by Stöhr on the authority of Fremy,
Schwager, and Mottura (Tagebl. d. fünfzigsten Versamml. Deutsch. Naturf.
u. Aertzte in München, 1877, p. 163).

--------------------------+-------------+--------------+-------------
Composition. |Tripoli from | Tripoli from | Tripoli from
| Licata | Grotte | Caltanisetta
| (Fremy). | (Schwager). | (Mottura).
--------------------------+-------------+--------------+-------------
Silica, | 30.98 | 58.58 | 68.6
Alumina, | 17.54 | 11.51 | }
Oxide of iron, | 0.33 | 1.84 | } 3.6
| | |
Lime, } | 38.09 | { 8.49 | }
Magnesia, } | | { 0.41 | } 12.1
| | |
Water and organic matter,}| | { 11.26 | }
Carbonic acid, }| 13.06 | { 7.12 | } 15.2
+-------------+--------------+------------
| 100.00 | 99.21 | 99.5
--------------------------+-------------+--------------+-------------

B. The Radiolarian marl of the Mediterranean appears, judging by the
accounts already published, to stretch along a considerable part of the
coast in the earlier and middle Tertiary formations; thus it occurs of
similar composition in widely separated localities, in Sicily, Calabria,
Zante, and Greece; in North Africa from Tripoli to Oran and probably much
farther. So long ago as 1854 Ehrenberg, in his Mikrogeologie (L. N. 6)
gave a series of important, even if incomplete, communications regarding
the "chalky white calcareous marl of Caltanisetta" (Taf. xxii.), the
"Platten marl of Zante" (Taf. xx.), the "plastic clay of Ægina" (Taf
xix.), and the "polishing slate of Oran" (Taf. xxi.). In 1880 Stöhr
showed in his fundamental description of the Tripoli from {clxxii}Grotte
in Sicily (L. N. 35) that its Radiolarian fauna is much richer than
Ehrenberg supposed. The same is the case in the Tripoli of Caltanisetta,
and also in the Baden marl of the Vienna basin. The richest deposit
appears to be the pure Kieselguhr-like Tripoli from Oran; a small
specimen, which was recently sent to me by Professor Steinmann of
Freiburg, i. B., contained many hitherto undescribed species, and was at
least as rich as the purest Barbados marl.

247. _Radiolarian Clays._--Among the Radiolarian or Polycystine clays we
include the firm, often plastic, formations, which contain a larger
proportion of Radiolaria than of other organic remains. The first of these
to be mentioned is the Cainozoic formation of the Nicobar Islands in
Further India, which rises to a height of 2000 feet above the level of the
sea, and consists for the most part of coloured masses of clay of varying
constitution; on Car Nicobar these are mostly grey or reddish, on the
Island of Camorta they are partly strongly ferruginous and red and yellow
(_e.g._ at Frederickshaven), partly white and light, like meerschaum
(_e.g._ at Mongkata). The latter varieties appear to pass over into pure
loose Polycystine marl like that of Barbados, the former into calcareous
sandstone. Although the Polycystine clays of the Nicobar Islands are as yet
only very incompletely known, it may be concluded with great probability
that they are true deep-sea formations and nearly allied to those recent
forms of red clay, which by their abundance in Radiolaria most nearly
approach the Radiolarian ooze, such for example as the red clay of the
North Pacific between Japan and the Sandwich Islands (Stations 241 to 245,
compare §§ 229 and 239). With this view agrees also the greater or less
quantity of pumice dust and other volcanic products. Probably Radiolarian
clays like those of the Nicobar Islands occur also in other Tertiary rocks;
part of the Barbados marl passes by gradually increasing content of clay
into such; and in this case also the amount of included pumice is often
considerable. Many mixed Radiolarian marls of the Mediterranean (_e.g._, of
Greece and Oran) also appear to pass over at certain points into
Radiolarian clay.

The Radiolarian clays of the Nicobar Islands are unfortunately very
incompletely known both as regards their geological nature and their
palæontological composition. The communications of Rink (Die
Nikobaren-Inseln, eine geographische Skizze, Kopenhagen, 1847) and of
Ehrenberg (L. N. 6, p. 160 and L. N. 25, pp. 116 to 120) leave many
important questions unanswered. The latter has only figured twenty-three
species in his Mikrogeologie (L. N. 6, Taf. xxxvi.). In his tabular list
of names (L. N. 25, p. 120) he only incompletely records thirty-nine
species, although in 1850, immediately after the first examination of the
Nicobar clay, he had distinguished "more than a hundred species, partly
new, partly identical with those of Barbados" (L. N. 16, p. 8). I have
unfortunately been unable in spite of many efforts, to obtain for
investigation a specimen of Nicobar clay. The only microscopical
preparation (from Ehrenberg's collection), which I was able to examine,
contained several hitherto undescribed species. A thorough systematic
examination of these important Radiolarian clays is a pressing necessity,
especially as they seem to be markedly different from those of the
Mediterranean (from Ægina, Zante, &c.).

{clxxiii}248. _Radiolarian Quartzes._--Under the name Radiolarian or
Polycystine quartzes are included those hard, siliceous rocks, which
consist for the most part of the closely compacted shells of SPUMELLARIA
and NASSELLARIA. To these "cryptocrystalline quartzes," or better,
quartzites, belong more especially the pure Radiolarian formations of the
Jura, which have been described as flint, chert, jasper, as well as other
cryptocrystalline quartzites. Most of the rocks of this nature hitherto
examined are from Germany (Hanover, South Bavaria), Hungary, Tyrol, and
Switzerland; others are known from Italy (Tuscany). They occur both in the
upper and middle, but especially in the lower Jurassic formation (also in
the lower layers of the Alpine Lias). A small part of them has been
examined in their primary situation (the red jaspers of Allgäu and Tyrol),
the greater part, however, only as loose rolled stones in secondary
situations (thus in Switzerland in the breccia of the Rigi, in the
conglomerate of the Uetli-Berg, and in many boulders of the Rhine, the
Limmat, the Reuss, and the Aar). The greatest abundance, however, of
Jurassic Radiolaria has been yielded by the silicified coprolites from the
Lias of Hanover. These "Radiolarian coprolites" are roundish or cylindrical
bodies, which may attain the size of a goose-egg; they probably originated
from Fish or Cephalopods, which had fed upon Crustacea, Pteropoda, and
similar pelagic organisms, whose stomachs were already full of Radiolarian
skeletons. Next to the coprolites the richest is the red jasper, whose
colour varies from bright to dark red; it constitutes a true "silicified
deep-sea Radiolarian ooze." The "_Aptychus_ beds" also of South Bavaria and
Tyrol are very rich, and have furnished about one-third of all the
Radiolaria known from the Jura; most of the species too are very well
preserved (compare § 243).

Regarding the remarkable composition and manifold varieties of the
Jurassic Radiolarian quartz, the very full treatise of Dr. Rüst may be
consulted (L. N. 51). The very interesting Radiolarian coprolites, which
that author has discovered in the lower and middle Jura of Hanover, occur
in astonishing numbers in the iron mines at the village of Gross-Ilsede,
four and a half miles south of the town of Peine. They constitute from 2
to 5 per cent. by weight of the Liassic iron ore; of this latter, in the
year 1883 alone, not less than two hundred and eighty million kilograms
were excavated. It is very probable that the careful microscopic
examination of thin sections of coprolites, as well as of flints, chert,
jasper, and other quartzites, would yield a rich harvest of fossil
Radiolaria in other formations also. In Italy Dante Pantanelli has
discovered interesting Polycystine jaspers in Tuscany (L. N. 36, 45);
these also appear to occur in the Jura (compare § 243, and L. N. 51, pp.
3-10).

249. _Fossil Groups._--The preservation of Radiolaria in the fossil state
is, of course, primarily dependent on the composition of their skeleton.
Hence the ACANTHARIA, whose acanthin skeleton although firm is readily
soluble, are never found fossil. The same is true of the skeletons of the
PHÆODARIA, which consist of a silicate of carbon; here, however, a single
exception is found in the Dictyochida, a subfamily of the Cannorrhaphida,
the isolated parts of whose skeletons appear to consist of pure silica, and
{clxxiv}are often found fossil. Of the two other legions those families
which possess no skeleton are of course excluded; the Nassellida among the
NASSELLARIA, and the Thalassicollida and Collozoida among the SPUMELLARIA.
Thus of the 85 known families there remain scarcely 55 of which the
skeletons may be expected in the fossil state; and of these scarcely half
have been actually observed in this condition. Of the 20 orders of this
class enumerated in § 155, the following 9 may be, for palæontological and
geological purposes, completely excluded:--(A) The 4 orders of ACANTHARIA
(1, #Actinelida#; 2, #Acanthonida#; 3, #Sphærophracta#; 4, #Prunophracta#);
(B) 3 orders of PHÆODARIA (5, #Phæosphæria#; 6, #Phæogromia#; 7,
#Phæoconchia#); (C) 1 order of NASSELLARIA (8, #Nassoidea#); (D) 1 order of
SPUMELLARIA (9, #Colloidea#). From a geological point of view the following
6 orders, although occasionally found fossil, are of quite subordinate
importance:--(A) Among the SPUMELLARIA (10, #Beloidea#, and 11,
#Larcoidea#); (B) among the NASSELLARIA (12, #Plectoidea#; 13,
#Stephoidea#; 14, #Botryodea#); (C) among the PHÆODARIA (15, the
#Phæocystina#). On the other hand the following 5 orders, which are the
main constituents of Radiolarian rocks, are of pre-eminent geological
importance:--(A) Among the SPUMELLARIA (16, #Sphæroidea#; 17, #Prunoidea#;
18, #Discoidea#); (B) among the NASSELLARIA (19, #Spyroidea#, and 20,
#Cyrtoidea#). The numerical relation in which the different families of
these orders appear in the Radiolarian formations may be seen on consulting
§ 157.

250. _Fossil and Recent Species._--The fact that there are many Radiolaria
living at the present day, whose shells are found fossil in Tertiary rocks,
is of great phylogenetic and geological significance. This appeared to be
the case even from the older observations upon the Polycystina of the
Barbados marl (see note A), but more recent and extensive observations both
upon these and upon the Miocene Radiolaria of Sicily, have shown that the
number of these "living fossil" forms is much greater than was previously
supposed (see note B). Among the Miocene Radiolaria numerous species, both
of SPUMELLARIA (especially #Sphæroidea# and #Discoidea#) and of NASSELLARIA
(especially #Spyroidea# and #Cyrtoidea#) are not to be distinguished from
the corresponding still living forms (see notes C, D). On the other hand,
those genera, which are rich both in species and individuals (recent as
well as fossil), present continuous series of forms which lead gradually
and uninterruptedly from old Tertiary species to others still living, which
are specifically indistinguishable from them. These interesting
morphological facts are capable of direct phylogenetic application, and
furnish valuable proofs of the truth of the theory of descent.

A. Ehrenberg, in his list of fossil Polycystina (L. N. 25, pp. 64-85,
1875), records 325 species of which 26 are still living.

{clxxv}B. Stöhr, in his list of Miocene Radiolaria from Grotte (L. N. 35,
p. 84, 1880), records 118 species, of which 29 are still living.

C. Teuscher, who at my request has made a large number of comparative
measurements and drawings, both of fossil and living Radiolaria, comes to
the conclusion that numerous SPUMELLARIA and NASSELLARIA from Barbados
are to-day extant and unchanged in the Radiolarian ooze of the deep
Pacific Ocean (compare § 242A, and p. 1760, Note).

D. From the comparative investigations, which I have made during the last
ten years into the recent deep-sea Radiolaria of the Challenger
collection and the Miocene Polycystina of Barbados, it appears that about
a quarter of the latter are identical with living species of the former.

{clxxvi}BIBLIOGRAPHICAL SECTION.

CHAPTER XI.--LITERATURE AND HISTORY.

251. _List of Publications from 1834 to 1884_:--

_Note._--In the text the references to the following publications are
indicated by the letters L. N.

1. 1834. MEYEN, F., Palmellaria (Physematium, Sphærozoum), in Beiträge
zur Zoologie, gesammelt auf einer Reise um die Erde. _Nova Acta Acad.
Cæs. Leop.-Carol._, vol. xvi., Suppl., p. 160, Taf. xxviii. figs. 1-7.

2. 1838. EHRENBERG, G., Polycystina (Lithocampe, Cornutella, Haliomma) in
Ueber die Bildung der Kreidefelsen und des Kreidemergels durch
unsichtbare Organismen. _Abhandl. d. k. Akad. d. Wiss. Berlin_, p. 117.

3. 1839. EHRENBERG, G., Ueber noch jetzt lebende Thierarten der
Kreidebildung (Haliomma radians). _Abhandl. d. k. Akad. d. Wiss. Berlin_,
p. 154.

4. 1844-1873. EHRENBERG, G., Vorläufige Mittheilungen über Beobachtungen
von Polycystinen. _Monatsber. d. k. preuss. Akad. d. Wiss. Berlin_.
Republished with illustrations in the Mikrogeologie (L. N. 6) and in the
two treatises of 1872 (L. N. 24) and 1875 (L. N. 25). Compare the
_Monatsberichte_ of 1844 (pp. 57, 182, 257), of 1846 (p. 382), of 1847
(p. 40), of 1850 (p. 476), of 1854 (pp. 54, 205, 236), of 1855 (pp. 292,
305), of 1856 (pp. 197, 425), of 1857 (pp. 142, 538), of 1858 (pp. 12,
30), of 1859 (p. 569), of 1860 (pp. 765, 819), of 1861 (p. 222), of 1869
(p. 253), of 1872 (pp. 300-321), of 1873 (pp. 214-263). Only one of these
small papers is of permanent value, The First Systematic Arrangement of
the Polycystina in 7 families, 44 genera, and 282 species (_Monatsber. d.
k. preuss. Akad. d. Wiss. Berlin_, 1847, p. 54). Compare my Monograph
(1862, L. N. 16), pp. 3-12, 214-219.

5. 1851. HUXLEY, TH., Upon Thalassicolla, a new Zoophyte. _Ann. and Mag.
Nat. Hist._, ser. 2, vol. viii. pp. 433-442, pl. xvi.

6. 1854. EHRENBERG, G., Mikrogeologie. Figures of numerous Polycystina
on 8 plates; Taf. xviii. figs. 110, 111; Taf. xix. figs. 48-56, 60-62;
Taf. xx. Nr. i., figs. 20-25, 42; Taf. xxi. figs. 51-56; Taf. xxii. figs.
20-40; Taf. xxxv. A., Nr. xix. A. fig. 5; Taf. xxxv. B. figs. 16-23; Taf.
xxxvi. figs. 1-33.

7. 1855. BAILEY, J. W., Notice of Microscopic Forms of the Sea of
Kamtschatka. _Amer. Journ. Sci. and Arts_, vol. xxii. p. 1, pl. i.

8. 1855. MÜLLER, JOHANNES, Ueber Sphærozoum und Thalassicolla.
_Monatsber. d. k. preuss. Akad. d. Wiss. Berlin_, p. 229.

9. 1855. MÜLLER, JOHANNES, Ueber die im Hafen von Messina beobachteten
Polycystinen (Haliomma, Eucyrtidium, Dictyospyris, Podocyrtis).
_Monatsber. d. k. preuss. Akad. d. Wiss. Berlin_, p. 671.

10. 1856. MÜLLER, JOHANNES, Ueber die Thalassicollen, Polycystinen und
Acanthometren des Mittelmeeres. _Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin_, p. 474.

11. 1858. MÜLLER, JOHANNES, Erläuterung einiger bei St. Tropez am
Mittelmeer beobachteter Polycystinen und Acanthometren. _Monatsber. d. k.
preuss. Akad. d. Wiss. Berlin_, p. 154.

12. 1858. MÜLLER, JOHANNES, Ueber die Thalassicollen, Polycystinen und
Acanthometren des Mittelmeeres, _Abhandl. d. k. Akad. d. Wiss. Berlin_,
pp. 1-62, Taf. i.-xi. (The fundamental treatise on the Radiolaria.)

{clxxvii}13. 1858. SCHNEIDER, ANTON, Ueber zwei neue Thalassicollen von
Messina. _Archiv f. Anat. u. Physiol._, p. 38, Taf. iii. B, figs. 1-4.

14. 1858. CLAPARÈDE et LACHMANN, Echinocystida (Plagiacantha et
Acanthometra). Études sur les Infusoires et les Rhizopodes, p. 458, pl.
xxii. figs. 8, 9; pl. xxiii. figs. 1-6.

15. 1860. HAECKEL, ERNST, Ueber neue lebende Radiolarien des
Mittelmeeres. _Monatsber. d. k. preuss. Akad. d. Wiss. Berlin_, pp. 794,
835.

16. 1862. HAECKEL, ERNST, Die Radiolarien (Rhizopoda radiaria). Eine
Monographie. 572 pp. fol. with an Atlas of 35 Copperplates.

17. 1862. BURY, Mrs., Polycystins, figures of remarkable forms in the
Barbados Chalk Deposit. Ed. ii. By M. C. Cooke, 1868. 25 quarto plates,
photographed from drawings by hand, containing many forms overlooked by
Ehrenberg from Barbados.

18. 1863 HARTING, PAUL, Bijdrage tot de Kennis der mikroskopische Fauna
en Flora van de Banda-Zee (Diep-Zee-Polycystinen). _Verhandl. d. Kon.
Akad. van. Wetensch. Amsterdam_, vol. ix. p. 30, pls. i.-iii.

19. 1865. HAECKEL, ERNST, Ueber den Sarcode-Körper der Rhizopoden
(Actinelius, Acanthodesmia, Cyrtidosphæra, &c.). _Zeitschr. f. wiss.
Zool._, Bd. xv. p. 342, Taf. xxvi.

20. 1867. SCHNEIDER, ANTON, Zur Kenntniss des Baues der Radiolarien
(Thalassicolla). _Archiv f. Anat. u. Physiol._, 1867, p. 509.

21. 1870. HAECKEL, ERNST, Beiträge zur Plastiden Theorie (Myxobrachia;
Amylum in den gelben Zellen). _Jenaische Zeitschr. für Naturw._, Bd. v.
p. 519-540, Taf. xviii.

22. 1871. CIENKOWSKI, L., Ueber Schwärmer-Bildung bei Radiolarien.
_Archiv f. mikrosk. Anat._, Bd. vii. p. 372-381, Taf. xxix.

23. 1872. WAGNER, N., Myxobrachia Cienkowskii. _Bull. d. Acad. St.
Petersburg_, vol. xvii. p. 140.

24. 1872. EHRENBERG, GOTTFRIED, Mikrogeologische Studien über das
kleinste Leben der Meeres-Tiefgründe aller Zonen und dessen geologischen
Einfluss. _Abhandl. d. k. Akad. d. Wiss. Berlin_, 1872. Mit 12 Tafeln.
(The Latin diagnoses of 113 new species here mentioned are given in the
_Monatsberichte_ of April 25, 1872, pp. 300-321.)

25. 1875. EHRENBERG, GOTTFRIED, Polycystinen-Mergel von Barbados
(Fortsetzung der Mikrogeologischen Studien). _Abhandl. d. k. Akad. d.
Wiss. Berlin_, 1875, 168 pag. mit 30 Tafeln. (The Latin diagnoses of 265
species here recorded are given in Namensverzeichniss der fossilen
Polycystinen von Barbados. _Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin_, Jan. 30, 1873, pp. 213-263.)

26. 1876. HERTWIG, RICHARD, Zur Histologie der Radiolarien.
Untersuchungen über den Bau und die Entwickelung der Sphærozoiden und
Thalassicolliden. 91 pp. with 5 plates.

27. 1876. MURRAY, JOHN, Challengerida. Preliminary Reports on Work done
on board the Challenger. _Proc. Roy. Soc. Lond._, vol. xxiv. pp. 471-536,
pl. xxiv.

28. 1876. ZITTEL, KARL, Palæozoologie, Bd. i. pp. 114-126, figs. 46-56.

29. 1876. ZITTEL, KARL, Ueber fossile Radiolarien der oberen Kreide.
_Zeitschr. d. deutsch. geol. Gesellsch._, Bd. xxviii. pp. 75-96, Taf. ii.
(with figures of six Cretaceous species).

30. 1877. MIVART, ST. GEORGE, Notes touching recent researches on the
Radiolaria. _Journ. Linn. Soc. Lond._ (Zool.), vol. xiv. pp. 136-186.
(Historical sketch of previous literature.)

31. 1877. WYVILLE THOMSON, The Voyage of the Challenger--The Atlantic,
vol. i. pp. 231-237, figs. 51-54; vol. ii. pp. 340-343, figs. 58, 59, &c.

32. 1878. HAECKEL, ERNST, Das Protistenreich, eine populäre Uebersicht
über das Formengebiet der niedersten Lebewesen, pp. 101-104.

33. 1879. HERTWIG, RICHARD, Der Organismus der Radiolarien. _Jenaische
Denkschriften_, Bd. ii. Taf. vi.-xvi. pp. 129-277.

34. 1879. HAECKEL, ERNST, Ueber die Phæodarien, eine neue Gruppe
kieselschaliger mariner Rhizopoden. _Sitzungsb. med.-nat. Gesellsch.
Jena_, December 12, 1879.

35. 1880. STÖHR, EMIL, Die Radiolarien-Fauna der Tripoli von Grotte
(Provinz Girgenti in Sicilien). _Palæontographica,_ Bd. xxvi. pp. 71-124,
Taf. xvii.-xxiii. A preliminary communication regarding this fauna from
the tripoli is given in _Tagebl. d. Naturf. Versamml. München_, 1877.

{clxxviii}36. 1880. PANTANELLI, DANTE, I Diaspri della Toscana e i loro
fossili. _Real. Accad. dei Lincei_, ser. 3, vol. vii. pp. 13-34, Tab. i.
Radiolaria di Calabria. _Atti. Soc. Tosc._, p. 59.

37. 1881. HAECKEL, ERNST, Prodromus Systematis Radiolarium, Entwurf eines
Radiolarien-Systems auf Grund von Studien der Challenger-Radiolarien.
_Jenaische Zeitschr. für Naturw._, Bd. xv. pp. 418-472.

38. 1881. BRANDT, KARL, Untersuchungen an Radiolarien. _Monatsber. d. k.
preuss. Akad. d. Wiss. Berlin_, (April 21), pp. 388-404, Taf. i.

39. 1882. BRANDT, KARL, Ueber die morphologische und physiologische
Bedeutung des Chlorophylls bei Thieren. I. Artikel. _Archiv f. Anat. u.
Physiol._, pp. 125-151, Taf. i. II. Artikel. _Mittheil. a. d. Zool.
Station zu Neapel_, Bd. iv. pp. 193-302, Taf. xix., xx.

40. 1882. BÜTSCHLI, OTTO, Beiträge zur Kenntniss der
Radiolarien-Skelette, insbesondere der der Cyrtida. _Zeitschr. f. wiss.
Zool._, Bd. xxxvi. pp. 485-540, Taf. xxxi.-xxxiii.

41. 1882. BÜTSCHLI, OTTO, Radiolaria. In Bronn's Klassen und Ordnungen
des Thierreichs. Bd. i., Protozoa, pp. 332-478, Taf. xvii.-xxxii.

42. 1882. GEDDES, PATRICK, Further Researches on Animals containing
Chlorophyll. _Nature_, pp. 303-305.

43. 1882. GEDDES, PATRICK, On the Nature and Functions of the "Yellow
Cells" of Radiolarians and Coelenterates. _Proc. Roy. Soc. Edin._, p.
377.

44. 1882. DUNIKOWSKI, EMIL, Die Spongien, Radiolarien und Foraminiferen
der Unter-Liassischen Schichten vom Schafberg bei Salzburg. _Denkschr. d.
k. Akad. d. Wiss. Wien_, Bd. xlv. pp. 22-34. Taf. iv.-vi.

45. 1882. PANTANELLI, DANTE, Fauna miocenica di Radiolari del Appennino
settentrional. _Boll. Soc. Geol. Ital._

46. 1883. HAECKEL, ERNST, Die Ordnungen der Radiolarien (Acantharia,
Spumellaria, Nassellaria, Phæodaria). _Sitzungsb. med.-nat. Gesellsch.
Jena_, February 16, 1883.

47. 1883. HERTWIG, OSCAR, Die Symbiose oder das Genossenschaftsleben im
Thierreich. 56. _Versamml. Deutscher Naturf. u. Aerzte_, Freiburg i/B.

48. 1883. RÜST, WILHELM, Ueber das Vorkommen von Radiolarien-Resten in
kryptokrystallinischen Quarzen aus dem Jura und in Koprolithen aus dem
Lias. 56. _Versamml. Deutscher Naturf. u. Aerzte_, Freiburg i/B.

49. 1884. CAR, LAZAR, Acanthometra hemicompressa (= Zygacantha
semicompressa). _Zool. Anzeiger_, p. 94.

50. 1884. HAECKEL, ERNST, Ueber die Geometrie der Radiolarien
(Promorphologie). _Sitzungsb. med.-nat. Gesellsch. Jena_, November 22,
1883.

251 A. _Supplementary List of Works Published in_ 1885:--

51. 1885. D. RÜST, Beiträge zur Kenntniss der fossilen Radiolarien aus
Gesteinen des Jura. 45 pp. 4to, and 20 plates. _Palæontographica_, Bd.
xxxi. (oder iii. Folge, vii. Band).

52. 1885. KARL BRANDT, Die koloniebildenden Radiolarien (Sphærozoeen) des
Golfes von Neapel und der angrenzenden Meeres-Abschnitte. 276 pp. 4to,
and 8 plates.

53. 1885. JOHN MURRAY, Narrative of the Cruise of H.M.S. Challenger, with
a general account of the scientific results of the Expedition. Vol i.
First part, pp. 219-227, pl. A. Second part, pp. 915-926, pl. N. fig. 2.

54. 1885. ERNST HAECKEL, System der Acantharien. _Sitzungsb. med.-nat.
Gesellsch. Jena_, November 13.

Since the printing of this Report began in 1884 and was far advanced in
1885, it was impossible to include the important works of Rüst and
Brandt (L. N. 51, 52) in the descriptive portion, so that they are only
referred to in the Introduction.

251 B. _Phaulographic Appendix_:--

A list of absolutely worthless literature, which contains either only
long known facts or false statements, and may hence be entirely
neglected with advantage. Compare § 252, and also L. N. 26, p. 9.

55. 1865. WALLICH, G. C., On the structure and affinities of Polycystina.
_Trans. Micr. Soc. Lond._, vol. xiii. pp. 57-84. (Compare L. N. 26, p.
9.)

{clxxix}56. 1879. WALLICH, G. C., Observations on the Thalassicollidæ.
_Ann. and Mag. Nat. Hist._, ser. 4, vol. iii. p. 97.

57. 1866. STUART, ALEXANDER, Ueber Coscinosphæra ciliosa, eine neue
Radiolarie (= Globigerina echinoides!!). _Zeitschr. f. wiss. Zool._, Bd.
xvi. p. 328, Taf. xviii. (Compare L. N. 26, p. 9.)

58. 1870. STUART, ALEXANDER, Neapolitanische Studien. _Göttinger Nachr._,
p. 99, and _Zeitschr. f. wiss. Zool._, Bd. xxii. p. 290 ("Blue Siliceous
Crystals" in Collozoum inerme!).

59. 1871. MACDONALD, JOHN DENIS, Remarks on the Structure of Polycystina
(Astromma Yelvertoni = Euchitonia Mülleri). _Ann. and Mag. Nat. Hist._,
ser. 4, vol. viii. p. 226.

60. 1871. DOENITZ, W., Beobachtungen über Radiolarien. _Archiv f. Anat.
u. Physiol._, 1871, p. 71, Taf. ii. (Compare L. N. 26, p. 7.)

252. _Progress of our Knowledge of the Radiolaria from_ 1862 _to_
1885.--The history of our scientific knowledge of the Radiolaria extends
over about half a century (from 1834 to 1885). A historical and critical
discussion of the works which appeared within the first twenty-eight years
of this period (from 1834 to 1862) is contained in the historical
introduction to my Monograph (L. N. 16, pp. 1-24); I shall therefore
give here only a brief survey of the investigations published during
the last twenty-three years (from 1862 to 1885). The most important
steps in our progress during this period we owe to the following
naturalists:--Cienkowski (1871), Ehrenberg (1872 and 1875), Richard Hertwig
(1876 and 1879), Karl Brandt (1881 and 1885), Bütschli (1882), and Rüst
(1885). To the valuable works of these authors must be added a number of
smaller contributions, which are recorded in the foregoing Bibliography.
Some communications from dilettanti, written with insufficient knowledge of
the subject, and hence of no value, are mentioned for the sake of
completeness in the "Phaulographic Appendix" (compare L. N. 55-60, also L.
N. 26, p. 9).

The first important advance in our knowledge of the organisation of the
Radiolaria, made after the publication of my Monograph (1862), was the
demonstration of the nature of the extracapsular "yellow cells." In the
year 1870 I showed that these yellow cells contain starch (L. N. 21, p.
519). I regarded them, as did all authors up to that time, as integral
parts of the Radiolarian organism, and hence considered this to be
multicellular; for no doubt was possible regarding the true cellular nature
of these remarkable, nucleated, yellow globules, which I had thoroughly
studied in 1862. It was first shown by Cienkowski in 1871 that the yellow
cells of the #Collodaria# remain unchanged even after the death of these
organisms, "that they continue to grow uninterruptedly, and eventually
multiply by division" (L. N. 22, pp. 378-380, Taf. xix. figs. 30-36).
Cienkowski concluded from these important observations that the yellow
cells are not integral parts of the Radiolarian body, but "parasitic
structures," independent, unicellular organisms, which live only as
parasites in the body of the Radiolaria (compare § 90).

This important recognition underwent ten years later a further development
and complete establishment by the extensive investigations of Karl Brandt
(L. N. 38, 39) {clxxx}and Patrick Geddes (L. N. 42, 43). This arrangement
was compared by Brandt to the remarkable symbiosis of the Algoid gonidia
and Fungoid hyphæ in the organisation of the Lichens, which had been
recently discovered, and since he recognised the independent nature of the
yellow cells, as unicellular Algæ, in all divisions of the Radiolaria, he
founded for them the genus _Zooxanthella_. Geddes named them _Philozoon_,
and showed experimentally that they give out oxygen under the influence of
sunlight (compare § 90). The great physiological importance of the yellow
cells in the metastasis of the Radiolaria, and, when they are developed in
large quantities, in the economy of marine organisms in general, has
recently been insisted upon by Brandt (see § 205 and L. N. 52, pp. 65-71,
86-94).

The proof that the yellow cells do not belong to the Radiolarian organism
itself, but only live parasitically in it, was a necessary preliminary to
the very important step which next took place in our knowledge of the
organisation of the Radiolaria. This step consisted in the demonstration
that the whole body of the Radiolaria, like that of all other Protista, is
only a single cell. It was Richard Hertwig who in two remarkable works (L.
N. 26, 33) firmly established this fundamental theorem of the unicellular
nature of the Radiolaria. In his treatise on the histology of the
Radiolaria (L. N. 26, 1876) he published complete investigations into the
structure and development of the Sphærozoida and Thalassicollida. Since he
made use of the modern methods of histological examination, and especially
of staining fluids, which he was the first to apply to the study of the
Radiolaria, he was able to show that no true cells (apart from the
parasitic yellow cells) are to be found in their bodies, but rather that
all their morphological components are to be regarded as differentiated
parts of a single true cell, and in particular that the central capsule
includes a genuine nucleus.

A wider foundation for this important discovery and its applicability to
all divisions of this extensive class, was given by Hertwig in a second
work on the organisation of the Radiolaria (L. N. 33, 1879). Among the
numerous discoveries by which this work enriched the natural history of the
Radiolaria must be specially mentioned the recognition of the fundamental
differences exhibited by the main divisions of the class in the structure
of their central capsule. Hertwig first observed that the capsular membrane
is double in the PHÆODARIA but single in the other Radiolaria (§ 56); the
former he named "TRIPYLEA" because he discovered in their capsular membrane
a large, peculiarly constructed main opening and two small accessory
openings. The NASSELLARIA, in which he found a single porous area at the
basal pole of the main axis, with a cone of pseudopodia rising from it, he
called on this account "MONOPYLEA"; whilst the other Radiolaria, whose
capsular membrane is perforated on all sides with fine pores, were termed
"PERIPYLEA." Besides the central capsule, Hertwig laid stress upon the
significance of the gelatinous envelope as a constant and important
constituent of the body. He also devoted attentive consideration to the
morphology of the skeleton, and on the basis of certain
{clxxxi}phylogenetic conclusions which he drew from it, he arrived at an
improved systematic arrangement in which he distinguished six orders:--(1)
#Thalassicollea#, (2) #Sphærozoea#, (3) #Peripylea#, (4) #Acanthometrea#,
(5) #Monopylea#, (6) #Tripylea#. The numerous isolated discoveries with
which Hertwig enriched the morphology of the Radiolaria, have been already
alluded to in the appropriate paragraphs in the anatomical portion of this
Introduction (see L. N. 42, pp. 340, 341).

The new and interesting group, which was thus erected into an order under
the name TRIPYLEA, I had already a year previously separated from the other
Radiolaria as "_Pansolenia_" in my Protistenreich (L. N. 32, p. 102).
Since, however, neither the three capsular openings of the TRIPYLEA nor the
skeletal tubes of the Pansolenia are present in all the families of this
extensive order, I substituted in 1879 the more suitable name PHÆODARIA,
which is applicable to all members of the group (L. N. 34). In the
preliminary memoir then published regarding the Phæodaria, a New Group of
Siliceous Marine Rhizopods, I distinguished four orders, ten families, and
thirty-eight genera. The great majority of these new forms (among which
were no less than 465 different species) were first discovered by the
deep-sea investigations of the Challenger. John Murray was the first who
called attention to the great abundance in the deep sea of these remarkable
Rhizopods, and to the constant presence of their peculiar, dark,
extracapsular pigment body (phæodium); even in 1876 he described a portion
of them as Challengerida (L. N. 27, p. 536; L. N. 53, p. 226). The earliest
observations on the PHÆODARIA were made at Messina in 1859, where I
examined five genera of this remarkable group alive (compare p. 1522 and L.
N. 16).

By the discovery that the PHÆODARIA, although differing in important
respects from the other Radiolaria, still conform to the definition of the
class, a new and extensive series of forms was added to this latter, and by
their closer investigation a fresh source of interesting morphological
problems was disclosed. In other groups, however, morphology was advanced
by comparative anatomical studies. In addition to the smaller contributions
of various authors, mentioned in the foregoing bibliography, I may
specially refer to the valuable Beiträge zur Kenntniss der
Radiolarien-Skelete, insbesondere der der Cyrtida by O. Bütschli (L. N. 40,
1882). On the basis of careful comparative anatomical studies,
investigations into the skeletal structure of a number of fossil
#Cyrtoidea# and critical application of the recently published researches
of Ehrenberg into the Polycystina of Barbados (L. N. 25), Bütschli
attempted to derive the complicated relations of the Monopylean skeletons
phylogenetically from a simple primitive form,--the primary sagittal ring.
Even if this attempt did not actually solve the very difficult
morphological problem in question, still the critical and synthetic mode in
which it was carried out deserves full recognition, and furnishes the proof
that the comparative anatomy of the skeleton in the Radiolaria not less
than in the Vertebrata, is a most interesting and fruitful field of
phylogenetic investigation. A {clxxxii}further demonstration of this was
furnished by Bütschli in the general account of the organisation of the
Radiolaria which he published in 1882 in Bronn's Klassen und Ordnungen des
Thierreichs (L. N. 41).

In our knowledge of the developmental history of these Protista the last
two decades have witnessed less progress than in their comparative anatomy.
The most important advance in this direction has been the proof that in all
the main groups of the class the contents of the central capsule are used
in the formation of swarm-spores. The movements of these zoospores in the
central capsule had indeed been observed by several previous authors in the
case of the SPUMELLARIA and ACANTHARIA (L. N. 10, 13, 16; compare also §
142, Note A). The origin of the flagellate spores from the contents of the
central capsule and their peculiar constitution were, however, first
described fully by Cienkowski in 1871 (L. N. 22, p. 372). Soon after this,
R. Hertwig discovered that in the social Radiolaria (Polycyttaria or
Sphærozoea) two different forms of zoospores are formed, one with, the
other without crystals, and that the latter are also divided into
macrospores and microspores (compare L. N. 26, and § 142). Recently this
sexual differentiation has been shown by Karl Brandt to exist in all the
groups of Sphærozoea, and its regular interchange with the formation of
crystal-spores has been interpreted as a true "alternation of generations"
(compare L. N. 52 and also § 216). The other forms of development also,
especially reproduction by cell-division (§ 213) and gemmation (§ 214),
have been elucidated by the recent investigations of the same author.

The palæontology of the Radiolaria has of late made important and
interesting advances. Until ten years ago fossil remains of this class were
known exclusively from the Tertiary period; almost the only source of our
information was to be found in the researches of Ehrenberg, commenced in
1838, continued in his Mikrogeologie in 1854, and concluded in his last
work (L. N. 25) published in 1875 (compare L. N. 16, pp. 3-9, 191-193). In
the year 1876 a number of Mesozoic Radiolaria from the chalk were described
by Zittel (L. N. 28), and afterwards others from the Jura by Dunikowski (L.
N. 44). That fossil Radiolaria occur in Mesozoic formations, especially in
the Jura, as well preserved and as abundantly as in the Tertiary rocks of
Barbados, was shown in 1883 by Rüst (L. N. 48). By the examination of
numerous thin sections he discovered that in all the main divisions of the
Jurassic formation (Lias, Dogger, Malm) there are distributed jaspers,
flints, cherts, and other quartzites, which consist largely of the
siliceous shells of Polycystina; the same is true also of many Coprolites
found in the Jura. The full account of these and the descriptions and
figures of 234 Jurassic species, distributed in 76 genera, are contained in
the Beiträge zur Kenntniss der fossilen Radiolarien aus Gesteinen des Jura
(L. N. 51, 1885). But even in the older rocks, the Trias, the Permian, and
Carboniferous systems, and even as far downwards as the Silurian and
Cambrian formations, Rüst has recently shown the existence of fossil
Radiolaria, {clxxxiii}and thus increased the known period of the
developmental history of the class by many millions of years (§ 244).

The great significance of the Radiolaria in geology and palæontology has
been brought into new light not only by these extensive discoveries, but
also by the important relations which have been shown to exist between the
Radiolarian rocks and the deep-sea deposits of the present day. In this
direction the wonderful discoveries of the Challenger, and especially the
investigation of the deep-sea deposits by Wyville Thomson (L. N. 31) and
John Murray (L. N. 27), have furnished us with new and valuable information
(compare §§ 236-239, and §§ 245-250). The Tertiary Polycystine formations
of Barbados and the Nicobar Islands, with which we have been acquainted for
the last forty years, as also the Mesozoic Radiolarian quartzes, which have
only recently been made known to us from the Jura, are ascertained to be
fossil representatives of the same deep-sea deposits which now occur in the
form of Radiolarian ooze (§ 237), and to some extent also of Globigerina
ooze and red clay (§§ 238, 239), on the bottom of the ocean, at depths of
from 2000 to 4500 fathoms.

These investigations into fossil Radiolaria and their comparison with
recent deep-sea forms have a further general significance, inasmuch as the
identity of many living and fossil species from the Tertiary formation has
been shown beyond all doubt. In this direction the numerous measurements
and accurate comparisons which I have made during the last ten years of the
abyssal forms in the Challenger collection, and of fossil species from
Barbados and Caltanisetta, have brought to light many important facts. In
this I had the able assistance of my friend, Dr. Reinhold Teuscher (compare
§ 250, and p. 1760). Further valuable contributions in this direction are
found in the careful observations and comparative measurements recently
published by Emil Stöhr (L. N. 35, 1880), regarding the Radiolarian fauna
of the Tripoli of Grotte in the province of Girgenti, Sicily. From these it
appears that the number of Miocene species which are still extant, is much
greater than would appear from the results of Ehrenberg.

Ehrenberg himself, towards the end of his long and laborious life,
collected the results of the systematic and palæontological researches,
which he had begun thirty-seven years previously (L. N. 16, pp. 3-12) into
the Polycystina, in two large works (L. N. 24, 25). The first treatise (L.
N. 24, 1872) contains the Mikrogeologische Studien über das Kleinste Leben
der Meeres-Tiefgründe aller Zonen und dessen geologischen Einfluss, with a
list of 279 Polycystina observed by him from the deep-sea, as well as
figures of 127 species. The second work (L. N. 25, 1875) contains the
Fortsetzung der Mikrogeologischen Studien, mit specieller Rücksicht auf den
Polycystinen-Mergel von Barbados; the list of fossil Polycystina observed
by him includes 325 species, of which 26 are still extant; 282 of them are
figured on the thirty plates accompanying the memoir. By means of these
numerous figures, as well as by the appended systematic and chorological
tables, Ehrenberg furnished a welcome {clxxxiv}supplement to the numerous
communications regarding the Polycystina, which he had made to the Berlin
Academy since 1838, and which he had published in his Mikrogeologie in
1854. It will always be the merit of this zealous and indefatigable
microscopist that he first called attention to the great wealth of forms
existing in this class; he separated systematically about 500 species, and
published drawings of about 400; in addition to which he was the first to
lay stress upon the great chorological and geological importance of the
Radiolaria.

With these systematic and descriptive, chorological and palæontological
works, however, which relate exclusively to the Polycystina, the merits of
the famous naturalist of Berlin are exhausted as regards this class of
animals. Of the organisation of the Radiolaria, Gottfried Ehrenberg
remained entirely ignorant up till his death in 1876. All that a number of
famous naturalists had observed during a quarter of a century as to the
structure and life-history of the Radiolaria, all the important discoveries
of Huxley (1851), Johannes Müller (1858), Claparède (1858), Cienkowski
(1871), and many others (L. N. 1-22), and all that I had published in my
Monograph (1862) on the basis of three years' study of their anatomy and
physiology--all this Ehrenberg ignored, or rather, he regarded it all as
worthless rubbish of science, as a chaos of devious errors, resting upon
incomplete observations and false conclusions. His strange "special
considerations regarding the Polycystina" (L. N. 24, pp. 339-346) and the
general "concluding remarks" (L. N. 25, pp. 146-147) leave no room for
doubt on this point. Ehrenberg indeed doubted to the last whether any
observer had seen living Radiolaria at all (L. N. 25, p. 108).

The invincible obstinacy with which Ehrenberg maintained his preconceived
opinion of the high organisation of the Radiolaria, and entirely ignored
the contrary observations of other naturalists, is explained by the
consistency with which he held to the end the "principle peculiar to
himself of the universally equal development of the animal kingdom" (L. N.
16, p. 7). From the complicated arrangement of their siliceous shells he
concluded that the animals inhabiting them must possess a structure
correspondingly complex, and nearly related to that of the Echinodermata
(Holothuria). Like all other animals the Radiolaria must possess systems of
organs for locomotion, sensation, nutrition, circulation, and reproduction.
Whilst Ehrenberg originally interpreted the Polycystina as siliceous
Infusoria polygastrica, and regarded them as compound Arcellina, he
afterwards classed them sometimes with the Echinodermata (Holothuria),
sometimes with the Bryozoa, sometimes with the Oscillaria (see L. N. 41, p.
336). Although a decided opponent of the cell-theory he called them
"multicellular animalcules" (Polycystina), interpreting the pores of the
siliceous shell as cells. To-day the opposite term (Monocystina) might be
adopted to express their unicellular organisation. It was a remarkable
irony of fate that in the self-same year (1838) in which Schwann of Berlin
made by his foundation of the cell theory the greatest advance in the whole
{clxxxv}of Biological Science, that Ehrenberg, all his life the most
zealous opponent of that theory, published his great work on the Infusoria,
and at the same time established the "family of multicellular animalcules
or Polycystina" (L. N. 16, p. 4).

The "short systematic survey of the genera of cellular animalcules" given
by Ehrenberg in 1875 (L. N. 25, p. 157), is only a new edition, increased
by sixteen genera, of his first systematic arrangement of the Polycystina
of 1847 (L. N. 4, p. 53). Since I have already given a full discussion of
this in my Monograph (L. N. 16, pp. 214-219), I need only here remark that
a correct understanding of his very inadequate generic diagnoses is only
possible by the aid of his figures. Relying upon these I have retained
almost all Ehrenberg's genera, although entirely new definitions of most of
them have been necessary.

The same is true also of the two orders which Ehrenberg distinguished in
his class of "Zellenthierchen." The first order is constituted by his
"Netzkörbchen" (Monodictya or NASSELLARIA) formerly known as "Polycystina
solitaria"; they include our #Cyrtoidea#, the greater part of Hertwig's
Monopylea. Ehrenberg's second order is the "Schaumsternchen" (Polydictya or
SPUMELLARIA), previously called "Polycystina composita"; they include the
Peripylea of Hertwig, as well as the Spyridina (our #Spyroidea#), which
belong properly to the NASSELLARIA. Although Ehrenberg's statements
regarding the organisation of both these orders were quite erroneous, and
his knowledge even of the structure of their shells very defective, I still
thought it advisable to retain his names for the groups, since they
constituted his one successful effort in the systematic treatment of the
Radiolaria (compare L. N. 41, p. 336).

The sketch of a systematic arrangement of the Radiolaria (L. N. 37), which
I published in 1881 on the basis of the study of the Challenger Radiolaria,
resembles, in respect of seven orders being distinguished, the new system
which R. Hertwig founded in 1879, in consequence of the variations which he
discovered in the structural relations of the central capsule (L. N. 33, p.
133). It differs, however, inasmuch as his Sphærozoea (my Polycyttaria) are
here divided into two orders, Symbelaria (#Collosphærida#) and Syncollaria
(#Sphærozoida#). In that sketch too I separated for the first time the two
subclasses Holotrypasta (Porulosa) and Merotrypasta (Osculosa). The fifteen
families established by Hertwig were then raised to twenty-four. The six
hundred and thirty genera, which I then distinguished, are still for the
most part retained, some, however, in a restricted sense, or with amended
definitions.

The differential characters of the orders and families of the Radiolaria,
given in the Prodromus in 1881, were amended in a further communication
which I gave in 1883 regarding the orders of the Radiolaria (L. N. 46, p.
17). There I reduced the seven orders to four, the structural relations of
the central capsule being precisely the same in the Polycyttaria and
#Collodaria# as in the #Peripylea#. The survey of the affinities of the
class was thus rendered much simpler and clearer, and the
{clxxxvi}hypothetical genealogical tree, which I then published, has been
still further carried out in Chapter VI. of the present Introduction (see
§§ 153-200).

253. _General Survey of the Growth of our Systematic Acquaintance with the
Radiolaria from 1834 to 1885._

1834. MEYEN (L. N. 1) describes 2 genera and species of
#Collodaria#:--_Sphærozoum fuscum_ and _Physematium atlanticum_.

1838. EHRENBERG (L. N. 2) founds the family Polycystina upon 3 fossil
genera (with 6 species):--_Lithocampe_, _Cornutella_, _Haliomma_.

1847. EHRENBERG (L. N. 4) publishes his preliminary communications
regarding the fossil Polycystina of Barbados and distinguishes 282
species, distributed in 44 genera and 7 families. In the tabular view of
the genera he distinguishes two orders:--I. #Solitaria#--(1)
Halicalyptrina, (2) Lithochytrina, (3) Eucyrtidina; and II.
#Composita#--(4) Spyridina, (5) Calodictya, (6) Haliommatina, (7)
Lithocyclidina (compare L. N. 16, pp. 214-219).

1851. HUXLEY (L. N. 5) gives the first accurate account of living
Radiolaria, and describes 2 species of the genus _Thalassicolla_
(_nucleata_ and _punctata_); under the latter are included 4 genera of
#Sphærozoea#:--_Collozoum_, _Sphærozoum_, _Collosphæra_, _Siphonosphæra_
(compare L. N. 16, pp. 12-14).

1854. EHRENBERG (L. N. 6) publishes in his Mikrogeologie, figures of
seventy-two species of fossil Polycystina (without descriptions).

1855. JOHANNES MÜLLER (L. N. 8, p. 248) describes the first
_Acanthometra_, and elucidates its affinity to Huxley's _Thalassicolla_
and Ehrenberg's Polycystina.

1858. JOHANNES MÜLLER (L. N. 12) establishes the new group Radiolaria as
a special order of the Rhizopoda, and includes in it the Thalassicolla,
Polycystina, and Acanthometra as closely related families. He opposes
these radiate Rhizopoda to the Polythalamia, and describes 50 species
observed by him living in the Mediterranean, these he arranges in 20
genera, of which 10 are new. The figures are contained in eleven plates
(see L. N. 16, pp. 22-24).

1858. CLAPARÈDE (L. N. 14) describes the first #Plectoidean#
(_Plagiacantha arachnoides_) and two species of _Acanthometra_, which he
had observed living in Norway (see L. N. 16, p. 18).

1860. EHRENBERG (L. N. 4) gives a short diagnosis of 22 new genera of
Polycystina, based on the investigation of numerous deep-sea species
brought up by Brooke from the depths of the Pacific Ocean. The number of
his genera is thus increased to 66 (compare L. N. 16, pp. 10, 11).

1862. ERNST HAECKEL (L. N. 16) embraces in his Monograph of the
Radiolaria all the species hitherto known either by figures or
descriptions, and arranges them in 15 families and 113 genera; of which
latter 46 are new. The number of new species observed living amounts to
144. In a "survey of the Radiolarian fauna of Messina" (p. 565) he
records 72 genera and 169 species. Most of these are figured in the
accompanying atlas of thirty-five plates.

{clxxxvii}1862. BURY (L. N. 17) gives in an atlas of twenty-five plates,
photographed from drawings, the figures of numerous fossil Polycystina of
Barbados (without descriptions), of which many are new species overlooked
by Ehrenberg (compare § 242, above).

1872. EHRENBERG (L. N. 24) gives a list of names (without description) of
all the Polycystina observed by him from the bottom of the sea, 279
species, of which 127 are figured on twelve plates.

1875. EHRENBERG (L. N. 25) gives a list of names of all the fossil
Polycystina observed by him (from Barbados, the Nicobar Islands and
Sicily), 326 species, of which 282 are figured (compare § 242 above). In
a new "Systematic Survey of the Genera" the number of these is given as
63. The 7 families are the same as given in 1847 (see above), as also the
two orders (NASSELLARIA = Solitaria, SPUMELLARIA = Composita).

1876. ZITTEL (L. N. 29) describes the first fossil Radiolaria from the
chalk (6 species) and establishes the new Cyrtoid genus _Dictyomitra_.

1876. JOHN MURRAY (L. N. 27) establishes the new family Challengerida,
and figures 6 new generic types of PHÆODARIA.

1879. RICHARD HERTWIG (L. N. 33) first describes the fundamental
differences in the structure of the central capsule, and in accordance
with them divides the Radiolaria into six orders:--(1) Thalassicollea,
(2) Sphærozoea, (3) Peripylea, (4) Acanthometrea, (5) Monopylea, (6)
Tripylea (p. 133). These are subdivided into 18 families, and their
phylogenetic affinities discussed (p. 137). On the ten plates, several
new species from Messina are figured, among them the types of several new
genera (_Cystidium_, _Coelacantha_, _Echinosphæra_) (compare § 252).

1879. ERNST HAECKEL (L. N. 34) founds the order PHÆODARIA as a "new group
of marine siliceous Rhizopods," and distinguishes in it 4 suborders, 10
families and 38 genera.

1880. EMIL STÖHR (L. N. 35) describes the Miocene "Radiolarian fauna of
the tripoli from Grotte in Sicily," 118 species, of which 78 are new;
among them is the new genus _Ommatodiscus_, the type of a new family,
Ommatodiscida. The new species are figured on seven plates.

1880. DANTE PANTANELLI (L. N. 36) describes 30 species of fossil
Polycystina from the jaspers of Tuscany, which he regarded as Eocene, but
which were probably of Jurassic origin (compare § 243, note B, above).

1881. ERNST HAECKEL (L. N. 37) publishes a "Sketch of a classification of
the Radiolaria on the basis of the study of the Challenger Collection,"
and distinguishes in his "conspectus ordinum" (p. 421) 2 subclasses and 7
orders, and in the "prodromus systematis Radiolarium" (pp. 423-472) 24
families with 630 genera, among which are more than 2000 new species.

1882. BÜTSCHLI (L. N. 40) on the basis of studies of the fossil Monopylea
of Barbados, investigates the "mutual relations of the Acanthodesmida,
Zygocyrtida and Cyrtida," and gives a critical revision of the genera of
these "Cricoidea;" a number of new species are described and figured
(Tafs. xxxii., xxxiii.), and some new genera of Stichocyrtida established
(_Lithostrobus_, _Lithomitra_, &c.).

1882. DUNIKOWSKI (L. N. 44) describes 18 new fossil Polycystina from the
lower lias of the Salzburg Alps, among them the types of 3 new genera
(_Ellipsoxiphus_, _Triactinosphæra_, and _Spongocyrtis_).

{clxxxviii}1883. ERNST HAECKEL (L. N. 46) revises the 4 orders and 32
families of Radiolaria, and gives more accurate definitions of them, as
well as of the 2 subclasses (I. _Holotrypasta_ = ACANTHARIA and
SPUMELLARIA; II. _Merotrypasta_ = NASSELLARIA and PHÆODARIA).

1885. D. RÜST (L. N. 51) describes 234 new species of fossil Radiolaria
from the Jura, and illustrates them by twenty plates. Among them are 103
SPUMELLARIA, 130 NASSELLARIA, and 1 PHÆODARIA; these are contained in 35
genera, of which 20 belong to the Porulosa, and 15 to the Osculosa.

254. _Statistical Synopsis of the Twenty Orders_:--

(a) Number of Families.
(b) Number of Genera.
(c) Number of Species.
(d) Previously known Species.
(e) New Species.
(f) Fossil Species.
(g) Pelagic Abundance.
(h) Abyssal Abundance.
(i) Figured on Plates

+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | | | | | | | | Figured on |
| (a) | (b) | (c) | (d) | (e) | (f) | (g) | (h) | Plates. |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| I. Legion SPUMELLARIA (Porulosa peripylea) |
+-----+---------------------------------------------------------------+
| | I. COLLODARIA (Spumellaria palliata) |
+-----+-----+---------------------------------------------------------+
| | 1. Colloidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 2 | 6 | 36 | 9 | 27 | 0 | A | E | 1, 3 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 2. Beloidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 2 | 8 | 56 | 9 | 47 | 0 | A | D | 2, 4 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | II. SPHÆRELLARIA (Spumellaria loricata) |
+-----+-----+---------------------------------------------------------+
| | 3. Sphæroidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 6 | 107 | 660 | 105 | 555 | 66 | A | B | { 5-8 |
| | | | | | | | | { 11-30 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 4. Prunoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 7 | 53 | 280 | 35 | 245 | 36 | B | B | { 16, 17 |
| | | | | | | | | { 39, 40 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 5. Discoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 6 | 91 | 503 | 126 | 376 | 102 | B | A | { 31-38 |
| | | | | | | | | { 41-48 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 6. Larcoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 9 | 51 | 260 | 8 | 252 | 0 | E | B | { 9, 10 |
| | | | | | | | | { 49, 50 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| II. Legion ACANTHARIA (Porulosa actipylea) |
+-----+---------------------------------------------------------------+
| | III. ACANTHOMETRA (Acantharia palliata) |
+-----+-----+---------------------------------------------------------+
| | 7. Actinelida |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 6 | 22 | 6 | 16 | 0 | E | E | 129 (figs. 1-3) |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 8. Acanthonida |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 21 | 138 | 50 | 88 | 0 | A | C | 130-132 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | IV. ACANTHOPHRACTA (Acantharia loricata) |
+-----+-----+---------------------------------------------------------+
| | 9. Sphærophracta |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 27 | 149 | 9 | 140 | 0 | B | B | 133-138 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 10. Prunophracta |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 11 | 63 | 5 | 58 | 0 | D | B | 139, 140 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| III. Legion NASSELLARIA (Osculosa monopylea) |
+-----+---------------------------------------------------------------+
| | V. PLECTELLARIA (Nassellaria palliata) |
+-----+-----+---------------------------------------------------------+
| | 11. Nassoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 1 | 2 | 5 | 1 | 4 | 0 | E | E | 91 (fig. 1) |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 12. Plectoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 2 | 17 | 61 | 5 | 56 | 0 | D | C | 91 (figs. 2-12) |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 13. Stephoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 4 | 40 | 205 | 14 | 191 | 17 | C | B | { 81, 82 |
| | | | | | | | | { 92-94 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | VI. CYRTELLARIA (Nassellaria loricata) |
+-----+-----+---------------------------------------------------------+
| | 14. Spyroidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 4 | 45 | 239 | 51 | 188 | 53 | C | A | 83-90 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 15. Botryodea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 10 | 55 | 15 | 40 | 10 | E | C | 96 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 16. Cyrtoidea |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 12 | 160 | 1122 | 328 | 794 |250 | C | A | 51-80 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| IV. Legion PHÆODARIA (Osculosa cannopylea) |
+-----+---------------------------------------------------------------+
| | VII. PHÆOCYSTINA (Phæodaria palliata) |
+-----+-----+---------------------------------------------------------+
| | 17. Phæocystina |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 15 | 112 | 30 | 82 | 24 | C | B | 101-105 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | VIII. PHÆOCOSCINA (Phæodaria loricata) |
+-----+-----+---------------------------------------------------------+
| | 18. Phæosphæria |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 4 | 22 | 121 | 5 | 116 | 0 | C | A | 106-112 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 19. Phæogromia |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 5 | 27 | 159 | 5 | 154 | 0 | C | A | { 99, 100 |
| | | | | | | | | { 113-120 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| | 20. Phæoconchia |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 3 | 20 | 73 | 4 | 69 | 0 | D | B | 121-128 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| Total, |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+
| 85 | 739 | 4318 | 810 | 3508 | 558 | ... | ... | 140 |
+-----+-----+------+-----+------+-----+-----+-----+-------------------+

_Note._--In the tenth and eleventh columns the relative abundance of each
order at or near the surface and near the bottom is approximately indicated
by the letters A-E, which have the following significance:--A, abundant; B,
very numerous; C, many (medium quantity); D, few; E, very few.

{1}SYSTEMATIC PART.

----

CLASS RADIOLARIA.

RADIOLARIA, Johannes Müller, 1858.
RHIZOPODA RADIARIA, Johannes Müller, 1858.
POLYCYSTINA (_pro parte_), Ehrenberg, 1838.
ECHINOCYSTIDA, Claparède, 1858.
RHIZOPODA CAPSULARIA, Haeckel, 1861.
CYTOPHORA, Haeckel, 1862.

_Definition of the Class:_--#Rhizopoda with unicellular body, divided by a
porous membrane into an internal or intracapsular part (with nucleus), and
an external or extracapsular part (with calymma); propagating by
flagellated spores.#

The RADIOLARIA or CAPSULATE RHIZOPODA, first constituted by Johannes Müller
in the year 1858 as a separate group of the Rhizopoda, form a peculiar
class of the PROTISTA, or unicellular organisms. This class is exclusively
marine, and has in general the characteristic organisation of the
Rhizopoda, with the development of numerous _pseudopodia_ from the surface
of the cell; but it differs from all other Rhizopoda in the possession of a
peculiar _membrane_, dividing the cell-body into two different parts; the
_central capsule_ or the internal part with the _nucleus_, and the external
part or _extracapsulum_ with the _calymma_; propagation by flagellated
spores produced in the central capsule; the sarcode or the _protoplasm_ of
both parts communicates by fine _pores_, piercing the separating membrane,
which is called the _capsule-membrane_.

The _Central Capsule_ or the inner part of the Radiolarian body is
constantly composed of three essential parts, viz.:--

1. _The Central Nucleus_ (a true cell-nucleus).

2. _The Intracapsular Sarcode_ (endosarc) or surrounding internal
protoplasm.

3. _The Capsule Membrane_ or enveloping porous membrane.

{2}Besides these constant and essential elements, the central capsule
contains very commonly (but not constantly) some other enclosed structures,
viz.:--

4. An internal or intracapsular skeleton.

5. Intracapsular vacuoles or alveoli.

6. Fat-granules or oil-globules.

7. Crystals of different composition.

8. Pigment-granules.

_The Extracapsulum_, or the outer part of the Radiolarian body is also
constantly composed of three essential elements,--

1. _The Calymma_, or the thick extracapsular _jelly-veil_, completely
enveloping the whole central capsule.

2. _The Matrix_, or the maternal tissue of the external protoplasm,
enveloping immediately the capsule-membrane as a thin continuous layer of
_extracapsular sarcode_ (ectosarc).

3. _The Pseudopodia_, or the very numerous thread-like filaments of
protoplasm, which radiate from the matrix; whilst their inner part is
enclosed in the calymma, their outer part floats freely in the sea-water.

Besides these three constant and essential elements, the extracapsulum
contains very commonly (but not constantly) some other enclosed structures,
viz.:--

4. An external or extracapsular skeleton.

5. Extracapsular vacuoles or alveoli.

6. Fat-granules or oil-globules.

7. Pigment-granules or a peculiar large body of dark extracapsular
pigment, the "phæodium."

8. "Xanthellæ" or "zooxanthellæ," peculiar yellow cells, which contain
starch and are unicellular yellow Algæ, living as "Symbiontes" in true
Symbiosis with a great many Radiolaria.

_The Nucleus_ of the Radiolaria is a large true simple cell-nucleus,
originally a solid spherical, roundish or longish body of nuclein. It is
placed either in the centre of the capsule (in most Peripylea) or
excentrically (in most other Radiolaria). Originally solid, the nucleus is
commonly differentiated later into an outer dense nuclear-membrane and an
inner softer or fluid content; either with one single nucleolus or with a
variable number of nucleoli. Originally always simple, the nucleus becomes
afterwards constantly divided into numerous small nuclei, each of which,
together with a part of the surrounding {3}protoplasm, forms a
vibratile-spore or "flagellate-spore." This division in the Acantharia and
in the social (or colonial) Peripylea begins very early, in all other
Radiolaria much later, immediately before propagation.

_The Endoplasm_ or "endosarc," or "intracapsular protoplasm" or "inner
sarcode," in all Radiolaria originally fills that space within the capsule,
which is not taken up by the nucleus. It seems to be employed mainly for
the purpose of propagation, becoming divided earlier or later into numerous
small particles, each of which surrounds a small particle of the nucleus
and forms together with it a flagellate-spore. Besides this the endoplasm
of the Radiolaria seems to have a great significance for the nutrition,
mainly for the interchange of materials. It becomes very often vacuolate or
alveolate, filled with smaller or larger spherical drops of fluid; it
produces very commonly smaller fat-granules or larger oil-globules, and
further pigment-granules of different colours, more rarely crystals and
other peculiar enclosed parts.

_The Membrane_ or "capsule-membrane" is the most typical and characteristic
part of the body of a Radiolarian, sufficient of itself to separate this
class from all other Rhizopoda. At the same time, by its different shape it
presents the best means for the systematic distinction of the four
subclasses or "legions" of the class. The membrane is composed of a special
organic matter (probably nearly related to chitin) and combines density
with elasticity to a high degree. Observed with a high power of the
microscope its margin (or section) appears commonly simple-edged, but often
in larger forms distinctly double-edged.

The legion PHÆODARIA is distinguished by a double membrane (the thinner
inner and thicker outer membranes being separated by an interval); in the
three other legions it is simple. The membrane completely separates the
intracapsular from the extracapsular body, both communicating only by
certain pores or openings in the membrane. With reference to this important
communication, the whole class can be divided into two subclasses,
Holotrypasta and Merotrypasta: the HOLOTRYPASTA contain the Peripylea and
Actipylea, in which the membrane is pierced by innumerable very small
pores; the MEROTRYPASTA consist of the Monopylea and the Cannopylea, in
which the membrane exhibits only one large main opening, distinguished in
the former by a peculiar "porous area," in the latter by an "osculum" or a
prolonged tubule.

_The Calymma_ or "jelly-veil" is the most characteristic part of the
extracapsular body in all Radiolaria; in the majority of the class it is
the most voluminous part of the whole body, being much more voluminous than
all the other parts taken together. The calymma is a structureless, clear,
and transparent jelly-envelope which always includes the whole central
capsule and often also the whole extracapsular skeleton. Owing to the high
degree of its consistence, this jelly-veil takes a very important part in
the formation of the extracapsular skeleton, furnishing the matrix for the
deposition of its tangential parts.

{4}_The Matrix_ or the "maternal tissue of the pseudopodia" is formed in
all Radiolaria by the thin layer of exoplasm or of extracapsular sarcode,
which immediately envelops the central capsule and is itself enclosed by
the calymma. This continuous sarcode-cover of the capsule communicates by
its pores or openings with the endoplasm or the intracapsular sarcode;
whilst from its outer surface arise the pseudopodia. The morphological
signification of the matrix is very small, but the physiological importance
is very great, for it seems to be the chief organ of many vital functions.

_The Pseudopodia_ or the very fine, long, thread-like filaments of exoplasm
arise in all Radiolaria in very great numbers from the surface of the
matrix, and exhibit in general the same characteristic shape as in the
other Rhizopoda. Their inner or proximal part is enclosed within the
jelly-veil or calymma, whilst their outer or distal part floats freely in
the sea-water. Their special motions and modifications exhibit considerable
variations in different groups, their tendency to ramify, anastomose, and
form networks being in some cases very small, in others very great. Also
the characteristic motion of granules in the pseudopodia is very different.
In general those most important exoplasmatic filaments serve as organs both
for the vegetative functions of nutrition, and for the animal functions of
motion and sensation.

_The class_ Radiolaria can be divided according to its varying structure
into four different legions or subclasses, the characters of which are the
following:--

I. PERIPYLEA OR SPUMELLARIA.

Membrane of the central capsule simple, perforated by innumerable very
fine pores. Fundamental form originally homaxon or spherical. Skeleton
wanting or siliceous. No phæodium in the extracapsular calymma. The
Peripylea comprise two orders:--

A. COLLODARIA (without lattice-shell).
B. SPHÆRELLARIA (with lattice-shell).

II. ACTIPYLEA OR ACANTHARIA.

Membrane of the central capsule simple, perforated by innumerable fine
pores. Fundamental form originally homaxon or spherical. Skeleton
acanthinic (not siliceous). No phæodium in the extracapsular calymma. The
Actipylea consist of two orders:--

A. ACANTHOMETRA (without complete lattice-shell).
B. ACANTHOPHRACTA (with complete lattice-shell).

{5}III. MONOPYLEA OR NASSELLARIA.

Membrane of the central capsule simple, perforated by a porous-area, or
by one single large opening, divided into numerous very fine pores.
Fundamental form originally monaxon or egg-shaped. Skeleton siliceous. No
phæodium in the extracapsular calymma. The Monopylea comprise two
orders:--

A. PLECTELLARIA (without complete lattice-shell).
B. CYRTELLARIA (with complete lattice-shell).

IV. CANNOPYLEA OR PHÆODARIA.

Membrane of the central capsule double, perforated by one simple
main-opening, prolonged into a tubulus, and besides this commonly by one
or two (rarely more) small accessory openings. Fundamental form
originally monaxon or egg-shaped. Skeleton siliceous. Constantly a
peculiar dark pigment-body or "phæodium" in the extracapsular calymma.
The Cannopylea comprise two orders:--

A. PHÆOCYSTINA (without lattice-shell).
B. PHÆOCOSCINA (with lattice-shell).

_Synopsis of the four Subclasses or Legions of Radiolaria._

{6}LEGION I. #SPUMELLARIA#,

VEL PERIPYLEA, VEL PERIPYLARIA (PLS. 1-50).

_Spumellaria_ (_exclusis_ Spyridinis), Ehrenberg, 1875.
_Peripylea_ (_inclusis_ Thalassicollis et Sphærozois), Hertwig, 1879.
_Peripylaria_ (_inclusis_ Collodariis et Polycyttariis), Haeckel, 1881.

_Definition._--Radiolaria with simple membrane of the central capsule,
which is everywhere perforated by innumerable very fine pores.
Extracapsulum without phæodium. Skeleton wanting or siliceous. Fundamental
form originally spherical.

The legion SPUMELLARIA vel PERIPYLEA, in the extent here defined, was
constituted by me in 1883 in my paper on Die Ordnungen der Radiolarien.[2]
I propose to retain for this legion either the name SPUMELLARIA of
Ehrenberg (1875) or PERIPYLEA of Hertwig (1879), although both groups have
not quite the same extension. We exclude from the SPUMELLARIA the Spyridina
(united with them by Ehrenberg) and include the Collodaria. With the
Peripylea of Hertwig we unite his Thalassicollea and Sphærozoea. To avoid
any confusion it would perhaps be better to name this legion "Peripylaria."

The SPUMELLARIA agree with the ACANTHARIA in the structure of the simple
capsule-membrane, which is perforated by numerous small pores (but devoid
of the large main opening, which the NASSELLARIA and PHÆODARIA possess),
whence we unite both the former as Holotrypasta, both the latter as
Merotrypasta.

The difference between the two legions of Holotrypasta is determined by the
skeleton, which in the SPUMELLARIA is either siliceous or wanting, whilst
in the Acantharia it consists of the peculiar organic substance, acanthin.

The legion SPUMELLARIA is by far the largest and most important of the four
legions of Radiolaria, as well with respect to the number of different
forms, as to the enormous masses of individuals, which we encounter living
and fossil. We distinguish in this legion not less than thirty-two
different families, three hundred and sixteen genera, and more than
seventeen hundred species.

The classification of this large group requires for its better
comprehension a careful division into larger and smaller groups. We divide
it therefore first of all into two orders, #Collodaria# and #Sphærellaria#,
as proposed in the paper mentioned above.[3]

The #Collodaria# have no perfect latticed skeleton, and comprise two
suborders or sections: in the #Colloidea# the skeleton is entirely wanting,
in the #Beloidea# it is represented by a variable number of siliceous
needles or spicules, scattered in the calymma around the central capsule.

{7}The #Sphærellaria# differ from the #Collodaria# in the possession of a
perfect siliceous skeleton, which is originally a latticed spherical shell,
enveloping the central capsule. By modification of this fenestrated sphere
arises an enormous mass of different forms, which we dispose in
twenty-eight families, and these in four larger groups, suborders or
sections,--#Sphæroidea#, #Prunoidea#, #Discoidea#, and #Larcoidea#.

The #Sphæroidea#, the common ancestral group of the #Sphærellaria#, possess
a skeleton which is either a simple fenestrated sphere, or composed of two
or more concentric latticed spheres, which are united by radial beams; more
rarely it becomes more or less spongy.

The #Prunoidea# are derived from the #Sphæroidea# by prolongation of the
latticed sphere in one axis; the skeleton therefore becomes here
ellipsoidal or cylindrical (often with annular transversal constrictions).

The #Discoidea# on the contrary must be derived from the #Sphæroidea# by
shortening in one axis; here therefore the fenestrated shell becomes more
or less lenticular or iscoidal (often with radial spines or arms in the
equatorial plane, on the circular margin).

The #Larcoidea#, the fourth section, differ from the three foregoing
sections by the different growth of the shell in three different dimensions
of space; therefore here the fenestrated shell becomes "lentelliptical," or
a "triaxial ellipsoid," its length, breadth, and height being different.

_The Skeleton_ consists in all SPUMELLARIA either of pure _silica_ or of a
peculiar silicate. The siliceous bars and beams constituting it are
invariably _solid_, as also in the NASSELLARIA, never hollow, as in the
PHÆODARIA. Never is the skeleton composed of acanthin, as in all
ACANTHARIA. Whilst in the first order of SPUMELLARIA, #Collodaria#, the
form of the spicula, or the scattered needles, composing the skeleton, is
very simple, _never latticed_, in the second order, the #Sphærellaria#, it
is constantly latticed or fenestrated, often also spongy.

The geometrical fundamental form of the lattice-shell in the #Sphærellaria#
is originally spherical (homaxon), as preserved in all #Sphæroidea#; in the
#Prunoidea# and #Discoidea# it becomes monaxon, with one single axis
(prolonged in the former, shortened in the latter); in the #Larcoidea# it
becomes triaxon, by different growth in three principal axes, perpendicular
one to another. The further development of radial parts of the skeleton in
these three axes is very important for the "promorphology" of the
Radiolaria.

_The Malacoma_, or the whole soft body of the SPUMELLARIA as opposed to the
skeleton, exhibits some differences of structure in two different groups,
which were separated formerly (1862) as Monocyttaria and Polycyttaria,
corresponding to the "Radiolaria monozoa and polyzoa" of Johannes Müller
(1858).

The #Monocyttaria# (or the Spumellaria solitaria) live isolated as single
cells--like {8}all other Radiolaria--and are never aggregated in colonies;
the calymma includes one single central capsule, and this again one central
nucleus, which does not become divided until full maturity.

The #Polycyttaria# on the contrary (or the Spumellaria socialia) live
aggregated in large colonies; the calymma includes a variable number of
associated central capsules and each of these commonly one central
oil-globule, whilst the original simple nucleus commonly becomes very early
divided into numerous small nuclei.

_The Nucleus_ of the SPUMELLARIA is originally constantly _central_, placed
quite in the centre of the concentric capsule, and it retains this central
position in all Monocyttaria or solitary Peripylea; whereas in the
Polycyttaria--in consequence of its early division--its place is commonly
taken by a central oil-globule. Whilst the numerous nuclei of the latter
are very small, the single nucleus of the former is comparatively large,
extremely large (more than a millimeter in diameter) in some gigantic
#Collodaria#.

_The Endoplasm_ or the intracapsular sarcode exhibits in the greater number
of SPUMELLARIA a more or less distinct radial striation. It encloses a
great variety of different parts; vacuoles, oil-globules, pigment-granules,
crystals, &c.

_The Membrane_ of the capsule in all SPUMELLARIA is simple (never double as
in the _Phæodaria_) and everywhere equally perforated by innumerable small
pores; in the thick, double-edged membrane of many large #Collodaria# these
pores appear (in the optical section of the capsule-wall) as distinct fine
radial canals, very densely and regularly disposed.

_The Central Capsule_ in the SPUMELLARIA is originally a _geometrical
sphere_, and this simple globular form is preserved in all #Sphæroidea#,
and in the greatest part of #Colloidea# and #Beloidea#. By prolongation of
one axis the form becomes _ellipsoidal_ (or even cylindrical) in the
#Prunoidea#, and in some few forms of #Colloidea#. By shortening of one
axis it becomes _lenticular_ (or even discoidal) in the _Discoidea_, and in
some few forms of #Colloidea#. By unequal growth in three different axes,
perpendicular one to another, the capsule becomes _lentelliptical_ in all
#Larcoidea#. Very rarely the capsule assumes in the SPUMELLARIA a
polyhedral or irregular (sometimes even amoeboid) form, only in a few
#Colloidea#.

_The Calymma_, or the jelly-veil including the central capsule, is very
voluminous in many SPUMELLARIA of gigantic size, mainly in the large
#Colloidea#, and in all Polycyttaria or social Radiolaria. It includes here
a considerable number of large vacuoles or "alveoli." The calymma never
exhibits in this legion the dark voluminous phæodium, possessed by all
PHÆODARIA.

_Xanthellæ_ or "zooxanthellæ" are numerous in the calymma of most
SPUMELLARIA, but by no means constant; they are very variable in number and
size.

_The Matrix_, placed between the calymma and central capsule, is, in the
majority of the SPUMELLARIA, a rather thick layer of granular exoplasm.

_The Pseudopodia_ arising from it are very numerous, equally disposed over
the whole {9}surface, and are in general rather fluid, exhibiting a
considerable tendency to ramify, anastomose, and form networks. The
movement of granules is commonly lively. In the Polycyttaria all capsules
of one colony or "coenobium" are connected by the dense variable network of
anastomosing pseudopodia.

_Synopsis of the Orders and Suborders of_ SPUMELLARIA.

I. COLLODARIA. { Skeleton entirely wanting, 1. #Colloidea.#
Skeleton wanting or {
quite imperfect, { Skeleton represented by
not latticed. { numerous scattered spicules, 2. #Beloidea.#

II. SPHÆRELLARIA. { Lattice-shell spherical or
{ composed of concentric spheres, 3. #Sphæroidea.#
Skeleton a perfect {
shell of lattice { Lattice-shell ellipsoidal or
work, or spongy and { prolonged in one axis, 4. #Prunoidea.#
resembling {
wicker-work. { Lattice-shell discoidal or
{ shortened in one axis, 5. #Discoidea.#
{
{ Lattice-shell lentelliptical,
{ with different extent of growth
{ in three axes, 6. #Larcoidea.#

----

Order I. COLLODARIA, Haeckel, 1881.

_Collodaria_, Haeckel, Prodromus, 1881, p. 469.
_Collida_ et _Sphærozoida_, Haeckel, 1862, Monogr. d. Radiol., pp. 246,
522.

_Definition._--SPUMELLARIA without latticed shell.

The order #Collodaria#, the first order of Radiolaria, comprises all those
SPUMELLARIA in which the skeleton is either entirely wanting, or
represented by numerous single, solid, siliceous needles or spicules,
loosely scattered in the calymma around the central capsule. Never in this
order is there any trace of the latticed or fenestrated shell, which
characterises the second order, #Sphærellaria#. The skeleton exhibits no
trace of phylogenetic connection in the two orders.

In my monograph (1862) two families appertaining to this order are
described, the Collida (p. 244) and the Sphærozoida (p. 521). Both families
contain forms with and without a skeleton. Of the solitary or monozous
Collida the Thalassicollida are devoid of a skeleton, whilst the
Thalassosphærida are provided with a skeleton. Of the social or polyzous
Sphærozoida the Collozoida are without a skeleton, the Rhaphidozoida
provided with one. As the special form in both skeletophorous subfamilies
is exactly the same, I prefer now to associate them in the suborder
#Beloidea#, and to oppose them to the other two skeletonless subfamilies,
which are united under the name of #Colloidea#.

{10}_Synopsis of the four Families of Collodaria._

Suborder I. { Solitary cells, living
COLLOIDEA. { as isolated individuals
{ (_Colloidea monozoa_), 1. THALASSICOLLIDA.
Skeleton entirely {
wanting. { Associated cells, living
{ in colonies or coenobia
{ (_Colloidea polyzoa_), 2. COLLOZOIDA.

Suborder II. { Solitary cells, living as
BELOIDEA. { isolated individuals
{ (_Beloidea monozoa_), 3. THALASSOSPHÆRIDA.
Skeleton composed {
of numerous { Associated cells, living in
needles or { colonies or coenobia
spicula, scattered { (_Beloidea polyzoa_), 4. SPHÆROZOIDA.
in the calymma. {

----

Suborder I. COLLOIDEA, Haeckel.

_Definition._--SPUMELLARIA without skeleton.

The suborder #Colloidea# comprises all those SPUMELLARIA in which no
skeleton is developed. The whole body is therefore soft--a true
malacoma--and is composed only of two essential parts, the central capsule
and the enveloping extracapsulum. The suborder contains only two different
families, the solitary #Thalassicollida# (or Colloidea monozoa) and the
associated #Collozoida# (or Colloidea polyzoa). Both families are very
nearly allied, and differ only in one single essential character: the
solitary life of the former, the social union of the latter. It seems to be
merely in consequence of this difference that the cleavage of the nucleus
commonly takes place very late in the former, very early in the latter.

Therefore the full-grown Thalassicollida (till immediately before
propagation) commonly exhibit one single nucleus in the centre of the
capsule, whilst in the Collozoida the capsule is distended by numerous
small nuclei. In these latter the centre of the capsule usually contains
one large oil-globule, whilst in the former oil-globules are either
wanting, or scattered in large numbers in the endoplasm, or disposed in one
layer on the inside of the capsule membrane.

In the solitary Thalassicollida each capsule is enclosed in its own
peculiar spherical calymma, whilst in the associated Collozoida all
capsules of the colony are united in one common, very voluminous calymma.

Family I. #THALASSICOLLIDA#, Haeckel, 1862.

_Thalassicollida_, Haeckel, 1862, Monogr. d. Radiol., p. 246.
_Thalassicollida_, Haeckel, 1881, Prodromus, p. 469.

_Definition._--#Colloidea# solitaria.

The family Thalassicollida comprises all solitary SPUMELLARIA without a
skeleton. The oldest and best known form of this family is the genus
_Thalassicolla_, as restricted by {11}Johannes Müller.[4] The most common
representative of it, the cosmopolitan _Thalassicolla nucleata_, was first
described by Huxley in 1851. But as early as 1834 another large
Radiolarian, appertaining either to this or to a nearly allied family, had
been described by Meyen as _Physematium atlanticum_. A third genus was
detected by me in 1859 at Messina and figured under the name _Thalassolampe
margarodes_.[5] A very accurate histological description of these forms was
given in 1876 by Richard Hertwig.[6] The same author figured in his
Organismus in 1879 a very interesting simpler form under the name
_Thalassolampe primordialis_ (Taf. iii. fig. 5). Some similar forms had
already been observed by me, and are here united with it to form the first
genus _Actissa_.[7]

_Actissa_ is of the highest general interest as the most simple and typical
form of all Radiolaria, and as the common ancestral form, from which all
other forms of this large class may be derived. Its unicellular body
exhibits neither the extracapsular alveoli of _Thalassicolla_, nor the
intracapsular alveoli of _Thalassolampe_, and shows all essential
characters of the Radiolarian type in its most simple form (Pl. 1, figs. 1
to 1_c_). The simple cell-body is composed of a spherical central capsule
and a concentric, spherical, enveloping calymma, both separated by a thin
membrane which is perforated by innumerable pores. The capsule includes the
endoplasm and in the centre a simple spherical nucleus with nucleolus; at
the time of propagation this latter becomes cleft into numerous small
nuclei, each of which, together with a small piece of the surrounding
endoplasm, forms a flagellated zoospore (fig. 1_c_). The extracapsulum is
formed by the large, structureless, spherical calymma or concentric
jelly-veil enveloping the capsule, and by the thin granular matrix or the
layer of exoplasm which separates the calymma from the membrane. From this
matrix or maternal tissue arise innumerable very long and thin pseudopodia,
as simple radiating filaments, the proximal part of which is included in
the calymma, whilst the distal part floats freely in the sea-water (Pl. 1,
fig. 1).

The other Thalassicollida differ from their common ancestral form,
_Actissa_, mainly by the higher histological differentiation of the
unicellular body. Whilst in _Thalassicolla_ and _Thalassolampe_ the nucleus
remains a single sphere as in _Actissa_, it becomes branched or covered
with radial blind saccules in _Thalassopila_ and _Thalassophysa_; also the
intracapsular protoplasm develops here a great variety of peculiar
different corpuscles, as oil-globules, pigment-granules, concentric
concretions, crystals, &c. But the most striking peculiarity by which the
other Thalassicollida differ from _Actissa_ is the development of large
vesicular alveoli, either within or without the capsule; the unicellular
body reaches by this inflation the extraordinary size of 5 to 10 mm. or
more.

{12}_Synopsis of the Genera of Thalassicollida._

A. Alveoli neither { Nucleus spherical
within nor without { (sometimes ellipsoidal),
the central capsule. { not branched, 1. _Actissa_.

B. Numerous large { Nucleus spherical, 2. _Thalassolampe_.
alveoli within the {
central capsule (not { Nucleus branched or
in the calymma). { covered with radial
{ sacs, 3. _Thalassopila_.

C. Numerous large { Nucleus spherical, 4. _Thalassicolla_.
alveoli without the {
central capsule, { Nucleus branched, or
within the jelly-veil { covered with radial
or calymma. { sacs, 5. _Thalassophysa_.

Genus 1. _Actissa_,[8] n. gen.

_Definition._--#Thalassicollida# with simple spherical nucleus, without any
alveoli (either within or outside the central capsule).

The genus _Actissa_ is the most simple and typical form of all Radiolaria,
and may be regarded as the common ancestral form of the whole class. The
spherical body is composed of a simple spherical capsule and a concentric
spherical calymma or jelly-envelope. Neither the former nor the latter
contains alveoli. The central capsule possesses a strong membrane
perforated by small pores, and contains in the intracapsular sarcode
numerous small pellucid vacuoles, and in its middle a simple, concentric,
spherical nucleus (often with some nucleoli); sometimes also one or more
oil-globules. The extracapsularium forms a soft, voluminous, structureless
calymma or enveloping jelly-sphere, perforated by the numberless, fine
pseudopodia, which radiate outwards from the matrix or the thin granulated
sarcode-layer, surrounding the capsule. Often (but not constantly)
xanthellæ or yellow cells are scattered in it. _Actissa_ differs from the
following skeletonless genera in the absence of all alveoli; it has neither
intracapsular alveoli (like _Thalassolampe_) nor extracapsular alveoli
(like _Thalassicolla_). The first observed species of this genus is that
which I found in 1866 at the Canary Islands, _Actissa prototypus_; the
second is that which Hertwig accurately described in 1879, from the
Mediterranean (Messina), _Actissa primordialis_; the third I observed in
1881 at Ceylon, frequent and sporiparous, _Actissa princeps_. A fourth
species (_Actissa radiata_) exhibited a distinct radial segmentation of the
capsule-contents. These four species are quite spherical. Six other
species, occurring in different preparations from the Challenger, are
distinguished by modifications of the spherical capsule-form and may
represent three different subgenera (or, perhaps better,
genera?)--_Actiprunum_ ellipsoidal, _Actidiscus_ lenticular, _Actilarcus_
lentelliptical; perhaps these are the ancestral forms of the three
sections: #Prunoidea#, #Discoidea#, #Larcoidea#.

{13}Subgenus 1. _Procyttarium_, Haeckel, 1879.

_Procyttarium_, Haeckel, Natürl. Schöpfungsgeschichte, ed. vii. p. 705.

_Definition._--Central capsule spherical.

1. _Actissa princeps_, n. sp. (Pl. 1, fig. 1).

Central capsule spherical, colourless or a little reddish, transparent,
with a thick double-edged membrane. Nucleus central, spherical, one-third
as broad as the central capsule, containing a single, central, glossy
nucleolus. Protoplasm finely granulated, without oil-globules, with
numerous clear spherical vacuoles of equal size and at equal distances; the
superficial layer of protoplasm (immediately below the membrane) radially
striated (fig. 1). In some older specimens the nucleus was divided into
numerous small nuclei (fig. 1_a_), which by further division gave the
nuclei of the flagellated spores (fig. 1_b_); each spore with a very thin
lateral flagellum (fig. 1_c_). Jelly-like calymma twice as broad as the
enclosed capsule, without xanthellæ or yellow cells, pierced by
innumerable, very thin and long, undivided pseudopodia, which arise from
the sarcode-matrix on the outside of the membrane (six to eight times
longer than shown in fig. 1).

_Dimensions._--Diameter of the central capsule 0.1 to 0.12, of the nucleus
0.03 to 0.04, of the jelly calymma 0.2 to 0.3.

_Habitat._--Indian Ocean, Ceylon, Belligemma, Haeckel, 1881; also in a
preparation from Station 271, Central Pacific, surface.

2. _Actissa primordialis_, Haeckel.

_Thalassolampe primordialis_, R. Hertwig, 1879, Organismus der
Radiolarien, p. 32, Taf. iii. fig. 5.

_Procyttarium primordiale_, Haeckel, 1879, Natürl. Schöpf., ed. vii. p.
705, Taf. xvi. fig. 1.

Central capsule spherical, dim-yellowish, with a thin, simple-edged but
compact membrane. Nucleus large, central (about half as broad), with one or
two dark nucleoli; on its side an excentric oil-globule, nearly of the same
size. Protoplasm between nucleus and membrane, in the younger specimens
finely granulated and radially striped; in the older specimens with
numerous hyaline globules (vacuoles). Jelly-envelope or calymma very
voluminous, ten to twelve times as broad as the central capsule,
structureless, containing numerous yellow bodies (xanthellæ?), pierced by
very numerous simple pseudopodia.

_Dimensions._--Diameter of the central capsule 0.11 to 0.18, of the nucleus
0.04 to 0.09, of the jelly-like calymma 1.2 to 1.5.

_Habitat._--Mediterranean, Messina, Hertwig, 1878, surface.

3. _Actissa prototypus_, n. sp.

Central capsule spherical, red-coloured, with a thick, double-edged
membrane. Nucleus central, spherical, half as large as the radius of the
capsule, containing a large number (forty to sixty) of small {14}dark
nucleoli. Protoplasm filled up with numerous small clear vacuoles, and
between them fine red pigment granules; on the inside of the membrane one
layer of dark oil-globules. Jelly-like calymma four times as broad as the
enclosed capsule, with very numerous small xanthellæ.

_Dimensions._--Diameter of the capsule 0.2, of the nucleus 0.05, of the
calymma 0.8.

_Habitat._--Atlantic, Canary Islands (Lanzerote, Haeckel), 1866; also at
Station 348, surface.

4. _Actissa radiata_, n. sp.

Central capsule spherical, dark, with a thick, double-edged membrane.
Nucleus central, spherical, half as large as the capsule, transparent.
Protoplasm divided into numerous cuneiform radial pieces which are
separated by clear intervals, and filled with very fine dark granules
(darker in the distal half). The equatorial optical section exhibits around
the circular clear nucleus a coronal of twenty-five such wedge-shaped
pieces (mother-cells of spores?) No oil-globules in the central capsule.
Jelly-like calymma one and a half times as broad as the enclosed capsule,
with numerous small xanthellæ.

_Dimensions._--Diameter of the capsule 0.15, of the nucleus 0.07, of the
calymma 0.24.

_Habitat._--North Pacific, Station 241, surface.

Subgenus 2. _Actiprunum_, Haeckel, 1882.

_Definition._--Central capsule ellipsoidal, with one prolonged axis.

5. _Actissa prunoides_, n. sp.

_Actiprunum prunoideum_, Haeckel, 1882, Manuscript.

Central capsule ellipsoidal, colourless, with a thin, simple-edged
membrane. Proportion of its major axis to the minor 4 : 3. Nucleus
spherical, its diameter equal to one-third of the minor axis, in its centre
a large, dark, spherical nucleolus. Protoplasm clear, containing numerous
small vacuoles, separated by regular distances, and on the inside of the
capsule-membrane, numerous (forty to fifty) small oil-globules. Calymma (or
jelly-veil) ellipsoidal, with a thin sarcode-stratum on the outside of the
capsule; its diameter twice as large as that of the central capsule.

_Dimensions._--Major axis of the capsule 0.16, minor 0.12; diameter of the
nucleus 0.04; major axis of the calymma 0.32, minor 0.24.

_Habitat._--Central Pacific, Station 274, surface.

6. _Actissa ellipsoides_, n. sp.

_Actiprunum ellipsoides_, Haeckel, 1882, Manuscript.

Central capsule ellipsoidal, red-coloured, with a thick, double-edged
membrane. Proportion of its major axis to the minor 5 : 3. Nucleus
ellipsoidal, one-third as large as the capsule, containing eight small dark
nucleoli. Protoplasm dusky, filled with dark pink pigment-granules; in the
{15}major axis, on both poles of the nucleus-axis, two large oil-globules,
half as large as the nucleus. Calymma ellipsoidal, with numerous xanthellæ;
its diameter four times as large as that of the capsule.

_Dimensions._--Major axis of the capsule 0.2, minor 0.12; major axis of the
nucleus 0.07, minor 0.04; major axis of the calymma 0.8, minor 0.5.

_Habitat._--Mediterranean, Corfu, 1877, Haeckel, surface.

Subgenus 3. _Actidiscus_, Haeckel, 1882.

_Definition._--Central capsule lenticular, with one shortened axis.

7. _Actissa discoides_, n. sp.

_Actidiscus discoides_, Haeckel, 1882, Manuscript.

Central capsule lenticular, red-coloured, about twice as broad as high,
with a thick, double-edged membrane. Nucleus spherical, one-third as broad
as the capsule, with one single, large central nucleolus. Protoplasm dusky,
filled with scarlet pigment; granules and small oil-globules between them.
Calymma lenticular, three times as broad as the capsule.

_Dimensions._--Major axis of the capsule 0.16, minor 0.08; diameter of the
nucleus 0.05; breadth of the calymma 0.5.

_Habitat._--North Pacific, Station 236, surface.

8. _Actissa lenticularis_, n. sp.

Central capsule lenticular, flattened, about three times as broad as high,
with a thin, simple-edged membrane. Nucleus lenticular, one-third as large
as the capsule, with ten small dark nucleoli. Protoplasm transparent,
colourless, filled with small vacuoles at regular distances; on the inside
of the membrane in the circular periphery of the lens twenty dark
oil-globules. Calymma lenticular, twice as broad as the capsule.

_Dimensions._--Major axis of the capsule 0.15, minor 0.05; breadth of the
nucleus 0.05, height 0.02; breadth of the calymma 0.03.

_Habitat._--East Pacific, Station 272, surface.

9. _Actissa phacoides_, n. sp.

_Actidiscus phacoides_, Haeckel, 1882, Manuscript.

Central capsule lenticular, strongly flattened, about four times as broad
as high, with a thin, simple-edged membrane. Nucleus lenticular, one-fourth
as broad as the capsule, with numerous (twenty or more) small nucleoli.
Protoplasm filled with dark pigment-granules; on the inside of the membrane
in the circular periphery of the lens thirty-two dark oil-globules. Calymma
lenticular, three times as broad as the capsule.

{16}_Dimensions._--Major axis of the capsule 0.2, minor 0.05; breadth of
the nucleus 0.05, height 0.015; breadth of the calymma 0.6.

_Habitat._--Tropical Atlantic, Station 347, surface.

Subgenus 4. _Actilarcus_, Haeckel.

_Definition._--Central capsule lentelliptical, with three different axes.

10. _Actissa larcoides_, n. sp.

Central capsule lentelliptical; with three different axes, bearing the
proportion 4 : 3 : 2. Nucleus spherical; its diameter equal to the shortest
radius of the capsule. No nucleoli visible. Protoplasm transparent, with
small vacuoles; on the inside of the thin capsule-membrane numerous (fifty
to sixty) small oil-globules, regularly disposed. Calymma lentelliptical,
twice as large as the central capsule.

_Dimensions._--Major axis or length of the capsule 0.2, middle axis or
breadth 0.15, minor axis or height 0.1; diameter of the nucleus 0.05, of
the calymma 0.3-0.4.

_Habitat._--Central Pacific, Station 266, surface.

Genus 2. _Thalassolampe_,[9] Haeckel, 1862, Monogr. d. Radiol., p. 253.

_Definition._--#Thalassicollida# without extracapsular alveoles, but with
large roundish or globular alveoles within the central capsule, with a
simple spherical, not branched nucleus in the centre.

The genus _Thalassolampe_ is, next to _Actissa_, the most simple of all
Radiolaria, but differs from it by the large intracapsular alveoles. By
these the central capsule is inflated to an extraordinary size, which in
_Thalassolampe maxima_ exceeds that of most other Radiolaria. From the
nearly allied _Thalassopila_ it differs by the simple spherical nucleus,
from _Physematium_ by the absence of spicula. Of the two species of the
genus the first observed _Thalassolampe margarodes_, 1862, is
Mediterranean, the second, _Thalassolampe maxima_, 1882, is Indian.

1. _Thalassolampe margarodes_, Haeckel.

_Thalassolampe margarodes_, Haeckel, 1862, Monogr. d. Radiol., p. 253,
Taf. ii. figs. 4, 5.

_Thalassolampe margarodes_, R. Hertwig, 1876, Histologie d. Radiol., p.
68, Taf. iii. figs. 1-5.

Spherical body very soft and limpid, somewhat pearl-like opalescent,
yellowish or bluish. Central capsule with a very thin structureless
membrane, its diameter six to eight times as large as that of the central
spherical nucleus. Wall of the vesicular nucleus thick, perforated by fine
{17}pore-canals; on its inside often numerous oval nucleoli. In the movable
protoplasmic network between the large alveoles a considerable number of
large yellowish or orange oil-globules. Extracapsular jelly-envelope very
thin, contains small yellow bodies (zooxanthellæ). (Compare the accurate
description of this Mediterranean species in my monograph and in Hertwig's
work.) In the Canary Islands I found very often a large variety of it, of
double and triple the size, distinguished by the delicate orange colour of
the intracapsular oil-globules. This may be distinguished as _Thalassolampe
aurantiaca_.

_Dimensions._--Diameter of the whole jelly-sphere 2 to 4 mm., of the
central capsule 2 to 3 mm., of its nucleus 0.2 to 0.4 mm.

_Habitat._--Mediterranean, Messina, Haeckel, Hertwig; Canary Islands,
Lanzerote, Haeckel; surface.

2. _Thalassolampe maxima_, n. sp. (Pl. 1, fig. 2).

Spherical body quite pellucid, like a glass globule, colourless. Central
capsule with a moderately thick, but quite transparent, structureless
membrane, its diameter ten to twelve times as large as that of the central
spherical nucleus. Wall of the vesicular nucleus thick, perforated by fine
pore-canals; on its inside numerous small spherical nucleoli. No large
oil-globules in the movable protoplasmic network between the large
alveoles. Extracapsular jelly-envelope very thin, containing no yellow
bodies. This differs from the preceding nearly allied species in the want
of the intracapsular oil-globules and of the extracapsular yellow bodies.
It possesses the largest central capsule of all known Radiolaria. I found
them living and floating in water taken from the surface of the Indian
Ocean by a bucket.

_Dimensions._--Diameter of the whole jelly-body 12 to 15 mm., of the
central capsule 10 to 12 mm., of the nucleus 0.8 to 1.2 mm.

_Habitat._--Indian Ocean, near the Maldive Islands, Haeckel, 1882, surface.

Genus 3. _Thalassopila_,[10] Haeckel, 1881, Prodromus, p. 469.

_Definition._--Thalassicollida without extracapsular alveoles, but with
large roundish or globular alveoles within the central capsule, with a
papillate or branched nucleus in its centre.

The genus _Thalassopila_ has, like _Thalassolampe_, a voluminous foamy
central capsule, inflated by numerous large alveoles; but it differs in the
complicated form of the nucleus, which is like that of _Thalassophysa_, and
is either branched or occupied by conical or roundish papillæ.

1. _Thalassopila cladococcus_, n. sp. (Pl. 1, fig. 3).

Spherical body dark-spotted, with a thin yellowish jelly-envelope. Central
capsule with a thick and firm membrane, perforated by pores; its diameter
three times that of the central nucleus, {18}three-fourths that of the
whole jelly-sphere. Nucleus profusely branched or papillated, its spherical
surface covered with numerous (more than a hundred) finger-shaped obtuse
blind sacs, about as long as its radius. Protoplasm of the central capsule
forming a loose network between the large roundish alveoles, in the
cortical zone radially striped and containing one layer of large dark
oil-globules. These are regularly distributed on the inside of the
capsule-membrane and separated by intervals, twice as broad as its
diameter, giving to the capsule-surface a spotted appearance. Extracapsular
jelly-envelope thin, yellowish, with very numerous and small xanthellæ.

_Dimensions._--Diameter of the whole jelly-sphere 5 mm., of the central
capsule 4 mm., of the nucleus 1.3 mm.

_Habitat._--Antarctic Ocean, Station 154 (south of Kerguelen), surface.

Genus 4. _Thalassicolla_,[11] Huxley, 1851, Ann. and Mag. Nat. Hist., ser.
2, vol. viii. p. 433.

_Definition._--Thalassicollida without intracapsular alveoles, but with
large roundish or globular alveoles within the extracapsular calymma.
Nucleus in the centre of the capsule simple spherical, not branched.

The genus _Thalassicolla_ was proposed by Huxley in 1851, for a certain
number of different voluminous jelly-like Radiolaria, which he had observed
living during his voyage in the "Rattlesnake" through the tropical seas,
and of which he gives an excellent description--the first accurate
observations on living Radiolaria. Johannes Müller afterwards removed from
this genus the social genera _Sphærozoum_ and _Collosphæra_ (formerly
_Thalassicolla punctata_), and retained as type of the genus _Thalassicolla
nucleata_. In 1862 in my Monograph I added two other species,
_Thalassicolla pelagica_ and _Thalassicolla zanclea_, and later (1870)
_Thalassicolla sanguinolenta_. Now I think it better to separate the last
two species as a new genus, _Thalassophysa_, characterised by the papillate
or branched nucleus, and to retain in _Thalassicolla_ only those forms with
simple spherical nucleus. For both genera the extracapsular, voluminous,
spherical calymma or jelly-envelope, with numerous large alveoles, is
characteristic. The membrane of the central capsule in _Thalassicolla_ is
now structureless (subgenus _Thalassicollarium_, with three species), now
characterised by a peculiar structure, prominent ridges on the inside of
the membrane, which form a network with polygonal plates, resembling an
epithelium (Pl. 1, fig. 5_b_; subgenus _Thalassicollidium_, with four
species). Of the seven species here described, two are cosmopolitan, widely
distributed, and common; one is Mediterranean, one Atlantic, and three
Pacific.

Subgenus 1. _Thalassicollarium_, Haeckel.

_Definition._--Membrane of the central capsule structureless, only
perforated by innumerable very small radial pores.

{19}1. _Thalassicolla pellucida_, n. sp.

Spherical body very soft, transparent, clear and colourless, without any
pigment. Central capsule soft, hyaline, with a thin, structureless, not
areolated membrane. Diameter of the central capsule about three times that
of the nucleus, one-fourth to one-sixth that of the jelly-envelope. Nucleus
delicate, transparent, with one single central nucleolus, about one-third
its diameter. Protoplasm of the central capsule contains only small,
pellucid, densely packed globules (vacuoles?), no oil-globules.
Extracapsular body quite transparent, without pigment or oil-globules, only
composed of the large alveoles imbedded in the jelly-cover, and of the fine
protoplasmic network between them. No xanthellæ.

_Dimensions._--Diameter of the central capsule 0.8 to 1.2, of the nucleus
0.3 to 0.4, of the calymma 3 to 6 mm.

_Habitat._--Cosmopolitan, Canary Islands, Haeckel; Cape, Australia,
Pacific, Challenger; surface.

2. _Thalassicolla spumida_, n. sp.

Spherical body nearly transparent, yellowish, without dark pigment. Central
capsule pellucid, with a thick, structureless, not areolated membrane.
Diameter of the central capsule about twice that of the nucleus, one-sixth
to one-eighth that of the jelly-cover. Nucleus delicate, somewhat opaque,
with numerous small nucleoli. Protoplasm of the central capsule contains
small pellucid globules (vacuoles?), and immediately under its membrane (on
its inside) one single layer of large, dark, refractive oil-globules.
Extracapsular body very voluminous, foamy, with innumerable alveoles in the
jelly, and many xanthellæ between them.

_Dimensions._--Diameter of the central capsule 0.6 to 0.8, of the nucleus
0.3 to 0.4, of the calymma 3 to 5 mm.

_Habitat._--Atlantic, Canary Islands, Haeckel; Cape Verde Islands,
Challenger Station 350; Brazil, Rabbe; surface.

3. _Thalassicolla zanclea_, Haeckel.

_Thalassicolla zanclea_, Haeckel, 1862, Monogr. d. Radiol., p. 252, Taf.
ii. fig. 3.

Spherical body opaque, transparent only in the periphery, with colourless
central capsule, but with brown or black pigment-powder scattered
everywhere through the extracapsular alveolated jelly-cover. Central
capsule soft, transparent, colourless, with a thin structureless, not
areolated membrane. Diameter of the central capsule about one and a half
times that of the nucleus, one-half or one-third that of the jelly-cover.
Nucleus delicate, transparent, with a thin, finely punctated membrane, with
one or more nucleoli. Protoplasm of the central capsule contains only
small, pellucid, densely packed globules (vacuoles?), no oil-globules.
Extracapsular body very dark and opaque, with a great mass of brown or
blackish-brown pigment between the alveoles of the jelly-cover. Numerous
xanthellæ.

_Dimensions._--Diameter of the central capsule 0.1 to 0.12, of the nucleus
0.07 to 0.08, of the calymma 0.2 to 0.4.

_Habitat._--Mediterranean, Messina, Haeckel.

{20}Subgenus 2. _Thalassicollidium_, Haeckel.

_Definition._--Membrane of the central capsule areolated, with small
polygonal plates, resembling an epithelial cell-tissue, spotted by
innumerable fine radial pores.

4. _Thalassicolla australis_, n. sp.

Spherical body nearly transparent, without dark pigment. Central capsule
colourless, somewhat opaque, with a thick and firm, very elastic membrane,
areolated by polygonal, punctated figures resembling cells. Diameter of the
central capsule about three times that of the nucleus, one-third that of
the jelly-cover. Nucleus thin-walled, with many small nucleoli. Protoplasm
of the central capsule finely granulated, containing numerous hyaline
globules (vacuoles?) of different size, and in each of these one single
roundish, dark refringing corpuscle, concentrically lamellated like an
amylum grain. Extracapsular body without pigment or oil-globules, only
composed of the large alveoles imbedded in the jelly-cover, and of the fine
protoplasmic network between them. No xanthellæ.

_Dimensions._--Diameter of the central capsule 1 to 2, of its nucleus 0.3
to 0.4, of the nucleoli 0.12 to 0.16, of the hyaline globules in the
protoplasm of the capsule 0.02 to 0.05; calymma, 4 to 6 mm.

_Habitat._--South-west Pacific, east coast of Australia, New Zealand, &c.;
Challenger Stations 163, 171; surface.

5. _Thalassicolla nucleata_, Huxley.

_Thalassicolla nucleata_, Huxley, 1851, Ann. and Mag. Nat. Hist., ser. 2,
vol. viii. p. 435, pl. xvi. fig. 4.

_Thalassicolla nucleata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 28.

_Thalassicolla coerulea_, Schneider, 1858, Archiv f. Anat. u. Physiol.,
p. 40, Taf. iii. Bd. i. figs. 5-7.

_Thalassicolla nucleata_, Haeckel, 1862, Monogr. d. Radiol., p. 249, Taf.
iii. figs. 1-5.

_Thalassicolla nucleata_, R. Hertwig, 1876, Histologie d. Radiol., p. 43,
Taf. iii. figs. 61-5, Taf. iv., v.

_Thalassicolla nucleata_, R. Hertwig, 1879, Organismus d. Radiol., p. 34.

Spherical body in the central part opaque, black or dark coloured, in the
periphery transparent, whitish, or yellowish. Central capsule rather
compact, yellowish, opaque, with a thick and firm, very elastic membrane,
areolated by polygonal, punctated figures resembling cells. Diameter of the
central capsule about twice as large as that of the nucleus, one-half to
one-fourth that of the jelly-cover. Nucleus with a very thick, finely
punctated membrane, containing a viscous fluid (when coagulated finely
granular), and sometimes one large, central, spherical, or ramified
nucleolus, sometimes a variable number of smaller roundish nucleoli.
Protoplasm of the central capsule containing many very variable corpuscles,
mostly pellucid (albuminous?) spherules, containing oil-globules, or
concentric amyloid concretions, or crystals, &c. Extracapsular body with
dark pigment-powder of variable colour (black, brown, violet, blue, &c.),
densely aggregated around the central capsule, more loosely dissipated
between the alveoles of the outer jelly-cover. Xanthellæ very numerous.

{21}_Dimensions._--Diameter of the central capsule 0.4 to 0.8, of the
nucleus 0.02 to 0.05, of the calymma 1 to 5 mm.

_Habitat._--Cosmopolitan; common in all warmer seas; Mediterranean,
Atlantic, Indian Ocean, Pacific, mainly between lat. 40° N. and lat. 40°
S.; surface.

6. _Thalassicolla maculata_, n. sp. (Pl. 1, fig. 4).

Spherical body in the central part opaque, black or dark coloured, in the
periphery semi-transparent, spotted. Central capsule compact, yellowish,
opaque, with a thick and firm, very elastic membrane, areolated by
polygonal, punctated figures resembling cells. Diameter of the central
capsule about twice that of the nucleus, one-third to one-fifth that of the
jelly-cover. Nucleus thin-walled, with one large spherical nucleolus.
Protoplasm of the central capsule contains innumerable very small, hyaline,
spherical vesicles of equal size (or vacuoles?), two to four times as broad
as the separating bridges of protoplasm. Extracapsular body with dark
pigment-powder of black or brown colour, densely accumulated around the
central capsule (in the matrix), loosely scattered between the alveoles of
the outer jelly-cover. The latter appears spotted by numerous large,
roundish lumps of protoplasm, scattered between the alveoles. No xanthellæ.

_Dimensions._--Diameter of the central capsule 0.3 to 0.6, of the nucleus
0.2 to 0.3, of the hyaline globules in the protoplasm of the capsule 0.02
to 0.03; calymma, 2 to 3 mm.

_Habitat._--South Pacific, Challenger Station 289.

7. _Thalassicolla melacapsa_, n. sp. (Pl. 1, fig. 5).

Spherical body in the central part opaque, black or dark coloured, in the
periphery semi-transparent, spotted. Central capsule compact, black, with a
thick and firm, very elastic membrane, areolated by polygonal, punctated
figures resembling cells. Diameter of the central capsule about twice that
of the nucleus, one-third or half that of the jelly-cover. Nucleus
thin-walled, transparent, containing very numerous and small nucleoli.
Protoplasm of the central capsule filled with small black pigment-granules,
quite intransparent, contains densely packed hyaline (albuminous?) globules
of equal size; every pellucid globule includes a smaller globule (one-third
or one-fourth its diameter), which appears to be composed of aggregated
oil-bubbles. Extracapsular body without pigment, contains between its
alveoles in the inner half numerous, dark refractive oil-globules, in the
outer half numerous amoeboid lumps of protoplasm, irregularly scattered. No
xanthellæ.

_Dimensions._--Diameter of the central capsule 2 to 2.5, of the nucleus 1
to 1.5, of the hyaline globules in the protoplasm of the capsule 0.03 to
0.04; calymma, 3 to 5 mm.

_Habitat._--South-east Pacific (near Valparaiso), Challenger Station 300,
surface.

Genus 5. _Thalassophysa_,[12] Haeckel, 1881, Prodromus, p. 470.

{22}The genus _Thalassophysa_ contains those species of Thalassicollida
formerly associated with _Thalassicolla_, which are distinguished by the
complicated, ramose, or papillate form of the large nucleus. All three
species here described are found in the Mediterranean and the Atlantic. To
this genus appertains also that strange form of Radiolaria which I
described in 1870 as _Myxobrachia_ (compare _Thalassophysa sanguinolenta_).

1. _Thalassophysa papillosa_, n. sp.

_Thalassicolla papillosa_, Haeckel, 1867, Manuscript.

Spherical body transparent, colourless, or somewhat yellowish. Central
capsule soft, colourless, with a very thin but firm, elastic, structureless
membrane. Diameter of the central capsule about twice that of the nucleus,
one-fourth to one-sixth that of the jelly-envelope. Nucleus papillated, its
spherical surface covered with a great number (50 to 80) of conical or
finger-like protuberances or blind sacs, not longer than half its radius.
Protoplasm of the central capsule filled with very small and numerous
spherical vacuoles, without oil-globules. Extracapsular jelly-body, without
dark pigment, oil-globules, and large protoplasmic lumps, contains between
its alveoles very numerous xanthellæ.

_Dimensions._--Diameter of the whole jelly sphere 4 to 5 mm., of the
central capsule 0.8 to 1 mm., of its nucleus 0.4 to 0.5.

_Habitat._--Canary Islands, Lanzerote, common, Haeckel; Cape Verde Islands,
Challenger; surface.

2. _Thalassophysa sanguinolenta_, Haeckel.

_Thalassicolla sanguinolenta_, Haeckel, 1870, Jenaische Zeitschr., Bd. v.
p. 526, Taf. 18.

_Thalassicolla sanguinolenta_, Haeckel, 1870, Biolog. Studien, i. p. 113,
Taf. iv.

_Thalassicolla sanguinolenta_, R. Hertwig, 1879, Organismus d. Radiol.,
p. 37, Taf. iii. fig. 1.

_Myxobrachia rhopalum_, Haeckel, 1870, Jenaische Zeitschr., Bd. v. p.
519, Taf. 18 (et in Biol. Stud., _loc. cit._).

_Myxobrachia pluteus_, Haeckel, 1870, Jenaische Zeitschr., Bd. v. p. 520,
Taf. 18 (et in Biol. Stud., _loc. cit._).

Spherical body in the central part opaque, reddish, in the periphery
transparent, yellowish. Central capsule compact, white, red spotted, with a
thick elastic membrane, perforated by pores, but not areolated. Diameter of
the central capsule three times that of the nucleus, one-fifth to
one-eighth that of the jelly-envelope. Nucleus papillated, its spherical
surface covered with numerous (80 to 120) conical or finger-like
protuberances not longer than one-fourth or one-third of its radius. On the
inside of these blind sacs lie numerous small roundish nucleoli. Protoplasm
of the central capsule in the outer (cortical) zone (on the inside of the
membrane) radially striped, with one layer of very numerous red
oil-globules of equal size, producing its blood-spotted appearance; in the
inner (medullary) zone foamy, with numerous small spherical vacuoles.
Extracapsular jelly-body without dark pigment, contains between its
alveoles no large protoplasmic lumps (as in _Thalassophysa pelagica_), but
numerous small oil-globules and xanthellæ. This species sometimes amasses
in its jelly-envelope large accumulations of Coccoliths and Coccospheres,
{23}which are much heavier than the jelly-body, and produce arm-like
protuberances of it; this modified form, often of very regular and peculiar
appearance, I formerly described as _Myxobrachia_ (compare my Biolog.
Studien, _loc. cit._, and Hertwig, _loc. cit._, p. 37). Compare also
_Myxobrachia cienkowski_, Wagner, 1872, L. N. 23.

_Dimensions._--Diameter of the whole jelly-sphere 5 to 8 mm., of the
central capsule 1 to 1.2 mm., of its nucleus 0.3 to 0.4.

_Habitat._--Canary Islands, Lanzerote; common, Haeckel; Mediterranean,
Messina, Hertwig; surface.

3. _Thalassophysa pelagica_, Haeckel.

_Thalassicolla pelagica_, Haeckel, 1862, Monogr. d. Radiol., p. 247, Taf.
i.

_Thalassicolla pelagica_, R. Hertwig, 1879, Organismus d. Radiol., p. 35,
Taf. iii. fig. 4.

Spherical body in the central part opaque, yellowish, in the periphery
semi-transparent, spotted. Central capsule compact, yellowish-white, with a
thick and compact membrane, perforated by pores, but not areolated.
Diameter of the central capsule about twice that of the nucleus, one-half
to one-sixth that of the jelly-envelope. Nucleus papillated, its spherical
surface covered with numerous (20 to 60) conical, roundish, or finger-like
protuberances, not longer than its radius (commonly only one-half or
one-third as long). Enclosed in the semi-fluid substance of the nucleus are
very long and thin cylindrical nucleoli snake-like, contorted, and
penetrating into the protuberances of the nucleus. Protoplasm of the
central capsule in the outer (cortical) zone (on the inside of the
membrane) radially striped, with one layer of large oil-globules of
different sizes; in the inner (medullary) zone foamy, with numerous small
spherical vacuoles, mostly of equal size. Extracapsular jelly-body without
dark pigment, contains between its alveoles a large number of large
roundish or amoeboid lumps of protoplasm, and very numerous yellow cells or
xanthellæ (compare the detailed description in my Monograph, and in R.
Hertwig's work).

_Dimensions._--Diameter of the whole jelly-sphere 1 to 4 mm., of the
central capsule 0.5 to 0.6, of the nucleus 0.2 to 0.3.

_Habitat._--Mediterranean--Messina, Corfu, Nizza, Genoa, Haeckel; Messina,
R. Hertwig; surface.

Family II. #COLLOZOIDA#, Haeckel, 1862 (Pl. 3).

_Collozoida_, Haeckel, 1862, Monogr. d. Radiol., p. 522.

_Definition._--#Colloidea# socialia.

The family Collozoida comprises all associated or colony-building
Radiolaria without skeleton. We unite here all these skeletonless
Radiolarian colonies into one single genus _Collozoum_, constituted (1862)
in my Monograph (p. 522). The oldest known form of it was the _Collozoum
inerme_, described firstly by Johannes Müller (1856) as _Sphærozoum
inerme_. Two other species of the genus were figured (1862) in my Monograph
(p. 522, Tafn. xxxii., xxxv.). A most accurate description of its
histological structure and {24}development was given in 1876 by Richard
Hertwig in his Histologie der Radiolarien (pp. 12-42, Tafn. i., ii.). A
number of other very remarkable forms of _Collozoum_ have been observed by
me during the last few years, and partly figured in Pl. 3.

_Collozoum_, as the only representative of this family, is sufficiently
distinguished from all other Radiolaria by the definition "_Skeletonless
Radiolarian Colonies._" These occur floating on the surface of all warmer
seas, often in astonishing masses, and may be easily confounded, owing to
their external resemblance, with the jelly-like egg-masses of certain
Mollusca. _Collozoum_ is derived either from _Actissa_ or from
_Thalassicolla_, simply by multiplication of the unicellular body and by
reunion of the associated capsules in one common calymma or jelly-veil;
this is constantly alveolated, as in _Thalassicolla_. As in _Actissa_, the
form of the central capsule remains either spherical, or it becomes
ellipsoidal or discoidal, rarely polyhedral or amoeboid. In _Collozoum_ as
in all colonial Radiolaria, the original central nucleus commonly undergoes
cleavage very early into numerous small nuclei, whilst its place is usually
taken by a central oil-globule. This peculiarity may serve often (but not
constantly) for the distinction of isolated capsules of _Collozoum_ from
_Actissa_.

Genus 6. _Collozoum_,[13] Haeckel, 1862, Monogr. d. Radiol., p. 522.

_Definition._--Skeletonless colonies of Radiolaria.

The genus _Collozoum_, as already mentioned, is the only representative of
its family, and comprises all Radiolaria living associated in colonies, and
having no skeleton. Therefore _Collozoum_ possesses all the peculiarities
described above. Although the floating colonies of this genus occur in
enormous masses on the surface of all warmer seas, nevertheless the number
of different species in this genus is not great, and amounts only to
thirteen. If this number increase by further investigations, the subgenera
distinguished in the following description can be advanced to the range of
genera; in which case _Collodinium_ (or _Collozoum_ sensu restricto) will
be characterised by the spherical form of its central capsules,
_Colloprunum_ by the ellipsoidal form (Pl. 3, fig. 9), _Collophidium_ by
the cylindrical, very prolonged form (figs. 2, 3), _Collodiscus_ by the
lenticular or discoidal form, and _Collodastrum_ by the indefinite,
polyhedral, or amoeboid form (figs. 4, 5).

Subgenus 1. _Collodinium_, Haeckel.

_Definition._--Form of the central capsules spherical or subspherical,
never polyhedral, ellipsoidal, or cylindrical.

{25}1. _Collozoum inerme_, Haeckel (Pl. 3, figs. 10-12).

_Collozoum inerme_, Haeckel, 1862, Monogr. d. Radiol., p. 522, Taf. xxxv.

_Collozoum inerme_, Cienkowski, 1871, Archiv. f. mikrosk. Anat., vol.
vii. p. 376, Taf. xxix. figs. 18-36.

_Collozoum inerme_, R. Hertwig, 1876, Histologie der Radiol., p. 12, Taf.
i., ii.

_Collozoum inerme_, R. Hertwig, 1879, Organismus d. Radiol., p. 31, Taf.
iii. fig. 12.

_Sphærozoum inerme_, J. Müller, 1856, Monatsber. d. k. Akad. d. Wiss.
Berlin, p. 478; Abhandl., p. 54.

_Sphærozoum bicellulare_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 54, Taf. viii. fig. 5.

_Thalassicolla punctata_, Huxley (_pro parte_), 1851, Ann. and Mag. Nat.
Hist., ser. 2, vol. viii. p. 433.

Central capsules spherical, with thin, simple-edged membrane, with one
single oil-globule in the centre. (If the capsules multiply by division,
the spherical form becomes violin-shaped, constricted in the middle; and in
this condition the number of oil-globules increases; but in the ordinary
mature state the capsule of this species remains spherical, and its
oil-globule solitary. In quite young capsules the oil-globules are wanting;
Pl. 3, fig. 12.)

_Dimensions._--Diameter of the central capsules 0.04 to 0.16.

_Habitat._--Cosmopolitan, common in all warmer seas (Mediterranean,
Atlantic, Indian, and Pacific), surface.

2. _Collozoum nostochinum_, n. sp.

Central capsules spherical, very large, opaque, distended with red
pigment-granules and with very numerous (two hundred to three hundred)
small oil-globules. Membrane thick, double-edged.

_Dimensions._--Diameter of the central capsules 0.3 to 0.5.

_Habitat._--Indian Ocean, off Socotra, surface, Haeckel.

3. _Collozoum volvocinum_, n. sp.

Central capsules spherical, very large, opaque, containing a great number
(ten to thirty) of large oil-globules, and between them densely packed
masses of dark pigment. Membrane thick, double-edged. This species differs
from _Collozoum inerme_, mainly by the great size of the central capsules
(three to five times as big as in the former) and the great number of
oil-globules in them.

_Dimensions._--Diameter of the central capsules 0.2 to 0.3.

_Habitat._--Central Pacific, Station 272, surface.

Subgenus 2. _Colloprunum_, Haeckel.

_Definition._--Form of the central capsules ellipsoidal, with one prolonged
axis.

4. _Collozoum ovatum_, n. sp.

_Colloprunum ovatum_, Haeckel, 1882, Manuscript.

Central capsules ovate or ellipsoidal, its longer diameter twice to three
times as large as the shorter. In the centre of every capsule one single
oil-globule.

{26}_Dimensions._--Length of the central capsules 0.2 to 0.3, breadth of
them 0.1 to 15.

_Habitat._--North Pacific, Station 244, surface.

5. _Collozoum ellipsoides_, n. sp. (Pl. 3, figs. 8, 9).

_Colloprunum ellipsoides_, Haeckel, 1882, Manuscript.

Central capsules regularly ellipsoidal, very large; their longer diameter
once and a half to twice as large as the shorter. In every capsule fifty to
eighty oil-globules.

_Dimensions._--Length of the central capsules 0.3 to 4, breadth of them
0.2.

_Habitat._--North Atlantic, Færöe Channel (Gulf Stream), surface, John
Murray.

Subgenus 3. _Collophidium_, Haeckel.

_Definition._--Form of the central capsules cylindrical, often snake-like,
contorted, with one axis much prolonged, several times longer than the
transverse axis.

6. _Collozoum contortum_, n. sp.

_Collophidium contortum_, Haeckel, 1882, Manuscript.

Central capsules cylindrical, three to four times as long as broad, C- or
S-like curved, transparent, without oil-globules.

_Dimensions._--Length of the central capsules 0.2 to 0.3, breadth 0.06 to
0.08.

_Habitat._--Tropical Atlantic, Station 347, surface.

7. _Collozoum serpentinum_, n. sp. (Pl. 3, figs. 1-3).

_Collophidium serpentinum_, Haeckel, 1882, Manuscript.

Central capsules cylindrical, filiform, much elongated, ten to one hundred
times, sometimes two hundred to four hundred times as long as broad,
snake-shaped or worm-shaped, curved and contorted in the most irregular
manner, often spiral or twisted into a large nodule. Numerous oil-vesicles
constantly present, forming one series of globules in the axis of every
capsule; distance of the globules, one from another, and also from the
capsule-membrane, about equal to their diameter. (This interesting and very
curious form was very frequently observed living by me in the Canary
Islands, in January 1867; the jelly-colonies were commonly spherical, and
contained fifty to two hundred or more capsules of very different size and
form.)

_Dimensions._--Length of the central capsules 1 to 10, sometimes 20 to 40
mm.; average breadth 0.1 mm.

_Habitat._--Canary Islands, Lanzerote, Haeckel, surface.

{27}8. _Collozoum vermiforme_, n. sp. (Pl. 3, figs. 6, 7).

_Collophidium vermiforme_, Haeckel, 1882, Manuscript.

Central capsules cylindrical, much elongated, five to ten times (sometimes
twenty to fifty times) as long as broad, snake-shaped or worm-shaped, very
irregularly curved and contorted. Numerous oil-globules constantly present,
forming in the axis of every capsule a double series of alternating
rose-coloured globules. (This species is nearly allied to the preceding;
but its capsules are thicker and shorter, and the oil-vesicles in them are
arranged not in a single, but in a double row.)

_Dimensions._--Length of the central capsules 0.6 to 1.2 mm., sometimes 3
to 6 mm.; breadth 0.12.

_Habitat._--Tropical Atlantic, near the west coast of Africa, Station 349,
Canary Islands, surface.

Subgenus 4. _Collodiscus_, Haeckel.

_Definition._--Form of the central capsules discoidal or lenticular, with
one shortened axis.

9. _Collozoum coeruleum_, Haeckel.

_Collozoum coeruleum_, Haeckel, 1862, Monogr. d. Radiol., p. 523, Taf.
xxxii. figs. 6-8.

_Collodiscus coeruleus_, Haeckel, 1882, Manuscript.

Central capsule lenticular or discoidal, flattened, blue coloured, with one
single oil-globule in the centre. Protoplasm containing numerous crystals
and dark blue pigment-granules. Membrane very thick, double-edged. (Whilst
at Messina in 1859 I found this form not constantly discoidal, in 1867 in
the Canary Islands I observed it constantly lenticular.)

_Dimensions._--Breadth of the central capsules 0.1 to 0.15, height 0.04 to
0.08.

_Habitat._--Mediterranean (Messina), Atlantic (Canary Islands), surface.

10. _Collozoum discoideum_, n. sp.

_Collodiscus discoideus_, Haeckel, 1882, Manuscript.

Central capsule discoidal, flattened, transparent, with a ring of twenty to
twenty-five oil-globules in its circular periphery (on the inside of the
thin membrane).

_Dimensions._--Breadth of the central capsules 0.2, height 0.05.

_Habitat._--South Pacific (Juan Fernandez), Station 300, surface.

Subgenus 5. _Collodastrum_, Haeckel.

_Definition._--Form of the central capsules irregular and indefinite,
variable, commonly polyhedral or polygonal, or amoeboid, often with
irregular, finger-like processes.

{28}11. _Collozoum pelagicum_, Haeckel.

_Collozoum pelagicum_, Haeckel, 1862, Monogr. d. Radiol., p. 525, Taf.
xxxii. figs. 4, 5.

_Sphærozoum pelagicum_, Haeckel, 1860, Monatsber. d. k. Akad. d. Wiss.
Berlin, 1860, p. 845.

Central capsules small, quite irregularly formed, roundish-polyhedral or
depressed-polygonal, transparent, without oil-globules. Often many
extracapsular oil-vesicles in the common jelly-body between the central
capsules. Membrane very thin and delicate.

_Dimensions._--Diameter of the central capsules 0.02 to 0.08.

_Habitat._--Mediterranean, Messina, Haeckel; Naples, Brandt; surface.

12. _Collozoum stellatum_, n. sp.

_Collodastrum stellatum_, Haeckel, 1882, Manuscript.

Central capsules star-shaped, irregularly radiating, with a great number
(eight to twenty or more) of radial, short, conical, acute processes, very
variable in size and number. Membrane thin. In every capsule several (four
to eight) oil-globules.

_Dimensions._--Diameter of the central capsules 0.12 to 0.2.

_Habitat._--Central Pacific, Station 274, surface.

13. _Collozoum amoeboides_, n. sp. (Pl. 3, figs. 4, 5).

_Collodastrum amoeboides_, Haeckel, 1882, Manuscript.

Central capsules amoebiform, of moderate size, quite irregularly formed,
with a variable number of finger-like, obtuse, irregular prolongations
(commonly three to six), very variable in size and form. Membrane thin. In
the centre of every capsule one single oil-globule.

_Dimensions._--Diameter of the central capsules 0.04 to 0.08.

_Habitat._--Indian Ocean, Ceylon, Haeckel; Madagascar, Rabbe; surface.

----

Suborder II. BELOIDEA, Haeckel.

_Definition._--SPUMELLARIA with an imperfect skeleton, composed of numerous
solid needles or spicula, scattered irregularly in the calymma.

The suborder #Beloidea# comprises all those SPUMELLARIA which possess an
imperfect or rudimentary skeleton, composed of a variable number of
isolated spicula scattered in the extracapsulum. The suborder contains only
two different families, the solitary Thalassosphærida (or Beloidea monozoa)
and the associated Sphærozoida (or Beloidea polyzoa). Both families are
very nearly allied, and differ only in one single character: the solitary
life of the former, the social union of the {29}latter. It seems to be
merely a consequence of this difference that the cleavage of the nucleus
commonly takes place very late in the former, very early in the latter.

Commonly, therefore, the full-grown Thalassosphærida (until immediately
before their propagation) exhibit one single nucleus in the centre of the
capsule, whilst in the Sphærozoida the capsule is distended with numerous
small nuclei. In these latter the centre of the capsule usually contains
one large oil-globule, whilst in the former oil-globules are either wanting
or scattered in large numbers in the endoplasm, or disposed in one layer on
the inside of the capsule membrane.

In the solitary Thalassosphærida each capsule is enclosed in its own
peculiar spherical calymma, whilst in the associated Sphærozoida all the
capsules of the colony are united into one common, very voluminous,
alveolated calymma.

Family III. #THALASSOSPHÆRIDA#, Haeckel, 1862, (Pl. 2).

_Thalassosphærida_, Monogr. d. Radiol., p. 255.

_Definition._--#Beloidea# solitaria.

The family Thalassosphærida comprises all solitary SPUMELLARIA with an
imperfect skeleton, composed of numerous solid needles or spicula,
scattered around the central capsule in the calymma. The structure of the
unicellular soft body is quite the same as in the Thalassicollida; it
differs from these only in the possession of the extracapsular skeleton.
All needles of this skeleton are solid siliceous spicula, never hollow, as
in the similar Cannorrhaphida among the PHÆODARIA. In the special structure
and form of the skeleton the Thalassosphærida agree perfectly with the
well-known, colony-building Sphærozoida; they differ from these only by
their hermit-like life and by some peculiarities derived from this solitary
development.

The oldest known form of this family is probably the first Radiolarian,
observed in the living state, described in 1834 by Meyen as _Physematium
atlanticum_ (see p. 35). A second form was figured in my Monograph (1862)
as _Thalassosphæra bifurca_ (p. 260, Taf. xii. fig. 1). A third form was
there described under the name _Thalassosphæra morum_; this remarkable form
was first observed by Johannes Müller, and figured under the name
_Thalassicolla morum_ (1858, Abhandl., p. 28, Taf. vii. figs. 1, 2). The
same form was afterwards observed living by myself in the Mediterranean, as
well as in the Atlantic, and in great numbers by the late Sir Wyville
Thomson in the Pacific. The latter gave a good figure of it with some
valuable remarks in his excellent work, The Atlantic (1877, vol. i. p. 233,
fig. 51). He called this peculiar Rhizopod _Calcaromma calcarea_, on
account of the very peculiar _calcareous_ bodies "looking in outline like
the rowels of spurs," which are accumulated in great quantity around the
central capsule, in the calymma. Further investigations have convinced me
that these peculiar stellate {30}bodies of carbonate of lime, for which we
propose the name "Calcastrella," are not parts of the skeleton produced by
the Radiolarian, but foreign bodies picked up by its extracapsular sarcode
(in the same way as the Coccoliths are picked up by _Thalassicolla
sanguinolenta = Myxobrachia!_). These Calcastrella occur also in the
calymma of some Discoidea and other Radiolaria; they are either unicellular
calcareous Algæ, or foreign bodies of other origin. The _Collodarium_,
however, described as _Thalassicolla morum_ and _Calcaromma calcarea_,
seems to be a simple _Actissa_, which has picked up a number of
Calcastrella.

The Challenger collection has yielded a number of other true
Thalassosphærida, which partly agree with _Thalassosphæra_ in the simple
structure of the unicellular body (resembling _Actissa_), and partly differ
from it in the development of alveoles, either within or without the
central capsule (similar to _Thalassolampe_ and _Thalassicolla_). The solid
siliceous spicula, which occur in great numbers scattered in the calymma,
agree perfectly in form with the spicula of the colony-building
Sphærozoida. A characteristic difference between the social and the
solitary #Beloidea# seems to be determined by the cleavage of the nucleus,
which takes place in the latter very late, in the former very early.
Therefore in the large central capsule of the mature solitary
Thalassosphærida, we commonly find one large nucleus in the centre, and a
number of oil-globules around it in the endosarc, or disposed in one layer
on the inside of the capsule-membrane (Pl. 2, figs. 2, 5); whereas in the
much smaller associated capsules of the Sphærozoida one large oil-globule
is placed commonly in the centre, and a great number of small nuclei
scattered in the endoplasm (compare above, p. 24).

_Synopsis of the Genera of Thalassosphærida._

A. Alveoles neither within { Spicula simple, 7. _Thalassosphæra_.
nor without the {
central capsule. { Spicula branched, 8. _Thalassoxanthium_.

B. Numerous large alveoles }
within the central } Spicula simple, 9. _Physematium_.
capsule (not in the }
calymma). }

C. Numerous large alveoles { Spicula simple, 10. _Thalassoplancta_.
within the calymma (not {
in the central capsule). { Spicula branched, 11. _Lampoxanthium_.

Genus 7. _Thalassosphæra_,[14] Haeckel, 1862, Monogr. d. Radiol., p. 259.

_Definition._--Thalassosphærida without alveoles, with simple, unbranched,
needle-shaped spicula in the calymma.

The genus _Thalassosphæra_ was founded by me in 1862 for those solitary
#Collodaria# in which the simple central capsule is surrounded by scattered
solid spicula. {31}I described these two different species, the new
_Thalassosphæra bifurca_ and the _Thalassosphæra morum_, which J. Müller
had formerly called _Thalassicolla morum_. This latter form is
characterised by peculiar _calcareous_ bodies "looking in outline like the
rowels of spurs, scattered irregularly in the gelatinous envelope," and was
therefore afterwards called "_Calcaromma calcarea_" by Sir Wyville
Thomson.[15] As already mentioned above, these calcareous rowels are
foreign bodies, picked up by an _Actissa_ (see p. 29). I here confine the
genus _Thalassosphæra_ to those solitary #Beloidea# in which the body
exhibits no alveoles, and the siliceous solid spicula in the calymma are
quite simple needles.

_Thalassosphæra belonium_, n. sp.

Spicula thin cylindrical rods, more or less curved, pointed at both ends,
with smooth surface (similar to the needles of _Rhaphidozoum italicum_).
Central capsule spherical, three times as large as the central nucleus,
without larger oil-globules.

_Dimensions._--Diameter of the central capsule 0.1 to 0.12, length of the
spicula 0.04 to 0.08.

_Habitat._--Central Pacific, Station 272, surface.

_Thalassosphæra rhaphidium_, n. sp.

Spicula thick cylindrical rods, more or less curved, pointed at both ends,
covered with numerous strong conical thorns, perpendicular to the axis.
Central capsule spherical, four times as broad as the central nucleus, with
twenty to thirty large oil-globules on the inside of the membrane.

_Dimensions._--Diameter of the central capsule 0.2, length of the spicula
0.12 to 0.16.

_Habitat._--Tropical Atlantic, Station 347, surface.

Genus 8. _Thalassoxanthium_,[16] Haeckel, 1881, Prodromus, p. 470.

_Definition._--Thalassosphærida without alveoles, with numerous branched or
compound spicula in the calymma.

The genus _Thalassoxanthium_ differs from the foregoing _Thalassosphæra_,
by the ramification of the spicula, and has therefore the same relation to
it as _Sphærozoum_ to _Belonozoum_. The soft unicellular body is as simple
as in _Actissa_, and exhibits alveoles neither in the capsule nor in the
calymma.

Subgenus 1. _Thalassoxanthella_, Haeckel.

_Definition._--Spicula not geminate, but simply radiate, consisting of
three, four, or more needles or shanks, radiating in different directions
from one and the same point; shanks now simple or needle-like, now furcate
or branched.

{32}1. _Thalassoxanthium triactinium_, n. sp.

Spicula all (or nearly all) triradiate, composed of three (or sometimes in
a few spicula four) needle-like shanks of equal length, diverging from one
common point. Shanks straight or somewhat curved, smooth, pointed. Central
capsule pellucid, twice as broad as its dark nucleus, without larger
oil-globules. Jelly-envelope very thin, with numerous xanthellæ.

_Dimensions._--Diameter of the central capsule 0.1, of its nucleus 0.05,
length of the spicule-shanks 0.6 to 0.8.

_Habitat._--Central Pacific, Station 266, surface.

2. _Thalassoxanthium triradiatum_, n. sp.

Spicula all (or nearly all) triradiate, composed of three (or sometimes in
a few spicula four) needle-like shanks of different length, diverging from
one common point. Shanks curved or bent, covered with small conical thorns.
Central capsule dark, three times as large as the nucleus, with numerous
large oil-globules. Jelly-envelope thick, without xanthellæ.

_Dimensions._--Diameter of the capsule 0.2, of the nucleus 0.07, length of
the spicule-shanks 0.1 to 0.15.

_Habitat._--South Pacific, Station 302, surface.

3. _Thalassoxanthium medusinum_, n. sp. (Pl. 2, fig. 5).

Spicula all (or nearly all) quadriradiate, irregular, composed of four (or
sometimes in a few spicula three) needle-like shanks (mostly of unequal
length), diverging from one common point. Shanks curved, pointed, thorny,
covered with small spinules. Central capsule dusky, twice to three times as
large as its dark central nucleus. On the inside of the membrane numerous
large oil-globules. Jelly-envelope thin, with numerous xanthellæ.

_Dimensions._--Diameter of the central capsule 0.45 to 0.5, of its nucleus
0.18 to 0.2.

_Habitat._--North Pacific, Station 244, surface.

4. _Thalassoxanthium hexactinium_, n. sp.

Spicula all (or nearly all) hexaradiate, composed of six (or sometimes in a
few spicula three) needle-like shanks, diverging in two opposite
hemispheres (three needles upwards, three needles downwards). Shanks
somewhat curved, pointed, smooth. Central capsule yellowish-brown, dark,
four times as broad as its dark central nucleus. Jelly-envelope thick
(about twice as broad as the capsule), with very numerous xanthellæ.

_Dimensions._--Diameter of the central capsule 0.3 to 0.4, of its nucleus
0.1 to 0.12.

_Habitat._--South Atlantic, Station 325, surface.

{33}5. _Thalassoxanthium cervicorne_, n. sp. (Pl. 2, figs. 3, 4).

Spicula all triradiate, trichotomously branched, with three equal shanks,
which diverge from one common point, and are again provided each with three
branches on the distal end. These nine branches are commonly once or twice
forked (each fork rarely provided with three ramules). The ramules are
thin, unequal, curved, or bent, and the ramification nearly resembles a
stag's horn. Central capsule transparent, without oil-globules, two to
three times as broad as the dark nucleus, which contains one single long
central nucleolus. Calymma thin, scarcely as thick as the radius of the
nucleus.

_Dimensions._--Diameter of the central capsule 0.2 to 0.25, of the nucleus
0.08 to 0.1, length of the spicula 0.05 to 0.15.

_Habitat._--Central Pacific, Stations 271, 274, surface.

Subgenus 2. _Thalassoxanthomma_, Haeckel.

_Definition._--Spicula all or partly geminate, consisting of one middle or
axial rod, from the two poles of which diverge two, three, or more shanks
in different directions. Shanks or needle-rays sometimes simple,
needle-like, sometimes bifurcated or branched.

6. _Thalassoxanthium furcatum_, n. sp.

Spicula all (or nearly all) geminate and simply forked, composed of a
simple axial rod and two simple, needle-like shanks on each end of it.
Shanks straight, pointed, smooth, somewhat shorter than the middle rod.
Central capsule yellowish, dark, three times as broad as its central dark
nucleus; besides this a single oil-globule, nearly of the same size.

_Dimensions._--Diameter of the central capsule 0.1, of its nucleus 0.03;
length of the axial rod of the spicula 0.04, of its shanks 0.03.

_Habitat._--Cape Verde Islands.

7. _Thalassoxanthium bifurcum_, Haeckel.

_Thalassosphæra bifurca_, Haeckel, 1862, Monogr. d. Radiol., p. 260, Taf.
xii. fig. 1.

_Sphærozoum bifurcum_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 845.

Spicula all geminate and double forked, composed of a simple axial rod and
two forked branches on each end; these branches are again forked, so that
each spiculum exhibits eight thin distal ends. All branches are thin,
slender and straight. Central capsule red, four times as broad as the
central nucleus, containing between the red pigment-granules numerous,
peculiar, violin-shaped bodies (fat-corpuscles?). Compare fig. 1, _loc.
cit._

_Dimensions._--Diameter of the central capsule 0.08, of the nucleus 0.02,
length of the spicula 0.05 to 0.08.

_Habitat._--Mediterranean (Messina), surface, Haeckel.

{34}8. _Thalassoxanthium ovodimare_, n. sp.

Spicula all geminate, composed of a simple, very short axial rod and three
diverging shanks or branches on each end of it; the shanks are very thin,
straight, or little curved, and eight to ten times as long as the axial
rod. The spicula are quite smooth, as in the similar _Sphærozoum ovodimare_
(in which, however, the axial rod is much longer). Central capsule
transparent, without oil-globules, twice as broad as the nucleus.

_Dimensions._--Diameter of the capsule 0.4, of the nucleus 0.2, length of
the spicula 0.1 to 0.2.

_Habitat._--Central Pacific, Station 273, surface.

9. _Thalassoxanthium punctatum_, n. sp.

Spicula all geminate-triradiate, composed of a simple middle rod and of
three diverging shanks on each end of it; the shanks are thorny with small
spinules and shorter than the axial rod, very similar to the common
_Sphærozoum punctatum_. Central capsule dark, with numerous (twenty to
thirty) oil-globules on the inside of the membrane, three times as broad as
the nucleus.

_Dimensions._--Diameter of the capsule 0.3, of the nucleus 0.1, length of
the spicula 0.05 to 0.2.

_Habitat._--Central Pacific, Station 274, surface.

10. _Thalassoxanthium octoceras_, n. sp. (Pl. 2, fig. 6).

Spicula all geminate-quadriradiate, composed of a simple short middle rod
and of four diverging shanks on each end of it; the shanks are quite
smooth, irregularly curved or bent, and four to eight times as long as the
middle rod. Central capsule dark, filled with pigment-granules, without
oil-globules, four times as large as the nucleus.

_Dimensions._--Diameter of the capsule 0.5, of the nucleus 0.12, length of
the spicula 0.2 to 0.4.

_Habitat._--Indian Ocean, Madagascar, Rabbe.

Genus 9. _Physematium_,[17] Meyen, 1834, Nova Acta Acad. Nat. Curios., vol.
xvi., Suppl., p. 286 (p. 162).

_Definition._--#Thalassosphærida# with large numerous alveoles within the
central capsule (not in the calymma), and with numerous simple,
needle-shaped spicula in the calymma.

The genus _Physematium_ is, together with the colony-forming _Sphærozoum_,
the first Radiolarian which was observed in the living state, described in
1834 by Meyen. It is most nearly allied to _Thalassolampe_, and has the
same large roundish alveoles within the central capsule, which reaches
therefore an extraordinary size, 5 to 10 mm. It {35}differs from the latter
by the possession of spicula in the calymma. The peculiar "centripetale
Zell-gruppen" on the inside of the capsule-membrane are probably due to
radial cleavages of the endoplasm; as also occurs in other #Collodaria#.

1. _Physematium mülleri_, Schneider.

_Physematium mülleri_, Schneider, 1858, Archiv. f. Anat. u. Physiol., p.
38, Taf. iii. B, figs. 1-5.

_Physematium mülleri_, Haeckel, 1862, Monogr. d. Radiol., p. 256, Taf.
iii. figs. 6-9.

Spherical body limpid, somewhat opalescent, sometimes a little yellowish or
brownish, rather soft. Central capsule with a thin, but firm, transparent
membrane, its diameter eight to ten times as large as that of the central
spherical nucleus. Membrane of the nucleus thick, porous, on its inside
with some nucleoli. In the protoplasmic network between the large
intracapsular alveoles, numerous large, pale, yellowish, or orange
oil-globules. On the inside of the membrane numerous conical bodies,
centripetally directed with the apex towards the centre; every conical body
("kegelförmige centripetale Zellgruppe") composed of three to nine
(commonly four or five) nucleated cells (mother-cells of the spores?); in
the axis of the cone there is often a yellowish, orange, or brown
oil-globule. Extracapsular jelly-envelope thin, with short pseudopodia.
Xanthellæ scarce or wanting. Spicula scattered in the jelly numerous,
small, simple needles, commonly C- or S-like curved, smooth.

_Dimensions._--Diameter of the whole jelly-sphere 3 to 6 mm., of the
central capsule 1 to 5 mm., of the nucleus 0.1 to 0.5, length of the
spicula 0.05 to 0.1.

_Habitat._--Mediterranean, Messina; Schneider, Haeckel; surface.

2. _Physematium atlanticum_, Meyen.

_Physematium atlanticum_, Meyen, 1834, Nova Acta Acad. Nat. Curios., vol.
xvi., Suppl., p. 286 (162), Taf. xxxviii. (xxviii.) figs. 1-3.

Spherical body opalescent, pearl-like, with a violet or purple lustre, very
soft. Central capsule with a strong, semi-transparent membrane, its
diameter six to eight times as large as that of the central spherical
nucleus. Membrane of the nucleus very thick, porous, on its inside with
many small nucleoli. In the protoplasmic network between the alveoles very
numerous, small, purple oil-globules. On the inside of the membrane a
continuous simple layer of small nuclei, enclosed in radially striped
protoplasm (mother-cells of the spores?). No centripetal conical bodies.
Extracapsular jelly-envelope very thin, with short pseudopodia. No
xanthellæ. Spicule scattered in the jelly numerous, small needles, straight
or slightly curved, thorny owing to numerous small vertical spinules. This
species and _Thalassolampe maxima_ (p. 17) possess the largest central
capsule.

_Dimensions._--Diameter of the whole jelly-sphere 8 to 12 mm., of the
central capsule 5 to 10 mm., of the nucleus 1 to 2 mm., length of the
spicula 0.1 to 0.3.

_Habitat._--Eastern Atlantic, between Canary Islands and Cape Verde
Islands, Meyen; Lanzerote, Haeckel.

{36}Genus 10. _Thalassoplancta_,[18] Haeckel, 1862, Monogr. d. Radiol., p.
261.

_Definition._--#Thalassosphærida# with numerous large alveoles in the
calymma (but not in the central capsule), and with numerous simple,
needle-shaped spicula around the central capsule.

The genus _Thalassoplancta_ was founded by me in 1862 for a Radiolarian
with simple hollow needles in the calymma, which was afterwards recognised
as a Phæodarium, belonging to _Cannorrhaphis_. We here retain this name for
a true Thalassosphærid, very similar to the latter, but distinguished by
the absence of the phæodium and the solid--not hollow--needle-shaped
spicula, which are scattered in the alveolated calymma. _Thalassoplancta_
can be regarded as the solitary form of the social _Belonozoum_.[19]

1. _Thalassoplancta longispicula_, n. sp.

Spicula long and thin, cylindrical, smooth, more or less bent, pointed at
both ends, similar to those of _Thalassoplancta cavispicula_. Central
capsule thin-walled, without oil-globules, four times as broad as the
nucleus, which encloses one single nucleolus.

_Dimensions._--Diameter of the capsule 0.6, of the nucleus 0.15, of the
calymma 4 mm.

_Habitat._--North Atlantic, Færöe Channel (Gulf Stream), John Murray.

2. _Thalassoplancta brevispicula_, n. sp. (Pl. 2, fig. 2).

_Lampoxanthium brevispiculum_, Haeckel, 1882, Atlas.

Spicula short and thick, thorny, irregularly curved, pointed at both ends,
very numerous. In the observed specimen all spicula were aggregated in the
outer part of the voluminous calymma, whilst the inner alveolated part was
devoid of them. Central capsule thick walled, with a layer of large
oil-globules on its inner surface, twice as broad as the large nucleus
which contains numerous nucleoli.

_Dimensions._--Diameter of the central capsule 0.5, of the nucleus 0.2, of
the calymma 2.5.

_Habitat._--South Atlantic, Station 334, surface.

Genus 11. _Lampoxanthium_,[20] n. gen.

_Definition._--#Thalassosphærida# with numerous large alveoles in the
calymma (but not in the central capsule), and with numerous branched or
compound spicula in the calymma.

{37}The genus _Lampoxanthium_ differs from the foregoing,
_Thalassoplancta_, by the composite form of the spicula, which are not
simple needles, but radiate or geminate, or branched in different forms;
the former stands therefore in the same relation to the latter as the
social _Belonozoum_ to _Sphærozoum_. The spicula of some species of
_Lampoxanthium_ are identical with those of some species of _Sphærozoum_,
so that the latter may be derived from the former by forming colonies. The
large central capsule is enveloped by a very voluminous alveolated calymma,
and includes a large central nucleus with numerous nucleoli.

Subgenus 1. _Lampoxanthella_, Haeckel.

_Definition._--Spicula all (or nearly all) of one kind, radiate.

1. _Lampoxanthium tetractinium_, n. sp.

Spicula all (or nearly all) tetraradiate, with four thorny, straight,
pointed shanks, radiating from one common point. (Intermingled with these
are often some few, thorny, triradiate spicula.) On the inside of the
capsule a layer of large oil-globules as in _Thalassoplancta_, Pl. 2, fig.
2.

_Dimensions._--Diameter of the central capsule 0.2, of the nucleus 0.08, of
the calymma 0.8.

_Habitat._--South Pacific, Station 288, surface.

Subgenus 2. _Lampoxanthomma_, Haeckel.

_Definition._--Spicula all (or nearly all) of one kind, geminate-radiate,
with a simple middle rod and two to four diverging shanks on each end of
it.

2. _Lampoxanthium punctatum_, n. sp.

Spicula all geminate-triradiate, thorny, of the same form as in the common
_Sphærozoum punctatum_, of which this species is the large solitary
representative. The spicula are aggregated in a very condensed layer on the
surface of the large calymma.

_Dimensions._--Diameter of the capsule 0.8, of the nucleus 0.6, of the
calymma 2.0.

_Habitat._--North Pacific, Station 248, surface.

3. _Lampoxanthium octoceras_, n. sp.

Spicula all geminate-quadriradiate, with a very short simple middle rod and
four very long divergent shanks on both ends of it; the shanks are smooth,
five to ten times as long as the middle {38}rod, irregularly bent and
curved. (Differs from the similar _Thalassoxanthium octoceras_, Pl. 2, fig.
6, by slender, more curved shanks, and by the voluminous calymma, there
entirely wanting.)

_Dimensions._--Diameter of the capsule 0.5, of the nucleus 0.2, of the
calymma 3.0.

_Habitat._--South Atlantic, Station 331, surface.

Subgenus 3. _Lampoxanthura_, Haeckel.

_Definition._--Spicula of two or three different kinds, simple, radiate,
and geminate-radiate mixed.

4. _Lampoxanthium pandora_, n. sp. (Pl. 2, fig. 1).

Spicula mixed, of three different kinds--simple, radiate and
geminate-radiate; all three kinds partly smooth, partly thorny. The simple
needles short, thin spindle-shaped, often curved. The radiate spicula
commonly with three or four, rarely five or six, unequal rays, straight or
curved. The radiate-geminate spicula commonly with three, rarely four,
shanks on each end, often different on both ends of the middle rod. The
size, number, and form of the irregular spicula are here quite as variable
as in the social _Rhaphidozoum pandora_, of which it is the solitary
representative. The wall of the large central capsule is very thick, with
evident pore-canals, separated by a clear interval from the coagulated and
vacuolated endoplasm, which contains no oil-globules. Nucleus with numerous
nucleoli.

_Dimensions._--Diameter of the central capsule 0.5 to 0.6, of the nucleus
0.1 to 0.2, of the calymma 2 to 4 mm.

_Habitat._--North Pacific, Station 244, surface.

Family IV. #SPHÆROZOIDA#, Haeckel (Pl. 4).

_Sphærozoida_, Haeckel, 1862, Monogr. d. Radiol., p. 521.

_Definition._--#Beloidea# socialia.

The family #Sphærozoida# comprises all associated or colony-forming
Radiolaria, which are provided with an imperfect skeleton, composed of
numerous solid needles or spicula, scattered around the central capsule in
the calymma. The structure and form of this skeleton is quite the same as
in the preceding solitary Thalassosphærida, but on the other hand, the
structure and form of the colonies and of the included numerous central
capsules is the same as in the skeletonless Collozoida.

The oldest well-known form of Sphærozoida is the common cosmopolitan
_Sphærozoum punctatum_, probably first observed in 1834 by Meyen, and
called _Sphærozoum fuscum_, afterwards more accurately described by Huxley
in 1851.

{39}Other forms were afterwards described by Müller and by myself in
1862.[21] Further investigations have shown me that some species of this
family are among the most common Radiolaria, and occur in astonishing
numbers on the surface of all warmer seas. But the number of species is
comparatively small, and their distinction is very difficult, as all the
different forms are very variable and connected by intermediate forms--a
truly "_transformistic_" group.

The only character sufficient for the constitution of genera in this
transformistic group is found in the form and composition of the spicula;
the very variable form of the jelly-calymma and the enclosed central
capsule being without value for this purpose. But also the form of the
spicula is very variable, and not always constant. In some species the
particular form of the spicula is transmitted by constant heredity, whilst
in others it is very inconstant, even in one and the same individual.
(Compare the remarks on variability in the general introduction.)

As the number of various forms is rather great, it seems to be advisable to
distinguish the three following genera.

_Synopsis of the Genera of Sphærozoida._

A. Spicula all of one kind, simple or needle-shaped, 12. _Belonozoum_.
B. Spicula all of one kind, branched or radiate,
or geminate, 13. _Sphærozoum_.
C. Spicula of two more different kinds, partly
simple, partly branched, 14. _Rhaphidozoum_.

Genus 12. _Belonozoum_,[22] n. gen.

_Definition._--#Sphærozoida# with simple needle-shaped spicula, which are
neither radiate nor branched.

The genus _Belonozoum_ comprises the Sphærozoida with simple needle-shaped
spicula, and may be regarded as the colonial form of _Thalassosphæra_ or
_Thalassoplancta_, derived from these solitary #Beloidea# by multiplication
of the capsules and union in a common calymma.

1. _Belonozoum bacillosum_, n. sp.

_Sphærozoum bacillosum_, Haeckel, 1881, Manuscript.

Spicula all simple rods, straight cylindrical, obtuse at both ends, quite
smooth. Central capsule pellucid, with one single central oil-globule.

_Dimensions._--Diameter of the central capsule 0.08 to 0.12, length of the
spicula 0.05 to 0.08.

_Habitat._--Central Pacific, Station 271, surface.

{40}2. _Belonozoum spinulosum_, Haeckel.

_Sphærozoum spinulosum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 54, Taf. viii. fig. 4.

_Sphærozoum spinulosum_, Haeckel, 1862, Monogr. d. Radiol., p. 527, Taf.
xxxiii. figs. 3, 4.

Spicula all simple rods, straight cylindrical, obtuse on both ends, thorny
with numerous small spines, placed vertically on the rods.

_Dimensions._--Diameter of the central capsule 0.08 to 0.1, length of the
spicula 0.05 to 0.2.

_Habitat._--Mediterranean, Nice, J. Müller; Messina, Haeckel; Naples,
Brandt; surface.

3. _Belonozoum italicum_, Haeckel.

_Sphærozoum italicum_, Haeckel, 1862, Monogr. d. Radiol., p. 526, Taf.
xxxiii. figs. 1, 2.

Spicula all simple rods, more or less curved or bent, pointed at both ends,
quite smooth. Central capsule with a variable number (commonly five to
twenty) of oil-globules.

_Dimensions._--Diameter of the central capsule 0.1 to 0.3, length of the
spicula 0.05 to 0.2.

_Habitat._--Mediterranean, Nice, Naples, Messina, Haeckel, surface.

4. _Belonozoum atlanticum_, n. sp.

_Sphærozoum atlanticum_, Haeckel, 1881, Manuscript.

Spicula all together simple rods, more or less curved or bent, pointed at
both ends, thorny from numerous small spines, placed vertically on the
rods.

_Dimensions._--Diameter of the central capsule 0.1 to 0.2, length of the
spicula 0.07 to 0.15.

_Habitat._--Tropical Atlantic, Station 348, surface.

Genus 13. _Sphærozoum_,[23] Meyen, 1834, Nova Acta Acad. Nat. Curios., Bd.
xvi., Suppl., p. 287 (p. 163).

_Definition._--#Sphærozoida# with branched or radiate spicula of one kind.

The genus _Sphærozoum_, with _Physematium_ one of the two oldest
Radiolaria, observed in the living state, was founded 1834 by Meyen for one
of the social #Beloidea#, which was probably the common cosmopolitan
_Sphærozoum punctatum_, the true type of this genus. Johannes Müller
described a number of species, which were partly skeletonless
(_Collozoum_), partly armed with simple or with compound spicula. The
species with simple spicula we refer here to _Belonozoum_, the species with
two or more different kinds of spicula to _Rhaphidozoum_, while we unite in
_Sphærozoum_ all species with one kind of branched or compound spicula. The
two following species are incompletely known:--_Sphærozoum orientale_,
Dana, 1863, _Ann. and Mag. Nat. Hist._, vol. xii. p. 54. _Sphærozoum
sanderi_, Doenitz, 1871, L. N. 60, p. 71.

{41}Subgenus 1. _Sphærozonactis_, Haeckel.

_Definition._--Spicula radiate, not geminate, consisting of three, four, or
more needles or shanks, radiating in different directions from one common
central point.

1. _Sphærozoum triactinium_, n. sp.

Spicula all (or nearly all) triradiate, composed of three (or sometimes in
few spicula four) needle-like shanks, diverging from one common point.
Shanks straight or somewhat curved, smooth, pointed. Central capsules
spherical, with one central oil-vesicle. This species may be regarded as
the social form of _Thalassoxanthium triactinium_.

_Dimensions._--Diameter of the central capsules 0.1 to 0.12, length of the
spicula-shanks 0.05 to 0.1.

_Habitat._--South Pacific, Station 295, surface.

2. _Sphærozoum medusinum_, n. sp.

Spicula all (or nearly all) quadriradiate, composed of four (or sometimes
in few spicula three) needle-like shanks (mostly of unequal length),
diverging from one common point. Shanks slightly curved, pointed, thorny,
covered with small spinules. Central capsules ellipsoidal, containing
several (four to eight) oil-vesicles. This species may be regarded as the
social form of the solitary _Thalassoxanthium medusinum_ (Pl. 2, fig. 5).

_Dimensions._--Diameter of the central capsules 0.15 to 0.18, length of the
spicula-shanks 0.08 to 0.12.

_Habitat._--North Pacific, Station 236, surface.

3. _Sphærozoum hamatum_, n. sp.

Spicula all (or nearly all) quadriradiate, composed of four (or sometimes
in few spicula three) needle-like shanks, mostly of very different size,
diverging from one common point. Shanks strong, straight, curved, or
hook-like; thorny, covered with small spinules on the distal extremity.
Central capsules ellipsoidal, large, containing many (ten to twenty)
oil-globules. This large species is distinguished by the very irregular
form and size of the spicula.

_Dimensions._--Diameter of the central capsules 0.2 to 0.25, length of the
spicula-shanks 0.12 to 0.18.

_Habitat._--Central Pacific, Station 265, surface.

4. _Sphærozoum hexactinium_, n. sp.

Spicula all (or nearly all) hexaradiate, composed of six (or sometimes in
few spicula five or seven) needle-like shanks, mostly of equal size,
diverging from one common point in two opposite hemispheres (three needles
upwards, three needles downwards). Shanks somewhat curved, pointed,
{42}smooth. Central capsules spherical, small, with one central
oil-globule. This species may be regarded as the social form of
_Thalassoxanthium hexactinium_.

_Dimensions._--Diameter of the central capsule 0.06 to 0.08, length of the
spicula-shanks 0.05 to 0.06.

_Habitat._--North Atlantic, Færöe Channel (Gulf Stream), John Murray.

Subgenus 2. _Sphærozonoceras_, Haeckel.

_Definition._--Spicula all geminate-radiate, consisting of one middle rod,
which bears an equal and constant number of rays (two, three, or four) at
each end.

5. _Sphærozoum furcatum_, n. sp.

Spicula all (or nearly all) geminate and simply forked, composed of a
simple axial rod and two simple needle-like shanks on each end of it.
Shanks straight, pointed, smooth, commonly somewhat longer than the middle
rod.

_Dimensions._--Diameter of the central capsules 0.1 to 0.15, length of the
axial rod of the spicula 0.03, of its shanks 0.04 to 0.06.

_Habitat._--Tropical zone of the Atlantic, near Ascension Island, Station
344, surface.

6. _Sphærozoum furculosum_, n. sp.

Spicula all (or nearly all) geminate and simply forked, composed of a
simple axial rod and two simple needle-like shanks on each end of it.
Shanks curved or bent, pointed, thorny, with many small spinules, commonly
somewhat shorter than the middle rod.

_Dimensions._--Diameter of the central capsules 0.2 to 0.25, length of the
axial rod of the spicula 0.1, of its shanks 0.05 to 0.08.

_Habitat._--South Atlantic, near Tristan da Cunha, Station 334, surface.

7. _Sphærozoum ovodimare_, Haeckel.

_Sphærozoum ovodimare_, Haeckel, 1862, Monogr. d. Radiol., p. 527, Taf.
xxxiii. figs. 5, 6.

_Sphærozoum punctatum_, var., Brandt, 1881, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, Taf. i. fig. 54.

Spicula all (or nearly all) geminate and triradiate, composed of a long
simple axial rod and three simple needle-like shanks on each end of it.
Shanks straight, pointed, smooth, commonly shorter than the middle rod.
(Often few furcate or four-rayed spicula are intermixed, or few spicula are
not smooth, but thorny.)

_Dimensions._--Diameter of the central capsules 0.05 to 0.2, length of the
middle rod of the spicula 0.02 to 0.06, of its shanks 0.01 to 0.04.

_Habitat._--Mediterranean, Naples, Messina, Haeckel; Atlantic, Canary
Islands, Cape Verde Islands, West Coast of Africa, Stations 351 to 354;
surface.

{43}8. _Sphærozoum trigeminum_, n. sp.

Spicula all (or nearly all) geminate-triradiate, composed of a short simple
axial middle rod and three simple needle-like shanks on each end of it.
Shanks curved or bent, very thin, smooth, commonly much longer than the
middle rod. (Often few quadriradiate or few thorny triradiate spicules are
interspersed among the others.)

_Dimensions._--Length of the middle rod of the spicula 0.02 to 0.04, of its
shanks 0.03 to 0.09.

_Habitat._--North Pacific, Stations 244 to 248, surface.

9. _Sphærozoum punctatum_, J. Müller.

_Sphærozoum punctatum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 54, Taf. viii. figs. 1, 2.

_Sphærozoum punctatum_, Haeckel, 1862, Monogr. d. Radiol., p. 528, Taf.
xxxiii. figs. 7-9.

_Sphærozoum fuscum_, Meyen, 1834, Nova Acta Acad. Nat. Cur., vol. xvi.
Taf. xxxviii. fig. 7.

_Thalassicolla punctata_, Huxley, 1851, Ann. and Mag. Nat. Hist., ser. 2,
vol. viii. p. 434, pl. xvi. figs. 1, 2, 3.

Spicula all (or nearly all) geminate-triradiate, composed of a long simple
axial middle rod and three simple needle-like shanks on each end of it.
Shanks straight, pointed, thorny, with many small spines, commonly somewhat
shorter than the middle rod. (Often few furcate or four-rayed spicula are
intermingled, or some of the spicula are smooth.) This cosmopolitan species
is extremely variable, and produces interesting transitional forms to many
other species of the genus. Compare also the general remarks on the genus,
and the chapter on "Transformation" in the general introduction.

_Dimensions._--Length of the middle rod of the spicula 0.02 to 0.06, of its
shanks 0.01 to 0.05.

_Habitat._--Cosmopolitan, common in nearly all warmer seas, Mediterranean,
Atlantic, Indian Ocean, Pacific; surface.

10. _Sphærozoum armatum_, n. sp. (Pl. 4, fig. 9).

Spicula all geminate-triradiate, with a stout and short middle rod and
three arborescent shanks on each end of it. Shanks longer than the middle
rod, very stout, straight, pine-shaped, with six to twelve irregular,
spinulated, lateral branches.

_Dimensions._--Diameter of the central capsules 0.04 to 0.08, length of the
middle rod of the spicula 0.02 to 0.03, of its shanks 0.05 to 0.08.

_Habitat._--North Pacific, Japan, Station 239, surface.

11. _Sphærozoum alveolatum_, n. sp. (Pl. 4, figs. 2, 3).

Spicula all together geminate-triradiate, with a simple stout middle rod
and three arborescent shanks on each end of it. Shanks more or less curved,
slender, pine-shaped, with four to eight short, thorny lateral branches. In
all coenobia of this remarkable species the central capsules are enclosed
in large thick-walled alveoles (of three times their breadth), and in each
alveole is placed besides {44}the capsule one single very large spiculum,
whilst the others are much smaller (fig. 3). All the alveolated capsules
are placed in one single stratum on the surface of the jelly-like spherical
coenobium, comparable to the blastoderm-cells of a blastula.

_Dimensions._--Diameter of the central capsules 0.08 to 0.1, of the
alveoles 0.2 to 0.4, length of the spicula 0.1 to 0.3.

_Habitat._--South Pacific (Juan Fernandez), Station 300, surface.

12. _Sphærozoum verticillatum_, n. sp. (Pl. 4, fig. 7).

Spicula all geminate-triradiate, with a short simple middle rod and three
much longer arborescent shanks on each end of it. Shanks straight, slender,
pine-shaped, each in the distal half with three to four elegant verticils
of thorny lateral branches.

_Dimensions._--Diameter of the capsules 0.1 to 0.12, middle rod of the
spicula 0.03 to 0.05, shanks 0.1 to 0.15.

_Habitat._--Indian Ocean, Ceylon, Haeckel; Madagascar, Rabbe; surface.

13. _Sphærozoum octoceras_, n. sp.

Spicula all geminate-quadriradiate, with a short simple middle rod and four
diverging shanks on each end of it. Shanks smooth, irregularly curved or
bent, three to six times as long as the middle rod. It may be regarded as
the social form of _Thalassoxanthium octoceras_ (Pl. 2, fig. 6).

_Dimensions._--Diameter of the capsules 0.12 to 0.16, middle rod of the
spicula 0.02, shanks 0.01.

_Habitat._--Australia, south coast, Faber; Station 163, surface.

14. _Sphærozoum quadrigeminum_, n. sp.

Spicula all geminate-quadriradiate, with a long thick middle rod and four
shorter diverging shanks on each end of it. Shanks straight, thorny.

_Dimensions._--Diameter of the capsules 0.06 to 0.08, length of the spicula
0.05 to 0.15.

_Habitat._--North Atlantic, Azores, Station 354, surface.

15. _Sphærozoum araucaria_, n. sp.

Spicula all geminate-quadriradiate, with stout straight middle rod and four
longer diverging shanks on each end of it. Shanks arborescent, with six to
twelve thorny lateral branches.

_Dimensions._--Diameter of the capsules 0.1 to 0.15, length of the spicula
0.05 to 0.1.

_Habitat._--South Atlantic, coast of Brazil, Rabbe; surface.

16. _Sphærozoum arborescens_, n. sp. (Pl. 4, fig. 8).

Spicula all geminate-quadriradiate, with a stout straight middle rod and
four longer diverging shanks on each end of it. Shanks arborescent,
pine-shaped, with four to six verticils of lateral branches, which again
are ramified and thorny.

{45}_Dimensions._--Diameter of the capsules 0.16 to 0.18, length of the
spicula 0.1 to 0.2.

_Habitat._--South Atlantic (Tristan da Cunha), Station 332, surface.

Subgenus 3. _Sphærozonura_, Haeckel.

_Definition._--Spicula all geminate-radiate, but with a different and
variable number of shanks on each end of the middle rod.

17. _Sphærozoum stellatum_, n. sp.

Spicula all geminate-radiate, with a strong middle rod and a variable
number of shorter radiating shanks on the two ends of it. Shanks straight,
nearly conical, smooth; for the most part three or four shanks on each end,
but sometimes also five or six; very often this number is unequal on the
two ends.

_Dimensions._--Diameter of the central capsules 0.1 to 0.2, length of the
spicula 0.05 to 0.15.

_Habitat._--Central Pacific, Station 270, surface.

18. _Sphærozoum geminatum_, n. sp. (Pl. 4, fig. 4).

Spicula all geminate-radiate, with a strong middle rod and a variable
number of longer radiant shanks on each end of it. Shanks straight,
conical, in the distal half thorny; commonly either three or four shanks on
each end of the middle rod, often also three on one end, four on the other
end; rarely five or six rays on one end.

_Dimensions._--Diameter of the capsules 0.15 to 0.2, length of the spicula
0.05 to 0.1.

_Habitat._--Indian Ocean, Ceylon, Haeckel; surface.

19. _Sphærozoum circumtextum_, n. sp.

Spicula all geminate-radiate, with a very variable number of rays (two to
six) on each end of the thin middle rod. All spicula very thin and
delicate, smooth, with curved or bent shanks, densely covering the central
capsule like a cobweb. The number of rays on each end is usually different,
generally four or five, often also two or three, rarely six.

_Dimensions._--Diameter of the capsule 0.1 to 0.2, length of the spicula
0.04 to 0.12.

_Habitat._--Southeast part of the Indian Ocean, Station 160, surface.

20. _Sphærozoum variabile_, n. sp. (Pl. 4, fig. 5).

Spicula all geminate-radiate, with a short middle rod and a variable number
of shanks on each end of it. Shanks four to eight times as long as the
middle rod, curved or bent, in the distal half thorny; their number is
commonly different on the two ends of it, three or five, often also four or
six, rarely two; their size and form very variable.

{46}_Dimensions._--Diameter of the capsules 0.1 to 0.3, length of the
spicula 0.1 to 0.2.

_Habitat._--North Pacific, Station 248, surface.

Genus 14. _Rhaphidozoum_,[24] Haeckel, 1862, Monogr. d. Radiol., p. 529.

_Definition._--Sphærozoida with two or more different kinds of spicula; one
kind simple, needle-shaped; the other kinds compound, radiate, or branched.

The genus _Rhaphidozoum_ differs from _Sphærozoum_ by the composition of
the skeleton of two or more different kinds of spicula, and has therefore
the same relation to it as the solitary _Lampoxanthura_ to
_Lampoxanthella_.

In some species nearly all the different forms, which characterise the
numerous species of #Beloidea#, may be united in one and the same
individual.

Subgenus 1. _Rhaphidonactis_, Haeckel.

_Definition._--Spicula of two different kinds; one kind simple,
needle-shaped, the other radiate (composed of three, four, or more shanks,
diverging from one common point).

1. _Rhaphidozoum pelagicum_, n. sp.

Spicula of two different kinds; one kind simple thin needles, a little
curved or bent, the other kind triradiate, with three thin, curved shanks.
Both kinds smooth, without thorns. Resembles a combination of _Belonozoum
italicum_ and _Sphærozoum triactinium_.

_Dimensions._--Diameter of the central capsules 0.1 to 0.12, length of the
spicula 0.05 to 0.15.

_Habitat._--Central Pacific, Station 267, surface.

2. _Rhaphidozoum pacificum_, n. sp.

Spicula of two different kinds; one kind simple needles, stout and
straight, pointed at both ends, the other kind triradiate, with three
straight and stout shanks. Both kinds thorny.

_Dimensions._--Diameter of the central capsule 0.06 to 0.08, length of the
spicula 0.05 to 0.1.

_Habitat._--Central Pacific, Station 271, surface.

3. _Rhaphidozoum acuferum_, Haeckel.

_Rhaphidozoum acuferum_, Haeckel, 1862, Monogr. d. Radiol., p. 529, Taf.
xxxii. figs. 9-11.

_Sphærozoum acuferum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 54, Taf. viii. fig. 3.

_Thalassicolla acufera_, J. Müller, 1855, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 237.

Spicula of two different kinds, simple needles and quadriradiate; both
strong, thorny, covered with small spinules. Simple needles mostly curved,
C-shaped. Four shanks of the quadriradiate {47}spicula now straight, now
curved, commonly of very different size. (Often one single quadriradiate
spiculum is distinguished by its extraordinary size.) For the detailed
description of this species compare my Monograph (_loc. cit._).

_Dimensions._--Diameter of the central capsules 0.05 to 0.35, length of the
simple needles 0.05 to 0.25, shanks of the quadriradiate spicula 0.05 to
0.15.

_Habitat._--Mediterranean, Messina, Naples, Nice.

4. _Rhaphidozoum arachnoides_, n. sp.

Spicula of two different kinds; one kind simple, needle-like, the other
quadriradiate; both very thin and slender, smooth, without spicules. Simple
needles curved, C-shaped. Four shanks of the quadriradiate spicula also
curved, commonly of nearly equal size. The numerous thread-like spicula of
this species are so densely packed around the central capsule, that they
extend all around its surface like the network round a balloon.

_Dimensions._--Diameter of the central capsules 0.12 to 0.15, length of the
simple needles 0.1 to 0.12, shanks of the quadriradiate spicula 0.06 to
0.08.

_Habitat._--Tropical Atlantic, Station 345, surface.

5. _Rhaphidozoum asperum_, n. sp.

Spicula of two different kinds; one kind simple, needle-shaped, stout, and
straight, the other kind hexaradiate; its six shanks about half as long as
the former, conical. Both kinds very thorny, covered with short conical
spinules.

_Dimensions._--Diameter of the capsules 0.06 to 0.08, length of the simple
needles 0.05 to 0.07, shanks of the hexaradiate spicula 0.03 to 0.04.

_Habitat._--South Pacific, Station 288, surface.

Subgenus 2. _Rhaphidoceras_, Haeckel.

_Definition._--Spicula of two different kinds; one kind simple,
needle-shaped, the other kind geminate-radiate, with rays on both poles of
a middle rod.

6. _Rhaphidozoum neapolitanum_, Haeckel.

_Sphærozoum neapolitanum_, C. Brandt, 1881, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 390, Taf. i. figs. 14, 16-18.

Spicula mixed, of two different kinds; simple needles and geminate-forked.
Simple rods, like those of _Belonozoum italicum_, more or less curved,
pointed at both ends, smooth (sometimes a little thorny at both ends).
Geminate spicula simply forked, like those of _Sphærozoum furcatum_,
composed of a short, simple, axial rod, and two simple, smooth, straight
shanks on each end of it, commonly somewhat longer than the middle rod.
This species, which I have observed myself in Spezzia in great quantity, is
quite as variable as all the other species of the genus, and has not more
claim to specific rights than the others. Commonly the simple needles are
much more numerous {48}than the geminate-forked, but sometimes the contrary
is the case. On their variability compare the general remarks on the genus,
and the chapter on "Transformation" in the general introduction.

_Dimensions._--Length of the simple spicula 0.05 to 0.1, of the middle rod
of the forked spicule 0.05 to 0.08, of their shanks 0.01 to 0.03.

_Habitat._--Mediterranean, Naples, Spezzia, surface.

7. _Rhaphidozoum patagonicum_, n. sp.

Spicula mixed, of two different kinds, simple needles and
geminate-triradiate. Simple rods, like those of _Belonozoum spinulosum_,
straight, thorny, pointed at both ends. Geminate spicula double-triradiate,
like those of _Sphærozoum punctatum_, composed of a simple, short, axial
rod and three simple pointed shanks on each end of it. Shanks straight,
thorny, with many small spinules, commonly somewhat longer than the middle
rod. (Often some of the spicula of both kinds are smooth, not thorny, or
not straight, but a little curved, or a few forked or four-radiated
geminate spicula are mingled with the others.)

_Dimensions._--Diameter of the central capsules 0.08 to 0.2, length of the
simple spicula 0.1 to 0.15, of the geminate 0.08 to 0.16.

_Habitat._--South Pacific, west coast of Patagonia, Station 302, surface.

8. _Rhaphidozoum ascensionis_, n. sp.

Spicula mixed, of two different kinds, simple needles and
geminate-triradiate; both kinds thin, smooth, without spinules. Simple
needles somewhat curved, C- or S-shaped. Geminate spicula
double-triradiate, composed of a simple, short, straight axial rod and
three slender curved shanks on each end of it. Shanks two to four times
longer than the middle rod. (Sometimes few simple hexaradiate and geminate
tetraradiate spicula are mingled.)

_Dimensions._--Diameter of the central capsules 0.12 to 0.15, length of the
simple spicula 0.1, of the double-triradiate 0.05 to 0.2.

_Habitat._--South Atlantic, off Ascension Island, Station 342, surface.

9. _Rhaphidozoum capense_, n. sp.

Spicula of two different kinds; one kind simple, needle-shaped, straight,
pointed at both ends, the other kind geminate-quadriradiate, with a stout
short middle rod and four longer bent shanks on each end of it. Both kinds
smooth.

_Dimensions._--Diameter of the capsules 0.2 to 0.25, length of the simple
needles 0.01 to 0.3, of the geminate spicula 0.05 to 0.15.

_Habitat._--Cape of Good Hope (Agulhas), Station 142, surface.

10. _Rhaphidozoum australe_, n. sp.

Spicula of two different kinds; one kind simple, needle-shaped, curved,
thin; the other kind geminate, with a variable number of shanks on both
ends of the shorter middle rod, often {49}different on the two poles of it.
The prevalent number of rays on each end is three or four, often also two
or five, rarely six. All spicula smooth, more or less bent.

_Dimensions._--Diameter of the capsules 0.1 to 0.2, length of the spicula
0.05 to 0.15.

_Habitat._--South West Pacific, Station 165, surface.

Subgenus 3. _Rhaphidonura_, Haeckel.

_Definition._--Spicula of three different kinds: one kind simple,
needle-shaped; the second kind radiate, with three to six shanks radiating
from a common central point; the third kind geminate-radiate, with rays on
both poles of a middle rod.

11. _Rhaphidozoum polymorphum_, n. sp.

Spicula of three different kinds; simple needles, radiate, and geminate
mixed. The simple needles straight and stout. The radiate spicula commonly
with three or six, rarely four or five, rays. The geminate-radiate spicula
prevalent, with three or four, rarely two or five, shanks on each end of
the middle rod. Number very variable. All shanks straight and smooth.

_Dimensions._--Diameter of the capsule 0.1 to 0.2, length of the spicula
0.05 to 0.15.

_Habitat._--South Pacific, Station 295, surface.

12. _Rhaphidozoum pandora_, n. sp. (Pl. 4, fig. 6).

Spicula of three different kinds; simple needles, radiate and geminate
mixed. The simple needles thin spindle-shaped, often curved. The radiate
spicula commonly with three or four, rarely five or six, curved rays. The
geminate-radiate spicula commonly with three or four, rarely two or five,
shanks on each end, often different on the two ends of the middle rod.
Number and form very variable. All or most of the shanks more or less bent
and thorny.

_Dimensions._--Diameter of the capsule 0.1 to 0.3, length of the spicula
0.05 to 0.2.

_Habitat._--South Atlantic (near Ascension Island), Station 343, surface.

----

Order II. SPHÆRELLARIA, Haeckel, 1881.

_Sphærellaria_, Haeckel, 1881, Prodromus, p. 421.
_Sphæridea_ vel _Peripylea_, Hertwig, 1879, Organismus der Radiol., p.
133.

_Definition._--SPUMELLARIA with latticed or spongy shell.

The order #Sphærellaria#, the second order of Radiolaria, comprises all
those SPUMELLARIA in which the skeleton is a latticed or fenestrated, often
more or less spongy, siliceous shell. Originally this shell is a simple
extracapsular lattice-sphere, in which the central capsule is included;
from this simple ancestral form an enormous {50}mass of different and often
very complicated forms is derived; this order is by far the largest, and in
morphological respects the most important and most interesting, of all
Radiolaria. It contains not less than twenty-eight different families,
three hundred and five genera, and more than sixteen hundred species.

In my Monograph (1862) seven families appertaining to this group are
described--the Ethmosphærida, Cladococcida, Ommatida, Spongurida, Discida,
Lithelida, and Collosphærida. The astonishing increase of this group by the
detection of a large series of new and interesting forms, and particularly
of important connecting forms between very different branches of it, now
enables me to give a much better arrangement. I discern now four suborders
or sections of #Sphærellaria#, according to the different geometrical form
of the central capsule and of the latticed shell enveloping it. The first
of these, and the common ancestral group of the whole order, is the
#Sphæroidea#, with spherical capsule; in the #Prunoidea# it becomes
ellipsoidal or cylindrical by prolongation of one axis; in the #Discoidea#
lenticular or discoidal by shortening of one axis; in the #Larcoidea#
lentelliptical, or triaxon-ellipsoid, by different growth of the capsule in
three different "dimensive axes."

_Synopsis of the Four Suborders of_ #Sphærellaria#.

Central capsule { Shell a simple sphere or a
spherical. { system of concentric spheres, 1. #Sphæroidea#.

Central capsule { Shell a simple ellipsoid or a
ellipsoidal or { cylinder with annular
cylindrical. { transverse constrictions, 2. #Prunoidea#.

Central capsule
lenticular or { Shell a biconvex lens or a
discoidal. { flat disk, 3. #Discoidea#.

Central capsule
lentelliptical or { Shell a triaxon-ellipsoid, with
triaxon. { three different axes, 4. #Larcoidea#.

----

Suborder I. SPHÆROIDEA, Haeckel.

_Sphæroida_, _Sphæridea_, _Sphærida_, Haeckel, 1878, Protistenreich, p.
103.
_Sphæridea_, R. Hertwig, 1879, Organismus der Radiol., p. 39.

_Definition._--SPUMELLARIA with spherical central capsule (very rarely
somewhat modified, or allomorphous); with spherical fenestrated siliceous
shell (often an endospherical polyhedron, very rarely of more modified,
subspherical form or allomorphous). Growth of the shell in the three
dimensive axes equal.

The suborder #Sphæroidea#, the first and most important of the four of the
#Sphærellaria#, comprises those SPUMELLARIA in which the original
geometrical {51}spherical form is quite constantly preserved in the central
capsule, and commonly also in the fenestrated shell enveloping the latter,
although in many forms the sphere is more or less modified; very frequently
it is an "endospherical polyhedron," _i.e._, a polyhedron all the angles
(or the nodes of the network) of which lie upon the surface of a
geometrical sphere; more rarely the spherical form is more or less modified
and irregular. In the great majority of #Sphæroidea# there is no external
indication of the three dimensive axes; but in many forms they are
indicated by the regular position of certain external radial spines or
internal radial beams. However, in no case are those three axes expressed
in the form of the shell itself and of the enclosed spherical central
capsule; this is the main character by which the #Sphæroidea# differ from
the following sections:--#Prunoidea#, #Discoidea#, #Larcoidea#, all three
of which arise from them.

The section #Sphæroidea#, in the sense here restricted, was founded by me
in my Protistenreich (1878, p. 103) and adopted by Hertwig (1879) in his
Organismus der Radiolarien (p. 39). The different groups appertaining to
this large section were characterised more accurately in my Prodromus
(1881, pp. 448-456); there I gave the characters of six subfamilies with
thirty tribes, containing ninety-three genera. Formerly, in my Monograph
(1862), the #Sphæroidea# were disposed in five different
families:--Ethmosphærida, Cladococcida, Ommatida, Spongosphærida,
Collosphærida. At that time I could not separate them sufficiently from
some ACANTHARIA and PHÆODARIA, which have a similar spherical
lattice-shell.

As the number of different genera and species in the #Sphæroidea# is much
greater than in all other sections of SPUMELLARIA, many forms were already
described by former authors. In the oldest system of Ehrenberg (1847, _loc.
cit._, p. 53) they represent one part of his Haliommatina (with four
genera, _Haliomma_, _Chilomma_, _Stylosphæra_, _Spongosphæra_). Most
species, however, of these genera are #Discoidea#. Also in the latest
system of Ehrenberg (1875, _loc. cit._, p. 157) his Haliommatina are a
confused conglomeration of different SPUMELLARIA (#Sphæroidea#,
#Discoidea#, and #Prunoidea#).

The section #Sphæroidea# is the largest division of #Sphærellaria#,
comprising not less than one hundred and seven genera and six hundred and
fifty species. This enormous number (easily to be augmented by further
investigations) requires a careful disposition in different families and
subfamilies. For this disposition two different principles only can be
employed: firstly, the number and disposition of the _radial spines_;
secondly, the number of the _concentric latticed spheres_, which are
connected by radial beams. I give here the preference to the first
principle, whilst in my Prodromus (1881) I had preferred the second. The
question, which of the two principles is more important for the
classification of #Sphæroidea#, is very difficult to answer; probably in
many cases the former, in many the latter is more important for their
phylogeny.

{52}Regarding the number of the concentric shells which compose the
latticed carapace of the #Sphæroidea#, we can distinguish six families,
viz.:--

I. Monosphærida (with one single shell).
II. Dyosphærida (with two concentric shells).
III. Triosphærida (with three concentric shells).
IV. Tetrasphærida (with four concentric shells).
V. Polysphærida (with five or more concentric shells).
VI. Spongosphærida (with spongy shells).

On the other hand, regarding the number of the radial spines and their
regular disposition on the shell-surface, we can distinguish five families,
viz.:--

I. Liosphærida (without radial spines).
II. Stylosphærida (with two radial spines, opposite in one axis).
III. Staurosphærida (with four radial spines, opposite in pairs in two
axes, perpendicular one to another).
IV. Cubosphærida (with six radial spines, opposite in pairs in the
three dimensive axes).
V. Astrosphærida (with numerous--eight, twelve, twenty, or
more--radial spines, often more than a hundred).

All five latter groups contain representatives of all six former groups;
therefore we get together not less than thirty different subfamilies of
#Sphæroidea#, already enumerated in my Prodromus, 1881, p. 449. I repeat
them here to give a better survey of the system there employed.

{53}The #Monosphærida# comprise all those #Sphæroidea# in which the
carapace is represented only by one single lattice-shell. Originally this
shell is probably everywhere an extracapsular or "cortical shell," which is
developed on the outside of the jelly-veil enveloping the central capsule,
and serves as a protective carapace for these soft enclosed parts. But with
the progress of growth the central capsule becomes larger than the
including shell, and sends out through its pores club-shaped prolongations
or cæcal-sacs (Pl. 11, figs. 1, 5; Pl. 19, figs. 2, 3, 5; Pl. 20, fig.
1_a_; Pl. 27, fig. 3). These protruded sacs may fuse together again outside
the shell and form a spherical bladder, now enveloping the smaller shell;
the latter now becomes an intracapsular or "medullary shell."

As #Pliosphærida# (or _Sphæroidea concentrica_) we can oppose to the simple
Monosphærida all other #Sphæroidea#, the lattice-shell of which is composed
of two or more concentric shells, connected by radial beams. Probably all
Pliosphærida (or at least the greater part of them) arise from the
Monosphærida by centrifugal growth; two or more radial spines are developed
from the surface of the simple lattice-sphere, and are united together by
communicating lateral branches, developed at equal distances from the
centre; and this same process may be repeated, two, three, four, or more
times. In this way originate the characteristic systems of concentric
spheres, all united by piercing radial beams which arise from the surface
of the innermost sphere (not from its centre). Regarding this mode of
growth, we can distinguish the innermost as "original" or "primary" shell,
and all subsequent ones as "apposed" or "secondary" shells; if the number
of concentric shells amount to three or more, commonly both innermost
shells lie within the central capsule and are medullary shells, whilst all
others lie outside it and are therefore cortical shells. This difference
can be commonly recognised also in the isolated shell, without its central
capsule; the distance between the cortical and the medullary shells being
commonly much larger than the distance between the two medullary shells.

The #Dyosphærida#, or the #Sphæroidea# with two concentric shells, are the
most numerous among the Pliosphærida. Commonly in this group the inner or
primary shell lies within the central capsule as a true "medullary shell,"
whilst the outer lies outside it as a "cortical shell"; therefore the
radial beams, connecting both, pierce the wall of the capsule. But in
several forms, mainly in the peculiar group of Diplosphærida, both
concentric shells remain outside the central capsule, and both are
therefore "cortical shells."

The #Triosphærida#, or the #Sphæroidea# with three concentric shells, are
also very rich in different forms, though not so numerous by far as the
Dyosphærida. Commonly in the Triosphærida both inner shells lie within the
central capsule as "medullary shells," whilst the third lies outside it as
a "cortical shell"; therefore the central capsule remains intermediate in
size between the outer and the middle shell. But in some genera (_e.g._,
_Rhodosphæra_) both outer shells are cortical and only the {54}innermost is
a medullary shell. In this case the size of the capsule remains
intermediate between the inner and the middle shell.

The #Tetrasphærida#, or the #Sphæroidea# with four concentric shells, are
in general not frequent, and not rich in different forms. In most of the
observed species two inner shells are medullary, two outer cortical shells,
the former within, the latter without, the central capsule; and the wall of
the capsule, pierced by the connecting radial beams, lies between the two
middle shells. But there are some Tetrasphærida in which all four shells
seem to be external or cortical shells.

The #Polysphærida#, or the #Sphæroidea# with five or more concentric
shells, seem of course to offer the greatest possibility for the
development of very different forms; but in reality this group is the
poorest and smallest of all; and only one part of it, the Arachnosphærida,
is rather common. In this peculiar division the shell is composed of five
to ten or more, very delicate, cobweb-like concentric shells, which are
connected by radial beams; all are cortical shells, and lie outside the
central capsule. Much more rare are those Polysphærida, in which both
innermost shells, as true medullary shells, lie within the central capsule,
all others being outside it. The total number of concentric shells in this
group is commonly between five and ten, rarely more.

The #Spongosphærida# are distinguished from all other #Sphæroidea# by the
spongy structure of the spherical shell, which is composed wholly or
partially of an irregular spongy framework. The relation of this group to
the other groups of #Sphæroidea# is probably rather complicated, for in
some Spongosphærida the whole shell is composed of massive spongy
reticulation, whilst in others it contains a spherical central cavity, and
in a third group this cavity is filled up by one or two concentric
lattice-shells, connected by radial beams. Many of these Spongosphærida are
very common, and of considerable size.

The #Collosphærida# form a peculiar separate group of #Sphæroidea#,
distinguished from all others by their social life or aggregation in
colonies (coenobia). They represent the only group of #Sphærellaria# in
which this association of numerous individual capsules or cells is
realised. The shell is almost constantly simple, without regularly disposed
radial spines; therefore they may be called "social Monosphærida," or
better "polyzoic Ethmosphærida." Only in one small group (Clathrosphærida)
the shell, enveloping every central capsule, is double or surrounded by an
external mantle; these may be compared to the Diplosphærida (or better to a
part of the Carposphærida, _Liosphæra_, p. 76). In most of the
Collosphærida the lattice-shell is more or less irregular in form and
structure.

_The Lattice Work_ of the fenestrated shells is in the #Sphæroidea# of the
greatest variability, and its innumerable modifications serve mainly for
the distinction of species. In general we can distinguish as the most
important modifications a _regular_ network (with equal size, form, and
distance of the pores or meshes) and an _irregular_ network (with
{55}differences in the size, form, or distance of the meshes or pores). In
both groups the pores may be either angular or round; so that there may
exist together four different main forms of network--(A) regular lattice
with equal hexagonal pores; (B) regular lattice with equal circular pores;
(C) irregular lattice with unequal polygonal pores; (D) irregular lattice
with unequal roundish pores. Besides these modifications, the pores may be
prolonged into tubules which are directed radially towards the outside
(rarely towards the inside) of the sphere. In other cases they are
surrounded by elevated or honeycomb-like frames.

_The Radial Spines_ exhibit in the #Sphæroidea# the greatest variety in
form, size, disposition, &c., and their numerous modifications serve mainly
for the distinction of genera, their peculiar formation and size also for
the distinction of species. In general we may distinguish as the most
important modifications primary and secondary spines. The primary spines or
"main spines" are commonly direct outward prolongations of the internal
radial beams, connecting the concentric shells. The secondary or
"by-spines" arise only from the surface of the lattice-shell, without
reference to the internal beams. The by-spines are commonly smaller, and
much more numerous than the main spines. Regarding the form, the radial
spines are either roundish (cylindrical or conical, often also club-shaped,
rarely spindle-shaped) or angular (commonly three-sided, prismatic or
pyramidal). The spines are constantly solid, never hollow; the "internal
canals," described by some authors, are only microscopic views of the
transparent edges. In many cases the spines are branched or forked. The
most important difference in the variable shape of the spines is their
regular or irregular number and disposition, which afford characters for
the distinction of our five families.

_The Three Dimensive Axes_--or the three diameters of the sphere,
perpendicular one to another--are in the great majority of the #Sphæroidea#
significant in the promorphological consideration of the body, and are
indicated either by the position of the external radial spines, or at least
of the internal radial beams, connecting the concentric spheres. Commonly
two radial spines are placed opposite in each axis. The most perfect group
in this respect seems to be that of the Cubosphærida, in which the three
axes are represented by three pairs of spines. Next come the
Staurosphærida, in which two axes in cross-form are exhibited by two pairs
of spines. The most simple group are the Stylosphærida, in which only one
pair of spines is developed, indicating one single axis. These three
families form together a continuous natural series,--the #Sphæroidea# with
real dimensive axes,--and exhibit at the same time relations to the three
other suborders of #Sphærellaria#, the #Larcoidea#, #Discoidea#, and
#Prunoidea# respectively. At both ends of this series stand two other
families, on one side the Liosphærida, without any radial spines on the
surface of the sphere, on the other side the Astrosphærida, in which the
radial spines are developed in great and variable numbers, at least eight
to twelve, commonly twenty to forty, often more than a hundred or even a
thousand.

{56}The #Liosphærida# comprise all those #Sphæroidea# in which the surface
of the shell is smooth, without radial spines (Pls. 12, 20). The simplest
of these are the Ethmosphærida, with one single lattice-shell, enveloping
the spherical central capsule. _Cenosphæra_, the most simple form of the
Ethmosphærida, may be regarded as the common ancestral form of all
#Sphæroidea#, in an ontogenetical as well as in a phylogenetical and
morphological sense. From this simple lattice sphere all other #Sphæroidea#
can be derived either by radial or by tangential growth. If the radial
beams, arising from the surface of the simple fenestrated sphere, become
connected (at equal distances from the centre) by tangential beams, we get
the compound shells of the "Liosphærida concentrica" (with two, three,
four, or more concentric spheres). The radial beams connecting these
exhibit in many Liosphærida the same regular disposition and number as the
external radial spines in the Astrosphærida. Perhaps these forms in a
"natural system" would be better united (_e.g._, Liosphærida with twelve or
twenty internal radial beams, and Astrosphærida with twelve or twenty
external radial spines); but in many cases (mainly for higher numbers) the
certain determination of their number and disposition is very difficult or
quite impossible.

The #Cubosphærida# (Pls. 21-25) represent the large and very important
family of #Sphæroidea#, in which all three dimensive axes are equally
distinguished by pairs of spines, corresponding to three axes of a cube or
of a regular octahedron, agreeing therefore also with the three axes of the
cubic or regular crystalline system. In the majority of the Cubosphærida
the six radial spines are accurately opposite each other in pairs in three
axes, perpendicular one to another, and commonly they are of equal size and
form; but in some genera the three pairs of spines become differentiated,
whilst both spines of each pair remain equal. Either one pair is larger
than the two others (which are equal), corresponding to the axes of the
quadratic crystalline system; or all three pairs are different
(corresponding to the three unequal axes of the rhombic crystalline
system); the former nearer to the #Discoidea#, the latter to the
#Larcoidea#. We may suppose with some probability, that the Cubosphærida
are for the most part the common ancestral group of those #Sphæroidea#, in
which a certain number of radial spines or beams is disposed in a regular
order; the Staurosphærida may be derived from them by loss of one pair of
spines, the Stylosphærida by loss of two pairs of spines, and most
Astrosphærida by multiplying the radial spines, six to fourteen or more
secondary spines being added to the six primary "dimensive spines."
However, in many Astrosphærida (_e.g._, in those with eight spines,
_Centrocubus_, _Octodendron_, &c.) the regular geometrical disposition of
the radial spines seems to follow another mathematical order, quite
independent of the Cubosphærida.

The #Staurosphærida# (Pl. 15) are distinguished by the possession of four
radial spines, opposite in pairs in two axes, perpendicular one to another.
This rectangular cross determines a certain plane, the "equatorial plane,"
and this brings the Staurosphærida near {57}to the #Discoidea#, mainly to
those which also bear on the periphery of the circular equatorial plane
four crossed spines (such as _Staurodisculus_, _Stethostaurus_,
_Staurodictya_, &c.). But in these cruciform #Discoidea# the shell and the
enclosed central capsule are discoidal or lenticular, whilst in the
Staurosphærida they remain spherical. Commonly the cross is quite regular,
with four right angles and four equal beams; but often also it becomes more
or less irregular. In some genera one pair of equal opposite spines is
larger than the other pair. These forms represent the three different axes
of the rhombic crystal system, whilst the common regular Staurosphærida
represent those of the quadratic crystal system. The latter can be derived
from the Cubosphærida (representing the regular crystal system) by
reduction of one axis and loss of its pair of spines. In general the number
of species (and particularly of the individuals) is much smaller in the
Staurosphærida than in all other families of #Sphæroidea#.

The #Stylosphærida# (Pls. 13-17) can be derived from the Cubosphærida by
reduction of two dimensive axes and loss of two pairs of spines. Therefore,
here one pair of spines only is developed, opposite in one single axis.
This "monaxonial" form brings the Stylosphærida very near to the
ellipsoidal #Prunoidea# (mainly to many two-spined forms of Ellipsida and
Druppulida); but they differ from these by the spherical (not ellipsoidal)
form of the central capsule and of the enclosing lattice-shell. In the
greater part of the Stylosphærida both spines are of equal size and form,
accurately opposite in the "main axis." But in many forms both spines
become unequal in size or form, often very different. More rarely they are
not accurately opposed, but placed in two different axes, intersecting at a
small variable angle. The small group of Saturnalida presents a very
remarkable and peculiar structure, in which both spines (at equal distances
from the centre) are united by a circular or elliptical ring (Pl. 13, figs.
15, 16; Pl. 16, figs. 16, 17).

The #Astrosphærida# are distinguished from the other #Sphæroidea# by the
great and variable number of their external radial spines (Pls. 11, 18-20,
26-30). Commonly this number amounts to from twelve to twenty, rarely to
only eight to ten, very often to thirty-two to forty or more; in many
species more than one hundred are present. As already mentioned above, it
would be important to distinguish between primary spines (as outer
prolongations of the inner radial beams) and secondary spines (developed
from the surface of the shell), but in many cases this distinction is
difficult or impossible. More practical is the distinction between larger
"main spines" and smaller "by-spines." The size and form of the radial
spines is extremely variable. Much more important is their number and
disposition. In general we can here distinguish the following different
cases:--(A) radial spines are developed from all the nodal points of the
network on the shell surface; (B) the number of the spines is smaller than
that of the nodal points, but they are irregularly scattered; (C) the
radial spines exhibit a limited number and a certain regular disposition.
In this latter case the following modes of distribution seem to be the most
important:--(_a_) eight spines placed in the four diagonal axes of the
{58}regular cube (Pl. 18, figs. 1-3); (_b_) twelve spines (placed in the
corner axes of the regular icosahedron); (_c_) fourteen spines (six placed
in the three dimensive axes of the regular octahedron, eight in the centres
of its eight faces); (_d_) twenty spines (placed either in the same order
as in many #Larcoidea# and ACANTHARIA [?], or in the twenty corners of the
regular dodecahedron); (_e_) thirty-two spines (twelve placed in the twelve
corners of the regular icosahedron, twenty in the centre of its triangular
faces). Besides these most important and quite geometrical modes of
disposition there also seem to occur in the Astrosphærida the following
subregular (or symmetrical?) modes: 9, 10, 16, 18, 24, 40, 60, 80. But it
is very difficult to give a correct account of these modes. In every case
this manifold and regular disposition of the radial spines is of the
highest interest for the study of general "Promorphology."

_The Central Capsule_ is in all #Sphæroidea# (without any exception) a
perfect sphere in the geometrical sense, even in those forms in which the
enclosing lattice-shell is more or less irregular (_i.e._, many
Collosphærida). This is the most important character, which separates the
#Sphæroidea# from all other #Sphærellaria#. For in the #Prunoidea# the
capsule is ellipsoidal, with one prolonged axis; in the #Discoidea#
lenticular, with one shortened axis; in the #Larcoidea# lentelliptical,
with three different dimensive axes. The central capsule is originally
always enclosed by the lattice-shell; but in many cases with increasing
growth this relation becomes inverted; the capsule sending out many
club-shaped blind sacs through the meshes of the lattice-shell, and these
melting together outside the latter, a new membrane is formed, enclosing a
"medullary shell."

_The Nucleus_ of the cell exhibits a very different shape in the solitary
and the social #Sphæroidea#. In the solitary or monozoic #Sphæroidea# the
centre of the central capsule is occupied by a large spherical concentric
nucleus, with or without nucleoli; also this nucleus is originally always
within the innermost lattice-shell, but with increasing size may overgrow
and enclose it. A short time before the formation of the vibratile spores
the central nucleus becomes resolved into many small nuclei. In the social
or polyzoic #Sphæroidea#--the Collosphærida--commonly the simple central
nucleus very early (a long time before the formation of the spores) is
divided into a great number of small nuclei, whilst the centre of the
capsule becomes filled with a large oil-globule. Therefore we find the same
difference between the solitary and social forms in the #Sphæroidea# as in
the #Colloidea#. Here also the calymma, or the jelly-mantle, enveloping the
central capsule, is in the social forms very large and voluminous,
differentiated into alveoles, whilst in the solitary forms it is much
smaller, without alveoles.

{59}_Synopsis of the Families of_ #Sphæroidea#.

{ A. Liosphærida { Spherical shell
{ monozoa. { commonly
{ Single cells { quite regular,
{ (each with shell) { simple, or
Surface of { living solitary. { composed of
the spherical { { two or more
shell smooth, { { concentric
rough, or { { spheres, 5. LIOSPHÆRIDA.
thorny, but {
not armed { B. Liosphærida { Spherical shell
with radial { polyzoa. { commonly
spines. { Aggregated cells { more or less
{ (each with { irregular,
{ shell) living { simple (rarely
{ in colonies. { composed of
{ { two concentric
{ { spheres), 6. COLLOSPHÆRIDA.

Surface of { Two radial main-spines, opposite in
the spherical { one axis of the shell 7. STYLOSPHÆRIDA.
shell armed {
with two, { Four radial main-spines, opposite in
four, or six { pairs in two dimensive axes,
radial main { perpendicular one to another, 8. STAUROSPHÆRIDA.
spines, {
opposite in { Six radial main-spines, opposite in
pairs in one, { pairs in three dimensive
two, or three { axes (perpendicular one to
dimensive { another), 9. CUBOSPHÆRIDA.
axes (always {
solitary). {

Surface of the spherical shell covered with numerous
(commonly irregularly disposed) radial spines,
often also twelve to twenty, more or less regularly
disposed, 10. ASTROSPHÆRIDA.

Family V. #LIOSPHÆRIDA#, Haeckel, 1881.

_Liosphærida_, Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Sphæroidea# without radial spines on the surface of the
spherical shell; living solitary (not associated in colonies).

The family #Liosphærida# comprises all those solitary #Sphæroidea# in which
the surface of the spherical shell is not armed with radial spines. Nearly
the half of this large group is formed by the Ethmosphærida, in which the
carapace is a quite simple, spherical lattice-shell; this subfamily is
probably the common ancestral group from which all other #Sphæroidea#, or
even all #Sphærellaria#, can be derived in a phylogenetical as well as in a
morphological sense. The central capsule in this first subfamily is
constantly enclosed by the fenestrated shell, and separated from it by the
jelly-veil. The shell is therefore an extracapsular or medullary shell.

To these simple Ethmosphærida all other subfamilies can be opposed as
"Liosphærida concentrica," as their carapace is composed of two or more
concentric lattice-shells; two in the Carposphærida, three in the
Thecosphærida, four in the Cromyosphærida, five or more in the
Caryosphærida. In all these four subfamilies the concentric shells are
simple (not spongy) fenestrated shells. In a sixth subfamily, in the
Plegmosphærida, the shell is wholly or partially composed of spongy
wicker-work, with or without a latticed medullary shell in the centre.

The internal radial beams, in the "Liosphærida composita" connecting the
concentric spheres, exhibit in their number and disposition similar
important differences, such as the external radial spines in the
Astrosphærida. The following eight {60}different cases of regular
disposition were observed:--(A) two opposite beams in one axis; (B) four
beams, opposite in pairs in two axes perpendicular one to another; (C) six
beams, opposite in pairs in the three dimensive axes; (D) eight beams,
opposite in pairs in the four diagonals of the regular cube; (E) twelve
beams corresponding to the twelve axes of the regular icosahedron; (F)
fourteen beams quite regularly disposed (six corresponding to the three
axes of the regular octahedron, eight to the central points of its faces);
(G) twenty beams (probably corresponding to the twenty corners of a regular
dodecahedron); (H) thirty-two beams, regularly disposed. Rarely the number
of the radial beams is intermediate between these eight cases, and rarely
it is higher; then commonly the disposition is irregular. The regularity of
their disposition in the great majority of cases is very remarkable and
evident.

_Synopsis of the Genera of Liosphærida._

-------------------------------------------------------------------------
I. Subfamily Ethmosphærida.
(Shell one single latticed sphere.)
-------------------------------------------------------------------------
{Shell cavity
{ simple, 15. _Cenosphæra_.
Pores of the {
shell simple, {Shell cavity
not prolonged { with radial
into free tubuli. { beams united
{ in the
{ centre, 16. _Stigmosphæra_.

Pores prolonged {Tubuli external,
into free { centrifugal, 17. _Ethmosphæra_.
conical or {
cylindrical {Tubuli internal,
tubuli. { centripetal, 18. _Sethosphæra_.
-------------------------------------------------------------------------
II. Subfamily Carposphærida.
(Two concentric spheres.)
-------------------------------------------------------------------------
One shell medullary (intracapsular),
the other cortical (extracapsular), 19. _Carposphæra_.

Both shells cortical
(near together), 20. _Liosphæra_.
-------------------------------------------------------------------------
III. Subfamily Thecosphærida.
(Three concentric spheres.)
-------------------------------------------------------------------------
Two shells medullary
(intracapsular), one shell
cortical (extracapsular), 21. _Thecosphæra_.

One shell medullary (intracapsular),
two shells cortical
(extracapsular), 22. _Rhodosphæra_.
-------------------------------------------------------------------------
IV. Subfamily Cromyosphærida.
(Four concentric spheres.)
-------------------------------------------------------------------------
Two inner medullary shells
(intracapsular), and two outer
cortical shells (extracapsular), 23. _Cromyosphæra_.
-------------------------------------------------------------------------
V. Subfamily Caryosphærida.
(Five or more concentric spheres.)
-------------------------------------------------------------------------
Two inner medullary shells, and
three or more outer cortical
shells, 24. _Caryosphæra_.
-------------------------------------------------------------------------
VI. Subfamily Plegmosphærida.
(Spherical shell wholly or partially of spongy structure.)
-------------------------------------------------------------------------
Spongy sphere {Sphere solid, 25. _Styptosphæra_.
without latticed {
medullary shell {Sphere with a
in the centre. { central cavity, 26. _Plegmosphæra_.

Spongy sphere with {One single
one or two { medullary shell, 27. _Spongoplegma_.
latticed {
medullary shells {Two concentric
in the centre. { medullary
{ shells, 28. _Spongodictyon_.

{61}Subfamily ETHMOSPHÆRIDA,[25] Haeckel, 1862, Monogr. d. Radiol., p. 348
(_sensu restricto_).

_Definition._--#Liosphærida# with one single spherical lattice-shell;
living solitary, not aggregated in colonies.

Genus 15. _Cenosphæra_,[26] Ehrenberg, 1854, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 237.

_Definition._--#Liosphærida# with one single latticed sphere, with simple
shell-pores (not prolonged into free tubuli) and with simple shell-cavity
(without internal radial beams).

The genus _Cenosphæra_ is the most simple form of all SPHÆROIDEA, and may
be regarded as the common ancestral form of this order. The siliceous
latticed shell, in which the central capsule is enclosed, represents a
simple regular sphere, with simple cavity. The pores of the shell-wall are
simple, not prolonged into radial tubuli (as in _Ethmosphæra_ and
_Sethosphæra_). According to the different form of the pores, the numerous
species of this genus can be disposed in four different subgenera. Some
species may be easily confounded with isolated shells of the corresponding
social _Collosphæra_; but in this latter the spherical shell-form is
commonly more or less irregular, in _Cenosphæra_ quite regular.

Subgenus 1. _Phormosphæra_, Haeckel, 1881, Prodromus, p. 448.

_Definition._--Pores of the shell regular or subregular, hexagonal or
circular, with hexagonal frames or lobes; all nearly of equal size and
form.

1. _Cenosphæra primordialis_, n. sp.

Shell very thin walled, smooth. Pores hexagonal, regular, or subregular;
twelve to fifteen on the half meridian of the shell; bars between them
extremely delicate (only visible when three hundred or four hundred times
enlarged). Diameter of the shell nine to ten times that of the meshes. This
species is remarkable for the extreme delicacy of the arachnoidal network
of the simple spherical shell; it may be regarded as the common ancestral
form of all #Sphæroidea#. The shell equals that of _Heliosphæra tenuissima_
(figured in my Monograph, 1862, pl. ix. fig. 2), but differs from it by the
smooth surface and the absence of all spines or thorns. I observed this
species living in the Indian Ocean, near Ceylon, in 1882; the spherical
diameter of the central capsule is about one-third of that of the shell;
the contents of the central capsule are colourless {62}and transparent,
except the central dark globular nucleus. The same shells also occur in
some mounted preparations of surface organisms from the Challenger.

_Dimensions._--Diameter of the shell 0.12, of the pores 0.012.

_Habitat._--Indian Ocean, Ceylon, Haeckel; Central Pacific, Stations 266,
271, surface.

2. _Cenosphæra inermis_, Haeckel.

_Heliosphæra inermis_, Haeckel, 1862, Monogr. d. Radiol., p. 351, Taf.
ix. fig. 1.

Surface of the thin-walled shell smooth. Pores regular, hexagonal, twelve
to fifteen times as broad as the bars, seven to nine on the quadrant.

_Dimensions._--Diameter of the shell 0.08 to 0.12, pores 0.012 to 0.015,
bars 0.001.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

3. _Cenosphæra hexagonalis_, n. sp.

Surface of the thick-walled shell smooth. Pores regular, hexagonal, five to
six times as broad as the bars, six to eight on the quadrant.

_Dimensions._--Diameter of the shell 0.1 to 0.15, pores 0.01 to 0.012, bars
0.002.

_Habitat._--Central Pacific, Stations 265 to 274, at various depths.

4. _Cenosphæra mellifica_, n. sp. (Pl. 12, fig. 9).

Surface of the thick-walled shell smooth. Pores regular, circular, with
thin hexagonal frames, four times as broad as the bars, six to eight on the
quadrant.

_Dimensions._--Diameter of the shell 0.2, pores 0.12, bars 0.003.

_Habitat._--South Pacific, Station 288, surface.

5. _Cenosphæra favosa_, n. sp. (Pl. 12, fig. 10).

Surface of the thick-walled shell rough. Pores regular, circular, with thin
hexagonal frames, three times as broad as the bars, ten to twelve on the
quadrant.

_Dimensions._--Diameter of the shell 0.2, pores 0.09, bars 0.003.

_Habitat._--North Atlantic, Færöe Channel (Gulf Stream), John Murray,
surface.

6. _Cenosphæra vesparia_, n. sp. (Pl. 12, fig. 11).

Surface of the thick-walled shell smooth. Pores regular, circular, with
thick hexagonal frames, twice as broad as the bars, ten to twelve on the
quadrant.

_Dimensions._--Diameter of the shell 0.25, pores 0.016, bars 0.008.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms, and surface.

{63}7. _Cenosphæra bombus_, n. sp.

Shell thick walled, rough. Pores regular, circular, with thin hexagonal
frames, of the same breadth as the bars, twenty to twenty-two on the
quadrant.

_Dimensions._--Diameter of the shell 0.3, pores and bars 0.005.

_Habitat._--Tropical Atlantic, Station 347, depth 2250 fathoms.

8. _Cenosphæra melecta_, n. sp.

Shell thick walled, papillate. Pores regular, circular, double-edged, with
thick hexagonal frames, of the same breadth as the bars; a short conical
papilla in the corner of each hexagon; fourteen to sixteen pores on the
quadrant.

_Dimensions._--Diameter of the shell 0.2, pores and bars 0.008.

_Habitat._--Fossil in Barbados.

9. _Cenosphæra anthophora_, n. sp.

Shell thick walled, papillate. Pores regular, circular, with an elegant
six-lobed frame and a coronal of six short papillæ; the latter alternating
with the six lobes (quite as in Pl. 28, fig. 1_b_). Pores twice as broad as
the bars, ten to twelve on the quadrant.

_Dimensions._--Diameter of the shell 0.13, pores 0.01, bars 0.005.

_Habitat._--Central Pacific, Station 271, surface.

10. _Cenosphæra rosetta_, n. sp.

Shell thin walled, smooth. Pores regular, circular, with an elegant
six-lobed outer opening, without papillæ. Pores of the same breadth as the
bars, six to eight on the quadrant.

_Dimensions._--Diameter of the shell 0.1, pores and bars 0.006.

_Habitat._--South Atlantic, Station 323, depth 1900 fathoms.

Subgenus 2. _Circosphæra_, Haeckel.

_Definition._--Pores of the spherical shell regular or subregular,
circular, without hexagonal frames or lobes, all nearly of equal size and
form.

11. _Cenosphæra porophæna_, Ehrenberg.

_Cenosphæra porophæna_, Ehrenberg, 1858, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 31.

Shell thin walled, smooth. Pores regular, circular, six to eight times as
broad as the bars, five to six on the quadrant.

_Dimensions._--Diameter of the shell 0.1, pores 0.012, bars 0.002.

_Habitat._--Mediterranean (Crete, depth 1100 fathoms; Corfu, surface).

{64}12. _Cenosphæra setosa_, Ehrenberg.

_Cenosphæra setosa_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 287, Taf. vii. fig. 1.

Shell thin walled, covered with very short and numerous bristles. Pores
regular, circular, five to six times as broad as the bars, six to eight on
the quadrant.

_Dimensions._--Diameter of the shell 0.17, pores 0.2, bars 0.03.

_Habitat._--West Tropical Pacific, 3300 fathoms; Philippine Sea, Station
200, depth 250 fathoms.

13. _Cenosphæra plutonis_, Ehrenberg.

_Cenosphæra plutonis_, Ehrenberg, 1854, Mikrogeol., Taf. xxxv. B, B. iv.
fig. 20.

Shell thin walled, covered with short conical papillæ. Pores regular,
circular, twice as broad as the bars, eight to nine on the quadrant.

_Dimensions._--Diameter of the shell 0.09, pores 0.006, bars 0.003.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

14. _Cenosphæra proserpinæ_, n. sp.

Shell thin walled, smooth. Pores regular, circular, four times as broad as
the bars, five to six on the quadrant.

_Dimensions._--Diameter of the shell 0.08, pores 0.008, bars 0.002.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

15. _Cenosphæra eridani_, n. sp.

Shell thin walled, smooth. Pores regular, circular, three times as broad as
the bars, eleven to twelve on the quadrant.

_Dimensions._--Diameter of the shell 0.16, pores 0.01, bars 0.003.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

16. _Cenosphæra lethe_, n. sp.

Shell thick walled, smooth. Pores regular, circular, double-edged, five
times as broad as the bars, sixteen to eighteen on the quadrant.

_Dimensions._--Diameter of the shell 0.25, pores 0.01, bars 0.002.

_Habitat._--North Atlantic, Station 64, depth 2700 fathoms.

17. _Cenosphæra elysia_, n. sp. (Pl. 12, fig. 8).

Shell thick walled, rough. Pores regular, circular, double-edged, twice as
broad as the bars, twelve to fourteen on the quadrant.

_Dimensions._--Diameter of the shell 0.2, pores 0.01, bars 0.005.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

{65}18. _Cenosphæra nirwana_, n. sp.

Shell thick walled, smooth. Pores regular, circular, twice as broad as the
bars, twenty-four to twenty-five on the quadrant.

_Dimensions._--Diameter of the shell 0.25, pores 0.004, bars 0.002.

_Habitat._--Indian Ocean, Belligemma, Ceylon, surface; Haeckel.

19. _Cenosphæra maxima_, n. sp.

Shell thick walled, smooth. Pores regular, circular, twice as broad as the
bars, thirty to thirty-three on the quadrant.

_Dimensions._--Diameter of the shell 0.3 to 0.4, pores 0.012, bars 0.006.

_Habitat._--West Tropical Pacific, Station 225, depth 4475; also fossil in
Barbados.

20. _Cenosphæra compacta_, n. sp. (Pl. 12, fig. 7).

Shell very thick walled, rough (its wall one-fourth to one-third as thick
as the radius). Pores subregular, circular, of the same breadth as the
bars, seven to eight on the quadrant.

_Dimensions._--Diameter of the shell 0.15, pores and bars 0.012.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

_Cenosphæra radiata_, Zittel, 1876 (L. N. 29, p. 84, Taf. ii. figs. 7, 8),
a fossil Cretaceous species, is closely allied to _Cenosphæra compacta_.

21. _Cenosphæra crassa_, n. sp.

Shell very thick walled, rough (its wall nearly half as thick as the
radius). Pores tubular, double-edged, regular, circular, ten times as broad
as the thin united bars, twelve to fourteen on the quadrant.

_Dimensions._--Diameter of the shell 0.14, pores 0.01, bars 0.001.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

22. _Cenosphæra solida_, n. sp.

Shell very thick walled, covered with innumerable short bristles (its wall
one-third as thick as the radius). Pores regular, circular, four times as
broad as the bars, tubular, eight to ten on the quadrant.

_Dimensions._--Diameter of the shell 0.2, pores 0.02, bars 0.005.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Subgenus 3. _Cyrtidosphæra_, Haeckel, 1862, Monogr. d. Radiol., p. 348.

_Definition._--Pores of the spherical shell irregularly polygonal, of
unequal size or form, sometimes roundish with polygonal frames.

{66}23. _Cenosphæra reticulata_, Haeckel.

_Cyrtidosphæra reticulata_, Haeckel, 1862, Monogr. d. Radiol., p. 349,
Taf. xi. fig. 2.

Shell very thin walled, smooth. Pores irregular, polygonal, two to eight
times as broad as the bars, fifteen to twenty on the quadrant (groups of
four to eight smaller meshes are scattered on the surface, and separated by
reticular rows of larger meshes).

_Dimensions._--Diameter of the shell 0.16, pores 0.004 to 0.016, bars
0.002.

_Habitat._--Mediterranean (Messina), surface.

24. _Cenosphæra tenerrima_, n. sp.

Shell extremely thin walled, smooth, like a cobweb. Pores very irregular
and small, polygonal with thread-like bars, thirty to forty on the
quadrant.

_Dimensions._--Diameter of the shell 0.25, pores 0.002 to 0.008, bars under
0.001.

_Habitat._--Central Pacific, Station 271, surface.

25. _Cenosphæra polygonalis_, n. sp.

Shell thin walled, smooth, with irregular, polygonal pores, three to four
times as broad as the bars, eight to ten on the quadrant.

_Dimensions._--Diameter of the shell 0.2, pores 0.012 to 0.02, bars 0.005.

_Habitat._--North Pacific, Station 236, surface.

26. _Cenosphæra papillata_, n. sp.

Shell thick walled, covered with short conical papillæ. Pores irregular,
polygonal, three to five times as broad as the bars, fourteen to sixteen on
the quadrant.

_Dimensions._--Diameter of the shell 0.12, pores 0.004 to 0.007, bars
0.0015.

_Habitat._--South Atlantic, Station 325, surface.

27. _Cenosphæra cristata_, n. sp.

Shell thick walled, rough. Pores irregular, roundish, surrounded by
polygonal crested frames two to three times as broad as the bars, eight to
twelve on the quadrant.

_Dimensions._--Diameter of the shell 0.16, pores 0.006 to 0.01, bars 0.003.

_Habitat._--North Pacific, Station 254, surface.

28. _Cenosphæra perforata_, n. sp. (Pl. 26, fig. 10).

_Ceriosphæra perforata_, Haeckel, 1881, Prodromus et Atlas, _loc. cit._

Shell thick walled, rough. Pores irregular, roundish, surrounded by high
polygonal funnel-shaped frames, which are solid in the inner half,
perforated by numerous very small pores in the {67}outer half,
sieve-shaped. Pores one to three times as broad as the bars, of very
different size, four to six on the quadrant.

_Dimensions._--Diameter of the shell 0.15, pores 0.01 to 0.02, bars 0.005
to 0.01.

_Habitat._--Tropical Atlantic, Station 347, depth 2250 fathoms.

29. _Cenosphæra coronata_, n. sp. (Pl. 26, fig. 11).

Shell thick walled, rough. Pores irregular, roundish, surrounded by high
polygonal frames bearing on their sharp crest a series of small papillæ, so
that each pore is surrounded by a coronal of such spinules. Pores four to
eight times as broad as the bars, four to five on the quadrant.

_Dimensions._--Diameter of the shell 0.15, pores 0.01 to 0.03, bars 0.002
to 0.008.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

Subgenus 4. _Porosphæra_, Haeckel.

_Definition._--Pores of the spherical shell irregular, roundish, without
polygonal frames, of unequal size or form.

30. _Cenosphæra antiqua_, Haeckel.

_Cenosphæra plutonis_, var., Dunikowski, 1882, Denkschr. d. k. Akad. d.
Wiss. Wien, Bd. xlv. p. 25, Taf. iv. figs. 47, 48.

_Cenosphæra plutonis_, var., Stöhr, 1880, Palæontogr. xxvi. p. 85, Taf.
i. fig. 1.

Shell thin walled, smooth. Pores irregular, roundish, two to eight times as
broad as the bars, ten to fifteen on the quadrant.

_Dimensions._--Diameter of the shell 0.15 to 0.2, pores 0.007 to 0.015,
bars 0.002 to 0.008.

_Habitat._--Fossil in the Jurassic, Cretaceous, and Tertiary formations;
living in the depths of the Atlantic and Pacific; Station 332, depth 2200
fathoms; Station 225, depth 4475 fathoms, &c.

31. _Cenosphæra gigantea_, n. sp.

Shell thick walled, smooth. Pores irregular, roundish, two to five times as
broad as the bars, thirty to forty on the quadrant.

_Dimensions._--Diameter of the shell 0.4 to 0.5, pores 0.004 to 0.01, bars
0.002.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms; also fossil
in Barbados.

32. _Cenosphæra marginata_, n. sp.

Shell very thick walled, smooth. Pores irregular, roundish, double-edged,
three to eight times as broad as the bars, six to eight on the quadrant.

{68}_Dimensions._--Diameter of the shell 0.12, pores 0.01 to 0.03, bars
0.004.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

33. _Cenosphæra aspera_, Stöhr.

_Cenosphæra aspera_, Stöhr, 1880, Palæontogr. Bd. xxvi. p. 85, Taf. i.
fig. 2.

Shell thick walled, rough, covered with short conical papillæ. Pores
irregular, roundish, of about the same breadth as the bars, eight to twelve
on the quadrant.

_Dimensions._--Diameter of the shell 0.17, pores and bars 0.01 to 0.04.

_Habitat._--Fossil in Tertiary rocks of Sicily; Grotte, Stöhr.

34. _Cenosphæra hirsuta_, Ehrenberg.

_Cenosphæra hirsuta_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.,
Berlin, p. 287, Taf. x. fig. 18.

Shell thin walled, rough, covered with innumerable very short bristles.
Pores very irregular, roundish, of about the same breadth as the bars.

_Dimensions._--Diameter of the shell 0.11, pores and bars 0.002 to 0.012.

_Habitat._--Indian Ocean, Zanzibar, Pullen; depth 2200 fathoms.

Genus 16. _Stigmosphæra_,[27] Haeckel, n. gen.

_Definition._--#Liosphærida# with one single latticed sphere, with simple
shell-pores (not prolonged into free tubuli); in the central point of the
spherical shell-cavity are united a number of radial beams, which become
forked and inserted into the inner surface of the shell by their distal
ends.

The genus _Stigmosphæra_ differs from _Cenosphæra_ (and from all other
Monosphærida) by internal radial beams, which are united in the centre of
the simple spherical shell; these beams are branched, and the distal ends
of the branches inserted on the internal surface of the shell. I have
observed only two, nearly identical specimens of this genus, both with
regular, hexagonal pores and thin bars; the beams were implanted in the
corners of the hexagons. In one specimen the surface was covered with short
radial bristles, whilst these in the other specimen were prolonged into
radial spines (like _Acanthosphæra_). Possibly this peculiar genus is
derived from _Carposphæra_, by reduction and loss of a central medullary
shell.

_Stigmosphæra actinocentra_, n. sp.

Shell very thin walled, rough, with regular circular, hexagonally framed
pores, six times as broad as the bars, eight to ten on the quadrant; in the
corner of each hexagon a small bristle. In {69}the central point of the
shell are united about twelve (?) thin and straight radial beams, which are
forked, with dichotomous branches; the distal ends of the branches are
inserted in the corners of the hexagons on the inside of the shell.

_Dimensions._--Diameter of the shell 0.15, pores 0.02, bars 0.003.

_Habitat._--Central Pacific, Station 271, surface.

Genus 17. _Ethmosphæra_,[28] Haeckel, 1862, Monogr. d. Radiol., p. 349.

_Definition._--#Liosphærida# with one single latticed sphere, with simple
shell-cavity; with shell-pores which are prolonged on the outside in
centrifugal, conical, or cylindrical tubuli.

The genus _Ethmosphæra_ differs from the simple _Cenosphæra_, its ancestral
form, by the peculiar formation of the shell-pores; in all observed species
of the genus these are quite regular, of nearly equal size and form; their
base in the spherical shell-face is hexagonal, but on the outside prolonged
into centrifugal, external, radial tubuli, which are either conical or
cylindrical (in the latter case both openings of the tubes being equal, in
the former the outer opening being smaller than the inner). The solitary
_Ethmosphæra_ corresponds to the social _Siphonosphæra_; but in the former
the formation of the shell and of its tubuli is quite regular, in the
latter more or less irregular.

Subgenus 1. _Ethmosphærella_, Haeckel.

_Definition._--Tubuli conical, their outer opening smaller than the inner.

1. _Ethmosphæra siphonophora_, Haeckel.

_Ethmosphæra siphonophora_, Haeckel, 1862, Monogr. d. Radiol., p. 350,
Taf. xi. fig. 1.

Tubuli conical, their outer opening half as broad as the inner and three
times as broad as their height. Five to six pores on the quadrant. Diameter
of the outer pores one and a half times as large as their distance from
each other.

_Dimensions._--Diameter of the shell 0.1, outer pores 0.01, their distance
0.007.

_Habitat._--Mediterranean (Messina), surface, Haeckel.

2. _Ethmosphæra conosiphonia_, n. sp. (Pl. 12, figs. 5, 5_a_).

Tubuli conical, their outer opening two thirds as broad as the inner, and
scarcely broader than their height. Ten to twelve pores on the quadrant.
Diameter of the outer pores twice as large as their distance from each
other.

_Dimensions._--Diameter of the shell 0.17, outer pores 0.01, their distance
0.005.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

{70}3. _Ethmosphæra polysiphonia_, n. sp. (Pl. 12, fig. 6).

Tubuli conical, their outer opening three-fourths as broad as the inner and
three times as broad as their height. Sixteen to eighteen pores on the
quadrant. Diameter of the outer pores three times as large as their
distance apart.

_Dimensions._--Diameter of the shell 0.2, outer pores 0.008, their distance
0.003.

_Habitat._--West Tropical Pacific, Station 225, depth 4475 fathoms; also
fossil in Barbados and in Sicily.

Subgenus 2. _Ethmosphæromma_, Haeckel.

_Definition._--Tubuli cylindrical, their outer opening about as large as
the inner.

4. _Ethmosphæra stenosiphonia_, n. sp.

Tubuli cylindrical, short, quite contiguous, so that their diameter is six
times as large as their distance apart, but about equal to their height.
Nine to ten pores on the quadrant.

_Dimensions._--Diameter of the shell 0.14, outer pores 0.012, their
distance 0.002.

_Habitat._--Central Pacific, Station 271, surface.

5. _Ethmosphæra pachysiphonia_, n. sp.

Tubuli cylindrical, twice as long as broad, very thick-walled, and nearly
contiguous, so that their diameter is five times as large as their distance
apart. The thickness of their wall is equal to their lumen. Twelve to
fourteen pores on the quadrant.

_Dimensions._--Diameter of the shell 0.16, outer pores 0.01, their distance
0.002.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

6. _Ethmosphæra leptosiphonia_, n. sp.

Tubuli cylindrical, short, about as long as broad, very thin-walled and
fragile, separated by wide distances, which are three times as large as
their diameter. Six to seven pores on the quadrant. (Very similar to
_Siphonosphæra cyathina_, Pl. 6, fig. 10, but quite regular, all tubuli
retaining the same size and distance.)

_Dimensions._--Diameter of the shell 0.12, outer pores 0.01, their distance
0.03.

_Habitat._--North Atlantic, Færöe Channel, John Murray; surface.

7. _Ethmosphæra macrosiphonia_, n. sp.

Tubuli cylindrical, very elongated, four times as long as broad; their
bases separated by distances which are equal to their breadth. Eight to ten
tubuli on the quadrant. (The tubes are similar to those of _Siphonosphæra
serpula_, Pl. 6, fig. 6, but quite regular, straight, not curved, all of
the same size and at equal distances apart.)

{71}_Dimensions._--Diameter of the shell 0.1, length of the tubes 0.04,
breadth 0.01, basal distance 0.01.

_Habitat._--Indian Ocean, Cocos Islands, surface, Rabbe.

Genus 18. _Sethosphæra_,[29] Hæckel, 1881, Prodromus, p. 452 (_sensu
emendato_).

_Definition._--#Liosphærida# with one single latticed sphere, with simple
shell-cavity; with shell-pores, which are prolonged on the inside into
centripetal, conical, or cylindrical tubuli.

The genus _Sethosphæra_ differs from its ancestral form, _Cenosphæra_, by
the production of internal, centripetal, radial tubuli on the inside of the
shell (the contrary of the preceding genus _Ethmosphæra_). It corresponds
therefore to the social _Pharyngosphæra_; but in the latter the formation
of the shell and its tubes is more or less irregular, whilst in the former
each regular pore is prolonged into a regular tubule.

1. _Sethosphæra entosiphonia_, n. sp.

Shell with smooth surface and regular circular pores, separated by
hexagonal frames, twice as broad as the bars. Six to eight on the quadrant.
Each pore is prolonged on the inside of the shell in a short conical
centripetal tube, twice as long as its diameter.

_Dimensions._--Diameter of the shell 0.15, outer pores 0.008, bars 0.004,
length of the tubuli 0.015.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Sethosphæra entosolenia_, n. sp.

Shell with smooth surface and regular circular pores, without hexagonal
frames, of about the same breadth as the bars. Ten to twelve on the
quadrant. Each pore is prolonged on the inside of the shell into a thin
cylindrical centripetal tube, three times as long as its diameter.

_Dimensions._--Diameter of the shell 0.2, outer pores and bars 0.006,
length of the tubuli 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subfamily CARPOSPHÆRIDA,[30] Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Liosphærida# with two concentric spherical lattice-shells,
which are united by radial beams.

_Carposphæra_,[31] Haeckel, 1881, Prodromus, p. 451.

_Definition._--Liosphærida with one medullary (intracapsular) and one
cortical (extracapsular) shell, both connected by radial beams piercing the
central capsule.

{72}The genus _Carposphæra_ comprises a large number of double-shelled
#Sphæroidea#, formerly united with _Haliomma_, but different from this
genus by the absence of radial spines. The shell is composed of two
concentric latticed spheres, the inner of which (or the medullary shell) is
situated within the central capsule, the other (or the cortical shell)
outside it. Both shells are connected by radial beams which pierce the wall
of the central capsule. The distance between the shells is at least as
large as (commonly much larger than) the radius of the inner shell, whilst
in the following genus, _Liosphæra_, that distance is much smaller than
this radius.

Subgenus 1. _Melittosphæra_, Haeckel, 1881, Prodromus, p. 451.

_Definition._--Pores of the cortical shell regular, hexagonal (or circular,
with hexagonal frames or lobes), all of nearly equal size and form.

1. _Carposphæra capillacea_, n. sp.

Cortical shell very delicate, four times as broad as the similarly
constructed medullary shell, with regular hexagonal meshes (twenty to
twenty-five on the quadrant) and very thin thread-like bars. Both shells
connected by twenty (?) very thin radial beams. (Similar to _Heliosoma
radians_, Pl. 28, fig. 3, 3_a_, but with smooth surface, without any radial
spines.)

_Dimensions._--Diameter of the outer shell 0.2, inner 0.05, pores 0.01,
bars below 0.001.

_Habitat._--Central Pacific, Station 274, surface.

2. _Carposphæra cubaxonia_, n. sp.

Cortical shell smooth, three times as broad as the medullary shell, with
regular hexagonal pores, four times as broad as the bars. Eight to ten
pores on the quadrant. Medullary shell with regular circular pores, twice
as broad as the bars. Both shells connected by six radial beams, which are
three-sided prismatical, opposite in pairs in the three dimensive axes.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; outer pores
0.01, inner 0.005.

_Habitat._--South Pacific, Station 291, surface.

3. _Carposphæra infundibulum_, Haeckel.

_Haliomma infundibuliforme_, Stöhr, 1880, Palæontogr. Bd. xxvi. p. 87,
Taf. i. fig. 6.

Cortical shell very thick-walled, two and a half times as broad as the
medullary shell, with rough surface and regular hexagonal, funnel-shaped
pores, of about the same breadth as the bars. Five to six on the quadrant.

_Dimensions._--Diameter of the outer shell 0.1, inner 0.04, outer pores and
bars 0.01.

_Habitat._--North Atlantic, Station 354; fossil in Tertiary rocks (Barbados
and Sicily).

{73}4. _Carposphæra melissa_, n. sp.

Cortical shell thick walled, four times as broad as the medullary shell,
with regular circular, hexagonally framed pores, three times as broad as
the bars. Eight to ten pores on the quadrant. Medullary shell with simple
circular pores.

_Dimensions._--Diameter of the outer shell 0.16, inner 0.04, outer pores
0.012, bars 0.004.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

5. _Carposphæra melitomma_, n. sp. (Pl. 20, fig. 4).

_Melitomma melittosphæra_, Haeckel, 1881; Prodromus et Atlas, _loc. cit._

Cortical shell thick walled, with thorny surface, two and a half times as
broad as the medullary shell. Its pores regular, circular, twice as broad
as the bars, elegantly six-lobed, separated by crested hexagonal frames; in
each hexagon-corner a short conical papilla (alternating with a lobe).
Eight to ten pores on the quadrant. Medullary shell with small simple
circular pores.

_Dimensions._--Diameter of the outer shell 0.17, inner 0.07, outer pores
0.01, bars 0.005, inner pores 0.005.

_Habitat._--Central Pacific, Stations 266 to 274, in various depths.

Subgenus 2. _Cerasosphæra_, Haeckel, 1881, Prodromus, p. 451.

_Definition._--Pores of the cortical shell regular, circular, without
hexagonal frames, all of nearly equal size and form.

6. _Carposphæra cerasus_, n. sp.

Cortical shell thin walled, smooth, twice as broad as the medullary shell;
both with regular circular pores, six times as broad as the bars. Twelve to
fifteen pores on the quadrant. Outer pores twice as large as the inner.

_Dimensions._--Diameter of the outer shell 0.24, inner 0.12; outer pores
0.016, inner 0.008.

_Habitat._--Central Pacific, Station 271, surface.

7. _Carposphæra apiculata_, Haeckel.

? _Haliomma apiculatum_, Ehrenberg, 1872; Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 313.

Cortical shell thin-walled, covered with numerous short conical thorns,
three times as broad as the medullary shell. Pores regular, circular, four
times as broad as the bars. Six to eight pores on the quadrant.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05, outer pores
0.012, bars 0.03.

_Habitat._--North Pacific, California, depth 2000 fathoms; Station 254,
depth 3025 fathoms.

{74}8. _Carposphæra entactinia_, Haeckel.

_Haliomma entactinia_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxvi. fig. 4.

Cortical shell thick walled, rough, twice as broad as the medullary shell;
both shells with regular circular pores, twice as broad as the bars. Six to
eight pores on the quadrant. Outer pores half as broad as the inner. Both
shells connected by very numerous (twenty-four to forty-eight or more)
radial beams.

_Dimensions._--Diameter of the outer shell 0.12, inner 0.06, outer pores
0.008, bars 0.004, inner pores 0.004.

_Habitat._--Cosmopolitan; Atlantic and Pacific, in various depths; fossil
in Barbados and Sicily.

9. _Carposphæra modesta_, Haeckel.

_Haliomma modestum_, Stöhr, 1880, Palæontogr. Bd. xxvi. p. 86, Taf. i.
fig. 5.

Cortical shell thick walled, rough, three times as broad as the medullary
shell, with regular circular pores of the same breadth as the bars. Eight
to twelve pores on the quadrant. (Very common, like the preceding species,
and connected with it by numerous intermediate forms.)

_Dimensions._--Diameter of the outer shell 0.12 to 0.2, inner 0.04 to 0.07,
pores and bars 0.006 to 0.008.

_Habitat._--Atlantic and Pacific, from many Stations and at various depths;
fossil in Barbados and Sicily.

10. _Carposphæra belladonna_, n. sp.

Cortical shell thick walled, smooth, five times as broad as the medullary
shell, with regular circular pores of the same breadth as the bars. Twenty
to twenty-two pores on the quadrant.

_Dimensions._--Diameter of the outer shell 0.3, inner 0.06, outer pores and
bars 0.004.

_Habitat._--North Atlantic, Færöe Channel, John Murray.

11. _Carposphæra areca_, n. sp.

Cortical shell very thick walled, rough, twice as broad as the medullary
shell, with regular circular pores half as broad as the bars. Eight to ten
pores on the quadrant.

_Dimensions._--Diameter of the outer shell 0.12, inner 0.06, outer pores
0.03, bars 0.006.

_Habitat._--Indian Ocean, Ceylon, Haeckel, surface.

Subgenus 3. _Prunosphæra_, Haeckel, 1881, Prodromus, p. 451.

_Definition._--Pores of the cortical shell irregular polygonal, of unequal
size or dissimilar form.

12. _Carposphæra prunulum_, n. sp.

Cortical shell thin walled, smooth, four times as broad as the medullary
shell, with large irregular polygonal pores, four to eight times as broad
as the bars. Connecting beams between them numerous.

{75}_Dimensions._--Diameter of the outer shell 0.24, inner 0.06, outer
pores 0.008 to 0.016, bars 0.002.

_Habitat._--South Atlantic, Station 325, surface.

13. _Carposphæra corypha_, n. sp.

Cortical shell thin walled, rough, three times as broad as the medullary
shell, with irregular polygonal pores, three to six times as broad as the
bars. Connecting beams between the two shells twenty, regularly disposed.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05, outer pores
0.01 to 0.02, bars 0.003.

_Habitat._--South Pacific, Station 300, surface.

14. _Carposphæra borassus_, n. sp.

Cortical shell thick walled, smooth, three times as broad as the medullary
shell, with irregular polygonal pores, two to four times as broad as the
bars. Connecting beams between the two shells six, opposite by pairs in the
three dimensive axes. (Similar to _Hexalonche aristarchi_, Pl. 22, fig. 3,
but without external radial spines.)

_Dimensions._--Diameter of the outer shell 0.12, inner 0.04, outer pores
0.01 to 0.02, bars 0.005.

_Habitat._--Central Pacific, Station 268, surface.

Subgenus 4. _Phoenicosphæra_, Haeckel.

_Definition._--Pores of the cortical shell irregular roundish, of unequal
size or form.

15. _Carposphæra nobilis_, Haeckel.

_Haliomma nobile_, Ehrenberg, 1844, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 268; Abhandl., 1875, Taf. xxvii. fig. 6.

Cortical shell thin walled, rough, twice as broad as the medullary shell,
with irregular roundish pores, two to four times as broad as the bars.

_Dimensions._--Diameter of the outer shell 0.1, inner 0.05, outer pores
0.01 to 0.02, bars 0.006.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, at various depths;
fossil in Jurassic, Cretaceous, and Tertiary rocks.

16. _Carposphæra micrococcus_, n. sp.

Cortical shell thin walled, rough, seven times as broad as the medullary
shell, with irregular roundish pores, three to six times as broad as the
bars.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.03, outer pores
0.012 to 0.025, bars 0.004.

_Habitat._--South Atlantic, Station 330, surface.

{76}17. _Carposphæra maxima_, n. sp.

Cortical shell thin walled, smooth, five times as broad as the medullary
shell, with irregular roundish pores, of about the same breadth as the
bars.

_Dimensions._--Diameter of the outer shell 0.4, inner 0.08, pores and bars
0.004 to 0.008.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

18. _Carposphæra nodosa_, n. sp. (Pl. 28, figs. 2, 2_a_).

_Anthomma nodosum_, Haeckel, 1879, Atlas, _loc. cit._

Cortical shell thick walled, covered with forty to fifty scattered
pyramidal nodules, two and a half times as broad as the medullary shell,
connected with it by very numerous thin radial beams. Outer and inner pores
irregular roundish or polygonal, two to three times as broad as the bars.
(This species in consequence of the cortical nodules may represent a
peculiar genus, analogous to _Conosphæra_, called _Anthomma_.)

_Dimensions._--Diameter of the outer shell 0.13, inner 0.05, inner and
outer pores 0.008 to 0.012, bars 0.004.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Genus 20. _Liosphæra_,[32] Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Liosphærida# with two cortical (extracapsular) shells
(without a medullary or intracapsular shell).

The genus _Liosphæra_ agrees with the preceding _Carposphæra_ in the
possession of two concentric latticed spheres; but whilst in the latter
genus the inner sphere is a medullary one (intracapsular), the outer a
cortical shell (extracapsular), both connected by radial beams piercing the
capsule-wall, here in _Liosphæra_ the central capsule lies freely within
the inner lattice shell and is not pierced by radial beams. Therefore both
shells are here cortical shells, both separated by a distance, which is
constantly much smaller than the radius of the inner shell; whereas in
_Carposphæra_ this distance is at least as large as that radius (commonly
much larger). In _Carposphæra_ the number of pores in both shells is never
the same; in several species of _Liosphæra_ this number is the same, each
outer regular hexagonal pore exactly corresponding to an inner; the six
corners of each connected by six short radial beams.

Subgenus 1. _Melitomma_, Haeckel.

_Definition._--Pores of both shells regular, in each shell all of nearly
equal size and form.

1. _Liosphæra hexagonia_, n. sp. (Pl. 20, fig. 3).

Both shells with the same number of pores, exactly corresponding, about ten
on the quadrant. {77}All pores regular, or subregular, hexagonal; the outer
twice as broad as the inner. Outer bars very thin, thread-like; inner bars
thick, one-third as broad as the pores. Surface smooth. Both shells
connected by numerous radial beams, their distance one-third as large as
the radius of the inner shell.

_Dimensions._--Diameter of the outer shell 0.16, inner 0.12, distance of
both 0.02; outer pores 0.014, inner pores 0.007.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Liosphæra rhodococcus_, n. sp.

Both shells with the same number of pores, exactly corresponding, about
twelve on the quadrant. All pores regular or subregular; the inner
circular, with elegant six-lobed frames, twice as broad as the bars; the
outer hexagonal, with very thin thread-like bars. All corners of the outer
and inner hexagons connected by thin, bristle-shaped radial beams. (Similar
to _Haliomma rhodococcus_, Pl. 19, fig. 6; but with smooth surface and
regular hexagonal pores of the outer shell.)

_Dimensions._--Diameter of the outer shell 0.2, inner 0.16, distance of
both 0.02; outer pores 0.03, inner 0.01.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

3. _Liosphæra porulosa_, n. sp.

Both shells with regular hexagonal pores; their number in the outer shell
seven times as great as in the inner. Pores of the stout inner shell large,
three times as broad as the bars, about eight on the quadrant. From each
hexagon-corner arises one bristle-shaped radial beam; their distal ends are
united by threads (three from each), forming the large meshes of the
delicate outer shell. Each of these is divided by very thin threads into
seven small circular porules, one central and six around it.

_Dimensions._--Diameter of the outer shell 0.25, inner 0.2, distance of
both 0.025; outer pores 0.04, their porules 0.012, inner pores 0.15.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe; surface.

Subgenus 2. _Craspedomma_, Haeckel.

_Definition._--Pores of both shells irregular, in each shell differing
either in form or size.

4. _Liosphæra peridromium_, n. sp.

Both shells with the same number of large, polygonal, very irregular pores,
exactly corresponding (about eight to ten on the quadrant); both with a
very delicate thin framework. From the thread-like bars of the inner, very
large and thin-walled, sphere arise perpendicularly innumerable short
bristles of equal length, which are united at equal distances by tangential
thread-like bars, parallel to the former, composing the outer shell. Each
mesh is, therefore, surrounded by a delicate ballister or rail.

{78}_Dimensions._--Diameter of the outer shell 0.42, inner 0.4, distance of
both 0.01; diameter of the meshes 0.02 to 0.06.

_Habitat._--North Pacific, Station 250, surface.

5. _Liosphæra polypora_, n. sp. (Pl. 20, fig. 2).

Both shells with small irregular roundish pores, of about the same size as
the bars between them; twenty to thirty on the quadrant. The pores of the
outer shell somewhat smaller, therefore much more numerous than the pores
of the inner shell. Distance between the two shells about one-third as
great as the radius of the inner. Both shells connected by numerous thin
radial beams. Surface smooth or a little rough.

_Dimensions._--Diameter of the outer shell 0.18, inner 0.14, distance of
both 0.02; pores and bars 0.003 to 0.005.

_Habitat._--West Tropical Pacific, Station 225, depth 4475.

Subfamily THECOSPHÆRIDA,[33] Haeckel, 1881, Prodromus, pp. 449, 452.

_Definition._--#Liosphærida# with three concentric spherical
lattice-shells, which are united by radial beams.

Genus 21. _Thecosphæra_,[34] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Liosphærida# with two medullary (intracapsular) shells and
one cortical (extracapsular) shell.

The genus _Thecosphæra_ comprises a large number of triple-shelled
#Sphæroidea#, formerly united with _Actinomma_, but different from this
genus in the absence of radial spines. The latticed shell is composed of
three concentric spheres, two of which lie within the central capsule
(medullary shells), and one outside (cortical shell). This latter is
connected with the former by radial beams piercing the wall of the central
capsule. From the following _Rhodosphæra_ (with one medullary and two
cortical shells) _Carposphæra_ differs also by the distance of the three
shells. In the former the distance between the two outer shells is much
smaller, in the latter much larger, than the distance between the inner
shells.

Subgenus 1. _Thecosphærantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, hexagonal, or circular,
with hexagonal frames or lobes, all of nearly equal size and form.

{79}1. _Thecosphæra triplodictyon_, n. sp.

Cortical shell thin walled, smooth, with regular, hexagonal pores, four
times as broad as the bars. Radial proportion of the three spheres =
1 : 2 : 8. Both medullary shells with regular circular pores, twice as
broad as the bars, the inner half as broad as the outer. All three shells
connected by six thin radial beams, opposite in pairs in the three
dimensive axes.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.05, inner 0.025;
cortical pores 0.012, bars 0.003.

_Habitat._--Central Pacific, Station 271, surface.

2. _Thecosphæra phænaxonia_, n. sp.

Cortical shell thick walled, rough, with regular, circular, hexagonally
framed pores, twice as broad as the bars. Radial proportion of the three
spheres = 1 : 2 : 6. Both medullary shells with regular hexagonal pores and
thin bars. All three shells connected by six prismatic radial beams,
opposite in pairs in the three dimensive axes. (Shell similar to
_Hexacontium sceptrum_, Pl. 24, fig. 1, 1_a_, but without external spines.)

_Dimensions._--Diameter of the outer shell 0.12, middle 0.04, inner 0.02;
cortical pores 0.01, bars 0.005.

_Habitat._--North Pacific, Station 253, surface.

3. _Thecosphæra favosa_, n. sp.

Cortical shell thick walled, thorny, with regular, circular, hexagonally
framed pores, of the same breadth as the bars. Radial proportion of the
three spheres = 1 : 3 : 10. Both medullary shells with regular circular
pores, connected with the former by twelve short prismatic, regularly
disposed radial beams.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.06, inner 0.02;
cortical pores and bars 0.008.

_Habitat._--Central Pacific, Station 268, surface; also fossil in Barbados.

4. _Thecosphæra floribunda_, n. sp.

Cortical shell thick walled, smooth, with regular, elegantly six-lobed
pores, three times as broad as the bars. Radial proportion of the three
spheres = 1 : 2 : 4. Both medullary shells with simple regular circular
pores, connected with the former by six dimensive radial beams. (Similar to
_Hexacontium floridum_, Pl. 24, fig. 4, but without external spines.)

_Dimensions._--Diameter of the outer shell 0.12, middle 0.06, inner 0.03;
cortical pores 0.01, bars 0.0033.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

{80}Subgenus 2. _Thecosphærella_, Haeckel.

_Definition._--Pores of the cortical shell regular, circular, without
hexagonal frames or lobes, all of nearly equal size and form.

5. _Thecosphæra inermis_, Haeckel.

_Actinomma inerme_, Haeckel, 1862, Monogr. d. Radiol., p. 440, Taf. xxiv.
fig. 5.

_Haliomma inerme_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, p. 815.

Cortical shell thin walled, rough, with regular circular pores, twice as
broad as the bars. Radial proportion of the three spheres and of their
circular regular pores = 1 : 2 : 4. All three spheres connected by twelve
regularly disposed radial beams.

_Dimensions._--Diameter of the outer shell 0.1, middle 0.05, inner 0.025;
cortical pores 0.006, bars 0.003.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific, at many
Stations and at various depths.

6. _Thecosphæra æquorea_, Haeckel.

_Haliomma æquorea_, Ehrenberg, 1844, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 83; Mikrogeol., 1854, Taf. xxii. fig. 35.

_Actinomma æquorea_, Haeckel, 1862, Monogr. d. Radiol., p. 443.

Cortical shell thick walled, smooth, with regular circular pores of the
same breadth as the bars. Radial proportion of the three spheres and of
their regular pores = 1 : 2 : 6 or = 1 : 3 : 9; they are connected by six
radial beams, opposite by pairs in the three dimensive axes.

_Dimensions._--Diameter of the outer shell 0.08 to 0.12, middle 0.03 to
0.04, inner 0.09 to 0.12; cortical pores and bars about 0.006.

_Habitat._--Mediterranean, Corfu, surface; fossil in Greece and Sicily.

7. _Thecosphæra medusa_, Haeckel.

_Haliomma medusa_, Ehrenberg, 1838, Abhandl. d. k. Akad. d. Wiss. Berlin,
p. 130; Mikrogeol., 1854, Taf. xxii. figs. 33, 34.

_Actinomma medusa_, Haeckel, 1862, Monogr. d. Radiol., p. 444.

_Actinomma medusa_, Stöhr, 1880, Palæontogr., Bd. xxvi. p. 90, Taf. ii.
fig. 3.

Cortical shell thick walled, rough or thorny, with regular circular pores
of the same breadth as the bars. Radial proportion of the three spheres =
1 : 2 : 4 (or 1 : 2.5 : 6); they are connected by four radial beams,
crossed by pairs in two diameters, perpendicular one to another.

_Dimensions._--Diameter of the outer shell 0.08 to 0.12, middle 0.04 to
0.06, inner 0.02 to 0.025; cortical pores and bars in average 0.005.

_Habitat._--Fossil in Tertiary rocks of Barbados and the Mediterranean.

{81}8. _Thecosphæra entactinia_, n. sp.

Cortical shell thick walled, smooth, with regular circular pores of the
same breadth as the bars. Radial proportion of the three spheres =
1 : 3 : 12; they are connected by very numerous (forty to fifty, or more)
thin radial beams.

_Dimensions._--Diameter of the outer shell 0.24, middle 0.06, inner 0.02;
cortical pores and bars 0.008.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

9. _Thecosphæra micropora_, n. sp.

Cortical shell thin walled, smooth, with very small and numerous, regular,
circular pores, half as broad as the bars. Radial proportion of the three
shells = 1 : 2 : 5; they are connected by twenty regularly disposed radial
beams.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.08, inner 0.04;
cortical pores 0.002, bars 0.004.

_Habitat._--South Pacific, Station 288, surface.

Subgenus 3. _Thecosphærina_, Haeckel.

_Definition._--Pores of the cortical shell irregular polygonal, of unequal
size or dissimilar form.

10. _Thecosphæra capillacea_, n. sp.

Cortical shell thin walled, smooth, with irregular polygonal pores, three
to six times as broad as the bars. Both medullary shells with similar, but
smaller, pores. Radial proportion of the three spheres = 1 : 3 : 8; they
are connected by very numerous (sixty to eighty or more) thin radial beams.
(Similar to _Actinomma capillaceum_, Pl. 29, fig. 6, but without external
spines.)

_Dimensions._--Diameter of the outer shell 0.2, middle 0.075, inner 0.025;
cortical pores 0.006 to 0.012, bars 0.002.

_Habitat._--North Pacific, Station 250, surface.

11. _Thecosphæra diplococcus_, n. sp.

Cortical shell thick walled, rough, with large irregular polygonal pores,
two to three times as broad as the bars. Both medullary shells with small
regular circular pores. Radial proportion of the three spheres = 1 : 2 : 6;
they are connected by twenty (?) stout radial beams.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.04, inner 0.02;
cortical pores 0.008 to 0.012, bars 0.004.

_Habitat._--South Atlantic, Station 330, surface.

Subgenus 4. _Thecosphæromma_, Haeckel.

_Definition._--Pores of the cortical shell irregular, roundish, of unequal
size or dissimilar form.

{82}12. _Thecosphæra dodecactis_, n. sp.

Cortical shell thin walled, smooth, with large irregular roundish pores,
two to eight times as broad as the bars. Both medullary shells with regular
circular pores, twice as broad as the bars. Radial proportion of the three
spheres = 1 : 2 : 5; they are connected by twelve regularly disposed stout
radial beams.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.08, inner 0.04;
cortical pores 0.004 to 0.016, bars 0.002.

_Habitat._--Central Pacific, Station 263, depth 2650 fathoms.

13. _Thecosphæra icosactis_, n. sp.

Cortical shell thin walled, with small irregular roundish pores, two to
four times as broad as the bars. Both medullary shells with similar but
smaller pores. Radial proportion of the three spheres = 1 : 3 : 8; they are
connected by twenty thin radial beams.

_Dimensions._--Diameter of the outer shell 0.32, middle 0.12, inner 0.04;
cortical pores 0.005 to 0.012, bars 0.003.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

14. _Thecosphæra maxima_, n. sp.

Cortical shell thin walled, with small irregular roundish pores, two to six
times as broad as the bars. Both medullary shells with similar, but
smaller, pores. Radial proportion of the three spheres = 1 : 3 : 9; they
are connected by numerous (forty to sixty or more) thin radial beams.

_Dimensions._--Diameter of the outer shell 0.45, middle 0.15, inner 0.05;
cortical pores 0.008 to 0.024, bars 0.004.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

Genus 22. _Rhodosphæra_,[35] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Liosphærida# with one medullary (intracapsular) shell and
two cortical (extracapsular) shells.

The genus _Rhodosphæra_ differs from the preceding _Thecosphæra_ in the
same manner in which, among the Dyosphærida, _Liosphæra_ differs from
_Carposphæra_. The cortical shell is double, composed of two not far
distant shells, lying outside the central capsule. The distance between the
shells is much smaller than the radius of the inner shell. This is
connected by radial beams (piercing the central capsule) with the small
central medullary shell.

Subgenus 1. _Rhodosphærella_, Haeckel.

_Definition._--Pores of both cortical shells regular, in each shell all of
nearly equal size and form.

{83}1. _Rhodosphæra hexagonia_, n. sp.

Both cortical shells with the same number of regular hexagonal pores; the
inner four times as broad as the bars, and half as broad as the outer
pores, which are separated by thread-like bars. Medullary shell only
one-fourth as broad as the inner cortical shell, with regular hexagonal
pores of half the size. (Differs from the similar _Liosphæra hexagonia_,
Pl. 20, fig. 3, by the possession of a medullary shell.)

_Dimensions._--Diameter of the outer shell 0.2, middle 0.16, inner 0.04;
outer pores 0.013, middle 0.008, inner 0.004.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

2. _Rhodosphæra melitomma_, n. sp.

Both cortical shells with the same number of regular pores; the inner
regular, circular, twice as broad as the bars, with elegant hexagonal
frames and six roundish lobes alternating with the six radial spines which
arise from the hexagon-corners; these short conical spines are connected at
the distal end (at equal distances from the centre) by delicate threads
(three from each spine), which form the delicate external shell. Medullary
shell one-third as broad as the inner cortical shell, with small, simple,
regular circular pores. (If in _Carposphæra melitomma_, Pl. 20, fig. 4, the
distal ends of the spines became united by a cobweb-like outer shell, this
species would be formed.)

_Dimensions._--Diameter of the outer shell 0.22, middle 0.18, inner 0.06;
outer pores 0.025, middle 0.0125, inner 0.005.

_Habitat._--Central Pacific, Station 270, depth 2925 fathoms.

Subgenus 2. _Rhodosphæromma_, Haeckel.

_Definition._--Pores of both cortical shells irregular, in each shell of
unequal size or dissimilar form.

3. _Rhodosphæra palliata_, n. sp.

Both cortical shells with an unequal number of irregular roundish pores;
the outer pores somewhat smaller and much more numerous than the inner
pores; the bars between the latter are thicker. Medullary shell about one
quarter as broad as the inner cortical shell, with regular circular pores.

_Dimensions._--Diameter of the outer shell 0.4, middle 0.36, inner 0.08;
outer pores on an average 0.008, middle 0.012, inner 0.004.

_Habitat._--Fossil in Barbados.

4. _Rhodosphæra pentaphylla_, n. sp.

Both cortical shells with unequal number of irregular roundish pores; the
inner pores large, three to four times as broad as the bars; to each inner
pore corresponds a group of five smaller {84}outer pores, like the five
petals of a flower. Medullary shell half as broad as the inner cortical
shell, with regular circular pores.

_Dimensions._--Diameter of the outer shell 0.25, middle 0.2, inner 0.1;
outer pores on an average 0.006, middle 0.012, inner 0.004.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subfamily CROMYOSPHÆRIDA,[36] Haeckel, 1881, Prodromus, pp. 449, 453.

_Definition._--#Liosphærida# with four concentric spherical latticed
shells, united by radial beams.

Genus 23. _Cromyosphæra_,[37] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Liosphærida# with two intracapsular (medullary) shells and
two extracapsular (cortical) shells; the former united with the latter by
radial beams piercing the wall of the central capsule.

The genus _Cromyosphæra_ is the only known genus of Cromyosphærida, or of
such #Sphæroidea#, the smooth shell of which is composed of two medullary
and two cortical shells. There may possibly be other Cromyosphærida, in
which the shell is composed of one simple medullary and three cortical
shells, or only of four extra-capsular cortical shells; but such have not
as yet been observed. _Caryosphæra polysphærica_, Bütschli, 1882 (L. N. 41,
Taf. xxiii. fig. 12) is probably a _Cromyosphæra_ (fossil in Barbados).

1. _Cromyosphæra quadruplex_, n. sp. (Pl. 30, fig. 9).

Radial proportion of the four spheres = 1 : 2 : 4 : 5. Outer cortical shell
smooth, with large regular hexagonal pores, ten times as broad as the bars;
inner cortical shell with irregular polygonal pores, five times as broad as
the bars. Both medullary shells with regular circular pores of the same
breadth as the bars.

_Dimensions._--Diameter of the four spheres--(A) 0.16, (B) 0.12, (C) 0.06,
(D) 0.03.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

2. _Cromyosphæra rosetta_, n. sp.

Radial proportion of the four spheres = 1 : 2 : 8 : 10. Outer cortical
shell smooth, with regular hexagonal pores and very thin bars; inner
cortical shell with the same number of exactly corresponding, regular
circular, hexagonally framed pores, twice as broad as the bars; the corners
of the outer and inner hexagons united by radial bristles. Both medullary
shells with regular circular pores, twice as broad as the bars.

_Dimensions._--Diameter of the four spheres--(A) 0.2, (B) 0.16, (C) 0.04,
(D) 0.02.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

{85}3. _Cromyosphæra bigemina_, n. sp.

Radial proportion of the four spheres = 1 : 2 : 7 : 8. Outer cortical shell
smooth, with regular hexagonal pores and very thin bars; inner cortical
shell with the same number of exactly corresponding, regular circular
pores. Both medullary shells with regular circular pores of the same
breadth as the bars. (Somewhat similar to _Hexacromyon elegans_, Pl. 24,
fig. 9, also with six inner radial beams, but without external radial
spines.)

_Dimensions._--Diameter of the four spheres--(A) 0.2, (B) 0.17, (C) 0.05,
(D) 0.025.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

4. _Cromyosphæra cepa_, n. sp.

Radial proportion of the four spheres = 1 : 2 : 4 : 5. All four shells of
the same structure, thick-walled, with regular circular pores, two to four
times as broad as the bars; the size of the pores increases gradually from
the inner to the outer shell. Surface thorny. Distance between the second
and third shells twice as great as that between the others.

_Dimensions._--Diameter of the four spheres--(A) 0.125, (B) 0.1, (C) 0.05,
(D) 0.025.

_Habitat._--Fossil in Barbados.

5. _Cromyosphæra scorodonium_, n. sp.

Radial proportion of the four spheres = 1 : 2 : 3 : 4. All four shells of
the same structure, thin-walled, with irregular roundish pores, two to four
times as broad as the bars; the size of the pores increasing gradually from
the inner to the outer shell. Surface smooth. Distance between each two
shells equal to the diameter of the innermost.

_Dimensions._--Diameter of the four spheres--(A) 0.12, (B) 0.09, (C) 0.06,
(D) 0.03.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms; also fossil
in Barbados.

6. _Cromyosphæra antarctica_, n. sp..

Radial proportion of the four spheres = 1 : 2 : 5 : 7. Both cortical shells
with irregular polygonal roundish pores; the outermost with thinner bars
and rough surface, the inner with thicker bars. Both medullary shells with
irregular roundish pores.

_Dimensions._--Diameter of the four spheres--(A) 0.18, (B) 0.12, (C) 0.05,
(D) 0.025.

_Habitat._--Antarctic Ocean; in very large number, together with
_Rhizosphæra antarctica_, in the diatomaceous ooze of Station 157 (3rd
March 1874); depth 1950 fathoms.

Subfamily CARYOSPHÆRIDA,[38] Haeckel, 1881, Prodromus, pp. 449, 454.

_Definition._--#Liosphærida# with numerous (five or more) concentric
spherical latticed shells, united by radial beams.

{86}Genus 24. _Caryosphæra_,[39] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Liosphærida# with two intracapsular (medullary) shells and
three or more extracapsular (cortical) shells; the former united with the
latter by radial beams piercing the wall of the central capsule.

The genus _Caryosphæra_, the only observed form of this subfamily,
comprises those #Liosphærida# in which the shell is composed of two
medullary and three or more cortical shells. Such forms (without radial
spines) are very rare; I observed only two species, one with five, the
other with six shells. They are derived from _Cromyosphæra_ by further
apposition of outer cortical shells.

1. _Caryosphæra pentalepas_, n. sp.

Shell composed of five concentric spheres, with radial proportion =
1 : 2 : 8 : 10 : 12. Both medullary shells with regular circular pores,
twice as broad as the bars. First cortical shell with regular, circular,
hexagonally framed pores, three times as broad as the bars; second cortical
shell with regular hexagonal pores, four times as broad as the bars; third
(outermost) cortical shell with regular hexagonal pores and very thin
thread-like bars. Surface smooth.

_Dimensions._--Diameter of the five shells--(A) 0.02, (B) 0.04, (C) 0.16,
(D) 0.2, (E) 0.24.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

2. _Caryosphæra hexalepas_, n. sp.

Shell composed of six concentric spheres, with the radial proportion =
1 : 2 : 4 : 5 : 6 : 8. All six shells with regular circular pores, two to
four times as broad as the bars, with increasing size from the centrum
against the smooth surface.

_Dimensions._--Diameter of the six shells--(A) 0.025, (B) 0.05, (C) 0.1,
(D) 0.13, (E) 0.16, (F) 0.2.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms; also fossil
in Barbados.

Subfamily PLEGMOSPHÆRIDA,[40] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Liosphærida# with spongy spherical shell, with or without
latticed medullary shell in the centre.

Genus 25. _Styptosphæra_,[41] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Liosphærida# forming a solid sphere of spongy framework,
without enclosed medullary shell, and without central cavity.

{87}The genus _Styptosphæra_ presents a spherical shell with smooth or
rough surface (without radial spines), the whole mass of which is composed
of looser or denser spongy wicker-work.

1. _Styptosphæra spumacea_, n. sp.

Spongy framework of the solid sphere loose, with large polygonal meshes of
slightly different size, ten to twenty times as broad as the bars.
Structure of the whole spongy sphere the same. Central capsule filled with
crystals. Surface smooth.

_Dimensions._--Diameter of the sphere 0.32, of the central capsule 0.26,
meshes 0.01 to 0.02, bars 0.001.

_Habitat._--North Pacific, Station 236, surface.

2. _Styptosphæra spongiacea_, n. sp.

Spongy framework in the central part of the solid sphere much more compact
than in the peripheral part, becoming gradually looser towards the rough
surface. Meshes in the centre three to five times, in the periphery fifteen
to twenty times as broad as the bent bars.

_Dimensions._--Diameter of the sphere 0.45, inner meshes 0.006 to 0.01,
outer meshes 0.03 to 0.04, bars 0.002.

_Habitat._--Central Pacific, Station 271, surface.

3. _Styptosphæra stupacea_, n. sp.

Spongy framework of the solid sphere rather compact, everywhere of the same
structure, with roundish, nearly equal meshes, six to eight times as broad
as the bars. Surface rough with prominent thorns.

_Dimensions._--Diameter of the sphere 0.22, meshes 0.01 to 0.012, bars
0.0015.

_Habitat._--South Pacific, Station 291, surface.

Genus 26. _Plegmosphæra_, Haeckel,[42] 1881, Prodromus, p. 455.

_Definition._--#Liosphærida# forming a hollow sphere of spongy framework,
without a medullary shell in the central cavity.

The genus _Plegmosphæra_ develops a large hollow sphere, the wall of which
is composed of looser or denser spongy wicker-work. On the inner as well as
on the outer face of the spongy shell-wall may be present a simple
lattice-sphere from which the threads of the framework arise; but in some
species these lattice-plates are quite absent.

{88}Subgenus 1. _Plegmosphærantha_, Haeckel.

_Definition._--Inside and outside of the spongy shell-wall smooth, closed
by a lattice-plate with polygonal meshes.

1. _Plegmosphæra maxima_, n. sp.

Radius of the spherical shell-cavity eight to ten times as great as the
thickness of the thin spongy wall. Inside and outside of the wall smooth,
closed by a lattice-plate with irregular polygonal meshes. Only three or
four meshes in the thickness of the wall.

_Dimensions._--Diameter of the spongy sphere 0.8 to 1.0 mm., of its inner
cavity 0.7 to 0.8, meshes 0.01 to 0.02, bars 0.002 to 0.003, central
capsule 0.5 to 0.6, nucleus 0.1 to 0.15.

_Habitat._--Central Pacific, Station 271, surface.

2. _Plegmosphæra coelopila_, n. sp.

Radius of the spherical shell-cavity eight to ten times as great as the
thickness of the spongy wall. Inside and outside of the wall closed by a
smooth lattice-plate with irregular polygonal meshes, five to ten times as
broad as the bars.

_Dimensions._--Diameter of the shell 0.32, of its cavity 0.26, meshes 0.01
to 0.02, bars 0.002.

_Habitat._--North Atlantic, Færöe Channel, Gulf Stream, John Murray.

3. _Plegmosphæra pachypila_, n. sp.

Radius of the spherical shell-cavity about equal to the thickness of the
spongy wall. Inside and outside of the wall closed by a smooth
lattice-plate with irregular polygonal meshes, three to six times as broad
as the bars.

_Dimensions._--Diameter of the shell 0.24, of its cavity 0.12.

_Habitat._--North Pacific, Station 250, surface.

Subgenus 2. _Plegmosphærella_, Haeckel.

_Definition._--Inside of the spongy shell-wall closed by a smooth
lattice-plate, outside rough, spongy, with prominent thorns.

4. _Plegmosphæra entodictyon_, n. sp.

Radius of the spherical shell-cavity half as great as the thickness of the
spongy wall. Inside of the wall closed by a smooth lattice-plate, outside
rough, spongy.

_Dimensions._--Diameter of the shell 0.24, of its cavity 0.08.

_Habitat._--South Pacific, Station 300, surface.

{89}5. _Plegmosphæra leptodictyon_, n. sp.

Radius of the spongy shell-cavity six times as great as the thickness of
the spongy wall. Inside of the wall closed by a smooth lattice-plate,
outside rough spongy.

_Dimensions._--Diameter of the shell 0.44, of its cavity 0.36.

_Habitat._--Central Pacific, Station 266, surface.

Subgenus 3. _Plegmosphæromma_, Haeckel.

_Definition._--Inside of the spongy shell-wall rough spongy, without
lattice-plate, outside closed by a smooth lattice-plate.

6. _Plegmosphæra exodictyon_, n. sp. (Pl. 18, fig. 8).

Radius of the spongy shell-cavity only one-fourth as great as the thickness
of the spongy shell-wall. Outside of the wall closed by a smooth
lattice-plate, inside rough, spongy.

_Dimensions._--Diameter of the shell 0.4, of its cavity 0.08.

_Habitat._--South Atlantic, Station 325, surface.

Subgenus 4. _Plegmosphærusa_, Haeckel.

_Definition._--Inside and outside of the spongy shell-wall rough, with
spongy or spiny surface, without lattice-plate.

7. _Plegmosphæra leptoplegma_, n. sp.

Radius of the spherical shell-cavity half as great as the thickness of the
loose spongy shell-wall. Inside and outside of the wall rough spongy, not
closed by a lattice-plate. Meshes ten to twenty times as broad as the bars.

_Dimensions._--Diameter of the shell 0.3, of its cavity 0.088.

_Habitat._--North Atlantic, Station 253, surface.

8. _Plegmosphæra pachyplegma_, n. sp.

Radius of the spherical shell-cavity about equal to the thickness of the
dark and dense spongy shell-wall. Inside and outside of the wall rough
spongy, not closed by a lattice-plate. Meshes three to five times as broad
as the bars.

_Dimensions._--Diameter of the shell 0.2, of its cavity 0.1.

_Habitat._--Central Pacific, Station 270, surface.

Genus 27. _Spongoplegma_,[43] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Liosphærida# forming a sphere of spongy framework, which
encloses in the centre one single latticed medullary shell.

{90}The genus _Spongoplegma_ may be regarded as a _Carposphæra_, in which
the simple latticed cortical shell is represented by an irregular spongy
framework, immediately enclosing the simple latticed medullary shell.

1. _Spongoplegma antarcticum_, n. sp.

Cortical shell with loose spongy framework and rough surface, four to six
times as broad as the enclosed simple medullary shell. Pores of the latter
regular circular, twice as broad as the bars. From its surface arise
numerous (forty to fifty or more) short radial beams, which become forked
and compose, by communication of lateral branches, the spongy cortical
shell.

_Dimensions._--Diameter of the spongy cortical shell 0.2 to 0.3 of the
latticed medullary shell 0.05 to 0.06.

_Habitat._--Antarctic Ocean, in large number, together with _Cromyosphæra
antarctica_; in the Diatom ooze of Station 157 (depth 1950 fathoms).

Genus 28. _Spongodictyon_,[44] Haeckel, 1862, Monogr. d. Radiol., p. 459.

_Definition._--#Liosphærida# forming a sphere of spongy framework, which
encloses in the centre a double latticed concentric medullary shell.

The genus _Spongodictyon_ can be regarded as a _Thecosphæra_, in which the
simple latticed cortical shell is represented by an irregular spongy
framework, which immediately encloses the double latticed medullary shell.
Sometimes this latter appears triple, the inner surface of the spongy
cortical shell forming a smooth spherical lattice-plate, separated by an
interval from the double medullary shell.

Subgenus 1. _Dictyoplegma_, Haeckel, 1862, Monogr. d. Radiol, p. 458.

_Definition._--Spongy cortical shell enveloping immediately the double
medullary shell.

1. _Spongodictyon spongiosum_, Haeckel.

_Dictyosoma spongiosum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 31, Taf. ii. figs. 9-11.

_Dictyoplegma spongiosum_, Haeckel, 1862, Monogr. d. Radiol., p. 458.

Spongy framework of the cortical shell loose, with large, polygonal
roundish meshes, on an average as large as the double medullary shell,
which is immediately enveloped by it. Both concentric medullary shells with
subregular roundish pores, twice as broad as the bars.

_Dimensions._--Diameter of the cortical shell 0.2 to 0.3 or more; of the
outer medullary shell 0.03, inner 0.01.

_Habitat._--Mediterranean (French south coast, surface), J. Müller.

{91}2. _Spongodictyon cavernosum_, n. sp.

Spongy framework of the cortical shell rather compact in the inner part,
which immediately envelops the double medullary shell; very loose, with
large caverns in the outer part, caverns of the surface larger than the
medullary shell. Both medullary shells with regular circular pores, three
times as broad as the bars.

_Dimensions._--Diameter of the cortical shell 0.3 to 0.4, outer medullary
shell 0.1, inner 0.03.

_Habitat._--Tropical Atlantic, Station 338, surface.

Subgenus 2. _Spongodictyoma_, Haeckel.

_Definition._--Spongy cortical shell on the inner surface with a smooth
lattice-plate (or third medullary shell), which is connected by radial
beams with the inner double medullary shell.

3. _Spongodictyon trigonizon_, Haeckel.

_Spongodictyon trigonizon_, Haeckel, 1862, Monogr. d. Radiol., p. 459,
Taf. xxvi. figs. 4-6.

_Dictyosoma trigonizon_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 841.

Spongy framework of the cortical shell very loose, with very large, for the
most part triangular meshes, which are two to six times as large as the
enclosed double medullary shell. From the surface of the latter arise
numerous radial beams, which are connected by a spherical lattice-plate,
forming the smooth inner surface of the spongy sphere (or a third medullary
shell). The structure of the framework reminds one of the PHÆODARIUM
_Sagena_ (Pl. 108). Pores of both medullary shells regular circular, twice
as broad as the bars.

_Dimensions._--Diameter of the cortical shell 0.5 to 1.15, outer medullary
0.05, inner 0.035.

_Habitat._--Mediterranean, Messina, surface.

4. _Spongodictyon arcadophoron_, n. sp.

Spongy framework of the cortical shell in the inner part very loose, in the
outer part more compact; outer meshes scarcely as large as the inner
medullary shell (or only half as large), inner meshes two to four times as
large. From the surface of the double medullary shell arise numerous radial
beams, which are forked at equal distances from the centre; the fork
branches are curved and united together by dichotomous branches, like
elegant arcades; and these arcades form together the large polygonal meshes
on the inside of the cortical shell (or a third medullary shell). Both
medullary shells with regular circular pores, of the same breadth as the
bars.

_Dimensions._--Diameter of the cortical shell 0.2, outer medullary shell
0.04, inner 0.02.

_Habitat._--Tropical Atlantic, Station 349, surface.

{92}Family VI. #COLLOSPHÆRIDA#, J. Müller[45] (Pls. 5-8).

_Definition._--#Sphæroidea# living associated in colonies, united by an
alveolar jelly-body, and connected by the network of anastomosing
pseudopodia.

The family #Collosphærida# comprises all polyzous or social #Sphæroidea#,
and constitutes the only polyzoic group among the SPHÆRELLARIA. This group
was first constituted by J. Müller as "_Radiolaria polyzoa_ with
shells."[46] Formerly following his authority, in my Monograph I had
separated them from the other #Sphæroidea# and united them with the social
Collodaria (Sphærozoida).[47] Also R. Hertwig in his Organismus der
Radiolarien[48] united them with his Sphærozoea. In my Prodromus[49] I had
retained this isolated position. But a further careful study has convinced
me that this isolation is not truly natural, and that the Collosphærida are
only "social Ethmosphærida" which have arisen from this solitary subfamily
by adaptation to colonial life. There are some forms of Collosphærida which
are nearly identical with some forms of Ethmosphærida, only differing from
the latter by their association in colonies; and in some forms of both
groups it is quite impossible to decide whether the isolated shells
appertain to one or to the other family.

The isolated shell of the Collosphærida is almost constantly (with few
exceptions) a simple extracapsular lattice-shell, as in the Monosphærida;
only the small group of Clathrosphærida (with the genera _Clathrosphæra_
and _Xanthiosphæra_) exhibit an exception, the simple lattice-shell being
overgrown by an external mantle or veil of very thin, cobweb-like,
irregular lattice-work (Pl. 8, figs. 6-11). Therefore these Clathrosphærida
bear to the Acrosphærida (or the common simple Collosphærida) a relation
similar to that which _Liosphæra_ (p. 76) bears to _Cenosphæra_; both
shells are extracapsular "cortical shells" at a very short distance apart.
In the Collosphærida true concentric medullary shells never occur; the
central capsule always lies quite freely in the simple or double cortical
shell, separated from it by a jelly-veil.

Although a well marked difference in the simple lattice-shell of the social
Collosphærida and the solitary Ethmosphærida does not exist, nevertheless
in most cases the two shells can be distinguished by a practiced observer.
The simple fenestrated shells of the monozoic Ethmosphærida are commonly
quite regular spheres in a mathematical sense, or regular "endospherical
polyhedra"; whereas in the Collosphærida they are commonly more or less
irregular, often to an extraordinary degree (Pls. 5-8). Some species of
Collosphærida, however, also possess quite regular spherical shells.
Another difference is often shown in the lattice-work of the shells, which
in the Collosphærida is nearly always very irregular, and exhibits a
peculiar tendency to the {93}production of radial, conical, or cylindrical
tubules. These occur as well on the inside as on the outside of the shell,
and the tubules are now more conical, now more cylindrical; their wall
either solid or pierced by pores (Pls. 5-8). The tubules are commonly very
irregular in form, size, and disposition; distinguished, however, by a
number of hereditary peculiarities, which are sufficient for the
distinction of genera. Similar tubules occur also in some genera of
solitary Ethmosphærida (_Coscinomma_, _Ethmosphæra_, _Conosphæra_, &c., Pl.
12); but the tubules are here much more regular and not so highly
developed.

Besides the tubules of the fenestrated shells, in some genera of
Collosphærida the surface is armed with irregular thorns, rarely with more
regular radial spines. But these spines obtain constantly the character of
accessory by-spines, and remain short and thin. In this family typical
radial spines never occur in a regular and characteristic disposition,
corresponding to dimensive axes, as is the case in nearly all solitary
#Sphæroidea#, only excepting the Liosphærida. Commonly these spines or
thorns serve as protective arms for the shell-meshes, surrounding them
often in the form of coronels. Often the lattice-plate of the irregular
roundish shell is tubercular, elevated into irregular protuberances,
bearing on the top a short spine or thorn (Pl. 8).

The _Central Capsule_ of the Collosphærida is always a regular sphere, as
in all other #Sphæroidea#; it is constantly placed within the
lattice-shell, and commonly much smaller than it, separated from it by a
thick jelly-veil. A remarkable difference from the solitary #Sphæroidea# is
shown in the early division of the nucleus. Commonly the central capsule of
the Collosphærida contains in its centre a large oil-globule, surrounded by
very numerous small nuclei. R. Hertwig estimated this difference as so
important, that he separated the social "Sphærozoea" and the solitary
"Peripylea" as two different orders. As already shown above (p. 7, 24), we
cannot support this separation, and are now convinced that this difference
in the development of the spores--just as in the #Collodaria#--is the
consequence of an adaptation to social life.

The common jelly-body, in which the numerous central capsules and their
enveloping shells are united, exhibits in the Collosphærida quite the same
characters as in the other social Radiolaria, the Collozoida and
Sphærozoida. The jelly-body is very voluminous, commonly spherical, often
cylindrical, of considerable size; constantly containing numerous large
alveoles. Often each shell is enclosed in a separate alveole with rather
solid wall (Pl. 6, fig. 2). Sometimes in the dead colonies all shells are
united in the central part of the jelly-body, whilst its peripheral part is
composed of a stratum of large alveoles (Pl. 8, fig. 11); at other times no
alveoles are visible (Pl. 7, fig. 11). In many living colonies I found a
very large spherical alveole with thick wall in the centre of the spherical
colony, surrounded by many strata of delicate thin-walled alveoles (Pl. 5,
fig. 1). In this case often the inner younger capsules were naked,
{94}without shells, the outer only surrounded by shells. Already in my
Monograph I had described the same peculiar formation.[50]

_Synopsis of the Genera of Collosphærida._

-------------------------------------------------------------------------
I. Subfamily Acrosphærida. (Lattice-shell simple, without an external
mantle of network.)
-------------------------------------------------------------------------
{Inside {Inside smooth, 29. _Collosphæra_.
Outside of { without {
the shell { tubuli. {Inside spiny, 30. _Tribonosphæra_.
smooth, {
without { {Tubuli
spines or {Inside with { imperforated, 31. _Pharyngosphæra_.
tubuli. { centripetal {
{ tubuli. {Tubuli
{ { fenestrated, 32. _Buccinosphæra_.

{Spines
{ irregularly
{ scattered on
{ the surface, 33. _Acrosphæra_.
{
Outside of the shell armed {Each larger
with solid spines, but { pore with one
with hollow tubuli. { single spine, 34. _Odontosphæra_.
{
{Each larger
{ pore with
{ a coronal of
{ spines, 35. _Choenicosphæra_.

{Mouth of the
{ tubuli
{ truncated,
{ smooth, 36. _Siphonosphæra_.
{
Outside of {Tubuli {Mouth with
the shell { simple, not { one single
with { branched. { large tooth, 37. _Mazosphæra_.
irregular { {
radial { {Mouth with a
tubuli, { { coronal of
the wall of { { teeth, 38. _Trypanosphæra_.
which is {
solid, not {Tubuli irregularly branched,
fenestrated.{ each with two to four or
{ more openings, 39. _Caminosphæra_.

{Mouth of the
{ tubuli
{ truncated,
{ smooth, 40. _Solenosphæra_.
Outside of the shell with {
irregular radial tubuli, {Mouth with one
open on both ends, { single large
with fenestrated wall. { tooth, 41. _Otosphæra_.
{
{Mouth with a
{ coronal of
{ teeth, 42. _Coronosphæra_.
-------------------------------------------------------------------------
II. Subfamily Clathrosphærida. (Lattice-shell double, with an external
mantle of network.)
-------------------------------------------------------------------------
Surface of the outer shell smooth, 43. _Clathrosphæra_.

Surface of the outer shell thorny, 44. _Xanthiosphæra_.

Subfamily ACROSPHÆRIDA, Haeckel, 1881, Prodromus, p. 471.

_Definition._--#Collosphærida# with one simple lattice-shell around every
central capsule of the coenobium.

{95}Genus 29. _Collosphæra_,[51] J. Müller, 1855.

_Definition._--#Collosphærida# with simple shells, smooth on the inside and
on the outside, without any spines or tubuli.

The genus _Collosphæra_ is the most simple form of all Collosphærida, and
must be regarded as the common ancestral form of this family. As the
lattice-shell is quite a simple sphere, without any spines, tubules, or
other peculiar productions, it agrees perfectly with _Cenosphæra_, and
represents the social or polyzoid aggregate of this solitary or monozoid
genus. Therefore a certain distinction between the isolated shells of the
two genera is often very difficult or even impossible; but commonly this
distinction is possible owing to the circumstance, that in the majority of
the _Collosphæræ_ the shell is more or less irregular roundish or
polyhedral, not quite spherical, as in _Cenosphæra_. _Dermatosphæra_,
Ehrenberg, is a _Collosphæra_ with small pores (compare L. N. 16, p. 533).

Subgenus 1. _Eucollosphæra_, Haeckel.

_Definition._--Shell a regular or subregular sphere.

1. _Collosphæra primordialis_, n. sp.

Shell a regular sphere, with very delicate and regular network of hexagonal
meshes. Six to eight meshes in the half meridian of the shell. Diameter of
the meshes ten to fifteen times as broad as the thin bars between them. Can
be regarded as social form of _Cenosphæra primordialis_.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.008.

_Habitat._--Central Pacific, Stations 271 to 274, surface.

2. _Collosphæra regularis_, n. sp.

Shell a regular sphere, with a perfectly regular network of circular
meshes, all of the same size. Ten to twelve meshes in the half meridian of
the shell. Diameter of the meshes the same as that of the bars between
them.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.005 to
0.006.

_Habitat._--Indian Ocean, Madagascar, surface, Rabbe.

3. _Collosphæra globularis_, n. sp.

Shell a regular sphere, with subregular network of circular meshes of
different sizes; few large pores between many smaller pores. Ten to twenty
meshes in the half meridian of the shell. Diameter of the meshes from half
to twice as broad as that of the bars.

{96}_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.002 to
0.008, breadth of the bars 0.004 to 0.008.

_Habitat._--Tropical and subtropical zone of both hemispheres, widely
distributed; Canaries, Azores, Cape Verde Islands, Guinea Coast, Brazil
Coast, Indian Ocean, Madagascar, Ceylon, surface.

Subgenus 2. _Dyscollosphæra_, Haeckel.

_Definition._--Shell not a regular sphere, but irregular roundish, in all
degrees of variation between subspherical and polyhedral or quite irregular
forms.

4. _Collosphæra huxleyi_, J. Müller.

_Collosphæra huxleyi_, J. Müller, 1855, Abhandl. d. k. Akad. d. Wiss.
Berlin, pp. 55-59, Taf. viii. figs. 6-9.

_Collosphæra huxleyi_, Haeckel, 1862, Monogr. d. Radiol., p. 534, Taf.
xxxiv.

_Collosphæra huxleyi_, Cienkowski, 1871, Archiv f. mikrosk. Anat., Bd.
vii. p. 374, Taf. xxix. figs. 1-6.

_Collosphæra ligurina_, J. Müller, 1856, Monatsber. d. k. Akad. d. Wiss.
Berlin, p. 481.

_Thalassicolla punctata_, var., Huxley, 1851, Ann. and Mag. Nat. Hist.,
ser. 2, vol. viii. p. 434, pl. xvi. fig. 6.

Shell subspherical, somewhat irregular, sometimes with more or less
superficial impressions, with irregular network of roundish meshes. Eight
to sixteen meshes in the half meridian of the shell, one to three times as
broad as their bars. Very variable, with direct transition-forms to other
species of this genus, especially to _Collosphæra globularis_, _Collosphæra
tuberosa_, _Collosphæra pyriformis_, and _Collosphæra polyedra_.

_Dimensions._--Diameter of the shell 0.1 to 0.16, of the pores 0.004 to
0.012, of the bridges 0.003 to 0.006.

_Habitat._--Cosmopolitan; common in the greater part of the warmer seas,
surface.

5. _Collosphæra polygona_, n. sp. (Pl. 5, fig. 13).

_Collosphæra huxleyi_ var., Haeckel, 1862, Monogr. d. Radiol., Taf.
xxxiv. fig. 5.

Shell irregular polygonal, with very delicate, irregular network of
polygonal meshes, four to twelve times as broad as the bars. Ten to twenty
pores on the half meridian of the shell.

_Dimensions._--Diameter of the shell 0.1 to 0.2, of the pores 0.012 to
0.004, of the bars 0.001 to 0.002.

_Habitat._--Mediterranean, Atlantic, surface; Stations 348 to 354.

6. _Collosphæra pyriformis_, Haeckel, n. sp.

Shell irregular, rounded, ovate or pear-shaped, with irregular network of
rounded or nearly polygonal meshes. Ten to twenty meshes in the half
meridian of the shell, one to three times as broad as the bars. Commonly
one large opening (two to three times as broad as the largest {97}meshes)
on the thinner end of the ovate shell (corresponding to the insertion of a
pear-stalk); sometimes two or three such large openings.

_Dimensions._--Diameter of the shell 0.1 to 0.15, of the pores 0.008 to
0.016, of the bridges 0.004 to 0.008.

_Habitat._--Tropical zone--Cape Verde Islands, Ceylon; Central Pacific,
Stations 266 to 272, 348 to 352, &c.

7. _Collosphæra polyedra_, n. sp.

_Trisolenia zanguebarica_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 301, Taf. x. fig. 11.

Shell irregular, polyhedral, with even or somewhat vaulted sides, and
obtuse ridges between them. Network more or less irregular, with small
rounded meshes, quite as broad or twice as broad as their bars. Besides
these small pores constantly some large round openings (commonly three to
six), situated on the corners of the polyhedral shell, four to six times as
large as the pores. Often an acute tooth on the edge of each large opening.
Transition-form between _Collosphæra_ and _Solenosphæra_ or _Odontosphæra_.

_Dimensions._--Diameter of the shell 0.1 to 0.15, of the pores 0.004 to
0.008, of their bridges 0.004, of the large openings 0.24 to 0.032.

_Habitat._--Tropical zone of the Pacific and the Indian Ocean; Stations 266
to 272, surface.

8. _Collosphæra tuberosa_, n. sp.

_Collosphæra huxleyi_, var., Haeckel, 1862, Monogr. d. Radiol., Taf.
xxxiv. figs. 3, 9.

Shell very irregular, between subspherical and polyhedral in form, but
always with irregular impressions, boils or bosses, and between these
different rounded prominent tubercles and ridges. Network irregular,
strong, with rounded, subcircular or nearly polygonal meshes. Ten to thirty
pores in the half meridian of the shell. Diameter of the meshes half to
four times as broad as that of the thick bars.

_Dimensions._--Diameter of the shell very variable in the same coenobium,
0.05 to 0.2, of the pores 0.002 to 0.008, breadth of the bridges 0.004 to
0.006.

_Habitat._--Cosmopolitan, common in all warmer seas, surface.

9. _Collosphæra irregularis_, n. sp.

_Collosphæra huxleyi_, var., Haeckel, 1862, Monogr. d. Radiol., Taf.
xxxiv. fig. 8.

Shell quite irregular, knotty or bulbous, with irregular impressions, and
prominent knobs or bulbs between them. Network thin, fragile, quite
irregular, with polygonal meshes of most unequal size and form. Five to
twenty pores in the half meridian of the shell. Diameter of the meshes two
to ten times as broad as that of the thin bars.

_Dimensions._--Diameter of the shell very variable in the same coenobium,
0.04 to 0.24, of the pores 0.005 to 0.05, of the bridges 0.002 to 0.004.

_Habitat._--Mediterranean, Atlantic, not common; Stations 348, 352, &c.,
surface.

{98}Genus 30. _Tribonosphæra_,[52] Haeckel, 1881, Prodromus, p. 471.

_Definition._--#Collosphærida# with simple shells, on the inside with
radial centripetal beams.

The genus _Tribonosphæra_ differs from _Collosphæra_ by a very peculiar and
rare character, the development of centripetal radial sticks on the
internal face of the shell; these beams are not united in the centrum, but
finish freely in a certain distance from it.

1. _Tribonosphæra centripetalis_, n. sp. (Pl. 5, fig. 12).

Shell roundish or subspherical, with numerous small circular or roundish
pores, about twice as broad as the bars. Twenty to thirty pores on the half
meridian of the shell. Outside of the shell smooth, inside a variable
number (ten to twenty) of thin, radial, centripetal sticks or spines,
one-third or one-half as long as the radius of the shell. (In the central
capsule many very large crystals, resting after the destruction of the
capsule.)

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.003 to
0.005, of the bridges 0.001 to 0.002; length of the inner centripetal
sticks 0.02 to 0.03.

_Habitat._--Central Pacific, Station 271, surface.

Genus 31. _Pharyngosphæra_,[53] n. gen.

_Definition._--#Collosphærida# with simple shells, having on the inside
radial centripetal tubes, the walls of which are solid.

The genus _Pharyngosphæra_ differs from _Collosphæra_ by the development of
radial tubules on the inside of the shell, which are directed centripetally
towards its centre. The wall of the tubule is solid, not latticed as in the
following genus.

1. _Pharyngosphæra stomodæa_, n. sp. (Pl. 5, fig. 10).

Shell irregular polyhedral, with ten to fifteen polygonal faces and rounded
edges. Pores very small, circular, irregularly scattered, smaller than the
bars. Twelve to fifteen pores on the half meridian of the shell. On the
inside of every shell-face one short, nearly cylindrical, centripetal
tubule, twice as long as broad, and about one-third as long as the shell
radius. Outer umbilical mouth of the tubules somewhat wider than the inner
truncated mouth.

_Dimensions._--Diameter of the shell 0.11 to 0.12, of the pores 0.003 to
0.005, of the bars 0.01 to 0.02; length of the inner tubuli 0.02, breadth
of them 0.01.

_Habitat._--South Pacific, Station 288, depth 2600 fathoms.

{99}Genus 32. _Buccinosphæra_,[54] n. gen.

_Definition._--#Collosphærida# with simple shells, having on the inside
radial centripetal tubes, the walls of which are fenestrated.

The genus _Buccinosphæra_ exhibits, on the inner surface of the shell,
radial centripetal tubules similar to those of the foregoing
_Pharyngosphæra_; but the walls of these tubes are here latticed, not
solid; they represent therefore true invaginations of the whole shell-wall.

1. _Buccinosphæra invaginata_, n. sp. (Pl. 5, fig. 11).

Shell irregular roundish or nearly polyhedral, with a variable number of
umbilical depressions, which are prolonged on the inside into cylindrical
or somewhat conical, centripetal, fenestrated tubes, about one-third as
long as the shell radius. Inner mouth of the tubes narrower, scarcely half
as broad as the outer mouth, about equal to one-fourth the shell radius,
truncated. Pores of the tubes and of the shell small, roundish, irregular
in size and distribution, about as broad as the bars. Twenty-five to thirty
pores in the half meridian of the shell. In all observed specimens the
spherical central capsule (half as broad as the shell) contained a large
number of crystals.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.001 to
0.003, of the bars 0.002 to 0.003; length of the tubuli 0.02; outer mouth
0.026, inner mouth 0.013; crystals in the central capsule 0.002 to 0.004,
sometimes 0.008.

_Habitat._--Philippine Islands (Samboangan), Station 213, surface.

2. _Buccinosphæra tubaria_, n. sp.

Shell irregular polyhedral with rounded edges, with a variable number of
umbilical depressions, which are prolonged on the inside into large, nearly
cylindrical, centripetal, fenestrated tubes, half as long as the shell
radius. In the middle the tubes are somewhat constricted and narrower.
Inner mouth of the tubes dilated, nearly as broad as the outer mouth, about
equal to one-half the shell-radius, truncated. Pores of the tubes and of
the shell large, roundish polygonal, irregular in size and distribution,
three to four times as broad as the bars. Fifteen to twenty pores in the
half meridian of the shell.

_Dimensions._--Diameter of the shell 0.12 to 0.14, of the pores 0.008 to
0.012, of the bars 0.002 to 0.004; length of the tubuli 0.03; outer mouth
0.04, inner mouth 0.03.

_Habitat._--North coast of New Guinea, Station 217, surface.

Genus 33. _Acrosphæra_,[55] Haeckel, 1881, Prodromus, p. 471.

_Definition._--#Collosphærida# with simple shells, the outer surface of
which is covered with radial, irregularly scattered spines.

{100}The genus _Acrosphæra_ differs from its ancestral genus _Collosphæra_
by the development of spines on the outer surface of the shell. These are
either short, straight, radial spines, or oblique and often curved; their
base is often inflated; they are irregularly scattered on the whole surface
between the pores.

1. _Acrosphæra erinacea_, n. sp.

Shell a regular sphere, everywhere covered with small, very numerous,
straight radial spines, regularly scattered between the pores. In the half
meridian of the shell ten to twelve circular pores, all of the same form
and size, double as broad as the bars. Spines bristle-shaped, very thin,
solid, about as long as the diameter of the pores.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.008 to
0.012; length of the spines 0.01.

_Habitat._--Tropical zone of the Atlantic, coast of Brazil, Rabbe, surface.

2. _Acrosphæra echinoides_, n. sp. (Pl 8, fig. 1).

Shell a regular sphere, covered with numerous, straight, radial spines,
irregularly scattered over the whole surface. In the half meridian of the
shell twenty to thirty irregular roundish pores of variable size, one to
four times as broad as the bars. Spines conical, strong, quite radial, at
the top of small conical elevations, which are perforated by from three to
six pores.

_Dimensions._--Diameter of the shell 0.12 to 0.15, of the pores 0.002 to
0.008; length of the spines 0.015, of their basal zones 0.01.

_Habitat._--South-east corner of the Pacific, Valparaiso, Station 298,
surface.

3. _Acrosphæra setosa_, Haeckel.

_Polysolenia setosa_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 299, Taf. viii. fig. 10.

Shell a regular sphere, covered with numerous bristle-shaped radial spines,
irregularly scattered between the pores. In the half meridian of the shell
two to four very large circular pores (equal to one-third the radius), and
between them numerous very small, point-like pores.

_Dimensions._--Diameter of the shell 0.05 to 0.08, of the large pores 0.01,
of the small 0.001; length of the spines 0.01 to 0.02.

_Habitat._--West Tropical Pacific, Philippine Sea, Station 206, depth 2100
fathoms.

4. _Acrosphæra spinosa_, Haeckel.

_Collosphæra spinosa_, Haeckel, 1862, Monogr. d. Radiol., p. 536, Taf.
xxxiv. figs. 12, 13.

_Collosphæra spinosa_, Cienkowsky, 1871, Archiv f. mikrosk. Anat., vii.
p. 374, Taf. xxix. figs. 7-17.

Shell a regular or subregular sphere, covered with numerous, obliquely
standing spines, irregularly scattered over the surface. In the half
meridian of the shell fifteen to twenty irregular {101}roundish pores of
very different form and size, one to four times as broad as their bars.
Spines conical, irregularly diverging and curved, their hollow base
perforated by several pores, not longer than the diameter of the largest
pores.

_Dimensions._--Diameter of the shell 0.1 to 0.2, of the pores 0.001 to
0.04; length of the spines 0.01 to 0.02.

_Habitat._--Mediterranean, Messina; Canary Islands, Haeckel.

5. _Acrosphæra collina_, n. sp. (Pl. 8, fig. 2).

Shell quite irregular, polyhedrical, hilly, with a variable number (eight
to sixteen) of large conical hill-like prominences; every cone or hill
about as high as broad, perforated by the same pores as the shell, on its
top bearing a larger irregular roundish pore, and on its edge one single
bristle-like spine, not larger than the diameter of this pore, obliquely
inserted. In the half meridian of the shell twenty to thirty irregular
roundish pores of very different size, one to six times as broad as the
bars. A very characteristic species, closely resembling the following
_Odontosphæra_.

_Dimensions._--Diameter of the shell 0.15 to 0.25, of the pores 0.005 to
0.02; length of the spines 0.01 to 0.02, height of the hills from which
they rise 0.03 to 0.04.

_Habitat._--North coast of New Guinea, Station 218, surface.

6. _Acrosphæra inflata_, n. sp. (Pl. 5, fig. 7).

_Mazosphæra inflata_, Haeckel, 1879, Atlas, _loc. cit._

Shell more or less irregular, polyhedral, hilly, with a variable number
(six to twelve) of large pyramidal, hill-like prominences; every hill about
as high as broad, on the top a strong conical, radial, or obliquely
inserted spine, inflated, with three to six very large polygonal meshes,
much larger than the other pores between the hills, which are also
polygonal, two to six times as broad as the bars. Ten to fifteen pores on
the half meridian.

_Dimensions._--Diameter of the shell 0.1 to 0.14, of the largest pores
0.05, of the smallest 0.005; length of the spines 0.02 to 0.03.

_Habitat._--North Atlantic, Station 64, surface.

Genus 34. _Odontosphæra_,[56] n. gen.

_Definition._--#Collosphærida# with simple shells, the outside of which
bears single scattered spines, one single spine on the margin of each
larger pore.

The genus _Odontosphæra_ is distinguished from the foregoing _Acrosphæra_
by the peculiar disposition of the spines, which are not scattered on the
outside of the shell between the pores, but so disposed that each larger
pore is protected by one single spine, obliquely placed over it.

{102}1. _Odontosphæra monodon_, n. sp. (Pl. 5, fig. 5).

Shell spherical or subspherical, with very small and numerous circular
pores, much smaller than the bars. Twelve to fifteen pores on the half
meridian of the shell. Between them a variable number of larger roundish
apertures (mostly twelve to sixteen) irregularly scattered, one-fourth to
one-fifth as broad as the shell radius. On the margin of every larger
aperture a single (rarely two or three) sharp conical tooth, about as long
as the diameter of the aperture, and obliquely laid over them.

_Dimensions._--Diameter of the shell 0.1 to 0.13, of the pores 0.001 to
0.003, of the bars 0.01 to 0.02, of the larger apertures 0.01 to 0.02.

_Habitat._--Sunda Archipelago, Station 192, surface.

2. _Odontosphæra cyrtodon_, n. sp. (Pl. 5, fig. 6).

Shell spherical or subspherical, with numerous roundish pores of very
irregular size and distribution, mostly larger than the bars. Ten to twelve
pores on the half meridian of the shell. Between them a variable number
(mostly six to nine) of large roundish pores, about half as broad as the
shell radius, armed on one side of the margin with one single large tooth,
about as long as the diameter of the aperture, hook-like, curved, and
obliquely laid over them.

_Dimensions._--Diameter of the shell 0.12 to 0.14, of the pores 0.01 to
0.02, of the bars 0.003 to 0.006, of the larger apertures 0.03 to 0.04.

_Habitat._--Indian Ocean, near the Cocos Islands, Rabbe, surface.

Genus 35. _Choenicosphæra_,[57] n. gen.

_Definition._--#Collosphærida# with simple shells, armed on the outside
with radial spines, forming elegant coronals around the larger pores.

The genus _Choenicosphæra_ is characterised by the peculiar disposition of
its radial spines, which form protective coronals around the larger pores,
or even around all pores of the shell.

Subgenus 1. _Choenicosphærula_.

_Definition._--A coronal of spines around every pore of the shell.

1. _Choenicosphæra murrayana_, n. sp. (Pl. 8, fig. 4).

Shell spherical, with large circular or roundish pores of unequal size, two
to four times as broad as the bars. Ten to twelve pores in the half
meridian of the shell. Margin of every pore {103}with a coronal of six to
nine short and acute spines, not longer than the half diameter of the pore.
No spines between the pores.

_Dimensions._--Diameter of the shell 0.16 to 0.2, of the pores 0.02 to
0.03; length of the spines 0.008 to 0.012.

_Habitat._--Færöe Channel (Gulf Stream), common. Expedition of H.M.S.
"Triton," John Murray.

2. _Choenicosphæra flosculenta_, n. sp.

Shell spherical, with large circular or roundish pores of different size,
three to six times as broad as the bars. Six to eight pores in the half
meridian of the shell. Margin of every pore somewhat prominent, with a
coronal of ten to twenty parallel acute spines of different length, the
largest somewhat longer than the diameter of the pore. No spines between
the pores.

_Dimensions._--Diameter of the shell 0.12 to 0.15, of the pores 0.02 to
0.04; length of the spines 0.02 to 0.05.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

Subgenus 2. _Choenicosphærium_.

_Definition._--A coronal of spines only around the larger pores, not around
the smaller.

3. _Choenicosphæra nassiterna_, n. sp. (Pl. 8, fig. 3).

Shell spherical, with circular or roundish pores of very different size.
The smaller pores very numerous, without coronal of spines, roundish, about
as broad as the bridges. Twenty to thirty pores in the half meridian of the
shell. Between them, irregularly scattered, a small number (eight to
twelve) of very large circular pores, one-third to one-half as broad as the
radius of the shell, armed with a coronal of six to nine parallel,
straight, acute spines, about half as long as the radius of the shell.

_Dimensions._--Diameter of the shell 0.1 to 0.13, of the smaller pores
0.002 to 0.004, of the larger armed pores 0.02 to 0.03; length of the
spines 0.03 to 0.04.

_Habitat._--Philippine Islands, Mindanao, Station 213, depth 2050 fathoms.

4. _Choenicosphæra flammabunda_, n. sp. (Pl. 8, fig. 5).

Shell spherical, with circular or roundish pores of very different size.
The smaller pores very numerous and unequal, very irregularly scattered,
hardly one-fourth to one-half as broad as the bridges between them. Twelve
to twenty-four pores in the half meridian of the shell. Between them,
irregularly scattered, a variable number (ten to twenty) of very large
circular pores, about one-fourth as broad as the radius of the shell. The
margin of these large pores is armed with a very irregular coronal of four
to twelve unequal, curved acute spines, partly simple, partly branched like
{104}a deer-horn, one-fourth to one-half as long as the radius of the
shell. Some other small spines irregularly scattered over the shell.

_Dimensions._--Diameter of the shell 0.12 to 0.16, of the smaller pores
0.001 to 0.004, of the larger armed pores 0.016 to 0.024; length of the
spines 0.02 to 0.04.

_Habitat._--Central area of the Tropical Pacific, Stations 266 to 272,
depth 2425 to 2925 fathoms.

Genus 36. _Siphonosphæra_,[58] J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 59.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external simple radial tubuli with solid wall; outer mouth
of the tubuli truncated, smooth.

The genus _Siphonosphæra_ is, next to _Collosphæra_, the most common of all
Collosphærida, and rich in different species; all agreeing in the tubular
prolongation of the pores, and corresponding therefore to _Ethmosphæra_
among the simple Liosphærida.

Subgenus 1. _Holosiphonia_, Haeckel.

_Definition._--All the pores or apertures of the shell prolonged into
tubules.

1. _Siphonosphæra pansiphonia_, n. sp.

Shell a regular sphere, everywhere occupied by short, regular cylindrical
tubes, all of the same size and form. Length of the tubules equal to their
breadth and to the intervals between them. Ten to twelve tubules in the
half meridian of the shell. This most regular species is nearly allied to
_Collosphæra regularis_, and may be derived from it by tubular prolongation
of all the regular pores.

_Dimensions._--Diameter of the shell 0.1 to 0.12, length and breadth of the
tubules 0.005 to 0.006.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe, surface.

2. _Siphonosphæra marginata_, n. sp.

Shell a regular or subregular sphere, occupied by numerous short
cylindrical tubules of different sizes. Six to eight tubules in the half
meridian of the shell. Diameter of the tubules about equal to their
distance apart, but two to four times as large as their length.

_Dimensions._--Diameter of the shell 0.1 to 14; length of the tubules 0.004
to 0.006, breadth of the tubules and the intervals 0.01 to 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

{105}3. _Siphonosphæra tubulosa_, J. Müller (Pl. 6, fig. 4).

_Siphonosphæra tubulosa_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 59.

_Siphonosphæra tubulosa_, Haeckel, 1862, Monogr. d. Radiol., p. 532.

_Collosphæra tubulosa_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 59.

_Thalassicolla punctata_, var., Huxley, 1851, Ann. and Mag. Nat. Hist.,
ser. 2, vol. viii. p. 435, pl. xvi. fig. 5.

Shell subspherical or roundish, somewhat irregular, occupied by a small
number (five to ten) of short cylindrical tubules, irregularly scattered at
great distances; intervals between the tubules very large, two to four
times as broad as the length of the tubules, which is equal to their
breadth and about one-fifth or one-sixth of the shell diameter. Only two or
three tubules in the half meridian of the shell.

_Dimensions._--Diameter of the shell 0.12 to 0.15, length and breadth of
the tubules 0.02 to 0.03.

_Habitat._--Equatorial zone of the Pacific, Stations 225, 265, 268,
surface.

4. _Siphonosphæra cyathina_, n. sp. (Pl. 6, fig. 10).

Shell a regular sphere, everywhere occupied by short cup-like tubules of
somewhat different size. In the half meridian of the shell about ten to
twelve tubules, nearly cylindrical, but constricted in the middle. Inner
and outer aperture of the tubule of the same size; their diameter equal to
their length and distance.

_Dimensions._--Diameter of the shell 0.1 to 0.12, length and breadth of the
tubules 0.005 to 0.01.

_Habitat._--West coast of Tropical Africa, Stations 348 to 352, surface.

5. _Siphonosphæra patinaria_, n. sp. (Pl. 6, figs. 7, 8).

Shell a regular or subregular sphere, occupied by numerous broad cup-like
tubules of very different sizes. In the half meridian of the shell about
six to eight tubules, very flat, circular or subcircular, much constricted
in the middle. Diameter of the inner aperture larger than that of the
middle stricture, smaller than that of the outer aperture of the tubule;
outer diameter two to four times as great as their length.

_Dimensions._--Diameter of the shell 0.1 to 0.15, of the pores 0.01 to
0.04; length of the tubules 0.01, breadth 0.02 to 0.04.

_Habitat._--Tropical Atlantic, Station 348, depth 2450 fathoms.

6. _Siphonosphæra infundibulum_, n. sp.

Shell subspherical or irregular rounded, occupied by a small number (six to
twelve) of large funnel-like tubules of very different size, scattered
irregularly at great distances. Intervals between the tubules larger than
their length, which surpasses the radius of the shell. Outer opening of the
funnels three to four times as broad as the inner.

_Dimensions._--Diameter of the shell 0.15; length of the tubules 0.05 to
0.09.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

{106}7. _Siphonosphæra conifera_, n. sp. (Pl. 6, fig. 9).

Shell subspherical, everywhere occupied by irregular conical tubules, the
length of which nearly reaches the shell radius. Inner aperture of the
cones two to three times as large as the outer. Four to five tubules in the
half meridian of the shell. Distances between the bases of the cones small
and irregular.

_Dimensions._--Diameter of the shell 0.06 to 0.08, length of the tubules
0.03 to 0.04; inner aperture of the cones 0.01 to 0.02, outer aperture
0.008 to 0.01.

_Habitat._--Indian Ocean, Cocos Islands, Rabbe, surface.

8. _Siphonosphæra fragilis_, n. sp.

Shell quite irregular roundish or nearly ovate, very thin and fragile,
everywhere occupied by irregular, short, and broad cylindrical or conical
tubes. Six to ten tubes in the half meridian of the shell. Diameter of the
tubules about one-eighth that of the shell, three to four times as large as
the length of the tubules, and the distance apart of their bases.

_Dimensions._--Diameter of the shell 0.16 to 0.24, of the tubules 0.02 to
0.03; length of the tubules 0.006 to 0.008, distance of them 0.005 to
0.009.

_Habitat._--East coast of Australia, Sydney, Faber; Station 165, surface.

Subgenus 2. _Merosiphonia_, Haeckel.

_Definition._--Only part of the shell apertures prolonged in tubules, the
others simple.

9. _Siphonosphæra socialis_, n. sp. (Pl. 6, figs. 1, 2).

Shell a regular or subregular sphere, bearing only a small number (one to
four, commonly two to three) of short and broad cylindrical tubules,
irregularly scattered. Between them many small circular or subcircular
pores of different sizes, double as broad as their bars. Eight to ten pores
in the half meridian of the shell. Tubules three to six times as broad as
the pores, about as long as broad, now quite cylindrical, now somewhat
dilated at the outer opening. (Although the shells and cells of this common
species are among the smallest, their colonies are among the largest, often
containing more than one hundred social individuals, often enclosed in
alveoles.)

_Dimensions._--Diameter of the shell 0.04 to 0.05, of the pores 0.002 to
0.004, of the tubules 0.015 to 0.02; length of them about the same.

_Habitat._--Tropical and subtropical part of the Eastern Atlantic, Cape
Verde Islands, Canaries, very common, Haeckel; Stations 338 to 353,
surface.

10. _Siphonosphæra polysiphonia_, n. sp.

Shell a regular or subregular sphere, bearing twelve to sixteen circular
pores in its half meridian. Nearly one half the pores simple, very small;
the other half prolonged into short cylindrical tubules, {107}half as long
as broad, two to four times as broad as the pores and their intervals. This
species is nearly related to the foregoing, which it represents in the
western tropical part of the Atlantic, but differs constantly in the double
size of the shell and the much larger number of the tubules.

_Dimensions._--Diameter of the shell 0.06 to 0.09, of the pores 0.002 to
0.004, of the tubules 0.01 to 0.012; their length 0.006 to 0.008, surface.

_Habitat._--Tropical and subtropical part of the Western Atlantic, coast of
Brazil, &c., Rabbe.

11. _Siphonosphæra macrosiphonia_, n. sp.

Shell a regular sphere, with numerous very small pores of equal size and
distribution. Twelve to sixteen pores in its half meridian. Bars (between
the pores) three to four times as broad as their diameter. Only a small
number (two to four) of very long cylindrical tubes, irregularly scattered,
nearly as long or somewhat longer than the shell diameter; now quite
straight, now somewhat curved. Diameter of the cylinders four to six times
as large as that of the pores.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.002 to
0.004, of the tubules 0.015 to 0.018; length of the tubules 0.08 to 0.16.

_Habitat._--Central Tropical Pacific, Station 266 to 272, surface.

12. _Siphonosphæra serpula_, n. sp. (Pl. 6, fig. 6).

Shell a regular sphere, with numerous very small pores of equal size and
distribution. Eight to ten pores in its half meridian; bars between them
three to four times as broad as their diameter. Only a small number (six to
eight) of very long and snake-like, contorted, cylindrical tubes,
irregularly scattered. The shape of the latter is very much like that of
the calcareous tubes of some species of _Serpula_; they are nearly as long
as, or longer than, the shell diameter, and four to six times as broad as
the pores.

_Dimensions._--Diameter of the shell 0.12 to 0.14, of the pores 0.003 to
0.005, of the tubules 0.02 to 0.022; length of the tubules 0.1 to 0.2.

_Habitat._--North Eastern Pacific, Station 252 to 262, Sandwich Islands,
Haltermann.

13. _Siphonosphæra chonophora_, n. sp. (Pl. 6, fig. 5).

Shell a regular or subregular sphere, with numerous very small pores of
equal size and distribution, ten to twelve in its half meridian. Bars
between the pores four to six times their diameter. Only a small number
(two to six) of very large funnel-like tubules, irregularly scattered. The
inner half of these tubules is a short cylindrical tube, of the same
thickness as the shell, three to four times as broad as the pores; the
outer half is an irregular funnel, suddenly expanded, with siliceous walls
of the utmost tenuity and fragility, often irregularly folded and
contorted, like a decayed flower-calyx, often half as large as the shell.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.003 to
0.005, inner half of the tubules 0.015 to 0.02, outer funnel-like half 0.05
to 0.08.

_Habitat._--South Pacific, Stations 285 to 295, surface.

{108}14. _Siphonosphæra pipetta_, n. sp. (Pl. 6, fig. 3).

Shell more or less irregularly roundish or subspherical, occupied in part
by very small pores, in part by very large cylindrical tubules, inflated in
the middle. Number, form, and size of the tubes very irregular; commonly
there are five to ten, half as long or as long as the shell radius; their
inner and outer aperture about half as broad as their inflated middle part;
three to nine times as broad as the pores. A very irregular and variable
species.

_Dimensions._--Diameter of the shell 0.1 to 0.15, of the pores 0.001 to
0.005, of the tubules 0.015 to 0.03; length of the tubules 0.03 to 0.08.

_Habitat._--North Pacific, Stations 242 to 253, surface.

Genus 37. _Mazosphæra_,[59] Ehrenberg, 1860.

_Mazosphæra_, Ehrenberg, 1860, Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, p. 833.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external simple radial tubuli with solid wall; outer mouth
of each tubulus armed with a single tooth.

The genus _Mazosphæra_ is intermediate between _Siphonosphæra_ and
_Odontosphæra_, agreeing with the former in the tubular prolongation of the
pores, with the latter in the possession of a single large protective tooth
on the outer opening.

1. _Mazosphæra hippotis_, n. sp. (Pl. 5, fig. 8).

Shell spherical, with circular pores of irregular size and distribution,
scarcely half as broad as the bars; fifteen to twenty on the half meridian
of the shell. Between them, irregularly distributed, a variable number (ten
to fifteen) of short cylindrical radial tubules, about twice as long as
broad, and half as long as the shell radius. Mouth of the tubuli obliquely
truncated, having on one side a strong acute tooth.

_Dimensions._--Diameter of the shell 0.11 to 0.12, of the pores 0.002 to
0.004, of the bridges 0.006 to 0.009; length of the tubuli 0.03, breadth of
them 0.01 to 0.015.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

2. _Mazosphæra lagotis_, n. sp. (Pl. 5, fig. 9).

Shell spherical, with circular pores of irregular size and distribution,
about as broad as the bars; twelve to sixteen on the half meridian. Between
them, irregularly distributed, a variable number (eight to twelve) of long,
cylindrical, curved tubules, three to six times as long as broad, and about
as long as the shell radius; their external mouth lateral, obliquely
truncated, ovate, having on one side a strong conical tooth.

{109}_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.002
to 0.01; length of the tubuli 0.05 to 0.07.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

3. _Mazosphæra lævis_, Ehrenberg.

_Mazosphæra lævis_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 297, Taf. vii. fig. 7.

Shell spherical, with very small pores, scarcely one-fourth as broad as the
bars. Fifteen to twenty pores on the half meridian of the shell. Between
them, irregularly distributed, a variable number (fifteen to twenty) of
short conical tubules, about as long as broad, only one-fifth to one-sixth
as long as the shell radius. Mouth of the tubuli truncated, with an obtuse
short tooth on one side.

_Dimensions._--Diameter of the shell 0.08 to 0.09, of the pores 0.001 to
0.002, of the bridges 0.005 to 0.008; length of the tubuli 0.01 to 0.02,
breadth of them the same.

_Habitat._--Philippine Islands (depth 3300 fathoms), Ehrenberg; Station
206, depth 2100 fathoms; Station 225, depth 4575 fathoms.

4. _Mazosphæra apicata_, Ehrenberg.

_Mazosphæra apicata_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 316.

Shell spherical, without small pores, only with a variable number (ten to
twenty) of short conical tubules, twice as long as broad, and half as long
as the shell radius. Mouths of the tubuli obliquely truncated, with a
strong acute tooth on one side. (This species differs from the two
preceding by the want of the small pores between the tubules.)

_Dimensions._--Diameter of the shell 0.08 to 0.1, of the tubules 0.01.

_Habitat._--Philippine Islands (depth 3300 fathoms), Ehrenberg; north coast
of New Guinea, depth 2000 fathoms; Station 217.

Genus 38. _Trypanosphæra_,[60] n. gen.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external simple radial tubuli with solid walls; outer mouth
of each tubulus armed with a coronal of spines.

The genus _Trypanosphæra_ is intermediate between _Siphonosphæra_ and
_Choenicosphæra_ agreeing with the former in the tubular prolongation of
the pores, with the latter in the possession of a coronal of teeth on their
outer opening.

Subgenus 1. _Trypanosphærula_, Haeckel.

_Definition._--All pores of the shell prolonged into short coronated
tubules.

{110}1. _Trypanosphæra trepanata_, n. sp. (Pl. 5, fig. 4).

Shell regular spherical, with regular circular pores of nearly equal size,
at unequal distances, one to four times as broad as the bars. Eight to ten
pores on the half meridian. All the pores prolonged into short cylindrical
tubuli about as long as broad, armed on the external mouth with an elegant
coronal of twenty to thirty straight bristle-shaped, parallel teeth.

_Dimensions._--Diameter of the shell 0.12 to 0.14, of the pores 0.015 to
0.02; length of the tubuli 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

2. _Trypanosphæra dentata_, n. sp.

Shell regular spherical, with regular circular pores of equal size, but at
very different distances. Only three to four pores on the half meridian.
All the pores prolonged into irregular curved, cylindrical tubuli, about as
long as the shell radius, with a coronal of ten to twelve short conical
teeth on the distal end.

_Dimensions._--Diameter of the shell 0.08 to 0.09; length of the tubuli
0.04, breadth 0.02.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

Subgenus 2. _Trypanosphærium_, Haeckel.

_Definition._--Only part of the shell-pores prolonged into coronated
tubules.

3. _Trypanosphæra coronata_, n. sp. (Pl. 5, fig. 3).

Shell regular spherical, with irregular roundish pores of very different
sizes. On the half meridian four to six large and twelve to sixteen very
small pores. About half of the large pores prolonged into short cylindrical
tubuli, the outer mouth of each being armed with an elegant coronal of ten
to twenty thin irregular teeth.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the large pores 0.01
to 0.02, of the small pores 0.001; length of the tubuli 0.012.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

4. _Trypanosphæra terebrata_, n. sp.

Shell irregular roundish, with unequal, small, roundish pores. Sixteen to
twenty pores on the half meridian. Six to eight larger pores are prolonged
into curved cylindrical tubuli, about as long as the shell radius, with a
coronal of ten to twelve strong conical straight teeth on the distal end.

_Dimensions._--Diameter of the shell 0.15, of the pores 0.001 to 0.003;
length of the tubuli 0.08, breadth 0.02.

_Habitat._--West Tropical Pacific, Station 225, depth 4575 fathoms.

{111}5. _Trypanosphæra transformata_, n. sp. (Pl. 5, figs. 1, 2).

Shell quite irregular, of very variable, roundish, or polyhedral form, with
small irregular roundish pores, two to four times as broad as the bars. Ten
to thirty on the half meridian. The different form of the shell depends
upon the variable number of tubuli, which arise at irregular distances from
the shell; commonly three to four, often also five to six, more rarely one
or two. The tubuli are now more conical, now more cylindrical, about as
long as the shell radius, at other times scarcely one-half or one-third as
long, with a coronal of ten to twenty more or less curved teeth on the
narrower distal mouth. All the different forms are to be found in one and
the same colony, as shown in fig. 1. This coenobium, which I observed
living in Ceylon, exhibited the same peculiar formation as I figured in
_Collosphæra huxleyi_ in my Monograph 1862 (Taf. xxxiv. fig. 1). In the
centre of the jelly-sphere lies a large globular alveole, surrounded by
numerous small, young central capsules without shell; whilst in the surface
lies one layer of older capsules, enclosed in shells. Some of the younger
capsules exhibit self-division.

_Dimensions._--Diameter of the shells 0.08 to 0.12, pores 0.002 to 0.006;
length and breadth of the tubuli 0.03 to 0.05.

_Habitat._--Indian Ocean, Belligemma, Ceylon, surface.

Genus 39. _Caminosphæra_,[61] n. gen.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external branched radial tubuli with solid wall.

The genus _Caminosphæra_ differs from _Siphonosphæra_ (and from all other
Collosphærida) in the ramification of the tubuli, which arise from the
pores; the walls of the tubuli are solid, not fenestrated.

1. _Caminosphæra furcata_, n. sp.

Shell spherical or subspherical, with a variable number (four to eight) of
short cylindrical tubes, irregularly scattered, about as long as the radius
of the shell. Every tube forked, with two cylindrical branches of the same
size as the simple basal part of the tube. Mouth of the branches truncated,
not dilated. Pores of the shell between the tubes very small, all of the
same size, half as broad as their bars. Fifteen to twenty pores in the
half meridian of the shell.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.001 to
0.002; length of the tubules 0.05 to 0.06, breadth of them 0.012 to 0.015.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

2. _Caminosphæra elongata_, n. sp.

Shell spherical, with a large number (twelve to twenty) of long cylindrical
tubes, irregularly formed and scattered, somewhat longer than the diameter
of the shell. Every tube forked at {112}the distal end, with two or three
short irregular branches of unequal size and form; branches much shorter
than the undivided basal part of the tube. Mouth of the branches narrowed,
truncated. Pores of the shell between the tubes about half as broad,
irregularly roundish or polygonal, two to three times as broad as their
bars. Ten to twelve pores in the half meridian of the shell.

_Dimensions._--Diameter of the shell 0.05 to 0.07, of the pores 0.006 to
0.009; length of the tubules 0.06 to 0.09, breadth of them 0.015 to 0.02.

_Habitat._--Tropical Central Pacific, Station 271, depth 2425 fathoms.

3. _Caminosphæra dichotoma_, n. sp. (Pl. 7, fig. 2).

Shell spherical, with a variable number (ten to fifteen) of cylindrical
tubes, irregularly scattered, about as long as the radius of the shell.
Every tube furcated, with two cylindrical branches of the same size as the
simple basal part of the tube. Mouth of the branches dilated, funnel-like,
twice as broad as the tube; the edges irregularly dentated or lacerated.
Pores between the tubes small, one-third to one-sixth as broad as these,
half as broad as their bars. Ten to twelve pores in the half meridian of
the shell.

_Dimensions._--Diameter of the shell 0.12 to 0.14, of the pores 0.003 to
0.005; length of the tubules 0.06 to 0.08, breadth of them 0.02 to 0.03.

_Habitat._--Southern Pacific, Station 295, depth 1500 fathoms.

4. _Caminosphæra dendrophora_, n. sp. (Pl. 7, fig. 1).

Shell spherical, with a variable number (eight to twelve) of long
cylindrical tubes, irregularly branched and scattered, nearly as long as
the diameter of the shell. Every tube with two to six (commonly three to
four) branches of different sizes. Mouth of the branches dilated,
funnel-like; the edges irregularly dentated or lacerated. Pores between the
tubes half as broad as these, irregularly roundish, twice as broad as their
bars. Ten to twelve pores in the half meridian of the shell.

_Dimensions._--Diameter of the shell 0.12 to 0.15, of the pores 0.006 to
0.01; length of the tubules 0.1 to 0.13, breadth of them 0.02 to 0.025.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

Genus 40. _Solenosphæra_,[62] n. gen.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external simple radial tubuli with fenestrated wall; outer
mouth of the tubuli truncated, smooth.

The genus _Solenosphæra_ differs from _Siphonosphæra_ in the fenestration
of the external radial tubes. A large number of shells, appertaining to
this genus, were already described by Ehrenberg, and disposed in five
different genera corresponding to the {113}different numbers of the
tubuli:--_Disolenia_ with two tubes, _Trisolenia_ with three tubes,
_Tetrasolenia_ with four tubes, _Pentasolenia_ with five tubes,
_Polysolenia_ with six or more tubes. All these five genera are without
value, as those different numbers of tubes occur frequently intermingled in
the individual cells of one and the same colony, wherever the form and
structure of the tubes is inherited with sufficient constancy to determine
the species.

Subgenus 1. _Solenosphactra_, Haeckel.

_Definition._--Tubuli of the shell cylindrical or nearly cylindrical, the
outer and inner apertures nearly of the same size.

1. _Solenosphæra variabilis_, Haeckel.

_Tetrasolenia quadrata_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, Taf. x. fig. 20.

Shell quite irregular roundish or polyhedral, with roundish pores of
different size. Ten to fifteen pores in the half meridian of the shell, two
to three times as broad as the bars. Porous tubuli of the shell in variable
number (in one and the same colony), three to nine, mostly four to six;
cylindrical or subcylindrical or somewhat conical, two to three times as
broad as long, not longer than the half radius of the shell. Inner aperture
of the tubuli commonly as broad as the half radius of the shell (or
somewhat smaller), about as large as the truncated outer aperture. This
species is closely related to _Collosphæra polyedra_ (p. 97), and may be
derived from it by a short tube-like prolongation of the larger apertures.

_Dimensions._--Diameter of the shell 0.1 to 0.16, of the pores 0.005 to
0.015; length of the tubules 0.02 to 0.03, breadth of them 0.04 to 0.06.

_Habitat._--Central area of the Tropical Pacific, Stations 270, 271, 272,
depths 2425 to 2925 fathoms.

2. _Solenosphæra pandora_, n. sp. (Pl. 7, figs. 10, 11).

Shell irregular roundish or subglobular, with roundish pores of different
sizes, mostly somewhat broader than the bars. About twelve to sixteen pores
on the half meridian of the shell. Porous tubuli of the shell of variable
number (in one and the same colony), one to six, mostly three to four;
cylindrical or nearly cylindrical, somewhat longer than broad, not longer
than the radius of the shell. Inner aperture of the tubuli commonly as
broad as the half radius of the shell, and a little smaller than the
truncated outer aperture.

_Dimensions._--Diameter of the shell 0.07 to 0.1, of the pores 0.003 to
0.006; length of the tubuli 0.03 to 0.05, breadth of them 0.02 to 0.03.

_Habitat._--Central area of the Tropical Pacific, Stations 266 to 274,
depths 2350 to 2925 fathoms.

{114}3. _Solenosphæra megalactis_, Haeckel.

_Trisolenia megalactis_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 301, Taf. viii. fig. 19.

Shell irregularly polyhedrical, with very small roundish pores, scarcely
half as broad as the bars. Only eight to ten pores on the half meridian of
the shell. Porous tubuli of the shell of variable number (in one and the
same colony), two to five, mostly three or four; cylindrical, about as long
as the radius of the shell. Inner aperture of the tubuli commonly as broad
as the half radius of the shell, and quite as broad as the truncated outer
aperture.

_Dimensions._--Diameter of the shell 0.07 to 0.09, of the pores 0.002 to
0.004, of the bars 0.005 to 0.009; length of the tubuli 0.03 to 0.04,
breadth of them 0.02.

_Habitat._--Pacific; California, Philippine Sea, Ehrenberg; Stations 256 to
285, depths 310 to 3000 fathoms.

4. _Solenosphæra serpentina_, n. sp. (Pl. 7, fig. 7).

Shell nearly spherical, with very small circular pores, scarcely one-third
or one-fourth as broad as the bars. Only five to seven pores in the half
meridian of the shell. Porous tubuli of the shell of variable number (in
one and the same colony), two to nine, mostly seven or eight; cylindrical,
somewhat curved or contorted, once and a half or twice as long as the
diameter of the shell, with few very small and widely scattered pores.
Inner and outer aperture of the tubuli have the same diameter, about
one-fifth or one-fourth that of the shell. (This species is closely allied
to _Siphonosphæra serpula_, but is distinguished from it by the long
tortuous tubuli and the small scarce pores.)

_Dimensions._--Diameter of the shell 0.08 to 0.1, of the pores 0.001 to
0.002, of the bars 0.004 to 0.008; length of the tubuli 0.12 to 0.18,
breadth of them 0.02 to 0.025.

_Habitat._--North-eastern Pacific, between Sandwich Islands and California,
Haltermann, surface.

Subgenus 2. _Solenosphenia_, Haeckel.

_Definition._--Tubuli of the shell more or less conical, the inner aperture
much larger than the outer aperture.

5. _Solenosphæra venosa_, Haeckel.

_Tetrasolenia venosa_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 301, Taf. vii. fig. 22.

Shell irregular polyhedral or roundish, with a delicate network of large
irregular polyhedral meshes, five to ten times as broad as the thin bars.
Eight to twelve meshes on the half meridian of the shell. Fenestrated
tubuli of the shell of variable number (in one and the same colony), one to
five, commonly three or four, shaped like a short truncated cone, about
half as long as broad on its base, shorter than the radius of the shell.
Inner aperture of the cone nearly as broad as the half radius of the shell,
about twice as broad as the truncated outer aperture.

{115}_Dimensions._--Diameter of the shell 0.07 to 0.12, of the pores 0.008
to 0.016, of the bars 0.001; length of the tubuli 0.02 to 0.03, inner
aperture 0.03 to 0.04, outer aperture 0.02 to 0.03.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe.

6. _Solenosphæra ascensionis_, n. sp. (Pl. 7, fig. 9).

Shell somewhat irregular, subspherical, with polygonal pores of different
size. Twelve to fifteen pores in the half meridian of the shell, two to
eight times as broad as their bars. Porous tubuli of the shell of variable
number (in one and the same colony), three to nine, mostly five to seven;
conical or nearly cylindrical, irregular, about as long as broad at their
base. Inner aperture of the tubuli two to four times as broad as the
broadest pores, and double as broad as the truncated circular outer
aperture.

_Dimensions._--Diameter of the shell 0.1 to 0.12, of the pores 0.004 to
0.018, of the bars 0.002, length of the tubuli 0.04, inner aperture 0.04,
outer 0.02.

_Habitat._--South Atlantic, near Ascension Island, Station 343, surface.

Subgenus 3. _Solenosphyra_, Haeckel.

_Definition._--Tubuli of the shell funnel-like, the outer aperture much
larger than the inner.

7. _Solenosphæra cornucopia_, n. sp. (Pl. 7, fig. 8).

Shell spherical or subspherical, with roundish pores of different size. Ten
to twelve pores in the half meridian of the shell, two to three times as
broad as the bars. Porous tubuli of the shell of variable number (in one
and the same colony), four to eight, mostly five to seven, funnel-like,
about as long as the diameter of the shell. Inner aperture of the tubuli
commonly two-thirds or three-fourths as broad as the radius of the shell
(or somewhat smaller), only one-half or two-thirds as broad as the dilated
and truncated outer aperture.

_Dimensions._--Diameter of the shell 0.07 to 0.09, of the pores 0.006 to
0.018; length of the tubuli 0.06 to 0.08, diameter of the inner aperture
0.04 to 0.05, of the outer 0.06 to 0.08.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

8. _Solenosphæra amalthea_, n. sp.

Shell irregular roundish or spherical, with small circular pores of
different size. Fifteen to twenty pores in the half meridian of the shell,
but still not as broad as the bars. Porous tubuli of the shell of variable
number (in one and the same colony), three to six, commonly four or five,
funnel-like, about as long as the radius of the shell. Inner aperture of
the tubuli about half as broad as the radius of the shell, only one-half or
one-third as broad as the truncated outer aperture. (This species is
intermediate between the preceding and _Siphonosphæra chonophora_, Pl. 6,
fig. 5.)

{116}_Dimensions._--Diameter of the shell 0.09 to 0.11, of the pores 0.002
to 0.004, of the bars 0.003 to 0.006; length of the tubuli 0.05 to 0.06;
diameter of the inner aperture 0.02 to 0.03, of the outer aperture 0.05 to
0.07.

_Habitat._--Western part of the South Atlantic, Station 325, surface.

Genus 41. _Otosphæra_,[63] n. gen.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external simple radial tubuli with fenestrated walls; outer
mouth of the tubuli armed with a single tooth.

The genus _Otosphæra_ differs from _Solenosphæra_ by the single tooth on
the external mouth of the tubuli, from _Mazosphæra_ by the fenestration of
the walls of the tubuli.

1. _Otosphæra polymorpha_, n. sp. (Pl. 7, fig. 6).

Shell quite irregular, polyhedral or roundish, very variable in size and
form, with numerous very small pores, much smaller than the bars. Twenty to
thirty pores in the half meridian of the shell. Porous tubuli of the shell
commonly in variable number (one to four), but sometimes constant in number
(one, two, three, or four) in the one and same colony. Tubuli irregular
conical, commonly about as long as the radius of the shell; their outer
aperture obliquely truncated, on one side prolonged into one large,
prominent, bill-like, curved, acute tooth.

_Dimensions._--Diameter of the shell 0.12 to 0.15, of the pores 0.001 to
0.002; length of the tubuli 0.06 to 0.08, inner aperture 0.03, outer
aperture 0.02.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

2. _Otosphæra auriculata_, n. sp. (Pl. 7, fig. 5).

Shell quite irregular, of extremely variable form, now inclining to
roundish, now to polyhedral, with very numerous small pores, irregularly
formed and distributed. Twelve to twenty-four pores in the half meridian of
the shell, of very different size, for the most part larger than the bars.
Porous tubuli of the shell of variable number (in one and the same colony),
one to five, mostly three or four, of conical form, irregularly formed and
scattered, commonly about half as long as the radius of the shell. Outer
aperture of the tubuli obliquely truncated, with one large prominent, often
curved, acute tooth.

_Dimensions._--Diameter of the shell 0.1 to 0.2, of the pores 0.003 to
0.005; length of the tubuli 0.01 to 0.05, inner aperture 0.04, outer
aperture 0.03.

_Habitat._--Central Tropical Pacific, Stations 268 to 272, depths 2425 to
2925 fathoms.

{117}Genus 42. _Coronosphæra_,[64] n. gen.

_Definition._--#Collosphærida# with simple shells, the pores of which are
prolonged into external simple radial tubuli with fenestrated walls; outer
mouth of the tubuli armed with a coronal of spines.

The genus _Coronosphæra_ differs from _Solenosphæra_ by the coronated mouth
of the tubuli, from _Trypanosphæra_ by the fenestration of the walls of the
tubuli.

1. _Coronosphæra diadema_, n. sp. (Pl. 7, fig. 3).

Shell spherical or subspherical, with a variable number (fifteen to twenty)
of short, coronal-like tubules, irregularly scattered, about half as long
as the radius of the shell. Outer aperture of the tubuli irregularly
dentated, a little dilated, and not much broader than the inner aperture,
one-half or one-third as broad as the shell radius. Pores of the shell and
of the tubuli circular or roundish, very irregularly scattered, mostly
one-half or one-third as broad as the bars.

_Dimensions._--Diameter of the shell 0.11 to 14, of the pores 0.002 to
0.004, of the bars 0.006 to 0.012; length of the tubuli 0.03, inner
aperture 0.02 to 0.03, outer aperture 0.03 to 0.04.

_Habitat._--Central Pacific, Stations 268 to 270, depths 2550 to 2925
fathoms.

2. _Coronosphæra calycina_, n. sp. (Pl. 7, fig. 4).

Shell spherical or subspherical, with a variable number (eight to twelve)
of large, funnel-like tubules, irregularly scattered, about as long as the
radius of the shell. Outer aperture of the tubuli irregularly dentated,
much dilated, somewhat broader than the shell radius, three to four times
as broad as the inner circular aperture. Pores of the shell and of the
tubuli circular or roundish, of very different size, one to three times as
broad as the bars.

_Dimensions._--Diameter of the shell 0.1 to 0.3, of the pores 0.003 to
0.01, of the bars 0.002 to 0.004; length of the tubuli 0.1, inner aperture
0.02 to 0.03, outer aperture 0.06 to 0.18.

_Habitat._--Central Pacific, Stations 271, 272, depths 2425 and 2600
fathoms respectively.

3. _Coronosphæra convolvulus_, n. sp.

Shell irregular roundish, with a variable number (five to ten) of long,
curved tubules, about as long as the shell diameter. The inner half of the
tubuli is narrow, cylindrical; the outer half funnel-like dilated, similar
to the flower of _Convolvulus_. The outer aperture is elegantly dentated,
five to six times as broad as the inner aperture. Pores of the shell and of
the tubuli very irregular roundish, about as broad as the bars.

_Dimensions._--Diameter of the shell 0.08 to 0.09, of the pores and bars
0.004 to 0.008; length of the tubuli 0.07 to 0.1, inner aperture 0.01,
outer aperture 0.05.

_Habitat._--Tropical Atlantic, Station 347, surface.

{118}Subfamily CLATHROSPHÆRIDA, Haeckel, 1881, Prodromus, p. 472.

_Definition._--#Collosphærida# with a double lattice-shell around every
central capsule of the coenobium; both concentric shells connected by
irregular or subradial beams, commonly solid or lamellar staffs, rarely
hollow tubes.

Genus 43. _Clathrosphæra_,[65] Haeckel, 1881, Prodromus, p. 472.

_Definition._--#Collosphærida# with a double lattice-shell around every
central capsule of the coenobium; surface of the outer shell smooth.

The genus _Clathrosphæra_ (with smooth surface) and the following
_Xanthiosphæra_ (with spiny surface) form together the small subfamily,
Clathrosphærida, different from the other Collosphærida by the double
lattice-shell. From the surface of the inner primary shell arise either
solid spines or hollow tubes, which unite by the anastomosis of irregular
branches and so form the outer secondary shell, often very incomplete and
irregular. All Clathrosphærida seem to inhabit great depths.

Subgenus 1. _Clathrosphærula_, Haeckel.

_Definition._--The connecting staffs between both shells are hollow tubes
(derived from _Siphonosphæra_).

1. _Clathrosphæra circumtexta_, n. sp. (Pl. 8, fig. 6).

Inner shell spherical, with irregular roundish large meshes, now broader
now smaller than their bars. Eight to ten meshes in the half meridian of
the shell. All these meshes are prolonged into short cylindrical hollow
tubes, about as long as broad, somewhat constricted in the middle. From the
margins of the outer openings of these tubes proceed very numerous and
delicate siliceous filaments, which all lie on the same spherical face,
branch, anastomose, and twine over the openings and the intervals between
them, forming a very thin, arachnoid spherical outer shell. The meshes of
this are quite irregular polygonal, of very different size and form. The
radius of the inner shell bears to that of the outer a ratio = 5 : 6.

_Dimensions._--Diameter of the inner shell 0.11 to 0.13, of the outer 0.13
to 0.16; meshes of the inner shell 0.005 to 0.02, of the outer 0.005 to
0.04.

_Habitat._--North Pacific, Stations 238 to 253, depths 2050 to 3950
fathoms.

Subgenus 2. _Clathrosphærium_, Haeckel.

_Definition._--The connecting staffs between the two shells are solid rods
or lamellar spines (derived from _Acrosphæra_).

{119}2. _Clathrosphæra arachnoides_, n. sp. (Pl. 8, fig. 7).

Inner shell spherical, with irregular roundish meshes, two or three times
as broad as the bars. Ten to twelve meshes in the half meridian of the
shell. From its surface arise numerous conical radial spines (with base
often fenestrated), which at equal distances from the surface send out
lateral branches. All these branches lie on a spherical face, and form by
communications the irregular, very delicate, arachnoid network of the outer
shell, quite unlike that of the inner, with large polygonal meshes of very
different size. Eight to sixteen meshes in the half meridian of the shell.
Surface of the outer shell nearly spherical, somewhat uneven, like a
spider's web. The radius of the inner shell bears to that of the outer a
ratio = 3 : 4.

_Dimensions._--Diameter of the inner shell 0.12 to 0.14, of the outer 0.15
to 0.18; pores of the inner shell 0.003 to 0.02, of the outer 0.01 to 0.04.

_Habitat._--Central area of the Tropical Pacific, Station 268, depth 2900
fathoms.

3. _Clathrosphæra lamellosa_, n. sp. (Pl. 8, fig. 8).

Inner shell spherical or subspherical, with irregular roundish meshes,
about half as broad as the bars. Twelve to sixteen meshes in the half
meridian of the shell. From its surface arise numerous oblique irregular
staffs or broad and thin lamellæ, which branch quite irregularly, and by
communications of the branches form the thin outer shell. This is quite
irregular roundish or subspherical, very unlike the inner, with large
polygonal meshes of different size, six to twelve in the half meridian of
the shell. Bridges between the meshes very variable, now very thin
filamentous, now very broad lamellar. Outer surface very uneven or
tuberculated, but not spinous. The radius of the inner shell bears to that
of the outer a ratio = 5 : 6.

_Dimensions._--Diameter of the inner shell 0.1 to 0.13, of the outer 0.12
to 0.18; pores of the inner shell 0.003 to 0.009, of the outer 0.01 to
0.04.

_Habitat._--Central area of the Tropical Pacific, Stations 270 to 274,
depths 2350 to 2925 fathoms.

Genus 44. _Xanthiosphæra_,[66] Haeckel, 1881, Prodromus, p. 472.

_Definition._--#Collosphærida# with a double lattice-shell around every
central capsule of the coenobium; surface of the outer shell thorny or
spiny.

The genus _Xanthiosphæra_ differs from the foregoing _Clathrosphæra_ by
spines or thorns arising from the surface of the outer shell, commonly very
irregular.

1. _Xanthiosphæra capillacea_, n. sp.

Inner shell spherical, with irregular polygonal meshes, three to five times
as broad as their narrow bars. Six to eight meshes in the half meridian of
the shell. From its surface arise at the nodes of the network numerous thin
radial spines, which, at equal distances from the surface, {120}send out
lateral branches. All these branches lie on a spherical face, and form by
communications the irregular delicate network of the outer shell, very like
that of the inner, with large polygonal meshes, six to eight meshes in the
half meridian of the shell. Surface of the outer shell covered with
numerous straight spines, prolongations of the inner spines, but only half
as long as these. The radius of the inner shell bears to that of the outer
a ratio = 3 : 5.

_Dimensions._--Diameter of the inner shell 0.1 to 0.12, of the outer 0.15
to 0.19; pores of the inner shell 0.02 to 0.04 to 0.06, of the outer 0.04
to 0.06 to 0.08; length of the outer spines 0.01 to 0.02.

_Habitat._--Central area of the Tropical Pacific, Station 263, depth 2650
fathoms.

2. _Xanthiosphæra erinacea_, n. sp. (Pl. 8, fig. 9).

Inner shell spherical, with irregular roundish meshes, one-half to two
times as broad as the bars. Fifteen to twenty meshes in the half meridian
of the shell. From its surface arise numerous thin radial spines, which at
equal distances from the surface send out lateral branches. All these
branches lie on the face of a sphere, and form by communications the
irregular delicate network of the outer shell, very unlike that of the
inner, with large polygonal meshes, twelve to twenty-four in the half
meridian of the shell. Surface of the outer shell covered with numerous
straight spines, prolongations of the inner spines, and of the same length.
The radius of the inner shell bears to that of the outer a ratio = 3 : 4.

_Dimensions._--Diameter of the inner shell 0.1 to 0.12, of the outer 0.13
to 0.16; pores of the inner shell 0.002 to 0.008, of the outer 0.01 to
0.03; length of the outer spines 0.02 to 0.03.

_Habitat._--Central area of the Tropical Pacific, Stations 270, 272, depth
2925 and 2600 fathoms respectively.

3. _Xanthiosphæra lappacea_, n. sp. (Pl. 8, figs. 10, 11).

Inner shell spherical or subspherical, with very small roundish pores,
quite irregularly scattered, one-fourth to three-fourth as broad as their
bars. Ten to twenty pores in the half meridian of the shell. From its
surface arise in an extremely irregular and variable manner numerous
oblique spines, often curved, often lamellar, and perforated by pores,
sometimes hollow, fenestrated cones. At different distances from the
surface these spines send out lateral curved branches, which by
communications form the delicate and very irregular network of the outer
shell. This network is often incomplete and very unlike that of the inner
shell, with large polygonal meshes, six to eighteen in the half meridian of
the shell. Surface of the outer shell covered with numerous small, curved,
and oblique spines, prolongations of the inner spines, but scarcely
one-third to one-half as long as these. The radius of the inner shell bears
to that of the outer a ratio = 3 : 4.

_Dimensions._--Diameter of the inner shell 0.08 to 0.12, of the outer 0.11
to 0.15; pores of the inner shell 0.001 to 0.009, of the outer 0.01 to
0.04; length of the outer spines 0.005 to 0.009.

_Habitat._--Central area of the Tropical Pacific, Stations 263 to 274,
depths 2350 to 3000 fathoms.

{121}Family VII. #STYLOSPHÆRIDA#, Haeckel (Pls. 13-17).

_Stylosphærida_, Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Sphæroidea# with two radial spines on the surface of the
spherical shell, opposite in one axis; living solitary (not associated in
colonies).

The family #Stylosphærida# comprises a large number of very common
#Sphæroidea#, and is distinguished from all others by the possession of two
radial spines which are placed in one axis of the spherical shell.[67] By
the expression of this "main axis" as a solid rod they form the transition
to the #Prunoidea#, in which the whole shell is more or less transformed
according to this "monaxial growth." But in these latter the shell, as well
as the central capsule, becomes ellipsoidal, prolonged in one axis, whilst
in the former they remain spherical. However, the distinction of both
nearly allied groups is sometimes difficult.

The most simple Stylosphærida are the _Xiphostylida_, with one single
spherical lattice-shell. To this ancestral group all other subfamilies can
be opposed as "Stylosphærida concentrica," as their carapace is composed of
two or more concentric lattice-shells: two in the Sphærostylida, three in
the Amphistylida, four in the Cromyostylida, five or more in the
Caryostylida. In all these four subfamilies the concentric shells are
simple (not spongy) fenestrated spheres. In a sixth subfamily, in the
Spongostylida, the shell is wholly or partially composed of a spongy
irregular wicker-work, with or without a medullary shell in the centre.

Both the radial spines in all Stylosphærida are opposed normally in one
axis; but in many species besides the normal form occur individual
abnormalities, in which the two spines are not accurately opposed in this
main axis, but placed in two different axes, intersecting at a smaller or
larger angle. In the majority of the Stylosphærida both opposite spines
have the same size and form; but in some genera they are more or less
different, often in a very striking degree. The same differences occur in
the nearly allied groups of #Prunoidea#, in the Ellipsida and Druppulida.

The distal ends of both spines are commonly free; but in the small group of
Saturnalida (_Saturnalis_ with one single shell, _Saturnulus_ with two
concentric shells, _Saturninus_ with three concentric shells) the distal
ends of both spines are united, at equal distances from the centre, by a
circular or elliptical ring. This remarkable peculiarity occurs in no other
group of #Sphæroidea#, and consequently brings the Saturnalida into close
relation with the #Discoidea#.

{122}_Synopsis of the Genera of Stylosphærida._

{ {Both spines
{Polar spines { equal, 45. _Xiphosphæra_.
{ free, without{
I. Subfamily { connecting {Spines
Xiphostylida. { ring on the { different
(Spherical shell{ distal ends. { in size
simple.) { { or form, 46. _Xiphostylus_.
{
{Both polar spines united by
{ a circular or elliptical
{ ring, 47. _Saturnalis_.

{ {Both spines
{ { equal, 48. _Stylosphæra_.
{Polar spines {
II. Subfamily { free. {Spines
Sphærostylida.{ { different in
(Two concentric { { size or
spheres.) { { form, 49. _Sphærostylus_.
{
{Both polar spines united by
{ a circular or elliptical
{ ring, 50. _Saturnulus_.

{ {Both spines
{ { equal, 51. _Amphisphæra_.
{Polar spines {
III. Subfamily { free. {Spines
Amphistylida. { { different in
(Three { { size or
concentric { { form, 52. _Amphistylus_.
spheres.) {
{Both polar spines united by
{ a circular or elliptical
{ ring, 53. _Saturninus_.

IV. Subfamily } {Both spines
Cromyostylida.}Polar spines { equal, 54. _Stylocromyum_.
(Four concentric} free. {
spheres.) } {Spines
} { different, 55. _Cromyostylus_.

V. Subfamily }
Caryostylida. }Polar spines {Both spines
(Five or more } free. { equal, 56. _Caryostylus_.
concentric }
spheres.) }

{Shell a solid spongy sphere
{ without central medullary
{ shell, 57. _Spongolonchis_.
VI. Subfamily {
Spongostylida.{ {One central
(Spherical shell{ { medullary
partially or {In the centre { shell, 58. _Spongostylus_.
wholly of a { of the spongy{
spongy { sphere one or{Two
structure.) { two medullary{ concentric
{ shells. { medullary
{ { shells, 59. _Spongostylidium_.

Subfamily XIPHOSTYLIDA, Haeckel, 1881, Prodromus, pp. 449, 450.

_Definition._--#Stylosphærida# with one simple spherical lattice-shell.

Genus 45. _Xiphosphæra_,[68] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Stylosphærida# with one single lattice-sphere and two free
spines of equal size and form.

The genus _Xiphosphæra_ is the most simple form of all Stylosphærida, and
may be regarded as the common ancestral form of this family. On the surface
of a simple {123}spherical lattice-shell, enclosing the central capsule,
arise two equal, free, radial spines, opposite to each other on the poles
of one axis.

Subgenus 1. _Xiphosphærantha_, Haeckel.

_Definition._--Pores of the spherical shell regular, of nearly equal size
and form; surface smooth or a little rough, without spines or thorns (other
than the two polar spines).

1. _Xiphosphæra planeta_, n. sp.

Pores regular, hexagonal, eight to nine times as broad as the thin bars.
Ten to twelve pores on the half equator. Shell very thin walled; surface
smooth. Polar spines three-sided pyramidal, about as long as the axis of
the sphere, as broad at the base as one pore.

_Dimensions._--Diameter of the sphere 0.12 to 0.13, pores 0.016 to 0.018,
bars 0.002; length of the polar spines 0.1 to 0.15, basal thickness 0.02.

_Habitat._--Pacific, central area, surface; Stations 271 to 274, depths
2425 to 2750 fathoms.

2. _Xiphosphæra gæa_, n. sp. (Pl. 14, fig. 5).

Pores regular, circular, with prominent hexagonal crests between them. On
the half equator ten to twelve pores, of the same breadth as the crested
bars. Shell thin walled; surface smooth. Polar spines three-sided
prismatic, about twice as long as the axis of the sphere, twice as broad at
the base as one pore.

_Dimensions._--Diameter of the sphere 0.07 to 0.09, pores and bars 0.005;
length of the polar spines 0.15 to 0.2, basal thickness 0.01.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

3. _Xiphosphæra venus_, n. sp. (Pl. 14, fig. 2).

Pores regular, circular, with prominent hexagonal frames. On the half
equator fifteen to eighteen pores, of the same breadth as the bars. Shell
very thick walled; surface smooth, honeycomb-like. Polar spines conical,
smooth, about as long as the axis of the shell, twice as broad at the base
as one pore.

_Dimensions._--Diameter of the sphere 0.12 to 0.13, pores and bars 0.005;
thickness of the shell wall 0.013; length of the polar spines 0.12 to 0.15,
basal breadth 0.01.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

4. _Xiphosphæra luna_, n. sp.

Pores regular, circular, hexagonally lobed or rosette-shaped, three times
as broad as the bars. Ten to twelve pores on the half equator. Shell thick
walled; surface smooth. Polar spines three-sided pyramidal, one to two
times as long as the axis of the shell, as broad at the base as one pore
{124}(very similar to _Xiphostylus phasianus_, Pl. 13, fig. 9, but
different in the equal size and similar form of the two large polar
spines).

_Dimensions._--Diameter of the sphere 0.12, pores 0.015, bars 0.005; length
of the polar spines 0.1 to 0.2, basal breadth 0.02.

_Habitat._--Indian Ocean, Cocos Islands, surface, Rabbe.

5. _Xiphosphæra hebe_, n. sp.

Pores regular, circular, three times as broad as the bars. On the half
equator sixteen to twenty pores. Shell thick walled; surface smooth. Polar
spines conical or nearly cylindrical, about as long as the axis of the
sphere, as broad at the base as two pores.

_Dimensions._--Diameter of the sphere 0.1 to 0.13, pores 0.006, bars 0.002;
polar spines 0.1 to 0.15 long, 0.01 thick.

_Habitat._--Pacific, central area, Stations 265 to 268, depths 2700 to 2900
fathoms.

6. _Xiphosphæra maxima_, n. sp.

Pores regular, circular, twice as broad as the bars, funnel-shaped. Twenty
to thirty pores on the half equator. Shell very thick walled; surface
smooth. Polar spines three-sided pyramidal, about as long as the radius of
the sphere, as broad at the base as two pores.

_Dimensions._--Diameter of the sphere 0.22 to 0.35, pores 0.008 to 0.01,
bars 0.005; polar spines 0.1 to 0.15 long, 0.02 thick.

_Habitat._--Equatorial Atlantic, Station 347, depth 2250 fathoms.

7. _Xiphosphæra euphrosyne_, n. sp.

Pores regular, circular, about as broad as the bars, double contoured.
Eight to ten on the half equator. Shell thin walled; surface smooth. Polar
spines conical, about as long as the radius of the sphere, as broad at the
base as one pore.

_Dimensions._--Diameter of the sphere 0.12 to 0.15, pores and bars 0.02;
polar spines 0.06 to 0.09 long, 0.02 thick.

_Habitat._--South Atlantic, Station 323, depth 1900 fathoms.

Subgenus 2. _Xiphosphærella_, Haeckel.

_Definition._--Pores of the spherical shell regular, of nearly equal size
and form; surface thorny or spiny, covered with regularly distributed
papillæ or thorns (in addition to the two large polar spines).

8. _Xiphosphæra pallas_, n. sp. (Pl. 14, fig. 4).

Pores regular, circular, separated by hexagonal elevated frames, the sharp
crest of which is elegantly denticulated; in each corner of the hexagons
(between three pores) is a short radial spine, {125}about as long as one
pore. On the half equator sixteen to twenty pores, of the same breadth as
the bars. Shell thick walled; whole surface spiny. Polar spines
cylindrical, at the apex conical, about as long as the axis of the sphere,
three to four times as broad as one pore.

_Dimensions._--Diameter of the sphere 0.1, pores and bars 0.005; length of
the polar spines 0.07 to 0.11, thickness 0.015 to 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

9. _Xiphosphæra flora_, n. sp.

Pores regular, circular, with hexagonal frames, twice as broad as the bars.
Ten to twelve pores on the half equator. Shell thin walled, with spiny
surface; in each corner of the hexagons is one bristle-like radial spine
twice as long as one pore. Polar spines three-sided prismatic, at the apex
pyramidal, nearly twice as long as the axis of the sphere, as broad at the
base as two pores (similar to _Ellipsoxiphus palliatus_, Pl. 14, fig. 7).

_Dimensions._--Diameter of the sphere 0.15, pores 0.01, bars 0.005; length
of the polar spines 0.2 to 0.25, breadth 0.02.

_Habitat._--Tropical Atlantic, Station 342, depth 1445 fathoms.

10. _Xiphosphæra juno_, n. sp.

Pores regular, circular, as broad as the bars, funnel-shaped. Fifteen to
twenty pores on the half equator. Shell thick walled, covered with
bristle-like spines, about twice as long as one pore. Polar spines conical,
thick, about as long as the axis of the sphere, twice as broad at the base
as one pore.

_Dimensions._--Diameter of the sphere 0.12, pores and bars 0.01; length of
the polar spines 0.14, basal breadth 0.02.

_Habitat._--Fossil in the Barbados rocks; living in the greatest depth of
the Tropical Pacific, Station 225, depth 4475.

11. _Xiphosphæra gigantea_, n. sp.

Pores regular, circular, two to three times as broad as the bars;
twenty-eight to thirty-two on the half equator. Shell thick walled, covered
with short conical thorns. Polar spines three-sided pyramidal, about as
long as the radius of the sphere, as broad at the base as three pores.

_Dimensions._--Diameter of the sphere 0.25 to 0.3, pores 0.01, bars 0.004;
polar spines 0.1 to 0.15 long, 0.03 broad.

_Habitat._--Fossil in the Tertiary rocks of Barbados and Sicily
(Cattanisetta).

Subgenus 3. _Xiphosphærissa_, Haeckel.

_Definition._--Pores of the spherical shell irregular, of different size or
form; surface smooth or a little rough, without spines or thorns (other
than the polar spines).

{126}12. _Xiphosphæra ceres_, n. sp.

Pores irregular, roundish, of different sizes, two to four times as broad
as the bars. Sixteen to twenty pores on the half equator. Shell thin
walled, with smooth surface. Polar spines conical, about as long as the
axis of the sphere, very thick at the base.

_Dimensions._--Diameter of the sphere 0.15 to 0.2, pores 0.004 to 0.008,
bars 0.002; polar spines 0.18 to 0.24 long, at the base 0.02 thick.

_Habitat._--North Atlantic, Station 353, surface.

13. _Xiphosphæra clavigera_, n. sp.

Pores irregular, roundish, double contoured, of very unequal size, two to
seven times as broad as the bars; ten to twelve on the half equator. Shell
thick walled; surface a little rough. Polar spines club-shaped, with
prominent edges, about half as long as the axis of the sphere; thinner at
both ends than in the middle. (Differs from _Ellipsoxiphus claviger_, Pl.
14, fig. 3, in the spherical shell and shorter spines.)

_Dimensions._--Diameter of the sphere 0.2, pores 0.005 to 0.02, bars 0.003;
polar spines 0.06 long, 0.02 broad.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

Subgenus 4. _Xiphosphæromma_, Haeckel.

_Definition._--Pores of the spherical shell irregular, of different size or
form; surface thorny or spiny (besides the two large polar spines).

14. _Xiphosphæra vesta_ n. sp. (Pl. 14, fig. 6).

Pores irregular, roundish, three to five times as broad as the bars;
fourteen to sixteen on the half equator. Scattered on the surface of the
thick-walled shell are from twenty to thirty strong three-sided pyramidal
spines of unequal size, the largest twice as long as the largest pores.
Polar spines very strong, nearly three-sided prismatic, with curved edges,
nearly as long as the axis of the sphere and twice as broad as the largest
pores.

_Dimensions._--Diameter of the sphere 0.17, pores 0.01 to 0.02, bars 0.004;
length of the polar spines 0.13, thickness 0.02 to 0.03.

_Habitat._--Pacific, central area. Station 266, depth 2750 fathoms,

15. _Xiphosphæra astræa_, n. sp.

Pores irregular, roundish, one to two times as broad as the bars; ten to
twelve on the half equator. Surface of the thick-walled shell covered with
numerous short conical thorns. Polar spines cylindro-conical, one and a
half to two times as long as the axis of the sphere.

{127}_Dimensions._--Diameter of the sphere 0.18, pores 0.01 to 0.015, bars
0.008; length of the polar spines 0.25 to 0.3, thickness 0.02.

_Habitat._--Indian Ocean, surface; Ceylon, Haeckel.

Genus 46. _Xiphostylus_,[69] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Stylosphærida# with one single lattice-sphere and two free
spines of different size or form.

The genus _Xiphostylus_ differs from the foregoing _Xiphosphæra_ in the
unequal size or form of both polar spines, which become more or less
differentiated.

Subgenus 1. _Xiphostylantha_, Haeckel.

_Definition._--Pores of the spherical shell regular, of nearly equal size
and form; surface smooth or a little rough, without spines or thorns.

1. _Xiphostylus alcedo_, n. sp. (Pl. 13, fig. 4).

Pores regular, circular, with elevated hexagonal frames, twice as broad as
the bars. Eight to ten pores on the half equator. Surface smooth. Polar
spines three-sided pyramidal, as broad at the base as one hexagon; the
major spine four to five times as long as the minor, which is about equal
to the radius of the sphere.

_Dimensions._--Diameter of the sphere 0.12, pores 0.012, bars 0.006; length
of the major polar spine 0.16 to 0.2, of the minor 0.04 to 0.06, basal
breadth 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475.

2. _Xiphostylus phasianus_, n. sp. (Pl. 13, fig. 9).

Pores regular, circular, twice as broad as the bars. Eight to ten pores on
the half equator. Outer opening of each pore elegantly lobed, with eight
indentations. Surface a little rough. Polar spines very unequal; major
spine sword-like, sharply edged, about as long as the diameter of the
sphere; minor spine scarcely half so long, pommel-shaped, with nine (?)
wing-like edges.

_Dimensions._--Diameter of the sphere 0.13, inner circular opening of the
pores 0.01, outer eight-lobed opening 0.015, bars 0.005; length of the
major polar spine 0.14, of the minor 0.06, breadth 0.03.

_Habitat._--Australian Sea, Station 162, surface.

3. _Xiphostylus motacilla_, n. sp.

Pores regular, circular, three times as broad as the bars; sixteen to
twenty on the half equator. Surface smooth. Polar spines compressed,
two-edged, at the base three to four times as broad as {128}one pore; the
major spine somewhat longer than the diameter of the shell, the minor
scarcely one-third or one-half as long.

_Dimensions._--Diameter of the sphere 0.14, pores 0.006, bars 0.002; length
of the major spine 0.16 to 0.18, of the minor 0.05 to 0.07, basal breadth
0.02.

_Habitat._--Indian Ocean, Zanzibar, 2200 fathoms, Pullen.

4. _Xiphostylus gallus_, n. sp.

Pores regular, circular, five times as broad as the bars. Twelve to sixteen
pores on the half equator. Surface smooth. Polar spines very unequal; the
major conical spine one and a half to three times as long as the diameter
of the sphere; the minor pommel-shaped, scarcely one-third as long (length
of both spines very variable).

_Dimensions._--Diameter of the sphere 0.13, pores 0.01, bars 0.002; length
of the major spine 0.2 to 0.4, of the minor 0.05 to 0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

5. _Xiphostylus alauda_, n. sp. (Pl. 14, fig. 15).

_Lithomespilus alauda_, Haeckel, 1881, Prodrom. et Atlas, _loc. cit._

Pores subregular, circular, three to four times as broad as the bars;
fifteen to eighteen on the half equator. Surface a little rough. Polar
spines irregularly conical or pyramidal, scarcely as long as the radius of
the sphere; one spine simple, the other composed of a bunch of four or five
spines united at the base.

_Dimensions._--Diameter of the sphere 0.11, pores 0.01, bars 0.003; length
of the polar spines 0.03 to 0.05, basal breadth 0.02.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

6. _Xiphostylus anhinga_, Haeckel.

_Rhabdolithis pipa_, Bury, 1862, Polycystins of Barbados, pl. iii. fig.
4.

Pores subregular, circular, about the same breadth as the bars; eight to
ten on the half equator. Surface smooth or a little rough. Polar spines
cylindrical, very irregularly curved like S or contorted, the major three
to six times as long as the diameter of the sphere, the minor scarcely
one-fourth as long as the former, at the end truncated.

_Dimensions._--Diameter of the sphere 0.07, pores and bars 0.005; length of
the major polar spine 0.2 to 0.4, of the minor 0.06 to 0.09, basal breadth
0.01.

_Habitat._--Fossil in the Barbados rocks.

Subgenus 2. _Xiphostyletta_, Haeckel.

_Definition._--Pores of the spherical shell regular, of nearly equal size
and form; surface thorny or spiny (other than the two large polar spines).

{129}7. _Xiphostylus cuculus_, n. sp.

Pores regular, circular, hexagonally framed, three times as broad as the
bars; ten to twelve on the half equator. Surface thorny, between every
three pores a short conical thorn. Polar spines three-sided prismatic, the
major somewhat longer than the diameter of the sphere, the minor scarcely
one-third as long, pommel-shaped.

_Dimensions._--Diameter of the sphere 0.17, pores 0.012, bars 0.004; length
of the major polar spine 0.2, of the minor 0.05, basal breadth 0.015.

_Habitat._--South Atlantic, surface; Station 335, depth 1425 fathoms.

8. _Xiphostylus trochilus_, n. sp. (Pl. 13, fig. 10).

Pores regular, circular, four times as broad as the bars; eight to nine on
the half equator. Polar spines cylindrical, the major somewhat longer than
the axis of the sphere, the minor shorter, surrounded by a group of from
four to eight shorter conical spines. Surface of the opposite hemisphere
smooth, without by-spines.

_Dimensions._--Diameter of the sphere 0.07 to 0.08, pores 0.01, bars
0.0025.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

9. _Xiphostylus picus_, n. sp. (Pl. 14, fig. 13).

_Lithomespilus picus_, Haeckel, 1881, Prodrom. et Atlas.

Pores regular, circular, twice as broad as the bars; sixteen to eighteen on
the half equator. Polar spines cylindrical, conical at the apex, the major
once and a half to twice as long as the diameter of the shell, the minor
scarcely half so long; around the latter a group of twelve to twenty
shorter conical spines, irregularly scattered. Surface of the other
hemisphere smooth.

_Dimensions._--Diameter of the sphere 0.13, pores 0.006, bars 0.003; length
of the major polar spine 0.2 to 0.24, of the minor 0.08 to 0.09, basal
breadth 0.02.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Subgenus 3. _Xiphostylissa_, Haeckel.

_Definition._--Pores of the spherical shell irregular, of unequal size or
form; surface smooth or a little rough, without thorns.

10. _Xiphostylus trogon_, n. sp. (Pl. 14, fig. 12).

_Lithomespilus trogon_, Haeckel, 1881, Prodrom. et Atlas.

Pores irregular, roundish or subcircular, two to three times as broad as
the bars; ten to twelve on the half equator. Surface smooth. Major polar
spine three-sided prismatic, once and a half to twice as long as the axis
of the sphere; minor spine quite rudimentary, scarcely longer than broad,
but surrounded by a group of from three to six similar short spines.

{130}_Dimensions._--Diameter of the sphere 0.1, pores 0.005 to 0.015, bars
0.005 to 0.008; length of the major spine 0.15 to 0.18, of the minor 0.01
to 0.02, basal breadth 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

11. _Xiphostylus falco_, n. sp. (Pl. 13, fig. 14).

Pores irregular, roundish, two to five times as broad as the bars; sixteen
to eighteen on the half equator. Surface smooth. Polar spines cylindrical,
very stout, nearly half as thick as the radius of the shell; major spine
two to four times as long as the diameter of the shell; minor spine
obliquely inserted, scarcely longer than the diameter, divided at the end
into two short, hook-shaped, curved branches.

_Dimensions._--Diameter of the sphere 0.08, pores 0.002 to 0.005, bars
0.001; breadth of the spines 0.02, length of the major spine 0.15 to 0.2,
of the minor 0.09.

_Habitat._--South Pacific, Station 302, depth 1450 fathoms.

12. _Xiphostylus alca_, n. sp. (Pl. 13, fig. 13).

Pores irregular, roundish, two to six times as broad as the bars; six to
eight on the half equator. Each pore with three to six lobes, composed of
three to six confluent smaller pores. Surface smooth. Major spine conical,
curved, somewhat longer than the axis of the sphere; minor spine somewhat
shorter, pommel-like, edged.

_Dimensions._--Diameter of the sphere 0.07, pores 0.01 to 0.02, bars 0.003;
length of the major spine 0.08, of the minor 0.06, basal thickness 0.02.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe, surface.

13. _Xiphostylus edolius_, n. sp. (Pl. 13, fig. 5).

Pores irregular, roundish, composed of two to six smaller confluent pores.
On the half equator six to eight large pores, and twenty to thirty small
pores; bars between the smaller very thin. Surface a little rough. Major
polar spine conical, S-shaped, about twice as long as the axis of the
shell; minor spine pommel-shaped, edged, scarcely as long as its radius.

_Dimensions._--Diameter of the sphere 0.12, large pores 0.01 to 0.03, small
pores 0.004 to 0.008, bars 0.001 to 0.004; length of the major spine 0.2,
of the minor 0.05, basal breadth 0.02.

_Habitat._--Central Pacific, Station 273, surface.

Subgenus 4. _Xiphostylomma_, Haeckel.

_Definition._--Pores of the spherical shell irregular, of different size or
form; surface thorny or spiny.

{131}14. _Xiphostylus emberiza_, n. sp. (Pl. 13, fig. 11).

Pores irregular, roundish, one to four times as broad as the bars; six to
eight on the half equator. Polar spines very unequal; major cylindrical,
twice as long as the axis of the sphere; minor scarcely half as long,
obliquely inserted, like a bird's head, surrounded by a group of ten to
twenty smaller conical spines. Opposite hemisphere smooth.

_Dimensions._--Diameter of the sphere 0.005, pores 0.002 to 0.008, bars
0.002; length of the major spine 0.09, of the minor 0.05, basal breadth
0.01.

_Habitat._--South Atlantic, Station 332, surface.

15. _Xiphostylus ardea_, n. sp.

Pores irregular, roundish, one to three times as broad as the bars; twelve
to sixteen on the half equator. Whole surface spiny. Major polar spine
three-sided pyramidal, somewhat longer than the diameter of the sphere;
minor scarcely so long as its half radius, pommel-like, edged.

_Dimensions._--Diameter of the sphere 0.12, pores 0.003 to 0.01, bars
0.003; length of the major polar spine 0.15, of the minor 0.03, basal
breadth 0.02.

_Habitat._--North Atlantic, Station 64, surface.

Genus 47. _Saturnalis_,[70] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Stylosphærida# with one single lattice-sphere and two equal
opposite spines, connected at the distal end by a circular or elliptical
ring.

The genus _Saturnalis_ (with simple lattice-sphere) and the two similar
genera _Saturnulus_ (with two concentric spheres) and _Saturninus_ (with
three spheres) form together the small peculiar group of Saturnalida,
distinguished by a remarkable circular or elliptical ring, connecting the
distal ends of the two equal opposite polar spines. This ring indicates a
certain equatorial plane, and therefore brings these #Sphæroidea# into
relation with the #Discoidea#.

Subgenus 1. _Saturnalina_, Haeckel.

_Definition._--Ring smooth, without spines or thorns.

1. _Saturnalis circularis_, n. sp.

Pores of the spherical shell regular, circular, hexagonally framed, twice
as broad as the bars. Ten to twelve pores on the half equator. Ring
circular, smooth, its diameter three times as great as that of the sphere.

{132}_Dimensions._--Diameter of the sphere 0.07, pores 0.005, bars 0.0025;
diameter of the circular ring 0.2, thickness of the axial beams and the
ring 0.01.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

2. _Saturnalis annularis_, n. sp. (Pl. 13, fig. 16).

Pores of the spherical shell regular, circular, with elevated hexagonal
frames, of the same breadth as the bars. Sixteen to twenty pores on the
half equator. Ring elliptical, smooth, somewhat constricted at the poles of
the axis, its diameter three times as great as that of the sphere.

_Dimensions._--Diameter of the sphere 0.09, pores and bars 0.005, major
axis of the elliptical ring 0.27 to 0.3, minor axis 0.19 to 0.2; thickness
of the ring and of the axial beams 0.01.

_Habitat._--Pacific, central area, Stations 270 to 274, surface.

3. _Saturnalis cyclus_, n. sp.

_Lithocircus mesocena_, Bury, 1862, Polycystins of Barbados, pl. iii.
fig. 1.

Pores of the spherical shell regular, circular, without hexagonal frames,
twice as broad as the bars. Eight to ten pores on the half equator. Ring
circular, smooth, its diameter four times as great as that of the sphere.

_Dimensions._--Diameter of the sphere 0.07, pores 0.006, bars 0.003;
diameter of the circular ring 0.28, thickness of the ring and both axial
beams 0.01.

_Habitat._--Fossil in the Barbados rocks.

4. _Saturnalis circoides_, n. sp. (Pl. 13, fig. 12).

Pores of the spherical shell irregular, roundish, often somewhat lobed, one
to three times as broad as the bars; fifteen to twenty on the half equator.
Ring circular, smooth, with four prominent edges, its diameter twice as
great as that of the sphere. (The figured specimen is a young or not fully
developed one; afterwards I found in the same locality other specimens with
quite perfect rings, similar to the edged ring of _Saturnulus annulus_, Pl.
16, fig. 17.)

_Dimensions._--Diameter of the sphere 0.09 to 0.1, pores 0.003 to 0.01,
bars 0.004; diameter of the circular ring 0.2 to 0.24, thickness of the
ring and the polar beams 0.01.

_Habitat._--Indian Ocean; fossil in the Nicobar rocks; living at great
depths near Zanzibar, 2200 fathoms, Pullen.

Subgenus 2. _Saturnalium_, Haeckel.

_Definition._--Ring armed on the periphery with numerous spines or thorns.

5. _Saturnalis trochoides_, n. sp.

_Haliomma_ species, Bury, 1862, Polycystins of Barbados, pl. xx. fig. 2.

Pores of the spherical shell subregular, circular, twice as broad as the
bars. Twelve to sixteen pores on the half equator. Ring circular, armed
with ten to twelve strong conical, irregular spines, its diameter twice as
great as that of the sphere.

{133}_Dimensions._--Diameter of the sphere 0.08, pores 0.006, bars 0.003;
diameter of the circular ring 0.16; length of the radial spines 0.02 to
0.04; thickness of the ring and the axial beams 0.01.

_Habitat._--Fossil in the Barbados rocks.

6. _Saturnalis rotula_, n. sp. (Pl. 13, fig. 15).

Pores of the spherical shell regular, circular, twice as broad as the bars;
sixteen to twenty on the half equator. Ring circular, armed with fifteen to
twenty strong, conical, irregular spines, partly simple, partly divided
into two or three irregular branches; diameter of the ring two and a half
times as great as that of the sphere.

_Dimensions._--Diameter of the sphere 0.08, pores 0.004, bars 0.002;
diameter of the circular ring 0.2, length of its spines 0.02 to 0.03;
thickness of the ring and the radial beams 0.01.

_Habitat._--North Pacific, Station 244, surface.

Subfamily SPHÆROSTYLIDA, Haeckel, 1881, Prodromus, pp. 449, 451.

_Definition._--#Stylosphærida# with two concentric, spherical
lattice-shells.

Genus 48. _Stylosphæra_,[71] Ehrenberg, 1847, Monatsber. d. Berlin Akad.,
p. 54.

_Definition._--#Stylosphærida# with two concentric lattice-spheres and two
free spines of equal size and similar form.

The genus _Stylosphæra_, the most simple form of the Sphærostylida, can be
derived either from _Xiphosphæra_ by duplication of the spherical shell, or
from _Carposphæra_ by development of two opposite polar spines. The inner
or medullary shell is enclosed in the central capsule, whilst the outer or
cortical shell lies outside it; the two are connected by two or more radial
beams, piercing the wall of the capsule.

Subgenus 1. _Stylosphærantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; surface smooth or a little rough, without spines or
thorns.

1. _Stylosphæra musa_, n. sp.

Radial proportion of the two concentric spheres = 3 : 1. Cortical shell
thin walled, smooth, with regular, hexagonal pores, three times as broad as
the thin bars; twelve on the half equator. Polar spines three-sided
pyramidal, as long as the axis of the cortical shell, one-tenth as broad at
the base.

_Dimensions._--Diameter of the outer shell 0.2, pores 0.01, bars 0.003;
diameter of the inner shell 0.06; length of the polar spines 0.2, basal
breadth 0.02.

_Habitat._--Tropical Atlantic, Station 347, depth 2250 fathoms.

{134}2. _Stylosphæra urania_, n. sp.

Radial proportion of the two shells = 4 : 1. Cortical shell thin walled,
smooth; pores regular, circular, hexagonally framed, twice as broad as the
bars; ten on the half equator. Polar spines conical, as long as the radius
of the outer shell.

_Dimensions._--Diameter of the cortical shell 0.24, pores 0.012, bars
0.006; medullary shell 0.06; length of the polar spines 0.12, basal breadth
0.024.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

3. _Stylosphæra calliope_, n. sp. (Pl. 16, fig. 6).

Radial proportion of the two shells = 3 : 1. Cortical shell thick walled,
smooth; pores regular, circular, three times as broad as the bars. Each
pore on its outer opening with eight regular lobules, flower-like. Nine to
ten pores on the half equator. Polar spines three-sided pyramidal, with
three strong prominent edges, about as long as the axis, as broad as one
pore. (Sometimes, as in the figured specimen, one spine is smaller than the
other; this variety, otherwise identical, may be called _Sphærostylus
calliope_.)

_Dimensions._--Diameter of the outer shell 0.12, pores 0.015, bars 0.005;
inner shell 0.04; length of the polar spine 0.08 to 0.12, breadth 0.02.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

4. _Stylosphæra clio_, n. sp. (Pl. 16, fig. 7).

Radial proportion of the two shells = 2 : 1. Cortical shell thick walled,
smooth; pores regular, circular, three times as broad as the bars; fourteen
to sixteen on the half equator. Polar spines three-sided pyramidal, very
robust, with thick prismatic edges, about as long as the axis of the
cortical shell, one-third as broad at the base. (Sometimes, as in the
figured specimen, one spine is greater than the other; this form may be
called _Sphærostylus clio_.)

_Dimensions._--Diameter of the outer shell 0.12, pores 0.01, bars 0.003;
inner shell 0.06; length of the polar spines 0.08 to 0.12, basal breadth
0.03 to 0.04.

_Habitat._--Pacific, central area; Station 272, depth 2600 fathoms.

5. _Stylosphæra polyhymnia_, n. sp.

Radial proportion of the two spheres = 3 : 1. Cortical shell very thin
walled, smooth, with regular, circular pores, three times as broad as the
bars; sixteen to twenty on the half equator. Polar spines cylindrical,
pointed, once and a half to twice as long as the axis of the outer sphere,
scarcely broader than one pore. The two spheres are connected only by the
two opposite beams.

_Dimensions._--Diameter of the outer shell 0.12 to 0.16, pores 0.006 to
0.009, bars 0.002 to 0.003; inner shell 0.04 to 0.05; length of the polar
spines 0.18 to 0.22, breadth 0.01.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

{135}6. _Stylosphæra dixyphos_, Haeckel.

_Haliomma dixyphos_, Ehrenberg, 1854, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 83; Mikrogeol., Taf. xxii. fig. 31.

_Haliomma dixyphos_, Haeckel, 1862, Monogr. d. Radiol. p. 433.

Radial proportion of the two spheres = 2 : 1. Cortical shell thin walled,
smooth, with regular, circular pores, twice as broad as the bars; ten to
twelve on the half equator. Polar spines about as long as the axis of the
outer shell, three-sided pyramidal, at the base twice as broad as one pore.
(The two spheres connected by four beams, two opposite in the main axis,
two opposite in the equatorial axis.)

_Dimensions._--Diameter of the outer shell 0.1, pores 0.01, bars 0.05;
inner shell 0.05; length of the polar spines 0.08 to 0.1, basal breadth
0.02.

_Habitat._--South Atlantic, Station 332, surface; fossil in Tertiary rocks
of Sicily.

Subgenus 2. _Stylosphærella_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; surface thorny or spiny.

7. _Stylosphæra setosa_, Ehrenberg, 1872.

_Stylosphæra setosa_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 320; Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. viii.
fig. 15.

Radial proportion of the two shells = 2 : 1. Cortical shell thin walled,
spiny; pores regular, hexagonal, four times as broad as the bars. Six to
eight pores on the half equator. Polar spines conical, thin, scarcely as
long as the radius of the cortical shell.

_Dimensions._--Diameter of the cortical shell 0.1, pores 0.002, bars 0.005;
medullary shell 0.05; length of the polar spines 0.04, basal breadth 0.01.

_Habitat._--Philippine Sea, depth 3300 fathoms, Ehrenberg; Station 206,
depth 2100 fathoms.

8. _Stylosphæra euterpe_, n. sp.

Radial proportion of the two shells = 3 : 1. Cortical shell thin walled,
spiny; pores regular, circular, with hexagonal frames, twice as broad as
the bars; eight to ten on the half equator. Polar spines conical, as thick
as one pore at the base, about as long as the axis of the cortical shell.

_Dimensions._--Diameter of the cortical shell 0.12, pores 0.012, bars
0.006; medullary shell 0.04; length of the polar spines 0.1, basal breadth
0.012.

_Habitat._--South Pacific, Station 302, depth 1450 fathoms.

9. _Stylosphæra melpomene_, n. sp. (Pl. 16, fig. 1).

Radial proportion of the two shells = 3 : 1. Cortical shell thin walled,
spiny, with regular, circular pores, four times as broad as the bars; eight
to ten on the half equator. Polar spines three-sided prismatic, pointed, as
broad as one pore, only one-third as long as the axis of the sphere (the
two shells connected by four thin beams, two opposite in the main axis, two
in the equatorial axis).

{136}_Dimensions._--Diameter of the outer shell 0.12, pores 0.012, bars
0.003; inner shell 0.04; length of the polar spines 0.04, thickness 0.013.

_Habitat._--Indian Ocean, Cocos Islands, Rabbe.

10. _Stylosphæra hispida_, Ehrenberg, 1854.

_Stylosphæra hispida_, Ehrenberg, 1854, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 246; Mikrogeol, Taf. xxxvi. fig. 26.

_Haliomma hispidum_, Haeckel, 1862, Monogr. d. Radiol., p. 433.

Radial proportion of the two spheres = 3 : 1. Cortical shell thick walled,
spiny, with regular, circular pores of the same breadth as the bars; ten to
fifteen on the half equator. Polar spines three-sided prismatic, pointed,
about as long as the axis of the outer sphere, nearly as broad at the base
as the inner sphere. (Compare _Sphærostylus hispidus_; also Ehrenberg,
Monatsber. d. k. preuss. Akad. d. Wiss. Berlin, 1874, p. 259.)

_Dimensions._--Diameter of the outer shell 0.1 to 0.12, pores and bars
0.004; inner shell 0.04; length of the polar spines 0.1 to 0.15, basal
breadth 0.03.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Barbados, Nicobars, &c.

11. _Stylosphæra liostylus_, Ehrenberg, 1875.

_Stylosphæra liostylus_, Ehrenberg, Abhandl. d. k. Akad. d. Wiss. Berlin,
p. 84, Taf. xxv. fig. 3.

Radial proportion of the two spheres = 3 : 1. Cortical shell thick walled,
thorny, with regular, circular pores, three times as broad as the bars;
eight to ten on the half equator. Polar spines conical, once and a half to
twice as long as the axis of the outer sphere, half as broad at the base as
its radius. (This species, common in the Barbados rocks, is different from
_Sphærostylus liostylus_, _loc. cit._, fig. 2, which Ehrenberg believed
identical.)

_Dimensions._--Diameter of the outer shell 0.1, pores 0.01, bars 0.0035;
inner shell 0.03; length of the polar spines 0.14 to 0.18, basal breadth
0.02.

_Habitat._--Fossil in the Barbados rocks; living in the depths of the North
Atlantic, Gulf Stream, Florida.

Subgenus 3. _Stylosphærissa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface smooth or a little rough, without thorns or spines.

12. _Stylosphæra nana_, n. sp. (Pl. 16, figs. 12, 13).

Radial proportion of the two spheres = 2 : 1. Cortical shell thick walled,
somewhat irregular, smooth, with irregular, roundish pores, one to three
times as broad as the bars; eight to ten on the half equator. Polar spines
three-sided pyramidal, scarcely as long as the axis of the outer sphere,
and nearly as broad at the base as its radius. (A very variable and
irregular form.)

_Dimensions._--Diameter of the outer shell 0.07 to 0.09, pores 0.003 to
0.009, bars 0.003; inner shell 0.03 to 0.04; length of the polar spines
0.04 to 0.07, basal breadth 0.03.

_Habitat._--North Pacific, Stations 241 to 253, surface.

{137}13. _Stylosphæra jugata_, n. sp.

Radial proportion of the two shells = 2 : 1. Cortical shell thick walled,
smooth, with irregular, roundish, double-contoured pores, confluent in
groups of two to six. On the half equator six to nine groups and fifteen to
twenty pores; bars between them of very variable breadth. Polar spines very
strong, three-sided pyramidal, twice as long as the axis of the outer
sphere, half as broad at the base as its radius. (Nearly allied to
_Lithatractus jugatus_, Pl. 16, fig. 2, but differs in the truly spherical
form of both shells and the double length of the polar spines.)

_Dimensions._--Diameter of the outer shell 0.15, pores 0.005 to 0.02; inner
shell 0.07; length of the polar spines 0.25 to 0.3, basal breadth 0.03.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

14. _Stylosphæra terpsichore_, n. sp.

Radial proportion of the two shells = 3 : 1 or 4 : 1. Cortical shell thick
walled, smooth, with irregular, roundish pores, one to three times as broad
as the bars; fifteen to twenty-five on the half equator. Polar spines
conical, about as long as the axis of the outer sphere, as broad at the
base as the inner shell.

_Dimensions._--Diameter of the outer shell 0.15 to 0.2, pores 0.005 to
0.02, bars 0.004 to 0.008; inner shell 0.05; length of the polar spines
0.15 to 0.25, basal breadth 0.05.

_Habitat._--Western Indian Ocean, Zanzibar, depth 2200 fathoms, Pullen.

Subgenus 4. _Stylosphæromma_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface spiny or thorny.

15. _Stylosphæra thalia_, n. sp.

Radial proportion of the two shells = 2 : 1. Cortical shell thin walled,
thorny, with irregular, roundish pores, two to four times as broad as the
bars; eight to twelve on the half equator. Polar spines conical, one to one
and a half times as long as the axis of the outer sphere, one-fourth to
one-sixth as thick at the base. (Resembles _Sphærostylus ophidium_, Pl. 16,
fig. 14, but differs in the straight regular conical polar spines, both of
equal length and similar form.)

_Dimensions._--Diameter of the outer shell 0.12, pores 0.01 to 0.02, bars
0.005; inner shell 0.06; polar spines 0.1 to 0.16 long, 0.03 broad.

_Habitat._--South Pacific, Station 302, depth 1450 fathoms.

16. _Stylosphæra erato_, n. sp.

Radial proportion of the two shells = 3 : 1. Cortical shell thick walled,
thorny, with irregular, roundish pores, two to five times as broad as the
bars; fourteen to eighteen on the half equator. {138}Polar spines
three-sided pyramidal, about as long as the axis of the outer sphere,
one-fourth as broad as its radius. (Similar to _Xiphosphæra vesta_, Pl. 14,
fig. 6.)

_Dimensions._--Diameter of the outer shell 0.15, pores 0.008 to 0.015, bars
0.003; inner shell 0.05; polar spines 0.12 long, 0.02 broad.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

Genus 49. _Sphærostylus_,[72] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Stylosphærida# with two concentric lattice-spheres and two
free spines, of different size or form.

The genus _Sphærostylus_ differs from _Stylosphæra_ in the different size
or form of the two polar spines, and therefore has the same relation to it
that _Xiphostylus_ bears to _Xiphosphæra_.

Subgenus 1. _Sphærostylantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; surface smooth or a little rough, without thorns.

1. _Sphærostylus liostylus_, Haeckel.

_Stylosphæra liostylus_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, Taf. xxv. fig. 2.

Cortical shell thin walled, with rough surface, three times as broad as the
medullary shell. Pores of the cortical shell regular, circular, twice as
broad as the bars; ten to twelve on the half equator. Polar spines
cylindrical, as broad as one pore, with conical apex; the minor spine about
as long as the axis of the outer shell, the major three to four times as
long.

_Dimensions._--Diameter of the outer sphere 0.12, pores 0.12, bars 0.006;
diameter of the inner sphere 0.04; length of the major polar spine 0.3 to
0.4, of the minor 0.1 to 0.15, breadth 0.012.

_Habitat._--Fossil in the Barbados rocks.

2. _Sphærostylus flexuosus_, Haeckel.

_Stylosphæra flexuosa_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, Taf. xxv. fig. 5.

Cortical shell thick walled, with rough surface, three times as broad as
the medullary shell. Pores of the cortical shell regular circular, of the
same breadth as the bars; eight to ten on the half equator. Polar spines
cylindrical, S-like curved, irregular; the minor scarcely as long as the
axis of the outer sphere, the major two to three times as long. (In the
figure of Ehrenberg the spines are broken off; I have found them myself
constantly irregular and of unequal length, sometimes with conical apex.)

{139}_Dimensions._--Diameter of the outer sphere 0.08, pores and bars
0.006; inner sphere 0.03; length of the major polar spine 0.15 to 0.25, of
the minor 0.07 to 0.09, breadth 0.02.

_Habitat._--Fossil in the Barbados rocks.

3. _Sphærostylus clio_, n. sp.

Cortical shell thick walled, with smooth surface, twice as broad as the
medullary shell; its network has regular, circular pores, three times as
broad as the bars; sixteen to twenty on the half equator. Polar spines very
strong, three-sided pyramidal; the major nearly twice as long as the axis
of the outer sphere, the minor scarcely as long as its radius. (Nearly
related to _Stylosphæra clio_, Pl. 16, fig. 7, but differs in the slender
form and unequal length of the polar spines.)

_Dimensions._--Diameter of the outer sphere 0.14, pores 0.01, bars 0.03;
inner sphere 0.07; length of the major spine 0.25, of the minor 0.06, basal
breadth 0.02.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

4. _Sphærostylus hippocampus_, n. sp. (Pl. 16, figs. 10, 11).

Cortical shell thick walled, with smooth surface and regular network; the
pores circular, hexagonally-lobed, three times as broad as the bars; ten to
twelve on the half equator. Medullary shell half as large, with very small
circular pores, eight to ten on the half equator. Major polar spine
scarcely as long as the axis of the outer shell, curved like a horn; minor
spine scarcely half as long, pommel-like, edged. (Sometimes, by
prolongation of the main axis, the spherical shells become ellipsoidal and
thus the species is transformed into _Druppatractus hippocampus_.)

_Dimensions._--Diameter of the outer sphere 0.08, pores 0.009, bars 0.003;
inner sphere 0.04; length of the major spine 0.07, of the minor 0.03,
breadth 0.02.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

Subgenus 2. _Sphærostyletta_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and form; surface spiny or thorny.

5. _Sphærostylus diadema_, n. sp.

Cortical shell thick walled, spiny, twice as broad as the medullary shell,
with regular, circular pores, twice as broad as the bars; eight to ten on
the half equator. Major polar spine about as long as the axis of the outer
sphere, three-sided pyramidal; minor spine scarcely half as long,
pommel-shaped, edged. (Similar in general form to _Xiphatractus glyptodon_,
Pl. 17, figs. 9, 10; but differs in the simple medullary shell, the
spherical form of both shells, and the simple circular regular pores.)

{140}_Dimensions._--Diameter of the outer sphere 0.12, pores 0.014, bars
0.007; inner sphere 0.06; length of the major spine 0.13, of the minor
0.05, greatest breadth 0.04.

_Habitat._--Southern Pacific, surface, Station 289.

Subgenus 3. _Sphærostylissa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface smooth or a little rough.

6. _Sphærostylus cottus_, n. sp.

Cortical shell thick walled, smooth, about twice as broad as the medullary
shell, with irregular, roundish pores, scarcely larger than the bars;
fifteen to twenty on the half equator. Polar spines conical, the major once
and a half to twice as long as the axis of the outer sphere, the minor
scarcely as long as its radius.

_Dimensions._--Diameter of the outer sphere 0.14, pores and bars 0.008 to
0.012; inner sphere 0.065; length of the major spine 0.2 to 0.3, of the
minor 0.05 to 0.07, basal breadth 0.03.

_Habitat._--Northern Atlantic, Færöe Channel, surface, John Murray.

7. _Sphærostylus trigla_, n. sp.

Cortical shell thin walled, smooth, three times as broad as the medullary
shell, with irregular, roundish pores, two to three times as broad as the
bars; ten to twelve on the half equator. Polar spines very unequal; the
major three-sided pyramidal, one and a half times as long as the axis of
the outer sphere; the minor scarcely as long as its radius, edged,
pommel-like.

_Dimensions._--Diameter of the outer sphere 0.12, pores 0.01 to 0.015, bars
0.005; inner sphere 0.04; length of the major spine 0.2, of the minor 0.05,
breadth 0.03.

_Habitat._--Northern Pacific, Station 241, depth 2300 fathoms.

Subgenus 4. _Sphærostylomma_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface spiny or thorny.

8. _Sphærostylus ophidium_, n. sp. (Pl. 16, figs. 14, 15).

_Stylosphæra ophidium_, Haeckel, 1878, Atlas, _loc. cit._

Cortical shell thin walled, thorny, twice as broad as the medullary shell,
with irregular, roundish pores; eight to ten on the half equator. Polar
spines conical, more or less curved or S-shaped; the minor scarcely as long
as the axis of the outer sphere, the major two to three times as long.

{141}_Dimensions._--Diameter of the outer sphere 0.11, pores 0.01 to 0.017,
bars 0.003 to 0.007; diameter of the inner sphere 0.06, pores 0.005 to
0.008, bars 0.002 to 0.004; length of the major polar spine 0.25, of the
minor 0.1, basal breadth 0.03.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

Genus 50. _Saturnulus_,[73] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Stylosphærida# with two concentric lattice-spheres and two
equal opposite spines, the distal ends of which are connected by a circular
or elliptical ring.

The genus _Saturnulus_ differs from the similar _Saturnalis_ by the
duplication of the spherical lattice-shell; the inner lies within, the
outer without the central capsule.

1. _Saturnulus circulus_, n. sp.

Cortical shell smooth, twice as broad as the medullary shell, with regular,
circular, hexagonally framed pores, of the same breadth as the bars;
fourteen to sixteen on the half equator. Ring circular, smooth, without
edges, its diameter three times as great as that of the sphere.

_Dimensions._--Diameter of the outer sphere 0.08, pores and bars 0.004; of
the inner sphere 0.04, of the ring 0.24.

_Habitat._--Central Pacific, Station 272, surface.

2. _Saturnulus annulus_, n. sp.

Cortical shell smooth, three times as broad as the medullary shell, with
regular, circular, hexagonally framed pores, of the same breadth as the
bars; eighteen to twenty on the half equator. Ring elliptical, smooth,
without edges, somewhat constricted at the poles of the minor axis, its
major diameter four times as great as that of the sphere. Differs from
_Saturnalis annularis_, Pl. 13, fig. 16, mainly in the possession of a
medullary shell.

_Dimensions._--Diameter of the outer sphere 0.1, pores and bars 0.005;
inner sphere 0.033; major axis of the ring 0.4, minor 0.3.

_Habitat._--North Pacific, Station 244, surface.

3. _Saturnulus ellipticus_, n. sp. (Pl. 16, fig. 16).

Cortical shell smooth, three times as broad as the medullary shell, with
regular, circular pores, three times as broad as the bars; sixteen to
eighteen on the half equator. Ring elliptical, smooth, without edges, its
major diameter three times as great as that of the sphere.

_Dimensions._--Diameter of the outer sphere 0.09, pores 0.006, bars 0.002;
inner sphere 0.03; major axis of the elliptical ring 0.28, minor 0.24;
thickness of the ring and the axial beams 0.008.

_Habitat._--South Pacific, Station 300, surface.

{142}4. _Saturnulus planetes_, n. sp. (Pl. 16, fig. 17).

Cortical shell smooth, twice as broad as the medullary shell, with regular,
circular pores, of the same breadth as the bars; sixteen to eighteen on the
half equator. Ring elliptical, smooth, with strong prominent edges,
constricted at the poles of the minor axis, its major diameter three times
as great as that of the outer sphere.

_Dimensions._--Diameter of the outer sphere 0.08, pores and bars 0.005;
inner sphere 0.035; major axis of the ring 0.25, minor axis 0.2; thickness
of the ring and the axial beams 0.012.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe; Station 200, surface.

Subfamily AMPHISTYLIDA, Haeckel, 1881, Prodromus, pp. 449, 452.

_Definition._--#Stylosphærida# with three concentric spherical
lattice-shells.

Genus 51. _Amphisphæra_, Haeckel,[74] 1881, Prodromus, p. 452.

_Definition._--#Stylosphærida# with three concentric lattice-spheres and
two free spines of equal size and similar form.

The genus _Amphisphæra_ differs from its probable ancestral form,
_Stylosphæra_, in the triple spherical lattice-shell. Commonly two of these
lie within the central capsule (medullary shell), whilst the third lies
outside it (cortical shell). But sometimes this order is inverted, the
cortical shell being double, the medullary shell simple; and perhaps these
forms may better represent a peculiar genus, _Amphisphæridium_.

Subgenus 1. _Amphisphærantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; surface smooth or a little rough, without spines or
thorns.

1. _Amphisphæra neptunus_, n. sp.

Radial proportion of the three concentric spheres = 4 : 2 : 1. Cortical
shell thick walled, smooth, with regular, circular, hexagonally framed
pores, of the same breadth as the bars; twelve to fifteen on the half
equator. Polar spines three-sided pyramidal, with strong prominent edges,
about as long as the radius of the outer shell, half as broad at the base.
(Similar to _Stylatractus neptunus_, Pl. 17, fig. 6, but differs in the
purely spherical form of the three concentric shells and the regular form
of the network and of the polar spines.)

_Dimensions._--Diameter of the outer shell 0.16, middle shell 0.08, inner
shell 0.04; pores and bars of the cortical shell 0.008; length of the polar
spines 0.08, basal breadth 0.04.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{143}2. _Amphisphæra uranus_, n. sp.

Radial proportion of the three spheres = 4 : 2 : 1. Cortical shell thick
walled, smooth, with regular, circular pores, three times as broad as the
bars; ten to twelve on the half equator. Polar spines three-sided
pyramidal, about as long as the diameter of the outer shell, twice as broad
at the base as one pore.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.06, inner 0.03;
pores of the cortical shell 0.012, bars 0.004; length of the polar spines
0.1, basal breadth 0.024.

_Habitat._--Western Indian Ocean, Zanzibar, 2200 fathoms, Pullen.

3. _Amphisphæra jupiter_, n. sp.

Radial proportion of the three spheres = 10 : 3 : 2. Cortical shell thick
walled, smooth, with regular, circular pores, twice as broad as the bars;
twenty to twenty-five on the half equator. Polar spines cylindro-conical,
nearly as long as the axis of the outer shell, twice as broad at the base
as one pore.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.06, inner 0.04;
pores of the cortical shell 0.01, bars 0.005; length of the polar spines
0.15 to 0.18, breadth 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Subgenus 2. _Amphisphærella_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; surface thorny or spiny.

4. _Amphisphæra apollo_, n. sp.

Radial proportion of the three spheres = 3 : 1.5 : 1. Cortical shell thick
walled, spiny, with regular, circular, hexagonally framed pores, of the
same breadth as the bars; twenty to twenty-two on the half equator. In each
hexagon-corner (between three pores) a bristle-like radial spine. Polar
spines cylindrical, with conical apex; one to one and a half times as long
as the axis of the outer shell, as broad as three pores at the base.
(Resembles closely _Xiphosphæra pallas_, Pl. 14, fig. 4, but differs in the
presence of two medullary shells and the absence of the fine denticulations
on the hexagonal crests.)

_Dimensions._--Diameter of the outer shell 0.12, middle 0.06, inner 0.04;
pores and bars of the cortical shell 0.006; length of the polar spines
0.15, breadth 0.02.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

5. _Amphisphæra mercurius_, n. sp.

Radial proportion of the three spheres = 3 : 2 : 1. Cortical shell thin
walled, spiny, with regular, circular pores, twice as broad as the bars;
fifteen to eighteen on the half equator; between {144}them short
bristle-like radial spines. Polar spines three-sided pyramidal, about as
long as the radius of the outer shell, one-third as broad at the base.

_Dimensions._--Diameter of the outer shell 0.15, middle 0.09, inner 0.05;
pores of the cortical shell 0.01, bars 0.005; length of the polar spines
0.09, basal breadth 0.03.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

Subgenus 3. _Amphisphærissa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface smooth or a little rough, without spines or thorns.

6. _Amphisphæra cronos_, n. sp. (Pl. 17, fig. 5).

Radial proportion of the three spheres = 3 : 2 : 1. Cortical shell thin
walled, smooth, with irregular, roundish pores, two to four times as broad
as the bars; eight to ten on the half equator. Outer medullary shell
similar, but with pores of half the size, connected with the cortical shell
by numerous radial beams; inner medullary shell with very small pores.
Polar spines three-sided pyramidal, as long as the axis of the inner
medullary shell, half as broad at the base.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.08, inner 0.04;
pores of the cortical shell 0.01 to 0.02, bars 0.05; length of the polar
spines 0.04, basal breadth 0.02.

_Habitat._--South Atlantic, Station 330, surface.

7. _Amphisphæra pluto_, n. sp. (Pl. 17, figs. 7, 8).

Radial proportion of the three spheres about = 4 : 2 : 1 (or 11 : 7 : 3).
Cortical shell thick walled, smooth, with very irregular, roundish pores,
two to four times as broad as the bars; eight to ten on the half equator;
often two to four pores confluent. Margin of their outer aperture double.
Polar spines conical, double contoured, as long as the radius of the outer
shell, one-third as broad at the base.

_Dimensions._--Diameter of the outer shell 0.11, middle shell 0.07, inner
shell 0.03; pores of the cortical shell 0.01 to 0.02, bars 0.06, length of
the polar spines 0.06, basal breadth 0.02.

_Habitat._--Central Pacific, Station 268, surface.

Subgenus 4. _Amphisphæromma_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface spiny or thorny.

8. _Amphisphæra mars_, n. sp.

Radial proportion of the three spheres = 10 : 3 : 2. Cortical shell thin
walled, thorny, with irregular, roundish pores, one to three times as broad
as the bars; sixteen to twenty on the {145}half equator. Irregularly
scattered between them short conical thorns. Polar spines conical, about as
long as the axis of the outer shell, as broad at the base as the inner
shell.

_Dimensions._--Diameter of the outer shell 0.2, middle shell 0.06, inner
shell 0.04; pores of the cortical shell 0.005 to 0.02, bars 0.006; length
of the polar spines 0.17, basal breadth 0.04.

_Habitat._--North Atlantic, Gulf Stream, near Florida, depth 1500 fathoms,
Schaffner.

Genus 52. _Amphistylus_,[75] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Stylosphærida# with three concentric lattice-spheres and
two free spines of different size or form.

The genus _Amphistylus_ differs from its ancestral form, _Amphisphæra_, by
the differentiation of both polar spines, and exhibits therefore the same
relation to it that _Sphærostylus_ bears to _Stylosphæra_.

1. _Amphistylus clio_, n. sp.

Radial proportion of the three spheres = 4 : 2 : 1. Cortical shell thick
walled, smooth; its pores regular, circular, three times as broad as the
bars; eighteen to twenty on the half equator. Polar spines three-sided
pyramidal, very stout, as broad at the base as the inner medullary shell;
major spine somewhat longer than the diameter of the cortical shell; minor
spine scarcely half as long. (Similar to _Stylosphæra clio_, Pl. 16, fig.
7, but different in the double medullary shell and the different length of
the polar spines.)

_Dimensions._--Diameter of the outer sphere 0.15, middle 0.08, inner 0.04;
pores of the outer shell 0.01, bars 0.003; length of the major spine 0.17,
minor 0.07, basal breadth 0.04.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

2. _Amphistylus hippocampus_, n. sp.

Radial proportion of the three spheres = 3 : 2 : 1. Cortical shell thick
walled, smooth; its pores regular, circular, hexagonally lobed, three times
as broad as the bars; twelve to fifteen on the half equator. Major polar
spine three-sided pyramidal, about as long as the axis of cortical shell;
minor spine pommel-like, edged, scarcely one-third as long. (Similar to
_Sphærostylus hippocampus_, Pl. 16, figs. 10, 11, but differs in the larger
size, the double medullary shell, and the straight, not curved, major
spine.)

_Dimensions._--Diameter of the outer sphere 0.11, middle 0.07, inner 0.035;
pores of the outer shell 0.011, bars 0.004; length of the major spine 0.12,
of the minor 0.04, breadth 0.03.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

3. _Amphistylus glyptodon_, n. sp.

Radial proportion of the three spheres = 6 : 3 : 1. Cortical shell thick
walled, spiny, with irregular, roundish pores, two to four times as broad
as the bars; ten to twelve on the half equator. {146}Inner aperture of each
pore closed by a thin fenestrated lamella with five to seven small pores.
Major polar spine three-sided pyramidal, longer than the diameter of the
outer sphere; minor polar spine scarcely as long as its radius,
pommel-shaped. (Similar to _Xiphatractus glyptodon_, Pl. 17, figs. 9, 10,
but differs in the spherical, not ellipsoidal, form of the three shells and
the size of the polar spines.)

_Dimensions._--Diameter of the outer sphere 0.12, middle 0.06, inner 0.02;
pores and spines of the outer shell 0.01 to 0.02, bars and porules 0.006;
length of the major polar spine 0.15, of the minor 0.05, breadth 0.04.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Genus 53. _Saturninus_,[76] n. gen.

_Definition._--#Stylosphærida# with three concentric lattice-spheres and
two equal opposite spines, connected at the distal end by a circular or
elliptical ring.

The genus _Saturninus_ differs from the similar _Saturnulus_ by the
triplication of the spherical lattice-shell; the inner shell is enclosed in
the central capsule, whilst both the others lie outside it.

1. _Saturninus triplex_, n. sp.

Radial proportion of the three spheres = 4 : 3 : 1. Inner cortical shell
with regular, circular pores, of the same breadth as the bars, sixteen to
eighteen on the half equator; outer cortical shell connected with the inner
by numerous bristle-like radial spines, network very delicate,
cobweb-shaped, with irregular polygonal meshes. Ring elliptical, two and a
half times as broad as the outer shell.

_Dimensions._--Diameter of the inner sphere 0.03, middle 0.09, outer 0.12;
pores and bars of the middle shell 0.004; major axis of the elliptical ring
0.3, minor 0.25; thickness of the ring and the axial beams 0.008.

_Habitat._--Central Pacific, Station 272, surface.

Subfamily CROMYOSTYLIDA, Haeckel, 1881, Prodromus, pp. 449, 453.

_Definition._--#Stylosphærida# with four concentric spherical
lattice-shells (two medullary and two cortical).

Genus 54. _Stylocromyum_,[77] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Stylosphærida# with four concentric lattice-spheres and two
free spines of equal size and similar form.

{147}The genus _Stylocromyum_ differs from its probable ancestral form,
_Amphisphæra_, by the duplication of the cortical shell; two opposite
radial beams, piercing the wall of the central capsule, connect it with the
double medullary shell, and are prolonged outside into two equal spines.

1. _Stylocromyum amphiconus_, n. sp.

Surface of the shell smooth. Radial proportion of the four spheres =
1 : 2 : 8 : 9. Two medullary shells inside the central capsule, two
cortical shells outside it. Interval between the second and third shells
the greatest. Pores of all the shells regular, circular, two to four times
as broad as the bars. Both polar spines equal, conical, about as long as
the axis of the outermost sphere, as broad at the base as the innermost.

_Dimensions._--Diameter of the four spheres--(A) inner medullary shell
0.03, (B) outer medullary shell 0.06, (C) inner cortical shell 0.25, (D)
outer cortical shell 0.28; length of the polar spines 0.3, basal breadth
0.03.

_Habitat._--Central Pacific, Station 273, depth 2350 fathoms.

2. _Stylocromyum amphipyramis_, n. sp.

Surface of the shell smooth. Radial proportion of the four spheres =
2 : 3 : 6 : 8. Both medullary shells with regular, circular pores, twice as
broad as the bars. Both cortical shells with irregular, roundish, much
larger pores, three to six times as broad as the bars. Both polar spines
equal or nearly equal, three-sided pyramidal, about as long as the axis of
the third shell.

_Dimensions._--Diameter of the four spheres--(A) 0.04, (B) 0.06, (C) 0.12,
(D) 0.16; length of the polar spines 0.11, basal breadth 0.02.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Genus 55. _Cromyostylus_,[78] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Stylosphærida# with four concentric lattice-spheres and two
free spines of different size or form.

The genus _Cromyostylus_ differs from its ancestral form, _Stylocromyum_,
in the differentiation of the two unequal polar spines.

1. _Cromyostylus gladius_, n. sp.

Surface of the shell smooth. Radial proportion of the four spheres =
1 : 3 : 10 : 12. Both medullary shells with regular, circular, simple
pores. Inner cortical shell with regular, circular, hexagonally framed
pores, twice as broad as the bars. From each hexagon-corner arises a
{148}bristle-shaped radial spine, which at the distal end gives off three
thread-like branches; by communication of these threads (at equal distances
from the centre) the delicate outer medullary shell is formed. The polar
spines very different; major spine six-sided pyramidal, longer than the
diameter of the shell; minor pommel-shaped, shorter than the radius
(similar to _Xiphatractus glyptodon_, Pl. 17, figs. 9, 10, but different in
the double spherical cortical shell).

_Dimensions._--Diameter of the four spheres--(A) 0.02, (B) 0.06, (C) 0.2,
(D) 0.24; length of the major spine 0.3, of the minor 0.1.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Subfamily CARYOSTYLIDA, Haeckel, 1881, Prodromus, pp. 449, 454.

_Definition._--#Stylosphærida# with five or more concentric, spherical
lattice-shells.

Genus 56. _Caryostylus_, Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Stylosphærida# with five to six or more concentric
lattice-shells and two free opposite spines of equal size and similar form.

The genus _Caryostylus_ differs from its ancestral form, _Stylocromyum_, by
the multiplication of the concentric spheres, the number of which amounts
to five or six or more. I have only observed one single species of this
genus. Some similar forms which in my Prodromus (1881, p. 454) were annexed
to it, and disposed in three nearly allied genera (_Caryoxiphus_,
_Caryodoras_, _Caryolonche_), have now been proved to belong to other
groups, mainly ellipsoidal Druppulida.

1. _Caryostylus hexalepas_, n. sp.

Surface of the spherical shell smooth. Radial proportion of the component
six concentric shells = 1 : 2 : 7 : 9 : 12 : 15. Both medullary shells
connected only by six radial beams, opposite in pairs in the three
dimensive axes. Between second and third shell numerous (twenty regularly
disposed?) radial beams. Four cortical shells connected by very numerous
(sixty to eighty or more?) short radial beams. Pores of all six shells
regular, circular, the size increasing towards the surface, two to three
times as broad as the bars. Two opposite polar spines very large, of equal
size, three times as long as the shell radius, cylindrical, club-shaped at
the thicker distal end. (The whole shell structure is similar to Pl. 15,
fig. 2, but the shells are spherical, not ellipsoidal.)

_Dimensions._--Diameter of the six spheres--(A) 0.02, (B) 0.04, (C) 0.15,
(D) 0.18, (E) 0.24, (F) 0.3; length of the spines 0.5.

_Habitat._--West Tropical Pacific, Station 225, depth 4475 fathoms.

Subfamily SPONGOSTYLIDA, Haeckel, 1881, Prodromus, pp. 449, 455.

_Definition._--#Stylosphærida# with spherical spongy shell (with or without
enclosed latticed medullary shells).

{149}Genus 57. _Spongolonchis_,[79] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Stylosphærida# with a solid sphere of spongy framework, and
with two opposite free radial spines.

The genus _Spongolonchis_ differs from its probable ancestral form,
_Styptosphæra_, by the development of two opposite radial spines situated
in one axis.

1. _Spongolonchis compacta_, n. sp.

Spongy framework of the spherical shell very compact, with small meshes,
three to four times as broad as the bars. Surface rough, but not spiny. Two
polar spines pyramidal, only as long as the radius of the shell, one-third
as broad at the base.

_Dimensions._--Diameter of the shell 0.2; length of the spines 0.1, basal
breadth 0.03.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

2. _Spongolonchis laxa_, n. sp.

Spongy framework loose, with large meshes, ten to twelve times as broad as
the bars. Surface spiny. Two polar spines three-sided prismatic, longer
than the diameter of the shell (broken off in the observed specimen).

_Dimensions._--Diameter of the shell 0.5; length of the spine 0.6 and more,
breadth 0.02.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Genus 58. _Spongostylus_,[80] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Stylosphærida# with spongy spherical cortical shell,
enclosing in the centre a simple latticed medullary shell, and with two
opposite free radial spines.

The genus _Spongostylus_ may probably be derived from _Spongoplegma_ by
development of two opposite radial spines in one axis.

1. _Spongostylus hastatus_, n. sp.

Spongy cortical shell with compact framework, twice as broad as the
enclosed medullary shell, the pores of which are regular, circular, twice
as broad as the bars. Two polar spines, three times as long as the radius
of the shell, in the basal two-thirds cylindrical, in the distal third
compressed, two-edged, spear-shaped.

_Dimensions._--Diameter of the cortical shell 0.1, medullary shell 0.05;
length of the spines 0.15.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

{150}2. _Spongostylus gladiatus_, Haeckel.

_Stylosphæra holosphæra_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 299, Taf. viii. fig. 14.

Spongy cortical shell with rather compact framework, four times as broad as
the enclosed medullary shell, the pores of which are irregular, roundish,
three to four times as broad as the bars. Two polar spines sword-shaped,
two-edged, as long as the shell diameter.

_Dimensions._--Diameter of the cortical shell 0.24, medullary shell 0.06;
length of the spines 0.25.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms; Philippine
Sea, depth 3300 fathoms.

3. _Spongostylus serratus_, n. sp.

Spongy cortical shell with lax framework, ten to twelve times as broad as
the enclosed medullary shell, the pores of which are irregular, roundish,
once and a half to twice as broad as the bars. Two polar spines longer than
the shell diameter (broken off in the observed specimen), three-sided
prismatical, with three straight serrated edges.

_Dimensions._--Diameter of the cortical shell 0.5, medullary shell 0.04;
length of the spines 0.6 or more.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Genus 59. _Spongostylidium_,[81] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Stylosphærida# with spongy spherical cortical shell,
enclosing two concentric spherical latticed medullary shells, and with two
opposite, free radial spines.

The genus _Spongostylidium_ differs from _Spongostylus_ by duplication of
the latticed medullary shell, and therefore bears the same relation to it
as _Spongodictyon_ to _Spongoplegma_.

1. _Spongostylidium streptacanthum_, n. sp.

Both medullary shells spherical, with small, regular, circular pores, twice
as broad as the bars (outer twice as broad as the inner). Spongy cortical
shell enclosing it with dense framework, four times as broad as the outer
medullary shell. Two polar spines very large, four times as long as the
diameter of the outer shell; as broad as the inner medullary shell, with
three dentated, spirally contorted edges. (Very similar to the common
_Spongosphæra streptacantha_ and to _Hexadoridium streptacanthum_, but with
only two opposite spines in one axis.)

_Dimensions._--Diameter of the cortical shell 0.16, of the outer medullary
shell 0.04, inner 0.02; length of the spines 0.7 and more, breadth 0.02.

_Habitat._--Indian Ocean, Cocos Islands, Rabbe.

{151}Family VIII. #STAUROSPHÆRIDA#, Haeckel (Pl. 15).

_Staurosphærida_, Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Sphæroidea# with four radial spines on the surface of the
spherical shell, forming a regular cross, being opposite in pairs in two
axes perpendicular to one another; living solitary (not associated in
colonies).

The family #Staurosphærida# is distinguished from the other #Sphæroidea# by
the possession of four radial spines, which are opposite in pairs in two
perpendicularly crossed axes. By these "two main axes" an equatorial plane
is determined, which approximates them to the #Discoidea#. But in the
latter the shells as well as the central capsule become more or less
flattened, lenticular, or discoidal, whilst in the former they remain
spherical. However, some forms of both groups are very similar, and
inspection from different sides (and mainly from the margin of the
equatorial plane) is required to determine the spherical (not compressed)
shell-form of the Staurosphærida. As a rule the species of this family are
much rarer, and much less numerous, than those of all the other
#Sphæroidea#.

The most simple Staurosphærida are the Staurostylida, with one single
spherical lattice-shell. To this ancestral group all other subfamilies can
be opposed as "Staurosphærida concentrica," since their carapace is
composed of two or more concentric lattice-shells; two in the
Staurolonchida, three in the Stauracontida, four in the Staurocromyida,
five or more in the Staurocaryida. In all these four subfamilies the
concentric shells are simple (not spongy) fenestrated spheres. In a sixth
subfamily, in the Staurodorida, the shell is wholly or partially composed
of irregular spongy wickerwork, with or without a medullary shell in the
centre.

_The Four Radial Spines_ in all Staurosphærida are normally opposed in
pairs in two axes perpendicular one to another, and therefore together form
a rectangular cross. But in many species besides this normal form
individual abnormalities occur, in which the four spines in the equatorial
plane are not quite accurately opposed, so that the four angles between
them are not right angles, but more or less unequal. More rarely also their
position in the equatorial plane is not accurately retained, so that they
are placed in two, three, or four different meridian planes, intersecting
at very small variable angles.

In the greater part of Staurosphærida all four radial spines are quite
equal, and of the same size and form. But in some genera there takes place
a more or less considerable differentiation of the four spines, commonly in
pairs, so that the two opposite spines of each pair are equal, but the
pairs different (_Staurostylus_, _Staurolonchidium_). More rarely also both
spines of one pair become unequal, whilst those of the other pair remain
equal (_Stauroxiphos_). Very rarely all four spines assume a different size
or form.

{152}_Synopsis of the Genera of Staurosphærida._

I. Subfamily {All four spines equal, 60. _Staurosphæra_.
Staurostylida. {
(Shell one single {Four spines different
lattice-sphere.) { in pairs, 61. _Staurostylus_.
{
{One spine larger than the
{ three others, 62. _Stylostaurus_.

II. Subfamily {All four spines {Simple, 63. _Staurolonche_.
Staurolonchida. { equal, {
(Shell with { {Branched, 64. _Staurancistra_.
two concentric {
lattice-spheres.) {Four spines different 65.
{ in pairs, _Staurolonchidium_.
{
{One spine larger than
{ the three others, 66. _Stauroxiphos_.

III. Subfamily }All four spines equal,
Stauracontida. } simple, 67. _Stauracontium_.
(Shell with three }
concentric spheres.) }

IV. Subfamily }All four spines {Simple, 68. _Staurocromyum_.
Staurocromyida. } equal, {
(Shell with four } {Branched, 69. _Cromyostaurus_.
concentric spheres.) }

V. Subfamily }All four spines equal, 70. _Staurocaryum_.
Staurocaryida. }
(Shell with five }
or more concentric }
spheres.) }

VI. Subfamily }Solid spongy sphere
Staurodorida. } without medullary shell, 71. _Staurodoras_.
(Shell a spongy }
sphere.) }

Subfamily STAUROSTYLIDA,[82] Haeckel, Prodromus, 1881, pp. 449, 450.

_Definition._--#Staurosphærida# with one single spherical lattice-shell.

Genus 60. _Staurosphæra_,[83] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Staurosphærida# with a single lattice-sphere and four
crossed equal spines.

The genus _Staurosphæra_ may be regarded as the common ancestral form of
this subfamily, since it represents their most simple and primitive form.
From the surface of the simple lattice-sphere, enclosing the central
capsule, arise four equal, simple, radial spines, opposite in pairs in two
diameters, perpendicular one to another. _Staurosphæra_ may be derived
phylogenetically either from _Cenosphæra_ by production of the four spines,
or from _Hexastylus_ by reduction of two opposite spines.

{153}Subgenus 1. _Staurosphærantha_, Haeckel.

_Definition._--Pores regular, all of nearly equal size and similar form;
surface smooth.

1. _Staurosphæra cruciata_, n. sp.

Shell thin walled, smooth, with regular, hexagonal pores, four times as
broad as the bars; ten to twelve on the quadrant. Four crossed radial
spines three-sided pyramidal, as long as the diameter of the shell, as
broad at the base as one pore (very similar to _Hexastylus phænaxonius_,
Pl. 21, fig. 3, but with only four spines).

_Dimensions._--Diameter of the shell 0.12, of the pores 0.008, bars 0.002.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

2. _Staurosphæra christiana_, n. sp.

Shell thick walled, smooth, with regular, circular, hexagonally framed
pores, three times as broad as the bars; six to eight on the quadrant. Four
crossed spines six-sided pyramidal, as long as the radius, as broad as one
mesh.

_Dimensions._--Diameter of the shell 0.13, pores 0.012, bars 0.004.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

3. _Staurosphæra johannis_, n. sp.

Shell thick walled, smooth, with regular, circular pores, four times as
broad as the bars; five to six on the quadrant. Four spines six-sided
pyramidal, half as long as the radius, as broad as one mesh.

_Dimensions._--Diameter of the shell 0.15, pores 0.02, bars 0.005.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

4. _Staurosphæra pauli_, n. sp.

Shell very thick walled, smooth, with regular, circular, double-edged
pores, four times as broad as the bars; eight to ten on the quadrant. Four
spines conical, as long as the radius, as broad as one mesh.

_Dimensions._--Diameter of the shell 0.15, pores 0.016, bars 0.004.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

5. _Staurosphæra petri_, n. sp.

Shell thin walled, smooth, with regular, circular pores, three times as
broad as the bars; twenty to twenty-two on the quadrant. Four spines
conical, half as long as the radius, as broad as one mesh.

_Dimensions._--Diameter of the shell 0.22, pores 0.006, bars 0.002.

_Habitat._--South Atlantic, Station 325, depth 2650 fathoms; also fossil in
Barbados.

{154}6. _Staurosphæra jacobi_, n. sp.

Shell thick walled, smooth, with regular, circular pores, twice as broad as
the bars; eleven to twelve on the quadrant. Four spines cylindrical, three
times as long as the radius, three times as broad as one mesh.

_Dimensions._--Diameter of the shell 0.1, pores 0.006, bars 0.003.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

7. _Staurosphæra simonis_, n. sp.

_Haliomma_ with four spines, Bury, 1862, Polycystins of Barbados, pl. iv.
fig. 4.

Shell thick walled, smooth, with regular, circular pores, of the same
breadth as the bars; eight to ten on the quadrant. Four spines cylindrical,
twice as long as the radius, five times as broad as one mesh.

_Dimensions._--Diameter of the sphere 0.1, pores and bars 0.003.

_Habitat._--Fossil in Barbados.

Subgenus 2. _Staurosphærella_, Haeckel.

_Definition._--Pores regular, all of nearly equal size and similar form;
surface covered with by-spines or accessory thorns.

8. _Staurosphæra philippi_, n. sp. (Pl. 15, fig. 6).

Shell thin walled, covered with bristle-shaped by-spines, as long as the
radius. Pores regular, circular, twice as broad as the bars; six to eight
on the quadrant. Four main spines cylindrical, five to ten times as long as
the radius, as broad as one mesh.

_Dimensions._--Diameter of the shell 0.08, pores 0.006, bars 0.003.

_Habitat._--Central Pacific, Station 272, surface.

9. _Staurosphæra andreæ_, n. sp.

Shell thick walled, with regular, circular, hexagonally framed pores, three
times as broad as the bars; nine to ten on the quadrant. From each
hexagon-corner arises a bristle-shaped by-spine, half as long as the
radius. Four main spines three-sided pyramidal, with spirally contorted
edges, as long as the radius (very similar to _Hexastylus solonis_, Pl. 21,
fig. 11, but with only four spines).

_Dimensions._--Diameter of the shell 0.16, pores 0.01, bars 0.003.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

{155}10. _Staurosphæra thomæ_.

Shell thick walled, with regular, circular pores, twice as broad as the
bars; twelve to fourteen on the quadrant; surface covered with short
conical by-spines. Four main spines conical, twice as long as the radius,
twice as broad as one mesh.

_Dimensions._--Diameter of the sphere 0.2, pores 0.012, bars 0.006.

_Habitat._--South Atlantic, Station 332, depth, 2200 fathoms.

Subgenus 3. _Staurosphærissa_, Haeckel.

_Definition._--Pores irregular, of different size or form; surface smooth.

11. _Staurosphæra judæ_, n. sp.

Shell thin walled, smooth, with irregular, polygonal pores, twice to four
times as broad as the bars; six to ten on the quadrant. Four main spines
three-sided pyramidal, as long as the radius.

_Dimensions._--Diameter of the sphere 0.15, pores 0.006 to 0.012, bars
0.003.

_Habitat._--Central Pacific, Station 273, depth, 2350 fathoms.

12. _Staurosphæra crassa_, Dunikowski.

_Staurosphæra crassa_, 1882, Denkschr. d. k. Akad. d. Wiss. Wien, Bd.
xlv. p. 27, Taf. v. figs. 52-55.

Shell thick walled, smooth, with irregular, roundish pores, scarcely
broader than the bars; eight to ten on the quadrant. Four spines
three-sided pyramidal, nearly as long as the shell diameter.

_Dimensions._--Diameter of the sphere 0.19, pores and bars 0.015.

_Habitat._--Fossil in the Alpine Lias (Schafberg near Salzburg,
Dunikowski).

13. _Staurosphæra apostolorum_, Haeckel.

? _Cenosphæra megapora_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 66, Taf. iii. fig. 1.

? _Cenosphæra micropora_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 66, Taf. iii. fig. 2.

Shell thin walled, smooth, with large, irregular, roundish pores, twice to
six times as broad as the bars; four to six on the quadrant. Four spines
conical, very stout, about as long as the shell diameter, often more or
less irregularly disposed.

_Dimensions._--Diameter of the sphere 0.12 to 0.2, pores 0.01 to 0.03, bars
0.005.

_Habitat._--Fossil in Barbados.

{156}Subgenus 4. _Staurosphæromma_, Haeckel.

_Definition._--Pores irregular, of different size or form; surface covered
with by-spines or accessory thorns.

14. _Staurosphæra bartholomæi_, n. sp.

Shell thin walled, with irregular, polygonal pores, three times as broad as
the bars; six to eight on the quadrant; surface covered with short
bristle-shaped by-spines. Four main spines three-sided pyramidal, twice as
long as the radius.

_Dimensions._--Diameter of the shell 0.14, pores 0.008 to 0.016, bars 0.003
to 0.005.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

15. _Staurosphæra thaddæi_, n. sp.

Shell thick walled, with irregular, roundish, polygonally framed pores,
twice to three times as broad as the bars; five to seven on the quadrant;
surface covered with bristle-shaped spines, half as long as the radius.
Four main spines pyramidal, as long as the radius.

_Dimensions._--Diameter of the shell 0.12, pores 006 to 0.012, bars 0.004.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms; also fossil
in Barbados.

16. _Staurosphæra matthæi_, n. sp.

Shell thick walled, with irregular, roundish pores, twice to five times as
broad as the bars; ten to twelve on the quadrant; surface covered with
short conical thorns or by-spines. Four main spines conical, as long as the
radius.

_Dimensions._--Diameter of the shell 0.15, pores 0.004 to 0.01, bars 0.002.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

Genus 61. _Staurostylus_,[84] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Staurosphærida# with one single lattice-sphere and four
crossed spines which are arranged in opposite pairs, one pair opposite
being larger than the other.

The genus _Staurostylus_ has been developed from _Staurosphæra_ by
differentiation of the four crossed spines in pairs; two opposite spines
growing much more strongly than the other two.

1. _Staurostylus græcus_, n. sp.

Shell thick walled, smooth, with regular, circular, hexagonally framed
pores, twice as broad as the bars; six to eight on the quadrant. Spines
three-sided prismatic, pointed, as broad as {157}one mesh; two opposite
larger spines as long as the diameter of the shell, two smaller only as
long as the radius.

_Dimensions._--Diameter of the shell 0.15, pores 0.01, bars 0.005; length
of the major spines 0.16, minor 0.08.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

2. _Staurostylus latinus_, n. sp.

Shell thin walled, smooth, with regular, circular, double-edged pores,
three times as broad as the bars; twelve to fourteen on the quadrant.
Spines conical, twice as broad at the base as one mesh; two opposite larger
spines as long as the radius of the shell, two smaller only one-third as
long.

_Dimensions._--Diameter of the shell 0.24, pores 0.012, bars 0.004; length
of the major spines 0.12, minor 0.04.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

3. _Staurostylus germanicus_, n. sp.

Shell thick walled, covered with bristle-shaped by-spines, half as long as
the radius. Pores regular, circular, hexagonally framed, twice as broad as
the bars; eight to ten on the quadrant. From each hexagon-corner arises one
short by-spine. Four main spines three-sided pyramidal, as broad at the
base as one mesh; two opposite larger spines one and a half times as long
as the radius, two smaller two-thirds as long.

_Dimensions._--Diameter of the shell 0.16, pores 0.008, bars 0.004; length
of the major spines 0.12, minor 0.08.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

Genus 62. _Stylostaurus_,[85] 1881, Prodromus, p. 450.

_Definition._--#Staurosphærida# with one single lattice-sphere and four
crossed spines, one of which is much larger than the other three.

The genus _Stylostaurus_ differs from the preceding in the extraordinary
development of one of the four spines, which is much longer than the other
three; these may be equal or different.

1. _Stylostaurus caudatus_, n. sp. (Pl. 13, fig. 7).

Shell thick walled, smooth, with regular, circular, hexagonally framed
pores, twice as broad as the bars; five to six on the quadrant. Spines
three-sided pyramidal, as broad at the base as {158}one mesh; one of the
spines longer than the shell diameter, the opposite spine nearly as long as
the shell radius; both lateral spines scarcely one-third as long.

_Dimensions._--Diameter of the shell 0.1, pores 0.01, bars 0.005; length of
the major spine 0.14, of the opposite 0.04, of both lateral spines 0.012.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

2. _Stylostaurus gladiatus_, n. sp.

Shell thick walled, smooth, with regular, circular pores, three times as
broad as the bars; nine to ten on the quadrant. Spines three-sided
prismatic, as broad at the base as one mesh; one of the spines longer than
the shell diameter, and much larger than the other three, which are nearly
equal (half as long as the radius).

_Dimensions._--Diameter of the shell 0.2, pores 0.015, bars 0.005; length
of the major spine 0.25, of the three others 0.005.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Subfamily STAUROLONCHIDA,[86] Haeckel, 1881, Prodromus, pp. 449, 451.

_Definition._--#Staurosphærida# with two concentric spherical
lattice-shells.

Genus 63. _Staurolonche_,[87] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Staurosphærida# with two concentric lattice-spheres and
four crossed, equal, simple spines.

The genus _Staurolonche_ may be derived either from _Staurosphæra_ by the
duplication of the lattice-sphere, or from _Carposphæra_ by the production
of four crossed radial spines, lying in one meridional plane, or from
_Hexalonche_ by the reduction of two opposite spines.

Subgenus 1. _Staurolonchantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, and of nearly equal
size and similar form; surface smooth.

1. _Staurolonche hexagona_, n. sp.

_Haliomma hexagonum_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvb., Bd. iv.
fig. 17.

_Haliomma hexagonum_, Haeckel, 1862, Monogr. d. Radiol., p. 434.

Cortical shell thin walled, smooth, three times as broad as the medullary
shell, with regular, hexagonal pores, four times as broad as the bars;
seven to eight on the quadrant. Four spines three-sided pyramidal, somewhat
longer than the radius, as broad at the base as one mesh.

{159}_Dimensions._--Diameter of the outer shell 0.12, inner 0.04; cortical
pores 0.012, bars 0.003; length of the spines 0.08.

_Habitat._--North Atlantic, Station 64, depth 2700 fathoms.

2. _Staurolonche spinozæ_, n. sp.

Cortical shell thick walled, smooth, twice as broad as the medullary shell,
with regular, circular, hexagonally framed pores, three times as broad as
the bars; five to six on the quadrant. Four spines three-sided pyramidal,
half as long as the radius, as broad at the base as one mesh.

_Dimensions._--Diameter of the outer shell 0.16, inner 0.08; cortical pores
0.012, bars 0.004; length of the spines 0.04.

_Habitat._--Central Pacific, Station 271, surface.

3. _Staurolonche aperta_, Haeckel.

_Haliomma apertum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxviii. fig. 5.

Cortical shell thin walled, smooth, three times as broad as the medullary
shell; pores regular, circular, five times as broad as the bars; four to
five on the quadrant. Four spines conical, as long as the radius, as broad
as one mesh.

_Dimensions._--Diameter of the outer shell 0.1, inner 0.03; cortical pores
0.015, bars 0.003; length of the spines 0.05.

_Habitat._--Fossil in Barbados.

4. _Staurolonche brunonis_, n. sp.

Cortical shell thick walled, smooth, twice as broad as the medullary shell,
with regular, circular pores, three times as broad as the bars; eight to
ten on the quadrant. Four spines cylindro-conical, longer than the
diameter, twice as broad as one mesh.

_Dimensions._--Diameter of the outer shell 0.08, inner 0.04; cortical pores
0.006, bars 0.002; length of the spines 0.12.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

5. _Staurolonche pertusa_, n. sp. (Pl. 15, figs. 5, 5_a_).

Cortical shell thin walled, smooth, three times as broad as the medullary
shell, with regular, circular pores, three times as broad as the bars;
eight to ten on the quadrant. Pores of the medullary shell only one-third
as large, also circular. Four spines three-sided prismatic, three times as
long as the radius; each of their three thin wings perforated by a single
row of small pores.

_Dimensions._--Diameter of the outer shell 0.1, inner 0.033; cortical pores
0.01, bars 0.003; length of the spines 0.15.

_Habitat._--North Atlantic, Station 353, surface.

{160}Subgenus 2. _Staurolonchella_, Haeckel.

_Definition._--Pores of the cortical shell regular, and of nearly equal
size and similar form; surface covered with by-spines or thorns.

6. _Staurolonche straussii_, n. sp.

Cortical shell thin walled, four times as broad as the medullary shell, and
covered with numerous bristle-shaped by-spines, half as long as the radius.
Pores regular, circular, hexagonally framed, twice as broad as the bars;
six to eight on the radius. On each hexagonal frame twelve by-spines (six
at the corners, six in the middle between them). Four main spines six-sided
pyramidal, about as long as the radius, twice as broad as one mesh.

_Dimensions._--Diameter of the outer shell 0.16, inner 0.04; cortical pores
0.016, bars 0.008; length of the spines 0.1.

_Habitat._--West Tropical Pacific, Station 225, depth 4475 fathoms.

7. _Staurolonche feuerbachii_, n. sp.

Cortical shell thick walled, five times as broad as the medullary shell,
and covered with short conical by-spines. Pores regular, circular, twice as
broad as the bars; twenty to twenty-two on the quadrant. Four main spines
three-sided prismatic, with pyramidal apex, two to three times as long as
the radius, twice as broad as one mesh.

_Dimensions._--Diameter of the outer shell 0.27, of the inner 0.055;
cortical pores 0.008, bars 0.004; length of the spines 0.3 to 0.4.

_Habitat._--Fossil in Barbados.

8. _Staurolonche moleschottii_, n. sp.

Cortical shell thick walled, three times as broad as the medullary shell,
and covered with short conical by-spines. Pores regular, circular, of the
same breadth as the bars; fourteen to sixteen on the quadrant. Four main
spines conical, as long as the radius, three times as broad at the base as
one mesh.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; cortical pores
and bars 0.004; length of the spines 0.08.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subgenus 3. _Staurolonchissa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface smooth.

9. _Staurolonche holbachii_, n. sp.

Cortical shell thin walled, with smooth surface, three times as broad as
the medullary shell. Pores irregular, polygonal, twice to four times as
broad as the bars. Four main spines three-sided prismatic, longer than the
shell diameter.

{161}_Dimensions._--Diameter of the outer shell 0.12, inner 0.04; cortical
pores 0.006 to 0.012, bars 0.003; length of the spines 0.15.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

10. _Staurolonche gassendii_, n. sp.

Cortical shell thick walled, with smooth surface, four times as broad as
the medullary shell. Pores irregular, roundish, three to five times as
broad as the bars. Four main spines conical, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.05; cortical pores
0.01 to 0.02, bars 0.004; length of the spines 0.1.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

Subgenus 4. _Staurolonchura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface covered with by-spines or thorns.

11. _Staurolonche epicurii_, n. sp.

Cortical shell thick walled, five times as broad as the medullary shell,
and covered with numerous bristle-shaped by-spines (half as long as the
radius). Pores irregular, roundish, polygonally framed, twice to three
times as broad as the bars. Four main spines pyramidal, as long as the
radius.

_Dimensions._--Diameter of the outer shell 0.22, inner 0.044; cortical
pores 0.02 to 0.04, bars 0.012; length of the spines 0.1.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

12. _Staurolonche lucretii_, n. sp.

Cortical shell thick-walled, three times as broad as the medullary shell,
and covered with numerous short, conical thorns. Pores irregular, roundish,
twice to four times as broad as the bars. Four main spines conical, half as
long as the radius.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; cortical pores
0.015 to 0.03, bars 0.008; length of the spines 0.04.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Genus 64. _Staurancistra_,[88] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Staurosphærida# with two concentric lattice-spheres and
four crossed, equal, branched spines.

{162}The genus _Staurancistra_ differs from its ancestral form,
_Staurolonche_, in the ramification of the four crossed spines.

1. _Staurancistra quadricuspis_, n. sp.

Cortical shell thin walled, with rough, thorny surface, and irregular,
roundish pores, twice to four times as broad as the bars; six to eight on
the quadrant. Medullary shell one-third as large, with regular, circular
pores; connected with the cortical shell by four crossed radial beams.
These are prolonged outside into four strong three-sided prismatic spines,
nearly as long as the shell diameter, each having three curved branches
below the distal end. (Similar to _Hexancistra quadricuspis_, Pl. 22, fig.
11, but with only four spines.)

_Dimensions._--Diameter of the cortical shell 0.16, of the medullary shell
0.05; pores of the former 0.06 to 0.012, bars 0.003; length of the spines
0.14, breadth 0.012.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Genus 65. _Staurolonchidium_,[89] n. gen.

_Definition._--#Staurosphærida# with two concentric lattice-spheres and
four crossed simple spines which are disposed in two different pairs, two
opposite being larger than the other two.

The genus _Staurolonchidium_ has arisen from _Staurolonche_ by the stronger
growth of the two opposite spines, the other two remaining stationary; both
spines of each pair equal.

1. _Staurolonchidium artioscelides_, n. sp.

Cortical shell thin walled, with smooth surface, three times as broad as
the medullary shell. Pores regular, circular, hexagonally framed, twice as
broad as the bars; eight to ten on the quadrant. Four radial spines
three-sided pyramidal, as broad as one mesh; two opposite larger spines
three times as long as the two smaller, which are about equal to the
radius.

_Dimensions._--Diameter of the cortical shell 0.16, medullary shell 0.05;
pores of the former 0.012, bars 0.006; length of the major spines 0.24,
minor 0.08.

_Habitat._--Central Pacific, Station 265, surface.

2. _Staurolonchidium perspicuum_, Haeckel.

_Haliomma perspicuum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxix. fig. 1.

Cortical shell thin walled, covered with small conical by-spines, five
times as broad as the medullary shell. Pores regular, circular, eight times
as broad as the bars; three to four on the quadrant. Four radial spines
three-sided pyramidal, scarcely one-third as broad as one mesh; two
opposite larger spines four times as long as the two smaller, which are
about equal to half the radius.

{163}_Dimensions._--Diameter of the cortical shell 0.12, medullary shell
0.025; pores of the former 0.03, bars 0.004; length of the major spines
0.12, minor 0.03.

_Habitat._--Fossil in Barbados.

Genus 66. _Stauroxiphos_,[90] n. gen.

_Definition._--#Staurosphærida# with two concentric lattice-spheres and
four crossed simple spines, one of which is much larger than the other
three.

The genus _Stauroxiphos_ differs from its ancestral form, _Staurolonche_,
in the greater development of one single spine, and exhibits therefore the
same relation to it that _Stylostaurus_ bears to _Staurosphæra_.

1. _Stauroxiphos gladius_, n. sp. (Pl. 15, fig. 7).

Cortical shell thick walled, smooth, three times as broad as the medullary
shell. Pores of the outer shell regular, circular, regularly six-lobed on
the outer opening, four times as broad as the bars; about six on the
quadrant. Pores of the inner shell only one-third as large, simple,
circular. Three of the four radial spines of nearly equal size,
pommel-shaped, with three prominent, dentated wings, somewhat shorter than
the shell radius and about half as broad as long; the fourth spine much
larger, sword-like, about three times as long as the shell radius.

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.04; pores
of the former 0.01, bars 0.0025; pores of the latter 0.003, bars 0.001;
length of the major spine 0.18, of the three minor 0.04.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

Subfamily STAURACONTIDA,[91] Haeckel, 1881, Prodromus, p. 52.

_Definition._--#Staurosphærida# with three concentric spherical
lattice-shells.

Genus 67. _Stauracontium_,[92] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Staurosphærida# with three concentric lattice-spheres and
four crossed, equal, simple spines.

The genus _Stauracontium_ differs from its probable ancestral form,
_Staurolonche_, in the duplication of the cortical shell.

Subgenus 1. _Stauracontarium_, Haeckel.

_Definition._--Pores of the cortical shell regular, all of nearly equal
size and similar form; surface smooth.

{164}1. _Stauracontium cruciferum_, n. sp.

Cortical shell thin walled, smooth, with regular, hexagonal pores, four
times as broad as the bars; eight to ten on the quadrant. Radial proportion
of the three spheres = 1 : 3 : 9. Connecting radial beams between them six
(opposite in pairs in the three dimensive axes), but only four of them are
prolonged outside into four stout three-sided prismatic spines, lying in
one equatorial plane, as long as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.22, middle 0.07, inner 0.025;
cortical pores 0.012, bars 0.003; length of the spines 0.2.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

2. _Stauracontium tetracanthum_, Haeckel.

_Haliomma tetracanthum_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 295, Taf. x. figs. 7, 8.

Cortical shell thin walled, smooth, with regular, circular pores, three
times as broad as the bars; five to six on the quadrant. Radial proportion
of the three spheres = 1 : 3 : 12. Radial spines three-sided pyramidal,
about as long as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.03, inner 0.01;
cortical pores 0.01, bars 0.003; length of the spines 0.1.

_Habitat._--Indian Ocean, Zanzibar, depth 2200 fathoms, Pullen.

3. _Stauracontium tetracontium_, Haeckel.

_Actinomma tetracanthum_, Stöhr, 1880, Palæontogr. 26, p. 91, Taf. ii.
fig. 6.

Cortical shell thin walled, rough, with regular, circular pores, of the
same breadth as the bars; six to seven on the quadrant. Radial proportion
of the three spheres = 1 : 2.5 : 8. Radial spines three-sided pyramidal,
nearly as long as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.1, middle 0.03, inner 0.013;
cortical pores and bars 0.006; length of the spines 0.08.

_Habitat._--Fossil in Tertiary rocks of Sicily; Grotte, Caltanisetta.

Subgenus 2. _Stauracontellium_, Haeckel.

_Definition._--Pores of the cortical shell regular, and of nearly equal
size and similar form; surface covered with numerous small thorns or
by-spines.

4. _Stauracontium daturæforme_, Haeckel.

_Actinomma daturæforme_, Stöhr, 1880, Palæontogr. 26, p. 93, Taf. ii.
fig. 13.

Cortical shell thick walled, covered with short thorns, and with regular,
hexagonal pores, five times as broad as the bars; six to seven on the
quadrant. Radial proportion of the three spheres = 1 : 3 : 3.5. Radial
spines six-sided pyramidal, about half as long as the radius and one-fourth
as broad.

{165}_Dimensions._--Diameter of the outer shell 0.15, middle 0.13, inner
0.04; cortical pores 0.017, bars 0.003; length of the spines 0.03.

_Habitat._--Fossil in Tertiary rocks of Sicily; Grotte, Stöhr.

5. _Stauracontium sparganium_, n. sp.

Cortical shell thick walled, covered with very numerous, short, conical
spines, and with regular, circular pores, five times as broad as the bars;
sixteen to eighteen on the quadrant. Radial proportion of the three spheres
= 2 : 3 : 9. Radial main spines three-sided prismatic, as long as the
radius or longer.

_Dimensions._--Diameter of the outer shell 0.275, middle 0.09, inner 0.06;
cortical pores 0.01, bars 0.002; length of the spines 0.15.

_Habitat._--Fossil in Barbados.

Subgenus 3. _Stauracontidium_, Haeckel.

_Definition._--Pores irregular, of different size or form; surface smooth.

6. _Stauracontium antarcticum_, n. sp.

Cortical shell thin walled, smooth. Pores irregular, roundish, two to four
times as broad as the bars. Radial proportion of the three spheres =
1 : 2 : 8. Radial spines conical, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.05, inner 0.025;
cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.1.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Subgenus 4. _Stauracontonium_, Haeckel.

_Definition._--Pores irregular, of different size or form; surface covered
with thorns or by-spines.

7. _Stauracontium setosum_, n. sp.

Cortical shell thin walled, covered with thin bristle-shaped by-spines,
half as long as the radius. Pores irregular, polygonal, three to five times
as broad as the bars. Proportion of the three spheres = 1 : 2 : 6. Radial
main spines pyramidal, nearly as long as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.15, middle 0.05, inner 0.025;
cortical pores 0.01 to 0.015, bars 0.003; length of the spines 0.12.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

8. _Stauracontium papillosum_, n. sp.

Cortical shell thick walled, covered with short conical papillæ or
by-spines. Pores irregular, roundish, two to four times as broad as the
bars. Proportion of the three spheres = 1 : 3 : 12. Radial main spines
conical, as long as the radius.

{166}_Dimensions._--Diameter of the outer shell 0.24, middle 0.06, inner
0.02; cortical pores 0.007 to 0.015, bars 0.004; length of the spines 0.12.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

Subfamily STAUROCROMYIDA,[93] Haeckel, 1881, Prodromus, pp. 449, 453.

_Definition._--#Staurosphærida# with four concentric spherical
lattice-shells.

Genus 68. _Staurocromyum_,[94] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Staurosphærida# with four concentric lattice-spheres and
four crossed, equal, simple spines.

The genus _Staurocromyum_ has arisen probably from _Stauracontium_ by
duplication of the cortical shell, two concentric shells lying within, two
others outside the central capsule.

1. _Staurocromyum quadruplex_, n. sp.

Radial proportion of the four spheres = 1 : 2 : 8 : 10. Both medullary
shells with very small, regular, circular pores. Inner cortical shell with
regular, circular, hexagonally-framed pores, twice as broad as the bars;
from each hexagon-corner arises a small by-spine, and these, connected by
tangential branches at equal distances from the centre, form the delicate
outer cortical shell, with spiny surface. Four main spines three-sided
pyramidal, as long as the shell radius.

_Dimensions._--Diameter of the four shells--(A) 0.25, (B) 0.2, (C) 0.05,
(D) 0.025; length of the spines 0.12.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

2. _Staurocromyum quadrispinum_, n. sp.

Radial proportion of the four spheres = 1 : 3 : 9 : 12. Both medullary
shells with small, regular, circular pores. Both cortical shells with
irregular, roundish pores. Surface covered with short, conical by-spines.
Four main spines cylindro-conical, somewhat longer than the shell diameter.

_Dimensions._--Diameter of the four shells--(A) 0.24, (B) 0.18, (C) 0.06,
(D) 0.02.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Genus 69. _Cromyostaurus_,[95] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Staurosphærida# with four concentric lattice-spheres and
four crossed, equal, branched spines.

{167}The genus _Cromyostaurus_ differs from the preceding _Staurocromyum_,
its ancestral form, in the ramification of the four crossed spines.

1. _Cromyostaurus verticillatus_, n. sp.

Radial proportion of the four spheres = 1 : 3 : 11 : 13. Both medullary
shells with small, regular, circular pores; inner cortical shell with
regular, hexagonal pores; from the hexagon-corners arise small, radial
by-spines, which at equal distances from the centre send out forked
tangential branches, three from each spine, and by communication of these
form the outer, delicate, cortical shell. Four main spines nearly as long
as the shell diameter, three-sided prismatic, with four to six verticils of
ramified lateral branches, each verticil composed of three forked branches,
which ramify again.

_Dimensions._--Diameter of the four shells--(A) 0.26, (B) 0.22, (C) 0.06,
(D) 0.02; length of the spines 0.24.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subfamily STAUROCARYIDA,[96] Haeckel, 1881, Prodromus, pp. 449, 454.

_Definition._--#Staurosphærida# with five or more concentric spherical
lattice-shells.

Genus 70. _Staurocaryum_,[97] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Staurosphærida# with five or more concentric
lattice-spheres and four crossed, equal spines.

The genus _Staurocaryum_ has arisen from the preceding _Staurocromyum_ by
the further multiplication of the concentric spheres; in the only observed
form there are six, at nearly equal distances apart.

1. _Staurocaryum arborescens_, n. sp. (Pl. 15, fig. 8).

Shell composed of six concentric latticed spheres, at nearly equal
distances apart, and with somewhat regular, circular pores, the size of
which gradually increases from the first to the sixth shell. The surface of
the outermost shell is densely covered with numerous arborescent by-spines,
which bifurcate from three to four times, and are three-sided pyramidal at
the base, and twice as long as the distance between each two shells. The
six shells are connected only by four crossed, conical, radial beams, which
increase in diameter from the centre, and are prolonged outside into very
stout cylindrical, radial spines, irregularly covered with small thorns and
forked ramules, and nearly as long as the shell diameter. Only a single
specimen was observed.

{168}_Dimensions._--Diameter of the whole shell 0.22; distance between each
two shells 0.02; length of the by-spines 0.05, of the main spines 0.2.

_Habitat._--Indian Ocean, Cocos Islands, surface, Rabbe.

Subfamily STAURODORIDA,[98] Haeckel, 1881, Prodromus, pp. 449, 455.

_Definition._--#Staurosphærida# with spongy, spherical shell (with or
without enclosed concentric lattice-shells).

Genus 71. _Staurodoras_,[99] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Staurosphærida# with solid, spongy, spherical shell and
four crossed simple spines.

The genus _Staurodoras_ may be developed from _Styptosphæra_ by the
production of four crossed spines on the simple, spongy, spherical shell,
which is composed of looser or denser irregular wicker-work, without
enclosed medullary shell.

1. _Staurodoras spongosphæra_, n. sp.

Four crossed spines, two to three times as long as the diameter of the
spongy sphere, three-sided prismatic, with three dentated and spirally
contorted edges. (Form of _Spongosphæra streptacantha_, but without
medullary shell and with four equal spines, crossed regularly at right
angles.)

_Dimensions._--Diameter of the sphere 0.25; length of the spines 0.4 to
0.7.

_Habitat._--Central Pacific, Station 271, surface.

2. _Staurodoras mojsisovicsi_, Dunikowski.

_Staurodoras mojsisovicsi_, Dunikowski, 1882, Denkschr. d. k. Akad. d.
Wiss. Wien, Bd. xlv. p. 28, Taf. v. fig. 56.

Four crossed spines, one and a half times as long as the diameter of the
spongy sphere, three-sided pyramidal, with three smooth edges. (What
Dunikowski describes as "inner canals" of the spines are their edges.)

_Dimensions._--Diameter of the sphere 0.14; length of the spines 0.18 to
0.2. basal breadth 0.04.

_Habitat._--Fossil in the Alpine Lias (Schafberg near Salzburg).

3. _Staurodoras liassica_, Dunikowski.

_Staurodoras liassica_, Dunikowski, 1882, Denkschr. d. k. Akad. d. Wiss.
Wien, Bd. xlv. p. 28, Taf. v. fig. 57.

Four crossed spines, shorter than the diameter of the spongy sphere,
conical or pyramidal (?).

_Dimensions._--Diameter of the sphere 0.19; length of the spine 0.13, basal
breadth 0.045.

_Habitat._--Fossil in the Alpine Lias (Schafberg near Salzburg).

{169}4. _Staurodoras wandae_, Dunikowski.

_Staurodoras wandae_, Dunikowski, 1882, Denkschr. d. k. Akad. d. Wiss.
Wien, Bd. xlv. p. 28, Taf. v. fig. 58.

Four crossed spines shorter than the radius of the spongy sphere, conical.
(May be the young form of the preceding species.)

_Dimensions._--Diameter of the sphere 0.14; length of the spines 0.06,
basal breadth 0.05.

_Habitat._--Fossil in the Alpine Lias (Schafberg near Salzburg).

Family IX. #CUBOSPHÆRIDA#, Haeckel (Pls. 21-25).

_Cubosphærida_, Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Sphæroidea# with six radial spines on the surface of the
spherical shell, opposite in pairs in the three dimensive axes,
perpendicular one to another; living solitary (not associated in colonies).

The family #Cubosphærida# is distinguished from the other #Sphæroidea# by
the possession of six radial spines, which are opposite in pairs in three
different axes, one perpendicular to the other two. These three axes are
the typical "dimensive axes," which are more or less differentiated in the
#Larcoidea#. But in these latter the shell itself and the enclosed central
capsule become affected by the unequal growth in the three axes, whilst in
the former the capsule constantly, and commonly also the shell, remains
spherical. Sometimes the shell assumes the form of a regular octahedron,
from the six corners of which arise the six radial spines, indicating its
three axes.

The most simple Cubosphærida are the Hexastylida, with one single,
spherical lattice-shell. To this ancestral group all other subfamilies can
be opposed as "Cubosphærida concentrica," as their carapace is composed of
two or more concentric lattice-shells--two in the Hexalonchida, three in
the Hexacontida, four in the Hexacromyida, five or more in the Hexacaryida.
In all these four subfamilies the concentric shells are simple (not
spongy), fenestrated spheres. In a sixth subfamily, in the Hexadorida, the
shell is wholly or partially composed of irregular, spongy wicker-work or
loose reticulations, with or without a medullary shell in the centre.

The _Six Radial Spines_ of the Cubosphærida are normally opposite in pairs
in the three dimensive axes, each of which is perpendicular to the other
two. But in many species besides this normal form occur individual
abnormalities, in which the six spines are not quite accurately opposed,
but more or less divergent; and often also the three dimensive planes
(determined each by two axes) are not quite regular, but more or less
uneven. More rarely the six spines appear disposed in quite an irregular
manner.

In the greater part of the Cubosphærida all six spines are quite equal, of
the same size and form. But in some genera a more or less considerable
differentiation takes place, so {170}that two pairs or all three pairs of
spines become different; very rarely, also both spines of one pair become
unequal (probably only an individual abnormality). Those variations
correspond to the differences between the crystalline systems. The common
Cubosphærida, with three equal spine-pairs, correspond to the regular or
cubic system, with three equal axes. The rarer forms (_Hexastylarium_,
_Hexaloncharium_, _Hexacontarium_) exhibit two equal pairs and one
different pair; they correspond to the quadratic system, with three
perpendicular axes, two of which are equal, the third unequal. Still more
rare are those forms (_Hexastylidium_, _Hexalonchidium_), in which all
three pairs of spines are different, corresponding to the three unequal
axes of the rhombic crystalline system.

_Synopsis of the Genera of Cubosphærida._

{All six radial spines simple,
I. Subfamily { of equal size, 72. _Hexastylus_.
Hexastylida. {
(Shell one {Six spines of {Two pairs equal,
simple { different sizes,{ one pair
latticed { all six simple. { different, 73. _Hexastylarium_.
sphere.) { {
{ {All three pairs
{ { different, 74. _Hexastylidium_.

{All six radial {Spines simple, 75. _Hexalonche_.
{ spines of {
II. Subfamily { equal size. {Spines branched, 76. _Hexancistra_.
Hexalonchida. {
(Shell composed {Six spines {Two pairs equal,
of two { simple, in { one pair
concentric { pairs of { different, 77. _Hexaloncharium_.
latticed { different sizes.
spheres.) { {All three pairs
{ { different, 78. _Hexalonchidium_.

{All six radial {Spines simple, 79. _Hexacontium_.
III. Subfamily { spines of {
Hexacontida. { equal size. {Spines branched, 80. _Hexadendron_.
(Shell composed {
of three {Six spines Two pairs equal,
spheres.) { simple, of one pair
{ different sizes. different, 81. _Hexacontarium_.

IV. Subfamily }All six radial Spines simple,
Hexacromyida. } spines of not branched, 82. _Hexacromyum_.
(Four concentric} equal size.
spheres.) }

V. Subfamily }All six radial {Spines simple, 83. _Cubosphæra_.
Hexacaryida. } spines of {
(Five or more } equal size. {Spines branched, 84. _Hexacaryum_.
spheres.) }

{Without latticed
{ medullary
VI. Subfamily }All six radial { shell, 85. _Cubaxonium_.
Hexadorida. } spines of {
(Shell a spongy } equal size, {With one single
sphere, with } simple (not { medullary
or without an } branched). { shell, 86. _Hexadoras_.
enclosed } {
central } {With two
medullary } { medullary
shell.) } { shells, 87. _Hexadoridium_.

{171}Subfamily HEXASTYLIDA,[100] Haeckel, 1881, Prodromus, pp. 449, 450.

_Definition._--#Cubosphærida# with one single spherical lattice-shell.

Genus 72. _Hexastylus_,[101] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Cubosphærida# with one simple lattice-sphere and six simple
spines of equal size.

The genus _Hexalonche_ is the most simple form of all Cubosphærida, and may
be regarded as the common ancestral form of this family. It can be derived
phylogenetically from _Cenosphæra_, by development of six radial spines on
the surface of the simple spherical lattice-shell. These six simple spines
are of equal size and opposite in pairs in the three dimensive axes,
corresponding to the three equal axes of a cubic crystal.

Subgenus 1. _Hexastylanthus_, Haeckel.

_Definition._--Pores regular or subregular, of nearly equal size and form;
surface of the cortical shell smooth, without radial by-spines (other than
the six main spines).

1. _Hexastylus phænaxonius_, n. sp. (Pl. 21, fig. 3).

Shell thin walled, with smooth surface. Pores subregular, hexagonal, five
to six times as broad as the bars; nine to ten on the radius. Six spines
triangular pyramidal, as long as the radius of the shell, as broad at the
base as one pore.

_Dimensions._--Diameter of the shell 0.13, pores 0.008 to 0.01, bars
0.0015; length of the spines 0.07, basal breadth 0.008 to 0.012.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Hexastylus sapientum_, n. sp.

Shell thin walled, with smooth surface. Pores regular, hexagonal, eight to
ten times as broad as the bars; six to seven on the radius. Six spines
bristle-shaped, longer than the diameter of the shell. (Lattice-work and
spines similar to those of _Heliosphæra actinota_, Monogr. d. Radiol., Taf.
ix. fig. 3.)

_Dimensions._--Diameter of the shell 0.15, pores 0.016, bars 0.002; length
of the spines 0.2, breadth 0.002.

_Habitat._--North Atlantic, Station 354, surface.

{172}3. _Hexastylus thaletis_, n. sp. (Pl. 21, fig. 4).

Shell thin walled, with smooth surface. Pores subregular, circular,
hexagonally framed, somewhat funnel-shaped, of the same breadth as the
bars; eight to nine on the radius. Six spines triangular pyramidal, with
prominent edges, as long as the radius, three to four times as broad at the
base as one pore.

_Dimensions._--Diameter of the shell 0.1, pores and bars 0.005; length of
the spines 0.05, basal breadth 0.02.

_Habitat._--Tropical Pacific, Station 225, depth 4575 fathoms.

4. _Hexastylus favosus_, n. sp.

Shell thick walled, with smooth surface. Pores regular, circular,
hexagonally framed, deep funnel-shaped, of the same breadth as the bars;
six to seven on the radius. Six spines triangular pyramidal, as long as the
diameter, twice as broad at the base as one pore.

_Dimensions._--Diameter of the shell 0.12, pores and bars 0.01; length of
the spines 0.12, basal breadth 0.02.

_Habitat._--South Atlantic, Station 320, surface.

5. _Hexastylus longissimus_, n. sp.

Shell thick walled, with smooth surface. Pores regular, circular,
hexagonally framed, of the same breadth as the bars; five to six on the
radius. Six spines triangular prismatic, extremely elongated, ten to twenty
times as long as the diameter of the shell, twice as broad as one pore.

_Dimensions._--Diameter of the shell 0.08, pores and bars 0.008; length of
the spines 1.0 to 1.5, breadth 0.015.

_Habitat._--Tropical Atlantic, Station 347, surface.

6. _Hexastylus minimus_, n. sp. (Pl. 21, fig. 5).

Shell thin walled, with smooth surface. Pores subregular, circular, twice
as broad as the bars; five to six on the radius. Six spines triangular
pyramidal, scarcely as long as the radius, at the base half as broad as
long.

_Dimensions._--Diameter of the shell 0.05, pores 0.004, bars 0.002; length
of the spines 0.02, basal breadth 0.01.

_Habitat._--Tropical Pacific, Station 225, depth 4575 fathoms.

7. _Hexastylus biantis_, n. sp.

? _Haliphormis hexacantha_, Ehrenberg, 1872, L. N. 24, Taf. x. fig. 6.

Shell thin walled, smooth. Pores regular, circular, four times as broad as
the bars; eight to nine on the radius. Six spines conical, as long as the
radius, at the base as broad as one pore.

{173}_Dimensions._--Diameter of the shell 0.12, pores 0.012, bars 0.003;
length of the spines 0.06, basal breadth 0.012.

_Habitat._--Indian Ocean, Madagascar, surface, Rabbe.

8. _Hexastylus pittaci_, n. sp.

Shell thin walled, smooth. Pores regular, circular, ten to twelve times as
broad as the bars; six to seven on the radius. Six spines triangular
pyramidal, as long as the radius, as broad as one pore.

_Dimensions._--Diameter of the shell 0.12, pores 0.02, bars 0.002; length
of the spines 0.06, basal breadth 0.02.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

9. _Hexastylus maximus_, n. sp.

Shell thick walled, smooth. Pores regular, circular, twice as broad as the
bars; twelve to sixteen on the radius. Six spines six-sided pyramidal, half
as long as the radius, three times as broad as one pore.

_Dimensions._--Diameter of the shell 0.3, pores 0.01, bars 0.005; length of
the spines 0.08, basal breadth 0.03.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

10. _Hexastylus periandri_, n. sp.

Shell thick walled, smooth. Pores regular, circular, double-edged, four
times as broad as the bars; six to seven on the radius. Six spines conical,
as long as the radius, twice as broad as one pore.

_Dimensions._--Diameter of the shell 0.2, pores 0.012, bars 0.003; length
of the spines 0.1, basal breadth 0.025.

_Habitat._--Tropical Atlantic, Station 335, depth 1425 fathoms.

Subgenus 2. _Hexastylettus_, Haeckel.

_Definition._--Pores regular or subregular, of nearly equal size and form;
surface of the spherical shell spiny, covered with numerous conical or
bristle-shaped radial by-spines.

11. _Hexastylus solonis_, n. sp. (Pl. 21, fig. 11).

Shell thick walled, covered with numerous bristle-shaped radial spines
(one-third to one-half as long as the six main spines, arising from the
lattice-knots). Pores regular, circular, enclosed by prominent, hexagonal
frames, four to six times as broad as the bars; seven to eight on the
radius. {174}Six main spines triangular pyramidal, with spirally contorted
prominent edges, as long as the radius, as broad at the base as one pore.

_Dimensions._--Diameter of the shell 0.13, pores 0.01, bars 0.002; length
of the spines 0.06 to 0.8, basal breadth 0.01.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

12. _Hexastylus cochleatus_, n. sp. (Pl. 21, fig. 1).

Shell thick walled, covered with numerous short, conical spines (about as
large as one pore). Pores regular, circular, three times as broad as the
bars; eight to nine on the radius. Six main spines triangular pyramidal,
with prominent, spirally-twisted edges, as long as the diameter of the
shell, and twice as broad at the base as one pore.

_Dimensions._--Diameter of the shell 0.17, pores 0.015, bars 0.005; length
of the spines 0.16, breadth 0.03.

_Habitat._--Mediterranean, Corfu, Haeckel, surface.

13. _Hexastylus setosus_, Haeckel.

_Acanthosphæra setosa_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 285, Taf. ix. fig. 11.

Shell thin walled, covered with numerous short, bristle-shaped spines (not
larger than one pore). Pores regular, circular, four times as broad as the
bars; thirteen to sixteen on the radius. Six main spines three-sided
pyramidal, scarcely one-third as long as the radius. (In the figure of
Ehrenberg, _loc. cit._, only four spines are in regular, crossed
disposition, two others opposite in oblique direction; this is either an
individual abnormality, or an error of drawing; the same species occurs
with six spines exactly regularly disposed in the three dimensive axes.)

_Dimensions._--Diameter of the shell 0.15; pores 0.008, bars 0.002; length
of the spines 0.02 to 0.03, basal breadth 0.005.

_Habitat._--Indian Ocean, Zanzibar, Pullen; Madagascar, Rabbe.

14. _Hexastylus chilonis_, n. sp.

Shell thin walled, covered with numerous bristle-shaped spines (as long as
the radius). Pores regular, circular, twice as broad as the bars; eight to
nine on the radius. Six main spines triangular pyramidal, with straight
edges, as long as the diameter of the shell, as broad as two pores.

_Dimensions._--Diameter of the shell 0.2, pores 0.02, bars 0.01; length of
the spines 0.2, basal breadth 0.05.

_Habitat._--South Atlantic, Station 325, surface.

15. _Hexastylus cleobuli_, n. sp.

Shell thick walled, covered with numerous short, conical spines (twice as
large as one pore). Pores regular, circular, of the same breadth as the
bars; twelve to thirteen on the radius. Six main spines conical, as long as
the radius, three times as broad as one pore.

{175}_Dimensions._--Diameter of the shell 0.24, pores and bars 0.005;
length of the spines 0.12, basal breadth 0.016.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

Subgenus 3. _Hexastylissus_, Haeckel.

_Definition._--Pores irregular, of unequal size or form; surface of the
spherical shell smooth, without radial by-spines (other than the six main
spines).

16. _Hexastylus triaxonius_, n. sp. (Pl. 21, fig. 2).

Shell thin walled, with smooth surface. Pores irregular, polygonal, three
to six times as broad as the bars; four to six on the radius. Six spines
triangular pyramidal, as long as the diameter of the shell, as broad at the
base as one pore.

_Dimensions._--Diameter of the shell 0.05, pores 0.004 to 0.008, bars
0.0015; length of the spines 0.04, basal breadth 0.006.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

17. _Hexastylus dimensivus_, n. sp. (Pl. 21, fig. 6).

Shell thin walled, with smooth surface. Pores irregular, polygonal, four to
eight times as broad as the bars; eleven to thirteen on the radius. Six
spines hexagonal pyramidal, as long as the radius, about three times as
broad at the base as one pore.

_Dimensions._--Diameter of the shell 0.13, pores 0.006 to 0.009, bars
0.0012; length of the spines 0.06, basal breadth 0.025.

_Habitat._--North Pacific, Station 256, depth 2950 fathoms.

18. _Hexastylus brevispinus_, n. sp.

Shell thin walled, with smooth surface. Pores irregular, roundish, two to
four times as broad as the bars; six to eight on the radius. Six spines
triangular pyramidal, half as long as the radius, twice as broad as one
pore.

_Dimensions._--Diameter of the shell 0.2, pores 0.008 to 0.016, bars 0.004;
length of the spines 0.05, basal breadth 0.025.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

19. _Hexastylus longispinus_, n. sp.

Shell thick walled, smooth. Pores irregular, roundish, two to three times
as broad as the bars; three to four on the radius. Six spines triangular
prismatical, two to three times as long as the diameter of the shell.

{176}_Dimensions._--Diameter of the shell 0.08, pores 0.008 to 0.012, bars
0.004; length of the spines 0.2, breadth 0.01.

_Habitat._--North Atlantic, Station 353, surface.

20. _Hexastylus marginatus_, n. sp. (Pl. 21, fig. 10).

Shell thick walled, rough. Pores irregular, roundish, somewhat
funnel-shaped double-edged, two to three times as broad as the bars; twelve
to fourteen on the radius. Six spines three-sided pyramidal, somewhat
longer than the radius, three times as broad as one pore.

_Dimensions._--Diameter of the shell 0.15, pores 0.007 to 0.01, bars 0.004;
length of the spines 0.1, basal breadth 0.025.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

21. _Hexastylus conifer_, n. sp.

Shell thick walled, rough. Pores irregular, roundish, scarcely broader than
the bars; fifteen to sixteen on the radius. Six spines conical, as long as
the radius, five to seven times as broad as one pore.

_Dimensions._--Diameter of the shell 0.18, pores and bars 0.004 to 0.006;
length of the spines 0.1, basal breadth 0.03.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe, surface.

Subgenus 4. _Hexastylurus_, Haeckel.

_Definition._--Pores irregular, of unequal size or form; surface of the
spherical shell spiny, covered with numerous conical or bristle-shaped
by-spines.

22. _Hexastylus dictyotus_, n. sp. (Pl. 21, figs. 8, 9).

Shell thin walled, spiny; short spines conical, smaller than the
pore-breadth. Pores irregular, polygonal, five to seven times as broad as
the bars; four to six on the radius. Six spines triangular pyramidal,
longer than the radius, about as broad as one smaller pore.

_Dimensions._--Diameter of the shell 0.09, pores 0.01 to 0.015, bars 0.002;
length of the spines 0.06, basal breadth 0.01.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

23. _Hexastylus hirsutus_, n. sp.

Shell thin walled, densely covered with bristle-shaped, radial spines, half
as long as the six main spines. Pores irregular, polygonal, three to four
times as broad as the bars; eight to ten on the radius. Six spines
triangular pyramidal, as long as the radius, twice as broad as one pore.

{177}_Dimensions._--Diameter of the shell 0.14, pores 0.005 to 0.008, bars
0.002; length of the spines 0.06, basal breadth 0.01.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

24. _Hexastylus contortus_, n. sp. (Pl. 21, fig. 12).

Shell thick walled, covered with bristle-shaped, radial spines, half as
long as the radius. Pores irregular, roundish, two to three times as broad
as the bars; seven to eight on the radius. Six spines triangular prismatic,
in the distal half spirally twisted like a cork-screw (fig. 12); longer
than the diameter of the shell, about as broad as one pore.

_Dimensions._--Diameter of the shell 0.12, pores 0.006 to 0.008, bars
0.003; length of the spines 0.15, breadth 0.015.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

25. _Hexastylus spiralis_, n. sp. (Pl. 21, fig. 7).

Shell thick walled, covered with short conical spines. Pores irregular,
roundish, two to three times as broad as the bars; five to six on the
radius. Six spines triangular prismatic, with three thin, spirally
contorted edges, two to three times as long as the diameter of the shell,
about as broad as one large pore.

_Dimensions._--Diameter of the shell 0.16, pores 0.01 to 0.02, bars 0.006;
length of the spines 0.3 to 0.5, basal breadth 0.02.

_Habitat._--Tropical Western Pacific, Station 225, depth 4475 fathoms.

Genus 73. _Hexastylarium_,[102] n. gen.

_Definition._--#Cubosphærida# with one simple lattice-sphere and six simple
spines of different sizes; one opposite pair larger than the other two.

The genus _Hexastylarium_ differs from its probable ancestral form,
_Hexastylus_, by the unequal growth of the six simple spines; two opposite
spines of one pair being more strongly developed than the four others,
which are equal. They correspond therefore to the three axes of a quadratic
crystal.

1. _Hexastylarium heteraxonium_, n. sp.

Cortical shell thin walled, smooth, with regular, hexagonal pores, three
times as broad as the bars; eight to ten on the radius. Six spines
three-sided pyramidal, at the base as broad as one pore. Two opposite major
spines longer than the shell diameter; four others scarcely as long as
{178}the radius. (Similar to _Hexastylus phænaxonius_, Pl. 21, fig. 3, but
differing in the unequal length of the spines.)

_Dimensions._--Diameter of the shell 0.15, pores 0.12, bars 0.004; length
of the two major spines 0.2, of the four minor 0.07.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

2. _Hexastylarium quadratum_, n. sp.

Shell very delicate, with smooth surface, and irregular, polygonal pores,
separated by very thin bars. The form of the shell is not, as commonly, a
sphere, but a geometrical square octahedron, one axis (with two opposite
major spines) being nearly twice as long as the other two dimensive axes;
four spines, opposite by pairs in the latter, are only half as long. The
eight sides of the octahedral shell are even, equilateral-triangular.
Spines angular, thin.

_Dimensions._--Diameter of the shell in the major axis 0.18, in the minor
0.1; length of the major spines 0.24, minor 0.12.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

3. _Hexastylarium elongatum_, n. sp.

Shell thick walled, with spiny surface, and with irregular, roundish pores,
two to four times as broad as the bars; eight to ten on the radius. Two
opposite major spines, three to four times as long as the shell diameter,
whilst the four other spines are very short, scarcely as long as the
radius. All six spines at the base three-sided pyramidal, the two longer
being cylindrical.

_Dimensions._--Diameter of the shell 0.1; length of the major spines 0.3 to
0.4, of the minor 0.04, basal breadth 0.02.

_Habitat._--North Pacific, Station 244, surface.

Genus 74. _Hexastylidium_,[103] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Cubosphærida# with one simple lattice-sphere and six simple
spines in pairs different; the two opposite spines of each pair equal, the
three pairs unequal.

The genus _Hexastylidium_ differs from its probable ancestral form,
_Hexastylus_, by the unequal growth of the six simple spines; the two
spines of each pair reaching the same dimensions, whilst the three pairs
are different. They correspond therefore to the three axes of a rhombic
crystal.

1. _Hexastylidium rhomboides_, n. sp.

Shell thin walled, smooth, with irregular, polygonal pores and very thin
bars. Its form is not, as commonly, a sphere, but a rhombic octahedron. The
radial proportion of the three unequal {179}dimensive axes = 1 : 2 : 4. The
radial proportion of the three pairs of spines = 1 : 3 : 8. Spines thin
cylindrical, at the base angular.

_Dimensions._--Diameter of the major shell axis 0.2, middle 0.1, minor
0.05; length of the major spines 0.3, middle 0.12, minor 0.04.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

2. _Hexastylidium spirale_, n. sp.

Shell thick walled, spherical, thorny, with irregular, roundish pores,
three to five times as broad as the bars; six spines very stout, prismatic
(as broad as one large pore), with three ring-like, spirally convoluted
edges. Two opposite major spines of extraordinary length, ten to twelve
times as long as the shell diameter, the two middle opposite spines being
about as long as the latter, the two minor scarcely one-third as long.
(Similar to _Hexastylus spiralis_, Pl. 21, fig. 7, but distinguished by the
very unequal length of the spines.)

_Dimensions._--Diameter of the shell 0.12; length of the major spines 1 to
1.5 mm., middle 0.15, minor 0.04.

_Habitat._--Central Pacific, Station 266, surface.

Subfamily HEXALONCHIDA,[104] Haeckel, 1881, Prodromus, pp. 449, 451.

_Definition._--#Cubosphærida# with two concentric spherical lattice-shells.

Genus 75. _Hexalonche_,[105] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Cubosphærida# with two concentric lattice-spheres and six
simple spines of equal size.

The genus _Hexalonche_ is the most simple form, and probably the common
ancestral form, of all Hexalonchida, or those Cubosphærida which possess
two concentric latticed spheres, connected by six radial beams. Commonly
one shell is intracapsular (medullary shell) and the other extracapsular
(cortical shell); but sometimes also both shells are extracapsular, and
these forms may perhaps be better separated as a peculiar genus
_Hexadilemma_. In _Hexalonche_ all six simple spines are of equal size, and
opposite by pairs in three equal dimensive axes, corresponding to the three
equal axes of a tesseral crystal. It can be derived from _Hexastylus_ by
duplication of the lattice-shell.

Subgenus 1. _Hexalonchara_, Haeckel.

_Definition._--Pores of the cortical shell regular or subregular, of nearly
equal size and similar form; surface smooth, without radial by-spines
(other than the six main spines).

{180}1. _Hexalonche phænaxonia_, n. sp.

Cortical shell thin walled, smooth; its pores regular, hexagonal, six to
eight times as broad as the bars; eight to ten to twelve on the radius.
Medullary shell one-third as broad, with regular, hexagonal pores of half
size. Six spines triangular pyramidal, as long as the radius of the shell,
at the base as broad as one pore. (Differs from _Hexastylus phænaxonius_,
Pl. 21, fig. 3, only in the medullary shell and the six inner radial beams,
connecting it with the cortical shell.)

_Dimensions._--Diameter of the outer shell 0.15, pores 0.01 to 0.015, bars
0.015 to 0.02; inner shell 0.05; length of the spines 0.08, basal breadth
0.01.

_Habitat._--Central Pacific, Stations 266 to 271, depth 2425 to 2925
fathoms.

2. _Hexalonche rosetta_, n. sp. (Pl. 25, figs. 3, 3_a_, 3_b_).

Cortical shell thick walled, smooth, two and a half time as broad as the
medullary shell. Pores of the latter (fig. 3_a_) regular circular; eight to
ten on the half meridian, about as broad as the bars between them. Pores of
the outer shell regular, hexagonal, remarkable for a very peculiar form and
arrangement. In the transverse section of the shell (fig. 3_b_) they appear
as narrow, hexagonal prismatic canals, twice as high as broad, and four to
five times as broad as the thin elevated bars between them. Every seven
meshes form together a larger, regular hexagon (six pores surrounding one
central pore). The periphery of these larger, rosette-like hexagons
projects more strongly from the surface than the walls between the smaller
hexagons. On the half meridian of the shell may be counted six to seven
larger and eighteen to twenty smaller hexagons. The six radial beams
between the two shells are thin, three-sided prismatic, not broader than
the bars of the network, the prominent prolongations of which form six very
strong spines of peculiar club-like shape (fig. 3), as long as the radius
of the outer shell. The three wings of the club are lower in the inner,
higher in the outer half; the broadest part of the spine (at the base and
in the outer third) is as broad as a hexagonal rosette (equal to three
meshes of the outer shell); its outer apex is pyramidal.

_Dimensions._--Diameter of the outer shell 0.13, pores 0.008 to 0.01, bars
0.02; inner shell 0.05; length of the spines 0.07, distal breadth 0.03.

_Habitat._--Tropical West Pacific, Station 225, depth 4475 fathoms.

3. _Hexalonche favosa_, n. sp.

Cortical shell thick walled, smooth, four times as broad as the medullary
shell. Pores of the former regular circular, hexagonally framed, deep
funnel-shaped, of the same breadth as the bars; six to eight on the radius.
Six spines triangular-pyramidal, as long as the radius, at the base twice
as broad as one pore. (Differs from the similar _Hexastylus favosus_ mainly
in the possession of a medullary shell.)

_Dimensions._--Diameter of the outer shell 0.16, pores and bars 0.012;
inner shell 0.04; length of the spines 0.08, basal breadth 0.025.

_Habitat._--Equatorial Atlantic, Station 348, depth 2450 fathoms; also
fossil in Barbados.

{181}4. _Hexalonche octahedra_, n. sp. (Pl. 22, figs. 8, 8_a_).

Cortical shell a regular octahedron, thin walled, with twelve more or less
rounded edges (between the bases of the spines), and with smooth surface,
three times as broad as the spherical medullary shell. Pores of the former
regular circular (three times as large as those of the latter), four times
as broad as the bars; five to seven on the radius. Six spines three-sided
prismatic, with thickened base and cuspidated end, somewhat longer than the
radius, and once to twice as broad as one pore.

_Dimensions._--Diameter of the outer shell 0.11, pores 0.01, bars 0.0025;
inner shell 0.04; length of the spines 0.08, bars 0.01.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

5. _Hexalonche conicornis_, n. sp. (Pl. 22, fig. 2).

Cortical shell thick walled, smooth, three times as broad as the medullary
shell. Pores of both spheres regular circular, three to four times as broad
as the bars; those of the thick walled outer shell six to seven on the
radius, twice as large as those of the thin walled inner shell; six radial
beams between the two spheres, very thin, cylindrical; six spines short,
conical, scarcely as long as the radius of the outer shell, at the base
twice as broad as one pore.

_Dimensions._--Diameter of the outer shell 0.12 to 0.14, pores 0.01, bars
0.003; inner shell 0.04 to 0.05; length of the spines 0.04 to 0.06, basal
breadth 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

6. _Hexalonche curvicornis_, n. sp. (Pl. 25, fig. 4).

Cortical shell thick walled, smooth, not much larger than the medullary
shell (= 4 : 3). Pores of both spheres regular circular, nearly of the same
size, three times as broad as the bars between them; seven to nine on the
radius. Six spines three-sided prismatic, inside and outside of the
exterior shell of equal thickness, twice as broad as one pore, longer than
the diameter of the outer shell, and in a singular manner curved like an ox
horn; the three edges of each horn somewhat spirally twisted.

_Dimensions._--Diameter of the outer shell 0.16, pores 0.01, bars 0.003;
inner shell 0.12; length of the spines 0.2, breadth 0.02.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

7. _Hexalonche brevicornis_, n. sp.

Cortical shell thin walled, smooth, three times as broad as the medullary
shell. Pores of the former regular circular, double-edged, six to eight
times as broad as the thin bars, five to six on the radius. Six spines
triangular pyramidal, as broad as one pore and only twice as long.

_Dimensions._--Diameter of the outer shell 0.14, pores 0.02, bars 0.003;
inner shell 0.045; length of the spines 0.05, basal breadth 0.02.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

{182}8. _Hexalonche grandis_, n. sp.

Cortical shell thick walled, smooth, four times as broad as the medullary
shell. Pores of the outer shell regular circular, four times as broad as
the bars; thirteen to fifteen on the radius. Six spines conical, scarcely
half as long as the radius, at the base as broad as one pore.

_Dimensions._--Diameter of the outer shell 0.2 to 0.32, pores 0.01 to 0.02,
bars 0.003 to 0.005; inner shell 0.06 to 0.08; length of the spines 0.06,
breadth 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subgenus 2. _Hexalonchetta_, Haeckel.

_Definition._--Pores of the cortical shell regular or subregular, of nearly
equal size and similar form; surface covered with numerous conical or
bristle-shaped radial by-spines.

9. _Hexalonche amphisiphon_, n. sp. (Pl. 25, figs. 2, 2_a_, 2_b_).

Cortical shell thin walled, armed with very numerous bristle-shaped, radial
by-spines, a quarter to a half as long as the six main spines. Pores
regular hexagonal, twelve to fourteen on the radius, with very thin bars,
prolonged on the outer as well as the inner surface into a short truncated
conical tube (Pl. 25, fig. 2_b_). Medullary shell octahedral, with
irregular polygonal meshes and very thin bars between them (fig. 2_a_),
connected with the outer (six to eight times larger) shell by six very thin
radial beams. These are prolonged outside into six strong pyramidal spines,
nearly as long as the diameter of the outer shell, with sharp straight
edges, at the base twice as broad as one pore.

_Dimensions._--Diameter of the outer shell 0.15, pores 0.01, bars 0.001;
inner shell 0.02; length of the spines 0.12, basal breadth 0.02.

_Habitat._--Central Pacific, Station 271, surface.

10. _Hexalonche anaximandri_, n. sp. (Pl. 22, fig. 5).

Cortical shell thin walled, covered with short conical spines (smaller than
the pores), three times as broad as the medullary shell. Pores of the outer
shell subregular hexagonal, four times as broad as the bars; five to seven
on the radius. Inner shell with regular hexagonal pores of half the size,
connected with the outer by six strong, three-sided prismatic beams, which
are prolonged outside into six very stout pyramidal spines, with three
prominent edges, longer than the radius and twice as broad as one pore.

_Dimensions._--Diameter of the outer shell 0.12, pores 0.01, bars 0.0025;
inner shell 0.04; length of the spines 0.07, basal breadth 0.02.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

{183}11. _Hexalonche octocolpa_, n. sp. (Pl. 22, figs. 6, 6_a_).

Cortical shell thin walled, nearly octahedral, with eight hemispherical or
bosom-shaped vaultings, corresponding to the eight faces of a regular
octahedron, the three axes of which are indicated by the six spines.
Surface covered with short bristle-shaped spines. Pores regular hexagonal,
ten to fifteen times as broad as the thin bars; seven to nine on the
radius. Medullary shell (fig. 6_a_) spherical, with regular circular pores,
one-third as broad as the cortical shell, and connected with it by six
strong triangular radial beams, which are prolonged outside into pyramidal
spines, one-third as long as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.2, pores 0.03, bars 0.002;
inner shell 0.06; length of the spines 0.06, breadth 0.02.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

12. _Hexalonche cristata_, n. sp.

Cortical shell thick walled, four times as broad as the medullary shell,
and covered with short conical spines arising from the nodal points of
elevated crests which form regular hexagonal frames around the
funnel-shaped circular pores. These are two to three times as broad as the
crested bars; eight to ten on the radius. Six spines three-sided pyramidal,
with strong prominent edges, about as long as the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.16 to 0.2, pores 0.01 to
0.015, bars 0.003 to 0.005; inner shell 0.04 to 0.05; length of the spines
0.08 to 0.12, basal breadth 0.01 to 0.03.

_Habitat._--Central Pacific, Station 266 to 274, depth 2350 to 2925
fathoms.

13. _Hexalonche serrata_, n. sp.

Cortical shell thick walled, four times as broad as the medullary shell,
with spiny surface. Pores regular circular, twice as broad as the bars,
five to seven on the radius, funnel-shaped, separated by hexagonal frames,
the sharp crests of which are serrated; at the nodal-points longer
bristle-shaped by-spines. Six main spines triangular prismatic, longer than
the diameter of the shell, twice as broad as one pore.

_Dimensions._--Diameter of the outer shell 0.15, pores 0.01, bars 0.005;
inner shell 0.04; length of the spines 0.2, breadth 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

14. _Hexalonche anaximenis_, n. sp. (Pl. 25, fig. 5).

Cortical shell thin walled, twice as broad as the medullary shell, and
covered with short conical by-spines. Pores regular circular, twice as
broad as the bars; five to six on the radius. Circular pores of the
medullary shell one-third as broad. Radial main spines conical, nearly as
long as the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.1, pores 0.01, bars 0.005;
inner shell 0.05; length of the spines 0.04, basal breadth 0.015.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

{184}15. _Hexalonche aspera_, n. sp.

Cortical shell thin walled, three times as broad as the medullary shell,
and covered with short conical spines. Pores regular circular, four to six
times as broad as the bars; eight to ten on the radius. Radial spines
conical, about as long as the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.12, pores 0.08, bars 0.0015;
inner shell 0.04; length of the spines 0.1, basal breadth 0.02.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

16. _Hexalonche castanella_, n. sp.

Cortical shell thick walled, five times as broad as the medullary shell,
and covered with numerous short conical spines. Pores regular circular,
twice as broad as the bars; ten to twelve on the radius. Radial spines
triangular pyramidal, about as long as the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.22, pores 0.01, bars 0.005;
inner shell 0.045; length of the spines 0.12, basal breadth 0.02.

_Habitat._--North Atlantic, Station 353, depth 3125 fathoms.

Subgenus 3. _Hexalonchilla_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of unequal size or
form; surface smooth, without radial by-spines (other than the six main
spines).

17. _Hexalonche hexacantha_, Haeckel.

_Haliomma hexacanthum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 35, Taf. iv. fig. 5.

_Haliomma hexacanthum_, Haeckel, 1862, Monogr. d. Radiol., p. 430.

Cortical shell thin walled, smooth, with irregular polygonal pores
(commonly hexagonal or pentagonal), two to three times as broad as the
bars; eight to ten on the radius. Medullary shell one quarter as broad,
connected with the former by six thin radial beams which are prolonged
outside into six triangular pyramidal spines (not quadrangular, as Müller
describes), longer than the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.2, pores 0.01 to 0.015, bars
0.005; inner shell 0.05; length of the spines 0.15, basal breadth 0.015.

_Habitat._--Mediterranean (Messina, Nice, Genoa).

18. _Hexalonche geometrica_, n. sp.

Cortical shell thick walled, smooth, with very peculiar geometrical
formation of its network, the pores of which are of very different size and
form, but highly regular disposition. Each of the six {185}triangular
spines is surrounded at the base by three small roundish pores (between the
three wings of its base), and further by a coronal of six very large pores,
three of which are pentagonal, and the other three (alternating)
heptagonal. The six basal coronals are separated by irregular smaller
pores. Medullary shell thin walled, with regular hexagonal pores (three on
the radius) and thin bars, connected with the outer shell by six thin
triangular prismatic radial beams, which are prolonged outside into short
pyramidal spines (half as long as the radius).

_Dimensions._--Diameter of the outer shell 0.2, thirty-six larger pores of
the coronals 0.03 to 0.04, smaller pores between them 0.01 to 0.02, bars
0.005; inner shell 0.05 (with pores of 0.008); length of the spines 0.05,
basal breadth 0.03.

_Habitat._--South Atlantic, off Buenos Ayres, Station 323, depth 1900
fathoms.

19. _Hexalonche pythagoræa_, n. sp. (Pl. 22, fig. 1).

Cortical shell thick walled, smooth, three times as broad as the medullary
shell. Pores irregular roundish, twice to eight times as broad as the bars,
of very different size; usually in the space between every three spines are
three or six larger pores, separated by numerous smaller pores (often the
disposition of the larger pores is much more regular than in the figured
specimen). Medullary shell with regular circular pores, connected with the
outer by six very thin radial beams, which are prolonged outside into six
short triangular pyramidal spines about as long and broad as the half
radius.

_Dimensions._--Diameter of the outer shell 0.12, pores 0.008 to 0.03, bars
0.004; inner shell 0.04; length of the spines 0.04, basal breadth 0.03.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

20. _Hexalonche aristarchi_, n. sp. (Pl. 22, fig. 3).

Cortical shell thin walled, smooth, four times as broad as the medullary
shell. Pores irregular polygonal, of very variable size and form, twice to
six times as broad as the bars; six to ten on the radius. Medullary shell
with regular hexagonal pores, connected with the outer by six very thin
radial beams, which are prolonged outside into six triangular pyramidal
spines, nearly as long as the radius.

_Dimensions._--Diameter of the outer shell 0.1, pores 0.005 to 0.02, bars
0.003; inner shell 0.025; length of the spines 0.05, basal breadth 0.01.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

21. _Hexalonche ekphantæa_, n. sp.

Cortical shell thin walled, smooth, four times as broad as the medullary
shell. Pores irregular roundish, of very variable size and form, twice to
six times as broad as the bars, ten to fifteen on the radius. Six spines
cylindro-conical, longer than the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.2, pores 0.004 to 0.012, bars
0.002; inner shell 0.05; length of the spine 0.3, basal breadth 0.01.

_Habitat._--Indian Ocean, Sunda Strait, surface, Rabbe.

{186}Subgenus 4. _Hexalonchusa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface covered with numerous conical or bristle-shaped, radial
by-spines.

22. _Hexalonche philosophica_, n. sp. (Pl. 22, fig. 4).

Cortical shell thin walled, covered with short conical spines, three times
as broad as the medullary shell. Pores irregular polygonal, or more
roundish, twice to six times as broad as the bars; six to eight on the
radius. Inner shell of the same structure, pores three times smaller,
connected with the outer by six very thin radial beams, which are prolonged
outside into six strong, triangular pyramidal spines, as long as the
radius. (Similar to _Hexalonche anaximandri_, Pl. 22, fig. 5, but different
in the irregular network and the shorter by-spines.)

_Dimensions._--Diameter of the outer shell 0.11, pores 0.005 to 0.015, bars
0.0025; inner shell 0.04; length of the spines 0.06, breadth 0.02.

_Habitat._--North Atlantic, Færöe Channel, John Murray; Iceland, Krabbe,
surface.

23. _Hexalonche seleuci_, n. sp.

Cortical shell thick walled, covered with numerous short, bristle-shaped
spines. Pores of very different size and form, and of a peculiar,
subregular disposition, similar to those of _Hexalonche geometrica_ (though
in this case more regular). Each of the six triangular main spines (which
reach nearly the length of the radius) is surrounded at the base by three
small roundish pores, and these are further supplemented by a coronal of
six very large polygonal pores; the six coronals are separated by irregular
rows of smaller pores. Inner shell equal to one-third of the outer. Six
main spines pyramidal, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.16, larger pores 0.03 to 0.04,
smaller pores 0.01 to 0.02, bars 0.05 to 0.01; inner shell 0.05; length of
the spines 0.08, basal breadth 0.03.

_Habitat._--North Atlantic, Station 354, surface.

24. _Hexalonche sexaculeata_, Haeckel.

_Haliomma sexaculeatum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 87, Taf.
i. fig. 8.

Cortical shell thick walled, twice as broad as the medullary shell, and
covered with short conical spines. Pores irregular polygonal (mostly
hexagonal), twice to three times as broad as the bars; six to eight on the
radius. Six spines triangular-pyramidal (not quadrangular), somewhat longer
than the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.11, pores 0.06 to 0.08, bars
0.03; inner shell 0.06; length of the spines 0.07, basal breadth 0.015.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Caltanisetta.

{187}25. _Hexalonche heracliti_, n. sp. (Pl. 22, fig. 7).

Cortical shell thick walled, covered with bunches of small spines, each
bunch at the nodule point between every three pores composed of four to
eight conical spinules. Pores irregular roundish, twice to four times as
broad as the bars; four to six on the radius. Inner shell equal to
one-third of the outer, connected with it by six thin radial beams, which
are prolonged outside into six strong triangular spines about as long as
the radius.

_Dimensions._--Diameter of the outer shell 0.15, pores 0.015 to 0.03, bars
0.08; inner shell 0.05; length of the spines 0.06, basal breadth 0.02.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

26. _Hexalonche xenophanis_, n. sp.

Cortical shell thin walled, covered with numerous short conical spines,
four times as broad as the medullary shell. Pores irregular circular, twice
to eight times as broad as the bars; five to seven on the radius. Six
spines conical or more cylindrical, as long as the diameter of the shell or
longer.

_Dimensions._--Diameter of the outer shell 0.16 to 0.24, pores 0.01 to
0.04, bars 0.005; inner shell 0.04 to 0.06; length of the spines 0.2 to
0.3, breadth 0.02.

_Habitat._--Central Pacific, Stations 266 to 272, surface.

27. _Hexalonche setosa_, n. sp.

Cortical shell thin walled, covered with numerous bristle-shaped spines,
half as long as the six main spines. Pores irregular roundish, twice to
five times as broad as the bars; ten to twelve on the radius. Inner shell
equal to one-fifth of the outer. Six spines conical, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.25, pores 0.008 to 0.02, bars
0.004; inner shell 0.05; length of the spines 0.12, basal breadth 0.03.

_Habitat._--South Pacific, Station 291, surface.

28. _Hexalonche hystricina_, n. sp. (Pl. 25, fig. 6).

Cortical shell thick walled, covered with numerous bristle-shaped spines,
half as long as the six main spines. Pores irregular roundish, twice to
four times as broad as the bars; five to seven on the radius. Medullary
shell with very small circular pores, one-fourth of the cortical shell,
connected with it by six thin prismatic radial beams, which are prolonged
outside into six strong, short, three-sided pyramidal, cuspidated spines,
only half as long as the radius of the outer shell.

_Dimensions._--Diameter of the outer shell 0.2, pores 0.015 to 0.03, bars
0.008; inner shell 0.05; length of the spines 0.05, basal breadth 0.02.

_Habitat._--Indian Ocean, Sunda Strait, surface, Rabbe.

{188}Genus 76. _Hexancistra_,[106] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Cubosphærida# with two concentric lattice-spheres and six
branched spines of equal size.

The genus _Hexancistra_ differs from its ancestral form, _Hexalonche_, in
the ramification of the six radial spines. These are very different in the
two subgenera; in _Hexancora_ each spine bears only three simple lateral
branches, while in _Hexapitys_ there are three rows of verticillate lateral
branches on each spine.

Subgenus 1. _Hexancora_, Haeckel.

_Definition._--Each radial spine with three simple lateral branches only
(one branch from each edge of the triangular spine).

1. _Hexancistra tricuspis_, n. sp. (Pl. 22, fig. 9).

Cortical shell thin walled, covered with short conical by-spines, three
times as broad as the medullary shell; its pores regular circular, three
times as broad as the bars; ten to twelve on the radius. Pores of the
medullary shell half as large, also regular circular. The two shells
connected by six thin prismatic radial beams, which are prolonged outside
into six very stout main spines, three-sided prismatic, as long as the
shell diameter, with three thin wing-like edges. Each edge at the distal
end prolonged into a strong curved lateral branch.

_Dimensions._--Diameter of the outer shell 0.13, inner 0.4; cortical pores
0.01, bars 0.003; medullary pores 0.005, bars 0.003; length of the six
spines 0.13, breadth 0.02.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

2. _Hexancistra ancorata_, n. sp.

Cortical shell thick walled, smooth, twice as broad as the medullary shell;
its pores regular circular, twice as broad as the bars; six to eight on the
radius. Six radial spines, three-sided prismatic, as long as the shell
radius, with three recurved lateral branches at the distal end like the
three teeth of an anchor.

_Dimensions._--Diameter of the outer shell 0.1, inner 0.05; cortical pores
0.006, bars 0.003; length of the six spines 0.05, breadth 0.02.

_Habitat._--South Pacific, Station 296, depth 1825 fathoms.

3. _Hexancistra triserrata_, n. sp. (Pl. 22, fig. 10).

Cortical shell thin walled, thorny, twice to three times as broad as the
medullary shell; the two shells connected by six strong triangular beams.
Inner shell spherical, with very small, regular, circular {189}pores; ten
to twelve on the half meridian. Bars as broad as the pores. Outer shell
regularly octahedral, with eight triangular perfectly regular even faces,
separated by eight prominent edges; the meshes circular, regular, about
twelve to fourteen on the half meridian, two to three times as broad as the
bars between them. Six strong radial spines, about as long as the diameter
of the outer shell, three-sided prismatic, cuspidate; their three edges
prominent, serrate, spirally twisted. Each spine bears about its middle
three strong flattened lateral branches, nearly perpendicular to it, not
serrated, and slightly curved.

_Dimensions._--Diameter of the outer shell 0.13, inner 0.045; pores of the
former 0.01, of the latter 0.004; length of the spine 0.12, breadth 0.014.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

4. _Hexancistra quadricuspis_, n. sp. (Pl. 22, fig. 11).

Cortical shell thin walled, covered with short bristle-shaped by-spines,
three and a half times as broad as the medullary shell. Pores of the latter
regular circular, small; pores of the former three to six times as large,
very irregular, roundish, double-edged. Six radial spines, three-sided
prismatic, about as long as the shell diameter, with three wing-like,
slightly twisted edges, which are prolonged towards the distal end into
three curved horn-shaped branches.

_Dimensions._--Diameter of the outer shell 0.17, inner 0.05; cortical pores
0.01 to 0.03, bars 0.005; medullary pores 0.003, bars 0.001; length of the
spines 0.15, breadth 0.013.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Subgenus 2. _Hexapitys_, Haeckel, 1881, Prodromus, p. 451.

_Definition._--Each radial spine with three rows of verticillate lateral
branches (a row arising from each edge of the spine).

5. _Hexancistra mirabilis_, n. sp. (Pl. 23, fig. 3).

_Hexapitys mirabilis_, Haeckel, 1881, Prodromus, p. 451.

Cortical shell very thin walled, three times as broad as the medullary
shell. Inner shell spherical, with very delicate, subregular hexagonal
meshes; seven to eight on the half diameter. Outer shell octahedral, with
irregular polygonal meshes of very different size; on the surface numerous
thin accessory radial spines, equal in length to its radius. Six main
spines, extremely long and stout, many times longer than the diameter of
the outer shell, nearly as broad as the radius of the inner shell,
three-sided prismatic, with sharp, prominent, spirally twisted edges; on
every edge a great number of thin lateral branches, arranged
perpendicularly to it, as long as the diameter of the outer shell, and
pinnated by ten to twenty pairs of delicate secondary spinules, biserial
and perpendicular to the primary branches. (In the figured specimen the
spherical central capsule, between both shells, was well preserved; its
nucleus nearly filled the medullary shell. The thick jelly-veil around it
was radially striped and octahedral.)

{190}_Dimensions._--Diameter of the outer shell 0.13, of the inner 0.05;
length of the spines 0.5 to 0.8 or more, breadth 0.02.

_Habitat._--Central Pacific, Station 271, surface.

Genus 77. _Hexaloncharium_,[107] n. gen.

_Definition._--#Cubosphærida# with two concentric lattice-spheres and six
simple spines of different sizes; one opposite pair larger than the two
others.

The genus _Hexaloncharium_ exhibits the same relation to its ancestral
form, _Hexalonche_, that _Hexastylarium_ bears to _Hexastylus_. Two
opposite spines of one pair are larger than the four others, and correspond
to the three axes of a quadratic crystal.

1. _Hexaloncharium octahedrum_, n. sp.

Cortical shell smooth, three to four times as broad as the spherical
medullary shell, each having regular circular pores, twice to three times
as broad as the bars. Form of the outer shell not a sphere, but a regular
octahedron, with eight congruent, equilateral triangular even faces. Two
opposite spines twice as long as the shell diameter, whilst the four others
are scarcely equal to it. Basal breadth of all six the same (three times as
large as one pore); form, three-sided prismatic, with cuspidate distal end.
(Similar to _Hexalonche octahedra_, Pl. 22, fig. 8, but distinct in the
unequal length of the spines.)

_Dimensions._--Diameter of the outer shell 0.16, inner 0.045; length of the
major spine 0.3, minor 0.12, basal breadth 0.03.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

2. _Hexaloncharium philosophicum_, n. sp.

Cortical shell spherical, covered with short conical by-spines, twice as
broad as the medullary shell; both with regular circular pores three to
four times as broad as the bars. Two major spines cylindrical, with conical
apex, three times as long as the four others, which are conical and about
as long as the shell radius; basal breadth of all six the same (equal to
one pore). Similar to _Hexalonche anaximenis_ (Pl. 25, fig. 5), but
distinct in the enormous prolongation of two opposite spines.

_Dimensions._--Diameter of the outer shell 0.12, inner 0.06; length of the
two major spines 0.2, of the four minor 0.06, basal breadth 0.02.

_Habitat._--Central Pacific, Station 267, depth 2700 fathoms.

{191}3. _Hexaloncharium hystricinum_, n. sp.

Cortical shell spherical, three times as broad as the medullary shell, and
densely covered with oblique bristle-shaped by-spines. Pores irregular
roundish. Two opposite major spines three-sided prismatic, longer than the
shell diameter; four minor spines pyramidal, scarcely half as long as the
shell radius. (Somewhat similar to _Hexalonche hystricina_, but distinct in
the prolongation of two major spines.)

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; length of the
two major spines 0.2, four minor 0.03, basal breadth 0.02.

_Habitat._--South Pacific, Station 288, surface.

Genus 78. _Hexalonchidium_,[108] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Cubosphærida# with two concentric lattice-spheres and six
simple spines of different sizes in pairs; the two opposite spines of each
pair equal, the three pairs unequal.

The genus _Hexalonchidium_ exhibits the same relation to _Hexalonche_ that
_Hexastylidium_ bears to _Hexastylus_; the growth of the three spine-pairs
is different, whilst both spines of each pair are equal; they correspond
therefore to the three axes of a rhombic crystal.

1. _Hexalonchidium axonometrum_, n. sp.

Cortical shell thin walled, covered with short bristle-shaped by-spines,
twice as broad as the medullary shell; both with regular hexagonal meshes,
twice to three times as broad as the bars (inner meshes half as broad as
the outer). All three spine-pairs three-sided prismatic, of very different
length but of equal breadth (equal to three pores). Major spine-pair twice
as long as the shell diameter; middle pair about equal to the latter, minor
scarcely half as long. (Similar to _Hexalonche anaximandri_, Pl. 22, fig.
5, but distinct in the different length of the spines.)

_Dimensions._--Diameter of the outer shell 0.12, inner 0.06; length of the
major spines 0.25, middle 0.1, minor 0.04.

_Habitat._--Indian Ocean, Madagascar, surface, Rabbe.

Subfamily HEXACONTIDA,[109] Haeckel, 1881, Prodromus, pp. 449, 452.

_Definition._--#Cubosphærida# with three concentric, spherical, or
octahedral lattice-shells.

{192}Genus 79. _Hexacontium_, Haeckel,[110] 1881, Prodromus, p. 452.

_Definition._--Shell with three concentric lattice-spheres and six simple
spines of equal size.

The genus _Hexacontium_, the ancestral form of the Hexacontida, is probably
derived from _Hexalonche_ by duplication of the medullary shell. As in the
latter, all six spines are of equal size, opposite in pairs in the three
dimensive axes, and correspond therefore to the three equal axes of a
tesseral crystal.

Subgenus 1. _Hexacontanna_, Haeckel.

_Definition._--Pores of the cortical shell regular or subregular, of nearly
equal size and similar form; surface smooth, without radial spines or
papillæ (other than the six main spines).

1. _Hexacontium phænaxonium_, n. sp.

Cortical shell thin walled, smooth; its pores regular hexagonal, six to
nine times as broad as the bars; twelve to sixteen on the radius. Radial
proportion of the three spheres = 1 : 2 : 4. Both medullary shells of the
same structure as the cortical shell, only with smaller pores. The three
spheres connected by six thin radial beams, which are prolonged on the
outside into six strong triangular pyramidal spines, as long as the radius
of the cortical shell, and, at the base, as broad as one of its pores.
(Differs from _Hexastylus phænaxonius_ and from _Hexalonche phænaxonia_,
Pl. 21, fig. 3, in the larger size and the triple shell.)

_Dimensions._--Diameter of the outer sphere 0.2, middle 0.1, inner 0.05;
cortical pores 0.008, bars 0.0012; length of the spines 0.1.

_Habitat._--Central Pacific, Station 270, surface.

2. _Hexacontium axotrias_, n. sp. (Pl. 24, fig. 3).

Cortical shell thin walled, smooth; its pores regular circular, five to six
times as broad as the bars; eleven to thirteen on the radius. Radial
proportion of the three spheres = 1 : 3 : 8. Outer medullary shell with
hexagonal frames around the regular circular pores (five to six on the
radius). Inner medullary shell with simple small circular pores (three on
the radius). The two outer shells connected by six triangular prismatic
beams, which are prolonged outside to the length of the cortical radius or
more.

_Dimensions._--Diameter of the outer sphere 0.2, middle 0.07, inner 0.025;
cortical pores 0.016, bars 0.003; length of the spines 0.1 to 0.015, basal
breadth 0.02.

_Habitat._--North Pacific, Station 253, surface.

3. _Hexacontium hexactis_, Haeckel.

_Actinomma hexactis_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 91, Taf.
ii. fig. 7.

Cortical shell thick walled, smooth, or a little rough. Pores regular
circular, of the same breadth as the bars; five to seven on the radius.
Radial proportion of the three spheres {193}= 1 : 3 : 6. Both medullary
shells of the same structure, but with smaller pores. Six spines triangular
pyramidal, nearly as long as the diameter of the outer shell, three times
as broad at the base as one pore.

_Dimensions._--Diameter of the outer sphere 0.1, middle 0.05, inner 0.016;
cortical pores and bars 0.008; length of the spines 0.08, basal breadth
0.025.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

4. _Hexacontium lævigatum_, n. sp. (Pl. 24, fig. 6).

Cortical shell thick walled, quite smooth. Pores regular circular, with
double margins, eight to ten on the radius, of the same breadth as the
smooth bars. Radial proportion of the three spheres = 1 : 2 : 6. All three
spheres connected by six very thin radial beams, which are prolonged
outside into six short, stout, triangular, pyramidal spines, half as long
as the radius of the outer shell.

_Dimensions._--Diameter of the outer shell 0.13, middle 0.04, inner 0.02;
cortical pores and bars 0.008; length of the spines 0.04, basal breadth
0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

5. _Hexacontium triplosphærium_, n. sp.

Cortical shell thick walled, smooth. Pores regular circular, three times as
broad as the bars; ten to twelve on the radius. Radial proportion of the
three spheres = 1 : 3 : 10. Six spines conical, about as long as the radius
of the cortical shell.

_Dimensions._--Diameter of the outer shell 0.16, middle 0.05, inner 0.016;
cortical pores 0.012, bars 0.004; length of the spines 0.07, basal breadth
0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

6. _Hexacontium octahedrum_, n. sp.

Cortical shell a regular octahedron, thin walled, with twelve more or less
rounded edges (between the spine-bases) and with smooth surface; its pores
regular circular, five to seven on the radius, four times as broad as the
bars. Radial proportion of the three shells = 1 : 3 : 9. Both medullary
shells spherical, with very small circular pores. Six inner bars very thin;
six outer spines (their prolongations) triangular pyramidal, as long as the
radius of the outer shell. (Differs from _Hexalonche octahedra_, Pl. 22,
fig. 8, almost solely in the duplication of the medullary shell.)

_Dimensions._--Diameter of the outer shell 0.12, middle 0.04, inner 0.013.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

7. _Hexacontium circumtextum_, n. sp. (Pl. 25, figs. 7, 7_a_).

Cortical shell double, enclosing a simple medullary shell. Radial
proportion of the three spheres = 3 : 10 : 12. Inner cortical shell very
thick walled, with regular circular, double-edged pores, four times as
broad as the bars; seven to nine on the radius. From each nodal-point
between {194}every six pores arises a small, bristle-shaped, radial spine,
as long as the diameter of one pore. The distal ends of all these spines
are connected by very delicate tangential threads, and consequently form by
their attachment an outer cortical shell, with regular hexagonal meshes and
smooth surface. Six main spines short, three-sided prismatic, scarcely as
long as half the radius of the outer shell.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.1, inner 0.03;
pores of the middle shell 0.004, bars 0.001; length of the spines 0.025,
basal breadth 0.01.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

Subgenus 2. _Hexacontella_, Haeckel.

_Definition._--Pores of the cortical shell regular or subregular, of nearly
equal size and similar form; surface covered with numerous conical or
bristle-shaped, radial spines (other than the six main spines).

8. _Hexacontium hexagonale_, n. sp.

Cortical shell thin walled, bristly, with very delicate network; its pores
regular hexagonal, twelve to sixteen on the radius, ten to twelve times as
broad as the thin thread-like bars. At each nodal-point of the network
(between every three meshes) arises a bristle-shaped, radial spine, as long
as the diameter of one mesh. Six main spines hexagonal, as long as the
radius of the outer shell, as broad at its base as one mesh. Radial
proportion of the three spheres = 1 : 2 : 4. Pores of both medullary shells
also regular hexagonal, but much smaller, in the middle shell eleven to
twelve, in the inner, six to seven on the half meridian.

_Dimensions._--Diameter of the outer shell 0.12, of the middle 0.06, of the
inner 0.03; meshes of the outer shell 0.008; length of the six spines 0.05,
basal thickness 0.008.

_Habitat._--Indian Ocean, Ceylon, surface, Haeckel.

9. _Hexacontium favosum_, n. sp. (Pl. 24, figs. 2, 2_a_).

Cortical shell very thick walled, spiny, with regular, honeycomb-like
network; its meshes funnel-shaped, with circular inner, hexagonal outer
aperture, twice as broad as the bars; five to seven on the radius. Between
every three meshes (at each corner of the hexagon) arises a short radial
thorn, not so long as the thickness of the shell-wall. Six radial spines
very short and stout, three-sided pyramidal, scarcely half so long as the
radius of the outer shell. Radial proportion of the three spheres =
1 : 2 : 5. Pores of both medullary shells regular, circular, about as broad
as the bars, six to eight on the half meridian.

_Dimensions._--Diameter of the outer shell 0.1 to 0.12, of the middle 0.04
to 0.05, of the inner 0.02; meshes of the outer shell 0.008; length of the
six spines 0.2 to 0.3, basal breadth 0.015.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

10. _Hexacontium sceptrum_, n. sp. (Pl. 24, figs. 1, 1_a_).

Cortical shell thick walled, spiny, with regular, honeycomb-like network;
its meshes funnel-shaped, with circular inner, hexagonal outer aperture,
three to four times as broad as the bars; {195}five to six on the radius.
Between every three meshes arises a short radial thorn, as long as the
thickness of the shell-wall. Six radial spines sceptre-shaped, six-sided,
somewhat constricted towards their middle part, strong, as long as the
radius of the outer shell, as broad as one of its meshes. Radial proportion
of the three spheres = 1 : 2 : 6. Pores of the two inner shells regular
hexagonal, with thin bars, six to eight on the half meridian (fig. 1_a_).

_Dimensions._--Diameter of the outer shell 0.13, of the middle 0.04, of the
inner 0.02; pores of the outer shell 0.01, of the middle 0.008, of the
inner 0.004; length of the six spines 0.06, basal breadth 0.015.

_Habitat._--Equatorial Atlantic, Station 347, surface.

11. _Hexacontium prionacanthum_, n. sp. (Pl. 24, figs. 7, 7_a_).

Cortical shell thick walled, thorny; its pores regular circular, with
elevated hexagonal frames six to nine on the radius, twice as broad as the
crest-shaped bars. At each nodal-point of the hexagon arises one short
conical papilla or thorn. Radial proportion of the three spheres =
1 : 3 : 10. Pores of both medullary shells much smaller, regular circular.
Six main spines three-sided prismatic, longer than the radius of the outer
shell, as broad as one of its meshes; their three edges serrated, with ten
to twelve teeth.

_Dimensions._--Diameter of the outer shell 0.15, middle 0.045, inner 0.015;
cortical pores 0.01, bars 0.005; length of the spines 0.1, breadth 0.012.

_Habitat._--West Tropical Pacific, Station 225, depth 4475 fathoms.

12. _Hexacontium clavigerum_, n. sp. (Pl. 23, fig. 5).

Cortical shell very thick walled, thorny; its pores regular circular,
hexagonally framed, three times as broad as the bars, nine to twelve on the
radius; between them short conical papillæ or thorns arising at the
nodal-points. Radial proportion of the three spheres = 1 : 2 : 5. Outer
medullary shell with regular circular, much smaller pores; connected with
the cortical shell by numerous (twenty to thirty) thin radial beams. Six
main spines very stout, club-shaped, scarcely as long as the radius of the
outer shell, three times as broad as one of its pores, three-sided, with
three to four teeth on each edge.

_Dimensions._--Diameter of the outer shell 0.2, of the middle 0.08, of the
inner 0.04; cortical pores 0.012, bars 0.004; length of the six spines
0.08, breadth 0.03.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

13. _Hexacontium floridum_, n. sp. (Pl. 24, fig. 4).

Cortical shell papillose, thick walled. Pores regular, six-lobed, four
times as broad as the bars, five to seven on the radius; each pore with six
(sometimes also five or seven) concave indentations or lobules; between the
pores at the nodal-points arise short conical papillæ or thorns. Radial
proportion of the three spheres = 1 : 2 : 3. Radial main spines
pommel-shaped, three-sided prismatic, longer than the radius of the outer
shell, two to three times as broad as one pore.

{196}_Dimensions._--Diameter of the outer shell 0.12, middle 0.06, inner
0.03; cortical pores 0.012, bars 0.003; length of the six spines 0.08,
breadth 0.03.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

14. _Hexacontium hexaconicum_, n. sp.

Cortical shell spiny, thick walled. Pores regular circular, of the same
breadth as the bars, ten to twelve on the radius; between them long
bristle-shaped by-spines, half as long as the six main spines. Radial
proportion of the three spheres = 1 : 3 : 9. Six main spines conical, as
long as the radius of the outer shell, four times as broad at the base as
one pore.

_Dimensions._--Diameter of the outer shell 0.18, middle 0.06, inner 0.02;
cortical pores and bars 0.005; length of the spines 0.1, basal breadth
0.02.

_Habitat._--Antarctic Sea, Station 157, depth 1950 fathoms.

15. _Hexacontium asteracanthion_, Haeckel.

_Haliomma asteracanthion_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 816.

_Actinomma asteracanthion, Haeckel_, 1862, Monogr. d. Radiol., p. 441,
Taf. xxiii. figs. 5, 6.

Cortical shell thin walled, spiny; its pores circular, without hexagonal
frame (as a rule very regular, but in other specimens more or less
irregular), often very variable in size, three to six times as broad as the
bars, commonly seven to eight on the radius. Radial proportion of the three
spheres = 1 : 2 : 4, or sometimes 1 : 3 : 8. Pores of the two inner shells
also circular, but much smaller; those of the middle shell about half as
broad, those of the inner shell one-fourth to one-sixth as broad as the
pores of the outer shell. Six radial spines strong, three-sided prismatic,
about as long as (or somewhat longer than) the radius of the outer shell,
as broad as one of its large meshes. Accessory spines very numerous and
very thin, bristle-like, usually half as long as the six main spines. (For
the variability of this common species compare my Monograph, p. 442.)

_Dimensions._--Diameter of the outer shell 0.1 to 0.12, of the middle 0.04
to 0.06, of the inner 0.02 to 0.03; cortical pores 0.005 to 0.01; length of
the six spines 0.06 to 0.08.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

Subgenus 3. _Hexacontosa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface smooth, without radial by-spines or papillæ (other than the
six main spines).

16. _Hexacontium axophænum_, n. sp.

Cortical shell thin walled, smooth. Pores irregular polygonal, three to
nine times as broad as the bars; seven to ten on the radius. Radial
proportion of the three spheres = 1 : 3 : 8. Both {197}medullary shells
with smaller, irregular roundish pores. Six spines three-sided pyramidal,
longer than the radius of the outer shell, one to two times as broad as one
larger pore.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.045, inner 0.015;
cortical pores 0.006 to 0.018, bars 0.002; length of the spines 0.06 to
0.12, basal breadth 0.02 to 0.03.

_Habitat._--Central Pacific, Stations 266 to 272, at various depths.

17. _Hexacontium polygonale_, n. sp.

Cortical shell thick walled, smooth. Pores irregular roundish, with
polygonal frames, two to four times as broad as the bars; eight to twelve
on the radius. Radial proportion of the three spheres = 1 : 3 : 10. Six
spines pyramidal, nearly as long as the diameter of the outer shell, with
six to nine prominent edges (along the corner number of the polygonal
meshes).

_Dimensions._--Diameter of the outer shell 0.2, middle 0.06, inner 0.02;
cortical pores 0.008 to 0.016, bars 0.004; length of the spines 0.16 to
0.18, basal breadth 0.02.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

18. _Hexacontium antarcticum_, n. sp.

Cortical shell thin walled, smooth, with irregular roundish pores, eight to
ten on the radius, two to four times as broad as the bars. Radial
proportion of the three spheres = 1 : 3 : 9. Both medullary shells with
smaller roundish irregular pores. Six spines conical, pyramidal at their
origin, about as long as the radius, as broad as one larger mesh.

_Dimensions._--Diameter of the outer shell 0.18, middle 0.06, inner 0.02;
cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.1, basal
breadth 0.12.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Subgenus 4. _Hexacontura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form; surface covered with numerous conical papillæ or bristle-shaped,
radial by-spines (other than the six main spines).

19. _Hexacontium papillosum_, n. sp. (Pl. 24, fig. 5).

Cortical shell thick walled, papillose. Pores irregular roundish, often
somewhat lobed, five to six on the radius, two to four times as broad as
the bars. Radial proportion of the three spheres = 1 : 2.5 : 5. Both
medullary shells with subregular circular pores (inner much smaller). Six
radial spines short and stout, pommel or club shaped, with three prominent
wings, about as long as the shell radius, and as broad as the medullary
shell.

_Dimensions._--Diameter of the outer shell 0.1, middle 0.05, inner 0.02,
cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.05, bars
0.02.

_Habitat._--Central Pacific, Stations 266 to 272, at various depths.

{198}20. _Hexacontium gladiatum_, n. sp. (Pl. 25, fig. 8).

Cortical shell very thick walled, spiny, with irregular network; its meshes
roundish, two to five times as broad as the bars, of very different size,
ten to fifteen on the half meridian. Bars between them very strong,
three-sided prismatic, armed with a great number of small thorns, and with
large spines at the nodal-points between every three meshes. Radial
proportion of the three spheres = 1 : 3 : 12. Six strong radial main spines
three-sided prismatic, with three prominent, somewhat contorted wings,
acute, broader than the diameter of the inner shell and as long as the
diameter of the outer shell. These six main spines are situated in the same
three dimensive axes as the six thin radial beams connecting the two
medullary shells. But the six radial beams which connect the middle with
the outer shell alternate with the former and lie in three other dimensive
axes,--a very rare and remarkable disposition.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.05, inner 0.016;
cortical pores 0.01 to 0.03, bars 0.006; length of the six spines 0.2,
breadth 0.02 to 0.03.

_Habitat._--Tropical Atlantic, Station 342, depth 1445 fathoms.

21. _Hexacontium setosum_, n. sp.

Cortical shell thin walled, with numerous bristle-shaped, simple, radial
by-spines of variable length. Pores irregular roundish, five to seven on
the radius, two to eight times as broad as the bars. Radial proportion of
the three spheres = 1 : 3 : 9. Main spines three-sided pyramidal, as long
as the radius.

_Dimensions._--Diameter of the outer shell 0.1 to 0.015, middle 0.035 to
0.05, inner 0.013 to 0.016; cortical pores 0.01 to 0.03, bars 0.003 to
0.004; length of the six spines 0.05 to 0.08, basal breadth 0.012 to 0.015.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface; also
fossil in Barbados.

22. _Hexacontium furcatum_, n. sp.

Cortical shell thin walled, covered with numerous thin, bristle-shaped
by-spines, which are forked and nearly as long as the main spines. Pores
irregular roundish, eight to ten on the radius, two to eight times as broad
as the bars. Radial proportion of the three spheres = 1 : 2.5 : 10. Main
spines triangular pyramidal, shorter then the radius.

_Dimensions._--Diameter of the outer shell 0.18, middle 0.045, inner 0.018;
cortical pores 0.005 to 0.015, bars 0.002; length of the six spines 0.07,
basal breadth 0.015.

_Habitat._--North Atlantic, Station 354, surface.

23. _Hexacontium drymodes_, Haeckel.

_Actinomma drymodes_, Haeckel, 1862, Monogr. d. Radiol., p. 442, Taf.
xxiv. fig. 9.

Cortical shell thin walled, covered with numerous thin, bristle-shaped
spines, which are double forked and half as long as the main spines. Pores
irregular roundish, eight to ten on the radius, {199}two to eight times as
broad as the bars. Radial proportion of the three spheres = 1 : 2 : 4. Main
spines three-sided prismatic, with prominent, often somewhat contorted
edges, at the distal end cuspidated; longer than the radius of the outer
shell.

_Dimensions._--Diameter of the outer shell 0.15, middle 0.07, inner 0.035;
cortical pores 0.01 to 0.03, bars 0.04; length of the six spines 0.11,
basal breadth 0.03.

_Habitat._--Mediterranean (Messina), surface; Canary Islands (Lanzerote);
Haeckel.

24. _Hexacontium periplectum_, n. sp.

Cortical shell double, enclosing a simple medullary shell. Radial
proportion of the three shells = 1 : 4 : 5. Inner cortical shell thick
walled, with irregular roundish pores, two to eight times as broad as the
bars; five to seven on the radius. Numerous radial spines, arising from it,
are connected below their distal ends (at equal distances from the centre)
by delicate branched threads, and so form an outer, irregular, thin,
cortical shell, with spiny surface. Six main spines three-sided pyramidal,
about as long as the radius.

_Dimensions._--Diameter of the outer shell 0.22, middle 0.2, inner 0.045;
cortical pores (of both outer shells) 0.01 to 0.03, bars 0.001 to 0.005;
length of the spines 0.1, basal breadth 0.02.

_Habitat._--North Pacific, Station 241, surface.

Genus 80. _Hexadendron_,[111] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Cubosphærida# with three concentric lattice-spheres and six
branched spines of equal size.

The genus _Hexadendron_ differs from _Hexacontium_ in the ramification of
the six dimensive spines, and from the similar _Hexancistra_ in the
duplication of the medullary shell. As in the latter instance, each spine
can bear either three simple lateral branches or three rows of pinnate
lateral branches.

1. _Hexadendron quadricuspis_, n. sp.

All three shells spherical, with radial proportion = 1 : 2 : 6. Pores of
both medullary shells regular circular, twice as broad as the bars. Pores
of the cortical shell irregular roundish, four to six times as broad as the
bars; surface a little thorny. Six radial spines prismatic, with three
prominent wing-like edges, which are prolonged below the distal end in
three curved lateral branches. (Differs from _Hexalonche quadricuspis_, Pl.
22, fig. 11, mainly in the double medullary shell.)

_Dimensions._--Diameter of the outer shell 0.15, middle 0.05, inner 0.025;
length of the spines 0.12, breadth 0.01.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

{200}2. _Hexadendron bipinnatum_, n. sp. (Pl. 23, fig. 1).

All three shells regular octahedral, with very delicate network of
irregular polygonal meshes, and very thin, thread-like bars between them;
their radial proportion = 1 : 2.5 : 7.5. Surface of the cortical shell
covered with numerous bristle-shaped by-spines, as long as the radius. Six
main spines very large, three-sided prismatic, with three rows of pinnate,
lateral branches, on the three wing-like, spirally twisted edges (similar
to those of _Hexancistra mirabilis_, p. 189, Pl. 23, fig. 3).

_Dimensions._--Diameter of the outer shell 0.12, middle 0.04, inner 0.016.

_Habitat._--Central Pacific, Station 272, surface.

Genus 81. _Hexacontarium_,[112] n. gen.

_Definition._--#Cubosphærida# with three concentric lattice-spheres and six
simple spines of different size; one opposite pair larger than the two
others.

The genus _Hexacontarium_ exhibits to its ancestral form _Hexacontium_ the
same relation that _Hexaloncharium_ bears to _Hexalonche_; the former is
developed from the latter by duplication of the medullary shell. As two
opposite spines of one pair are larger than the four others, they
correspond to the three axes of a quadratic crystal.

1. _Hexacontarium dentatum_, n. sp.

Cortical shell with regular circular, hexagonally framed pores, twice as
broad as the bars, with smooth surface. Radial proportion of the three
spheres = 1 : 2 : 5. Six radial spines three-sided prismatic, half as broad
as the inner medullary shell, with three dentated edges. Two opposite major
spines longer than the shell diameter; four other minor spines only half as
long as the radius. (Similar to _Hexacontium clavigerum_, Pl. 23, fig. 5,
but distinct by the prolongation of the spines of one axis.)

_Dimensions._--Diameter of the outer shell 0.15, middle 0.06, inner 0.03;
length of the major spines 0.2, minor 0.04.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

2. _Hexacontarium clavatum_, n. sp.

Cortical shell covered with short conical by-spines and irregular roundish
pores, three to four times as broad as the bars. Radial proportion of the
three spheres = 1 : 3 : 8. Six radial spines three-sided prismatic, at the
distal end club-shaped; two major spines three times as long as the four
others, which are equal to the shell radius.

_Dimensions._--Diameter of the outer shell 0.16, middle 0.06, inner 0.02.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

{201}Subfamily HEXACROMYIDA,[113] Haeckel, 1881, Prodromus, pp. 449, 453.

_Definition._--#Cubosphærida# with four concentric spherical
lattice-shells.

Genus 82. _Hexacromyum_,[114] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Cubosphærida# with four concentric lattice-spheres and six
simple spines of equal size.

The genus _Hexacromyum_ possesses four concentric, spherical, or octahedral
lattice-shells; two inner medullary shells within the central capsule, two
outer cortical outside it. The four spheres are connected by six radial
beams, which are prolonged outside into simple spines of equal size,
opposite in pairs in the three dimensive axes. This genus can be derived
from _Hexacontium_ by duplication of the cortical shell.

1. _Hexacromyum elegans_, n. sp. (Pl. 24, fig. 9).

Shell composed of four concentric shells, with radial proportion =
1 : 2.5 : 7.5 : 10. First (innermost) shell with very small circular pores,
second shell with larger circular pores. Third shell (inner cortical shell)
with large, subregular, circular, hexagonally framed pores (eight to nine
on the radius), twice as broad as the bars; from the elevated nodal-points
of the hexagonal frames (between every three pores) arise thin
bristle-shaped radial beams, which are united at the distal end by vaulted
branches forming the delicate fourth shell. Surface smooth. Radial spines
three-sided pyramidal, as long as the radius, as broad at the base as the
innermost shell.

_Dimensions._--Diameter of the four shells--(A) 0.02, (B) 0.05, (C) 0.15,
(D) 0.2; length of the six radial spines 0.1, basal breadth 0.02.

_Habitat._--Central Pacific, Station 271, surface.

2. _Hexacromyum quadrigatum_, n. sp.

Shell composed of four concentric shells, with radial proportion =
1 : 3 : 8 : 10. Structure of all four shells the same, with regular,
circular pores, twice to three times as broad as the bars; size of the
pores gradually increasing from the innermost to the outermost shell;
surface smooth. Radial spines three-sided pyramidal, as long as the radius,
half as broad at the base as the innermost shell.

_Dimensions._--Diameter of the four shells--(A) 0.025, (B) 0.08, (C) 0.20,
(D) 0.25; length of the spines 0.12, basal breadth 0.012.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

{202}3. _Hexacromyum arachnoides_, n. sp.

_Hexacromidium arachnoides_, Haeckel, 1881, Prodromus, p. 453.

Shell composed of four concentric shells, with radial proportion =
1 : 2 : 3 : 4. Innermost shell with regular, hexagonal meshes; the three
other shells with irregular, polygonal meshes; bars between the large
meshes in all four shells very thin, cobweb-like. Surface covered with thin
bristle-shaped by-spines, as long as the radius. Six radial main spines
three-sided prismatic, longer than the diameter of the whole shell.

_Dimensions._--Diameter of the four shells--(A) 0.08, (B) 0.16, (C) 0.24,
(D) 0.32; length of the spines 0.4, breadth 0.01.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

4. _Hexacromyum octahedrum_, n. sp. (Pl. 23, fig. 2).

Shell composed of four concentric shells which are not spherical (as in the
three preceding species), but regular octahedral. Radial proportion =
1 : 2.5 : 6 : 9. Network in all four shells delicate, with irregular
polygonal meshes and thin bars; the thickness of the bars and size of the
meshes increasing from the innermost to the outermost shell. Six radial
spines three-sided prismatic, increasing slowly in breadth towards the
distal end, much longer than the shell diameter.

_Dimensions._--Diameter of the four shells--(A) 0.02, (B) 0.05, (C) 0.12,
(D) 0.18; length of the radial spines 0.2 to 0.3 and more, breadth 0.01.

_Habitat._--Central Pacific, Station 263, depth 2650 fathoms.

Subfamily HEXACARYIDA,[115] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Cubosphærida# with five or more concentric lattice-shells.

Genus 83. _Cubosphæra_,[116] n. gen.

_Definition._--#Cubosphærida# with five to six or more concentric
lattice-shells and six simple spines of equal size.

The genus _Cubosphæra_ is developed from the preceding _Hexacromyum_ by
further addition of the concentric lattice-shells, their number amounting
to five, six, or more. The innermost two of these are medullary shells, the
others being cortical shells. All are connected by six radial beams,
prolonged outside into six simple spines of equal size; these lie opposite
in pairs in three dimensive axes, corresponding to the three axes of a
cube.

{203}1. _Cubosphæra cubaxonia_, n. sp. (Pl. 24, fig. 8).

_Hexacromyon cubaxonium_, Haeckel, 1881, Prodrom. et Atlas.

Shell composed of five concentric spheres, with the radial proportion =
1 : 3 : 8 : 10 : 13. The two medullary shells with small regular, circular
pores of the same breadth as the bars. Inner cortical shell (third shell)
with large regular, circular pores, four times as broad as the bars,
hexagonally framed. From each hexagon-corner arises a thin bristle-shaped
radial by-spine, which at a fixed distance from the centre is united with
the middle cortical shell (fourth shell), which has very small circular
pores. The beginning of the fifth shell (outermost) is indicated by six
small reticula, produced by the six main spines at equal distances from the
centre. All five shells are united by six prismatic radial beams, ending
outside in pyramidal furrowed spines.

_Dimensions._--Diameter of the five shells--(A) 0.2, (B) 0.15, (C) 0.12,
(D) 0.045, (E) 0.015.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Cubosphæra concentrica_, n. sp.

Shell composed of six concentric spheres, with the radial proportion =
1 : 2 : 6 : 7.5 : 9 : 11. The two medullary shells with regular, circular
pores, twice as broad as the bars. The four cortical shells of the same
structure, with irregular, roundish pores, three to four times as broad as
the bars. The size of these pores and the breadth of their bars gradually
increase towards the smooth surface. All six shells are connected by six
thin three-sided prismatic radial beams, which are prolonged outside in six
stronger spines, angular pyramidal, with smooth edges, as long as the shell
radius.

_Dimensions._--Diameter of the six shells--(A) 0.02 (B) 0.04, (C) 0.12, (D)
0.15, (E) 0.18, (F) 0.22.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Genus 84. _Hexacaryum_,[117] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Cubosphærida# with five to six or more concentric
lattice-shells and six branched spines of equal size.

The genus _Hexacaryum_ is distinguished from the foregoing _Cubosphæra_ by
ramification of the six radial spines, and therefore exhibits the same
relation to it that _Hexancistra_ bears to _Hexastylus_, &c.

1. _Hexacaryum arborescens_, n. sp. (Pl. 23, figs. 4, 4_a_).

Shell composed of five, six, or more concentric shells, which are united by
six very large radial spines. The two medullary shells spherical, inner
with regular, circular, outer with irregular polygonal pores. All cortical
shells (third and following) not spherical, but regular octahedral, with
irregular polygonal meshes and thin bars. Radial spines prismatic, with
three wing-like, spirally twisted edges, which at equal distances send out
thin forked lateral branches (six on each {204}verticil); by further
ramification and communication of these branches the triangular net-plates
arise, filling out the sides of the octahedral cortical shells. Diameter of
all shells little different. The outer free parts of the six spines are
arborescent, twice to three times as long as the enclosed inner parts, and
bear six to eight verticils of free lateral branches, decreasing in size
towards the distal end (similar to _Arachnosphæra_).

_Dimensions._--Diameter of the six shells--(A) 0.015, (B) 0.04, (C) 0.1,
(D) 0.16, (E) 0.22, (F) &c.; average distance of the concentric octahedra =
0.06; length of the radial spines 0.3 to 0.4 and more, breadth 0.02.

_Habitat._--Central Pacific, Station 274, surface.

Subfamily HEXADORIDA,[118] Haeckel, 1881, Prodromus, pp. 449, 455.

_Definition._--#Cubosphærida# with spongy spherical or octahedral shell
(with or without enclosed concentrical lattice-shells).

Genus 85. _Cubaxonium_,[119] n. gen.

_Definition._--#Cubosphærida# with solid spongy spherical or octahedral
shell, without latticed medullary shell in the centre, and with six simple
radial spines of equal size.

The genus _Cubosphæra_ may be regarded as a form of _Styptosphæra_, which
develops six radial spines, opposite in pairs in the three dimensive axes.
The solid spongy framework of the shell assumes the outer form either of a
sphere, or of a regular octahedron.

1. _Cubaxonium spongiosum_, n. sp.

Spongy shell spherical, composed of a very dense spongy framework of nearly
uniform structure; the meshes three to four times as broad as the bars;
surface almost smooth. Six spines cylindrical, twice to three times as long
as the diameter of the spongy sphere, about as broad as one half mesh.

_Dimensions._--Diameter of the sphere 0.2, meshes 0.006 to 0.008, bars
0.002; length of the spines 0.4 to 0.5, breadth 0.004.

_Habitat._--South Pacific, Station 288, surface.

2. _Cubaxonium octahedrum_, n. sp.

Spongy shell octahedral, composed of a loose spongy framework of nearly
uniform structure; the meshes ten to twelve times as broad as the bars;
surface thorny. Six spines three-sided {205}pyramidal, longer than the
diameter of the octahedron, arising from its six corners, as broad at the
base as one mesh.

_Dimensions._--Diameter of the shell 0.15, meshes 0.01 to 0.012, bars
0.001; length of the spines 0.2, basal breadth 0.01.

_Habitat._--South Pacific, Station 300, surface.

Genus 86. _Hexadoras_,[120] Haeckel, 1881, Prodromus, p. 455.

_Definition._--#Cubosphærida# with spongy spherical shell and one simple
latticed medullary shell in its centre, having six simple spines of equal
size.

The genus _Hexadoras_ exhibits the same structure of the spongy shell as
_Spongoplegma_, but differs from it in the production of six dimensive
spines. In the centre lies one simple, latticed, medullary shell, which is
either spherical or octahedral. In the latter case the six spines arise
from the six corners of the octahedron.

1. _Hexadoras axophæna_, n. sp.

Medullary shell spherical, with regular hexagonal meshes, twice as broad as
the bars. Cortical shell entirely enclosing it, with loose irregular
framework and thorny surface. Diameter of the outer shell three times as
large as that of the inner. Six radial spines arising from the medullary
shell, four to five times as long as the radius of the cortical shell,
three-sided prismatic, with straight dentated edges.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; length of the
spines 0.3 to 0.4.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

2. _Hexadoras lychnosphæra_, n. sp.

Medullary shell spherical, with regular, circular, hexagonally framed
pores, three times as broad as the bars. Cortical shell enveloping it, with
loose irregular framework and spiny surface. Diameter of the outer shell
eight times as large as that of the inner. Six radial spines arising from
the inner shell, three-sided prismatic, with dentated, spirally contorted
edges, seven times as long as the diameter of the medullary shell, with
three lateral branches at the distal end (similar to _Lychnosphæra_, Pl.
11, fig. 1).

_Dimensions._--Diameter of the outer shell 0.5, inner 0.06; total length of
the spines 0.4, breadth 0.01.

_Habitat._--Central Pacific, Station 270, depth 2925 fathoms.

3. _Hexadoras octahedrum_, n. sp.

Medullary shell regular octahedral, with irregular polygonal meshes, five
times as broad as the bars. Cortical shell enveloping it, with dense spongy
framework, also octahedral with rough surface. {206}Diameter of the outer
shell five times as large as that of the inner. Six radial spines very
long, arising from the six corners of the inner and piercing the spongy
mass of the outer shell, considerably exceeding it at the free distal end,
three-sided prismatic, with elegantly denticulate edges.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.04; total length of
the spines 0.3 or more, breadth 0.02.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

Genus 87. _Hexadoridium_,[121] Haeckel, 1881, Prodromus, p. 455.

_Definition_.--#Cubosphærida# with spongy spherical shell and two
concentric latticed medullary shells in its centre, having six simple
spines of equal size.

The genus _Hexadoridium_ differs from _Hexadoras_ in the duplication of the
medullary shell, and exhibits therefore the same relation to it that
_Spongodictyon_ bears to _Spongoplegma_. In the only known species the six
spines are opposite, arranged quite regularly in pairs in the three
dimensive axes, and consequently represent the three axes of a regular
crystal or cube.

1. _Hexadoridium streptacanthum_, n. sp. (Pl. 25, figs. 1, 1_a_).

Both medullary shells spherical, with small regular, circular pores, twice
as broad as the bars; outer twice as broad as the inner. Spongy cortical
shell enclosing it with dense framework, five times as broad as the outer
medullary shell, regular octahedral. Six radial spines, arising from the
latter, are thinned at the inner end, three to five times as long as the
diameter of the cortical shell, and nearly as broad as the inner medullary
shell, with three dentated and spirally contorted edges. (Very similar to
the common _Spongosphæra streptacantha_, with irregular and variable number
and dispositions of spines; possibly its ancestral form?).

_Dimensions._--Diameter of the cortical shell 0.2, of the outer medullary
shell 0.04, inner 0.02; length of the spines 1 mm. and more, breadth 0.02.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Family X. #ASTROSPHÆRIDA# (Pls. 11, 18-20, 26-30).

_Astrosphærida_, Haeckel, 1881, Prodromus, p. 449.

_Definition._--#Sphæroidea# with numerous (eight to twelve or more,
commonly between twenty and sixty) radial spines on the surface of the
spherical shell; living solitary (not associated in colonies).

The family #Astrosphærida#, the largest and most varied of all
#Sphæroidea#, is distinguished from the other members of this group by the
possession of numerous {207}radial spines, which are either regularly or
irregularly disposed on the surface of the spherical shell. The extreme
variability and richness of form in this family is mainly due to the
different size, shape, and disposition of these radial spines.

The simplest Astrosphærida are the Coscinommida, with a single spherical or
polyhedral lattice-shell. To this ancestral group all other subfamilies can
be opposed as "Astrosphærida composita," since their skeleton is composed
of two or more concentric lattice-shells: two in the Haliommida, three in
the Actinommida, four in the Cromyommida, five or more in the Caryommida.
In these four subfamilies the concentric shells are all simple (not spongy)
fenestrated spheres or endospherical polyhedra. In the sixth subfamily, the
Spongiommida, the shell is wholly or partially composed of spongy irregular
wicker-work, with or without a medullary shell in the centre.

_The Number of the Radial Spines_ in the Astrosphærida is extremely
variable, and ranges from eight to forty or more; in many cases more than
one hundred. Often each nodal-point of the network develops on the shell
surface one spine. Still more frequently the number of the spines is less
than that of the nodal-points. In all concentric Astrosphærida, having two
or more concentrical shells, we can distinguish "primary spines," as outer
prolongations of the inner radial beams connecting the shells, and
"secondary spines," developed only on the outer surface of the shell.
Naturally the former are of much greater importance than the latter. But we
can also often distinguish among the latter larger "main spines" and
smaller "by-spines," the latter commonly much more numerous than the
former.

_The Disposition of the Radial Spines_, either regular or irregular, is a
subject of great morphological interest, and remains to be exhausted by
further observations. The following cases of regular disposition have been
observed by me--(A) eight spines, opposite in pairs in four axes
corresponding to the four diagonal axes of a cube; (B) nine spines,
regularly disposed at equal distances (?) (not opposed in pairs); (C) ten
spines, disposed at equal distances (?); (D) twelve spines, regularly
disposed, corresponding to the twelve corners of the regular icosahedron;
(E) fourteen spines, quite regularly disposed (six corresponding to the
three axes of a regular octahedron, eight to the central points of its
eight faces); (F) sixteen spines, regularly disposed (?); (G) twenty spines
(very common!), either disposed in the same manner (after the law of
Johannes Müller) as in the ACANTHARIA (?), or corresponding to the twenty
corners of the regular or pentagonal dodecahedron, or disposed in the same
manner as in many #Larcoidea# (Tholonida, &c., to be described afterwards);
(H) twenty-four spines, regularly disposed (?); (I) thirty-two spines,
quite regularly disposed (twenty corresponding to the twenty corners of the
regular dodecahedron, twelve to the central points of its twelve faces);
(K) forty spines, nearly regularly (or quite symmetrically?) disposed. If
the number of the spines amounts to more than forty, it is as a rule
impossible to determine their regular disposition in a satisfactory manner.

{208}_Synopsis of the Genera of Astrosphærida._

-------------------------------------------------------------------------
I. Subfamily Coscinommida.
(One single latticed shell.)
-------------------------------------------------------------------------
{ All spines of the same
Spines all simple, not { kind, 88. _Acanthosphæra_.
branched and not {
tubulous. { Larger main spines and
{ smaller by-spines, 89. _Heliosphæra_.

Spines hollow conical tubes with porous walls, 90. _Conosphæra_.

Between simple spines the pores prolonged in hollow
tubes, 91. _Coscinomma_.

{ Spines with lateral
{ branches, 92. _Cladococcus_.
Spines branched or forked.{
{ Spines forked or
{ dichotomous, 93. _Elaphococcus_.
-------------------------------------------------------------------------
II. Subfamily Haliommida.
(Two concentric latticed shells.)
-------------------------------------------------------------------------
{ All spines equal,
{ simple, 94. _Haliomma_.
{
A. Elatommida, one { All spines simple, of
medullary and one cortical{ two different kinds, 95. _Heliosoma_.
shell. {
{ Spines branched, 96. _Elatomma_.

{ No by-spines, 97. _Leptosphæra_.
{
B. Diplosphærida, both { Inner by-spines, 98. _Diplosphæra_.
shells cortical. {
{ Outer by-spines, 99. _Drymosphæra_.
{
{ Outer and inner
{ by-spines, 100. _Astrosphæra_.
-------------------------------------------------------------------------
III. Subfamily Actinommida.
(Three concentric latticed shells.)
-------------------------------------------------------------------------
{ All spines equal, 101. _Actinomma_.
Radial spines not {
branched. { Larger main spines and
{ smaller by-spines, 102. _Echinomma_.

Radial spines branched or forked, 103. _Pityomma_.
-------------------------------------------------------------------------
IV. Subfamily Cromyommida.
(Three concentric latticed shells.)
-------------------------------------------------------------------------
{ All spines equal, 104. _Cromyomma_.
Radial spines not {
branched. { Larger main spines and
{ smaller by-spines, 105. _Cromyechinus_.

Radial spines branched or forked, 106. _Cromyodrymus_.
-------------------------------------------------------------------------
V. Subfamily Caryommida.
(Numerous, five to ten or more, concentric latticed shells.)
-------------------------------------------------------------------------
Two medullary shells and three or more cortical
shells (lattice work ordinary), 107. _Caryomma_.

{ Triangular meshes,
No medullary shells { simple bars, without
(lattice work { diagonal bars, 108. _Arachnopila_.
arachnoidal); three-sided {
prismatic spines, { Triangular meshes,
with verticils of three { simple bars; diagonal
forked branches. { bars between the 109. _Arachnopegma_.
(Arachnosphærida.) { shells,
{
{ Polygonal meshes, bars
{ branched, 110. _Arachnosphæra_.

-------------------------------------------------------------------------
VI. Subfamily Spongiommida.
(Spherical shell whole or partly spongy, with or without enclosed
latticed medullary shells in the centre.)
-------------------------------------------------------------------------
I. Tribe {Spongy {Spines simple, 111. _Spongiomma_.
Spongodrymida, { sphere {
without { solid. {Spines branched, 112. _Spongodrymus_.
latticed {
medullary {Spongy {Spines simple, 113. _Spongechinus_.
shell. { sphere {
{ hollow. {Spines branched, 114. _Spongothammus_.

{Framework arising
{ from the medullary
{ shell, 115. _Spongopila_.
{
{Medullary {Framework {No medullary by-spines,
II. Tribe { shell { separate { 116. _Rhizoplegma_.
Rhizoplegmida, { spherical. { from the {
with { { medullary {Medullary by-spines,
one single { { shell. { 117. _Lychnosphæra_.
latticed {
medullary {Medullary {Framework arising
shell. { shell a { immediately from
{ simple { the medullary
{ cube. { shell, 118. _Centrocubus_.
{ {
{ {Framework separate
{ { from the medullary
{ { shell, 119. _Octodendron_.

{Framework arising
III. Tribe Rhizosphærida, { from the medullary
with two concentric { shell, 120. _Spongosphæra_.
latticed medullary shells. {
{Framework separate
{ from the medullary
{ shell, 121. _Rhizosphæra_.

Subfamily COSCINOMMIDA, Haeckel.

_Heliosphærida_, Haeckel, 1881, Prodromus, pp. 449, 450.

_Definition._--#Astrosphærida# with one single spherical lattice-shell.

Genus 88. _Acanthosphæra_,[122] Ehrenberg, 1858, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 12.

_Definition._--#Astrosphærida# with one simple lattice-sphere, covered with
simple radial spines of the same kind.

The genus _Acanthosphæra_ exhibits the most simple form of all
_Astrosphærida_; a simple spherical lattice-shell, the surface of which is
covered by radial spines of one and the same kind. The number of the latter
is very variable, often twelve to twenty, regularly disposed; in other
cases forty to sixty or more; and sometimes at each nodal-point of the
network a spine is developed.

{210}Subgenus 1. _Rhaphidococcus_, Haeckel, 1862, Monogr. d. Radiol., p.
365 (_sensu emendato_).

_Definition._--Pores of the spherical shell regular or subregular, all of
nearly equal size and similar form. Radial spines arising from all the
nodal-points of the network.

1. _Acanthosphæra tenuissima_, Haeckel.

_Heliosphæra tenuissima_, Haeckel, 1862, Monogr. d. Radiol., p. 351, Taf.
ix. fig. 2.

Shell extremely thin walled, eight to ten times as broad as one pore.
Meshes or pores regular, hexagonal, with thread-like bars; five to seven on
the radius. At each nodal-point (between every three meshes) arises a
bristle-shaped radial spine, as long as the diameter of one pore.

_Dimensions._--Diameter of the shell 0.2 to 0.25, of the meshes or pores
0.025 to 0.03, bars below 0.0001; length of the spines 0.03.

_Habitat._--Mediterranean (Messina), North Atlantic (Canary Islands).

2. _Acanthosphæra tenuis_, n. sp.

Shell very thin walled, about twenty times as broad as one pore. Meshes
subregular, hexagonal, with thread-like bars; twelve to fourteen on the
radius. At each nodal-point arises a bristle-shaped radial spine, about as
long as the radius.

_Dimensions._--Diameter of the shell 0.3 to 0.35, pores 0.015 to 0.018,
bars below 0.001; length of the spines 0.12 to 0.16.

_Habitat._--Central Pacific, Stations 266 to 274, depth 2350 to 2925
fathoms.

3. _Acanthosphæra macropora_, n. sp.

Shell thin walled, five to six times as broad as one mesh. Pores regular,
circular, hexagonally framed, ten to twelve times as broad as the bars.
Radial spines bristle-shaped, as long as the diameter of one pore, arising
from all the nodal-points.

_Dimensions._--Diameter of the shell 0.07 to 0.08, pores 0.012, bars 0.001;
length of the spines 0.012.

_Habitat._--Central Pacific, Station 271, surface.

4. _Acanthosphæra micropora_, n. sp.

Shell thick walled, forty to fifty times as broad as one pore. Pores
regular, circular, hexagonally framed, half as broad as the bars. Radial
spines bristle-shaped, as long as the radius of the shell, arising from all
the nodal-points.

_Dimensions._--Diameter of the shell 0.22, pores 0.003, bars 0.006; length
of the spines 0.1.

_Habitat._--South Pacific, Station 288, surface.

{211}5. _Acanthosphæra dentata_, Haeckel.

_Cladococcus dentatus_, Haeckel, 1862, Monogr. d. Radiol., p. 367, Taf.
xiii. fig. 10.

Shell thin walled, eight to ten times as broad as one pore. Pores regular,
circular, hexagonally framed, three times as broad as the bars. Radial
spines arising from all the nodal-points of the network, three-sided
prismatic, with dentated or serrated edges, longer than the shell diameter.

_Dimensions._--Diameter of the shell 0.08, pores 0.01, bars 0.003; length
of the spines 0.1, breadth 0.003.

_Habitat._--Mediterranean (Messina), surface.

6. _Acanthosphæra acufera_, Haeckel.

_Rhaphidococcus acufer_, Haeckel, 1862, Monogr. d. Radiol., p. 366, Taf.
xiv. fig. 1.

_Cladococcus acufer_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 800.

Shell thin walled, eight to ten times as broad as one mesh. Pores regular,
circular, three times as broad as the bars. Radial spines bristle-shaped,
arising with thicker conical bases from all the nodal-points, about as long
as the diameter of the shell.

_Dimensions._--Diameter of the shell 0.08, pores 0.01, bars 0.003; length
of the spines 0.08.

_Habitat._--Mediterranean (Messina); North Atlantic, Station 354, surface.

7. _Acanthosphæra castanea_, n. sp. (Pl. 26, fig. 3).

Shell thick walled, thirty times as broad as one mesh. Pores regular,
circular, nearly of the same breadth as the bars. Radial spines
bristle-shaped, arising with thicker conical bases from all the
nodal-points, scarcely half as long as the radius of the shell.

_Dimensions._--Diameter of the shell 0.15, pores and bars 0.005; length of
the spines 0.03.

_Habitat._--North Pacific, Station 244, surface.

8. _Acanthosphæra flosculenta_, n. sp.

Shell thick walled, thirteen times as broad as one mesh. Pores regular,
circular, three times as broad as the bars, with an elegant six-lobed frame
(Pl. 28, fig. 1_b_). In the intervals between the six lobes of each mesh
arise six conical radial spines (half as long as the radius), six around
each pore. (Differs from the similar _Haliomma flosculentum_, Pl. 28, fig.
1, in the absence of an enclosed medullary shell and the stronger
development of the spines.)

_Dimensions._--Diameter of the shell 0.13, pores 0.01, bars 0.003; length
of the spines 0.03.

_Habitat._--Indian Ocean, Cocos Islands, surface, Rabbe.

Subgenus 2. _Rhaphidocapsa_, Haeckel.

_Definition._--Pores of the spherical shell regular or subregular, all of
nearly equal size and similar form. Radial spines scattered at some
distance apart, not at all the nodal-points.

{212}9. _Acanthosphæra insignis_, Haeckel.

_Heliosphæra insignis_, R. Hertwig, 1879, Organismus der Radiol., p. 40,
Taf. v. fig. 7.

Shell thin walled, about ten times as broad as one mesh. Pores regular,
hexagonal, ten to twelve times as broad as the bars. Radial spines about
one hundred and twenty, arising from certain nodal-points of the network,
being as long as the diameter of the sphere, three-sided prismatic, with
three thin denticulated edges.

_Dimensions._--Diameter of the shell 0.2, pores 0.02, bars 0.002; length of
the spines 0.18, breadth 0.003.

_Habitat._--Cosmopolitan; Mediterranean (Messina), Atlantic (Stations 325,
347), Indian (Ceylon), Pacific (Stations 270 to 274), surface.

10. _Acanthosphæra fortispina_, n. sp.

Shell thin walled, about six times as broad as one mesh. Pores subregular,
hexagonal, with thread-like bars; three to four on the radius. Radial
spines about twenty, three-sided pyramidal, as long as the diameter, and
one-third as broad at the base as one pore.

_Dimensions._--Diameter of the shell 0.06 to 0.09, pores 0.01 to 0.014,
bars below 0.001; length of the spines 0.07 to 0.08, basal breadth 0.03 to
0.04.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

11. _Acanthosphæra mucronata_, n. sp. (Pl. 26, fig. 7).

Shell thick walled, fifteen times as broad as one pore. Pores regular,
circular, hexagonally framed, funnel-shaped, three times as broad as the
bars. Radial spines twenty to thirty, dagger-shaped or spindle-shaped,
angular, twice as broad in the middle as one pore, about as long as the
radius.

_Dimensions._--Diameter of the shell 0.11, pores 0.0075, bars 0.0025;
length of the spines 0.06, breadth 0.015.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

12. _Acanthosphæra clavata_, n. sp. (Pl. 26, fig. 8).

Shell thick walled, ten to twelve times as broad as one mesh. Pores
subregular, circular, three times as broad as the bars. Radial spines
twenty, club-shaped, angular, with prominent edges, twice as broad at the
distal end as one pore, about as long as the radius.

_Dimensions._--Diameter of the shell 0.11, pores 0.01, bars 0.003; length
of the spines 0.06, breadth 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

13. _Acanthosphæra marginata_, n. sp.

Shell thick walled, twelve times as broad as one mesh. Pores regular,
circular double-edged, four times as broad as the bars. Radial spines
conical, fourteen in number, about as long as the {213}radius, and as broad
at the base as one mesh. Six spines correspond to the six corners of a
regular octahedron, eight to the centre of the eight faces.

_Dimensions._--Diameter of the shell 0.1, pores 0.008, bars 0.002; radial
spines 0.06, basal breadth 0.01.

_Habitat._--Central Pacific, Station 274, surface.

14. _Acanthosphæra florida_, n. sp.

Shell thick walled, fifteen times as broad as one mesh. Pores regular,
circular, hexagonally lobed, separated by prominent funnel-shaped crests of
half the breadth. Twenty conical radial spines, half as long as the radius,
as broad at the base as one funnel.

_Dimensions._--Diameter of the shell 0.15, pores 0.01, bars 0.005; length
of the spines 0.04, basal breadth 0.02.

_Habitat._--South Atlantic, Station 325, depth 2650 fathoms.

15. _Acanthosphæra enneacantha_, n. sp.

Shell thin walled, fourteen times as broad as one mesh. Pores regular,
circular, twice as broad as the bars. Nine radial spines, regularly
disposed, as long as the shell diameter, three-sided prismatic, with
pyramidal apex, as broad as one mesh.

_Dimensions._--Diameter of the shell 0.14, pores 0.01, bars 0.005; length
of the spines 0.15, breadth 0.01.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

16. _Acanthosphæra octahedralis_, n. sp.

Shell thick walled, octahedral, fourteen times as broad as one mesh. Pores
regular, circular, three times as broad as the bars. Fourteen radial
spines, regularly disposed, conical, as long as the radius of the shell,
twice as broad at the base as one mesh. Six spines correspond to the six
corners of a regular octahedron, eight to the central points of its eight
faces.

_Dimensions._--Diameter of the shell 0.14, pores 0.01, bars 0.003; length
of the spines 0.08, basal breadth 0.02.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

17. _Acanthosphæra compacta_, n. sp.

Shell thick walled, about sixty times as broad as one mesh. Pores regular,
circular, twice as broad as the bars. Thirty to forty radial spines,
three-sided pyramidal, scarcely half as long as the radius, five to six
times as broad at the base as one pore.

_Dimensions._--Diameter of the shell 0.24, pores 0.004, bars 0.002; length
of the radial spines 0.05, basal breadth 0.02.

_Habitat._--Fossil in Barbados.

{214}Subgenus 3. _Raphidodrymus_, Haeckel.

_Definition._--Pores of the spherical shell irregular, of different size or
form. Radial spines arising from all the nodal-points of the network.

18. _Acanthosphæra capillaris_, n. sp.

Shell thin walled, with irregular polygonal meshes, twelve to twenty times
as broad as the bars; eight to ten on the radius. Radial spines
bristle-shaped, arising from all the nodal-points of the network, about as
long as the diameter of the largest meshes.

_Dimensions._--Diameter of the shell 0.15 to 0.2, pores 0.012 to 0.02, bars
0.001; length of the spines 0.02.

_Habitat._--Central Pacific, Stations 260 to 274, surface.

19. _Acanthosphæra arctica_, n. sp.

Shell thin walled, with irregular roundish, polygonally framed meshes,
three to four times as broad as the bars. Radial spines arising from all
the nodal-points of the network, pyramidal at the base; in the distal half
bristle-shaped, as long as the radius.

_Dimensions._--Diameter of the shell 0.12, pores 0.006 to 0.008, bars
0.002; length of the spines 0.07.

_Habitat._--Arctic Ocean, Greenland (in the stomach of _Periphylla
hyacinthina_).

20. _Acanthosphæra antarctica_, n. sp.

Shell thick walled, with irregular, roundish pores, about as broad as the
bars. Radial spines arising from all nodal-points of the network, conical
at the base, half as long as the radius.

_Dimensions._--Diameter of the shell 0.15, pores and bars 0.005 to 0.008;
length of the spines 0.04.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Subgenus 4. _Rhaphidosphæra_, Haeckel, 1881, Prodromus, p. 450.

_Definition._--Pores of the spherical shell irregular, of different size or
form. Radial spines scattered at intervals, not at all the nodal-points.

21. _Acanthosphæra echinoides_, Haeckel.

_Cyrtidosphæra echinoides_, Haeckel, 1865, Zeitschr. f. wiss. Zool., xv.
p. 367, Taf. xxvi. fig. 5.

Shell thin walled, with irregular polygonal or more roundish pores of very
different size. Forty to fifty very large meshes, separated by rows of much
smaller meshes. Radial spines forty to sixty, half as long as the shell
radius, bristle-shaped, with conical bases.

{215}_Dimensions._--Diameter of the shell 0.13, larger pores 0.03, smaller
0.003; length of the spines 0.03.

_Habitat._--Mediterranean (Nice), surface.

22. _Acanthosphæra longispina_, n. sp.

Shell thin walled, with irregular polygonal meshes, four to six times as
broad as the bars; six to eight on the radius. Thirty to forty radial
spines, three-sided prismatic, twice as broad as the bars, twice to three
times as long as the diameter of the shell.

_Dimensions._--Diameter of the shell 0.12 to 0.15, pores 0.012 to 0.02,
bars 0.003; length of the radial spines 0.2 to 0.4, breadth 0.006.

_Habitat._--North Pacific, Station 253, surface.

23. _Acanthosphæra brevispina_, n. sp.

Shell thick walled, with irregular polygonal meshes, twice to four times as
broad as the bars; twelve to sixteen on the radius. Sixty to eighty radial
spines, pyramidal, half as long as the radius of the shell, one-fourth as
broad at the base.

_Dimensions._--Diameter of the shell 0.2 to 0.22, pores 0.006 to 0.012,
bars 0.003; length of the spines 0.05, basal breadth 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

24. _Acanthosphæra acanthica_, Haeckel.

_Cenosphæra acanthica_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 86, Taf.
i. fig. 3.

Shell thick walled, with irregular, roundish pores, twice to three times as
broad as the bars; seven to nine on the radius. Ten to twenty radial spines
pyramidal, shorter than the radius, twice as broad at the base as one mesh.

_Dimensions._--Diameter of the shell 0.12 to 0.16, pores 0.006 to 0.009,
bars 0.003; length of the spines 0.04 to 0.06, basal breadth 0.01 to 0.015.

_Habitat._--Fossil in Tertiary rocks of Sicily and Barbados.

25. _Acanthosphæra haliphormis_, Ehrenberg.

_Acanthosphæra haliphormis_, Ehrenberg, 1861, Abhandl. d. k. Akad. d.
Wiss. Berlin, 1872, Taf. ii. fig. 1.

Shell thick walled, with irregular, roundish pores, twice to four times as
broad as the bars; four to five on the radius. Twelve to twenty radial
spines pyramidal, longer than the radius, scarcely as broad as one mesh at
the base.

_Dimensions._--Diameter of the shell 0.11, pores 0.01 to 0.02, bars 0.005;
length of the spines 0.06 to 0.08, basal breadth 0.01.

_Habitat._--Arctic Ocean, near Greenland, depth 1000 fathoms.

{216}26. _Acanthosphæra angulata_, n. sp. (Pl. 26, fig. 4).

Shell thin walled; its pores irregular, roundish, with angular,
double-edged margin, two to four times as broad as the bars; six to eight
on the radius. Twenty to thirty radial spines pyramidal, angular, with
prominent edges, shorter than the radius, as broad at the base as one small
mesh.

_Dimensions._--Diameter of the shell 0.2, pores 0.02 to 0.04, bars 0.01;
length of the spines 0.06, basal breadth 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

27. _Acanthosphæra conifera_, n. sp.

Shell thick walled, with irregular, roundish pores, twice to five times as
broad as the bars; ten to twelve on the radius. Twenty radial spines
conical, regularly disposed, half as long as the radius, as broad at the
base as one of the largest meshes.

_Dimensions._--Diameter of the shell 0.2, pores 0.008 to 0.02, bars 0.004;
length of the spines 0.05, basal breadth 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

28. _Acanthosphæra maxima_, n. sp.

Shell thick walled, with irregular, roundish pores, twice to eight times as
broad as the bars; twelve to twenty on the radius. Radial spines very
numerous (two to three hundred), short, conical, scarcely as long as the
diameter of the largest meshes, and one-third as broad.

_Dimensions._--Diameter of the shell 0.3 to 0.4, pores 0.008 to 0.03, bars
0.004; length of the radial spines 0.03, basal breadth 0.01.

_Habitat._--Tropical Atlantic, Station 348, depth 2450 fathoms.

29. _Acanthosphæra simplex_, Haeckel.

_Rhaphidococcus simplex_, Haeckel, Monogr. d. Radiol., 1862, p. 366,
figs. 5, 6.Taf. xiii.

_Cladococcus simplex_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 800.

Shell thick walled, with irregular, roundish pores, three to six times as
broad as the bars; eight to nine on the radius. Forty to sixty radial
spines, about as long as the diameter of the shell, three-sided prismatic,
not straight, but more or less bent.

_Dimensions._--Diameter of the shell 0.12, pores 0.005 to 0.012, bars
0.0015 to 0.02; length of the spines 0.12, breadth 0.003.

_Habitat._--Mediterranean (Messina), surface, Haeckel.

30. _Acanthosphæra gibbosa_, n. sp.

Shell thin walled, rugged or tuberculate, covered by about twenty
hill-shaped tubercles or protuberances with flat valleys between them.
Network very delicate, with thread-like bars and {217}irregular, polygonal
pores; twenty to thirty on the radius. Radial spines very numerous,
bristle-shaped, twice to three times as long as the diameter of the pores.

_Dimensions._--Diameter of the shell 0.3, pores 0.01 to 0.02; length of the
spines 0.02 to 0.05.

_Habitat._--Central Pacific, Station 274, surface.

31. _Acanthosphæra reticulata_, n. sp. (Pl. 26, fig. 5).

_Rhaphidosphæra reticulata_, Haeckel, 1881, Prodromus.

Shell thick walled, with irregular, roundish pores, twice to four times as
broad as the bars; six to eight on the radius. Surface of the bars covered
with a peculiar delicate network of very fine crests. Twenty to forty
radial spines, angular, pyramidal, scarcely one-third as long as the radius
of the shell, as broad at the base as the bars.

_Dimensions._--Diameter of the shell 0.22, pores 0.02 to 0.04, bars 0.01;
length of the spines 0.04, basal breadth 0.01.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Genus 89. _Heliosphæra_,[123] Haeckel, 1862, Monogr. d. Radiol., p. 350
(_sensu emendato_).

_Definition._--#Astrosphærida# with one simple lattice-sphere, covered with
simple radial spines of two different kinds: larger main spines and smaller
by-spines.

The genus _Heliosphæra_ (in the mended definition here employed) differs
from the foregoing _Acanthosphæra_ in the possession of two different kinds
of radial spines: larger main spines scattered on the surface or disposed
regularly in limited numbers (twelve to twenty, sometimes forty to fifty or
more), and smaller by-spines in much larger numbers, arising from all the
nodal-points of the network (or sometimes also from its bars).

Subgenus 1. _Heliosphærella_, Haeckel.

_Definition._--Pores of the shell regular or subregular, all of nearly
equal size and similar form.

1. _Heliosphæra hexagonaria_, n. sp. (Pl. 26, fig. 2).

Shell very thin walled, about twenty times as broad as one pore. Meshes or
pores subregular, hexagonal, with thread-like bars; fifteen to seventeen on
the radius. Radial spines at the nodal-points of the network; about forty
main spines three-sided pyramidal, half as broad at the base as one pore,
and twice as long as the bristle-shaped by-spines, which are very numerous,
and as long as the diameter of one pore.

{218}_Dimensions._--Diameter of the shell 0.25 to 0.3, of the meshes or
pores 0.012 to 0.015, bars below 0.001; length of the main spines 0.03,
basal breadth 0.007.

_Habitat._--Central Pacific, Stations 272 to 274, depth 2350 to 2750
fathoms.

2. _Heliosphæra actinota_, Haeckel.

_Heliosphæra actinota_, Haeckel, 1862, Monogr. d. Radiol., p. 352, Taf.
ix. fig. 3.

Shell very thin walled, about ten times as broad as one mesh. Pores
regular, hexagonal, with thread-like bars; six to eight on the radius.
Radial spines at the nodal-points of the network, bristle-shaped, scarcely
broader than the bars; about twenty main spines as long as the diameter of
the shell, and numerous by-spines, only one-third to one-half as long as
the former.

_Dimensions._--Diameter of the shell 0.2 to 0.25, of the meshes 0.02 to
0.03, bars below 0.001; length of the main spines 0.2 to 0.3.

_Habitat._--Mediterranean (Messina), Atlantic (Canaries, Azores), surface.

3. _Heliosphæra echinoides_, Haeckel.

_Heliosphæra echinoides_, Haeckel, 1862, Monogr. d. Radiol., p. 352, Taf.
ix. fig. 4.

Shell thin walled, about six times as broad as one mesh. Pores regular,
hexagonal, eight times as broad as the bars. Radial spines arising, not
from the nodal-points of the network, but from the midst of the bars (very
rare disposition!); twenty main spines regularly disposed, as long as the
radius, four times as long as the numerous by-spines; all spines
bristle-shaped, of the same breadth as the bars.

_Dimensions._--Diameter of the shell 0.09, pores 0.015, bars 0.002; length
of the main spines 0.04.

_Habitat._--Mediterranean (Messina), surface, Haeckel.

4. _Heliosphæra elegans_, Haeckel.

_Heliosphæra elegans_, Haeckel, 1862, Monogr. d. Radiol., p. 353, Taf.
ix. fig. 5.

Shell very thin walled, about ten times as broad as one mesh. Pores
regular, hexagonal, with thread-like bars, which are crossed by tangential
bars, so that each side of a hexagon exhibits a regular rectangular cross
(exactly the same as in Pl. 19, fig. 5). All radial spines bristle-shaped,
as thin as the bars, and arising from the nodal-points; twenty main spines
as long as the radius, numerous by-spines scarcely one-sixth as long.

_Dimensions._--Diameter of the shell 0.26, pores 0.026, bars below 0.001;
length of the main spines 0.13.

_Habitat._--Mediterranean (Messina), surface, Haeckel.

5. _Heliosphæra pectinata_, n. sp. (Pl. 26, fig. 9).

_Acanthosphæra pectinata_, Haeckel, 1881, Atlas.

Shell thick walled, combed, about fourteen times as broad as one mesh.
Pores subregular, circular, with elevated hexagonal frames, three times as
broad as the bars. Radial spines very {219}numerous and stout; thirty to
forty main spines, three-sided pyramidal, nearly as long as the radius, as
broad as one mesh; by-spines small, conical, everywhere scattered at the
nodal-points of the network and on the high combs of the bars.

_Dimensions._--Diameter of the shell 0.16, pores 0.012, bars 0.004; length
of the main spines 0.07, basal breadth 0.016.

_Habitat._--West Tropical Pacific, Station 225, depth 4475 fathoms.

6. _Heliosphæra coronata_, n. sp. (Pl. 26, figs. 6, 6_a_).

_Acanthosphæra coronata_, Haeckel, 1881, Atlas.

Shell thick walled, about ten times as broad as one mesh. Pores regular,
circular, five times as broad as the bars; each pore surrounded by a
regular coronal of six short, conical by-spines (fig. 6_a_); twenty to
thirty main spines, also conical, half as long as the radius, as broad as
one pore.

_Dimensions._--Diameter of the shell 0.15, pores 0.015, bars 0.003; length
of the main spines 0.04, basal breadth 0.015.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

7. _Heliosphæra floribunda_, n. sp.

Shell thick walled about ten times as broad as one mesh. Pores regular,
six-lobed, twice as broad as the bars; each pore surrounded by six small
conical by-spines (as in Pl. 28, figs. 1, 1_b_); twenty main spines
regularly disposed cylindro-conical, as long as the diameter of the shell
or longer.

_Dimensions._--Diameter of the shell 0.15, pores 0.015, bars 0.008; length
of the main spines 0.16, breadth 0.008.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

8. _Heliosphæra cristata_, n. sp.

Shell thick walled, about twelve times as broad as one mesh. Pores
subregular, circular, twice as broad as the bars; each pore surrounded by
an elegant coronal of ten to twenty small, conical by-spines; twenty main
spines regularly disposed, conical, only one-third as long as the radius,
as broad at the base as one mesh.

_Dimensions._--Diameter of the shell 0.25, pores 0.02, bars 0.01; length of
the main spines 0.04, basal breadth 0.02.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

9. _Heliosphæra castanella_, n. sp.

Shell thick walled, about twenty times as broad as one mesh. Pores regular,
circular, of the same breadth as the bars. Whole surface densely covered
with innumerable bristle-shaped by-spines, half as long as the radius;
fifty to eighty main spines, conical, nearly as long as the diameter,
{220}twice as broad at the base as one pore. (Very similar to some species
of Castanella, Pl. 113, but without the osculum characteristic of this
Phæodarian; may be easily confounded with it.)

_Dimensions._--Diameter of the shell 0.3, pores 0.015, bars 0.015; length
of the main spines 0.25, basal breadth 0.03.

_Habitat._--North Pacific (Japan), Station 234, surface.

Subgenus 2. _Heliosphæromma_, Haeckel.

_Definition._--Pores of the shell irregular, of different size or form.

10. _Heliosphæra polygonaria_, n. sp.

Shell very thin walled, with thread-like bars and irregular, polygonal
pores (having four to eight angles, commonly five to seven); eight to ten
on the radius. Radial spines at all the nodal-points of the network,
bristle-shaped; forty to sixty main spines, as long as the radius, twice as
thick as the numerous by-spines, which are not larger than one mesh.

_Dimensions._--Diameter of the shell 0.15 to 0.2, pores 0.012 to 0.02, bars
0.001; length of the main spines 0.08 to 0.1.

_Habitat._--Central Pacific, Stations 266 to 272, surface.

11. _Heliosphæra heteracantha_, n. sp.

Shell thin walled, with irregular, polygonal pores, twice to four times as
broad as the bars; six to eight on the radius. Twenty radial main spines,
three-sided pyramidal, nearly as long as the diameter of the shell, as
broad as a larger mesh; innumerable bristle-shaped by-spines variously
distributed on the bars and at the nodal-points of the net; half as long as
the radius.

_Dimensions._--Diameter of the shell 0.12, pores 0.006 to 0.012, bars
0.003; length of the main spines 0.1, basal breadth 0.012.

_Habitat._--Equatorial Atlantic, Station 347, surface.

12. _Heliosphæra hyperionis_, n. sp.

Shell thick walled. Pores irregular, roundish, with polygonal frames, three
to six times as broad as the bars; twelve to fourteen on the radius. Thirty
to forty main spines, angular, pyramidal, scarcely as long as the radius
and twice as long as the numerous bristle-shaped by-spines.

_Dimensions._--Diameter of the shell 0.24, pores 0.006 to 0.012, bars
0.002; length of the main spines 0.1, basal breadth 0.012.

_Habitat._--North Pacific, Station 256, surface.

13. _Heliosphæra elector_, n. sp.

Shell thick walled, with irregular, roundish pores, twice to three times as
broad as the bars; eight to ten on the radius. Twenty main spines,
three-sided pyramidal, somewhat longer than the radius and four times as
long as the short bristle-shaped by-spines.

{221}_Dimensions._--Diameter of the shell 0.12, pores 0.005 to 0.01, bars
0.003; length of the main spines 0.08, basal breadth 0.01.

_Habitat._--South Atlantic, Station 325, surface.

14. _Heliosphæra solaris_, n. sp.

Shell thick walled, with irregular, roundish pores, about the same breadth
as the bars; six to eight on the radius. Fifty to eighty main spines,
conical, as long as the radius; by-spines very numerous, also conical, but
only as large as one pore.

_Dimensions._--Diameter of the shell 0.15, pores and bars 0.006 to 0.008;
length of the main spines 0.08, basal breadth 0.012.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Genus 90. _Conosphæra_,[124] Haeckel, 1881, Prodromus, p. 451.

_Definition._--#Astrosphærida# with one simple latticed sphere, covered
with radial spines having the form of hollow cones with porous walls.

The genus _Conosphæra_ differs from _Acanthosphæra_ in the peculiar
formation of the radial spines, which are not simple solid sticks, but
hollow cones with porous walls, as immediate elevations of the hollow
sphere.

1. _Conosphæra platyconus_, n. sp.

Pores of the shell regular, circular, twice as broad as the bars; ten to
twelve on the radius. Conical spines about sixty, regular, broader than
they are high, with six to nine pores in the wall.

_Dimensions._--Diameter of the shell 0.16, pores 0.008, bars 0.004; length
of the spines 0.012, basal breadth 0.024.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Conosphæra orthoconus_, n. sp. (Pl. 12, fig. 2).

Pores of the shell irregular, roundish, twice to three times as broad as
the bars; fourteen to sixteen on the radius. Conical spines about forty,
regular, higher than they are broad, as long as the radius, with sixteen to
twenty pores in the wall.

_Dimensions._--Diameter of the shell 0.2, pores 0.008 to 0.012, bars 0.004;
length of the spines 0.05, basal breadth 0.03.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

{222}3. _Conosphæra plagioconus_, n. sp. (Pl. 12, fig. 4).

Pores of the shell irregular, polygonal, twice to five times as broad as
the bars; twenty to twenty-four on the radius. Conical spines about eighty,
irregularly formed and scattered, with oblique (not radial) axes; about as
high as broad, with eight to twelve pores in the wall.

_Dimensions._--Diameter of the shell 0.25, pores 0.005 to 0.015, bars
0.003; length of the spines 0.02, basal breadth 0.02.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Genus 91. _Coscinomma_,[125] n. gen.

_Definition._--#Astrosphærida# with one simple lattice-sphere, covered with
simple radial spines; the pores between them prolonged into hollow,
conical, or cylindrical tubuli.

The genus _Coscinomma_ exhibits among the Astrosphærida the same peculiar
formation that distinguishes _Ethmosphæra_ and _Sethosphæra_ among the
Liosphærida; each pore of the simple shell is prolonged into a short
conical or cylindrical tubulus, as a rule either on the outside or on the
inside of the shell, but sometimes on both sides.

Subgenus 1. _Coscinommarium_, Haeckel.

_Definition._--Pores prolonged into short tubes both on the inside as well
as the outside of the shell.

1. _Coscinomma amphisiphon_, Haeckel (Pl. 26, figs. 1, 1_a_, 1_b_).

Pores regular, circular, hexagonally framed, twice as broad as the bars,
prolonged on the inside as well as on the outside of the shell into a short
truncated conical tube; fifteen to eighteen pores on the radius. In each
hexagon-corner arises a bristle-shaped radial spine, half as long as the
radius.

_Dimensions._--Diameter of the shell 0.22, pores 0.01, bars 0.005; length
of the spines 0.1.

_Habitat._--Central Pacific, Station 271, surface.

Subgenus 2. _Coscinommidium_, Haeckel.

_Definition._--Pores prolonged into external tubes on the outside of the
shell.

2. _Coscinomma ectosiphon_, n. sp.

Pores regular, circular, hexagonally framed, twice as broad as the bars,
prolonged on the outside of the shell into a short truncated conical tube;
ten to twelve pores on the radius. In each {223}hexagon-corner arises a
short bristle-shaped spine, twice as long as the tube, one-third as long as
the radius. (Very similar to _Ethmosphæra conulosa_, Pl. 12, fig. 5, but
differs in the possession of radial spines.)

_Dimensions._--Diameter of the shell 0.2, pores 0.012, bars 0.006; length
of the spines 0.04, of the tubes 0.02.

_Habitat._--North Pacific, Station 253, surface.

3. _Coscinomma macrosiphon_, n. sp.

Pores regular, circular, without hexagonal frames, of the same breadth as
the bars, prolonged on the outside of the shell into a long cylindrical
tube, half as long as the radius (eight to nine pores on the radius);
between them bristle-shaped, at the base conical, radial spines of double
length.

_Dimensions._--Diameter of the shell 0.16, pores and bars 0.008; length of
the spines 0.08, of the tubes 0.04.

_Habitat._--Central Pacific, Station 274, surface.

Subgenus 3. _Coscinommonium_, Haeckel.

_Definition._--Pores prolonged into internal tubes on the inside of the
shell.

4. _Coscinomma endosiphon_, n. sp.

Pores regular, circular, hexagonally framed, twice as broad as the bars
(fourteen to sixteen on the radius), prolonged on the inside of the shell
into a short truncated cylindrical tube. In each hexagon-corner arises a
thin, bristle-shaped, radial spine with pyramidal base, half as long as the
radius, twice as long as the tube.

_Dimensions._--Diameter of the shell 0.25, pores 0.012, bars 0.006; length
of the spines 0.066, of the tubes 0.03.

_Habitat._--South Pacific, Station 288, surface.

Genus 92. _Cladococcus_,[126] J. Müller, 1856, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 485.

_Definition._--#Astrosphærida# with one simple lattice-sphere, covered with
branched radial spines (the stem of the spine never forked).

The genus _Cladococcus_, together with the following _Elaphococcus_, is
distinguished from the other _Coscinommida_ by the ramification of the
radial spines covering the surface of the simple hollow lattice-sphere. In
_Cladococcus_ each spine sends out three or more lateral branches, which
are either simple or again ramified; but the stem of the spine itself is
not forked, as in _Elaphococcus_.

{224}Subgenus 1. _Cladococcalis_, Haeckel.

_Definition._--Branches of the spines simple, not ramified. Pores regular,
all of nearly equal size and similar form.

1. _Cladococcus arborescens_, J. Müller.

_Cladococcus arborescens_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 31, Taf. i. fig. 2.

Pores of the spherical shell regular, hexagonal, three times as broad as
the bars; three to four on the radius. Ten to twenty spines, three-sided
prismatic, two to three times as long as the shell diameter; towards the
distal end each spine with three branches (one lateral simple straight
branch on each edge of the spine).

_Dimensions._--Diameter of the shell 0.1, pores 0.012, bars 0.004; length
of the spines 0.2 to 0.3, breadth 0.02.

_Habitat._--Mediterranean (Nice); North Atlantic, Canary Islands, surface.

2. _Cladococcus spinifer_, Haeckel.

_Cladococcus spinifer_, Haeckel, 1862, Monogr. d. Radiol., p. 368, Taf.
xiii. fig. 9.

Pores regular, circular, hexagonally framed, three times as broad as the
bars; five to six on the radius. Radial spines, arising from all the
nodal-points of the network, three-sided, longer than the shell diameter,
with six to twelve simple verticillate branches (two to four branches on
each edge).

_Dimensions._--Diameter of the shell 0.08, pores 0.01, bars 0.003; length
of the spines 0.1, breadth 0.03.

_Habitat._--Mediterranean (Messina), surface.

3. _Cladococcus penicillus_, n. sp.

Pores subregular, hexagonal, twice as broad as the bars; eight to ten on
the radius. Sixty to eighty radial spines, three-sided prismatic,
pencil-shaped, longer than the shell diameter; each at the distal end with
a brush or pencil composed of nine to twenty-one short, simple, curved
branches (three to seven on each edge).

_Dimensions._--Diameter of the shell 0.1, pores 0.006, bars 0.003; length
of the spines 0.12 to 0.16, breadth 0.006.

_Habitat._--Central Pacific, Station 271, surface.

Subgenus 2. _Cladococcinus_, Haeckel.

_Definition._--Branches of the spines simple, not ramified. Pores
irregular, of different size or form.

{225}4. _Cladococcus antarcticus_, n. sp.

Pores irregular, polygonal, twice to four times as broad as the bars; five
to six on the radius. Forty to fifty radial spines, angular, curved, of
variable size and form, with three to nine irregular, simple, blunt curved
branches.

_Dimensions._--Diameter of the shell 0.14, pores 0.01 to 0.02, bars 0.05;
length of the spines 0.1 to 0.18, breadth 0.007.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

5. _Cladococcus japonicus_, n. sp.

Pores irregular, roundish, of the same breadth as the bars; six to eight on
the radius. Twenty to thirty radial spines, angular, longer than the shell
diameter, with thirty to forty simple branches, decreasing in size from the
middle part of the spine to the distal end (ten to thirteen branches on
each edge).

_Dimensions._--Diameter of the shell 0.12, pores and bars 0.007; length of
the spines 0.15 to 0.2, breadth 0.004.

_Habitat._--North Pacific (Japan), Station 240, surface.

6. _Cladococcus quadricuspis_, n. sp.

Pores irregular, roundish, twice to eight times as broad as the bars; eight
to ten on the radius. About twenty radial spines, three-sided prismatic, as
long as the shell diameter; in the proximal half simple, with smooth edges;
in the distal half with three diverging simple branches, half the length of
the spine, with dentated edges.

_Dimensions._--Diameter of the shell 0.15, pores 0.004 to 0.016, bars
0.002; length of the spines 0.17, breadth 0.01.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Subgenus 3. _Cladococcodes_, Haeckel.

_Definition._--Branches of the spines again ramified. Pores regular, of
nearly equal size and similar form.

7. _Cladococcus scoparius_, n. sp. (Pl. 27, fig. 2).

Pores regular, circular, three times as broad as the bars; five to six on
the radius. Twenty radial spines, broom-shaped, three times as long as the
shell diameter; in the proximal half simple, in the distal half branched,
with three to nine branches, which are again ramified; spines and their
branches with three smooth edges, not dentated.

_Dimensions._--Diameter of the shell 0.055, pores 0.0075, bars 0.0025;
length of the spines 0.15, basal thickness 0.007.

_Habitat._--Central Pacific, Station 271, surface.

{226}8. _Cladococcus viminalis_, Haeckel.

_Cladococcus viminalis_, Haeckel, 1862, Monogr. d. Radiol., p. 369, Taf.
xiv. figs. 2, 3.

Pores regular, circular, hexagonally framed, four times as broad as the
bars; five to six on the radius. Radial spines, arising from all the
nodal-points of the network, twice as long as the shell diameter; in the
basal half simple, in the distal half with three to six long, thin, curved
branches, which are partly forked; three edges of the spines dentated.

_Dimensions._--Diameter of the shell 0.08, pores 0.01, bars 0.003; length
of the radial spines 0.16, basal breadth 0.003.

_Habitat._--Mediterranean (Messina).

9. _Cladococcus bifurcus_, Haeckel.

_Cladococcus bifurcus_, Haeckel, 1862, Monogr. d. Radiol., p. 368, Taf.
xiii. figs. 7, 8.

Pores regular, circular, hexagonally framed, four times as broad as the
bars; five to seven on the radius. Radial spines arising from all the
nodal-points, three-sided prismatic, with dentated edges, longer than the
shell diameter. On each spine six to nine branches, which are for the most
part forked, the distal branches only being simple. (May be regarded as a
further developmental stage of _Cladococcus spinifer_ and _Cladococcus
viminalis_.)

_Dimensions._--Diameter of the shell 0.08, pores 0.01, bars 0.0025; length
of the spines 0.1, basal breadth 0.003.

_Habitat._--Mediterranean (Messina), Canary Islands, Azores, surface.

10. _Cladococcus pinetum_, n. sp. (Pl. 27, fig. 1).

Pores regular, circular, polygonally framed, about the same breadth as the
bars: two to three on the radius. About twenty radial spines, very large,
three to four times as long as the shell diameter, branched like a pine
tree, with straight, stout, three-sided prismatic stem; three prominent
edges dentated. On each edge five to seven lateral branches, the distal
ends of which are simple, the proximal again ramified, with numerous
ramules. The figured specimen is a young one, with branches but little
developed.

_Dimensions._--Diameter of the shell 0.06, pores and bars 0.008; length of
the spines 0.2 to 0.25 breadth 0.01.

_Habitat._--Central Pacific, Station 266 to 274, surface.

Subgenus 4. _Cladococcurus_, Haeckel.

_Definition._--Branches of the spines again ramified. Pores irregular, of
different size and form.

11. _Cladococcus abietinus_, n. sp. (Pl. 27, fig. 3).

Pores irregular, roundish, twice to four times as broad as the bars; six to
ten on the radius. About twenty radial spines, two to three times as long
as the shell diameter, branched like a pine {227}tree, with straight,
stout, three-sided pyramidal stem. From the dentated edges arise numerous
verticillate branches, the proximal ends of which are thickly ramified.
(Differs from the preceding and similar species mainly in the large size
and irregular lattice-work of the shell.)

_Dimensions._--Diameter of the shell 0.12 to 0.14, pores 0.006 to 0.012,
bars 0.003; length of the spines 0.25 to 0.4, breadth 0.012.

_Habitat._--North Pacific, Station 256, surface.

12. _Cladococcus tricladus_, n. sp.

Pores irregular, polygonal, twice to eight times as broad as the bars;
eight to ten on the radius. About twenty radial spines, three-sided
prismatic, twice as long as the shell diameter; in the proximal half
simple, with smooth edges; in the distal half with three diverging curved
branches, which bifurcate two to three times or ramify irregularly.
(Closely related to the simpler _Cladococcus quadricuspis_.)

_Dimensions._--Diameter of the shell 0.16, pores 0.04 to 0.016, bars 0.002;
length of the spines 0.3, breadth 0.012.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

13. _Cladococcus stalactites_, n. sp. (Pl. 27, fig. 4).

Pores irregular, roundish, once to five times as broad as the bars; eight
to ten on the radius. About twenty radial spines, very stout, longer than
the shell diameter, with three wing-like, prominent smooth edges. At the
middle, or in the distal half, each spine bears a verticil of three strong,
irregularly formed ramified branches.

_Dimensions._--Diameter of the shell 0.14, pores 0.003 to 0.015, bars
0.003; length of the spines 0.16 to 0.2, breadth 0.02.

_Habitat._--West Tropical Pacific, Station 220, depth 1100 fathoms.

14. _Cladococcus dendrites_, n. sp. (Pl. 27, fig. 5).

Pores irregular, roundish, with denticulate margin, twice to four times as
broad as the bars; eight to twelve on the radius. Fifty to eighty radial
spines, three-sided prismatic, with elegantly denticulated edges; in the
proximal two-thirds simple, in the distal third with a bunch of ten to
twenty short simple spines (three to seven on each edge).

_Dimensions._--Diameter of the shell 0.16 to 0.2, pores 0.006 to 0.012,
bars 0.003; length of the spines 0.2 to 0.3, breadth 0.01.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Genus 93. _Elaphococcus_,[127] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Astrosphærida# with one simple lattice-sphere, covered with
forked or dichotomously branched radial spines (the stem of the spine
always forked).

{228}The genus _Elaphococcus_ differs from the preceding _Cladococcus_ in
the mode of ramification of the radial spines. These are forked; and the
forked branches are either simple, again forked, or dichotomously ramified.

Subgenus 1. _Elaphococcinus_, Haeckel.

_Definition._--Pores of the shell regular, of nearly equal size and similar
form.

1. _Elaphococcus furcatus_, n. sp.

Pores of the spherical shell regular, hexagonal, four times as broad as the
bars; four to five on the radius. Radial spines, arising from all the
nodal-points of the network, cylindrical, as long as the radius, simply
forked at the distal end; both branches half as long as the undivided part.

_Dimensions._--Diameter of the shell 0.08, pores 0.01, bars 0.0025; length
of the spines 0.04, breadth 0.004.

_Habitat._--Tropical Atlantic, Station 348, surface.

2. _Elaphococcus cervicornis_, Haeckel.

_Cladococcus cervicornis_, Haeckel, 1862, Monogr. d. Radiol., p. 370,
Taf. xiv. figs. 4-6.

Pores regular, hexagonal, ten to twenty times as broad as the bars; five to
seven on the radius. Fifty to ninety radial spines (or more), arising not
only from the nodal-points of the network but also from the bars between
them. Each spine is cylindrical, longer than the shell diameter, and
repeatedly forked (three to six times), having, therefore, numerous (thirty
to sixty or more) curved branches. The distal ends of all branches fall in
one spherical face.

_Dimensions._--Diameter of the shell 0.07 to 0.09, pores 0.01 to 0.015,
bars 0.001 to 0.015; length of the spines 0.1 to 0.15, breadth 0.005.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

3. _Elaphococcus elaphoceras_, n. sp.

Pores regular, circular, hexagonally framed, three to four times as broad
as the bars; six to eight on the radius. At each nodal-point of the hexagon
arises a short bristle-shaped, simple by-spine. In addition, there arise
from the bars thirty to sixty large main spines, longer than the shell
diameter, repeatedly forked in the same way as in the preceding species.

_Dimensions._--Diameter of the shell 0.1, pores 0.008, bars 0.002; length
of the spines 0.12, breadth 0.002.

_Habitat._--Central Pacific, Station 272. depth 2600 fathoms.

4. _Elaphococcus umbellifer_, n. sp.

Pores regular, circular, three times as broad as the bars; ten to twelve on
the radius. Twenty to forty straight cylindrical spines, as long as the
shell diameter, having at the distal end a regular {229}umbel composed of
nine to twelve curved branches of equal length, which are again ramified
and resemble the inflorescence of an umbelliferous plant, the distal ends
of all ramules falling in a spherical face.

_Dimensions._--Diameter of the shell 0.12, pores 0.006, bars 0.002; length
of the spines 0.14, breadth 0.002.

_Habitat._--South Atlantic, Station 325, surface.

Subgenus 2. _Elaphococculus_, Haeckel.

_Definition._--Pores of the shell irregular, of different size or form.

5. _Elaphococcus dichotomus_, n. sp.

Pores irregular, polygonal, twice to four times as broad as the bars; six
to eight on the radius. Thirty to sixty radial spines cylindrical, curved,
as long as the shell diameter, simply forked at the distal end; both
branches one-third as long as the undivided part.

_Dimensions._--Diameter of the shell 0.15, pores 0.007 to 0.015, bars
0.004; length of the spines 0.2, breadth 0.004.

_Habitat._--Arctic Ocean, Greenland, surface (Koch).

6. _Elaphococcus umbellatus_, n. sp.

Pores irregular, roundish, or polygonal, twice to five times as broad as
the bars; six to eight on the radius. Forty to sixty radial spines
cylindrical, curved, as long as the shell radius, having at the distal end
an irregular umbel, composed of six to twelve short branches, which are
irregularly ramified or forked. (Differs from the regular _Elaphococcus
umbellifer_ mainly in the irregularity.)

_Dimensions._--Diameter of the shell 0.14, pores 0.007 to 0.015, bars
0.003; length of the spines 0.12, breadth 0.01.

_Habitat._--South-east Pacific (Juan Fernandez), Station 299, surface.

7. _Elaphococcus drymodes_, n. sp.

Pores irregular, roundish, little broader than the bars; ten to twelve on
the radius. Eighty to one hundred and twenty (or more) radial spines,
cylindrical, three to four times as long as the shell diameter, irregularly
forked or repeatedly dichotomous (each spine with forty to sixty forked
branches); the distal ends of all branches fall in a spherical plane.
(Differs from the regular _Elaphococcus cervicornis_ mainly in the
irregularity.)

_Dimensions._--Diameter of the shell 0.1, pores and bars 0.003 to 0.008;
length of the spines 0.3 to 0.4, breadth 0.01.

_Habitat._--Arctic Ocean, Iceland (Steenstrup).

{230}Subfamily HALIOMMIDA,[128] Haeckel.

_Diplosphærida_, Haeckel, 1881, Prodromus, pp. 449, 451.

_Definition._--#Astrosphærida# with two concentric spherical
lattice-shells, united by radial beams.

Genus 94. _Haliomma_,[129] Ehrenberg, 1838, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 128.

_Definition._--#Astrosphærida# with one medullary (intracapsular) and one
cortical (extracapsular) shell, which are connected by radial beams,
piercing the central capsule. Shell surface covered with simple radial
spines of the same kind.

The genus _Haliomma_, one of the oldest known Radiolarian genera, contained
in the catalogue of its discoverer, Ehrenberg, a large number of very
different #Sphærellaria#, belonging to at least sixteen different genera.
We limit here the conception of the genus to those Haliommida which bear
simple radial spines of one kind on the surface of the cortical shell (the
latter being separated from the medullary shell by the central capsule).

Subgenus 1. _Haliommantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; spines on the whole surface (commonly one spine at each
nodal-point).

1. _Haliomma hexagonium_, n. sp.

Cortical shell four times as broad as the medullary shell, both having very
thin thread-like bars, and regular, hexagonal pores (eighteen to twenty on
the radius of the outer, five to six on the radius of the inner shell). The
two shells connected by twenty thin thread-like radial beams. At each
nodal-point of the outer shell arises a bristle-shaped radial spine, half
as long as the radius. (Similar to _Heliosoma radians_, Pl. 28, fig. 3, but
with all the spines equal.)

_Dimensions._--Diameter of the outer shell 0.2, inner 0.05; pores of the
outer 0.015, of the inner 0.01; length of the spines 0.05.

_Habitat._--Equatorial Atlantic, Station 347, surface.

2. _Haliomma arachnium_, n. sp.

Cortical shell three times as broad as the medullary shell; pores of the
former regular, hexagonal, with very thin thread-like bars (fourteen to
sixteen on the radius); pores of the latter regular, {231}circular, three
times as broad as the bars (four to five on the radius). The two shells
connected by about forty radial beams. At each nodal-point of the outer
shell arises one bristle-shaped radial spine, twice as long as the diameter
of one hexagonal mesh.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; pores of the
outer 0.012, of the inner 0.006; length of the spines 0.025.

_Habitat._--Central Pacific, Station 266, surface.

3. _Haliomma favosum_, n. sp.

Cortical shell thick walled, three times as broad as the medullary shell,
with regular, circular, hexagonally framed pores, twice as broad as the
bars; eight to ten on the radius. Medullary shell with simple, circular
pores, of the same breadth as the bars. At each nodal-point of the outer
shell arises a short triangular spine three times as long as one pore.

_Dimensions._--Diameter of the outer shell 0.12, inner 0.04; pores of the
former 0.01, of the latter 0.005; bars 0.005; length of the spines 0.015.

_Habitat._--Central Pacific, Station 271 to 274, surface.

4. _Haliomma regulare_, n. sp.

Cortical shell thin walled, nearly twice as broad as the medullary shell,
and connected with it by forty to sixty thin radial beams. Both shells of
the same perfectly regular structure, with an identical number of regular,
circular pores, which are hexagonally framed, four times as broad as the
bars, nine to eleven on the radius. The outer pores are twice as broad as
the inner, exactly corresponding pores. Between every three pores of the
outer surface (in each corner of the cortical hexagon) arises one short
three-sided pyramidal spine, twice as long as the diameter of one pore.

_Dimensions._--Diameter of the outer shell 0.2, of the inner 0.11; pores of
the former 0.018, of the latter 0.009; length of the spines 0.04.

_Habitat._--South Atlantic (Tristan da Cunha), Station 332, depth 2200
fathoms.

5. _Haliomma melitomma_, n. sp. (Pl. 20, fig. 4).

_Melitomma formosum_, Haeckel, 1879, Atlas, _loc. cit._

Cortical shell thick walled, two and a half times as broad as the medullary
shell, with regular pores of very elegant structure, twice as broad as the
bars; eight to ten on the radius. The inner opening of each pore is simple,
circular, the outer regular, six-lobed; corresponding to the six lobes are
six short conical spines, which arise from the six corners of the regular
hexagonal frames separating the pores. Pores of the medullary shell simple,
circular, of the same breadth as the bars. The two shells connected only by
six radial beams (in three dimensive axes).

_Dimensions._--Diameter of the outer shell 0.17, of the inner 0.07; pores
of the former 0.012, of the latter 0.004; length of the spines 0.005 to
0.01.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

{232}6. _Haliomma lirianthus_, n. sp. (Pl. 28, figs. 1, 1_a_, 1_b_).

Cortical shell thick walled, three times as broad as the medullary shell,
with regular, rosette-shaped pores, twice as broad as the bars; eight to
ten on the radius. The regular structure of the elegant pores is the same
as in the preceding species, but without prominent crested frames (fig.
1_b_). Also the medullary shell (fig. 1_a_) is different, much thinner and
smaller, with simple, circular pores, which are three times as broad as the
bars. The two shells are connected by numerous (twenty?) radial beams.

_Dimensions._--Diameter of the outer shell 0.15, of the inner 0.05; pores
of the former 0.01, of the latter 0.005; length of the spines 0.005 to
0.02.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

7. _Haliomma castanea_, Haeckel.

_Haliomma castanea_, Haeckel, 1862, Monogr. d. Radiol., p. 428, Taf.
xxiv. fig. 4.

Cortical shell thick walled, three times as broad as the medullary shell;
pores of both regular, circular, twice as broad as the bars (seven to eight
on the radius of the outer, four to five on the radius of the inner shell),
the two connected by six to twelve (?) strong radial beams. Radial spines
bristle-shaped, with conical bases, twice as long as the diameter of the
cortical pores (one spine at the nodal-point between every three pores).

_Dimensions._--Diameter of the outer shell 0.1 to 0.15, inner 0.03 to 0.05;
pores of the former 0.005, of the latter 0.003; length of the spines 0.005
to 0.01.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

8. _Haliomma horridum_, Stöhr.

_Haliomma horridum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 87, Taf. i.
fig. 10.

Cortical shell thick walled, two and a half times as broad as the medullary
shell. Pores regular, circular, of the same breadth as the bars; eight to
ten on the radius. Radial spines conical, stout, nearly half as long as the
radius. (Differs from the closely allied _Haliomma castanea_ in the smaller
pores and larger spines.)

_Dimensions._--Diameter of the outer shell 0.14, of the inner 0.06; pores
of the former 0.004, of the latter 0.02.

_Habitat._--Fossil in Tertiary rocks of Barbados and Sicily; living in the
Atlantic, Station 9, depth 3150 fathoms, and Station 353, depth 2965
fathoms.

9. _Haliomma datura_, n. sp.

Cortical shell thin walled, only one and a half times as broad as the
medullary shell, both having regular, circular pores, four to six times as
broad as the bars (five to six on the radius of the outer, three to four on
the radius of the inner shell), the two connected by numerous (forty to
sixty?) thin, short, radial beams. Radial spines conical, twice as long as
the diameter of one cortical pore, arising from all the nodal-points
between them.

{233}_Dimensions._--Diameter of the outer shell 0.08 to 0.12, inner 0.06 to
0.08; pores of the former 0.015 to 0.02, of the latter 0.007 to 0.012;
length of the spines 0.02.

_Habitat._--Central Pacific, Stations 268 to 274, surface.

Subgenus 2. _Haliommetta_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; the spines not covering the entire surface, but scattered
at intervals (their number smaller than that of the nodal-points of the
network).

10. _Haliomma circumtextum_, n. sp. (Pl. 28, figs. 7, 7_a_).

Cortical shell very delicate, with thin thread-like bars, and regular,
hexagonal pores, little larger than the thick-walled medullary shell (=
7 : 6). Pores of the latter regular, circular, double-edged, with hexagonal
frames, of the same breadth as the bars; from all the hexagon-corners arise
thin bristle-shaped, radial spines, twice as long as the diameter of the
pores, becoming connected at equal distances from the centre by tangential
threads, regularly disposed, forming the cortical shell. Twelve strong,
three-sided pyramidal, radial spines, as broad at the base as one mesh, and
about half as long as the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.14, of the inner 0.12; pores
of the former 0.012, of the latter 0.005; length of the radial spines 0.04,
basal breadth 0.012.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

11. _Haliomma duodecinum_, n. sp.

Cortical shell thick walled, three times as broad as the medullary shell.
Both shells with regular, circular pores, twice as broad as the bars; eight
to ten on the radius of the outer, five to six on the radius of the inner
shell. Twelve conical, regularly disposed radial spines, as long as the
radius, and as broad at the base as one mesh.

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.04; pores
of the former 0.005, of the latter 0.002; length of the spines 0.05, basal
breadth 0.005.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

12. _Haliomma megaporum_, Ehrenberg.

_Haliomma megaporum_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 313.

Cortical shell thin walled, three times as broad as the medullary shell;
the pores regular, circular, three to four on the radius, eight times as
broad as the bars, quite as broad as the medullary shell, the pores of
which are much smaller, twice as broad as the bars. Eight radial spines
regularly disposed, conical, as long as the radius. (As the diagnosis of
Ehrenberg is very {234}incomplete, and no figure is given with it, it
remains doubtful whether his Mediterranean species be identical with my
Atlantic variety.)

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.04; pores
of the former 0.04, of the latter 0.008; length of the spines 0.06.

_Habitat._--Mediterranean (Grecian shore); North Atlantic, Station 354,
surface.

13. _Haliomma oculatum_, Ehrenberg.

_Haliomma oculatum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxviii. figs. 2, 3.

Cortical shell thick walled, four times as broad as the medullary shell;
its pores regular, circular, four times as broad as the bars, six to eight
on the radius. Pores of the medullary shell regular, hexagonal, with very
thin bars, three to four on the radius. Nine radial spines, regularly
disposed, conical, as long as the radius, as broad at the base as one
cortical pore. (In the specimen figured by Ehrenberg, only two spines were
preserved, seven being accidentally broken off.)

_Dimensions._--Diameter of the outer shell 0.16, of the inner 0.04; pores
of the former 0.01, of the latter 0.005.

_Habitat._--Fossil in Barbados.

14. _Haliomma enneaxiphos_, n. sp.

Cortical shell thick walled, four times as broad as the dark medullary
shell, with regular, circular, hexagonally framed pores, twice as broad as
the bars; eight to ten on the radius. Nine radial spines regularly
disposed, three sided pyramidal, as long as the radius, as broad at the
base as one mesh.

_Dimensions._--Diameter of the outer shell 0.11, of the inner 0.03;
cortical pores 0.008, bars 0.004; length of the radial spines 0.05, basal
breadth 0.012.

_Habitat._--Central Pacific, Station 272, surface.

15. _Haliomma tenuispinum_, J. Müller.

_Haliomma tenuispinum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 39, Taf. iv. fig. 9.

_Haliomma tenuispinum_, Haeckel, 1862, Monogr. d. Radiol., p. 431.

Cortical shell thin walled, three times as broad as the medullary shell.
Both shells with very fine, thread-like bars, and regular, hexagonal pores;
five to seven on the radius. Twenty radial spines, very thin,
bristle-shaped, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.04; pores
of the former 0.016, of the latter 0.006.

_Habitat._--Mediterranean (Nice); North Atlantic, Station 353, surface.

{235}16. _Haliomma longispinum_, J. Müller.

_Haliomma longispinum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 39, Taf iv. fig. 8.

_Haliomma longispinum_, Haeckel, 1862, Monogr. d. Radiol., p. 431.

Cortical shell thick walled, three times as broad as the dark medullary
shell. Pores regular, circular, hexagonally framed, twice as broad as the
bars; four to six on the radius. Twenty radial spines, very long, regularly
disposed, four to five times as long as the diameter of the shell,
three-sided prismatic, with elegantly dentated edges, as broad as one
cortical mesh.

_Dimensions._--Diameter of the outer shell 0.08, of the inner 0.03; pores
of the former 0.01, bars 0.005; length of the spines 0.3 to 0.4.

_Habitat._--Mediterranean (Nice); Central Pacific, Station 272, surface.

17. _Haliomma capense_, n. sp.

Cortical shell thick walled, twice as broad as the medullary shell, both
with regular, circular pores, twice as broad as the bars; eight to ten on
the radius of the outer, four to six on the radius of the inner shell.
Forty to sixty radial spines, conical, as long as the radius, as broad as
one cortical pore.

_Dimensions._--Diameter of the outer shell 0.11, of the inner 0.05; pores
of the former 0.01, bars 0.005; length of the spines 0.006, basal breadth
0.012.

_Habitat._--Cape of Good Hope, Station 142, surface.

18. _Haliomma denticulatum_, n. sp.

Cortical shell thick walled, three times as broad as the dark medullary
shell, with regular, circular, double-edged pores, of the same breadth as
the bars; eight to ten on the radius. Thirty to fifty radial spines,
three-sided pyramidal, half as long as the radius, with three denticulated
edges. (Very similar to _Actinomma denticulatum_, Pl. 29, fig. 3, but with
simple medullary shell.)

_Dimensions._--Diameter of the outer shell 0.12, inner 0.04; pores and bars
of the former 0.008; length of the spines 0.04, basal breadth 0.01.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

19. _Haliomma grande_, n. sp.

Cortical shell thick walled, five times as broad as the dark medullary
shell, with small, regular, circular pores, half as broad as the bars;
twenty to twenty-four on the radius. One hundred to one hundred and twenty
conical radial spines, five times as long as broad, only half as long as
the radius.

_Dimensions._--Diameter of the outer shell 0.4, inner 0.08; pores of the
outer 0.006, bars 0.012; length of the spines 0.1, basal breadth 0.02.

_Habitat._--Equatorial Atlantic, Station 348, depth 2450 fathoms.

{236}Subgenus 3. _Haliommilla_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size and
form; spines covering the entire surface (commonly one spine at each
nodal-point).

20. _Haliomma capillaceum_, Haeckel.

_Haliomma capillaceum_, Haeckel, 1862, Monogr. d. Radiol., p. 426, Taf.
xxiii. fig. 2.

Cortical shell very thin walled, seven to eight times as broad as the
medullary shell, both with irregular, polygonal pores, and very thin
thread-like bars; outer pores twice to three times as broad as the inner.
Radial spines very numerous, straight, bristle-shaped, about as long as the
diameter of the medullary shell.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.025 to 0.03; outer
pores 0.02 to 0.04, inner 0.008 to 0.016, bars 0.001; length of the spines
0.02 to 0.03.

_Habitat._--Mediterranean (Messina), Atlantic (Canary Islands), surface.

21. _Haliomma erinaceum_, Haeckel.

_Haliomma erinaceum_, Haeckel, 1862, Monogr. d. Radiol., p. 427, Taf.
xxiii. figs. 3, 4.

Cortical shell thin walled, seven to eight times as broad as the medullary
shell, both with irregular, polygonal pores, two to ten times as broad as
the bars, outer pores much larger than the inner. Radial spines very
numerous, bristle-shaped, as long as the diameter of the medullary shell,
the majority or all being either curved or obliquely depressed, the greater
part neither straight nor radial.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.025 to 0.03; outer
pores 0.005 to 0.03, inner 0.008 to 0.016, bars 0.002 to 0.004; length of
the spines 0.03.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

22. _Haliomma tenellum_, Haeckel.

_Haliomma tenellum_, Haeckel, 1862, Monogr. d. Radiol., p. 428.

_Haliomma spinuloso affine_, J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 40, Taf. iv. fig. 7.

Cortical shell thin walled, three times as broad as the medullary shell,
with irregular, roundish pores, and very thin bars. Pores of the inner
shell regular, circular. Radial spines very numerous, bristle-shaped,
straight, as long as the radius of the outer shell.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; outer pores
0.008 to 0.016, inner 0.005; length of the spines 0.07.

_Habitat._--Mediterranean (Nice); Atlantic, Stations 349 to 354, surface.

{237}23. _Haliomma spinulosum_, J. Müller.

_Haliomma spinulosum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 39, Taf. iv. fig. 6.

Cortical shell thin walled, twice as broad as the medullary shell, with
irregular, polygonal pores, and very thin bars. Pores of the inner shell
subregular, hexagonal, ten to twelve times as broad as the bars. Radial
spines very numerous, bristle-shaped, one-sixth to one-fourth as long as
the radius of the outer shell.

_Dimensions._--Diameter of the outer shell 0.16, inner 0.08; outer pores
0.01 to 0.03, inner 0.02; length of the spines 0.02 to 0.03.

_Habitat._--Mediterranean (Nice); Central Pacific, Station 266, surface.

24. _Haliomma rhodococcus_, n. sp. (Pl. 19, fig. 6).

_Sethosphæra rhodococcus_, Haeckel, 1879, Atlas, pl. xix. fig. 6.

Cortical shell very thin walled, little larger than the thick walled
medullary shell (= 10 : 9). Pores of the outer shell very irregular,
roundish, twice to six times as broad as the bars, about twenty on the
radius. Pores of the inner shell twice as broad as the bars, six to eight
on the radius, very regular, circular on the inner, six-lobed on the outer
opening, separated by prominent hexagonal crests; in the hexagon-corners
arise short conical radial spines, which at equal distances from the centre
are united by the outer shell, but are prominent over its surface. (A very
peculiar form; the two shells may better be regarded as inner and outer
cortical shell.)

_Dimensions._--Diameter of the outer shell 0.2, inner 0.18; outer pores
0.04 to 0.012, inner 0.01; length of the spines 0.01.

_Habitat._--Tropical Atlantic, Station 338, depth 1990 fathoms; also fossil
in Barbados.

25. _Haliomma boreale_, n. sp.

Cortical shell thick walled, four times as broad as the medullary shell,
with irregular, roundish pores, twice to four times as broad as the bars;
eight to ten on the radius. Pores of the inner shell regular, circular,
twice as broad as the bars, four to six on the radius. Radial spines very
numerous, conical, nearly as long as the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.24, inner 0.06; outer pores
0.008 to 0.02, bars 0.005, inner pores 0.006, bars 0.003; length of the
spines 0.2.

_Habitat._--Arctic Ocean, Greenland ("Alert" Expedition).

Subgenus 4. _Haliommura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size and
form; spines not covering the entire surface, but scattered at intervals
(their number smaller than that of the nodal-points in the network).

{238}26. _Haliomma macrodoras_, n. sp. (Pl. 28, figs. 6, 6_a_).

Cortical shell thin walled, twice as broad as the medullary shell, with
irregular, polygonal meshes, three to six times as broad as the bars. Inner
shell with regular, hexagonal meshes, ten times as broad as the bars.
Twenty radial spines, very long, stout, three-sided prismatic, two to four
times as long as the diameter of the shell, as broad as one medullary mesh,
with three wing-like, denticulated edges. At the base of each spine three
supporting curved beams.

_Dimensions._--Diameter of the outer shell 0.14 to 0.16, of the inner 0.07
to 0.08; outer pores 0.01 to 0.02, bars 0.003; inner pores 0.01, bars
0.001; length of the spines 0.3 to 0.6, breadth 0.01.

_Habitat._--Central Pacific, Station 271, surface.

27. _Haliomma antarcticum_, n. sp.

Cortical shell very thin walled, three times as broad as the medullary
shell, with irregular, polygonal pores, and very thin thread-like bars.
Inner shell with regular, hexagonal meshes, six times as broad as the bars.
About forty radial spines, angular, pyramidal, half as long as the radius
of the shell, as broad at the base as the largest mesh.

_Dimensions._--Diameter of the outer shell 0.2, of the inner 0.07; outer
pores 0.006 to 0.015, inner 0.008; length of the spines 0.05, basal breadth
0.012.

_Habitat._--Antarctic Ocean, Station 154, surface.

28. _Haliomma wyvillei_, Haeckel.

_Haliomma wyvillei_, Haeckel, 1878, Protistenreich, p. 44, fig. 31.

_Haliomma species_, Wyville Thomson, 1877, Atlantic, vol. i. p. 236, fig.
54.

Cortical shell thin walled, three times as broad as the medullary shell.
Both shells with irregular, large, polygonal pores, and very thin bars.
From the inner shell arise very numerous (eighty to one hundred and twenty
or more) radial spines, which pierce the outer shell, and are outside it,
as long as the radius of the inner shell, straight, bristle-shaped, and as
thick as the bars.

_Dimensions._--Diameter of the outer shell 0.18, of the inner 0.06; pores
0.01 to 0.02, bars 0.001 to 0.002; length of the free spines 0.06.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

29. _Haliomma beroes_, Ehrenberg.

_Haliomma beroes_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvB. B. iv. fig.
19.

_Haliomma beroes_, Haeckel, 1862, Monogr. d. Radiol., p. 434.

Cortical shell thin walled, three times as broad as the medullary shell,
with irregular, roundish pores, twice to four times as broad as the bars.
Inner shell with regular, circular pores, twice as broad as the bars. Both
shells connected by four (or six ?) radial beams, perpendicularly crossed.
Thirty to forty radial spines, conical, thin, shorter than the radius.

_Dimensions._--Diameter of the outer shell 0.11, inner 0.04; outer pores
0.007 to 0.012, inner 0.005; length of the spines 0.03, basal breadth
0.005.

_Habitat._--Atlantic, depth 2000 fathoms.

{239}30. _Haliomma compactum_, n. sp. (Pl. 28, figs. 5, 5_a_).

Cortical shell very thick walled, four times as broad as the thin walled
medullary shell; pores of the former irregular, roundish, with high
polygonal frames of very different size, twice to four times as broad as
the bars. Inner shell with simple, small, polygonal pores, connected with
the outer shell by eight (?) regularly disposed thin radial beams, opposite
in pairs, prolonged outside into strong conical spines, longer than the
radius.

_Dimensions._--Diameter of the outer shell 0.08, inner 0.02; outer pores
0.005 to 0.01, inner 0.005; length of the spines 0.06, basal breadth 0.012.

_Habitat._--Central Pacific, Stations 265 to 272, depth 2425 to 2925
fathoms.

31. _Haliomma permagnum_, n. sp.

Cortical shell thick walled, six times as broad as the medullary shell,
with irregular, roundish, double-edged pores, three to six times as broad
as the bars. Inner shell with regular, circular pores, twice as broad as
the bars. One hundred and twenty to one hundred and fifty radial spines,
conical, only one-fourth as long as the radius.

_Dimensions._--Diameter of the outer shell 0.42, inner 0.07; outer pores
0.012 to 0.025, bars 0.004; inner pores 0.008, bars 0.004; length of the
spines 0.05, basal breadth 0.03.

_Habitat._--Central Pacific, Station 268, surface.

32. _Haliomma patagonicum_, n. sp.

Cortical shell thick walled, four times as broad as the medullary shell,
with irregular, roundish pores, twice to three times as broad as the bars.
Inner shell also with irregular, roundish pores, of half the size. Forty to
fifty radial spines, cylindro-conical, about as long as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.24, inner 0.06; outer pores
0.012 to 0.02, bars 0.006; inner pores 0.005 to 0.01, bars 0.004; length of
the spines 0.3, breadth 0.02.

_Habitat._--South-east Pacific, west coast of Patagonia, Station 302,
surface.

33. _Haliomma clavatum_, n. sp.

Cortical shell thick walled, three times as broad as the medullary shell,
with irregular, roundish pores little larger than the bars; eight to twelve
on the radius. Inner shell with similar but smaller pores. About twenty
radial spines, club-shaped, as long as the radius, three-sided, with
prominent edges, twice as broad at the distal end as at the base. (Similar
to _Acanthosphæra clavata_, Pl. 26, fig. 8, but differs from it in the
possession of a medullary shell.)

_Dimensions._--Diameter of the outer shell 0.2, inner 0.06; outer pores and
bars 0.008 to 0.02, inner 0.006 to 0.01; length of the spines 0.1, distal
breadth 0.03.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

{240}Genus 95. _Heliosoma_,[130] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Astrosphærida# with one medullary (intracapsular) and one
cortical (extracapsular) shell, the two being connected by radial beams
piercing the central capsule. Shell surface covered with simple radial
spines of two different kinds; larger main spines and smaller by-spines.

The genus _Heliosoma_ differs from the preceding _Haliomma_ in the
possession of two different kinds of radial spines, and exhibits therefore
the same relation to it that _Heliosphæra_ bears to _Acanthosphæra_ among
the Coscinommida. The smaller by-spines are much more numerous than the
larger main spines.

Subgenus 1. _Heliosomantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form.

1. _Heliosoma radians_, n. sp. (Pl. 28, figs. 3, 3_a_).

Cortical shell very thin walled, with thread-like bars and regular,
hexagonal meshes; twenty to twenty-two on the radius. Medullary shell only
one-fifth as large, of the same structure. The two shells connected by
twenty very thin radial beams, which are prolonged outside into twenty
stout, three-sided pyramidal main spines, as long as the diameter of the
inner shell. Between these, arising from the surface, numerous
bristle-shaped by-spines.

_Dimensions._--Diameter of the outer shell 0.25, inner 0.05, pores 0.012;
length of the main spines 0.05, basal breadth 0.012.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

2. _Heliosoma elegans_, n. sp.

Cortical shell thick walled, with regular, circular, hexagonally framed
pores, twice as broad as the bars; ten to twelve on the radius. Medullary
shell half as large, with simple, regular, circular pores, twice as broad
as the bars. The two shells connected by twenty very thin radial beams,
which are prolonged outside into twenty slender, three-sided pyramidal main
spines as long as the radius. In each corner of the surface hexagons a
bristle-shaped by-spine one-fourth as long.

_Dimensions._--Diameter of the outer shell 0.12, inner 0.06; outer pores
0.006, bars 0.003; length of the main spines 0.05, basal breadth 0.01.

_Habitat._--Central Pacific, Station 271, surface.

3. _Heliosoma echinaster_, Haeckel.

_Haliomma echinaster_, Haeckel, 1862, Monogr. d. Radiol., p. 429, Taf.
xxiv. figs. 1-3.

Cortical shell thin walled, with regular, circular, hexagonally framed
pores, twice as broad as the bars; ten to twelve on the radius. Medullary
shell one-fourth as large, with few irregular, large, {241}polygonal pores
and very thin thread-like bars, connected with the former by nine similar
thin radial beams, which are prolonged outside into nine pyramidal main
spines, as long as the radius. From each corner of the surface hexagon
arises a bristle-shaped by-spine, half as long as the main spine.

_Dimensions._--Diameter of the outer shell 0.16, inner 0.04; outer pores
0.008, bars 0.004; length of the main spines 0.08, basal breadth 0.01.

_Habitat._--Mediterranean (Messina), surface.

4. _Heliosoma duodecilla_, n. sp.

Cortical shell thick walled, with regular, circular pores, of the same
breadth as the bars; sixteen to eighteen on the radius. Medullary shell
one-third as large, of the same structure. Twelve radial main spines,
regularly disposed, cylindro-conical, longer than the shell diameter, as
broad as two pores. Whole surface covered with short conical by-spines.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.05; outer pores
and bars 0.007; length of the main spines 0.2, breadth 0.02.

_Habitat._--South Atlantic, Station 333, surface.

Subgenus 2. _Heliosomura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size and
form.

5. _Heliosoma hastatum_, n. sp. (Pl. 28, fig. 4).

Cortical shell very thin walled, with large, irregular, polygonal meshes,
three to six times as broad as the bars; four to six on the radius.
Medullary shell one-third as large, with small, regular, circular pores.
The two shells connected by twelve regularly disposed radial beams, which
are prolonged outside into twelve very stout three-sided pyramidal main
spines, somewhat longer than the radius, spear-shaped; each of the three
wings in the middle part with one tooth. Scattered on the surface numerous
by-spines of the same form, half as long and only one-fourth as broad.

_Dimensions._--Diameter of the outer shell 0.1, inner 0.033; outer pores
0.006 to 0.013, inner pores 0.002, bars 0.002; length of the main spines
0.06, breadth 0.015.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

6. _Heliosoma indicum_, n. sp.

Cortical shell thick walled, with irregular, roundish pores, twice to five
times as broad as the bars. Medullary shell of the same structure, only
one-fourth as large. Thirty to forty conical main spines, as long as the
radius; between them numerous bristle-shaped by-spines, only half as long.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.05; outer pores
0.004 to 0.01, bars 0.002; length of the main spines 0.1, breadth 0.01.

_Habitat._--Indian Ocean, between Aden and Socotora, surface, Haeckel.

{242}Genus 96. _Elatomma_,[131] n. gen.

The genus _Elatomma_ differs from _Haliomma_ in the ramification of the
radial spines, and exhibits therefore the same relation to it that
_Cladococcus_, among the Coscinommida, bears to _Acanthosphæra_.

Subgenus 1. _Elatommella_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form.

1. _Elatomma pinetum_, n. sp.

Cortical shell thick walled, with regular, circular, hexagonally framed
pores, three times as broad as the bars. From its surface arise twenty to
thirty large, three-sided prismatic, radial spines as long as the shell
diameter, branched like a pine tree; on each edge of the spine five to six
ramified branches, decreasing in size towards the distal end. Similar to
_Cladococcus pinetum_ (Pl. 27, figs. 1, 3), but differing in the possession
of a medullary shell (one-third as large as the outer), with regular,
circular pores, as broad as the bars.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.07; cortical pores
0.012, bars 0.004; length of the spines 0.2 to 0.24.

_Habitat._--Central Pacific, Stations 266 to 272, surface.

2. _Elatomma scoparium_, n. sp.

Cortical shell thick walled, with regular, circular pores, twice as broad
as the bars, connected with the medullary shell by twenty thin radial
beams, which are prolonged outside into twenty large cylindrical radial
spines somewhat longer than the radius; in the proximal half simple, in the
distal half with six to nine dichotomous, irregular branches. Similar to
_Cladococcus scoparius_ (Pl. 27, fig. 2), but much larger, and with a large
medullary shell (one-third as broad as the outer), and regular, circular
pores.

_Dimensions._--Diameter of the outer shell 0.12, inner 0.04; cortical pores
0.01, bars 0.005; length of the spines 0.08.

_Habitat._--North Pacific, Station 240, surface.

Subgenus 2. _Elatommura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form.

{243}3. _Elatomma juniperinum_, n. sp. (Pl. 28, fig. 8).

Cortical shell thin walled, with regular, polygonal, or roundish pores,
twice to four times as broad as the bars, connected with the small
medullary shell by about twenty thin radial beams. Inner shell only
one-eighth as broad as the outer, with few irregular, polygonal meshes.
Surface covered with very numerous (one hundred to one hundred and fifty or
more) branched conical radial spines, one-third as long as the shell
radius, with six to twelve short lateral branches.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.025; cortical pores
0.08 to 0.016, bars 0.004; length of the spines 0.04.

_Habitat._--Central Pacific, Station 271, surface.

4. _Elatomma penicillus_, n. sp.

Cortical shell thin walled, with irregular, polygonal pores, twice to four
times as broad as the bars, connected with the medullary shell by twenty
(or more?) very thin beams. These are prolonged outside into twenty
straight, three-sided prismatic, radial spines as long as the radius, with
a brush-like bunch of six to nine short, irregularly ramified branches at
the end. Medullary shell very delicate (half as broad as the outer), with
regular, hexagonal meshes and thread-like bars.

_Dimensions._--Diameter of the outer shell 0.15, inner 0.08; length of the
spines 0.08.

_Habitat._--West coast of Norway, Bergen, Haeckel.

5. _Elatomma irregulare_, n. sp.

Cortical shell thin walled, with irregular, polygonal, or roundish meshes,
twice to six times as broad as the bars; the medullary shell of the same
structure, half as broad, with smaller irregular pores. Forty to sixty
curved, three-sided, radial spines, as long as the shell, with a bunch of
very irregular, curved, and partly ramified branches at the distal end.

_Dimensions._--Diameter of the outer shell 0.2, inner 0.1; length of the
spines 0.2, breadth 0.01.

_Habitat._--North Atlantic, Gulf Stream, Færöe Channel, surface, John
Murray.

Genus 97. _Leptosphæra_,[132] n. gen.

_Definition._--#Astrosphærida# with two extracapsular cortical shells
without by-spines, connected by long prismatic radial spines.

The genus _Leptosphæra_, together with the three following genera, forms
the very peculiar and typical small group of #Sphæroidea# which we call the
Diplosphærida (with four genera and twenty-four species); their shell is
composed of two concentric spheres as in the three foregoing genera; but
whilst in these the inner shell is an intracapsular medullary shell and the
outer an extracapsular cortical shell, in the Diplosphærida {244}both
shells are extracapsular or cortical shells, therefore the inner shell of
the latter corresponds to the outer of the former. The inner spherical
shell of all Diplosphærida is composed of very delicate beams and large
pores, which are either regular hexagonal or irregular polygonal (never
roundish). From its surface arise a variable number (twenty to thirty) of
stout long radial spines, which are invariably longer than the shell
diameter (often two to three times as long or more), and of three-sided
prismatic form, the three edges either smooth or serrate, often with three
rows of lateral branches (commonly three to five branches in each row); the
latter are invariably of the same form, concavely curved towards the spine,
and decrease in size towards the distal end. From the three edges of each
main spine in all Diplosphærida, at equal distances from the centre, arise
six very thin, thread-like lateral branches (a pair from each edge) and
connect the spine in a tangential direction with all neighbouring spines.
In this manner the polyhedral outer shell is formed, the meshes of which
therefore are always very large and triangular. Sometimes each of these
primary triangular meshes becomes filled up with a secondary network,
either of regular quadrangular or of irregular polygonal secondary meshes.
Besides the constant twenty to thirty large main spines, in the majority of
Diplosphærida bristle-shaped radial by-spines arise, either from the inner
shell (_Diplosphæra_) or from the outer (_Drymosphæra_), or from both
(_Astrosphæra_). They are absent only in _Leptosphæra_. Commonly the
by-spines are simple, rarely forked or branched. The central capsule in the
Diplosphærida is usually enclosed in the inner shell; often it completely
fills up the latter, or drives out a cæcal protuberance through each mesh;
but these processes rarely unite outside. The average size of the
Diplosphærida, which are all pelagic organisms, is much larger than that of
the other Haliommida.

Subgenus 1. _Leptosphærella_, Haeckel.

_Definition._--Radial spines simple, without lateral branches.

1. _Leptosphæra hexagonalis_, n. sp. (Pl. 19, fig. 2).

Inner shell with regular, hexagonal meshes, and very thin, thread-like
bars; outer shell twice as broad, with simple triangular meshes. Radial
spines with three smooth edges. (Fig. 2 represents the central capsule with
numerous club-shaped saccules, prominent externally through the meshes; in
the centre a large simple spherical nucleus, one-third as broad as the
capsule. The skeleton of this species is identical with that of
_Diplosphæra hexagonalis_, Pl. 19, fig. 3, but has no by-spines.)

_Dimensions._--Diameter of the outer shell 0.3, of the inner 0.15.

_Habitat._--Cosmopolitan; Mediterranean (Corfu), North Atlantic (Canary
Islands), Tropical Pacific, surface.

{245}2. _Leptosphæra serrata_, n. sp.

Inner shell with regular, hexagonal meshes, five to six times as broad as
the bars; outer shell three times as broad, with simple triangular meshes.
Radial spines with three serrated edges (of the same form as the main
spines of _Drymosphæra dendrophora_, Pl. 20, fig. 1).

_Dimensions._--Diameter of the outer shell 0.45, of the inner 0.15.

_Habitat._--Central Pacific, Station 271, surface.

3. _Leptosphæra polygonalis_, n. sp.

Inner shell with irregular, polygonal meshes and very thin thread-like
bars; outer shell twice as broad, with simple triangular meshes. Radial
spines with three smooth edges. (Resembles _Drymosphæra polygonalis_, Pl.
19, fig. 1, but has no by-spines.)

_Dimensions._--Diameter of the outer shell 0.35, inner 0.175.

_Habitat._--North Pacific, Station 256, surface.

Subgenus 2. _Leptosphæromma_, Haeckel.

_Definition._--Radial spines with three rows of lateral branches (one row
on each edge).

4. _Leptosphæra ciliata_, n. sp.

Inner shell with regular, hexagonal meshes and very thin thread-like bars;
outer shell three times as broad, with simple triangular meshes. Radial
spines with smooth edges and three rows of simple, smooth, curved, lateral
branches (six branches on each edge), similar to those of _Diplosphæra
gracilis_.

_Dimensions._--Diameter of the outer shell 0.6, inner 0.2.

_Habitat._--South Atlantic, Station 325, surface.

5. _Leptosphæra spinosa_, Haeckel.

_Diplosphæra spinosa_, R. Hertwig, 1879, Organismus d. Radiol., p. 40,
Taf. v. fig. 2.

Inner shell with regular, hexagonal meshes, five times as broad as the
bars; outer shell three times as broad, with simple triangular meshes.
Radial spines with three densely serrated edges, and with three rows of
serrated, simple, curved, lateral branches (three branches on each edge).
Differs from the preceding in the thicker bars and the elegantly
denticulated spines and branches.

_Dimensions._--Diameter of the outer shell 0.66, inner 0.22.

_Habitat._--Mediterranean (Messina), R. Hertwig.

6. _Leptosphæra stellata_, n. sp.

Inner shell with regular, hexagonal meshes, six times as broad as the bars,
each bar crossed by a transverse tangential rod, so that each mesh
represents an elegant six-rayed star (as {246}in _Astrosphæra stellata_,
Pl. 19, fig. 5, but without radial by-spines); outer shell twice as broad,
with simple triangular meshes. Radial spines with serrated edges and three
rows of simple lateral branches (four branches on each edge).

_Dimensions._--Diameter of the outer shell 0.44, inner 0.22.

_Habitat._--Tropical Atlantic (Ascension Island), Station 343, surface.

7. _Leptosphæra reticulum_, n. sp.

Inner shell with irregular, polygonal meshes and very thin thread-like
bars; outer shell four times as broad, also with irregular, polygonal
meshes, the sides of the triangular main meshes being connected by
irregular lateral ramules, forming an extremely delicate reticulum.

_Dimensions._--Diameter of the outer shell 0.64, inner 0.16.

_Habitat._--South Pacific, Station 291, surface.

Genus 98. _Diplosphæra_,[133] Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 804.

_Definition._--#Astrosphærida# with two extracapsular cortical shells,
connected by long, prismatic, radial spines; inner shell with thin radial
by-spines.

The genus _Diplosphæra_ differs from its ancestral form _Leptosphæra_ in
the development of radial by-spines on the surface of the inner shell, the
outer shell being smooth.

Subgenus 1. _Diplosphærella_, Haeckel.

_Definition._--Radial main spines simple, without lateral branches.

1. _Diplosphæra hexagonalis_, n. sp. (Pl. 19, fig. 3).

Inner shell with regular, hexagonal meshes and very thin thread-like bars;
at each nodal-point one bristle-shaped by-spine; outer shell twice as
broad, with simple triangular meshes. Radial spines with three smooth
edges. (The radially striped central capsule, enclosed in the inner shell,
envelops a large central nucleus one-third its size, fig. 3.)

_Dimensions._--Diameter of the outer shell 0.36, inner 0.18.

_Habitat._--Central Pacific, Stations 265 to 274, surface.

2. _Diplosphæra ornata_, n. sp.

Inner shell with regular, hexagonal meshes, four times as broad as the
bars; outer shell three times as broad, with simple triangular meshes.
Radial main spines as well as the bars of both {247}shells very elegantly
denticulated; radial by-spines (very numerous in the surface of the inner
shell) smooth.

_Dimensions._--Diameter of the outer shell 0.7, inner 0.24.

_Habitat._--Indian Ocean, Madagascar, surface, Rabbe.

3. _Diplosphæra dictyota_, n. sp.

Inner shell with regular, hexagonal meshes, six times as broad as the bars;
at each nodal-point one radial by-spine; outer shell two and a half times
as broad, with very delicate square meshes, separated by thread-like bars
which arise from the sides of the triangular main meshes. Radial main
spines with three serrated edges. (Differs from the similar _Diplosphæra
gracilis_ in the simple main spines, with delicate dentition of the edges.)

_Dimensions._--Diameter of the outer shell 0.5, inner 0.2.

_Habitat._--North Atlantic, Canary Islands, surface.

4. _Diplosphæra polygonalis_, n. sp.

Inner shell with irregular, polygonal meshes and very thin thread-like
bars, covered with very numerous bristle-shaped by-spines; outer shell
twice as broad, with simple triangular meshes. Radial main spines with
three smooth edges.

_Dimensions._--Diameter of the outer shell 0.44, inner 0.22.

_Habitat._--South Pacific, Station 288, surface.

Subgenus 2. _Diplosphæromma_, Haeckel.

_Definition._--Radial main spines with three rows of lateral branches (one
row on each edge).

5. _Diplosphæra gracilis_, Haeckel.

_Diplosphæra gracilis_, Haeckel, 1862, Monogr. d. Radiol., p. 354, Taf.
x. fig. 1.

Inner shell with regular, hexagonal meshes and very thin bars; at each
nodal-point one simple bristle-shaped radial by-spine; outer shell twice as
broad, with very delicate square meshes, separated by thread-like bars
which arise from the sides of the triangular main meshes. Radial spines
with smooth edges and with three rows of simple curved lateral branches
(four to six branches on each edge).

_Dimensions._--Diameter of the outer shell 0.54, inner 0.27.

_Habitat._--Mediterranean (Messina).

6. _Diplosphæra denticulata_, n. sp.

Inner shell with regular, hexagonal meshes, four times as broad as the
bars; at each nodal-point is a simple bristle-shaped by-spine; outer shell
three times as broad, with simple triangular meshes. {248}Radial spines
very strong, with dentated edges and with three rows of lateral branches
(six dentated curved branches on each edge).

_Dimensions._--Diameter of the outer shell 0.4, inner 0.13.

_Habitat._--North Atlantic, Azores, surface.

7. _Diplosphæra reticulata_, n. sp.

Inner shell with regular, hexagonal meshes, five times as broad as the
bars, which are densely covered with very numerous, simple bristle-shaped
by-spines; outer shell four times as broad, with very numerous small
irregular polygonal meshes, forming a very delicate network, and filling
out the large triangular main meshes. Radial spines with three serrated
edges and three rows of short curved branches (four to five branches on
each edge).

_Dimensions._--Diameter of the outer shell 0.6, inner 0.15.

_Habitat._--North Atlantic, Gulf Stream, Færöe Channel, surface, John
Murray.

8. _Diplosphæra triglochin_, n. sp.

Inner shell with irregular, polygonal meshes and very thin thread-like
bars; outer shell three times as broad, with similar irregular polygonal
meshes, filling out the large triangular main meshes. Radial spines with
three serrated edges, each of which bears one single large lateral branch,
concavely curved towards the distal end. By-spines of the inner shell very
numerous.

_Dimensions._--Diameter of the outer shell 0.5, inner 0.17.

_Habitat._--Central Pacific, Station 264, surface.

Genus 99. _Drymosphæra_,[134] Haeckel, 1881, Prodromus, p. 452.

_Definition._--#Astrosphærida# with two extracapsular cortical shells,
connected by long prismatic, radial spines; outer shell with thin radial
by-spines.

The genus _Drymosphæra_ differs from its ancestral form _Leptosphæra_ in
the development of radial by-spines on the surface of the outer shell, the
inner shell being smooth.

Subgenus 1. _Drymosphærella_, Haeckel.

_Definition._--Radial by-spines simple, not branched (main spines simple).

1. _Drymosphæra hexagonalis_, n. sp.

Inner shell with regular, hexagonal meshes and very thin thread-like bars;
outer shell twice as broad, with simple triangular meshes and thin
thread-like bars, which bear very numerous, {249}bristle-shaped radial
by-spines. Radial main spines with three smooth edges. (Differs from the
first species of the other three genera of Diplosphærida in the presence of
by-spines on the outer, and their absence on the inner shell.)

_Dimensions._--Diameter of the outer shell 0.32, inner 0.16.

_Habitat._--Central Pacific, Station 274, surface.

2. _Drymosphæra polygonalis_, n. sp. (Pl. 19, fig. 1).

_Diplosphæra polygonalis_, Haeckel, 1879, Atlas.

Inner shell with irregular, polygonal meshes and very thin thread-like
bars; outer shell one and a half times as broad, with simple triangular
meshes and thicker bars, which bear numerous bristle-shaped radial
by-spines (in the figure the majority of these are broken off). Radial main
spines with three smooth edges. (Similar to _Leptosphæra polygonalis_ and
_Diplosphæra polygonalis_, but differs from both in the presence of
by-spines on the outer, and their absence on the inner shell.)

_Dimensions._--Diameter of the outer shell 0.33, inner 0.22.

_Habitat._--North Pacific, Station 236 south of Japan, surface.

Subgenus 2. _Drymosphæromma_, Haeckel.

_Definition._--Radial by-spines forked or branched (main spines simple).

3. _Drymosphæra furcata_, n. sp.

Inner shell with irregular, polygonal meshes and thin bars, the former five
times as broad as the latter; outer shell two and a half times as broad,
with simple triangular meshes and smooth bars, bearing numerous long smooth
bristle-shaped by-spines, which in the distal half are forked. Radial main
spines with three smooth edges.

_Dimensions._--Diameter of the outer shell 0.4, inner 0.16.

_Habitat._--Indian Ocean, Ceylon, surface, Haeckel.

4. _Drymosphæra cladophora_, n. sp.

Inner shell with irregular, polygonal meshes, three to six times as broad
as the smooth bars; outer shell twice as broad, with simple triangular
meshes and very thin smooth bars, bearing numerous curved, irregularly
branched, smooth by-spines. Radial main spines with three serrated edges.

_Dimensions._--Diameter of the outer shell 0.4, inner 0.2.

_Habitat._--Central Pacific, Station 276, surface.

5. _Drymosphæra dendrophora_, n. sp. (Pl. 20, figs. 1, 1_a_, 1_b_).

Inner shell with irregular, polygonal meshes and very thin thread-like
bars; outer shell one and a half times as broad, with simple triangular
meshes and thicker bars, which bear a forest of {250}very numerous,
repeatedly dichotomous or irregularly branched, curved by-spines, longer
than the diameter of the outer shell. Radial main spines with three
dentated edges. All parts of the skeleton, the net bars as well as the
radial beams and spines, are very elegantly denticulated (fig. 1_b_). The
central capsule (fig. 1_a_) completely distends the inner shell and forces
out protuberances through all its pores; in its centre lies a nucleus
one-third its size.

_Dimensions._--Diameter of the outer shell 0.25, inner 0.16.

_Habitat._--Central Pacific, Station 271, surface.

Genus 100. _Astrosphæra_,[135] n. gen.

_Definition._--#Astrosphærida# with two extracapsular cortical shells,
connected by long, prismatic, radial spines; inner and outer shell with
thin radial by-spines.

The genus _Astrosphæra_ differs from its ancestral form, _Leptosphæra_, in
the development of radial by-spines on the surface of both shells.

Subgenus 1. _Astrosphærella_, Haeckel.

_Definition._--Radial main spines simple without lateral branches.

1. _Astrosphæra hexagonalis_, n. sp. (Pl. 19, fig. 4).

Inner shell with regular, hexagonal meshes and very thin bars, having a
bristle-shaped, short radial by-spine in each hexagon-corner; outer shell
twice as broad, with simple triangular meshes and thicker bars, bearing one
row of simple bristle-shaped, curved, radial by-spines. Radial main spines
with three smooth edges.

_Dimensions._--Diameter of the outer shell 0.32, inner 0.16.

_Habitat._--South Pacific (West Patagonia), Station 302, surface.

2. _Astrosphæra splendens_, n. sp.

Inner shell with regular, hexagonal meshes, four times as broad as the
bars, and covered with numerous curved, long, bristle-shaped by-spines;
outer shell three times as broad, with simple triangular meshes and thin
bars, bearing a row of very long, curved, bristle-shaped by-spines. Radial
main spines with three dentated edges. All parts of the skeleton elegantly
denticulated, as in _Drymosphæra dendrophora_ (Pl. 20, fig. 1).

_Dimensions._--Diameter of outer shell 0.7, inner 0.24.

_Habitat._--Tropical Atlantic, Station 347, surface.

{251}Subgenus 2. _Astrosphæromma_.

_Definition._--Radial main spines with three rows of lateral branches (one
row on each edge).

3. _Astrosphæra sideræa_, n. sp.

Inner shell with regular, hexagonal meshes, six times as broad as the bars,
and covered with numerous short bristle-shaped by-spines; outer shell four
times as broad, with simple triangular meshes, and rows of long
bristle-shaped by-spines arising from the bars. Radial main spines with
three serrated edges and three rows of lateral branches (four branches on
each edge).

_Dimensions._--Diameter of the outer shell 0.6, inner 0.15.

_Habitat._--Central Pacific, Station 266, surface.

4. _Astrosphæra stellata_, n. sp. (Pl. 19, fig. 5).

_Diplosphæra stellata_, Haeckel, 1881, Prodrom. et Atlas.

Inner shell with regular, hexagonal, six-rayed meshes, each bar being
crossed by a transverse tangential rod, at each nodal-point a long
bristle-shaped by-spine; outer shell three times as broad, with simple
triangular meshes and denticulated thin bars, bearing a row of
bristle-shaped radial by-spines. Radial main spines in the proximal half
with three serrated edges, in the distal half with three rows of curved
lateral branches (five branches on each edge).

_Dimensions._--Diameter of the outer shell 0.6, inner 0.2.

_Habitat._--Central Pacific, Station 274, surface.

Subfamily ACTINOMMIDA,[136] Haeckel, 1862, Monogr. d. Radiol., p. 440
(_sensu emendato_).

_Definition._--#Astrosphærida# with three concentric, spherical,
lattice-shells, united by radial beams.

Genus 101. _Actinomma_,[137] Haeckel, 1862, Monogr. d. Radiol., p. 440.

_Definition._--#Astrosphærida# with three concentric lattice-spheres and
numerous simple radial spines of one kind.

The genus _Actinomma_ is here restricted to those #Sphæroidea# which
combine the possession of three concentric lattice-shells with numerous
radial spines on the surface; the spines are all simple, being of one and
the same kind. Commonly two of the three shells are intracapsular medullary
shells, connected by radial beams (piercing {252}the central capsule) with
the outer, extracapsular, cortical shell. But in some species only one
medullary shell is enclosed in the central capsule, whilst both other
shells lie outside it. In such case the distance between these two cortical
shells is much smaller than their distance from the simple internal
medullary shell. These forms correspond more to aculeate _Rhodosphæræ_,
whilst the others resemble aculeate _Thecosphæræ_.

Subgenus 1. _Actinommantha_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form; spines on the entire surface (commonly one spine at each
nodal-point).

1. _Actinomma hexagonium_, n. sp.

Cortical shell, as well as both medullary shells, very thin walled, with
regular, hexagonal pores and thread-like bars between them. Pores of the
outer shell twice as broad as those of the middle, and three times as broad
as those of the inner shells. Radial proportion of the three spheres =
1 : 3 : 9; about twenty thin radial beams between them. At each nodal-point
of the surface arises one bristle-shaped radial spine, half as long as the
radius.

_Dimensions._--Diameter of the outer shell 0.22, middle 0.07, inner 0.025;
meshes of the cortical shell 0.01; length of the spines 0.05.

_Habitat._--Central Pacific, Stations 270 to 274, surface.

2. _Actinomma facetum_, n. sp.

Cortical shell thick walled, with regular, circular, hexagonally framed
pores, three times as broad as the bars. Pores of both medullary shells
regular, circular. Radial proportion of the three spheres = 1 : 2 : 4;
about forty thin radial beams between them. At each nodal-point of the
surface arises one short, three-sided pyramidal, radial spine, about
one-third as long as the radius.

_Dimensions._--Diameter of the outer shell 0.16, middle 0.08, inner 0.04;
cortical pores 0.012, bars 0.004; length of the spines 0.03, basal breadth
0.015.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

3. _Actinomma anthomma_, n. sp.

Cortical shell thick walled, with regular, six-lobed pores, twice as broad
as the bars. At each nodal-point of the surface is one short conical radial
spine, as long as the diameter of the pores; one corresponding to each
lobe, and there is therefore around each pore a regular corona of six
spines, as in _Haliomma lirianthus_ (Pl. 28, fig. 1_b_). Both medullary
shells with simple, circular, regular pores. Radial proportion of the three
spheres = 1 : 2.5 : 7; radial beams between them only six, opposite in
pairs in the three dimensive axes.

{253}_Dimensions._--Diameter of the outer shell 0.2, middle 0.075, inner
0.003; cortical pores 0.01, bars 0.005; length of the spines 0.01.

_Habitat._--Central Pacific, Station 271, surface.

4. _Actinomma castanomma_, n. sp.

Cortical shell as well as both medullary shells thick walled, with regular,
circular pores, twice as broad as the bars; between them at each
nodal-point one bristle-shaped radial spine, one-third as long as the
radius, with conical base. Radial proportion of the three spheres =
2 : 3 : 8; only six radial beams between them (opposed in pairs in the
three dimensive axes).

_Dimensions._--Diameter of the outer shell 0.15, middle 0.06, inner 0.04;
cortical pores 0.008, bars 0.004; length of the spines 0.025.

_Habitat._--South Atlantic, Station 333, surface.

5. _Actinomma entactinia_, Stöhr.

_Actinomma entactinia_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 93, Taf.
ii. fig. 12.

Cortical shell as well as both medullary shells thick walled, with regular,
circular pores, of the same breadth as the bars. Radial proportion of the
three spheres = 1 : 3 : 8; radial beams between them very numerous (thirty
to fifty or more?). Entire surface covered with short thick conical
spines, only one-fifth as long as the radius.

_Dimensions._--Diameter of the outer shell 0.16, middle 0.06, inner 0.02;
cortical pores and bars 0.007; length of the spines 0.015.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Caltanisetta.

Subgenus 2. _Actinommetta_, Haeckel.

_Definition._--Pores of the outer shell regular, of nearly equal size and
similar form; spines not over the entire surface, but scattered at
intervals (their number smaller than that of the nodal-points).

6. _Actinomma japonicum_, n. sp.

Cortical shell thick walled, with regular, circular, hexagonally framed
pores, three times as broad as the bars. Both medullary shells with simple,
circular, regular pores, of the same breadth as the bars. Radial proportion
of the three spheres = 2 : 5 : 9; radial beams between them twenty,
prolonged outside into twenty symmetrically disposed, three-sided pyramidal
spines, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.18, middle 0.1, inner 0.04;
cortical pores 0.01, bars 0.003; length of the spines 0.1, basal breadth
0.012.

_Habitat._--North Pacific, Station 240, east of Japan, surface.

{254}7. _Actinomma denticulatum_, n. sp. (Pl. 29, fig. 3).

Cortical shell thick walled, with regular, circular, double-edged pores,
twice as broad as the bars. Forty to fifty radial spines, three-sided
pyramidal, half as long as the radius, with three elegantly denticulated
edges. Radial proportion of the three spheres = 1 : 2 : 5. (Differs from
_Haliomma denticulatum_ only in the double medullary shell and larger
size.)

_Dimensions._--Diameter of the outer shell 0.14, middle 0.06, inner 0.03;
cortical pores 0.01, bars 0.005; length of the spines 0.04, basal breadth
0.01.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

8. _Actinomma trinacrium_, Haeckel.

_Actinomma trinacrium_, Haeckel, 1862, Monogr. d. Radiol., p. 441, Taf.
xxiv. figs. 6-8.

_Haliomma trinacrium_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 815.

Cortical shell as well as both medullary shells thin walled, with
subregular, circular pores, twice as broad as the bars. Radial proportion
of the three spheres = 1 : 3 : 9; radial beams between them twenty,
prolonged outside into strong, three-sided pyramidal spines, as long as the
radius; between them, on the surface, numerous similar spines. Sometimes
the latter remain smaller, the pores more irregular, and then this species
corresponds to _Echinomma trinacrium_.

_Dimensions._--Diameter of the outer shell 0.09, middle 0.03, inner 0.01;
cortical pores 0.008, bars 0.004; length of the spines 0.04, basal breadth
0.01.

_Habitat._--Mediterranean (Messina, Corfu, Haeckel), surface.

9. _Actinomma pachyderma_, n. sp. (Pl. 29, figs. 4, 5).

Cortical shell very thick walled, with regular, circular, double-edged
pores, twice as broad as the bars. Both medullary shells with simple,
small, circular pores, of the same breadth as the bars. Radial proportion
of the three spheres = 1 : 2 : 4; radial spines about twenty, conical,
shorter than the radius, prolonged inside into twenty very thin connecting
beams (fig. 4).

_Dimensions._--Diameter of the outer shell 0.08, middle 0.04, inner 0.02;
cortical pores 0.01; bars 0.005; length of the spines 0.03, basal breadth
0.01.

_Habitat._--South Pacific, Station 297, depth 1775 fathoms.

Subgenus 3. _Actinommilla_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size and
form; spines over the entire surface (commonly one spine at each
nodal-point).

10. _Actinomma spinigerum_, Stöhr.

_Actinomma spinigerum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 92, Taf.
ii. fig. 10.

Cortical shell thick walled, with irregular, roundish, or subcircular
pores, twice to four times as broad as the bars. Radial proportion of the
three spheres = 1 : 3 : 8; between them eight (?) {255}connecting radial
beams. Entire surface densely covered with small conical spines, not larger
than the pores.

_Dimensions._--Diameter of the outer shell 0.17, middle 0.07, inner 0.023;
cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.01.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

11. _Actinomma hirsutum_, n. sp.

Cortical shell thick walled, with irregular, roundish pores, about the same
breadth as the bars. Radial proportion of the three spheres = 1 : 2 : 8;
between them numerous (twenty to thirty or more) cylindrical connecting
beams. Entire surface densely covered with innumerable small bristle-shaped
spines, as long as the diameter of the inner shell.

_Dimensions._--Diameter of the outer shell 0.25, middle 0.06, inner 0.03;
cortical pores and bars 0.004 to 0.008; length of the spines 0.03.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Subgenus 4. _Actinommura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size and
form; spines not over the entire surface, but scattered at intervals (their
number smaller than that of the nodal-points).

12. _Actinomma capillaceum_, n. sp. (Pl. 29, fig. 6).

Cortical shell very thin walled, with irregular, polygonal meshes (sixteen
to eighteen on the radius), three to six times as broad as the bars. Both
medullary shells with smaller pores of the same structure. Radial
proportion of the three spheres = 1 : 2.5 : 7; radial connecting beams
between them very thin and numerous (one hundred and twenty to one hundred
and fifty or more), each prolonged outside into a short three-sided
pyramidal spine, as long as the radius of the inner shell. (Similar to
_Haliomma capillaceum_.)

_Dimensions._--Diameter of the outer shell 0.27, middle 0.1, inner 0.04;
cortical pores 0.01 to 0.02, bars 0.003; length of the spines 0.02, basal
breadth 0.005.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

13. _Actinomma arcadophorum_, n. sp. (Pl. 29, figs. 7, 8).

Cortical shell very thin walled, with irregular, polygonal meshes, ten to
twenty times as broad as the bars; twelve to sixteen on the radius. Both
medullary shells with similar delicate network. Radial proportion of the
three spheres = 1 : 2 : 6; numerous thin radial beams (forty to eighty or
more) connect both medullary shells and alternate with other beams, which
arise from arcade-shaped {256}protuberances of the outer medullary shell,
and connect it with the cortical shell. Outer prolongations of these form
the bristle-shaped spines of the surface.

_Dimensions._--Diameter of the outer shell 0.22, middle 0.09, inner 0.04;
cortical pores 0.01 to 0.02, bars 0.001 to 0.002; length of the radial
spines 0.2.

_Habitat._--Central Pacific, Station 266, surface.

14. _Actinomma schwageri_, Stöhr.

_Actinomma schwageri_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 92, Taf.
ii. fig. 9_a_, _b_.

Cortical shell thick walled, with irregular, large, polygonal meshes, five
to twelve times as broad as the bars; six to eight on the radius. Both
medullary shells with very small, circular, regular pores. Radial
proportion of the three spheres = 1 : 2 : 9; connecting radial beams
between them nine, prolonged outside into nine short stout, three-sided
pyramidal spines, as long as the diameter of the inner shell.

_Dimensions._--Diameter of the outer shell 0.17, middle 0.04, inner 0.02;
cortical pores 0.01 to 0.03, bars 0.006 and less; length of the spines
0.02, basal breadth 0.01.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

15. _Actinomma dodecomma_, n. sp.

Cortical shell thin walled, with irregular, roundish pores, three to six
times as broad as the bars. Both medullary shells with regular, circular
pores, twice as broad as the bars. Radial proportion of the three shells =
1 : 2.5 : 7; connecting radial beams between them twelve, regularly
disposed, prolonged outside into twelve strong, three-sided pyramidal
spines, as long as the radius.

_Dimensions._--Diameter of the outer shell 0.14, middle 0.05, inner 0.02;
cortical pores 0.008 to 0.018, bars 0.003; length of the spines 0.08,
breadth 0.02.

_Habitat._--Tropical Atlantic, Station 348, depth 2450 fathoms.

16. _Actinomma pachycapsa_, n. sp.

Cortical shell very thick walled, with irregular, roundish pores, twice to
four times as broad as the bars; twelve to sixteen on the radius. Radial
proportion of the three spheres = 1 : 3 : 12. Both medullary shells of
similar irregular structure, connected with the cortical shell by twelve
regularly disposed radial beams, which are prolonged outside into twelve
short stout conical spines, as long as the diameter of the middle shell.

_Dimensions._--Diameter of the outer shell 0.25, middle 0.06, inner 0.02;
cortical pores 0.006 to 0.012, bars 0.003; length of the spines 0.05,
breadth 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

17. _Actinomma giganteum_, n. sp.

Cortical shell thick walled, little larger than the outer medullary shell,
whilst the inner is only one-tenth as large. Pores of all three shells
irregular, roundish, or subcircular, twice to three times {257}as broad as
the bars, on an average twice as large in the outer and middle as in the
inner shell. Sixty to eighty short conical radial spines on the surface,
about as long as the diameter of the inner shell, and one-fourth as broad
at the base. Radial proportion of the three spheres = 1 : 9 : 10.

_Dimensions._--Diameter of the outer shell 0.4 to 0.5, middle 0.35 to 0.45,
inner 0.05 to 0.06.

_Habitat._--Fossil in Barbados.

Genus 102. _Echinomma_,[138] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Astrosphærida# with three concentric lattice-spheres and
numerous simple radial spines of two different kinds; larger main spines
and smaller by-spines.

The genus _Echinomma_, differs from its ancestral form, _Actinomma_, in the
differentiation of the radial spines. Whilst a large number of small
by-spines cover the entire surface, a smaller number of large main spines
are scattered over it, or limited to certain regularly distributed points.
It represents _Heliosoma_ among the Haliommida.

Subgenus 1. _Echinommetta_, Haeckel.

_Definition._--Pores of the cortical shell regular, of nearly equal size
and similar form.

1. _Echinomma echinidium_, n. sp.

Cortical shell thin walled, with regular, hexagonal pores, four times as
broad as the bars; ten to twelve on the radius. Both medullary shells with
regular, circular pores, twice as broad as the bars. Radial proportion of
the three spheres = 1 : 2 : 5. Surface covered with short bristle-shaped
by-spines (one in every hexagon-corner); twenty main spines regularly
disposed, three-sided pyramidal, as long as the diameter of the inner
shell.

_Dimensions._--Diameter of the outer shell 0.1, middle 0.045, inner 0.02;
cortical pores 0.008, bars 0.002; length of the main spines 0.02, breadth
0.01.

_Habitat._--South Pacific, Station 295, surface.

2. _Echinomma cidaris_, n. sp.

Cortical shell thin walled, with regular, circular, hexagonally framed
pores, twice as broad as the bars; six to eight on the radius. Both
medullary shells with simple, circular pores. Radial proportion of the
three spheres = 1 : 2 : 6. Surface covered with short bristle-shaped
by-spines, half as long as the forty to fifty main spines, which are
three-sided pyramidal, and reach the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.18, middle 0.06, inner 0.03;
cortical pores 0.01, bars 0.005; length of the main spines 0.08, basal
breadth 0.015.

_Habitat._--Central Pacific, Station 272, surface.

{258}3. _Echinomma aculeatum_, Haeckel.

_Actinomma aculeatum_, Stöhr, 1880 Palæontogr., vol. xxvi. p. 92, Taf.
ii. fig. 11.

Cortical shell thick walled with regular circular pores, of the same
breadth as the bars; eight to ten on the radius. Radial proportion of the
three spheres = 1 : 2 : 8. Surface covered with short conical by-spines
about twenty strong main spines, three-sided pyramidal, one-third as long
as the shell diameter.

_Dimensions._--Diameter of the outer shell 0.16, middle 0.04, inner 0.02;
cortical pores and bars 0.003; length of the main spines 0.05, basal
breadth 0.013.

_Habitat._--Fossil in the Tertiary rocks of Sicily and Barbados; living in
the Atlantic, Station 332, depth 2200 fathoms.

4. _Echinomma diadema_, n. sp.

Cortical shell thick walled with regular circular pores, twice as broad as
the bars; sixteen to eighteen on the radius. Radial proportion of the three
spheres = 1 : 3 : 11. Surface covered with short bristle-shaped by-spines
one third as long as the twelve stout conical main spines, which reach in
length the radius, and lie opposite in pairs in six axes.

_Dimensions._--Diameter of the outer shell 0.22, middle 0.06, inner 0.02;
cortical pores 0.006, bars 0.003 length of the main spines 0.12, basal
breadth 0.02.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subgenus 2. _Echinommura_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different size or
form.

5. _Echinomma sphærechinus_, n. sp. (Pl. 29, fig. 2).

_Actinomma sphærechinus_, Haeckel, 1879, Atlas (pl. xxix. fig. 2).

Cortical shell thin walled, with irregular, roundish pores, twice to five
times as broad as the bars; five to seven on the radius. Both medullary
shells with regular, circular pores, twice as broad as the bars. Radial
proportion of the three spheres = 1 : 2 : 4. Surface covered with thirty to
fifty pyramidal main spines, as long as the diameter of the inner shell,
and with numerous small conical by-spines of half the length.

_Dimensions._--Diameter of the outer shell 0.11, middle 0.05, inner 0.025;
cortical pores 0.01 to 0.02, bars 0.004; length of the main spines 0.03,
basal breadth 0.005.

_Habitat._--North Atlantic, Station 353, surface.

6. _Echinomma trinacrium_, Haeckel.

_Actinomma trinacrium_, Haeckel, 1862, Monogr. d. Radiol., p. 441, Taf.
xxiv. figs. 6-8.

Cortical shell thin walled, with irregular, roundish pores, twice to three
times as broad as the bars. Both medullary shells with regular, circular
pores. Radial proportion of the three spheres {259}= 1 : 3 : 9. On the
surface about twenty three-sided pyramidal main spines, as long as the
radius, and numerous (forty to sixty) by-spines of half the length.
(Compare with this species _Actinomma trinacrium_, with which it is
connected by transitional forms.)

_Dimensions._--Diameter of the outer shell 0.09, middle 0.03, inner 0.01;
cortical pores 0.008 to 0.012, bars 0.004; length of the spines 0.02 to
0.05, basal breadth 0.01.

_Habitat._--Mediterranean (Messina).

7. _Echinomma toxopneustes_, n. sp. (Pl. 29, fig. 1).

Cortical shell thin walled, with large, irregular roundish, polygonally
framed pores, twice to four times as broad as the crested bars. Both
medullary shells of similar structure. Radial proportion of the three
spheres = 1 : 2.5 : 6. Numerous (thirty to fifty or more) thin radial beams
connecting them, prolonged outside into strong three-sided pyramidal
spines, shorter than the radius; each of the three wings with two teeth.
Between these, numerous smaller angular by-spines of one-quarter to
one-half the length.

_Dimensions._--Diameter of the outer shell 0.12, middle 0.05, inner 0.02;
cortical pores 0.01 to 0.03, bars 0.008; length of the main spines 0.05,
basal breadth 0.01.

_Habitat._--South-east Pacific (South of Juan Fernandez), Station 300,
surface.

Genus 103. _Pityomma_,[139] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Astrosphærida# with three concentric lattice-shells and
numerous branched radial spines.

The genus _Pityomma_ differs from its ancestral form, _Actinomma_, in the
ramification of its radial spines, and exhibits therefore the same relation
to it that _Elatomma_, among the Haliommida, bears to _Haliomma_.

1. _Pityomma scoparium_, n. sp.

Cortical shell thick walled, connected with both concentric medullary
shells by twenty thin radial beams, which are prolonged outside into twenty
large cylindrical radial spines; these are somewhat shorter than the shell
radius, simple in the proximal inner half, irregularly branched in the
outer half. All three spheres with regular, circular pores, twice to three
times as broad as the bars; radial proportion = 1 : 2 : 6. (Similar to
_Cladococcus scoparius_, Pl. 27, fig. 2, and _Elatomma scoparium_, but
differs from both in the double medullary shell.)

_Dimensions._--Diameter of the outer shell 0.15, middle 0.05, inner 0.025;
length of the spines 0.1.

_Habitat._--Central Pacific, Station 273, surface.

{260}2. _Pityomma piniferum_, n. sp.

Cortical shell thin walled, with irregular, roundish pores, connected with
both medullary shells by twenty stout radial beams, which are prolonged
outside into twenty large three-sided prismatic spines. These are longer
than the shell diameter and have three dentate edges, being branched like a
pine tree (with four to six ramified branches on each edge, decreasing in
size towards the distal end); similar to those of _Cladococcus abietinus_
(Pl. 27, fig. 3). Radial proportion of the three spheres = 1 : 2 : 8. Both
medullary shells with regular, circular pores, twice as broad as the bars.

_Dimensions._--Diameter of the outer shell 0.2, middle 0.05, inner 0.025;
length of the spines 0.25.

_Habitat._--South Pacific, Station 299 (Juan Fernandez), surface.

3. _Pityomma drymodes_, n. sp. (Pl. 29, fig. 9).

Cortical shell thin walled, connected with both medullary shells by very
numerous (one hundred to two hundred or more) very thin radial beams, which
are prolonged outside into thin branched radial spines, scarcely half as
long as the shell radius, each spine bearing six to nine simple branches.
Outer and middle shell uneven, with hill-shaped protuberances and valleys
between them; the connecting radial beams, which arise from the top of the
protuberances of the middle shell, are inserted into the deepest part of
the valleys of the outer shell. Radial proportion of the three spheres =
1 : 3 : 8. Outer medullary shell with small, irregular, roundish, or
polygonal pores, not much broader than the bars; innermost shell with
polygonal pores and very thin bars.

_Dimensions._--Diameter of the outer shell 0.28, middle 0.1, inner 0.033;
length of the spines 0.07, breadth 0.004.

_Habitat._--Central Pacific, Station 271, surface.

Subfamily CROMYOMMIDA,[140] Haeckel, 1881, Prodromus, pp. 449, 453.

_Definition._--#Astrosphærida# with four concentric spherical
lattice-shells.

Genus 104. _Cromyomma_,[141] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Astrosphærida# with four concentric lattice-spheres and
numerous simple radial spines of one kind.

The genus _Cromyomma_ is the common ancestral form of the Cromyommida, or
of those Astrosphærida in which the shell is composed of four concentric
spheres, connected by radial beams. Usually two of these are intracapsular
or medullary {261}shells, two extracapsular or cortical shells; the
distance between the former and the latter is greater than the distance
between either the two inner or the two outer shells. In some cases,
however (_e.g._, _Cromyomma zonaster_), the distance between all four
shells is equal.

Subgenus 1. _Cromyommetta_, Haeckel.

_Definition._--Pores of the outer cortical shell regular, of nearly equal
size and similar form.

1. _Cromyomma villosum_, n. sp. (Pl. 30, fig. 2).

Radial proportion of the four spheres = 1 : 2 : 4 : 5. Outer cortical shell
thick walled, densely covered with innumerable bristle-shaped radial
spines, half as long as the radius. Pores regular, circular, double-edged,
of the same breadth as the bars; ten to twelve on the radius. The thick
outer shell is so dark, that the outlines only of the other three shells
can be seen.

_Dimensions._--Diameter of the four shells--(A) outer cortical shell 0.2,
(B) inner cortical shell 0.16, (C) outer medullary shell 0.08, (D) inner
medullary shell 0.04; pores and bars of the outer shell 0.008; length of
the spines 0.04.

_Habitat._--Central Pacific, Station 267, depth 2700 fathoms.

2. _Cromyomma zonaster_, Haeckel.

_Acanthosphæra zonaster_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 285, Taf. ii. fig. 2.

Radial proportion of the four spheres = 2 : 3 : 4 : 5. Outer cortical shell
thick walled, densely covered with numerous bristle-shaped radial spines,
as long as the radius. Pores regular, circular, twice as broad as the bars;
eight to ten on the radius. The thick outer shell is so dark, that the
outlines only of the other three shells can be seen.

_Dimensions._--Diameter of the four shells--(A) 0.15, (B) 0.12, (C) 0.09,
(D) 0.06; pores of the outer shell 0.01, bars 0.005; length of the spines
0.08.

_Habitat._--Arctic Ocean, Greenland (depth 1000 fathoms), "Alert"
Expedition.

3. _Cromyomma perplexum_, Stöhr.

_Cromyomma perplexum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 95, Taf.
iii. fig. 5.

Radial proportion of the four spheres = 1 : 2 : 3 : 4. Outer cortical shell
thick walled, densely covered with short conical spines, as long as the
diameter of the pores. These are regular, circular, three times as broad as
the bars; six to eight on the radius. The thick outer shell is so dark,
that the outlines only of the three other shells can be seen.

_Dimensions._--Diameter of the four shells--(A) 0.17, (B) 0.13, (C) 0.087,
(D) 0.043; pores of the outer shell 0.015, bars 0.005; length of the spines
0.02.

_Habitat._--Fossil in the Tertiary rocks of Sicily (Grotte) Stöhr.

{262}Subgenus 2. _Cromyommura_, Haeckel.

_Definition._--Pores of the outer cortical shell irregular, of different
size and form.

4. _Cromyomma perspicuum_, n. sp. (Pl. 30, fig. 8).

_Cromyosphæra perspicua_, Haeckel, 1879, Atlas (pl. xxx. fig. 8).

Radial proportion of the four shells = 1 : 2.5 : 6 : 9. Outer cortical
shell very thin walled, with thread-like bars, and irregular, polygonal
pores; inner cortical shell with similar pores, but with thicker bars. Both
medullary shells with regular, circular pores (the outer three times as
broad as the inner). Numerous thin radial beams connect the latter with the
former; other very thin and numerous beams connect the two cortical shells,
and are prolonged outside into short bristle-shaped spines (often longer
than in the figure).

_Dimensions._--Diameter of the four shells--(A) 0.18, (B) 0.12, (C) 0.05,
(D) 0.02; pores of the outer shell 0.01 to 0.02; length of the spines 0.01
to 0.03.

_Habitat._--Central Pacific, Station 274, surface.

5. _Cromyomma quadruplex_, Haeckel.

_Cromyomma quadruplex_, Haeckel, 1862, Monogr. d. Radiol., p. 446.

_Haliomma quadruplex_, Ehrenberg, 1854, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 243.

Radial proportion of the four spheres = 1 : 3 : 7 : 10. Outer cortical
shell thin walled, irregularly covered with numerous angular spines, as
long as the radius. Pores of all four shells irregular, roundish,
increasing in size from the first to the fourth.

_Dimensions._--Diameter of the four shells--(A) 0.15, (B) 0.1, (C) 0.045,
(D) 0.015; pores of the outer shell 0.01 to 0.02; length of the spines
0.07.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

6. _Cromyomma macroporum_, Stöhr.

_Cromyomma macroporum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 95, Taf.
iii. fig. 4.

Radial proportion of the four spheres = 1 : 3 : 6 : 9. Outer cortical shell
thick walled, densely covered with short conical spines, as long as the
diameter of the pores; the latter are irregular, roundish, or polygonal,
larger and smaller alternating.

_Dimensions._--Diameter of the four shells--(A) 0.1, (B) 0.066, (C) 0.033,
(D) 0.011; pores of the outer shell 0.033 to 0.016, bars 0.003; length of
the spines 0.01.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Stöhr.

7. _Cromyomma circumtextum_, n. sp. (Pl. 30, fig. 4).

Radial proportion of the four spheres = 1 : 2 : 5 : 7. Outer cortical
shell very delicate and thin walled, with thread-like bars and large,
irregular, polygonal meshes (larger than the innermost shell); {263}inner
cortical shell also with irregular, polygonal pores, three to nine times as
broad as the thick bars. Both medullary shells with subregular, circular
pores (the outer three times as broad as the inner). Radial spines twelve
to twenty, three-sided prismatic, stout, piercing the three outer shells,
inserted on the innermost; their outer part pyramidal, half as long as the
radius.

_Dimensions._--Diameter of the four shells--(A) 0.22, (B) 0.15, (C) 0.07,
(D) 0.03; meshes of the outer shell 0.02 to 0.04; length of the outer
spines 0.05.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

8. _Cromyomma mucronatum_, n. sp. (Pl. 30, figs. 5, 5_a_).

Radial proportion of the three spheres = 1 : 2 : 3 : 6. Outer cortical
shell thin walled, with large, irregular, roundish pores and crested
three-sided bars; inner cortical shell with regular, circular, hexagonally
formed pores (four times as broad as the bars). Both medullary shells with
simple, small, regular, circular pores (fig. 5_a_). Twelve radial spines
three-sided prismatic, dagger-shaped, with three teeth, as long as the
diameter of the innermost shell.

_Dimensions._--Diameter of the four shells--(A) 0.24, (B) 0.12, (C) 0.08,
(D) 0.04; meshes of the outer shell 0.02 to 0.04; length of the spines
0.04.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

Genus 105. _Cromyechinus_,[142] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Astrosphærida# with four concentric lattice-spheres and
numerous simple spines of two different kinds; larger main spines and
smaller by-spines.

The genus _Cromyechinus_ differs from _Cromyomma_ in the same way as
_Echinomma_ from _Actinomma_; the radial spines being differentiated into
two different kinds; a larger number of small by-spines, and a smaller
number of large main spines.

1. _Cromyechinus icosacanthus_, n. sp. (Pl. 30, fig. 1).

Radial proportion of the three spheres = 1 : 2 : 6 : 8. Outer cortical
shell very delicate, with very small, regular, circular pores, covered with
very numerous bristle-shaped by-spines, half as long as the radius; inner
cortical shell with very large, irregular, polygonal pores, three to eight
times as broad as the bars. Both medullary shells with regular, circular
pores, connected with the former by twenty regularly disposed, very thin
bars; these are prolonged outside into twenty very stout, three-sided
prismatic main spines, as long as the radius, which arise from the inner
cortical shell, and at the distal end are cuspidate.

_Dimensions._--Diameter of the four shells--(A) 0.16, (B) 0.13, (C) 0.04,
(D) 0.02; pores of the outer cortical shell 0.003, of the inner 0.03, bars
0.003; length of the spines 0.08.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

{264}2. _Cromyechinus dodecacanthus_, n. sp. (Pl. 30, figs. 3, 3_a_).

Radial proportion of the four spheres = 1 : 3 : 10 : 12. Outer cortical
shell very delicate, with very small, regular, circular pores, covered with
numerous short bristle-shaped by-spines, scarcely one-fourth as long as the
radius; inner cortical shell with very large, irregular, polygonal pores,
twice to six times as broad as the bars. Both medullary shells with very
small, regular, circular pores, connected with the former by twelve
regularly disposed, thin radial beams; these are prolonged outside into
twelve strong, three-sided pyramidal main spines, one-third as long as the
shell diameter. (Differs from the foregoing only in the number and form of
the radial spines.)

_Dimensions._--Diameter of the four shells--(A) 0.12, (B) 0.1, (C) 0.03,
(D) 0.01; pores of the outer cortical shell 0.002, of the inner 0.02, bars
0.002; length of the spines 0.04, basal breadth 0.01.

_Habitat._--South Atlantic, Station 325, surface.

3. _Cromyechinus polyacanthus_, n. sp.

Radial proportion of the four spheres = 1 : 2 : 8 : 10. Outer cortical
shell of the same structure as the inner, with irregular, roundish pores,
twice to six times as broad as the bars. Both medullary shells with small,
regular, circular pores, twice as broad as the bars. Connecting radial
beams forty to sixty, thin, cylindrical, prolonged outside into forty to
sixty strong conical main spines, about as long as the radius. Between
these, numerous thin bristle-shaped by-spines, nearly of the same length.

_Dimensions._--Diameter of the four shells--(A) 0.25, (B) 0.2, (C) 0.05,
(D) 0.025; pores of both cortical shells 0.01 to 0.03; length of the spines
0.12.

_Habitat._--North Pacific, Station 237, surface.

Genus 106. _Cromyodrymus_,[143] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Astrosphærida# with four concentric lattice-spheres and
numerous branched radial spines.

The genus _Cromyodrymus_ differs from its ancestral form, _Cromyomma_, in
the ramification of the radial spines, and exhibits therefore the same
relation to it that _Pityomma_ among the Actinommida bears to _Actinomma_.

1. _Cromyodrymus quadricuspis_, n. sp. (Pl. 30, figs. 7, 7_a_).

Radial proportion of the four shells = 1 : 3 : 6 : 12. All four shells with
regular, circular pores, about twice as broad as the bars. Radial
proportion of the pores in the four shells = 2 : 4 : 2 : 1. Radial spines
fifty to sixty, three-sided prismatic, half as long as the radius, with
three recurved teeth, each spine in this way bearing four points.

{265}_Dimensions._--Diameter of the four shells--(A) 0.16, (B) 0.08, (C)
0.04, (D) 0.013; length of the spines 0.03 to 0.04, breadth 0.007.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Cromyodrymus abietinus_, n. sp. (Pl. 30, fig. 6).

Radial proportion of the four shells = 1 : 2 : 5 : 11. All four shells with
irregular, roundish pores, twice to four times as broad as the bars. Radial
proportion of the pores in the four shells = 9 : 3 : 2 : 1. Radial spines
eighty to one hundred and twenty, about as long as the radius, branched
like a pine tree, with six to twelve ramified branches.

_Dimensions._--Diameter of the four shells--(A) 0.22, (B) 0.1, (C) 0.04,
(D) 0.02; length of the spines 0.12, breadth 0.006.

_Habitat._--South-east Pacific (Juan Fernandez), Station 299, surface.

Subfamily CARYOMMIDA,[144] Haeckel.

_Arachnosphærida_, Haeckel, 1862, Monogr. d. Radiol., p. 354; Prodromus,
1881, p. 454.

_Definition._--#Astrosphærida# with five or more concentric spherical
lattice-shells.

Genus 107. _Caryomma_,[145] n. gen.

_Definition._--#Arachnosphærida# with five to six or more concentric
spherical lattice-shells; two inner (intracapsular) medullary and three or
more outer (extracapsular) cortical shells, composed of ordinary
lattice-work; distance between the former and the latter greater than
between any other two shells.

The genus _Caryomma_ may be regarded as a _Cromyomma_, in which the number
of the cortical shells is increased to three, four, or more. These lie
outside the central capsule, whilst two medullary shells lie within it, and
are connected with the former by numerous radial spines piercing the wall
of the capsule. The ordinary lattice-work is not arachnoidal, as in the
three following genera.

1. _Caryomma regulare_, n. sp.

Radial proportion of the five shells = 1 : 2 : 6 : 8 : 10. All five shells
with regular, circular pores, twice to three times as broad as the bars,
and gradually increasing in size from the innermost to the outermost shell.
Radial beams connecting them twenty, prolonged on the surface into twenty
stout, three-sided pyramidal, radial spines, half as long as the radius,
regularly disposed.

_Dimensions._--Diameter of the five shells--(A) 0.25, (B) 0.2, (C) 0.15,
(D) 0.05, (E) 0.025.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

{266}2. _Caryomma irregulare_, n. sp.

Radial proportion of the six shells = 1 : 2.5 : 8 : 10.5 : 13 : 15. Both
inner (medullary) shells with regular, circular pores, the other four
(cortical) shells with irregular, roundish pores, gradually increasing in
size from the innermost to the outermost shell. Radial spines sixty to
eighty, pyramidal, irregularly disposed.

_Dimensions._--Diameter of the six shells--(A) 0.3, (B) 0.26, (C) 0.21, (D)
0.16, (E) 0.05, (F) 0.02.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

Genus 108. _Arachnopila_,[146] n. gen.

_Definition._--#Astrosphærida# with five to ten or more cortical,
concentric, polyhedral, or spherical lattice-shells, composed of a very
thin cobweb-like network; innermost shell with hexagonal (regular) or
polygonal (irregular) meshes; other shells with simple triangular meshes,
without diagonal threads between them.

The genus _Arachnopila_, together with the two following genera, forms the
peculiar small group of large Arachnosphærida, separated from the true
Caryommida by the totally different structure and disposition of the
numerous concentric shells; the former exhibits a similar relation to the
latter that the Diplosphærida bears to the Elatommida among the dispherical
Haliommida. The concentric shells (five to ten or more) lie outside the
central capsule, and are composed of very delicate, cobweb-like threads.
From the innermost shell arise numerous, three-sided prismatic, very long
spines, from which at equal regular distances arise lateral branches (three
pairs from each spine, and one pair from each corner). In _Arachnopila_
these threads pass directly from one spine to the other, and form simple,
large, triangular meshes between them. The concentric shells are not
connected by interwoven diagonal threads.

1. _Arachnopila hexagonella_, n. sp.

Innermost shell with regular hexagonal pores; its diameter twice as long as
the equal distance between every two concentric shells. Radial spines
twenty to forty, each with twenty to twenty-four verticils.

_Dimensions._--Diameter of the innermost shell 0.1; distance between the
concentric shells 0.05.

_Habitat._--Central Pacific, Station 271, surface.

2. _Arachnopila polygonella_, n. sp.

Innermost shell with irregular, polygonal pores; its diameter fully as long
as the equal distance between every two concentric shells. Radial spines
sixty to eighty, each with twelve to sixteen verticils.

_Dimensions._--Diameter of the innermost shell 0.04; distance between the
concentric shells 0.04.

_Habitat._--Central Pacific, Station 266, surface.

{267}Genus 109. _Arachnopegma_,[147] Haeckel, 1881, Prodromus, p. 454.

_Definition._--#Astrosphærida# with five to ten or more cortical,
concentric, polyhedral, or spherical lattice-shells, composed of a very
thin, cobweb-like network; innermost shell with hexagonal or polygonal
meshes; other shells with simple triangular meshes, connected to one
another by diagonal threads between them.

The genus _Arachnopegma_ differs from its ancestral form _Arachnopila_, in
the possession of peculiar diagonal threads, which connect the verticils or
nodal-points of every two neighbouring radial spines in two different
neighbouring concentric shells. In this case, therefore, not only do
triangular meshes lie in the spherical faces of the concentric spheres, but
also between them, in numerous oblique diagonal planes; a very rare and
remarkable structure, and forming a transition to spongy shells.

1. _Arachnopegma verticillatum_, n. sp.

Innermost shell with regular, hexagonal meshes; its diameter is the same as
the equal distance between each of the two shells. Threads of the network
smooth. Radial spines twenty to forty, each with fifteen to twenty
verticils.

_Dimensions._--Diameter of the innermost shell 0.04, distance between the
concentric shells 0.03 to 0.04.

_Habitat._--Central Pacific, Station 266, surface.

2. _Arachnopegma longispinum_, n. sp.

Innermost shell with regular, hexagonal meshes; its diameter half as large
as the equal distance between every two shells. Threads of the network
dentated or with small knots. Radial spines fifty to sixty, each with
twenty to twenty-five verticils.

_Dimensions._--Diameter of the innermost shell 0.02; distance between the
concentric shells 0.04 to 0.05.

_Habitat._--Central Pacific, Station 272, surface.

3. _Arachnopegma increscens_, n. sp.

Innermost shell with regular, hexagonal meshes; its diameter quite as large
as the distance between it and the second shell; only half as large as the
distance between the fourth and fifth shells. Forty to fifty radial spines,
each with fifteen to twenty verticils.

_Dimensions._--Diameter of the innermost shell (A) 0.025; distance between
the following shells--A, B = 0.025, B, C = 0.03, C, D = 0.04, D, E = 0.048,
E, F = 0.056, F, G = 0.064, &c.

_Habitat._--Central Pacific, Station 274, surface.

{268}Genus 110. _Arachnosphæra_,[148] Haeckel, 1862, Monogr. d. Radiol., p.
355.

The genus _Arachnosphæra_ (accurately described by me in 1862, _loc. cit._)
differs from the two preceding genera in the irregular form of the large
meshes in all shells (except often the innermost). This is caused by the
ramification of the thin threads, which on each shell connect the
neighbouring spines. In _Arachnopila_ and _Arachnopegma_ the threads pass
directly and undivided from each spine to the neighbouring spine (three
pairs from the three edges), and therefore all meshes are triangular. In
_Arachnosphæra_ they become polygonal by irregular ramification of the
threads. There are here no diagonal threads.

Subgenus 1. _Arachnosphærella_, Haeckel.

_Definition._--Pores of the innermost shell regular, hexagonal.

1. _Arachnosphæra oligacantha_, Haeckel.

_Arachnosphæra oligacantha_, Haeckel, 1862, Monogr. d. Radiol., p. 356,
Taf. x. fig. 2, Taf. xi. fig. 3.

Innermost shell with regular, hexagonal meshes; its diameter three times as
long as the equal distances between every two concentric shells. Fifteen to
twenty radial spines scattered at wide intervals, each with six to eight
verticils.

_Dimensions._--Diameter of the innermost shell 0.12; distance between the
concentric shells 0.04 to 0.05.

_Habitat._--Mediterranean (Messina), surface, Haeckel.

2. _Arachnosphæra myriacantha_, Haeckel.

_Arachnosphæra myriacantha_, Haeckel, 1862, Monogr. d. Radiol., p. 357,
Taf. x. fig. 3, Taf. xi. fig. 4.

Innermost shell with regular, hexagonal meshes; its diameter twice as long
as the equal distances between every two concentric shells. At each
nodal-point occurs one radial spine with six to eight verticils (spines in
all one hundred to one hundred and twenty or more).

_Dimensions._--Diameter of the innermost shell 0.1; distance between the
concentric shells 0.04 to 0.05.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

{269}3. _Arachnosphæra dolichacantha_, n. sp.

Innermost shell with regular, hexagonal meshes, very small; its diameter
only as long as the equal distances between every two concentric shells. At
each nodal-point occurs one radial spine with twelve to sixteen verticils
(altogether thirty to forty spines).

_Dimensions._--Diameter of the innermost shell 0.05; distance between the
concentric shells 0.04 to 0.05.

_Habitat._--Central Pacific, Station 265 to 274, surface.

4. _Arachnosphæra increscens_, n. sp.

Innermost shell with regular, hexagonal meshes; its diameter three times as
long as the distance between it and the second shell, quite as long as the
distance between the fifth and sixth shells; the distances between the
concentric shells gradually increasing from the centre. Radial spines about
fifty to sixty, each with six to eight verticils.

_Dimensions._--Diameter of the innermost shell (A) 0.75; distances between
the following shells--A, B = 0.025, B, C = 0.037, C, D = 0.05, D, E =
0.062, E, F = 0.075, &c.

_Habitat._--Central Pacific, Station 271, surface.

Subgenus 2. _Arachnosphæromma_, Haeckel.

_Definition._--Pores of the innermost shell irregular, polygonal.

5. _Arachnosphæra tenuissima_, n. sp.

Innermost shell with irregular, polygonal meshes; its diameter twice as
long as the equal distance between every two concentric shells. Forty to
fifty radial spines, each with twelve to sixteen verticils.

_Dimensions._--Diameter of the innermost shell 0.1, distance between the
concentric shells 0.04 to 0.05.

_Habitat._--South Atlantic, Station 332, surface.

6. _Arachnosphæra velaris_, n. sp.

Innermost shell with irregular, polygonal meshes; its diameter twice as
long as the distance between it and the second shell, quite as long as the
distance between the third and fourth shells; the distance between the
concentric shells gradually increasing from the centre. Radial spines
twenty to thirty, each with ten to twelve verticils.

_Dimensions._--Diameter of the innermost shell (A) 0.5; distances of the
following shells--A, B = 0.025, B, C = 0.037, C, D = 0.05, D, E = 0.062, E,
F = 0.075.

_Habitat._--Tropical Atlantic, Station 347, surface.

{270}Subfamily SPONGIOMMIDA,[149] Haeckel.

_Definition._--#Astrosphærida# with spongy spherical or polyhedral shell
(with or without enclosed concentric lattice-shells).

Genus 111. _Spongiomma_,[150] n. gen.

_Definition._--#Astrosphærida# with solid spongy sphere, with numerous
simple radial spines, but without latticed medullary shells.

The genus _Spongiomma_ differs from its ancestral form, _Styptosphæra_, in
the development of simple radial spines on the surface of the solid sphere,
the entire mass of which is composed of an irregular, spongy wicker-work.

Subgenus 1. _Spongiommella_, Haeckel.

_Definition._--Radial spines on the surface of the spongy sphere all of the
same shape.

1. _Spongiomma radiatum_, n. sp.

Spongy framework of the solid sphere of the same structure throughout, with
small, irregular, polyhedral meshes and very thin thread-like bars. From
the surface arise very numerous (one hundred and twenty to one hundred and
sixty or more) straight, bristle-shaped radial spines, as long as the
radius of the sphere (counting from the middle part of it).

_Dimensions._--Diameter of the spongy sphere 0.2; length of the radial
spines 0.1.

_Habitat._--Central Pacific, Station 272, surface.

2. _Spongiomma denticulatum_, n. sp.

Spongy framework of the solid sphere of the same structure throughout, with
small, almost equal meshes, four to eight times as broad as the thin,
elegantly denticulated bars; from the surface arise very numerous (two
hundred to three hundred or more) curved, radial spines, as long as the
radius, and of the same form as the bars.

_Dimensions._--Diameter of the sphere 0.25; length of the spines 0.12.

_Habitat._--Central Pacific, Station 274, surface.

3. _Spongiomma spathillatum_, n. sp.

Spongy framework in the central part of the sphere much denser and darker,
and with smaller meshes than in the peripheral part in which are very thin
bars. Entire surface covered with {271}innumerable short, bristle-shaped
radial spines, only one-eighth as long as the radius, of the same elegant
form as in _Octodendron spathillatum_ (Pl. 18, figs. 2, 4); each spine
developed in a zig-zag fashion, with very small beards, with a delicate
spathillum (or coronal of beard spines) at the distal end.

_Dimensions._--Diameter of the spheres 0.16; length of the spines 0.01.

_Habitat._--South Pacific, Station 295, surface.

4. _Spongiomma clavatum_, n. sp.

Spongy framework in the central part of the sphere much denser and darker
than in the peripheral part. On the surface are sixty to eighty stout,
club-shaped radial spines, as long as the radius of the sphere, in the
proximal half three-sided prismatic, with three dentated edges; they begin
at the middle of the radius (where the denser inner framework changes into
the looser outer) and are very thin at first but increase slowly in
thickness towards the truncated distal end. (Similar to _Centrocubus
rhopalophorus_, Pl. 18, fig. 1, but without the cubical medullary shell.)

_Dimensions._--Diameter of the sphere 0.4; length of the spines 0.2.

_Habitat._--North Pacific, Station 241, surface.

Subgenus 2. _Spongiommura_, Haeckel.

_Definition._--Radial spines on the surface of the spongy sphere of two
different kinds; large main spines and small by-spines.

5. _Spongiomma helioides_, n. sp.

Spongy framework of the sphere everywhere of the same structure, with
almost equal meshes, ten to twelve times as broad as the bars. Sixteen to
twenty radial main spines, longer than the shell diameter, three-sided
prismatic, with three serrated edges, beginning about the middle of the
radius and increasing in thickness to the truncated distal end. Between
them occur numerous thin, bent, bristle-shaped by-spines. (Very similar to
_Spongosphæra helioides_, Monogr. d. Radiol., Taf. xii. figs. 11-13, but
without medullary shells.)

_Dimensions._--Diameter of the sphere 0.3; length of the main spines 0.4,
of the by-spines 0.01.

_Habitat._--Tropical Atlantic, Station 352, surface.

6. _Spongiomma multiaculeum_, Haeckel.

_Spongechinus multiaculeatus_, Dunikowski, 1882, Denkschr. d. k. Akad. d.
Wiss. Wien, Bd. xlv. p. 29, Taf. v. figs. 60-63.

Spongy framework of the sphere very compact, with small meshes, scarcely
broader than the bars; four to eight large main spines, three-sided
pyramidal, longer than the shell radius; numerous (thirty to forty) thin
by-spines, scarcely half as long.

{272}_Dimensions._--Diameter of the sphere 0.16; length of the main spines
0.11, of the by-spines 0.03.

_Habitat._--Fossil in the Alpine Jura, Schafberg near Salzburg
(Dunikowski).

7. _Spongiomma asteroides_, n. sp.

Spongy framework in the central part of the sphere much denser and darker
than in the peripheral part. Sixty to eighty cylindro-conical main spines,
as long as the radius, between them are numerous straight, bristle-shaped
by-spines half that length.

_Dimensions._--Diameter of the sphere 0.28; length of the main spines 0.16,
of the by-spines 0.08.

_Habitat._--South Atlantic, Station 325, surface.

Genus 112. _Spongodrymus_,[151] Haeckel, 1881, Prodromus, p. 456.

_Definition._--#Astrosphærida# with solid spongy sphere, without latticed
medullary shell, with numerous branched radial spines.

The genus _Spongodrymus_ differs from the preceding _Spongiomma_ in the
ramification of the numerous radial spines, covering the surface of the
solid spongy sphere.

1. _Spongodrymus elaphococcus_, n. sp. (Pl. 18, fig. 9).

Spongy framework of the solid sphere equal throughout, on the surface
arising in the form of very numerous (sixty to eighty or more) spongy cones
with rather rhomboidal meshes. Each cone is prolonged into a very thin,
irregularly curved radial spine, which is twice as long as the shell
diameter, simple in the proximal half, branched like a tree in the distal
half; each arborescent spine has from sixteen to thirty-two terminal
branches, which fall in one spherical face. The branches of neighbouring
spines are partly connected by anastomoses, so that they begin to form an
outer spherical shell (cortical shell) with a looser spongy framework.

_Dimensions._--Diameter of the whole spherical skeleton (sphere formed by
the distal ends of the dichotomous branches) 0.9, of the inner solid spongy
sphere 0.02.

_Habitat._--Tropical Atlantic, Station 349, surface.

2. _Spongodrymus abietinus_, n. sp.

Spongy framework denser and darker in the central part of the sphere than
in the peripheral part, with rather coarse meshes and thick bars. From the
surface arise very numerous (sixty to eighty) stout, radial branches, as
long as the shell radius, branched like a pine tree (with six to twelve
ramified branches). Similar in structure to _Cromyodrymus abietinus_ (Pl.
30, fig. 6), but with a quite irregular spongy texture in the central
sphere.

_Dimensions._--Diameter of the sphere 0.3; length of the spines 0.015.

_Habitat._--Central Pacific, Station 272, surface.

{273}Genus 113. _Spongechinus_,[152] Haeckel, 1881, Prodromus, p. 456.

_Definition._--#Astrosphærida# with hollow, spongy sphere, without latticed
medullary shell in the central cavity, and with numerous simple radial
spines.

The genus _Spongechinus_ differs from its ancestral form, _Plegmosphæra_,
in the development of numerous radial spines on the surface of the spongy
sphere, within which is enclosed a large spherical central cavity.

1. _Spongechinus setosus_, n. sp.

Spongy sphere three times as broad as its inner cavity, with a very
delicate, equal framework. Entire surface covered with short, straight,
bristle-shaped radial spines, about half as long as the radius.

_Dimensions._--Diameter of the sphere 0.2, of its inner cavity 0.07; length
of the spines 0.05.

_Habitat._--North Atlantic, Azores, surface.

2. _Spongechinus serrulatus_, n. sp.

Spongy sphere twice as broad as its inner cavity, with a delicate, equal
framework. Entire surface covered with short, curved, radial spines, which
are elegantly denticulated, and as long as the shell radius.

_Dimensions._--Diameter of the sphere 0.3, of its cavity 0.15; length of
the spines 0.15.

_Habitat._--Central Pacific, Station 274, surface.

3. _Spongechinus cavus_, n. sp.

Spongy sphere only one-fourth broader than the large inner cavity, its
spongy wall being only one-fourth as thick as the radius, composed of three
to four strata of irregular, small meshes. Surface covered with short,
bristle-shaped, curved spines, one-third as long as the radius.

_Dimensions._--Diameter of the sphere 0.5, of its inner cavity 0.4; length
of the spines 0.08.

_Habitat._--North Atlantic, Station 354, surface.

Genus 114. _Spongothamnus_,[153] n. gen.

_Definition._--#Astrosphærida# with hollow, spongy sphere, without latticed
medullary shell in the central cavity, and with numerous branched radial
spines.

The genus _Spongothamnus_ differs from the preceding _Spongechinus_ in the
ramification of the numerous radial spines, covering the surface of the
hollow spongy sphere; {274}it therefore exhibits the same relation to the
latter that _Spongodrymus_ bears to _Spongiomma._

1. _Spongothamnus furcatus_, n. sp.

Spongy sphere twice as broad as its inner cavity, with very delicate bars,
and irregular, dense framework. From the surface arise one hundred and
fifty to one hundred and eighty thin, forked, bristle-shaped spines, half
as long as the radius; both fork branches one-third as long as the basal or
simple part.

_Dimensions._--Diameter of the sphere 0.3, of its inner cavity 0.15; length
of the spines 0.08.

_Habitat._--North Pacific, Station 241, surface.

2. _Spongothamnus scoparius_, n. sp.

Spongy sphere four times as broad as its inner cavity, with thick bars and
rather loose framework. From the surface arise sixty to eighty broom-shaped
radial spines, as long as the radius, each in the basal half simple, in the
distal half with six to twelve irregularly ramified branches (similar to
the spines of _Cromyodrymus abietinus_, Pl. 30, fig. 6).

_Dimensions._--Diameter of the sphere 0.4, of the inner cavity 0.1; length
of the spines 0.2.

_Habitat._--Central Pacific, Station 271, surface.

Genus 115. _Spongopila_,[154] Haeckel, 1881, Prodromus, p. 456.

_Definition._--#Astrosphærida# with a single, spherical, latticed medullary
shell, immediately enveloped by the spongy framework of the cortical shell;
on the surface of the latter are numerous radial spines.

The genus _Spongopila_ is a _Spongoplegma_ with radial spines. On the other
hand it may be derived either from _Elaphococcus_, by communication of the
branched spines, or from _Arachnosphæra_, by development of spongy branches
between the concentric spheres.

1. _Spongopila dichotoma_, n. sp.

Medullary shell with regular, hexagonal meshes, six times as broad as the
bars. From each nodal-point (between every three meshes) arises a
bristle-shaped radial spine, which is dichotomously branched. By
communication of the neighbouring branches the loose spongy framework of
the spherical cortical shell is formed, which is four times as broad as the
medullary shell. On the surface occur very numerous bristle-shaped radial
spines, as long as the diameter of the medullary shell. (May be derived
from _Elaphococcus_.)

_Dimensions._--Diameter of the spongy cortical shell 0.3, of the medullary
shell 0.08.

_Habitat._--Tropical Atlantic, Station 347, surface.

{275}2. _Spongopila verticillata_, n. sp.

Medullary shell with regular, hexagonal meshes, four times as broad as the
bars. From its surface arise forty to sixty, three-sided prismatic radial
spines, bearing eight to ten verticils of lateral branches, each verticil
with six forked branches (two from each edge). By irregular ramification of
these branches, and communication in all directions, the loose spongy
framework of the cortical shell originates, which is six times as broad as
the medullary shell. The free distal ends of the spines are as long as the
shell radius. (May be derived from _Arachnosphæra_.)

_Dimensions._--Diameter of the spongy cortical shell 0.6, of the medullary
shell 0.1.

_Habitat._--Tropical Pacific, Station 200, surface.

Genus 116. _Rhizoplegma_,[155] Haeckel, 1881, Prodromus, p. 456.

_Definition._--#Astrosphærida# with a single, spherical, latticed medullary
shell, which is not armed with by-spines, but connected by stout radial
main spines with the spongy cortical shell.

The genus _Rhizoplegma_ is immediately allied to the foregoing
_Spongopila_, but differs from it in the large interval separating both
shells. In this interval lies the wall of the central capsule, which is
only pierced by the radial spines connecting both shells.

Subgenus 1. _Rhizoplegmarium_, Haeckel.

_Definition._--No free lateral branches of the radial spines between the
two shells.

1. _Rhizoplegma polyacanthum_, n. sp.

Spongy cortical shell with a very fine dense framework and nearly smooth
surface (without superficial by-spines). Its inner cavity is four times as
broad as the medullary shell, which exhibits regular, hexagonal meshes.
Radial spines sixty to eighty, three-sided prismatic, with three smooth
edges, without lateral branches between both shells; their free outer part
as long as the shell radius.

_Dimensions._--Diameter of the spongy cortical shell 0.3, of its inner
cavity 0.2, of the medullary shell 0.05.

_Habitat._--Central Pacific, Station 274, surface.

2. _Rhizoplegma spirale_, n. sp.

Spongy cortical shell with a very delicate loose framework and bristly
surface. Its inner cavity three times as broad as the medullary shell,
which exhibits regular, hexagonal meshes. {276}Radial spines thirty to
forty, prismatic, with three smooth, spirally contorted edges, without
lateral branches between the two shells; their free outer part half as long
as the shell radius.

_Dimensions._--Diameter of the spongy cortical shell 0.2, of its inner
cavity 0.12, of the medullary shell 0.04.

_Habitat._--Central Pacific, Station 265, surface.

3. _Rhizoplegma trigonacantha_, n. sp.

Spongy cortical shell with a rather loose framework and coarse bars, with
nearly smooth surface (without by-spines). Its inner cavity twice as broad
as the medullary shell, which exhibits irregular, roundish pores. Radial
spines forty to sixty, prismatic, with three smooth straight edges, without
free branches between the two shells; their outer pyramidal part only
one-third of the shell radius. (Very similar to the common _Rhizosphæra
trigonacantha_, but with simple medullary shell.)

_Dimensions._--Diameter of the spongy shell 0.2, of its inner cavity 0.16,
of the medullary shell 0.08.

_Habitat._--North Atlantic, Station 353, surface.

Subgenus 2. _Rhizoplegmidium_, Haeckel.

_Definition._--Between the two shells free lateral branches arise from the
three edges of the radial spines.

4. _Rhizoplegma radicatum_, n. sp. (Pl. 15, figs. 9, 9_a_).

Spongy cortical shell with a very loose framework; on the surface are
innumerable thin, forked, or repeatedly dichotomous by-spines. Its inner
cavity three times as broad as the medullary shell, which exhibits regular,
hexagonal meshes (fig. 9_a_). Radial spines twelve, prismatic, with
straight dentated edges, their outer pointed part as long as the shell
radius. Each spine between the two shells has a verticil of three forked
lateral branches (fig. 9_a_). The central capsule completely distends the
medullary shell, and forces out through each mesh a hernia-shaped process
(fig. 9).

_Dimensions._--Diameter of the spongy shell 0.5, of its inner cavity 0.25,
of the medullary shell 0.08.

_Habitat._--Central Pacific, Station 271, surface.

5. _Rhizoplegma lychnosphæra_, n. sp. (Pl. 11, fig. 5).

_Lychnosphæra rhizoplegma_, Haeckel, 1879, Atlas (pl. xi. fig. 5).

Spongy cortical shell with a very loose framework, composed of long thin
beams as in _Lychnosphæra regina_ (Pl. 11, figs. 1-4). Surface covered with
short bristles. Its inner cavity six times as broad as the medullary shell,
which exhibits regular, hexagonal meshes. Radial spines twelve, prismatic,
with three smooth edges; their outer pointed part half as long as the shell
radius. Each spine has three verticils of three forked branches; the first
verticil is free {277}between the two shells, while the two following
verticils, by communication of their ramules, form the irregular framework.
Central capsule with many herniæ, forced out through the meshes of the
medullary shell (fig. 5).

_Dimensions._--Diameter of the spongy shell 0.7, of its inner cavity 0.45,
of the medullary shell 0.07.

_Habitat._--South Pacific, Station 284, surface.

Genus 117. _Lychnosphæra_,[156] Haeckel, 1881, Prodromus, p. 453.

_Definition._--#Astrosphærida# with a single, spherical, latticed medullary
shell, which is armed with free radial by-spines, and connected by stout
radial main spines with the spongy cortical shell.

The genus _Lychnosphæra_, known only by one single, large, and very
remarkable species, mainly differs from the foregoing in the development of
free radial by-spines on the surface of the medullary shell, and in the
free interval between it and the cortical shell; but beyond this the loose
spongy framework of the latter exhibits a very remarkable structure,
figured in Pl. 11.

1. _Lychnosphæra regina_ (Pl. 11, figs. 1-4).

Medullary shell (fig. 3) with regular, circular, hexagonally framed pores,
twice as broad as the bars; from each hexagon-corner arises a radial,
bristle-shaped by-spine, as long as the diameter. Twelve radial main spines
each as broad as one of the meshes, three-sided prismatic, six to eight
times as long as the medullary shell. From their three leaf-shaped (often
somewhat denticulated or spirally contorted) edges arise four verticils of
lateral branches, each composed of three forked, thin branches. The forked
branches of the first verticil end free between the two shells (figs. 2,
3), while the two following verticils are ramified, and, by anastomosis of
their branches, compose the loose spongy framework of the cortical shell.
On the surface of the latter arise numerous radial (zig-zag-shaped)
by-spines. The fourth verticil is terminal, with three shorter, thicker,
dentated, simple branches, which constitute, together with the distal apex
of the spine itself, a bunch of four terminal spines. The large central
capsule completely distends the medullary shell, and forces out by its
pores numerous club-shaped herniæ (fig. 1).

_Dimensions._--Diameter of the spongy cortical shell 0.6, of its inner
cavity 0.4, of the central capsule 0.22, of the medullary shell 0.06;
length of the radial spines 0.4, breadth 0.01.

_Habitat._--Central Pacific, Station 271, surface.

Genus 118. _Centrocubus_,[157] n. gen.

_Definition._--#Astrosphærida# with a single, cubical medullary shell,
immediately surrounded by the spongy framework of the cortical shell; from
the eight corners {278}of the central cube arise eight primary radial
spines, and often others from the framework between them.

The genus _Centrocubus_ and the following closely allied _Octodendron_ may
represent a peculiar small group of Spongiommida, remarkable for the
regular, cubical form of the medullary shell, which is composed of twelve
thin rods, corresponding to the twelve edges of a mathematical cube; from
the eight corners invariably arise eight primary radial spines, the
branches of which form the spongy cortical shell.

1. _Centrocubus octostylus_, n. sp.

Radial spines eight, arising from the eight corners of the cubiform,
regular, medullary shell, gradually increasing in thickness towards the
club-shaped distal end, which is five to six times as broad as the central
end. From the three denticulate edges of each spine arise six to eight
lateral branches, which ramify irregularly and form by their anastomosis
the spongy framework which is of nearly similar structure throughout, and
with large loose meshes. The free distal part of each spine is half as long
as the enclosed part.

_Dimensions._--Diameter of the sphere 0.6, of the central cube 0.02; length
of the spines 0.15, distal thickness 0.02.

_Habitat._--Central Pacific, Station 271, surface.

2. _Centrocubus cladostylus_, n. sp. (Pl. 18, fig. 1).

Radial spines thirty-two, club-shaped, at the distal end eight to ten times
as broad as at the basal end. Eight primary spines arise from the eight
corners of the regular, cubiform medullary shell, and from these, in the
form of lateral branches, twenty-four secondary spines arise with concavely
curved bases (three from the three denticulate edges of each spine, at
nearly equal distances from the centre). The free distal end of each of the
thirty-two spines is of the same shape, about half as long as the radius of
the spongy sphere; framework much looser in the outer than in the inner
part.

_Dimensions._--Diameter of the sphere 0.8, of the central cube 0.02; length
of the spines 0.2, distal thickness 0.02.

_Habitat._--North Pacific, Station 256, surface.

3. _Centrocubus polystylus_, n. sp.

Radial spines sixty to eighty, club-shaped, four to six times as broad at
the distal as at the basal end. Eight primary spines arise from the eight
corners of the regular, cubiform medullary shell, the remainder either
springing as lateral branches from the three denticulate corners of the
former, or arising within the spongy framework, which is much denser and
darker in the central than in the peripheral part. The free distal end of
each spine is one-third as long as the radius.

{279}_Dimensions._--Diameter of the sphere 0.9, of the central cube 0.02;
length of the spines 0.15, distal thickness 0.02.

_Habitat._--South Pacific, Station 295, surface.

Genus 119. _Octodendron_,[158] n. gen.

_Definition._--#Astrosphærida# with a single, cubical medullary shell, and
eight primary radial spines arising from its eight corners; these are
connected at equal distances by a latticed, spherical, cortical shell, from
which the spongy framework directly springs; often from the latter
secondary radial spines arise.

The genus _Octodendron_ has the same regular, cubical medullary shell as
the preceding _Centrocubus_, but differs from it in the wide interval
separating the medullary shell from the inner spherical face of the spongy
cortical shell, the two being connected only by eight radial beams, arising
from the eight corners of the cube.

Subgenus 1. _Octodendridium_, Haeckel.

_Definition._--Only eight primary radial spines, arising from the eight
corners of the central cube.

1. _Octodendron cubocentron_, n. sp. (Pl. 18, fig. 3).

Radial spines eight, club-shaped, with three denticulate, straight edges,
twice as long as the diameter of the central cavity of the cubical spongy
shell; the inner face of the latter (or the "inner cortical shell")
exhibits the form of a large cube, the spongy sides of which are parallel
with the simple square sides of the central cube. The thin eight radial
beams, connecting the corresponding corners of both cubes, are not thicker
than the edge-bars of the central cube, whilst their outer prolongations
are much thicker, with four to six verticils of lateral branches. Surface
of the thin walled, loose, spongy shell armed with short simple thorns.

_Dimensions._--Diameter of the spongy shell 0.2, of its inner cavity 0.1,
of the central cube 0.02; length of the spines 0.2 (from the centre 0.3).

_Habitat._--Central Pacific, Station 266, surface.

2. _Octodendron spirale_, n. sp.

Radial spines eight, three-sided prismatic, three times as long as the
diameter of the inner shell-cavity; their three edges denticulated and
spirally contorted around the spine axis. Thickness of the loose spongy
shell-wall equal to the radius of the inner cavity, three times as large as
the central cube. Surface covered, with numerous thin, bristle-shaped, bent
spinules.

{280}_Dimensions._--Diameter of the spongy shell 0.24, of its inner cavity
0.12, of the central cube 0.02; length of the spines 0.4, breadth 0.01.

_Habitat._--North Pacific, Station 254, surface.

3. _Octodendron pinetum_, n. sp.

Radial spines eight, three-sided prismatic, three to four times as long as
the diameter of the inner shell-cavity, with three spirally contorted
edges. From each spine arise at equal distances (equal to the half radius
of the cavity) fifteen to twenty verticils of branches which increase in
size from the base of the spine. Each verticil is composed of six forked
branches (two from each spine-edge); the inferior are richly branched, and
form by their connection the loose network of the spongy cortical shell,
the surface of which is covered with numerous bent threads. Each spine
bears a resemblance to a pine tree.

_Dimensions._--Diameter of the spongy shell 0.3, of its inner cavity 0.15,
of the central cube 0.02; length of the spines 0.5 to 0.6, breadth 0.01.

_Habitat._--Central Pacific, Station 272, surface.

4. _Octodendron araucaria_, n. sp.

Radial spines eight, three-sided prismatic, with three spirally contorted
edges, eight to ten times as long as the diameter of the inner shell-cavity
(the free distal portion twice to three times as long). From each spine
arise at equal distances thirty to thirty-five verticils of branches, which
decrease in size towards the distal end. Each verticil is composed of six
forked branches (two from each spine-edge); the inferior are richly
ramified and form by their connection the loose spongy framework; the
distal branches bear on the free end elegant spathillæ (as in the following
species). Surface covered with innumerable straight bristles, as long as
the cavity radius, and ending with a spathilla. Diameter of the central
capsule about equal to the radius of the spongy sphere, its membrane is
double-edged.

_Dimensions._--Diameter of the spongy shell 1.4 mm., of its inner cavity
0.14, of the central cube 0.02, of the central capsule 0.5 to 0.6; length
of the spines 0.9 to 1.2, breadth 0.02.

_Habitat._--South Pacific, Station 288, surface.

5. _Octodendron spathillatum_, n. sp. (Pl. 18, figs. 2, 4).

Radial spines eight, three-sided prismatic, with contorted edges, five to
eight times as long as the diameter of the shell-cavity (the free part
twice as long). From each spine arise ten to twelve verticils of lateral
branches; each verticil composed of three forked branches. The proximal
larger branches ramify richly, and form by their connection the loose
spongy framework of the spherical shell; the distal smaller branches are
simple or bifurcated, and the ramules are provided with an elegant
spathillum at the end (fig. 4). Entire surface of the spongy sphere covered
with innumerable bristle-shaped radial spines (half as long as the cavity
radius), zig-zag, bent, with beards, and with a spathillum at the end.

{281}_Dimensions._--Diameter of the spongy sphere 1 mm., of its central
cavity 0.15, of the central cube 0.02; length of the spines 1 to 1.2 mm.,
breadth 0.008.

_Habitat._--Central Pacific, Station 271, surface.

Subgenus 2. _Octodendronium_, Haeckel.

_Definition._--Radial spines thirty-two or more, eight primary (arising
from the eight corners of the central cube) and twenty-four or more
secondary (between them).

6. _Octodendron verticillatum_, n. sp.

Radial spines thirty-two, with three denticulate straight edges, six to
eight times as long as the diameter of the shell-cavity; the distal parts
half free. Eight primary spines arise from the eight corners of the central
cube, twenty-four secondary from the edges of these (a verticil of every
three from each primary spine). Distal free parts of all thirty-two spines
equal. Each spine with eight to ten verticils of forked lateral branches,
without spathillæ. Surface of the spongy sphere covered with short simple
bristles.

_Dimensions._--Diameter of the spongy shell 0.3, of its cavity 0.12, of the
central cube 0.02; length of the spines 0.7 to 1 mm., breadth 0.02.

_Habitat._--South Pacific, Station 291, surface.

7. _Octodendron contortum_, n. sp.

Radial spines thirty-two, disposed in a similar manner to those of the
former species (eight primary and twenty-four secondary); also the spongy
shell of the same shape. The difference arises in the form of the spines,
the three edges of which are much broader and spirally contorted around the
axis; and thus the corresponding branches of the verticil do not lie in the
same meridian-plane, but alternate one with another.

_Dimensions._--Diameter of the spongy shell 0.4, of its cavity 0.1, of the
central cube 0.02; length of the spines 0.8 to 1.2, breadth 0.04.

_Habitat._--Central Pacific, Station 274, surface.

8. _Octodendron arboretum_, n. sp.

Radial spines sixty to ninety, three-sided prismatic, with spirally
contorted and denticulate edges, six to eight times as long as the diameter
of the inner shell-cavity. Eight primary arise from the eight corners of
the central cube, twenty-four others from their three edges (as in the two
former species); the remaining thirty to sixty spines seem to arise between
the former and immediately from the dense spongy framework, which is twice
as thick as the diameter of the inner shell-cavity. The numerous verticils
of the free distal part are of equal shape in all the spines, composed of
three forked branches in the terminal, and of more ramified branches in the
inferior parts. Entire surface of the spongy shell covered with simple
radial bristles, without spathillæ.

{282}_Dimensions._--Diameter of the spongy shell 0.75, of its central
cavity 0.15, of the central cube 0.02; length of the spines 0.9 to 1.2,
breadth 0.02.

_Habitat._--Tropical Pacific, Station 225, surface.

Genus 120. _Spongosphæra_,[159] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Astrosphærida# with two concentric latticed medullary
shells, connected by radial beams; the outer is immediately enveloped by
the spongy framework, and bears numerous radial spines.

The genus _Spongosphæra_ (in the definition here restricted) differs from
_Spongopila_ in the double medullary shell, which in the latter is simple;
it exhibits to the latter the same relation that _Spongodictyon_ among the
Liosphærida bears to _Spongoplegma_. The outer medullary shell of
_Spongosphæra_ is immediately enveloped by the spongy wicker-work, which
everywhere pierces the wall of the central capsule.

1. _Spongosphæra streptacantha_, Haeckel.

_Spongosphæra streptacantha_, Haeckel, 1862, Monogr. d. Radiol., p. 455,
Taf. xxvi. figs. 1-3.

Spongy shell of polyhedric, irregular outline, the framework being
prolonged sheath-like into the eight to twelve radial spines, which are
quite irregularly distributed, very large, three-sided prismatic, with
three serrated, spirally contorted edges; their length is twice to four
times as great as the diameter of the spongy body; they arise with thinner
bases from the outer medullary shell, which is three times as broad as the
inner, both having roundish pores, twice to four times as broad as the
bars; surface without radial by-spines.

_Dimensions._--Diameter of the spongy shell 0.2 to 0.6, outer medullary
shell 0.04 to 0.06, inner 0.012 to 0.016.

_Habitat._--Cosmopolitan; common in all warmer seas, surface.

2. _Spongosphæra polyacantha_, J. Müller.

_Spongosphæra polyacantha_, J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 32, Taf. iv. figs. 1-4.

Spongy shell spherical, with ten to twenty large radial spines, which arise
with thinner bases from the medullary shell, and are prominent on the
surface at different lengths; they are three-sided prismatic (not
four-sided), with three leaf-shaped straight edges. Whilst the spongy
framework is much looser than in the preceding species, both medullary
shells have nearly the same shape.

_Dimensions._--Diameter of the spongy shell 0.2 to 0.5, outer medullary
shell 0.04 to 0.06, inner 0.012 to 0.016.

_Habitat._--Mediterranean (Nice); Atlantic, Station 353, surface.

{283}Subgenus _Spongosphæromma_, Haeckel.

_Definition._--Radial spines on the surface of the spongy shell of two
different kinds; large piercing main spines and small superficial
by-spines.

3. _Spongosphæra helioides_, Haeckel.

_Spongosphæra helioides_, Haeckel, 1862, Monogr. d. Radiol., p.456, Taf.
xii. figs. 11-13.

Spongy shell spherical, with numerous curved, bristle-shaped by-spines on
the surface, as long as the radius. Main spines ten to twenty, irregularly
disposed, prismatic, with three dentated, spirally contorted edges, broader
toward the distal end. Outer medullary shell three times as broad as the
inner, with polygonal meshes (of the same size as those in the spongy
framework) and fine bars.

_Dimensions._--Diameter of the spongy shell 0.2, outer medullary shell
0.02, inner 0.007.

_Habitat._--Mediterranean (Messina).

4. _Spongosphæra quadricuspis_, n. sp.

Spongy shell spherical, with numerous curved, bristle-shaped by-spines on
the surface, half as long as the radius. Main spines twenty to thirty,
irregularly disposed, prismatic, with three dentated, spirally contorted
edges, and at the distal end four strong pyramidal divergent teeth (three
as terminations of the edges, the fourth as end of the spine axis). Outer
medullary shell twice as broad as the inner, both having circular, regular
pores, twice to three times as broad as the bars.

_Dimensions._--Diameter of the spongy shell 0.3, outer medullary shell
0.04, inner 0.013.

_Habitat._--Central Pacific, Station 272, surface.

Genus 121. _Rhizosphæra_,[160] Haeckel, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 840.

_Definition._--#Astrosphærida# with two concentric latticed medullary
shells, connected by radial beams; from the outer arise numerous radial
spines, which at equal distances are connected by a latticed spherical
cortical shell, surrounded by a spongy framework.

The genus _Rhizosphæra_ exhibits the same relation to _Spongosphæra_ that
_Rhizoplegma_ bears to _Spongopila_; but in the latter the latticed
medullary shell is simple, in the two former double. The wall of the
central capsule is pierced only by the radial spines connecting the
medullary and the cortical shells.

1. _Rhizosphæra trigonacantha_, Haeckel.

_Rhizosphæra trigonacantha_, Haeckel, 1862, Monogr. d. Radiol., p. 452,
Taf. xxv. figs. 1-7.

Central cavity of the spongy cortical shell twice as broad as the diameter
of the outer medullary shell. Bars of all three shells of the same breadth
as the thirty to fifty (or more) radial spines, {284}which are three-sided
prismatic; their outer free distal end only as long as the diameter of the
inner medullary shell.

_Dimensions._--Diameter of the spongy shell 0.25, of its inner cavity 0.2,
outer medullary shell 0.1, inner 0.05.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific, surface
from many Stations.

2. _Rhizosphæra serrata_, n. sp. (Pl. 18, figs. 5-7).

Central cavity of the spongy shell five times as broad as the diameter of
the outer medullary shell. Bars of all three shells of the same breadth as
the forty to sixty (or more) thin radial beams between them. These are
three-sided prismatic, with denticulate edges, scarcely half as broad as
their outer prolongations, which are half as long as the shell radius, and
possess three spirally contorted serrated edges. (The figured specimen is a
young one; in the older specimens the spongy framework of the cortical
shell is much more developed.)

_Dimensions._--Diameter of the spongy shell 0.3, of its central cavity
0.22, outer medullary shell 0.06, inner 0.02.

_Habitat._--Central Pacific, Stations 270 to 274, surface.

3. _Rhizosphæra leptomita_, Haeckel.

_Rhizosphæra leptomita_, Haeckel, 1862, Monogr. d. Radiol., p. 453, Taf.
xxv. figs. 8-10.

Central cavity of the spongy cortical shell twice as broad as the diameter
of the outer medullary shell; bars of both very thin, only one-third as
broad as the bars of the inner medullary shell. Radial spines thirty to
fifty (or more), curved, three-sided prismatic; inside the spongy shell as
thin as their bars, outside three times as broad.

_Dimensions._--Diameter of the spongy shell 0.27, of its inner cavity 0.2,
outer medullary shell 0.1, inner 0.05.

_Habitat._--Mediterranean (Messina); Atlantic, Stations 348 to 354,
surface.

----

Suborder PRUNOIDEA, Haeckel, 1883 (Pls. 13-17, 39, 40).

_Definition._--SPUMELLARIA with an ellipsoidal or cylindrical central
capsule, prolonged into one axis (sometimes articulate by annular
transverse strictures); with an ellipsoidal or cylindrical, fenestrated
siliceous shell (often articulate by annular strictures), invariably
prolonged into one axis. Fundamental form monaxon, usually with the poles
of the prolonged dimensive main axis equal.

The suborder #Prunoidea# comprises those SPUMELLARIA in which the
fenestrated spherical shell appears prolonged into one axis. The geometric
fundamental form of the shell, which in the #Sphæroidea# was a sphere, in
this case therefore becomes an ellipsoid, and whilst in the former all axes
originally have the same value (Homaxonia), {285}here one main axis is
constantly larger than all other axes (Monaxonia). Usually both poles of
this main axis are equal (Haplopola); but in some genera both poles become
different (Diplopola).

In the #Sphæroidea# all planes going through the centre of the shell are
circular, whereas in the #Prunoidea# only those planes are circular which
are perpendicular to the main axis; all other planes going through the
centre are elliptical; the largest of these are the meridian planes, in
which is situated the main axis. Commonly all meridian planes are equal, as
no transverse axes (or cross axes) are differentiated.

In my Monograph (1862) only very few forms of #Prunoidea#, such as
_Didymocyrtis_ and _Spongurus_, are described, and the greater part of them
are distributed under different genera of #Sphæroidea#, such as _Haliomma_
and _Actinomma_. In my Prodromus (1881) I separated them as the family
Zygartida (p. 462). But it seems now much more convenient to restrict this
term to a particular family and to give a wider extension to the whole
suborder under the name #Prunoidea# (called after the characteristic
ellipsoidal form of a plum, or _Prunus_, with its stone).

The suborder #Prunoidea# comprises seven different families, of which the
Ellipsida constitutes the simplest and the probable common ancestral group.
In all Ellipsida the fenestrated shell is simple, and never composed of
concentric or twin shells. In their primitive genus _Cenellipsis_, the
whole shell is geometrically nothing more than a simple ellipsoid (Pl. 39,
figs. 1, 2). By development of radial spines it passes into _Ellipsidium_,
by development of spongy framework into _Spongellipsis_. In the greater
part of this family large spines are developed on both poles of the main
axis of the shell, but sometimes instead of these solid spines, two
opposite hollow fenestrated tubes are developed (_Pipettella_, Pl. 39, fig.
6).

The second family of #Prunoidea#, the Druppulida, is much richer in
different forms than the simple Ellipsida. In this case the ellipsoidal
shell is composed of two or three (rarely more) concentric shells.
Constantly one or two of these fenestrated shells are enclosed in the
central capsule, and may therefore be called "medullary shells"; and one or
two (rarely more) lie outside the central capsule, "cortical shells." The
inner medullary shells (one or two) are either spherical or ellipsoidal;
the outer cortical shells (one or two, rarely more) are always ellipsoidal.
All concentric shells are connected by radial beams. In the simplest form
of the subfamily, _Druppula_ (Pl. 39, fig. 3), one medullary shell is
connected with one cortical shell. By duplication of the medullary shell
arises _Prunulum_ (Pl. 39, fig. 4), and by duplication or multiplication of
the cortical shell _Cromyodruppa_ (Pl. 15, figs. 1-4) is formed. In by far
the greater portion of this subfamily large spines are developed on both
poles of the main axis of the cortical shell (Pls. 16, 17), but sometimes
also instead of these solid spines two opposite hollow fenestrated tubes
are developed (_Pipetta_, Pl. 39, figs. 7, 8).

{286}A third family of #Prunoidea#, closely allied to the two preceding
families, is the Spongurida, in which we include all #Prunoidea# with an
ellipsoidal or cylindrical, unjointed shell, in which the lattice-work of
the cortical shell is transformed into an irregular, siliceous framework.
In the simplest form, _Spongellipsis_, the simple lattice-shell of
_Cenellipsis_ is substituted by an external spongy envelope. In other cases
(_Spongurus_ and allied genera) the whole cavity of this external spongy
shell is distended with a fine spongy framework. The subfamily of
Spongodruppida is distinguished by the possession of a simple or double
latticed medullary shell; this lies in the midst of the central capsule,
and is connected by radial beams (perforating its membrane) with the
enveloping spongy cortical shell. The surface of the latter may bear either
radial spines, or two opposite strong polar spines, at the poles of the
main axis (Pl. 17, fig. 12).

Closely allied to the Ellipsida and Druppulida are two other families of
the #Prunoidea#, the Artiscida and Cyphinida, which differ from the former
by a circular constriction in the equatorial plane of the ellipsoidal
shell; and in this way assume a characteristic twin form, like a figure of
eight. In the Artiscida the shell is simple (as in the Ellipsida), whereas
in the Cyphinida it is composed of two or more concentric shells (as in the
Druppulida). The simplest form of the Artiscida is _Artiscus_ (Pl. 39, fig.
9), differing from _Cenellipsis_ in the ring-shaped, equatorial
constriction. In other Artiscida polar appendages are developed on both
poles of the main axis, either in the form of solid, strong spines
(_Stylartus_), or hollow fenestrated tubes (_Cannartus_, Pl. 39, fig. 10).

The family Cyphinida differs from the Druppulida in the equatorial
constriction of the shell, and from the Artiscida in the presence of two or
more concentric shells. One or two of these concentric fenestrated shells
are enclosed in the central capsule (and therefore may be called "medullary
shells"); the others (one or two, rarely more) lie outside of the central
capsule (therefore "cortical shells"). The internal "medullary shells" are
always spherical or somewhat lenticular, compressed from both sides; the
external "cortical shells" have constantly a ring-like constriction in the
equatorial plane, and "twin-shells" are therefore like a figure of eight.
The simplest form of this subfamily is _Cyphanta_, composed of a simple
medullary shell and a simple cortical shell, the two being connected in the
equatorial plane by radial beams. In Cyphonium (Pl. 39, fig. 12) the
medullary shell is doubled, and in _Cypassis_ (Pl. 39, fig. 13) the
cortical shell likewise. On both poles of the main axis strong spines are
often developed (_Cyphinus_, Pl. 39, fig. 14), or hollow fenestrated tubes
(_Cannartidium_, Pl. 39, figs. 16-19).

The equatorial constriction of the ellipsoidal shell, which characterises
the Artiscida and Cyphinida, is repeated or multiplied in the two following
families, in the Panartida and Zygartida; in the former we find three
ring-like strictures, in the latter five or more (lying in parallel
transverse planes); therefore the fenestrated shell is composed in the one
instance of four chambers, in the other of six or {287}more; all the
chambers form a single series and have a common main axis. All
constrictions lie in planes parallel to the equatorial plane of the
original ellipsoid; in the centre of the latter constantly lies a double
"medullary shell," composed of two concentric, either spherical or
lenticular, compressed shells. In all Panartida we call the two inner
chambers (on both sides of the equatorial constriction) "proximal
chambers," the two outer chambers (on the poles of the main axis) "distal
chambers." The four-chambered cortical shell of the Panartida is either
simple (in _Panartus_, Pl. 40, figs. 1-4) or double, with an external
mantle (as in _Peripanartus_, Pl. 40, figs. 5-7). The simplest form of the
subfamily is _Panartus_ (_loc. cit._). In this case also on both poles of
the main axis may be developed solid spines, or hollow fenestrated tubes
(_Panarium_, Pl. 40, fig. 9).

The seventh and last family of the #Prunoidea#, the Zygartida is most
nearly allied to the Panartida, and appears as a further developmental step
from that family. Whilst in the Panartida the cortical shell is constantly
four-chambered, with three parallel ring-like constrictions, in the
Zygartida it is always prolonged and composed of six or more chambers,
separated by five or more ring-shaped constrictions, in the middle of which
is the equatorial stricture. In the centre of the latter (as also in the
Panartida) always lies the double medullary shell, composed of two
concentric, spherical, or lenticular shells. The number of the chambers of
the cortical shells is commonly six or eight (with five to seven ring
strictures), but it often mounts to ten and sometimes to twenty (with
nineteen strictures), as in some species of _Zygartus_ (Pl. 40, fig. 13).
All the chambers lie in one series, one behind another, with a common main
axis. The cortical shell is usually simple (in _Ommatocampe_, Pl. 40, fig.
10), sometimes double (in _Desmocampe_, Pl. 40, fig. 12), rarely triple (in
ZYGOCAMPE, Pl. 40, fig. 13). In all three cases hollow fenestrated tubes
may be developed on the poles of the main axis.

The morphological references and the phylogenetic affinities of all
#Prunoidea# are so complex, that they seem to represent a quite natural
group; all forms of it may be derived from the common ancestral form
_Cenellipsis_. But a far more difficult question is the manner in which its
pedigree may be constructed. The oldest family is probably the simplest,
namely, Ellipsida. From this the Druppulida may be derived by production of
medullary shells, the Artiscida by equatorial constriction. The Cyphinida
can be produced either from the Druppulida by equatorial constriction or
from the Artiscida by development of medullary shells. The Panartida appear
as further developmental steps of the Cyphinida, by duplication of the
chamber number; and the Zygartida as further productions of the Panartida,
by increasing the number of the chambers.

The seven subfamilies of the #Prunoidea# can be arranged in two sections
according to the presence or absence of medullary shells. The Ellipsida,
Spongellipsida, and Artiscida possess a simple cortical shell, without a
medullary shell; they represent the {288}section Cenoprunida. All other
families possess medullary shells, and so represent the section
Coccoprunida.

Another character, which can be employed in the arrangement of the seven
subfamilies in some larger groups, is the presence or absence of ring-like
constrictions, by which the cortical shell is divided into chambers. I. The
Monoprunida comprise all forms without any constriction, of which the
Ellipsida are without a medullary shell, the Druppulida with one or two
medullary shells, and the Spongurida with a spongy cortical shell. II. The
Dyoprunida contain all forms with a cortical twin shell, or with two
chambers separated by one equatorial constriction, of which the Artiscida
are without a medullary shell and the Cyphinida have one or two medullary
shells. III. The Polyprunida comprise all forms with several (three or
more) constrictions, which separate four or more chambers, of which the
Panartida have three constrictions and four chambers, and the Zygartida
five or more constrictions and six or more chambers.

_The Central Capsule_ of the #Prunoidea# is originally ellipsoidal
(monaxial), and preserves this form in the greater part of the genera. In
some groups, where the axis of the ellipsoid is much prolonged, it passes
over to the cylindrical form (with hemispherical vaultings on both poles),
as in _Spongurus_ and _Spongocore_, in many Panartida and Zygartida. Very
often the ellipsoidal or cylindrical capsule gets annular transverse
constrictions, corresponding to those of the enveloping cortical shell (one
single, equatorial stricture in the Artiscida and Cyphinida, three
strictures in the Panartida, five or more in the Zygartida). In the
Cenoprunida (Ellipsida and Artiscida, also in _Spongellipsis_) the central
capsule lies freely in the cavity of the cortical shell, separated from its
inner surface by the jelly-envelope; in all other groups it contains a part
of the skeleton, the medullary shell and the beams which connect it with
the enveloping cortical shell.

_Synopsis of the Families of_ #Prunoidea#.

{a. Shell simple, latticed (not
{ spongy), without enclosed internal
{ enclosed internal shells, 1. ELLIPSIDA.
A. MONOPRUNIDA. {
{b. Shell composed of two or more
Shell without { concentric latticed shells
transverse stricture { (not spongy), 2. DRUPPULIDA.
{
{c. Shell partially or wholly
{ composed of an irregular spongy
{ framework, 3. SPONGURIDA.

B. DYOPRUNIDA. {
{d. Shell simple, without enclosed
Shell bilocular, { internal shells, 4. ARTISCIDA.
divided by an {
equatorial stricture {
into two communicating{e. Shell composed of two or more
hemi-ellipsoidal { concentric shells, 5. CYPHINIDA.
shells. {

C. POLYPRUNIDA. {f. Shell with three parallel
{ strictures and therefore four
Shell multilocular, { cameræ, 6. PANARTIDA.
divided by three {
or more parallel {g. Shell with five or more parallel
transverse strictures { strictures and therefore six
into four or more { or more cameræ, 7. ZYGARTIDA.
serial cameræ. {

{289}Family XI. #ELLIPSIDA#, Haeckel, 1882 (Pls. 13, 14, 39).

_Definition._--#Prunoidea# with simple ellipsoidal shell, without
equatorial stricture (without enclosed medullary shell); network a simple
lattice lamella, not spongy. Central capsule ellipsoidal or cylindrical,
without annular equatorial constriction.

The family #Ellipsida# comprises the simplest forms of #Prunoidea#, and
probably represents the ancestral forms of this whole suborder. The
fenestrated shell, which encloses the ellipsoidal central capsule, is a
perfectly simple "cortical shell" of the same form, without enclosed
"medullary shell." Its form is commonly a regular monaxial ellipsoid;
sometimes a little modified by unequal growth of the two poles of the main
axis. Two opposite large spines are often developed at these poles, or it
may be that instead of these, two hollow fenestrated tubes are present.

The ellipsoidal fenestrated shell exhibits in the regular Ellipsida all the
characters of a geometric ellipsoid; one main axis surpasses in length all
other possible axes. All sections going through this main axis are
"meridian sections," with elliptical periphery; all sections perpendicular
to the main axis are "transverse sections," with circular periphery. The
largest of these is the equatorial section, which divides the main axis
into halves. The diameter of this equatorial plane is the "minor axis" of
the ellipsoid.

The proportion of the two axes of the ellipsoidal shell, of the major
vertical or main axis and the minor horizontal or equatorial axis, is
commonly between 6 : 5 and 3 : 2. In the former case it approaches the
spherical shell, from which it is derived; in the latter case it becomes
almost fusiform or cylindrical. The network of silex, constituting the
shell, is constantly a simple latticed lamella, never composed of
concentric shells (as in Druppulida) or spongy (as in Spongurida). The
network is often very regular and elegant, in other cases irregular.

The simplest genus among the Ellipsida, and probably the common ancestral
form of the whole subfamily, is the genus _Cenellipsis_, possessing a
simple ellipsoidal shell without any appendages. It is derived from
_Cenosphæra_ (the simplest spherical shell) by the prolongation of one
axis. _Cenellipsis_ passes over into _Ellipsidium_ by the production of
radial spines on the surface (corresponding to _Heliosphæra_). _Axellipsis_
is a peculiar genus differing from _Cenellipsis_ in an axial rod, which
corresponds to the minor or equatorial axis. In all other genera of the
subfamily both poles of the main axis are distinguished by peculiar polar
prolongations, either hollow fenestrated tubes (as in _Pipettella_) or
strong solid spines. Both polar spines are of equal size and similar form
in _Ellipsoxiphus_, unequal in _Ellipsostylus_. From the latter is derived
_Lithapium_, by reduction and loss of one spine (so that only one remains);
_Lithomespilus_, by production of a bunch of several spines at one pole. In
the three latter genera both poles of the main axis are unequal, in all
others equal.

{290}The central capsule of the Ellipsida is in all cases ellipsoidal, and
occupies the largest part of the shell, being separated from its inner
surface by a thinner or thicker jelly-mantle.

_Synopsis of the Genera of Ellipsida._

-------------------------------------------------------------------------
Ellipsoidal shell without polar appendages (neither solid spines nor
hollow tubes at the poles of the axis).
-------------------------------------------------------------------------
Surface without {Shell cavity
radial spines. { simple, without
{ an axial rod, 122. _Cenellipsis_.
{
{Shell cavity
{ with a transverse
{ axial rod, 123. _Axellipsis_.

Surface covered with radial spines, 124. _Ellipsidium_.
-------------------------------------------------------------------------
Ellipsoidal shell with polar appendages (either solid spines or hollow
fenestrated tubes) at the poles of the main axis.
-------------------------------------------------------------------------
Solid spines of {Shell cavity
similar shape, { simple, without
at both poles of { axial rods, 125. _Ellipsoxiphus_.
the main axis. {
{Shell cavity
{ with a cross of
{ axial rods, 126. _Axoprunum_.

Solid spines of {Two polar spines
different shape, { of different
at both poles { shape, 127. _Ellipsostylus_.
of the main {
axis. {A bunch of spines
{ at one pole
{ only, 128. _Lithomespilus_
{
{Only a single
{ spine at one
{ one pole, 129. _Lithapium_.

Two hollow fenestrated tubes opposite,
at the poles of the main axis, 130. _Pipettella_.

Genus 122. _Cenellipsis_,[161] n. gen.

_Definition_.--#Ellipsida# with simple ellipsoidal shell, without radial
spines and without polar tubes.

The genus _Cenellipsis_ is the simplest and most primitive form, not only
among the Ellipsida, but also among the #Prunoidea#, and it may therefore
be regarded as the common ancestral form of the whole family. It
corresponds to _Cenosphæra_ among the #Sphæroidea#, to _Cenodiscus_ among
the #Discoidea#, to _Cenolarcus_ among the #Larcoidea#. Probably it is
derived from _Cenosphæra_ by prolongation of one axis.

Subgenus 1. _Cenellipsium_, Haeckel.

_Definition._--Network of the shell regular, with meshes of equal size and
similar form.

{291}1. _Cenellipsis primitiva_, n. sp.

Proportion of the longer axis of the ellipsoid to the shorter = 3 : 2.
Network of the thin wall very delicate and regular, with hexagonal pores.
All pores of nearly the same size and form, ten to twelve times as broad as
the bars; nine to twelve on the half equator of the shell. Surface smooth.

_Dimensions._--Longer axis of the ellipsoid 0.09 to 0.11, shorter axis 0.06
to 0.08; pores 0.01, bars between them 0.001.

_Habitat._--Pacific, central area, Station 272, surface.

2. _Cenellipsis faceta_, n. sp. (Pl. 39, fig. 1).

Proportion of the longer axis of the ellipsoid to the shorter = 4 : 3.
Network of the thin wall delicate and regular. All the pores of the same
size and form, circular, with a hexagonal frame, twice to three times as
broad as the bars; eight to ten on the half equator. Surface smooth or
slightly spiny.

_Dimensions._--Longer axis of the ellipsoid 0.12 to 0.13, shorter axis 0.09
to 0.1; pores 0.013, bars 0.005.

_Habitat._--Pacific, central area. Station 274, surface.

3. _Cenellipsis ehrenbergii_, Haeckel.

_Haliomma (?) cenosphæra_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf xxvi. fig. 5.

Proportion of the longer axis to the shorter = 2 : 1. Network of the thin
wall regular or subregular. All pores circular, nearly of the same size,
without hexagonal frame, three to four times as broad as the bars; eight to
nine on the half equator. Surface covered with small thorns.

_Dimensions._--Longer axis of the ellipsoid 0.16 to 0.2, shorter axis 0.08
to 0.1; pores 0.1, bars 0.003.

_Habitat._--Fossil in the Tertiary rocks of Barbados (Ehrenberg, Haeckel).

4. _Cenellipsis circopora_, n. sp.

Proportion of the longer axis to the shorter = 5 : 3. Network of the thin
wall delicate and regular. All pores circular, nearly of the same size,
without hexagonal frame, twice as broad as the bars; fifteen to twenty on
the half equator. Surface quite smooth.

_Dimensions._--Longer axis of the ellipsoid 0.15 to 0.2, shorter axis 0.09
to 0.12; pores 0.08, bars 0.4.

_Habitat._--North Pacific, Station 236, surface.

5. _Cenellipsis micropora_, n. sp.

Proportion of the longer axis to the shorter = 5 : 4. Network of the thick
wall quite regular. All pores circular, without hexagonal frame, very
small, scarcely half as broad as the bars; six to eight on the half
equator. Surface quite smooth.

{292}_Dimensions._--Longer axis of the ellipsoid 0.1, shorter axis 0.08;
pores 0.004, bars 0.01.

_Habitat._--South Pacific, Station 289, surface.

Subgenus 2. _Cenellipsula_, Haeckel.

_Definition._--Network of the shell irregular, with meshes of different
size or form.

6. _Cenellipsis heteropora_, n. sp.

Proportion of the longer axis to the shorter = 3 : 2 or = 5 : 3. Network of
the thin wall very irregular, with polygonal meshes of different size and
form (for the most part pentagonal or hexagonal, but also many tetragonal
or heptagonal). The largest pores four to six times as broad as the
smallest; ten to twenty pores on the half equator; bars between them thin.
Surface smooth or somewhat spiny.

_Dimensions._--Longer axis of the ellipsoid 0.18 to 0.22, shorter axis 0.1
to 0.12; pores 0.004 to 0.008, bars 0.001.

_Habitat._--Indian Ocean, near the Cocos Islands, surface, Rabbe.

7. _Cenellipsis maxima_, n. sp.

Proportion of the longer axis to the shorter = 3 : 2. Network of the thin
wall very irregular, with polygonal meshes of very different size and form,
twice to eight times as broad as the bars; fifty to sixty on the half
equator. Surface smooth.

_Dimensions._--Longer axis of the ellipsoid 0.36, shorter axis 0.25; pores
0.004 to 0.015, bars 0.002.

_Habitat._--Equatorial Atlantic, Station 347, surface.

8. _Cenellipsis ovulum_, n. sp.

Proportion of the longer axis to the shorter = 3 : 2. Network of the thin
wall very irregular, with roundish or longish pores of very different size
and form. Pores flat, with even margins, thirty to forty on the half
equator, twice to six times as broad as the bars. Surface smooth. (This
species being observed alive, showed a transparent central capsule of
ellipsoidal form, two-thirds as large as the shell.)

_Dimensions._--Longer axis of the ellipsoid 0.3, shorter axis 0.2; pores
0.04 to 0.012, bars 0.002.

_Habitat._--Indian Ocean, Ceylon, Haeckel; Pacific, central area, Station
266, surface.

9. _Cenellipsis infundibulum_, n. sp. (Pl. 39, fig. 2).

Proportion of the longer axis to the shorter = 3 : 2. Network of the thick
wall irregular, with roundish pores of different size and form. Pores
funnel-like, with prominent, irregular, spiny {293}crests between them,
their outer aperture three to four times as broad as the inner; twelve to
fifteen pores on the half equator. Surface prickly.

_Dimensions._--Longer axis of the ellipsoid 0.14, shorter axis 0.11; outer
aperture of the pores 0.01 to 0.012, inner aperture 0.003 to 0.004.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

10. _Cenellipsis oblonga_, n. sp.

Proportion of the longer axis to the shorter = 2 : 1. Network of the thick
wall irregular, with roundish pores of different size and form. Pores with
prominent conical edges, about as broad as the bars; twenty to twenty-five
on the half equator. Sometimes each pore is prolonged into a short conical
tubulus.

_Dimensions._--Longer axis of the ellipsoid 0.22, shorter axis 0.12; pores
and bars 0.008 to 0.012.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Genus 123. _Axellipsis_,[162] n. gen.

_Definition._--#Ellipsida# with simple ellipsoidal shell, without radial
spines or polar tubes, but with an inner transverse axial rod, which
corresponds to the shorter axis of the ellipsoid.

The genus _Axellipsis_ differs from the simple _Cenellipsis_ in a very
peculiar character, namely, the presence of an inner siliceous bar marking
the minor axis of the ellipsoidal shell. If in some forms of _Druppula_
(e.g., Pl. 39, fig. 3) we remove the central medullary shell and prolong
both beams (connecting it with the cortical shell) till they unite in the
centre, we shall arrive at _Axellipsis_. Therefore _Axellipsis_ may be
derived in the same way from _Druppula_ as _Axoprunum_ is from
_Lithatractus_, by phyletic loss of the medullary shell.

1. _Axellipsis perforata_, n. sp.

Proportion of the major axis of the ellipsoid to the minor = 3 : 2. Pores
of the shell regular, circular, twice as broad as the bars; ten to twelve
on the half equator. Surface smooth.

_Dimensions._--Longer axis of the shell 0.12, shorter axis 0.08; pores
0.006, bars 0.003.

_Habitat._--Central area of the Pacific, Station 266, depth 2750 fathoms.

2. _Axellipsis lobata_, n. sp.

Proportion of the major axis to the minor = 5 : 4. Meshes of the shell
irregular, roundish, lobed, with three to six indentations (each mesh
formed by the confluence of three to six, commonly {294}four or five,
smaller pores); eight to nine meshes on the half equator, these being twice
to four times as broad as the bars.

_Dimensions._--Longer axis of the shell 0.1, shorter axis 0.08; pores 0.007
to 0.01, bars 0.003.

_Habitat._--Tropical zone of the Western Pacific, Station 225, surface.

Genus 124. _Ellipsidium_,[163] n. gen.

_Definition._--#Ellipsida# with simple ellipsoidal shell, with numerous
radial spines on the surface, without polar spines or tubes.

The genus _Ellipsidium_ differs from _Cenellipsis_ solely in the
development of numerous radial spines on the surface, and exhibits the same
phylogenetic relation to it that _Heliosphæra_ bears to _Cenosphæra_.

1. _Ellipsidium pandanidium_, n. sp.

Proportion of the longer axis of the ellipsoid to the shorter = 5 : 4.
Shell thick walled, with regular and hexagonal meshes, twice as broad as
the bars; eighteen to twenty on the half equator. In each hexagon-corner
(between three pores) arises a short, three-sided pyramidal spine, half as
long as the equatorial radius, and as thick at the base as a single mesh.

_Dimensions._--Major axis of the ellipsoid 0.15, minor 0.12; pores 0.006,
bars 0.003; length of the radial spines 0.03, basal breadth 0.006.

_Habitat._--Fossil in the Tertiary rocks of Barbados (Haeckel).

2. _Ellipsidium datura_, n. sp.

Proportion of the longer axis of the ellipsoid to the shorter = 4 : 3.
Shell thick walled, with regular, circular meshes, twice as broad as the
thick bars between them eight to ten meshes on the half equator. Outer
surface of the shell thorny, covered with short, conical, radial spines,
which are regularly distributed (one spine between every three meshes), and
about as long as the diameter of the meshes. (The shell is similar to the
outer shell of _Haliomma castanea_, Haeckel, Monogr. d. Radiol., Taf. xxiv.
fig. 4.)

_Dimensions._--Major axis of the ellipsoid 0.12, minor axis 0.09; meshes
0.012, bars 0.006.

_Habitat._--Western part of the Tropical Pacific, Station 225, depth 4475
fathoms.

3. _Ellipsidium artocarpus_, n. sp.

Proportion of the longer axis to the shorter = 3 : 2. Shell thick walled,
with regular, circular meshes, separated by deep furrows, which represent a
regular, hexagonal framework; on the half equator twelve to fifteen meshes,
scarcely broader than the broad bars between them. Outer {295}surface
spiny; between every three meshes arises a strong radial spine, twice to
three times as long as the diameter of the meshes; the base of the spine is
like a three-sided pyramid.

_Dimensions._--Major axis of the ellipsoid 0.12, minor axis 0.08; meshes
0.006, bars 0.005.

_Habitat._--Central area of the Pacific, Station 268, depth 2900 fathoms.

4. _Ellipsidium opuntia_, n. sp.

Proportion of the longer axis to the shorter = 5 : 4. Shell thin walled,
with irregular, roundish meshes of different size and form, about twice to
three times as broad as the irregular, thin bars between them; ten to
fifteen meshes on the half equator. Between the meshes arise numerous thin,
bristle-like, radial spines, about as long as the shorter radius of the
shell. The number of the meshes may be three to four times as great as the
number of the spines.

_Dimensions._--Major axis of the ellipsoid 0.15, minor axis 0.12; pores
0.006 to 0.01, bars 0.003 to 0.004.

_Habitat._--Southern Pacific, Station 284, surface.

5. _Ellipsidium echinidium_, n. sp.

Proportion of the longer axis to the shorter = 4 : 3. Shell thick walled,
with irregular, roundish pores of different size and form, about as large
or somewhat smaller than the broad bars; twelve to sixteen pores on the
half equator. On the surface, irregularly scattered, twenty to thirty
strong, three-sided pyramidal, radial spines, one-fourth to one-half as
long as the main axis.

_Dimensions._--Major axis of the ellipsoid 0.16, minor 0.12; pores and bars
0.002 to 0.008; length of the radial spines 0.04 to 0.08, basal breadth
0.01.

_Habitat._--Equatorial Atlantic, Station 347, depth 2250 fathoms.

Genus 125. _Ellipsoxiphus_,[164] Dunikowski, 1882, Denkschr. d. k. Akad. d.
Wiss. Wien, vol. xlv. p. 25.

_Definition._--#Ellipsida# with simple ellipsoidal shell, the main axis of
which is prolonged at both poles into two strong opposite spines of equal
size and similar form.

The genus _Ellipsoxiphus_ was established by Dunikowski (in 1882, _loc.
cit._) for those simple amphistylous fenestrated shells, formerly united
with _Xiphosphæra_, in which the mathematical form of the shell itself is
not a true sphere, but an ellipsoid. It may therefore be derived from
_Xiphosphæra_ by prolongation of the axis in which lie both polar spines;
but it may also be derived from _Cenellipsis_ by the production of two
equal spines at the poles of the main axis.

{296}Subgenus 1. _Ellipsoxiphetta_, Haeckel.

_Definition._--Network of the shell regular, with pores of equal size and
similar form.

1. _Ellipsoxiphus elegans_, n. sp. (Pl. 14, fig. 7).

Proportion of the major axis of the ellipsoid to the minor = 5 : 4. Shell
thick walled, with regular, circular meshes, separated by a regular,
hexagonal elevated framework. From each hexagon-corner arises (between
every three meshes) a short thin spine, about as long as the diameter of a
mesh; twelve to fourteen meshes on the half equator, of the same breadth as
the bars. Polar spines three-sided prismatic, cuspidate, about as long as
the axis of the shell. (This elegant and not uncommon species is remarkable
for its variations. By unequal development of both spines it passes over to
_Ellipsostylus_.) In the figured variety (Pl. 14, fig. 7) the ends of the
surface spines are connected by a very delicate network, so as to form an
outer veil. This may further be separated as a peculiar genus
_Ellipsoxiphium palliatum_.

_Dimensions._--Longer axis 0.08 to 0.13, shorter axis 0.06 to 0.11; pores
and bars 0.008; length of the polar spines 0.06 to 0.12, basal breadth
0.01.

_Habitat._--Pacific, central area, Stations 270 to 274, from 2350 to 2925
fathoms.

2. _Ellipsoxiphus flosculus_, n. sp.

Proportion of the major axis to the minor = 6 : 5. Shell very thick walled,
with regular, hexagonal meshes, eight to nine on the half equator, three to
four times as broad as the bars. The inner contour of the meshes is
circular, the outer six-lobed (similar to _Xiphostylus phasianus_, Pl. 13,
fig. 9). Polar spines three-sided pyramidal, about as long as the major
axis of the shell; their basal thickness equals one mesh.

_Dimensions._--Longer axis 0.12, shorter axis 0.1; pores 0.012 to 0.015,
bars 0.004; length of the polar spines 0.12 to 0.15, basal breadth 0.015.

_Habitat._--Fossil in the Tertiary rocks of Barbados (Teuscher).

3. _Ellipsoxiphus fragilis_, n. sp.

Proportion of the major axis to the minor = 4 : 3. Shell thin walled, with
regular, circular meshes, twice as broad as the bars; ten to twelve on the
half equator. Surface smooth. Polar spines cylindrical, at the apex
conical, half as long as the major axis.

_Dimensions._--Longer axis 0.16, shorter axis 0.12; pores 0.008, bars
0.004; length of the polar spines 0.09, basal breadth 0.008.

_Habitat._--South Atlantic, Station 325, surface.

Subgenus 2. _Ellipsoxiphilla_, Haeckel.

_Definition._--Network of the shell irregular, with pores of different size
or form.

{297}4. _Ellipsoxiphus claviger_, n. sp. (Pl. 14, fig. 3).

_Xiphosphæra clavigera_, Haeckel, 1881, Prodromus et Atlas.

Proportion of the major axis to the minor = 5 : 4. Shell thick walled, with
irregular network; meshes roundish or circular, double contoured, of
unequal size, twice to four times as broad as the bars; eight to ten on the
half equator. Polar spines club-shaped, with prominent edges, about as long
as the minor axis; thicker in their middle part than at both ends (differs
from _Xiphosphæra clavigera_ by the ellipsoidal form of the shell, and the
double length of the spines).

_Dimensions._--Longer axis 0.15. shorter axis 0.12; pores 0.007 to 0.015,
bars 0.004; length of the polar spines 0.12, thickness in the middle part
0.02.

_Habitat._--Central area of the Pacific, Station 272, depth 2600 fathoms.

5. _Ellipsoxiphus suessi_, Dunikowski.

_Ellipsoxiphus suessi_, Dunikowski, 1882, Denkschr. d. k. Akad. d. Wiss.
Wien, Bd. xlv. p. 26, Taf. v. fig. 50.

Proportion of the major axis to the minor = 7 : 5. Shell thick walled, with
irregular, roundish meshes, six to eight on the half equator, twice to
three times as broad as the bars. Polar spines nearly cylindrical, blunt,
as long as the major axis.

_Dimensions._--Longer axis 0.14; shorter axis 0.1; pores 0.018 to 0.023,
bars 0.01; length of the polar spines 0.12; thickness of them 0.035.

_Habitat._--Fossil in the Alpine Lias, Schafberg near Salzburg, Dunikowski.

6. _Ellipsoxiphus parvoforaminus_, Dunikowski.

_Ellipsoxiphus parvoforaminus_, Dunikowski, 1882, Denkschr. d. k. Akad.
d. Wiss. Wien, Bd. xlv. p. 26, Taf. v. fig. 51.

Proportion of the major axis to the minor = 7 : 6. Shell thick walled,
with irregular, roundish or oval meshes, ten to twelve on the half equator,
scarcely as broad as the bars. Polar spines conical, shorter than the
minor axis.

_Dimensions._--Longer axis 0.14, shorter axis 0.12; pores 0.016 to 0.01,
bars 0.01 to 0.02; length of the polar spines 0.1, basal thickness 0.045.

_Habitat._--Fossil in the Alpine Lias, Schafberg near Salzburg, Dunikowski.

7. _Ellipsoxiphus bipolaris_, n. sp. (Pl. 14, fig. 11).

_Xiphosphæra bipolaris_, 1881, Prodromus et Atlas.

Proportion of the major axis to the minor = 8 : 7. Shell thick walled,
with irregular, roundish meshes, twice to three times as broad as the bars;
sixteen to twenty on the half equator. Polar spines short and thick,
three-sided pyramidal, scarcely half as long as the shorter radius of the
shell; surrounding the base of every spine is a circle of eight to nine
shorter spines.

_Dimensions._--Longer axis 0.08, shorter axis 0.07; pores 0.003 to 0.007,
bars 0.002; length of the polar spines 0.02 to 0.03, basal thickness 0.02.

_Habitat._--Tropical part of the Western Pacific, Station 225, depth 4475
fathoms.

{298}8. _Ellipsoxiphus atractus_, n. sp. (Pl. 14, fig. 1).

Proportion of the major axis to the minor = 3 : 2. Shell spindle-shaped,
very thick walled, with roundish, very irregular meshes, twice to five
times as broad as the bars; eight to twelve on the half equator. The meshes
are partly simple, oblong, partly lobed or composed of two to four
(commonly three) confluent meshes. Surface smooth. Polar spines very short
and thick, shorter than the minor radius of the shell, three-sided
pyramidal, with prominent, somewhat contorted edges.

_Dimensions._--Longer axis 0.15, shorter axis 0.1; meshes 0.007 to 0.02,
bars 0.004; length of the polar spines 0.04, basal thickness 0.03.

_Habitat._--Central area of the Pacific, Station 268, depth 2900 fathoms.

Genus 126. _Axoprunum_,[165] n. gen.

_Definition._--#Ellipsida# with simple ellipsoidal shell, the main axis of
which is prolonged at both poles into two opposite spines of equal size and
similar form. Within the cavity of the shell four radial rods arise from
its inner surface, two in the main axis, two in the smallest axis,
perpendicular to the former; their free inner ends are at the same distance
from the centre.

The genus _Axoprunum_ possesses precisely the same shell as the foregoing
_Ellipsoxiphus_, but differs from it in a very remarkable peculiarity. The
two polar spines are centripetally prolonged into two internal beams, and
perpendicular to these are two other, opposite, transverse beams, marking
the minor axis of the ellipsoid. The free inner ends of all four radial
rods bear little thickened knobs, and are at the same distance from the
centre. It therefore appears as though a central, spherical, medullary
shell had been lost, and this gives a strong support to the important
hypothesis, that in many #Sphærellaria#, where the medullary shell is
absent, it may have been lost by phylogenetic reduction or retrograde
metamorphosis. In this case _Axoprunum_ (and _Ellipsoxiphus_) would arise
from _Lithatractus_.

1. _Axoprunum stauraxonium_, n. sp. (Pl. 48, fig. 4).

Shell ellipsoidal, one and one-third times as long as broad, with smooth
surface. Network regular, with circular meshes four times as broad as the
bars. Two polar spines three-sided pyramidal, half as long as the shell, as
thick at the base as a single mesh. Four inner radial beams (lying, two in
the major and two in the minor axis of the ellipsoid) very thin, at the
central free ends knob-like, thickened. The distance between two opposite
beams equals one-third of the minor axis, and indicates probably the
diameter of the lost spherical medullary shell. (Three perfect and complete
specimens of the same size and shape were observed.)

{299}_Dimensions._--Major axis of the ellipsoid 0.14, minor axis 0.11;
pores 0.012, bars 0.003; length of the polar spines 0.07, basal breadth
0.012. (Diameter of the lost medullary shell 0.03?)

_Habitat._--South Atlantic, Station 338, depth 1990 fathoms.

Genus 127. _Ellipsostylus_,[166] n. gen.

_Definition._--#Ellipsida# with simple ellipsoidal shell, the main axis of
which is prolonged at both poles into two opposite spines of different size
or form.

The genus _Ellipsostylus_ differs from _Ellipsoxiphus_ only in the unequal
size or different form of the polar spines, which in the former are equal.
This difference does not seem important at first, but in the further
development it produces very singular and strange forms. Theoretically it
is always important, because the fundamental haplopolar form of the monaxon
body becomes diplopolar by this differentiation.

Subgenus 1. _Ellipsostyletta_, Haeckel.

_Definition._--Network of the shell regular, with pores of equal size and
similar form.

1. _Ellipsostylus ornithoides_, n. sp.

Proportion of the major axis of the ellipsoid to the minor = 4 : 3. Shell
thin walled, with regular, circular meshes and hexagonal framework between
them, like that of _Xiphostylus favosus_ (Pl. 13, fig. 4). Circular pores
about as broad as the bars; eight to nine on the half equator. Surface
smooth. Polar spines three-sided pyramidal, as broad at the base as a
single hexagon; the longer spine as long as the major axis, the shorter
half as long.

_Dimensions._--Longer axis of the ellipsoidal shell 0.16, shorter axis
0.12; pores and bars 0.006; length of the longer polar spine 0.16, of the
shorter 0.08.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

2. _Ellipsostylus avicularis_, n. sp.

Proportion of the major axis to the minor = 3 : 2. Shell thick walled, with
regular, rosette-shaped meshes, three times as broad as the bars; ten to
twelve on the half equator. Every mesh on the outer margin with five to six
rounded lobes, as in _Xiphostylus phasianus_ (Pl. 13, fig. 9). Surface
smooth. Polar spines sharp edged, thick; the longer straight and equal to
the longer axis, the shorter half as long, shaped like a bird's head.

_Dimensions._--Longer axis 0.18, shorter axis 0.12; pores 0.015, bars
0.005.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

{300}3. _Ellipsostylus psittacus_, n. sp. (Pl. 13, fig. 6).

Proportion of the major axis to the minor = 4 : 3. Shell thin walled, with
regular or subregular rosette-shaped meshes, twice as broad as the bars;
eight to nine on the half equator; each mesh with three to four rounded
lobes. Surface smooth. Polar spines sharp edged, very unequal; longer spine
about twice as long as the major axis, somewhat curved; shorter spine
scarcely longer than the radius, shaped like a bird's head.

_Dimensions._--Longer axis 0.08, shorter axis 0.06; pores 0.01, bars 0.005;
length of the polar spines--longer 0.15, shorter 0.05.

_Habitat._--Central area of the Pacific, Station 266, depth 2750 fathoms.

4. _Ellipsostylus aquila_, n. sp. (Pl. 13, fig. 1).

Proportion of the major axis to the minor = 7 : 6. Shell thick walled, with
subregular, circular meshes, five to six times as broad as the bars; ten to
twelve on the half equator. Surface smooth. Polar spines sharp edged
(six-sided?), very unequal; longer spine straight, about equal to the minor
axis; shorter spine obliquely inserted, scarcely half as long, shaped like
a bird's head.

_Dimensions._--Longer axis 0.15, shorter axis 0.13; pores 0.02, bars 0.004;
length of the polar spines--longer 0.1, shorter 0.05.

_Habitat._--Central area of the Pacific, Station 272, depth 2600 fathoms.

5. _Ellipsostylus columba_, n. sp. (Pl. 13, fig. 3).

Proportion of the major axis to the minor = 4 : 3. Shell egg-shaped, thin
walled, with regular, circular meshes, three times as broad as the bars;
ten to twelve on the half equator. Surface somewhat thorny. Polar spines
nearly of equal length (equal to the major axis), but of very different
form; one straight, pyramidal, and obliquely inserted, the other like a
bird's head.

_Dimensions._--Longer axis 0.08, shorter 0.06; pores 0.003, bars 0.01;
length of the spines 0.09.

_Habitat._--South Atlantic, Station 332, surface.

6. _Ellipsostylus ciconia_, n. sp. (Pl. 13, fig. 8).

Proportion of the major axis to the minor = 7 : 5. Shell egg-shaped, thick
walled; the meshes very small, regular, circular, three times as broad as
the bars; sixteen to twenty on the half equator. Surface uneven. Polar
spines nearly cylindrical, pointed; the smaller obliquely inserted, equal
to the major axis; the larger more than twice as long and thick, furrowed
at its base. (Compare with this and the allied species _Rhabdolithis pipa_,
Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. i. fig. 27.)

_Dimensions._--Longer axis 0.07, shorter 0.05; pores 0.003, bars 0.001;
length of the polar spines--longer 0.2, shorter 0.08.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

{301}Subgenus 2. _Ellipsostylissa_, Haeckel.

_Definition._--Network of the shell irregular, with pores of different size
or form.

7. _Ellipsostylus megadictya_, Haeckel.

_Stylosphæra megadictya_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 299, Taf. viii. fig. 13.

Proportion of the major axis to the minor = 5 : 4. Shell thin walled, with
irregular, roundish, large meshes, four to five times as broad as the bars;
only four to five on the half equator. Surface smooth. Polar spines
straight, thin, angular; the shorter equal to the minor axis, the longer
four times as large.

_Dimensions._--Longer axis 0.05, shorter axis 0.04; meshes 0.01, bars
0.002; length of the polar spines--longer 0.12, shorter 0.03.

_Habitat._--Philippine Sea, 3300 fathoms, Ehrenberg.

8. _Ellipsostylus gallinula_, n. sp.

Proportion of the major axis to the minor = 3 : 2. Shell thick walled, with
irregular, roundish meshes, twice to four times as broad as the bars; eight
to ten on the half equator. Surface thorny. Polar spines conical, straight;
the longer twice as long as the major axis, the minor scarcely half as
long.

_Dimensions._--Longer axis 0.15, shorter 0.12; pores 0.01 to 0.02, bars
0.002 to 0.004.

_Habitat._--North Pacific, Station 253, surface.

9. _Ellipsostylus hirundo_, n. sp. (Pl. 13, fig. 2).

Proportion of the major axis to the minor = 4 : 3. Shell thick walled, with
irregular, roundish meshes, three to five times as broad as the bars; eight
to ten on the half equator. The inner aperture of every mesh is fenestrated
by a delicate lamella of silex, perforated by six to eight very small
circular pores. Polar spines sharp edged, more or less curved, the shorter
equal to the minor axis, the longer twice as long.

_Dimensions._--Longer axis 0.16, shorter 0.12; pores 0.01 to 0.02, bars
0.003 to 0.006; length of the polar spines--longer 0.24, shorter 0.12.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms; the
same form also fossil in the rocks of Barbados.

Genus 128. _Lithomespilus_,[167] Haeckel, 1881, Prodromus, p. 450.

_Definition._--#Ellipsida# with simple ellipsoidal or oviform shell, the
main axis of which bears at one pole a single spine, at the other a bunch
of several spines.

{302}The genus _Lithomespilus_ differs from the closely allied
_Ellipsoxiphus_ in the further differentiation of both poles of the main
axis. One pole exhibits only a single polar spine, the other pole a group
of several spines, peculiarly grouped together. It differs from the similar
_Sphæromespilus_ (Pl. 14, figs. 12, 13) in the ellipsoidal form of the
shell.

1. _Lithomespilus phloginus_, n. sp. (Pl. 14, fig. 16).

Proportion of the major axis of the ellipsoid to the minor = 4 : 3. Shell
thick walled, with circular pores of different size, the breadth of which
equals that of the bars; twelve to fifteen pores on the half equator.
Surface smooth, with the exception of a circumpolar region covered with
numerous thick spines of unequal size, which surround the large single
polar spine and are curved like a bow against its axis. This larger polar
spine is straight, and equals in length the major axis of the shell. The
other and opposite polar spine is scarcely one-fourth as long, perfectly
simple, very stout, and of a three-sided pyramidal shape.

_Dimensions._--Longer axis of the ellipsoidal shell 0.08, shorter axis
0.06; pores 0.002 to 0.006, bars 0.003 to 0.005; length of the polar
spines--longer 0.08, shorter 0.02.

_Habitat._--Indian Ocean, near Madagascar, depth 1200 fathoms, Smith.

2. _Lithomespilus phlogoides_, n. sp. (Pl. 14, fig. 17).

Proportion of the major axis to the minor = 5 : 4. Shell very thick walled,
with roundish, double-contoured pores of different size and form, twice to
three times as broad as the bars; sixteen to eighteen pores on the half
equator. Surface smooth, with the exception of a circumpolar region covered
with numerous thin spines of unequal size, which surround the large single
polar spine and are curved like a bow against its axis. This larger polar
spine is straight, and equals half the size of the major axis of the shell;
the other polar spine (sometimes double, as in the figured specimen) is
much shorter, being scarcely one-fourth as long.

_Dimensions._--Longer axis of the shell 0.08, shorter axis 0.065; pores
0.003 to 0.006, bars 0.002 to 0.004; length of the polar spines--longer
0.04, shorter 0.01.

_Habitat._--Central area of the Pacific, Stations 270 to 272, depth 2425 to
2925 fathoms.

3. _Lithomespilus flammeus_, n. sp.

_Stylosphæra species_, Bury, 1862, Polycystins of Barbados, pl. xi. fig.
3.

Proportion of the major axis to the minor = 4 : 3. Shell thick walled, with
irregular, roundish pores, about as broad as the bars; ten to twelve pores
on the half equator. Surface spiny. Length of the conical straight spines
increasing towards the poles; each polar spine surrounded by a circumpolar
group of larger, somewhat curved spines. The larger polar spine equals in
length the major axis; the smaller is scarcely half as long. (The figure,
given by Bury, is not quite exact; there the spines are situated in the
pores, instead of between them. In my specimen the polar spines were not
branched.)

{303}_Dimensions._--Longer axis of the shell 0.12, shorter axis 0.09; pores
and bars 0.004 to 0.006; length of the polar spines--longer 0.12, shorter
0.06.

_Habitat._--Fossil in Barbados.

4. _Lithomespilus flammabundus_, n. sp. (Pl. 14, fig. 14).

Proportion of the major axis to the minor = 4 : 3. Shell thin walled, with
irregular, roundish pores, partly simple, partly composed of three to six
confluent pores; only six to eight pores on the half equator, twice to four
times as broad as the bars. Surface spiny. Length of the conical irregular
spines increasing towards the poles; each polar spine surrounded by a
flame-shaped, circumpolar area of longer spines; all large spines (also the
polar spines) curved or contorted at one pole and much stronger and more
numerous than at the other; length variable, often equal to the longer
axis.

_Dimensions._--Longer axis of the shell 0.12, shorter axis 0.09; pores
0.005 to 0.015, bars 0.003 to 0.005; length of the polar spines 0.1 to
0.15.

_Habitat._--Western part of the Tropical Atlantic, Station 347, depth 2250
fathoms.

Genus 129. _Lithapium_,[168] n. gen.

_Definition._--#Ellipsida# with simple ellipsoidal or pear-shaped shell;
with a single spine only situated at one pole of the main axis.

The genus _Lithapium_ represents a peculiar modification of
_Ellipsoxiphus_; one of the two opposite polar spines disappears by
reduction, and in this way only a single spine remains, at one pole of the
main axis. For this reason the shell assumes a characteristic pear-shape,
and may easily be confounded with some similar Monocyrtida (_Halicapsa_).

1. _Lithapium pyriforme_, n. sp. (Pl. 14, fig. 9).

Proportion of the longer axis to the shorter = 6 : 5. Shell thin walled,
with regular, circular pores, four times as broad as the bars; six to eight
on the half equator. Surface a little thorny. The single polar spines
three-sided pyramidal, as broad at the base as one mesh, about as long as
the radius of the shell. (In the specimen figured, there was on the
opposite pole a little rudiment of the other lost polar spine; it is
missing in other specimens.)

_Dimensions._--Major axis of the ellipsoidal shell 0.12, minor axis 0.1;
pores 0.02, bars 0.005; length of the single polar spine 0.05, basal
thickness 0.02.

_Habitat._--Central area of the Pacific, Station 266, depth 2750 fathoms.

2. _Lithapium halicapsa_, n. sp. (Pl. 14, fig. 8).

Proportion of the longer axis to the shorter = 6 : 5. Shell thin walled,
with irregular, lobed meshes, six to eight on the half equator, twice to
five times as broad as the bars; each mesh {304}composed of two to five
confluent roundish pores. Surface a little thorny. The single polar spine
pyramidal, as broad at the base as one mesh, one-third as long as the axis.

_Dimensions._--Major axis 0.12, minor axis 0.1; pores 0.01 to 0.02, bars
0.002 to 0.006; length of the single polar spine 0.03, basal thickness
0.02.

_Habitat._--Central area of the Pacific, Station 270, depth 2925 fathoms.

3. _Lithapium monocyrtis_, n. sp. (Pl. 14, fig. 10).

Proportion of the longer axis to the shorter = 7 : 6. Shell thick walled,
with irregular, lobed meshes, four to six on the half equator, three to six
times as broad as the bars; each mesh composed of three to six confluent
roundish pores. Surface quite smooth, without thorns. The single polar
spine pyramidal, as broad at the base as the largest mesh, half as long as
the radius. (This species is closely allied to the foregoing, but differs
in the smooth surface and the larger meshes.)

_Dimensions._--Major axis 0.15, minor axis 0.13; pores 0.01 to 0.03, bars
0.003 to 0.01; length of the single spine 0.04, basal thickness 0.03.

_Habitat._--Central area of the Pacific, Station 268, depth 2900 fathoms.

Genus 130. _Pipettella_,[169] n. gen.

_Definition._--#Ellipsida# with simple ellipsoidal shell, the main axis of
which is prolonged at the pole into two opposite hollow fenestrated tubes
of equal size and similar form.

The genus _Pipettella_ is distinguished from the other Ellipsida by two
hollow perforated tubes, which are directed in the longer axis of the
ellipsoidal shell and arise from opposite poles of this axis. It may be
derived from _Cenellipsis_ by prolongation of both poles of the main axis.
As the same peculiar production of two opposite latticed tubuli at the
poles of the main axis obtains in nearly all families of #Prunoidea#
(_Pipetta_, _Cannartus_, _Panarium_, _Zygartus_, &c.), it may possess a
peculiar value in this group.

1. _Pipettella fusiformis_, n. sp.

Shell spindle-shaped, thin walled, the two opposite tubes being conical and
not longer than the short transverse axis of the ellipsoidal shell; no
sharp demarcation between them. The two axes of the ellipsoid bear the
proportion of 3 : 2. Network of the shell and of the tubes equal, delicate,
regular, hexagonal, with circular apertures of equal size; fourteen to
sixteen on the half equator of the shell. Bars very thin. The shell of this
species is similar to that of _Cannartiscus amphiconiscus_ (Pl. 39, fig.
19), but possesses no equatorial stricture and no medullary shell.

_Dimensions._--Longer axis of the ellipsoid 0.18, shorter axis 0.12; length
of the tubes 0.1, their basal breadth 0.05; pores of the network 0.01, bars
0.001.

_Habitat._--Northern Pacific, Station 244, depth 2900 fathoms.

{305}2. _Pipettella tubulosa_, n. sp.

Shell ellipsoidal, thin walled, distinctly separated from the two opposite
tubes, which are cylindrical, longer than the main axis of the ellipsoid,
and one-sixth as broad as the shorter axis. The longer axis of the
ellipsoid bears to the shorter the proportion of 5 : 4. Network of the
shell and of the tubes equal, regular, with very small circular pores,
about as broad as the bars; sixteen to eighteen pores on the half equator
of the shell.

_Dimensions._--Longer axis of the ellipsoid 0.15, shorter axis 0.12; length
of the tubes 0.16 to 0.2, breadth of them 0.02; pores of the network 0.003,
bars 0.003.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

3. _Pipettella elongata_, n. sp.

Shell ellipsoidal, thick walled, distinctly separated from both opposite
tubes, which are cylindrical, much prolonged, twice to three times as long
as the main axis of the ellipsoid, and one-fifth as broad as the shorter
axis. Both axes of the ellipsoid bear the proportion of 3 : 2. Network of
the shell and of the tubes irregular with small rounded pores of different
size, separated by broader bars (often twice to three times as broad), four
to six pores on the half equator of the shell. (This species somewhat
recalls _Solenosphæra serpentina_, Pl. 7, fig. 7; but the tubes are
straight, not contorted.)

_Dimensions._--Longer axis of the ellipsoid 0.2, shorter axis 0.14; length
of the tubes 0.4 to 0.5, breadth 0.03; pores of the network 0.001 to 0.002,
bars 0.003 to 0.004.

_Habitat._--Central area of the Pacific, Station 271, depth 2425 fathoms.

4. _Pipettella prismatica_, n. sp. (Pl. 39, fig. 6).

Shell ellipsoidal, thick walled, distinctly marked off from the two
opposite tubes, which are longer than its main axis and as broad as
one-fifth of it; they are nearly four-sided prismatic, with four strong
ribs or edges; these are directed parallel to the main axis, in two
meridian planes, perpendicular to one another. The wall of the shell is
thickened in the equatorial plane, so as to form a slight stricture on the
inside, separating its two halves. Both axes of the ellipsoid bear the
proportion of 7 : 6. Network regular, with circular pores, somewhat broader
than the bars. The meshes of the shell (fourteen to sixteen on the half
equator) are twice as great as those of the tubes, which are arranged in
two longitudinal rows between every two ribs (there being eight
longitudinal rows on the whole tube). This species is very remarkable for
the rudimentary internal equatorial stricture of the shell (transition to
the genus _Cannartus_, Pl. 39, fig. 10), and by the four edges of the
tubes, which indicate two of the dimensive axes, the third being
represented by the main axis.

_Dimensions._--Longer axis of the ellipsoid 0.14, shorter axis 0.12; length
of the tubes 0.15 to 0.16, breadth 0.03; pores of the shell 0.008 to 0.01,
pores of the tubes 0.004.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

{306}Family XII. #DRUPPULIDA#, Haeckel, 1882 (Pls. 15, 16, 17, 39).

_Definition._--#Prunoidea# with ellipsoidal, latticed (not spongy) shell,
composed of two or more concentric shells; a simple or double cortical
shell enclosing one or two internal concentric shells (medullary shells),
without equatorial stricture. Central capsule ellipsoidal or cylindrical,
without annular equatorial constriction.

The family #Druppulida# differs from the Ellipsida only in the possession
of a simple or double medullary shell, which is enclosed in the centre of
the central capsule, and connected with the ellipsoidal cortical shell
(lying outside it) by radial beams, perforating the membrane of the
capsule. The Druppulida exhibit therefore the same relation to the
Ellipsida that the Dyosphærida among the #Sphæroidea# bear to the
Monosphærida. The cortical shell may be simple or multiple. The whole
fenestrated shell is thus composed of a variable number (two at least) of
concentric shells, which are connected by radial beams.

_The Medullary Shell_--enclosed in the centre of the central capsule--is
either simple or double, and composed of two small concentric shells. Their
form is either spherical, or ellipsoidal, or lenticular. If the medullary
shell be ellipsoidal, the main axis of the ellipsoid is the same as in the
cortical shell. Sometimes the inner medullary shell is spherical, the outer
ellipsoidal or lenticular. If the medullary shell be lenticular (arising
from both poles of the main axis) its vertical axis is also identical with
that of the cortical shell (Pl. 39, fig. 5).

_The Radial Beams_, which connect the medullary and cortical shell, are
either developed in all possible directions (Pls. 16, 17), or limited to
the equatorial plane, more rarely to the meridional plane. Sometimes the
connection is produced only by two opposite beams which lie in the minor or
equatorial axis (Pl. 39, figs. 3, 7, 8); more rarely in the major or
meridional axis (Pl. 17, figs. 7, 8).

_The Cortical Shell_ is constantly ellipsoidal, rarely with modifications,
similar to those which appear in the simple shell of some Ellipsida. As a
rule it is simple, rarely composed of two or more (sometimes six or more)
concentric ellipsoidal shells (in _Cromyodruppa_ and _Cromyocarpus_). The
outer surface is commonly smooth, more rarely covered with radial spines
(in _Druppocarpus_, _Prunocarpus_, &c.). In the majority peculiar polar
appendages are developed at both poles of the main axis, these being
prolongations of them, either in the form of hollow fenestrated tubes
(_Pipetta_, _Pipettaria_, Pl. 39, figs. 7, 8), or solid strong spines. The
two polar spines are either equal in size and similar in form (as in
_Lithatractus_, _Stylatractus_) or unequal (as in _Druppatractus_,
_Xiphatractus_, Pls. 16, 17).

The most primitive of all Druppulida is _Druppula_, with simple medullary
shell and simple cortical shell (Pl. 39, fig. 3); _Prunulum_ differs from
it only in the possession {307}of a double medullary shell (Pl. 39, fig.
4). From these two genera all other forms of the subfamily may be derived.

_The Central Capsule_ of the Druppulida is constantly ellipsoidal, larger
than the concentric enclosed medullary shells, smaller than the surrounding
cortical shell; it is separated from the inner surface of the latter by a
thinner or thicker jelly-mantle.

_Synopsis of the Genera of the Druppulida._

-------------------------------------------------------------------------
Cortical shell without polar appendages (neither solid spines nor hollow
tubes at the poles of the main axis).
-------------------------------------------------------------------------
{ {Surface
{Medullary { smooth, 131. _Druppula_.
{ shell simple.{
{ {Surface
Cortical shell { { spiny, 132. _Druppocarpus_.
simple, {
ellipsoidal. { {Surface
{Medullary { smooth, 133. _Prunulum_.
{ shell double.{
{ {Surface
{ { spiny, 134. _Prunocarpus_.

Cortical shell }Medullary {Surface
composed of } shell { smooth, 135. _Cromyodruppa_.
two or more } double. {
concentric } {Surface
shells. } { spiny, 136. _Cromyocarpus_.
-------------------------------------------------------------------------
Two opposite solid spines, arising from the poles of the main axis.
-------------------------------------------------------------------------
{ {Both polar
{Medullary { spines equal, 137. _Lithatractus_.
{ shell {
{ simple. {Both polar
Cortical { { spines
shell { { unequal, 138. _Druppatractus_.
simple, {
ellipsoidal.{ {Both polar
{ { spines equal, 139. _Stylatractus_.
{Medullary {
{ shell {Both polar
{ double. { spines
{ { unequal, 140. _Xiphatractus_.

Cortical shell composed of two or more
concentric shells; medullary shell
double; both polar spines equal, 141. _Cromyatractus_.
-------------------------------------------------------------------------
Two opposite hollow fenestrated tubes, arising from the poles of the
main axis.
-------------------------------------------------------------------------
Medullary shell simple, 142. _Pipetta_.

Medullary shell double, 143. _Pipettaria_.

Genus 131. _Druppula_,[170] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal, cortical shell, and
simple medullary shell, without spines or polar tubes.

The genus _Druppula_, as the simplest form of the Druppulida, may be
regarded as the common ancestral form of this subfamily. It may be derived
phylogenetically either {308}from _Carposphæra_ by prolongation of one
axis, or from _Cenellipsis_ by duplication of the fenestrated shell. The
outer (or cortical) shell is always more or less ellipsoidal; the inner (or
medullary) shell also is sometimes ellipsoidal, sometimes spherical. Both
shells are concentric, connected by a variable number of radial beams.
Compare _Haliomma oblongum_, Harting, 1863, L. N. 18, p. 15, pl. 2, fig.
42.

Subgenus 1. _Druppuletta_, Haeckel.

_Definition._--Pores of the cortical shell regular, hexagonal, or circular.

1. _Druppula drupa_, n. sp.

Cortical shell thin walled, with smooth surface and regular network; both
its axes bear the proportion of 4 : 3. Pores regular, hexagonal, three
times as broad as the bars; twelve to fifteen on the half equator.
Medullary shell spherical, about one-third as broad as the cortical shell.

_Dimensions._--Major axis of the ellipsoidal cortical shell 0.1 to 0.12,
minor axis 0.08 to 0.09; pores 0.06, bars 0.02; diameter of the medullary
shell 0.03.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms; also
fossil in Barbados.

2. _Druppula pandanus_, n. sp. (Pl. 39, fig. 3).

Cortical shell thin walled, somewhat rough, with regular network; both its
axes bear the proportion of 3 : 2. Pores subregular, circular, hexagonally
framed, one and a half times as broad as the elevated bars; ten to twelve
on the half equator. Medullary shell spherical, one-fourth as broad as the
cortical shell.

_Dimensions._--Major axis of the cortical shell 0.16 to 0.17, minor axis
0.11 to 0.12; pores 0.01, bars 0.007; diameter of the medullary shell
0.035.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Druppula cocos_, n. sp.

Cortical shell thick walled, somewhat rough, with regular network;
proportion of both axes = 6 : 5. Pores regular, circular, hexagonally
framed, quite as broad as the elevated bars; fourteen to sixteen on the
half equator. Medullary shell ellipsoidal (with longitudinal main axis),
nearly half as large as the cortical shell.

_Dimensions._--Major axis of the cortical shell 0.07 to 0.08, minor axis
0.6 to 0.65; pores and bars 0.003 to 0.004; diameter of the medullary shell
0.03 and 0.035.

_Habitat._--Western Tropical Pacific, Station 222, surface, Indian Ocean,
Ceylon, Haeckel.

4. _Druppula phoenix_, n. sp.

Cortical shell thick walled, smooth, with regular network; proportion of
both axes = 5 : 4. Pores regular, circular, hexagonally lobulate (in the
same manner as in _Stauroxiphus gladius_, {309}Pl. 15, fig. 7), three times
as broad as the bars; ten to twelve on the half equator. Medullary shell
ellipsoidal (with longitudinal main axis), one-third as broad as the
cortical shell.

_Dimensions._--Major axis of the cortical shell 0.1, minor axis 0.08; pores
0.009, bars 0.003; axis of the medullary shell 0.03 and 0.025.

_Habitat._--Mediterranean, Smyrna, Haeckel, surface.

5. _Druppula areca_, n. sp.

Cortical shell thin walled, with smooth surface and regular network; both
its axes bear the proportion of 4 : 3. Pores regular, circular, four times
as broad as the bars; fifteen to eighteen on the half equator. Medullary
shell spherical, half as broad as the cortical shell.

_Dimensions._--Major axis of the ellipsoidal cortical shell 0.12 to 0.14,
minor axis 0.1 to 0.11; pores 0.008, bars 0.002; diameter of the medullary
shell 0.05 to 0.06.

_Habitat._--Indian Ocean, off Bombay, Haeckel.

6. _Druppula ovata_, Haeckel.

_Haliomma ovatum_, Ehrenberg, 1854, Mikrogeol., pl. xx. I. fig. 20.

_Haliomma ovatum_, Haeckel, 1862, Monogr. d. Radiol., p. 432.

Cortical shell thin walled, with smooth surface and regular network;
proportion of both axes = 3 : 2. Pores regular, circular, three times as
broad as the bars; nine to ten on the half equator. Medullary shell
spherical, one-third as broad as the cortical shell.

_Dimensions._--Major axis 0.08 to 0.1, minor axis 0.05 to 0.06; pores
0.003, bars 0.001; diameter of the medullary shell 0.02.

_Habitat._--Fossil in the Tertiary rocks of the Mediterranean coast, Greece
(Zante), Ehrenberg; Sicily (Caltanisetta), Haeckel.

7. _Druppula caryota_, n. sp.

Cortical shell thick walled, with rough surface and regular network;
proportion of both axes = 5 : 4. Pores regular, circular, twice as broad as
the bars; sixteen to twenty on the half equator. Medullary shell
ellipsoidal, one-third as large as the cortical shell.

_Dimensions._--Major axis 0.2 to 0.22, minor 0.16 to 0.18, pores 0.008,
bars 0.004; axes of the medullary shell 0.07 and 0.055.

_Habitat._--Pacific, central area, Stations 266 to 268, depth 2700 to 2900
fathoms.

Subgenus 2. _Druppulissa_, Haeckel.

_Definition._--Pores of the cortical shell irregular, of different form or
size, usually subcircular or roundish, sometimes lobed or compound.

{310}8. _Druppula nucula_, n. sp.

Cortical shell thin walled, smooth, with irregular network; its two axes
bear the proportion 3 : 2. Pores subcircular or irregular, roundish, twice
to four times as broad as the bars; fifteen to twenty on the half equator.
Medullary shell spherical, about one-third as broad as the cortical shell.

_Dimensions._--Major axis of the cortical shell 0.15 to 0.2, minor axis 0.1
to 0.14; pores 0.006 to 0.012, bars 0.03; diameter of the medullary shell
0.04.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

9. _Druppula elliptica_, Haeckel.

_Haliomma ellipticum_, Stöhr, 1880, Palæontogr., _loc. cit._, p. 88, Taf.
i. fig. 11.

Cortical shell thin walled, rough, or thorny, with irregular network;
proportion of the axes = 3 : 2. Pores subcircular or irregular, roundish,
about as broad as the bars; ten to twelve on the half equator. Medullary
shell half as broad as the cortical shell, irregularly polyhedral (with
crooked beams in its interior).

_Dimensions._--Major axis 0.15, minor 0.06; pores and bars 0.006; diameter
of the medullary shell 0.03.

_Habitat._--Fossil in the Tertiary rocks of Sicily: Grotte (Stöhr),
Caltanisetta (Haeckel).

10. _Druppula prunum_, n. sp.

Cortical shell thick walled, smooth, with irregular network; proportion of
the axes = 5 : 4. Pores subcircular or irregular, roundish, three to four
times as broad as the bars; seven to eight on the half equator; every pore
is closed at the bottom by a thin membrane, perforated by four to six
irregular pores (like _Stylatractus sethoporus_, Pl. 17, fig. 3). Medullary
shell spherical, one-third as broad as the cortical shell.

_Dimensions._--Major axis 0.2, minor axis 0.16; pores 0.03, bars 0.01;
small enclosed porules 0.01; diameter of the medullary shell 0.05.

_Habitat._--Mediterranean, surface, Portofino near Genoa, Haeckel.

11. _Druppula oliva_, n. sp.

Cortical shell thick walled, rough, with irregular network; proportion of
the axes = 5 : 4. Pores irregular, roundish, three to four times as broad
as the bars; lobed or composed of several conjugated porules (as in
_Lithapium halicapsa_, Pl. 14, fig. 8); five to six large pores on the half
equator. Medullary shell ellipsoidal, about one-third as broad as the
cortical shell.

_Dimensions._--Major axis of the cortical shell 0.15, minor 0.12; pores
0.02 to 0.03, bars 0.007; diameter of the medullary shell 0.04.

_Habitat._--Mediterranean (Corfu), Canary Islands (Lanzerote), Haeckel.

{311}Genus 132. _Druppocarpus_,[171] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
simple medullary shell, with numerous radial spines, without polar tubes.

The genus _Druppocarpus_ differs from _Druppula_ only in the radial spines,
which arise between the pores of the cortical shell, and therefore exhibits
the same relation to it that _Ellipsidium_ bears to _Cenellipsis_, or
_Prunocarpus_ to _Prunulum_.

Subgenus 1. _Druppocarpetta_, Haeckel.

_Definition._--Network of the cortical shell regular, with meshes of equal
size and similar form.

1. _Druppocarpus ananassa_, n. sp.

Cortical shell thin walled, with regular, circular, hexagonally framed
pores, of about the same breadth as the elevated bars; fourteen to sixteen
on the half equator. From each hexagon-corner (between every three pores)
arises a short thin radial spine, about as long as two pores, three-sided
pyramidal at the base. Proportion of both cortical axes = 4 : 3. Medullary
shell spherical, half as broad as the cortical shell. (Greatly resembles
_Druppula pandanus_, Pl. 39, fig. 3, but differs in the larger medullary
shell and the spines on the surface.)

_Dimensions._--Major axis of the ellipsoidal cortical shell 0.16, minor
0.12; pores and bars 0.006; length of the radial spines 0.02; diameter of
the medullary shell 0.06.

_Habitat._--South Pacific, Station 289, surface.

2. _Druppocarpus castanea_, n. sp.

Cortical shell thin walled, with regular circular pores (without hexagonal
frames), of about the same breadth as the bars; twelve to fourteen on the
half equator. Between every three meshes arise a short conical spine with
bristle-like apex, twice to four times as long as one pore. Proportion of
both cortical axes = 3 : 2. Medullary shell spherical, one-third as broad
as the cortical. (Resembles very closely the spherical _Haliomma castanea_,
Haeckel, figured in my Monograph. pl. xxiv. fig. 4.)

_Dimensions._--Major axis 0.15, minor axis 0.1; pores and bars 0.005;
length of the radial spines 0.01 to 0.02; medullary shell 0.035.

_Habitat._--North Atlantic, surface; Canary Islands, Lanzerote, Haeckel.

Subgenus 2. _Druppocarpissa_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
different size or form.

{312}3. _Druppocarpus chamaerops_, n. sp.

Cortical shell thin walled, with irregular, roundish, or subcircular pores
of very different size, twice to six times as broad as the thin bars; eight
to twelve on the half equator. Between them arise numerous thin,
bristle-like radial spines, about half as long as the equatorial axis, and
equal to the diameter of the spherical medullary shell; pores of the latter
subregular, circular, very small. (Resembles _Prunocarpus artocarpus_, Pl.
39, fig. 5, but differs in the simple spherical medullary shell.)

_Dimensions._--Major axis 0.12, minor 0.09; meshes 0.005 to 0.02, bars
0.003; length of the radial spines 0.05; diameter of the medullary shell
0.04.

_Habitat._--Mediterranean, in the Strait of Gibraltar, Algesiras, Haeckel,
surface.

4. _Druppocarpus borassus_, n. sp.

Cortical shell thick walled, with irregular, roundish, or subcircular
pores, three to five times as broad as the bars; twelve to fifteen on the
half equator. Irregularly scattered on the surface about twenty to thirty
short conical spines; their length equals their basal breadth and the
diameter of the largest pores. Medullary shell ellipsoidal, half as large
as the cortical.

_Dimensions._--Major axis 0.1, minor 0.08; pores 0.006 to 0.01, bars 0.002;
length and thickness of the radial spines 0.01; axes of the medullary shell
0.05 and 0.04.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

5. _Druppocarpus corypha_, n. sp.

Cortical shell thick walled, with irregular, funnel-like, roundish pores,
scarcely as broad as the bars; sixteen to twenty on the half equator.
Irregularly scattered on the surface about fifteen to twenty three-sided
pyramidal radial spines, half as long as the equatorial axis, and as the
diameter of the medullary shell.

_Dimensions._--Major axis 0.17, minor 0.14; pores and bars 0.003 to 0.009;
length of the radial spines 0.08; diameter of the medullary shell 0.09.

_Habitat._--Fossil in the Tertiary rocks of the Nicobars, Haeckel.

Genus 133. _Prunulum_,[172] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
double medullary shell; without spines or polar tubes.

The genus _Prunulum_ differs from _Druppula_ in the double (not simple)
medullary shell, which is sometimes spherical, sometimes ellipsoidal; it
may be derived either from _Druppula_ by duplication of the medullary
shell, or from _Thecosphæra_ by prolongation of one axis.

{313}Subgenus 1. _Prunuletta_, Haeckel.

_Definition._--Network of the cortical shell regular, with meshes of equal
size and similar form.

1. _Prunulum frugulum_, n. sp.

Cortical shell thin walled, smooth, with regular, hexagonal pores, three
times as broad as the bars; ten to twelve on the half equator. Proportion
of the major axis of the ellipsoid to the minor = 4 : 3. Both medullary
shells spherical. (Differs from _Druppula drupa_ almost entirely in the
double medullary shell.)

_Dimensions._--Major axis of the ellipsoidal cortical shell 0.12, minor
0.09; pores 0.006, bars 0.002; diameter of both medullary shells 0.05 and
0.03.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Prunulum cerasum_, n. sp.

Cortical shell thick walled, smooth, with circular, hexagonally framed
pores, of the same breadth as the bars; sixteen to eighteen on the half
equator. Proportion of the two axes of the ellipsoid = 5 : 4. Both
medullary shells spherical. (The cortical shell resembles that of _Pipetta
tuba_, Pl. 39, fig. 7, without the polar tubes.)

_Dimensions._--Major axis 0.15, minor 0.12; pores and bars 0.007; diameter
of the medullary shells 0.08 and 0.04.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

3. _Prunulum amygdalum_, n. sp.

Cortical shell thick walled, smooth, with regular, circular, hexagonally
lobulated pores, three times as broad as the bars; twelve to fourteen on
the half equator (of the same form as in _Druppula phoenix_ and in
_Stauroxiphos gladius_, Pl. 15, fig. 7). Proportion of the two axes =
6 : 5. Inner medullary shell spherical, outer ellipsoidal.

_Dimensions._--Major axis 0.12, minor 0.1; pores 0.009, bars 0.003;
diameter of the medullary shells 0.07 and 0.04.

_Habitat._--Mediterranean (Corfu), surface.

4. _Prunulum coccymelium_ (Pl. 39, fig. 4).

Cortical shell thin walled, slightly rough, with regular, circular pores,
twice as broad as the bars; twelve to fifteen on the half equator.
Proportion of both axes = 4 : 3. Both medullary shells spherical.

_Dimensions._--Major axis 0.12, minor 0.09; pores 0.006, bars 0.003;
diameter of the medullary shells 0.06 and 0.03.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

{314}5. _Prunulum crenatum_, Haeckel.

? _Haliomma crenatum_, Ehrenberg, 1854, Mikrogeol., Taf. xxii. fig. 36.

_Actinomma crenatum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 94, Taf.
iii. fig. 3.

_Caryolithis crenata_, Ehrenberg, 1847, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 43.

Cortical shell thick walled, slightly rough or thorny, with small, regular,
circular pores, of the same breadth as the bars; sixteen to eighteen on the
half equator. Proportion of the two axes = 4 : 3. Both medullary shells
ellipsoidal. (The figure of Stöhr represents exactly the fossil form, as I
have observed it myself in the Caltanisetta rocks, whilst the figure of
Ehrenberg is inaccurate and doubtful. The same form, somewhat variable in
size and in the number of the pores, I have also observed in the Pacific
ooze.)

_Dimensions._--Major axis 0.14 to 0.17, minor 0.1 to 0.13; pores and bars
0.006; main axes of the medullary shells 0.09 and 0.03.

_Habitat._--Fossil in the Tertiary rocks of Sicily (Grotte and
Caltanisetta); living in the Central Pacific, Station 268, depth 2900
fathoms.

6. _Prunulum triplex_, Haeckel.

_Haliomma triplex_, Ehrenberg, 1854, Microgeol., Taf. xxxvB. fig. Biv.,
q.

_Actinomma triplex_, Haeckel, 1862, Monogr. d. Radiol., p. 444.

Cortical shell thin walled, covered with numerous very thin, short,
bristle-like spines. Pores small, regular, circular, of the same breadth as
the bars; fourteen to sixteen on the half equator. Proportion of the two
axes = 3 : 2. Both medullary shells ellipsoidal. The description of
Ehrenberg--as is very often the case--is quite incongruent with his figure.
From a combination of both I give here the diagnosis of a deep-sea species,
which is possibly identical with it. The velvet-like covering of very short
and thin bristles is peculiarly characteristic of this species.

_Dimensions._--Major axis 0.1, minor 0.065; pores and bars 0.004; main axes
of the medullary shells 0.04 and 0.02.

_Habitat._--North Atlantic, 1800 fathoms, Ehrenberg; Station 353, depth
2965 fathoms.

Subgenus 2. _Prunulissa_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
different form or size (usually roundish, but sometimes lobed or compound).

7. _Prunulum persicum_, n. sp.

Cortical shell thick walled, with smooth surface and irregular, roundish,
double-contoured pores, twice to four times as broad as the bars; eight to
ten on the half equator. Some of the pores are simple, often subcircular,
others lobed, i.e., composed of from two to three confluent pores as in
_Amphisphæra pluto_ (Pl. 17, fig. 7). Proportion of the two axes = 3 : 2.
Both medullary shells ellipsoidal (or the inner spherical).

{315}_Dimensions._--Major axis 0.12, minor 0.08; pores 0.06 to 0.012, bars
0.003; main axes of the medullary shells 0.05 and 0.025.

_Habitat._--Indian Ocean, between Socotra and Ceylon, surface, Haeckel.

8. _Prunulum fenestratum_, Haeckel.

_Actinomma fenestratum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 94, Taf.
iii. fig. 2.

Cortical shell thick walled, rough, with irregular, roundish pores, once to
three times as broad as the bars; twelve to fourteen on the half equator.
Proportion of the two axes = 9 : 8. Both medullary shells ellipsoidal. (The
pores in Stöhr's description are by mistake called "regular"; in the figure
they are very irregular, as also in the fossil specimens observed by
myself. Between the cortical pores arise very short irregular thorns.)

_Dimensions._--Major axis 0.12 to 0.15, minor 0.1 to 0.12; pores 0.003 to
0.01, bars 0.003; main axis of the outer medullary shell 0.07 to 0.09, of
the inner 0.03 to 0.04.

_Habitat._--Fossil in the Tertiary rocks of Sicily; Grotte, Stöhr,
Caltanisetta, Haeckel.

9. _Prunulum pyrenium_, n. sp.

Cortical shell very thick walled, smooth, with large irregular, roundish
pores, three to four times as broad as the bars; six to eight on the half
equator. The bottom of each funnel-like pore is closed by a thin siliceous
membrane, perforated by three to five irregular roundish pores. Proportion
of the two axes = 4 : 3. Both medullary shells ellipsoidal. (Closely
resembles _Stylatractus sethoporus_, Pl. 17, figs. 2, 3, but is devoid of
the polar spines.)

_Dimensions._--Major axis 0.16, minor 0.12; pores 0.015 to 0.02, bars
0.005; porules 0.01; main axes of the medullary shells 0.07 and 0.03.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 134. _Prunocarpus_,[173] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
double medullary shell, with numerous radial spines, but without polar
tubes.

The genus _Prunocarpus_ differs from _Prunulum_ only in the radial spines
of the cortical shell, and exhibits therefore the same relation to it that
_Druppocarpus_ bears to _Druppula_. While in the latter the medullary shell
is simple, in the former it is double.

Subgenus 1. _Prunocarpetta_, Haeckel.

_Definition._--Network of the cortical shell regular, with meshes of equal
size and similar form.

{316}1. _Prunocarpus datura_, n. sp.

Cortical shell thick walled, with regular, circular pores, twice as broad
as the bars; ten to twelve on the half equator. Between every three meshes
arises a short conical spine, twice to three times as long as one pore.
Both medullary shells spherical. (Differs mainly from _Ellipsidium datura_
and from _Druppocarpus castanea_ in the double medullary shell. The outer
network resembles _Haliomma castanea_, figured 1862 in my Monograph, Taf.
xxiv. fig. 4.)

_Dimensions._--Major axis of the ellipsoid 0.16, minor 0.12; pores 0.012,
bars 0.006; length of the radial spines 0.03; diameter of the medullary
shells 0.06 and 0.04.

_Habitat._--North Atlantic, Færöe Channel, John Murray, surface.

2. _Prunocarpus sparganium_, n. sp.

Cortical shell thick walled, with very small, numerous, regular, circular
pores, of the same size as the bars; forty to fifty on the half equator.
Between them over the entire surface occur small conical spines.
Irregularly scattered over the surface ten to twenty larger conical spines,
three to six times as thick at the base as one pore, one-fourth to one-half
as long as the main axis. Both medullary shells ellipsoidal.

_Dimensions._--Major axis of the cortical shell 0.16, of the outer
medullary shell 0.11, of the inner 0.06; equatorial axis of the first shell
0.12, of the second 0.08, of the third 0.04; pores and bars on an average
0.005; length of the spines 0.05 to 0.1, basal breadth 0.02 to 0.03.

_Habitat._--Fossil in the Barbados deposits (Haeckel).

Subgenus 2. _Prunocarpilla_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
different size or form.

3. _Prunocarpus artocarpium_, n. sp. (Pl. 39, fig. 5).

Cortical shell thin walled with irregular, roundish pores of very different
sizes, twice to nine times as broad as the thin bars; ten to fifteen on the
half equator. Between them arise numerous bristle-shaped, radial spines,
with conical base, on an average one-fourth to two-thirds as long as the
equatorial axis. The outer medullary shell, with irregular, roundish pores,
presents a transverse ellipsoid, its main axis lying in the equatorial axis
of the cortical shell, whilst its equatorial axis corresponds to the main
axis of the latter. Inner medullary shell very small, spherical, with very
small pores.

_Dimensions._--Major axis of the cortical shell 0.14, of the outer
medullary shell 0.05; minor axis of the former 0.1, of the latter 0.035;
diameter of the inner medullary shell 0.014; pores of the cortical shell
0.005 to 0.02, bars 0.002; length of the radial spines 0.02 to 0.06.

_Habitat._--Indian Ocean, surface; Ceylon, Belligemma, Haeckel.

{317}4. _Prunocarpus melocactus_, n. sp.

Cortical shell thick walled, with irregular, roundish pores, twice to five
times as broad as the bars; eighteen to twenty-four on the half equator.
Between them arise over the entire surface small conical thorns, not longer
than the largest pores. Irregularly scattered over the surface twenty to
thirty strong, conical, radial spines, about half as long as the main axis,
as broad at the base as a large pore. Both medullary shells ellipsoidal,
their main axis identical with that of the cortical shell.

_Dimensions._--Major axis of the outer shell 0.18, of the middle 0.09, of
the inner 0.05; minor axis of the first 0.14, of the second 0.07, of the
third 0.04; pores of the cortical shell 0.006 to 0.02, bars 0.004; length
of the spines 0.1, basal breadth 0.01.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms; also
fossil in the Tertiary rocks of Sicily; Caltanisetta, Haeckel.

Genus 135. _Cromyodruppa_,[174] n. gen.

_Definition._--#Druppulida# with four or more concentric shells (two
medullary and two or more cortical shells), without spines or polar tubes.

The genus _Cromyodruppa_ is characterised by the multiplication of the
concentric fenestrated shell, which is composed of two medullary shells
(enclosed in the central capsule) and two or more cortical shells (outside
it). The former may be either spherical or ellipsoidal. The latter are
always ellipsoidal, and in this it differs from _Cromyosphæra_. Probably
_Cromyodruppa_ has arisen from _Prunulum_ by secondary apposition of more
cortical envelopes.

Subgenus 1. _Cromyodruppium_, Haeckel.

_Definition._--Shell composed of four concentric shells, two medullary and
two cortical.

1. _Cromyodruppa cepa_, n. sp.

Shell composed of two ellipsoidal, cortical, and two spherical medullary
shells. Proportion of the main axes of the four shells = 1 : 2 : 4 : 5.
Network of all four shells nearly of the same form, subregular, with
circular pores of almost equal size in every shell. The absolute size of
the pores increases from the innermost to the outermost shell, but the
breadth of the bars does not increase in a similar degree. The bars of the
outermost shell are only twice as broad as those of the innermost; but the
pores are three to four times larger. Surface of all four shells smooth.
(The shell greatly resembles that of _Cromyatractus tetraphractus_, Pl. 15,
fig. 2, but is devoid of polar spines; it differs also in the spherical
form of both medullary shells and their relative size.)

{318}_Dimensions._--Main axes of the four shells--(A) innermost 0.035, (B)
second 0.08, (C) third 0.15, (D) outermost 0.2; their equatorial axes--(A)
0.03, (B) 0.08, (C) 0.12, (D) 0.16; pores 0.004 to 0.016, bars 0.002 to
0.004.

_Habitat._--Western Tropical Pacific, Station 198, depth 2150 fathoms.

Subgenus 2. _Caryodruppula_, Haeckel.

_Definition._--Shell composed of five or more concentric shells (two
medullary and three or more cortical).

2. _Cromyodruppa mango_, n. sp.

Shell composed of six ellipsoidal, concentric shells, two medullary and
four cortical. Proportion of their main axes = 2 : 3 : 8 : 11 : 14 : 18.
Network of the two medullary shells and of the innermost cortical shell
subregular, with subcircular pores, about the same breadth as the bars.
Network of the three outer cortical shells more lax, with larger,
irregular, roundish pores, twice to six times as broad as the bars. The
size of the pores and bars increases gradually from the innermost to the
outermost shell. Surface smooth.

_Dimensions._--Main axis of the six shells--(A) innermost 0.04, (B) 0.06,
(C) 0.16, (D) 0.22, (E) 0.28, (F) 0.35; their equatorial axes--(A)
innermost 0.03, (B) 0.05, (C) 0.12, (D) 0.16, (E) 0.2, (F) 0.25; pores
0.003 to 0.02, bars 0.003 to 0.005.

_Habitat._--Indian Ocean, surface; Matura, Ceylon, Haeckel.

Genus 136. _Cromyocarpus_,[175] n. gen.

_Definition._--#Druppulida# with four or more concentric shells (two
medullary and two or more cortical shells), with numerous radial spines,
but without polar tubes.

The genus _Cromyocarpus_ differs from _Cromyodruppa_ only in the
development of numerous large radial spines which start from the outer
surface of the shell. It exhibits therefore the same relation to the latter
that _Prunocarpus_ bears to _Prunulum_, or _Druppocarpus_ to _Druppula_. It
differs from both in the multiplication of the cortical shell.

1. _Cromyocarpus quadrifarius_, n. sp.

Shell composed of four concentric shells, two ellipsoidal cortical shells,
and two spherical medullary shells. Proportion of their main axes =
10 : 8 : 3 : 2. Pores of the two cortical shells irregular, roundish, twice
to four times as broad as the bars. Pores of the two medullary shells
subregular, circular, about the same size as the bars. Outer surface
covered with numerous conical radial spines, about half as long as the main
axis, half as broad at the base as the innermost shell.

{319}_Dimensions._--Major axis of the outer cortical shell 0.2, of the
inner 0.15; equatorial axis of the former 0.16, of the latter 0.12;
diameter of the outer medullary shell 0.06, of the inner 0.04; pores of the
cortical shells 0.01 to 0.02, of the medullary shells 0.004 to 0.008, bars
0.003 to 0.006; length of the radial spines 0.1, basal breadth 0.02.

_Habitat._--Indian Antarctic Ocean, Station 157, depth 1950 fathoms.

Genus 137. _Lithatractus_,[176] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
simple medullary shell, with two large opposite polar spines in the main
axis of equal size and similar form.

The genus _Lithatractus_, rich in common and widely distributed species,
begins the series of those Druppulida which are characterised by peculiar
polar spines at both poles of the main axis. It repeats the formation of
_Stylosphæra_ and _Ellipsostylus_, and differs from the former in the
ellipsoidal form of the cortical shell, from the latter in the possession
of a medullary shell. Formerly all these forms were united in the one genus
_Stylosphæra_ (see above, p. 121).

Subgenus 1. _Lithatractara_, Haeckel.

_Definition._--Network of the outer shell regular or subregular, with
meshes of nearly equal size and similar form; surface smooth, without
thorns or papillæ.

1. _Lithatractus hexagonalis_, n. sp.

Outer shell thin walled, smooth, without thorns or papillæ, with regular
delicate network; the meshes hexagonal, three to four times as broad as the
thin bars; ten to twelve on the half equator. Proportion of the major axis
of the ellipsoid to the minor = 4 : 3. Minor axis three times as large as
that of the inner spherical shell; pores of the latter, small, circular.
Two spines three-sided pyramidal, as long as the radius of the outer shell,
each as broad at its base as one mesh.

_Dimensions._--Longer axis of the ellipsoidal cortical shell 0.16, shorter
axis 0.12; pores 0.011, bars 0.003; diameter of the spherical medullary
shell 0.04; length of the polar spines 0.07; basal breadth 0.01.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

2. _Lithatractus fragilis_, n. sp. (Pl. 16, fig. 3).

_Stylosphæra fragilis_, Haeckel, 1881, Prodromus et Atlas, pl. xvi. fig.
3.

Outer shell thin walled, smooth, without thorns or papillæ, with regular
network; the meshes circular, of equal size, eight to ten times as broad as
the thin bars; six to eight on the half equator. {320}Proportion of the
major axis of the ellipsoid to the minor = 5 : 4. Minor axis once and
one-third as broad as that of the ellipsoidal inner shell, the pores of
which are also circular, but of half the size. Two spines strong,
three-sided pyramidal, acute, as broad as a large mesh, as long as the
equatorial axis. The stout inner prolongations of these form the only
connection between the two shells.

_Dimensions._--Longer axis of the ellipsoidal cortical shell 0.12, shorter
axis 0.1; pores 0.02, bars 0.002; longer axes of the ellipsoidal medullary
shell 0.09, shorter axis 0.07; pores 0.01, bars 0.002; length of the polar
spines 0.1, basal breadth 0.002.

_Habitat._--Central area of the Pacific, Station 272, depth 2600 fathoms.

3. _Lithatractus leptostylus_, n. sp.

Outer shell thin walled, smooth, with regular, circular pores of equal
size, three to four times as broad as the thin bars; ten to twelve on the
half equator. Proportion of the major axis to the minor = 7 : 6. Minor axis
three times as large as that of the inner spherical shell; pores of the
latter half as large. Polar spines cylindrical, blunt, longer than the
major axis, scarcely half as broad as one larger pore.

_Dimensions._--Longer axis of the cortical shell 0.14, shorter axis 0.12;
pores 0.015 to 0.02, bars 0.004; diameter of the medullary shell 0.04;
length of the polar spines 0.15 to 0.2, its thickness 0.01.

_Habitat._--Central area of the Pacific, Stations 270 to 272, depth 2425 to
2925 fathoms.

4. _Lithatractus pachystylus_, n. sp.

Outer shell thick walled, smooth, with regular, circular pores of equal
size; twelve to fifteen on the half equator. Each pore is deep,
funnel-shaped, its outer aperture double the size of the inner, its breadth
about three times that of the high bars. Proportion of the major axis to
the minor = 6 : 5. Major axis double as long as the diameter of the
spherical medullary shell. Polar spines very thick and short, tetrahedral,
one-fourth as long and broad as the major axis.

_Dimensions._--Longer axis of the cortical shell 0.17, shorter axis 0.14;
pores 0.01, bars 0.003; diameter of the medullary shell 0.08; length and
thickness of the polar spines 0.04.

_Habitat._--Central area of the Pacific, Station 271, depth 2425 fathoms.

5. _Lithatractus convallaria_, n. sp.

Outer shell thick walled, smooth, with elegant regular network; the meshes
circular, six-lobed, rosette-like (of the same form as _Stauroxiphos
gladius_, Pl. 15, fig. 7), twice to three times as broad as the bars; six
to eight on the half equator. Proportion of the major axis of the ellipsoid
to the minor = 4 : 3. Minor axis twice as long as the diameter of the
inner spherical shell, which has regular, simple, circular pores of half
the size. Polar spines short and thick, conical, only one-fourth to
one-sixth as long as the minor axis, and quite as thick.

_Dimensions._--Longer axis of the cortical shell 0.16, shorter axis 0.12;
pores 0.016, bars 0.006; diameter of the medullary shell 0.06; length of
the polar spines 0.02 to 0.03, basal thickness the same.

_Habitat._--Central area of the Pacific, Station 268, depth 2900 fathoms.

{321}Subgenus 2. _Lithatractylis_, Haeckel.

_Definition._--Network of the outer shell regular, with meshes of equal
size and similar form; surface thorny or papillose, covered with small
conical spines or tubercles.

6. _Lithatractus echiniscus_, n. sp.

Outer shell thorny, thin walled, with regular network; meshes circular,
with hexagonal frame, four to five times as broad as the thin bars;
eighteen to twenty on the half equator. From every corner between the three
meshes, where three hexagons unite, starts one short, straight, triangular
thorn (as in _Ellipsoxiphus elegans_, Pl. 14, fig. 7). Proportion of the
major axis of the ellipsoid to the minor = 4 : 3. Minor axis twice the
diameter of the inner spherical shell, the pores of which are half as
broad, circular. Polar spines three-sided pyramidal, about as long as the
minor axis, as broad at the base as one hexagonal frame.

_Dimensions._--Longer axis of the cortical shell 0.08, shorter axis = 0.06;
pores 0.004, bars 0.001; length of the polar spines 0.05, basal thickness
0.005; diameter of the inner shell 0.03.

_Habitat._--South Atlantic, off Tristan da Cunha, Station 334, surface.

7. _Lithatractus carduelis_, n. sp.

Outer shell thin walled, thorny, with regular network; meshes circular,
simple, four to six times as broad as the thin bars; ten to twelve on the
half equator. Between every three meshes is one short conical spine.
Proportion of the major axis to the minor = 6 : 5. Minor axis equals three
times the diameter of the inner spherical shell. Polar spines conical,
thick, half as long as the major axis, thicker at the base than one pore.
(This species differs from _Stylatractus carduus_ only by the simple
medullary shell.)

_Dimensions._--Longer axis of the ellipsoid 0.12 to 0.16, shorter axis 0.1
to 0.14; pores 0.02 to 0.03, bars 0.005; length of the polar spines 0.05 to
0.10, basal breadth 0.04; diameter of the medullary shell 0.03 to 0.04.

_Habitat._--Central area of the Pacific, Stations 268 to 274, depth 2350 to
2900 fathoms; also fossil in the rocks of Barbados.

8. _Lithatractus cirsium_, n. sp.

Outer shell thin walled, thorny, with regular network; pores circular,
simple, small, two to three times as broad as the thin bars; eighteen to
twenty-four on the half equator. Proportion of the major axis to the minor
= 4 : 3. Inner shell ellipsoidal, half as large as the outer. Polar spines
cylindrical, blunt, thin, very variable in length (one-fourth to
three-fourths of the major axis, not thicker than a mesh).

_Dimensions._--Longer axis of the ellipsoid 0.07 to 0.09, shorter axis 0.05
to 0.07; pores 0.005 to 0.006, bars 0.002; length of the polar spines 0.02
to 0.06, basal breadth 0.004; length of the inner shell 0.04, breadth 0.03.

_Habitat._--North Pacific, Station 244, surface.

{322}9. _Lithatractus rosetta_, n. sp.

Outer shell thick walled, thorny, with regular network; meshes circular,
funnel-shaped, its outer aperture elegant, rosette-like, with eight to ten
incisions (like _Stylosphæra calliope_, Pl. 16, fig. 6); eight to ten
meshes on the half equator, three to four times as broad as the bars.
Proportion of the longer axis to the shorter = 5 : 4. Shorter axis equals
three times the diameter of the inner spherical shell. Polar spines
three-sided pyramidal, about half as long as the shorter axis, as broad at
the base as a mesh.

_Dimensions._--Longer axis of the cortical shell 0.15, shorter 0.12; pores
0.02, bars 0.005; length of the polar spines 0.05, basal breadth 0.02;
diameter of the inner shell 0.04.

_Habitat._--South-eastern part of the Pacific, Station 302, depth 1450
fathoms.

Subgenus 3. _Lithatractona_, Haeckel.

_Definition._--Network of the outer shell irregular, with meshes of unequal
size or dissimilar form; surface smooth, without thorns or papillæ.

10. _Lithatractus conifer_, n. sp.

Outer shell thin walled, smooth, with irregular, roundish pores, two to
four times as broad as the thin bars; fifteen to twenty on the half
equator. Margin of the pores simple. Proportion of the major axis to the
minor = 3 : 2. Minor axis twice as large as the diameter of the inner
spherical shell, the pores of which are also irregular, roundish, but of
half the size. Polar spines conical, somewhat shorter than the main axis,
on the base two to three times as thick as a large pore.

_Dimensions._--Longer axis of the outer shell 0.15, shorter axis 0.1; pores
0.002 to 0.004, bars 0.001; diameter of the inner shell 0.05; length of the
polar spines 0.12, basal breadth 0.012.

_Habitat._--South Atlantic, Station 325, surface.

11. _Lithatractus lobatus_, n. sp.

Outer shell thick walled, smooth, with irregular, roundish pores, four to
eight times as broad as the bars; six to eight on the half equator. Margin
of the pores lobed, very irregular, bluntly dentate, by five to twenty
slight incisions. Proportion of the major axis to the minor very variable,
between 3 : 2 and 9 : 8. Diameter of the inner shell also variable, between
one-third and one-half of the outer; pores of the former scarcely half the
size of the latter, simple, roundish, or circular. The inner shell is at
some parts quite spherical, at other parts more or less ellipsoidal. Polar
spines conical, very variable in size and form, sometimes in the basal half
triangular; they are sometimes somewhat longer than the main axis, at other
times considerably shorter; their basal breadth is occasionally the same as
that of the largest pores, sometimes, however, scarcely half as large. This
deep-sea species is very common in the central area of the Tropical Pacific
(Stations 266 to 272), and occurs also fossil in the Barbados deposits. It
is interesting from its great variability, and sometimes constitutes a
transitional form to _Stylosphæra_.

{323}_Dimensions._--Longer axis of the outer shell 0.1 to 0.15, shorter
axis 0.05 to 0.12; pores 0.018 to 0.024, bars 0.002 to 0.004; diameter of
the inner shell 0.04 to 0.08; length of the polar spines 0.08 to 0.2, basal
thickness 0.01 to 0.02.

_Habitat._--Central area of the Pacific, common; Stations 266 to 272, depth
2425 to 2900 fathoms; also fossil in Barbados.

12. _Lithatractus jugatus_, n. sp. (Pl. 16, fig. 2).

_Stylosphæra jugata_, Haeckel, 1881, Prodromus et Atlas (pl. xvi. fig.
2).

Outer shell thick walled, smooth, or somewhat reticulated, with a peculiar,
irregular, double network. The pores are roundish, of very different sizes,
with double margin of the outer aperture, and so irregularly distributed in
polygonal groups that every group contains two to six pores immediately
touching each other; the groups are separated by broader bars. On the half
equator of the shell are about six to eight groups and twelve to eighteen
pores. Proportion of the longer axis of the ellipsoidal shell to the
shorter = 6 : 5 or 5 : 4. Inner ellipsoidal shell about half the size; its
pores are not easy to make out. Polar spines three-sided pyramidal, with
prominent edges, about half as long as the major axis, as broad at the base
as a group of pores. (Nearly allied to _Stylosphæra jugata_, p. 137.)

_Dimensions._--Longer axis of the cortical shell 0.12 to 0.16, shorter axis
0.1 to 0.13; pores 0.01 to 0.02; length of the polar spines 0.06 to 0.08,
basal breadth 0.03 to 0.4.

_Habitat._--Central area of the Pacific, Stations 266 to 272, depth 2425 to
2900 fathoms.

Subgenus 4. _Lithatractium_, Haeckel.

_Definition._--Network of the outer shell irregular, with meshes of unequal
size or dissimilar form; surface thorny or papillose, covered with small
conical spines or tubercles.

13. _Lithatractus conostylus_, n. sp.

Outer shell thorny, thin walled, with irregular network; pores roundish, of
unequal size, twice to four times as broad as the thin bars; sixteen to
twenty on the half equator. Proportion of the longer axis to the shorter =
5 : 4. Shorter axis twice the diameter of the inner spherical shell. Polar
spines conical, very stout, longer than the main axis, its base equals
one-third of the minor axis.

_Dimensions._--Major axis of the cortical shell 0.1, shorter axis 0.08;
pores 0.004 to 0.002, bars 0.001; diameter of the medullary shell 0.04;
polar spines--length 0.12, thickness 0.03.

_Habitat._--Central area of the Pacific, Station 271, depth 2425 fathoms.

14. _Lithatractus gamoporus_, n. sp.

Outer shell thorny, thick walled, with irregular network; pores roundish,
of unequal size, so irregularly distributed in polygonal groups that in
every group two to six pores (commonly three to four) are near together;
the groups are separated by broader bars. On the half equator {324}six to
eight groups and eighteen to twenty-four pores. Proportion of the longer
axis to the shorter = 6 : 5. Shorter axis equals twice the diameter of the
inner ellipsoidal shell. Polar spines three-sided pyramidal, about as long
as the equatorial axis, as broad at the base as a group of pores.

_Dimensions._--Major axis of the cortical shell 0.12, minor axis 0.1; pores
0.004 to 0.012, bars 0.003; length of the polar spines 0.1, basal breadth
0.03.

_Habitat._--Central area of the Pacific, Stations 271 to 274, depth 2350 to
2750 fathoms.

Genus 138. _Druppatractus_,[177] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
simple medullary shell, with two large opposite polar spines in the main
axis of different size or dissimilar form.

The genus _Druppatractus_ differs from its near relation _Lithatractus_ in
the differentiation of the two polar spines, which are different in size or
form, often to a very considerable degree. It has therefore the same
relation to the latter that _Ellipsoxiphus_ bears to _Ellipsostylus_.

Subgenus 1. _Druppatractara_, Haeckel.

_Definition._--Network of the cortical shell regular or subregular, with
meshes of nearly equal size and similar form; surface smooth, without
thorns or papillæ.

1. _Druppatractus ichthydium_, n. sp.

Cortical shell thin walled, with smooth surface and regular network.
Proportion of the major axis to the minor = 3 : 2. Pores circular,
hexagonally framed, twice as broad as the bars; seven to eight on the half
equator (as in _Xiphostylus alcedo_, Pl. 13, fig. 4). Medullary shell
spherical, one-third as broad as the cortical shell. Polar spines straight,
three-sided pyramidal, as broad at the base as one mesh; the longer equals
the main axis of the cortical shell, the shorter only its half.

_Dimensions._--Major axis of the ellipsoidal cortical shell 0.18, minor
axis 0.12; pores 0.01, bars 0.005; diameter of the medullary shell 0.04;
length of the major polar spine 0.16 to 0.2, of the minor 0.01 to 0.12,
basal breadth 0.01.

_Habitat._--Mediterranean (Corfu), surface, Haeckel.

2. _Druppatractus hippocampus_, n. sp. (Pl. 16, figs. 10, 11).

_Stylosphæra hippocampus_, Haeckel, 1881, Atlas, pl. xvi. figs. 10, 11.

Cortical shell thick walled, with smooth surface and regular network.
Proportion of both axes = 7 : 6. Pores circular, hexagonally lobed, three
times as broad as the bars; nine to ten on the half {325}equator. Medullary
shell ellipsoidal, nearly half as large as the cortical. Larger polar spine
horn-like curved, as long as the main axis, smaller scarcely half as long,
pommel-shaped.

_Dimensions._--Major axis 0.09, minor 0.075; pores 0.01, bars 0.003; axes
of the medullary shell 0.04 and 0.03; length of the larger polar spine 0.06
to 0.09, of the smaller 0.04, basal breadth 0.02.

_Habitat._--Pacific, central area, Stations 270 to 272, surface.

3. _Druppatractus belone_, n. sp.

Cortical shell thin walled, with smooth surface and regular network.
Proportion of both axes = 4 : 3. Pores circular, twice as broad as the
bars; ten to twelve on the half equator. Medullary shell spherical,
one-third as broad as the cortical shell. Polar spines strong, straight,
cylindrical, with conical apex, as broad at the base as two pores; the
larger once and a half to twice as long as the main axis, the smaller only
one-third to one-fourth as long.

_Dimensions._--Major axis 0.1 to 0.12, minor 0.08 to 0.09; pores 0.008,
bars 0.004; diameter of the medullary shell 0.03; length of the larger
polar spine 0.15 to 0.2, of the minor 0.04 to 0.05, basal thickness of them
0.02.

_Habitat._--Cosmopolitan; Atlantic, Indian, and Pacific, surface.

4. _Druppatractus testudo_, Haeckel.

_Stylosphæra testudo_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 299, Taf. viii. fig. 16.

Cortical shell thick walled, spindle-shaped, with smooth surface and
regular network. Proportion of the two axes = 3 : 2. Pores circular, five
times as broad as the bars; only six to seven on the half equator.
Medullary shell ellipsoidal, about one-third as large as the cortical
shell. Polar spines strong, short, three-sided pyramidal; the larger as
long as half the transverse axis, the smaller only one-fourth as long.

_Dimensions._--Major axis 0.12, minor 0.8; pores 0.015, bars 0.003; axes of
the medullary shell 0.04 and 0.03; length of the larger polar spine 0.04,
of the smaller 0.02, basal thickness 0.01.

_Habitat._--Pacific, Philippine and Californian Sea (Ehrenberg), Stations
244, 266, 289, &c., depth 2550 to 2900 fathoms.

Subgenus 2. _Druppatractylis_, Haeckel.

_Definition._--Network of the cortical shell regular or subregular, with
meshes of nearly equal size and similar form; surface thorny or papillose,
covered with small conical spines or tubercles.

5. _Druppatractus accipenser_, n. sp.

Cortical shell thick walled, thorny, with regular network. Proportion of
the two axes = 6 : 5. Pores circular, each with a six-lobed outer opening,
funnel-shaped, twice as broad as the bars; nine {326}to ten on the half
equator. Medullary shell spherical, one-third as broad as the cortical
shell. Polar spines very strong, three-sided pyramidal, as broad at the
base as three pores; the larger about as long as the main axis, the smaller
only one-third as long.

_Dimensions._--Major axis 0.12, minor 0.1; pores 0.01, bars 0.005;
medullary shell 0.035; length of the larger polar spine 0.11, of the
smaller 0.04, basal breadth 0.03.

_Habitat._--Indian Ocean, Madagascar, surface, Rabbe.

6. _Druppatractus ostracion_, n. sp. (Pl. 16, figs. 8, 9).

Cortical shell thick walled, thorny, with regular network. Proportion of
the two axes = 4 : 3. Pores circular, hexagonally framed, three to four
times as broad as the crested bars; from every corner of the hexagon
(between three meshes) arises a short papilla; the bottom of every
funnel-like mesh is closed by a very thin plate with three regular,
circular pores (occasionally four), sometimes confluent, at other times
separate. Medullary shell (fig. 9) half as large as the cortical,
ellipsoidal and papillate, with regular, circular pores. Polar spines very
strong, three-sided prismatic, often somewhat irregular, with short apex;
the larger once to twice as long as the main axis, the smaller scarcely
half as long.

_Dimensions._--Major axis 0.16, minor 0.12; pores 0.02 (porules at their
base 0.01), bars 0.006; axes of the medullary shell 0.07 and 0.06, its
pores 0.01, bars 0.003; length of the larger polar spine 0.15 to 0.3, of
the smaller 0.07 to 0.09, basal thickness 0.03.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

7. _Druppatractus coronatus_, Haeckel.

_Stylosphæra coronata_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxv. fig. 4.

Cortical shell thick walled, thorny, with regular network. Proportion of
the two axes = 5 : 4. Pores circular, simple, twice as broad as the bars;
eight to ten on the half equator. Medullary shell circular, one-third as
broad as the cortical shell. Polar spines furrowed and angular; the longer
and thinner pyramidal about as long as the main axis, the shorter and
thicker scarcely half as long, thickened towards the short conical apex.

_Dimensions._--Major axis 0.08 to 0.1, minor 0.06 to 0.08; pores 0.01, bars
0.005; diameter of the medullary shell 0.03; length of the large polar
spine 0.08 to 0.12, of the shorter 0.04 to 0.05.

_Habitat._--Fossil in the Tertiary rocks of Barbados; also living in the
depths of the Pacific, central area, Stations 265 to 268, depth 2700 to
2900 fathoms.

Subgenus 3. _Druppatractona_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
unequal size or dissimilar form; surface smooth, without thorns or papillæ.

{327}8. _Druppatractus lævis_, Haeckel.

_Stylosphæra lævis_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxv. fig. 6.

Cortical shell thin walled, smooth, with irregular network. Proportion of
the two axes = 3 : 2. Pores roundish or subcircular, of different size,
one-half to twice as broad as the bars; eight to ten on the half equator.
Medullary shell ellipsoidal, one-third as large as the cortical shell.
Polar spines conical, the larger and thinner nearly as long as the main
axis, the shorter and thicker scarcely one-third as long.

_Dimensions._--Major axis 0.07 to 0.1, minor 0.05 to 0.07; pores and bars
0.005 to 0.01; axes of the medullary shell 0.03 and 0.02; length of the
major polar spine 0.06 to 0.09, of the shorter 0.03 to 0.04.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, surface.

9. _Druppatractus xiphias_, n. sp.

Cortical shell thick walled, smooth, with irregular network. Proportion of
the two axes = 6 : 5. Pores funnel-shaped, composed of two to four
confluent smaller porules, twice to four times as broad as the bars; ten to
twelve on the half equator. Medullary shell spherical, half as broad. Polar
spines three-sided pyramidal, the larger about as long as the main axis,
the shorter one-third to one-half as long.

_Dimensions._--Major axis 0.12, minor 0.1; pores 0.012 to 0.018, bars
0.004; diameter of the medullary shell 0.05; length of the major polar
spine 0.1, of the minor 0.03 to 0.05.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

Subgenus 4. _Druppatractium_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
unequal size or dissimilar form; surface thorny or papillose, covered with
small spines or tubercles.

10. _Druppatractus diodon_, n. sp.

Cortical shell thin walled, thorny, with irregular network. Proportion of
the two axes = 5 : 4. Pores simple, irregular, roundish, twice to four
times as broad as the bars; eight to ten on the half equator. Medullary
shell spherical, half as broad. Polar spines conical, more or less curved;
the major longer than the main axis, the minor scarcely half as long.
(Resembles _Sphærostylus ophidium_, Pl. 16, figs. 14, 15, but differs from
it in the prolongation of the main axis.)

_Dimensions._--Major axis 0.15, minor 0.12; pores 0.01 to 0.02, bars 0.005;
diameter of the medullary shell 0.06; length of the larger polar spine 0.12
to 0.18, of the shorter 0.06 to 0.08, basal thickness 0.003.

_Habitat._--Pacific, central area, Station 274, surface.

{328}11. _Druppatractus pisciculus_, n. sp.

Cortical shell thick walled, spiny, with irregular network. Proportion of
the two axes = 3 : 2. Pores irregular, roundish, lobed, or composed of
three to five confluent porules, twice to six times as broad as the bars;
six to eight on the half equator. (Similar to _Ellipsoxiphus atractus_, Pl.
14, fig. 1.) Medullary shell ellipsoidal, nearly half as large as the
cortical shell. Polar spines three-sided pyramidal, the major longer than
the main axis, the minor scarcely one-third to one-fourth as long.

_Dimensions._--Major axis 0.18, minor 0.12; pores 0.01 to 0.03, bars 0.004;
axes of the medullary shell 0.08 and 0.05; length of the major polar spine
0.2, of the minor 0.05, basal breadth 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Genus 139. _Stylatractus_,[178] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
double medullary shell, in the main axis with two large opposite polar
spines of equal size and similar form.

The genus _Stylatractus_ differs from _Lithatractus_ in the double
medullary shell, from _Amphisphæra_ in the ellipsoidal form of the cortical
shell. As in these two genera, both opposite polar spines have the same
shape and size.

Subgenus 1. _Stylatractara_, Haeckel.

_Definition._--Network of the cortical shell regular or subregular, with
meshes of nearly equal size and similar form; surface smooth, without
thorns or papillæ.

1. _Stylatractus neptunus_, n. sp. (Pl. 17, fig. 6).

_Amphistylus neptunus_, Haeckel, 1878, Atlas, pl. xvii. fig. 6.

Cortical shell thick walled, smooth, with subregular, circular, polygonally
framed pores, quite as broad as the bars; ten to twelve on the half
equator. Polar spines three-sided pyramidal, about as long as the half main
axis, three times as broad at the base as the pores. (Much resembles
_Amphisphæra neptunus_, p. 142, but differs from it by the prolongation of
the main axis, which equals one and a half or one and a fourth the
equatorial axis, and by the somewhat irregular formation of the cortical
hexagonal network. Sometimes also, as in the figured specimen, the length
of both polar spines is somewhat different.)

_Dimensions._--Major axis of the cortical shell 0.15, minor axis 0.12;
pores and bars 0.007; main axes of both ellipsoidal medullary shells 0.08
and 0.05; length of the polar spines 0.08 to 0.1, basal breadth 0.02.

_Habitat._--Pacific, central area, Stations 266 to 268, depth 2700 to 2900
fathoms.

{329}2. _Stylatractus fusiformis_, n. sp.

Cortical shell thick walled, smooth, with regular, simple, circular pores,
quite as broad as the bars; fourteen to sixteen on the half equator. Polar
spines three-sided pyramidal, half as long as the main axis, as broad at
the base as the inner medullary shell. (Resembles very nearly _Xiphatractus
armadillo_, Pl. 17, fig. 11, but differs in the regular form and equal
length of the polar spines.)

_Dimensions._--Major axis 0.17, minor axis 0.13; pores and bars 0.007; main
axes of both ellipsoidal medullary shells 0.09 and 0.05; length of the
polar spines 0.08, basal breadth 0.04.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, at various depths.

3. _Stylatractus compactus_, n. sp. (Pl. 17, fig. 4).

Cortical shell thick walled, smooth, with subregular, circular,
double-contoured pores, smaller than the bars; sixteen to eighteen on the
half equator. The thickness of the shell-wall equals the radius of the
inner medullary shell. Polar spines short, three-sided pyramidal; their
length and basal thickness variable, but commonly equal to the diameter of
the inner medullary shell.

_Dimensions._--Major axis 0.16, minor axis 0.13; pores 0.007, bars 0.01;
main axes of both ellipsoidal medullary shells 0.09 and 0.04; length and
basal breadth of the polar spines 0.04 to 0.05.

_Habitat._--Pacific, central area, Stations 270 to 272, depth 2425 to 2925
fathoms.

Subgenus 2. _Stylatractylis_, Haeckel.

_Definition._--Network of the outer shell regular, with meshes of equal
size and similar form; surface thorny or papillose, covered with small
spinules or tubercles.

4. _Stylatractus giganteus_, n. sp. (Pl. 17, fig. 1).

_Amphistylus giganteus_, Haeckel, 1879, Atlas (pl. xvii. fig. 1).

Cortical shell papillose, very thick walled, with regular network; pores
circular, with double margin, about twice as broad as the bars; ten to
twelve on the half equator. The cortical shell is connected with the outer
medullary shell by numerous strong beams, and the inner prolongations of
both polar spines are much stronger. The circular pores of the outer
medullary shell are three times as large as those of the inner, and equal
to those of the cortical shell, but the bars are much thinner. Polar spines
very strong, three-sided pyramidal, with spirally contorted edges, as long
as the main axis, as broad at the base as the inner medullary shell.

_Dimensions._--Major axis of the cortical shell 0.3, minor axis 0.22; pores
0.02, bars 0.01; main axes of the ellipsoidal medullary shells 0.14 and
0.07; length of the polar spines 0.3, basal thickness 0.06.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

{330}5. _Stylatractus carduus_, Haeckel.

_Stylosphæra carduus_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxv. fig. 7.

Cortical shell thin walled, spiny, with regular, circular pores, five times
as broad as the bars; ten to twelve on the half equator. Polar spines
conical, half as long as the main axis, at the base broader than the pores.
(The cortical shell is ellipsoidal as well as both medullary shells; the
figure of Ehrenberg, as is generally the case, is more correct than his
description. This interesting species occurs in the Barbados rocks, with
double as well as with simple medullary shell, and may in the latter case
be distinguished as _Lithatractus carduelis_.)

_Dimensions._--Major axis of the cortical shell 0.12 to 0.16, minor 0.1 to
0.14; pores 0.01 to 0.02, bars 0.002 to 0.005; main axes of the ellipsoidal
medullary shells 0.05 and 0.03; length of the polar spines 0.06, basal
breadth 0.02.

_Habitat._--Fossil in the Tertiary rocks of Barbados.

Subgenus 3. _Stylatractona_, Haeckel.

_Definition._--Network of the outer shell irregular, with meshes of
different size or form; surface smooth, without thorns or papillæ.

6. _Stylatractus variabilis_, n. sp.

Cortical shell thin walled, smooth, with irregular, roundish pores, twice
to six times as broad as the bars; twelve to sixteen on the half equator.
Polar spines conical, about half as long as the main axis, and as broad at
the base as the inner medullary shell. (The size and form of the pores and
of the polar spines in this species are very variable, so that the
proportions given are to be understood as averages.)

_Dimensions._--Major axis 0.13, minor axis 0.11; pores 0.004 to 0.012, bars
0.002; main axes of the ellipsoidal medullary shells 0.07 and 0.04; length
of the polar spines 0.06, basal breadth 0.03.

_Habitat._--Pacific, central area. Stations 272 to 274, depth 2350 to 2750
fathoms.

7. _Stylatractus sethoporus_, n. sp. (Pl. 17, figs. 2, 3).

Cortical shell thick walled, smooth, with large irregular, roundish pores,
twice to four times as broad as the bars; seven to eight on the half
equator. At the bottom of each pore a thin lamella of silex, perforated by
four to six irregular, roundish, double-contoured porules. Medullary shells
resemble those of _Stylatractus giganteus_. Polar spines three-sided
pyramidal; their length and basal thickness scarcely equal the diameter of
the inner medullary shell.

_Dimensions._--Major axis 0.15, minor axis 0.13; pores 0.02, bars 0.005,
porules 0.01; length of the polar spines 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

{331}Subgenus 4. _Stylatractium_, Haeckel.

_Definition._--Network of the outer shell irregular, with meshes of
different size and form; surface thorny or papillose, covered with small
spinules or tubercles.

8. _Stylatractus papillosus_, n. sp.

Cortical shell thin walled, with papillose surface, and irregular, simple,
roundish pores, ten to sixteen on the half equator, twice to three times as
broad as the bars. Conical papillæ of the surface irregularly scattered.
Polar spines conical, about as long as the main axis, at the base half as
broad as the inner medullary shell.

_Dimensions._--Major axis 0.12, minor 0.1; pores 0.006 to 0.01, bars 0.003;
main axes of the medullary shells 0.09 and 0.06; length of the polar spines
0.13, basal breadth 0.04.

_Habitat._--Indian Ocean, surface (between Socotra and Ceylon), Haeckel.

9. _Stylatractus disetanius_, n. sp.

Cortical shell thin walled, with spiny surface, and irregular, roundish
pores, eight to ten on the half equator, twice to three times as broad as
the bars. Every pore is divided by thinner bars into four to six small
roundish porules. Polar spines three-sided pyramidal, nearly as long as the
main axis, as broad at the base as the inner medullary shell. (Resembles
_Xiphatractus glyptodon_, Pl. 17, figs. 9, 10, but differs from it by the
irregular network and the equal size of both polar spines.)

_Dimensions._--Major axis 0.14, minor 0.1; pores 0.02, bars 0.008, porules
0.007; main axes of the two medullary shells 0.08 and 0.05; length of the
polar spines 0.13, basal breadth 0.04.

_Habitat._--South Pacific, near New Zealand, Station 169, surface.

Genus 140. _Xiphatractus_,[179] n. gen.

_Definition._--#Druppulida# with a simple ellipsoidal cortical shell and
double medullary shell, in the main axis with two large opposite polar
spines of different size or form.

The genus _Xiphatractus_ differs from the nearly related _Stylatractus_ in
the differentiation of two polar spines (in the same manner as
_Druppatractus_ differs from _Lithatractus_). From the spherical
_Amphistylus_ it differs in the ellipsoidal form of the cortical shell.

Subgenus 1. _Xiphatractara_, Haeckel.

_Definition._--Network of the cortical shell regular or subregular, with
meshes of nearly equal size and similar form; surface smooth, without
thorns or papillæ.

{332}1. _Xiphatractus armadillo_, n. sp. (Pl. 17, fig. 11).

Cortical shell thick walled, smooth, with regular, circular pores, about as
broad as the bars; fifteen to sixteen on the half equator. Polar spines
three-sided pyramidal, with thick, prominent edges, of irregular form; the
longer about as long as the main axis, the shorter scarcely one-third as
long; their basal breadth equals the diameter of the inner medullary shell.

_Dimensions._--Major axis of the cortical shell 0.13, minor axis 0.1,
thickness of its wall 0.01; pores and bars 0.005; main axes of the two
medullary shells 0.07 and 0.03; length of the major polar spine 0.12, of
the minor 0.04, basal thickness 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Xiphatractus euphractus_, n. sp.

Cortical shell thick walled, smooth, with regular, circular pores, twice as
broad as the bars; eight to nine on the half equator. Each pore has six to
nine excisions (commonly eight), elegantly lobed, flower-like (as in
_Xiphostylus phasianus_, Pl. 13, fig. 9). Polar spines strong, angulate;
the major pyramidal, longer than the main axis, the shorter elegantly
pommel-like, scarcely one-third as long.

_Dimensions._--Major axis 0.15, minor 0.12; pores 0.014, bars 0.007; main
axes of the two medullary shells 0.09 and 0.05; length of the larger polar
spine 0.16, of the shorter 0.05, greatest thickness 0.03.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Subgenus 2. _Xiphatractylis_, Haeckel.

_Definition._--Network of the cortical shell regular or subregular, with
meshes of nearly equal size and similar form; surface thorny or papillose.

3. _Xiphatractus spinulosus_, Haeckel.

_Stylosphæra spinulosa_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxv. fig. 8.

Cortical shell thin walled, with spiny surface, and large, regular,
circular pores, four times as broad as the bars; seven to eight on the half
equator. Spines between the pores of the same size, conical. Polar spines
also conical, but much larger; the major about as long as the main axis,
the minor scarcely half as long; their basal thickness equals the largest
pores.

_Dimensions._--Major axis 0.1, minor 0.08; pores 0.012, bars 0.003;
diameter of the spherical medullary shells 0.06 and 0.03; length of the
major polar spine 0.09, of the minor 0.04, basal thickness 0.02.

_Habitat._--Fossil in the Tertiary rocks of Barbados.

{333}4. _Xiphatractus sulcatus_, Haeckel.

_Stylosphæra sulcata_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiv, fig. 6.

Cortical shell thick walled, with thorny surface, and regular, circular
pores, quite as broad as the bars; nine to ten on the half equator. Polar
spines cylindrical, with longitudinal basal furrows and conical apex, three
times as broad as the pores; the larger about as long as the main axis, the
shorter only one-half or two-thirds as long.

_Dimensions._--Major axis 0.01, minor 0.08; pores and bars 0.007; main axes
of the two medullary shells 0.08 and 0.04; length of the major polar spine
0.1, of the minor 0.06, breadth of them 0.02.

_Habitat._--Fossil in the Barbados deposits.

5. _Xiphatractus dasypus_, n. sp.

Cortical shell thick walled, with spiny surface, and regular, elegant
network; pores circular, hexagonally framed, about as broad as the bars;
from every corner of the network (between three pores) arises a short
bristle-like spine. Polar spines three-sided prismatic, as broad as one
hexagon; the major once to twice as long as the main axis, the minor only
one-half to one-fourth as long. (The cortical shell much resembles
_Ellipsoxiphus elegans_, Pl. 14, fig. 7.)

_Dimensions._--Major axis 0.15, minor 0.13; pores and bars 0.01; main axes
of the two medullary shells 0.1 and 0.06; length of the major polar spine
0.1 to 0.3, of the minor 0.05 to 0.1, thickness 0.01.

_Habitat._--Tropical Western Pacific, Station 222, surface.

Subgenus 3. _Xiphatractona_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
different size or form; surface smooth, without thorns or papillæ.

6. _Xiphatractus chlamydophorus_, n. sp.

Cortical shell thin walled, smooth, with irregular, roundish pores, twice
to four times as broad as the bars; eight to twelve on the half equator.
Polar spines angular, irregularly curved or contorted; the larger once to
twice as long as the main axis, the shorter and thicker pommel-shaped, only
one-third to one-half as long; their basal breadth equals two pores.

_Dimensions._--Major axis 0.14, minor 0.1; pores 0.006 to 0.012, bars
0.003; main axis of the two ellipsoidal medullary shells 0.09 and 0.04;
length of the major polar spine 0.1 to 0.2, of the minor 0.05 to 0.08,
breadth 0.02.

_Habitat._--Cosmopolitan; Atlantic, Pacific, many Stations, surface.

{334}Subgenus 4. _Xiphatractium_, Haeckel.

_Definition._--Network of the cortical shell irregular, with meshes of
unequal size or dissimilar form; surface thorny or papillose.

7. _Xiphatractus radiosus_, Haeckel.

_Stylosphæra radiosa_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiv. fig. 5.

Cortical shell very thick walled, with thorny surface, and irregular,
roundish pores, once to three times as broad as the bars; nine and ten on
the half equator. Polar spines conical, on the base about as broad as the
largest pores; the larger as long as the main axis, the smaller scarcely
one-fourth as long. (The radial striation, figured by Ehrenberg and applied
to the name of this species, is produced by the contours of the
funnel-shaped pores in the thick walls seen in optical section; the
ellipsoidal cortical shell is double.)

_Dimensions._--Major axis 0.13, minor 0.09; pores 0.003 to 0.01, bars
0.003; main axes of the two medullary shells 0.05 and 0.03; length of the
major polar spine 0.12, of the minor 0.03, basal breadth 0.01.

_Habitat._--Fossil in the Tertiary rocks of Barbados.

8. _Xiphatractus glyptodon_, n. sp. (Pl. 17, figs. 9, 10).

Cortical shell thick walled, covered with radial spines of the size of the
pores; on the half equator eight to ten irregular or subregular roundish
pores, twice to four times as broad as the bars. In the bottom of each pore
a thin lamella of silex, perforated by four to six smaller roundish pores.
Both medullary shells (fig. 10) spherical, with smaller, regular, circular
pores. Larger polar spine three-sided pyramidal, in length equals the main
axis; shorter spine only half as long, but twice as thick, of elegant
pommel-form (fig. 9), with nine prominent edges.

_Dimensions._--Major axis 0.12, minor, 0.1; pores and spines 0.01 to 0.02,
bars and porules 0.005; diameter of the two medullary shells 0.05 and 0.02;
length of the major polar spine 0.1, of the minor 0.06, greatest thickness
0.04.

_Habitat._--Pacific, central area, Station 263, surface.

Genus 141. _Cromyatractus_,[180] n. gen.

_Definition._--#Druppulida# with four or more concentric shells (two
medullary shells and two or more cortical shells), in the main axis with
two large opposite polar spines of equal size and similar form.

The genus _Cromyatractus_ is very nearly allied to all the foregoing
genera, which bear two equal spines opposite to one another on the poles of
the main axis; it differs {335}from them in the multiplication of the
cortical shell, and may be considered as the most highly developed form of
this amphistylous series, which proceeds from _Ellipsoxiphus_ to
_Lithatractus_ and _Stylatractus_.

Subgenus 1. _Cromyatractium_, Haeckel.

_Definition._--Shell composed of two medullary shells and two cortical
shells.

1. _Cromyatractus tetraphractus_, n. sp. (Pl. 15, fig. 2).

_Stylocromium tetraphractum_, Haeckel, 1879, Atlas (pl. xv. fig. 2).

Proportion of the main axes of the four concentric shells about =
1 : 3 : 5 : 7. Network of all four shells nearly of the same structure,
subregular, with circular pores of equal size (in one and the same shell).
The absolute size of the pores increases gradually from the innermost to
the outermost shell. The bars between the pores are smaller and quite
smooth. Surface of the outermost shell smooth. Polar spines cylindrical,
with conical apex, of variable length, nearly as broad as the innermost
shell.

_Dimensions._--Main axes of the four shells--(A) inner medullary shell
0.03, (B) outer medullary shell 0.08, (C) inner cortical shell 0.15, (D)
outer cortical shell 0.2; equatorial axes of them--(A) 0.025, (B) 0.06, (C)
0.12, (D) 0.15; pores of (A) 0.004, (B) 0.007, (C) 0.01, (D) 0.013, bars
0.002 to 0.004; length of the polar spines 0.1 to 0.3 (and more), breadth
0.015.

_Habitat._--Antarctic (Indian) Ocean, Station 157, depth 1950 fathoms.

2. _Cromyatractus tetralepas_, n. sp.

Proportion of the main axes of the four concentric shells about =
1 : 2 : 4 : 6. Network of the two medullary shells regular, with small
circular pores, little larger than the bars. Network of the two cortical
shells irregular, with much larger polygonal pores, three to nine times as
broad as the thin bars. (Somewhat similar to _Cromyatractus ceparius_, Pl.
15, fig. 4.) Surface of the outermost shell smooth or a little thorny.
Polar spines cylindrical, very large, twice to three times as long as the
main axis of the outermost shell, about as broad as the innermost shell,
with conical apex; smooth or a little thorny.

_Dimensions._--Main axes of the four shells--(A) 0.03, (B) 0.05, (C) 0.13,
(D) 0.18; equatorial axes of them--(A) 0.2, (B) 0.45, (C) 0.11, (D) 0.14;
pores of the two medullary shells 0.004 and 0.006, of both cortical shells
0.01 to 0.03, bars 0.003.

_Habitat._--Indian Ocean, Zanzibar, Pullen, 2200 fathoms.

3. _Cromyatractus tetracelyphus_, n. sp. (Pl. 15, figs. 1, 1_a_).

Proportion of the main axes of the four concentric shells about =
1 : 3 : 10 : 11. Network of the two spherical medullary shells (fig. 1_a_)
regular, with small circular pores, twice as broad as {336}the bars.
Network of the inner cortical shell regular with circular,
hexagonally-framed pores, twice as broad as the bars. From every corner of
the hexagons (between three pores) arises a very thin, hair-like, short
radial spinule. These spinules communicate with one another by tangential
branches (at equal distances from the inner cortical shell), and form
thereby an outer, delicate cortical network, with large polygonal meshes
and very thin bars. Polar spines very strong, pyramidal, with prominent
edges, as long as the main radius of the cortical shell, and half as broad
at the base.

_Dimensions._--Main axes of the four shells--(A) 0.02, (B) 0.06, (C) 0.2,
(D) 0.22; equatorial axes of them--(A) 0.02, (B) 0.06, (C) 0.13, (D) 0.15;
pores of the four shells--(A) 0.002, (B) 0.004, (C) 0.008, (D) 0.02; bars
0.001 to 0.003; length of the polar spines 0.1, basal breadth 0.05.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

Subgenus 2. _Caryatractus_, Haeckel.

_Definition._--Shell composed of two medullary shells and three or more
cortical shells.

4. _Cromyatractus cepicius_, n. sp. (Pl. 15, fig. 3).

_Caryostylus cepicius_, Haeckel, 1879, Atlas (pl. xv. fig. 3).

_Caryodoras cepicius_, Haeckel, 1881, Prodromus, p. 454.

Proportion of the main axes of the five concentric shells about =
1 : 2 : 5 : 7 : 8. Network of both spherical medullary shells regular, with
small circular pores, about as broad as the bars. Network of the three
ellipsoidal cortical shells irregular, with large polygonal meshes, five to
ten times as broad as the thin bars. Surface of the outermost shell smooth.
Polar spines cylindrical, thorny, about as thick as the innermost shell and
as long as the outermost shell, against the apex with a spindle-like
intumescence, formed by delicate spongy framework (fig. 3).

_Dimensions._--Main axes of the five concentric shells--(A) 0.03, (B) 0.05,
(C) 0.16, (D) 0.2, (E) 0.25; equatorial axes of them--(A) 0.03, (B) 0.05,
(C) 0.12, (D) 0.16, (E) 0.2; pores of the two medullary shells 0.002 to
0.004, of the three cortical shells 0.02 to 0.04, bars 0.002; length of the
polar spines 0.3, breadth 0.03.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

5. _Cromyatractus ceparius_, n. sp. (Pl. 15, fig. 4).

_Caryostylus ceparius_, Haeckel, 1881, Prodromus et Atlas (pl. xv. fig.
4).

Proportion of the main axes of the six concentric shells about =
1 : 2 : 7 : 10 : 13 : 18. Network of the three inner shells regular, with
small, circular pores, which are about as broad as the bars and in the
third shell hexagonally framed. Network of the three outer shells
irregular, with large polygonal meshes, six to twelve times as broad as the
bars. Surface smooth. Polar spines strong, spindle-shaped, thorny, outside
of the sixth shell about as long as inside of it; broader than {337}the
innermost shell; the thorns of their outer free part (arising at equal
distances) represent perhaps the beginnings of three to four further
shells.

_Dimensions._--Main axes of the six concentric shells--(A) 0.02, (B) 0.05,
(C) 0.14, (D) 0.2, (E) 0.26, (F) 0.37; equatorial axes--(A) 0.02, (B) 0.05,
(C) 0.09, (D) 0.15, (E) 0.21, (F) 0.3; pores of the three inner shells
0.003 to 0.006, of the three outer shells 0.02 to 0.04, bars 0.003 to
0.005; length of the polar spines, from the base of the innermost shell to
the apex, 0.35, greatest breadth 0.03.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Genus 142. _Pipetta_,[181] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
simple medullary shell, with two hollow fenestrated tubes opposite on both
poles of the main axis.

The genus _Pipetta_ differs from _Pipettella_ (p. 304) in the possession of
an inner (medullary) shell; it exhibits the same tubular prolongations of
the cortical shell at both poles of the main axis.

1. _Pipetta fusus_, n. sp. (Pl. 39, figs. 8, 8_a_).

Cortical shell spindle-shaped, the middle ellipsoidal part gently passing
over on both poles into the conical tubes, which attain about the same
length. Network regular, with circular, hexagonally-framed pores, twice as
broad as the bars, sixteen to eighteen on the half equator. In the middle
part of the shell the pores arise from their hexagonal bases in the form of
short conical funnels; on both tubes they are much smaller and simple,
without frame. Medullary shell (fig. 8_a_) spherical, scarcely one-third as
broad as the cortical, with regular, circular pores, twice as broad as the
bars, eight to nine on the half equator. The two shells are connected only
by two opposite beams, lying in the equatorial axis and ramified at the
distal insertion (fig. 8_a_).

_Dimensions._--Middle ellipsoidal part of the cortical shell 0.13 to 0.15;
polar tubes 0.15 long, 0.06 broad at the base; pores of the former 0.01,
bars 0.005; pores of the tubes 0.06, bars 0.003. Medullary shell 0.04,
pores 0.004, bars 0.002.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

2. _Pipetta tuba_, n. sp. (Pl. 39, fig. 7).

Cortical shell nearly spherical in the middle part, which is sharply
separated at both poles from the long, nearly cylindrical tubes; these are
longer than the main axis, at the distal end open (always broken off).
Network regular, with circular, hexagonally-framed pores, of the same
breadth as the bars, eighteen to twenty on the half equator. The pores of
the polar tubes are of the same shape, only much smaller, and arranged in
sixteen to eighteen longitudinal rows. Medullary shell {338}spherical,
about one-third as broad as the cortical, with simple circular pores. The
connection between the two shells is only made by two simple opposite
beams, lying in the equatorial axis.

_Dimensions._--Middle spheroidal part of the cortical shell 0.14 to 0.16;
polar tubes 0.15 to 0.2 or longer, 0.03 broad; pores and bars of the former
0.007, of the latter 0.003. Medullary shell 0.04, pores 0.006, bars 0.003.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

3. _Pipetta salpinx_, n. sp.

Cortical shell ellipsoidal, sharply separated from the cylindrical polar
tubes, which reach about the same length (or more). Network regular, with
circular pores, three times as broad as the bars, without hexagonal frames,
fourteen to fifteen on the half equator. Pores of the polar tubes smaller,
arranged in eight to ten longitudinal rows. Medullary shell spherical,
one-fourth as broad as the cortical, connected with it by a circle of four
to six radial beams, lying in the equatorial plane. Differs from
_Pipettella prismatica_ (Pl. 39, fig. 6) in the possession of a medullary
shell and the absence of the tube edges.

_Dimensions._--Main axis of the ellipsoid 0.14, equatorial axis 0.12, pores
0.01, bars 0.003; length of the tubes 0.15 or more, breadth 0.03. Medullary
shell 0.03, pores 0.002, bars 0.001.

_Habitat._--Pacific, central area, Stations 265 to 268, depth 2700 to 2900
fathoms.

4. _Pipetta conus_, n. sp.

Cortical shell spindle-shaped, the middle ellipsoidal part gently passing
over at both poles into the conical tubes, which attain only half its
length. Network irregular, with roundish or subcircular pores, twice to
three times as broad as the bars, twelve to sixteen on the half equator.
Medullary shell spherical, one-fourth as broad as the cortical. (Differs
from the foregoing species in the short conical tubes and the irregular
reticulation.)

_Dimensions._--Middle part of the cortical shell 0.16 long, 0.13 broad;
tubes 0.09 long, 0.05 on the base broad; pores 0.005 to 0.01, bars 0.003.
Medullary shell 0.04.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

Genus 143. _Pipettaria_,[182] n. gen.

_Definition._--#Druppulida# with simple ellipsoidal cortical shell and
double medullary shell, with two hollow fenestrated tubes opposite on both
poles of the main axis.

The genus _Pipettaria_ differs from the foregoing _Pipetta_ only in the
duplication of the medullary shell; as in this, the tubular prolongations
of the main axis of the cortical shell may be either conical (with closed
apex) or cylindrical (with apical opening?).

{339}1. _Pipettaria fusaria_, n. sp.

Cortical shell spindle-shaped, the middle ellipsoidal part gently passing
over on both poles into the conical tubes, which attain about half its
length. Pores regular, circular, twice as broad as the bars, sixteen to
eighteen on the half equator; pores of the tubes smaller. Both medullary
shells spheroidal, compressed. (The appearance of the cortical shell
resembles that of _Cannartiscus amphiconus_, Pl. 39, fig. 19, but without
the equatorial constriction.)

_Dimensions._--Main axis of the ellipsoid 0.15, equatorial axis 0.13;
length of the polar tubes 0.08, basal breadth 0.05; pores of the former
0.008, bars 0.004; diameter of the medullary shells 0.04 and 0.02.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

2. _Pipettaria tubaria_, n. sp. (Pl. 39, fig. 15).

_Cannartidium tubarium_, Haeckel, 1882, Atlas (pl. xxxix. fig. 15).

Cortical shell ellipsoidal, on both poles distinctly separated from the
short conical tubes, the length and breadth of which equal the outer
medullary shell. In the equatorial plane arises a circle of four to six
short conical protuberances, similar to the polar tubes. Pores subregular,
circular, or roundish, scarcely broader than the bars, sixteen to twenty on
the half equator. Both medullary shells spheroidal, somewhat compressed in
the direction of the two poles (as in fig. 18_a_).

_Dimensions._--Main axis of the ellipsoid 0.12, equatorial axis 0.09; pores
0.005, bars 0.004; size of the equatorial protuberances and of the polar
tubes 0.02.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Family XIII. #SPONGURIDA#, Haeckel (Pl. 48, figs. 6, 7).

_Spongurida_, Haeckel, 1862, Monogr. d. Radiol., p. 447 (_sensu emendato_).

_Definition._--#Prunoidea# with spongy ellipsoidal or cylindrical shell,
composed wholly or partially of a spongy framework, without equatorial
stricture, with or without an enclosed medullary shell.

The family #Spongurida# comprises, in the sense here restricted, all those
#Prunoidea# in which the ellipsoidal or cylindrical shell is composed
wholly or partially of an irregular siliceous framework, not of simple
lattice-work. It contains two subfamilies, differing in the absence or
presence of a latticed medullary shell in the middle of the central
capsule; in the Spongellipsida it is absent, in the Spongodruppida present;
the former are most nearly related to the Ellipsida, the latter to the
Druppulida, the difference consisting only in the spongy structure of the
cortical shell.

In my Monograph (1862, p. 447) the family Spongurida had a much wider
extent, comprising also a number of #Sphæroidea# and #Discoidea#, agreeing
in the spongy {340}structure of the shell. Here we restrict the definition
to those spongy #Sphærellaria# in which the central capsule and the
enclosing spongy cortical shell are ellipsoidal or cylindrical, therefore
each transverse section is a circle, and each meridional section an
ellipse, as in all #Prunoidea#.

_The Cortical Shell_ in all Spongurida is composed of a delicate framework
of irregularly branched and interwoven siliceous threads; commonly this
spongy structure is rather dense or compact, but sometimes also very loose.
In the simplest form, _Spongellipsis_, the spongy cortical shell contains a
large cavity, in which lies freely the central capsule. In _Spongurus_ this
cavity is completely distended by a spongy framework. In this case the
solid spongy shell becomes often prolonged, and its original ellipsoidal
form passes over into a cylindrical one (as in many Zygartida). Sometimes
(particularly in _Spongocore_, Pl. 48, fig. 6) the cylinder becomes
three-jointed by two more or less distinct annular constrictions.
_Spongocore_ is distinguished by an outer veil, a thin lattice-lamella,
which envelops the spongy shell and is connected with it by radial beams.

_The Medullary Shell_, absent in the Spongellipsida, is constant in the
second subfamily Spongodruppida. It is either a simple latticed shell
(_Spongodruppa_) or double, composed of two concentric latticed shells
(_Spongoliva_); its form is either spherical or ellipsoidal. It lies in the
middle of the central capsule, and is connected by radial beams
(perforating the membrane of the latter) with the enveloping spongy
cortical shell.

In many Spongurida the surface of the shell is armed with radial spines,
and in some genera (_Spongoprunum_, _Spongatractus_, &c., Pl. 17, fig. 12),
on both poles of the main axis, are developed two strong opposite polar
spines, as in many other #Prunoidea#.

_The Central Capsule_ of the Spongurida is either ellipsoidal or
cylindrical. Only in _Spongellipsis_ it lies freely in the internal cavity
of the spongy shell. In all other genera it is perforated by a part of the
skeleton; in the Spongodruppida it contains the simple or double medullary
shell, and the radial beams which perforate its membrane and connect the
latter with the external spongy cortical shell. In _Spongurus_ and the
allied genera (_Spongocore_, _Spongoprunum_) the whole central capsule is
filled with a spongy framework which also envelops its surface.

{341}_Synopsis of the Genera of Spongurida._

-------------------------------------------------------------------------
I. Subfamily Spongellipsida.
(Ellipsoidal shell composed only of a spongy framework without a
latticed medullary shell in the centre.)
-------------------------------------------------------------------------
Spongy shell }No polar spines, 144. _Spongellipsis_.
with internal }
cavity. }

{Without lattice
Spongy shell { mantle. No polar
solid, without { spines, 145. _Spongurus_.
internal {
cavity. {With lattice
{ mantle.
{ No polar spines, 146. _Spongocore_.
{
{Without lattice
{ mantle. Two
{ opposite spines
{ on the poles of
{ the axis, 147. _Spongoprunum_
-------------------------------------------------------------------------
II. Subfamily Spongodruppida.
(Ellipsoidal shell composed of an outer spongy cortical shell and an
inner latticed medullary shell.)
-------------------------------------------------------------------------
Medullary shell {No polar spines, 148. _Spongodruppa_.
simple. {
{Two opposite spines
{ on the poles of
{ the axis, 149. _Spongatractus_.

Medullary shell {No polar spines, 150. _Spongoliva_.
double. {
{
{Two opposite spines
{ on the poles of
{ the axis, 151. _Spongoxiphus_.

Subfamily 1. SPONGELLIPSIDA, Haeckel.

_Definition._--#Spongurida# with a spongy ellipsoidal or cylindrical shell,
without an internal latticed medullary shell.

Genus 144. _Spongellipsis_,[183] n. gen.

_Definition._--#Spongurida# with an ellipsoidal or cylindrical spongy
shell, containing an internal cavity, without a latticed medullary shell.
Polar spines absent.

The genus _Spongellipsis_ embraces those very simple Spongurida in which
the ellipsoidal central capsule is enclosed in a spongy cortical shell of
the same form. It corresponds, therefore, to _Plegmosphæra_ among the
#Sphæroidea#, to _Plegmodiscus_ among the #Discoidea#, and to
_Spongolarcus_ among the #Larcoidea#. In some species the ellipsoidal form
is prolonged and passes into a cylindrical one.

Subgenus 1. _Spongellipsarium_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

{342}1. _Spongellipsis lævis_, n. sp.

Shell ellipsoidal, with smooth surface, its spongy wall scarcely one-tenth
as thick as the minor axis of the inner cavity. Spongy framework very
compact, with small meshes, three to six times as broad as the bars.
Proportion of the major axis to the minor = 3 : 2.

_Dimensions._--Major axis (or length) of the shell 0.36, minor axis (or
breadth) 0.24.

_Habitat._--Pacific, central area, Station 265, surface.

2. _Spongellipsis aspera_, n. sp.

Shell ellipsoidal, with rough surface, its spongy wall nearly half as thick
as the minor axis of the inner cavity. Spongy framework very loose, with
large meshes, ten to twenty times as broad as the bars. Proportion of the
major axis to the minor = 4 : 3.

_Dimensions._--Length of the shell 0.24, breadth 0.18.

_Habitat._--North Pacific, Station 253, surface.

3. _Spongellipsis aplysina_, n. sp.

Shell nearly cylindrical, with rough surface, its spongy wall about
one-fourth as thick as the minor axis of the internal cavity. Spongy
framework loose, with large meshes, four to eight times as broad as the
bars. Proportion of both axes = 6 : 1. (Similar to a spongy cylinder of
Aplysina.)

_Dimensions._--Length of the shell 0.3, breadth 0.05.

_Habitat._--North Atlantic, Station 64, surface.

Subgenus 2. _Spongellipsidium_, Haeckel.

_Definition._--Surface of the shell covered with radial spines.

4. _Spongellipsis setosa_, n. sp.

Shell ellipsoidal, covered with numerous (sixty to eighty) thin,
bristle-shaped, radial spines, about half as long as the major axis. Spongy
framework loose, with large meshes, ten to twenty times as broad as the
bars. Minor axis of the inner cavity twice as long as the thickness of the
spongy wall. Proportion of both axes = 5 : 3.

_Dimensions._--Length of the shell 0.2, breadth 0.12.

_Habitat._--South Atlantic, Station 325, surface.

5. _Spongellipsis spinosa_, n. sp.

Shell nearly cylindrical, covered with numerous thorns and thirty to forty
larger conical radial spines, somewhat longer than the major axis. Spongy
framework compact, with small meshes, four {343}to eight times as broad as
the bars. Minor axis of the inner cavity about six times as large as the
thickness of the spongy wall. Proportion of both axes = 4 : 1.

_Dimensions._--Length of the shell 0.24, breadth 0.06.

_Habitat._--South Pacific, Station 300, surface.

Genus 145. _Spongurus_,[184] Haeckel, 1862, Monogr. d. Radiol., p. 465.

_Definition._--#Spongurida# with ellipsoidal or cylindrical (sometimes
three-jointed) shell, of solid spongy framework, without internal cavity
and without latticed medullary shell. Polar spines and outer lattice-mantle
absent.

The genus _Spongurus_ was founded by me in 1862 for the common cosmopolitan
_Spongurus cylindricus_, a massive spongy cylinder with radial spines. I
enlarge here the conception of the genus, in receiving also ellipsoidal,
massive spongy #Prunoidea#, with or without radial spines. Sometimes the
cylindrical shell is more or less distinctly three-jointed, with two
annular strictures, as also in the following and nearly allied genus.

Subgenus 1. _Spongurantha_, Haeckel.

_Definition._--Spongy framework everywhere of the same structure; surface
smooth or rough, but without radial spines.

1. _Spongurus stuparius_, n. sp.

Shell ellipsoidal, one and a half times as long as broad, with nearly
smooth surface. Spongy framework everywhere of equal structure, with small
meshes, four to six times as broad as the bars.

_Dimensions._--Length of the shell (or major axis) 0.2, breadth of it (or
minor axis) 0.14.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

2. _Spongurus stypticus_, n. sp.

Shell ellipsoidal, twice as long as broad, with thorny surface. Spongy
framework everywhere of equal structure, with large meshes, ten to twenty
times as broad as the bars.

_Dimensions._--Length of the shell 0.3, breadth 0.15.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

3. _Spongurus phalanga_, Haeckel.

_Spongurus cylindricus_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 119,
Taf. vii. fig. 3.

Shell nearly cylindrical, five times as long as broad, with thorny surface.
Spongy framework compact, everywhere of equal structure, with small meshes,
scarcely broader than the bars.

{344}_Dimensions._--Length of the shell 0.24, breadth 0.05.

_Habitat._--Pacific, central area, Station 268, depth 2600 fathoms; fossil
in the Tertiary rocks of Sicily, Stöhr.

Subgenus 2. _Spongurella_, Haeckel.

_Definition._--Spongy framework everywhere of the same structure; surface
armed with radial spines.

4. _Spongurus asper_, Haeckel.

_Haliomma asperum_, Joh. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 40, Taf. ii. fig. 2.

_Haliomma asperum_, Haeckel, 1862, Monogr. d. Radiol., p. 431.

Shell ellipsoidal, one and a third times as long as broad, with thorny
surface and twenty symmetrically disposed, thin, bristle-shaped, radial
spines, about as long as the shell. Spongy framework everywhere of the same
structure, very compact, with small meshes, three to four times as broad as
the bars.

_Dimensions._--Length of the shell 0.11, breadth 0.08.

_Habitat._--Mediterranean, Ligurian coast (J. Müller), Portofino (Haeckel).

5. _Spongurus cylindricus_, Haeckel.

_Spongurus cylindricus_, Haeckel, 1862, Monogr. d. Radiol., p. 465, Taf.
xxvii. fig. 1.

Shell cylindrical, four to five times as long as broad, with nearly smooth
surface, and twenty to thirty thin, bristle-shaped, radial spines, about
half as long as the shell. Spongy framework everywhere of the same
structure, very compact, with small meshes, scarcely broader than the bars.

_Dimensions._--Length of the shell 0.2, breadth 0.04 to 0.05.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, and Pacific, surface.

6. _Spongurus tricolus_, n. sp.

Shell nearly cylindrical, with two slight annular transverse strictures;
its middle part is somewhat broader. Surface thorny, with numerous (forty
to sixty or more) short, thin, radial spines, not longer than the breadth
of the shell. Spongy framework everywhere of the same structure, compact,
with small meshes, twice to three times as broad as the bars.

_Dimensions._--Length of the shell 0.24, breadth 0.04 to 0.06.

_Habitat._--Pacific, central area, Station 272, 2600 fathoms.

Subgenus 3. _Sponguroma_, Haeckel.

_Definition._--Spongy framework in the inner part of the shell very
compact, in the outer part very loose; surface armed with radial spines.

{345}7. _Spongurus radians_, n. sp.

Shell ellipsoidal, one and a half times as long as broad, with thorny
surface, and eighty to one hundred and twenty (or more) thin,
bristle-shaped, radial spines, somewhat longer than the shell. Spongy
framework in the inner part very compact, in the outer part very loose,
with a gradual transition between the two parts. Meshes in the central part
not broader than the bars, in the superficial part ten to twenty times as
broad.

_Dimensions._--Length of the shell 0.24, breadth 0.16.

_Habitat._--Mediterranean, Portofino, near Genoa, Haeckel, surface.

8. _Spongurus spongechinus_, n. sp.

Shell ellipsoidal, one and a third times as long as broad, with thorny
surface, and thirty to forty thick, conical radial spines, about half as
long as the shell. Spongy framework in the inner part of the shell compact,
in the outer loose, with a gradual transition between the two parts. Meshes
in the central part smaller than the bars, in the superficial part four to
eight times as broad.

_Dimensions._--Length of the shell 0.18, breadth 0.14.

_Habitat._--North Atlantic, Station 353, surface.

Genus 146. _Spongocore_,[185] n. gen.

_Definition._--#Spongurida# with ellipsoidal or cylindrical (sometimes
three-jointed) shell of solid spongy framework, without internal cavity and
without latticed medullary shell. Polar spines absent. An outer
lattice-mantle is connected with the spongy shell by radial beams.

The genus _Spongocore_ contains some widely distributed Spongurida, which
are distinguished from the nearly related _Spongurus_ by the development of
a peculiar veil or mantle of delicate lattice-work, which envelops either
the whole shell or only the middle part of it, and is connected with it by
numerous radial beams. The distance of the simple fine lattice-lamella from
the spongy shell is everywhere the same. Often the cylindrical shell is
three-jointed, with two annular strictures, as also in the foregoing
_Spongurus._ It is derived from the latter by development of the veil
connecting the points of the radial spines.

Subgenus 1. _Spongocorina_, Haeckel.

_Definition._--Shell without distinct annular strictures, not evidently
three-jointed.

{346}1. _Spongocore velata_, n. sp.

Shell ellipsoidal, one and a half times as long as broad, without annular
strictures. Spongy framework loose, with rather large meshes, four to six
times as broad as the bars. The whole spongy shell enveloped by a delicate
veil with smooth surface, connected with it by numerous thin radial beams.
Breadth of the spongy ellipsoid (or minor axis) six times as large as its
distance from the veil.

_Dimensions._--Length of the whole shell (with veil) 0.3, breadth 0.2;
distance of the veil from the spongy ellipsoid 0.035.

_Habitat._--South Atlantic, Station 325, surface.

2. _Spongocore diplocylindrica_, n. sp.

Shell cylindrical, three times as long as broad, without annular
strictures. Spongy framework compact, with small meshes, twice to three
times as broad as the bars. The whole surface of the spongy cylinder, with
exception of both rounded polar faces, enveloped by a delicate cylindrical
veil with smooth surface, connected with it by numerous radial beams.
Diameter of the spongy cylinder twice as large as its distance from the
veil.

_Dimensions._--Length of the shell 0.3, breadth (with veil) 0.1; distance
of the veil 0.025.

_Habitat._--South Pacific, Station 302, surface.

Subgenus 2. _Spongocorisca_, Haeckel.

_Definition._--Shell distinctly three-jointed, with two transverse annular
strictures.

3. _Spongocore chrysalis_, n. sp.

Spongy shell cylindrical, without the veil six times as long as broad,
three-jointed, with two annular strictures; all three joints of the same
length. The whole spongy shell enveloped by a thin veil with thorny
surface, connected with it by numerous radial beams. Breadth of the spongy
cylinder three times as large as its distance from the veil.

_Dimensions._--Length of the shell (with veil) 0.4, breadth 0.1; distance
of the veil from the spongy shell 0.02.

_Habitat._--South Atlantic (off Patagonia), Station 318, surface.

4. _Spongocore cincta_, n. sp.

Spongy shell cylindrical, without the veil four times as long as broad,
three-jointed, with two annular strictures; all three joints of the same
length. Only the middle joint enveloped by a thin veil with smooth surface,
connected with it by radial beams. Both terminal joints with long,
{347}bristle-shaped, radial spines. Breadth of the spongy cylinder twice as
large as its distance from the veil.

_Dimensions._--Length of the shell 0.24, breadth (with veil) 0.12; distance
of the veil 0.03.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

5. _Spongocore pupula_, n. sp.

Spongy shell cylindrical, without the veil six times as long as broad,
three-jointed, with two annular strictures; the middle joint half as long
as either terminal joint. Only the middle joint enveloped by a thin veil
with spiny surface, connected with it by twenty to thirty radial beams,
which are prolonged on the outside into short radial spines. Breadth of the
spongy cylinder about equal to its distance from the veil.

_Dimensions._--Length of the shell 0.2, breadth (with veil) 0.1; distance
of the veil 0.033.

_Habitat._--North Pacific, Station 244, surface.

6. _Spongocore puella_, n. sp. (Pl. 48, fig. 6).

Spongy shell cylindrical, without the veil five times as long as broad,
three-jointed, with two annular strictures; the middle joint twice as long
as either terminal joint. Only the middle joint enveloped by a thin veil
with delicate lattice-work and smooth surface; both terminal joints armed
with numerous thin, bristle-shaped, radial spines. Breadth of the spongy
cylinder twice as large as the distance from the veil.

_Dimensions._--Length of the shell 0.25 to 0.3, breadth (with veil) 0.1 to
0.12; distance of the veil 0.028 to 0.032.

_Habitat._--South Pacific, Stations 295 to 304, surface.

Genus 147. _Spongoprunum_,[186] n. gen.

_Definition._--#Spongurida# with ellipsoidal or cylindrical shell of solid
spongy framework, without internal cavity and without latticed medullary
shell. On the poles of the axis occur two opposite strong spines.

The genus _Spongoprunum_ differs from the most nearly allied _Spongurus_ by
the possession of two strong solid spines, lying in the axis of the
ellipsoidal or cylindrical massive spongy shell, on its two opposite poles.
_Spongoprunum_ bears therefore the same relation to _Spongurus_ that
_Ellipsoxiphus_ does to _Cenellipsis_.

1. _Spongoprunum amphilonche_, n. sp. (Pl. 48, fig. 7).

Shell ellipsoidal, two and a third times as long as broad, with nearly
smooth surface. Spongy framework very compact, in the whole solid shell of
the same structure, with very small meshes, {348}scarcely broader than the
bars. Two polar spines conical, very strong, about half as long as the main
axis of the shell and three times as long as broad.

_Dimensions._--Length of the shell (without spines) 0.2, breadth 0.08.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms; fossil in the
rocks of Barbados.

2. _Spongoprunum atractus_, n. sp.

Shell spindle-shaped, twice as long as broad, with thorny surface. Spongy
framework loose, in the whole shell of the same structure, with large
meshes, eight to twelve times as broad as the bars. Two polar spines
angular, pyramidal, very strong, about one-third as long as the shell, and
twice as long as broad.

_Dimensions._--Length of the shell 0.2, breadth 0.11.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

3. _Spongoprunum amphicylindrus_, n. sp.

Shell nearly cylindrical, four times as long as broad, with rough surface.
Spongy framework compact, in the whole shell of equal structure, with small
meshes, twice to three times as broad as the bars. Two polar spines
cylindrical, very large, longer than the shell, and about one-fourth as
thick as its diameter.

_Dimensions._--Length of the shell 0.2, breadth 0.05.

_Habitat._--South Atlantic, Station 333, surface.

Subfamily SPONGODRUPPIDA, Haeckel.

_Definition._--#Spongurida# with latticed medullary shell, enclosed by a
spongy cortical shell.

Genus 148. _Spongodruppa_,[187] n. gen.

_Definition._--#Spongurida# with spongy ellipsoidal cortical shell,
enclosing a simple, spherical or ellipsoidal, latticed medullary shell.
Polar spines absent.

The genus _Spongodruppa_ opens the series of Spongodruppida, or of those
Spongurida in which the ellipsoidal spongy cortical shell encloses a simple
or double, latticed, medullary shell. In _Spongodruppa_, the simplest and
probably the ancestral form of this subfamily, the medullary shell is
simple and the polar spines absent. It may be derived from _Druppula_ by a
spongy thickening of the simple latticed cortical shell.

{349}Subgenus 1. _Spongodruppula_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Spongodruppa terebintha_, n. sp.

Cortical shell one and a third times as long as broad, with smooth surface,
three times as broad as the spherical medullary shell. Spongy framework
very compact, with small meshes of the same breadth as the bars. Thickness
of the spongy wall equal to the radius of the medullary shell.

_Dimensions._--Length of the cortical shell (or major axis of the
ellipsoid) 0.16, breadth (or minor axis) 0.12; thickness of its wall 0.02;
medullary shell 0.04.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

2. _Spongodruppa pistacia_, n. sp.

Cortical shell one and a half times as long as broad, with thorny surface,
nearly five times as broad as the spherical medullary shell. Spongy
framework compact, its thickness equal to the diameter of the medullary
shell.

_Dimensions._--Length of the cortical shell 0.2, breadth 0.14; thickness of
its wall 0.03; medullary shell 0.03.

_Habitat._--Fossil in the rocks of Barbados.

3. _Spongodruppa lentisca_, n. sp.

Cortical shell twice as long as broad, with rough surface, four times as
large as the ellipsoidal medullary shell. Spongy framework loose, with
large meshes, its thickness equal to the length of the medullary shell.

_Dimensions._--Length of the cortical shell 0.25, breadth 0.13; thickness
of its wall 0.06; length of the medullary shell 0.06, breadth 0.04.

_Habitat._--Pacific, central area, Station 266, surface.

Subgenus 2. _Spongodruppium_, Haeckel.

_Definition._--Surface of the shell armed with radial spines.

4. _Spongodruppa frangula_, n. sp.

Cortical shell one and a third times as long as broad, three times as broad
as the spherical medullary shell. Spongy framework loose, with large
meshes, its thickness half as large as the diameter of the medullary shell.
Surface covered with numerous (forty to fifty) thin, bristle-shaped, radial
spines, about as long as the cortical shell.

{350}_Dimensions._--Length of the cortical shell 0.24, breadth 0.18;
thickness of its wall 0.035; diameter of the medullary shell 0.06.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

5. _Spongodruppa elliptica_, Haeckel.

_Acanthosphæra elliptica_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 285, Taf. vii. fig. 4.

Cortical shell nearly twice as long as broad, four times as broad as the
ellipsoidal medullary shell. Spongy framework compact, with small meshes.
Surface covered with very numerous short radial spines, about as long as
the medullary shell.

_Dimensions._--Length of the cortical shell 0.2, breadth 0.11; length of
the medullary shell 0.04, breadth 0.03.

_Habitat._--Philippine Sea, Ehrenberg, Station 206, depth 2100 fathoms.

6. _Spongodruppa polyacantha_, Haeckel.

_Haliomma polyacanthum_, J. Müller, 1858, Abhandl., p. 36, Taf. i. figs.
10, 11.

_Haliomma polyacanthum_, Haeckel, 1862, Monogr. d. Radiol., p. 430.

Cortical shell one and a third times as long as broad, twice as broad as
the ellipsoidal medullary shell. Spongy framework loose on the surface,
with large regular meshes. Surface covered with fifteen to twenty thick,
conical spines, about as long as the radius of the shell.

_Dimensions._--Length of the cortical shell 0.12, of the medullary shell
0.09; diameter of the medullary shell 0.05 to 0.06.

_Habitat._--Mediterranean (south coast of France); Atlantic (Canary
Islands), Station 354, surface.

Genus 149. _Spongatractus_,[188] n. gen.

_Definition._--#Spongurida# with spongy ellipsoidal cortical shell,
enclosing a simple, spherical or ellipsoidal, latticed medullary shell. On
the poles of the axis occur two opposite strong spines.

The genus _Spongatractus_ differs from _Spongodruppa_ by development of two
strong spines in the axis of the shell, on its two opposite poles;
therefore it bears the same relation to the latter that _Lithatractus_ does
to _Druppula_, and can be derived from _Lithatractus_ by a spongy
thickening of the cortical shell.

1. _Spongatractus pachystylus_, Haeckel.

_Spongosphæra pachystyla_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 82, Taf. xxvi. fig. 3.

Cortical shell one and a half times as long as broad, with rough surface.
Spongy framework compact, with small meshes, about as thick as the
spherical medullary shell. Polar spines very {351}stout, conical, slightly
sulcated, about as long as the breadth of the cortical shell, as broad at
the base as the medullary shell.

_Dimensions._--Length of the cortical shell 0.2, breadth 0.13; thickness of
the spongy wall 0.045; diameter of the medullary shell 0.04.

_Habitat._--Fossil in the rocks of Barbados, Ehrenberg; living in the
Equatorial Atlantic, Station 348, depth (2450) fathoms.

2. _Spongatractus fusiformis_, n. sp.

Cortical shell one and a third times as long as broad, with nearly smooth
surface. Spongy framework very compact, with very small meshes, about as
thick as the radius of the spherical medullary shell. Polar spines very
strong, three-sided pyramidal, about as long as the breadth of the cortical
shell, as broad at the base as the medullary shell.

_Dimensions._--Length of the cortical shell 0.16 to 0.18, breadth 0.12 to
0.14; thickness of the spongy wall 0.02; diameter of the medullary shell
0.04.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Spongatractus streptacanthus_, n. sp.

Cortical shell twice as long as broad, with thorny surface. Spongy
framework loose, with large meshes, about as thick as the ellipsoidal
medullary shell. Polar spines very long, three-sided prismatic, with three
spirally contorted edges, much longer than the cortical shell, half as
broad as the medullary shell.

_Dimensions._--Length of the cortical shell 0.25, breadth 0.13; thickness
of the spongy wall 0.03; diameter of the medullary shell 0.03 to 0.04.

_Habitat._--North Atlantic, off Canary Islands, Haeckel.

Genus 150. _Spongoliva_,[189] n. gen.

_Definition._--#Spongurida# with spongy ellipsoidal cortical shell,
enclosing a double, spherical or ellipsoidal, latticed medullary shell.
Polar spines absent.

The genus _Spongoliva_ differs from the nearest _Spongodruppa_ by
duplication of the medullary shell; it bears therefore the same relation to
this that _Prunulum_ does to _Druppula_, and can be regarded as a
_Prunulum_, in which the simple latticed cortical shell is replaced by a
spongy framework.

Subgenus 1. _Spongolivetta_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

{352}1. _Spongoliva cerasina_, n. sp.

Cortical shell one and a fourth times as long as broad, with smooth
surface, four times as broad as the outer spherical medullary shell. Spongy
framework very compact, with very small meshes, its thickness equal to the
diameter of the outer medullary shell.

_Dimensions._--Length of the cortical shell 0.2, breadth 0.16; thickness of
its wall 0.04; diameter of the outer medullary shell 0.04, of the inner
0.014.

_Habitat._--Fossil in the rocks of Barbados.

2. _Spongoliva prunulina_, n. sp.

Cortical shell one and a third times as long as broad, with smooth surface,
three times as large as the outer ellipsoidal medullary shell. Spongy
framework very compact, with small meshes, its thickness equal to the
diameter of the inner medullary shell.

_Dimensions._--Length of the cortical shell 0.24, breadth 0.18; thickness
of its wall 0.03; size of the outer medullary shell 0.08 to 0.06, of the
inner 0.03.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

3. _Spongoliva persicina_, n. sp.

Cortical shell one and a half times as long as broad, with rough surface,
five to six times as large as the ellipsoidal outer medullary shell. Spongy
framework loose, with large meshes, its thickness about equal to the outer
medullary shell.

_Dimensions._--Length of the cortical shell 0.26, breadth 0.18; thickness
of its wall 0.04; size of the outer medullary shell 0.045 to 0.035, of the
inner 0.015.

_Habitat._--Equatorial Atlantic, Station 348, depth (2450) fathoms.

4. _Spongoliva amygdalina_, n. sp.

Cortical shell twice as long as broad, with thorny surface, ten times as
long as the small spherical outer medullary shell. Spongy framework loose,
with large meshes, its thickness one and a half times as large as the outer
medullary shell.

_Dimensions._--Length of the cortical shell 0.3, breadth 0.16; thickness of
its wall 0.045; size of the outer medullary shell 0.03, of the inner 0.015.

_Habitat._--Fossil in the Tertiary rocks of Sicily (Caltanisetta).

Subgenus 2. _Spongolivina_, Haeckel.

_Definition._--Surface of the shell armed with radial spines.

5. _Spongoliva opuntina_, n. sp.

Cortical shell one and a third times as long as broad, armed with numerous
thin, bristle-shaped, radial spines, nearly as long as the shell. Spongy
framework loose, with large meshes, its thickness nearly equal to the
diameter of the outer spherical medullary shell.

{353}_Dimensions._--Length of the cortical shell 0.18, breadth 0.14;
thickness of its wall 0.035; diameter of the outer medullary shell 0.04, of
the inner 0.02.

_Habitat._--South Atlantic, Station 325, surface.

6. _Spongoliva daturina_, n. sp.

Cortical shell one and a half times as long as broad, armed with numerous,
thick and short, conical radial spines, about as long as the outer
medullary shell. Spongy framework compact, with small meshes, its thickness
equal to half the length of the outer ellipsoidal medullary shell.

_Dimensions._--Length of the cortical shell 0.24, breadth 0.16; thickness
of its wall 0.025; diameter of the outer medullary shell 0.045 to 0.035, of
the inner 0.025.

_Habitat._--Pacific, central area, Station 272, surface.

Genus 151. _Spongoxiphus_,[190] n. gen.

_Definition._--#Spongurida# with spongy ellipsoidal cortical shell,
enclosing a double, spherical or ellipsoidal, latticed medullary shell. On
the poles of the axis occur two opposite strong spines.

The genus _Spongoxiphus_ differs from _Spongoliva_ by the development of
two strong spines in the axis of the shell, on its two opposite poles. From
the similar _Spongatractus_ it differs in the duplication of the medullary
shell. It may also be regarded as a _Stylatractus_, in which the simple
latticed cortical shell is replaced by an irregular spongy framework.

1. _Spongoxiphus sphærococcus_, n. sp.

Cortical shell one and a half times as long as broad, with thorny surface.
Spongy framework compact, with small meshes, its thickness equal to the
radius of the outer medullary shell. Both medullary shells spherical, the
outer three times as broad as the inner. Polar spines very stout, conical,
slightly sulcated, about half as long as the cortical shell, as broad at
the base as the outer medullary shell. (Differs from _Spongatractus
pachystylus_ almost only in the double medullary shell.)

_Dimensions._--Length of the cortical shell 0.18, breadth 0.12; thickness
of the spongy wall 0.025; diameter of the outer medullary shell 0.05, of
the inner 0.016.

_Habitat._--Fossil in the rocks of Barbados (Haeckel); also living in the
Pacific, central area, Station 265, depth 2900 fathoms.

{354}2. _Spongoxiphus prunococcus_, n. sp. (Pl. 17, figs. 12, 13).

_Spongostylus prunococcus_, Haeckel, 1881, Prodromus, p. 455, et Atlas,
pl. xvii. figs. 12, 13.

Cortical shell one and a third times as long as broad, with smooth surface.
Spongy framework very compact, with very small meshes, its thickness about
equals the breadth of the outer medullary shell. Both medullary shells
ellipsoidal, the outer three times as large as the inner. Polar spines very
stout, three-sided pyramidal, about half as long as the cortical shell, on
the base about as broad as the outer medullary shell. (Differs from the
preceding in the ellipsoidal form of both medullary shells, and in the
finer structure of the spongy framework.)

_Dimensions._--Length of the cortical shell 0.18 to 0.2, breadth 0.14 to
0.16; thickness of the spongy wall 0.035 to 0.04; length of the outer
medullary shell 0.04 to 0.06, breadth 0.03 to 0.04.

_Habitat._--Pacific, central area, Stations 265 to 268, depths 2700 to 2900
fathoms.

Family XIV. #ARTISCIDA#, Haeckel (Pl. 39, figs. 9, 10; Pl. 48, fig. 5).

_Artiscida_, Haeckel, 1881, Prodromus, p. 462.

_Definition._--#Prunoidea# with an ellipsoidal twin-shell divided by an
equatorial stricture into two communicating hemiellipsoidal or
hemispherical chambers, without enclosed medullary shell. Central capsule
ellipsoidal, with or without equatorial stricture.

The family #Artiscida# has a simple fenestrated outer shell, like that of
the Ellipsida, but differs from these in the presence of an equatorial
constriction, by which it assumes a characteristic twin form, somewhat
similar to a violin (Pl. 39, figs. 9, 10). From the Cyphinida, which have
the same form of the cortical shell, the Artiscida differ in the absence of
the medullary shell. It is possible that the Artiscida are descended from
the Cyphinida (by loss of the medullary shell), but it is more probable
that they arise from the Ellipsida by an annular constriction in the
equatorial plane (perhaps the formation of the shell originally took place
while the central capsule was undergoing division). Both halves of the twin
shell are always of the same size and form. Its outer surface is either
smooth or covered with radial spines (_Artiscus_, Pl. 39, fig. 9).
Sometimes on the opposite poles of the main axis are developed solid spines
(_Stylartus_, Pl. 48, fig. 5) or hollow fenestrated tubes (_Cannartus_, Pl.
39, fig. 10).

_The Central Capsule_ is either simply ellipsoidal, or has also an
equatorial constriction, which divides it into two equal halves. It is
constantly smaller than the surrounding shell, and separated from its inner
surface by a thicker or thinner jelly-mantle, the calymma.

_Synopsis of the Genera of Artiscida._

On the poles of the main axis neither solid spines
nor hollow tubes, 152. _Artiscus_.

On the poles of the main axis two solid spines (or
bunches of spines), 153. _Stylartus_.

On the poles of the main axis two hollow fenestrated
tubes, 154. _Cannartus_.

{355}Genus 152. _Artiscus_,[191] Haeckel, 1881, Prodromus, p. 462.

_Definition._--#Artiscida# without peculiar polar appendages (without
strong solid spines or hollow fenestrated tubes on the poles of the main
axis).

The genus _Artiscus_ represents the most simple and primitive form of the
Artiscida, and may therefore be regarded as the ancestral form of this
subfamily. It resembles an ordinary double loaf, composed of two equal,
nearly ellipsoidal fenestrated shells separated by the stricture in the
equatorial plane. _Artiscus_ can be derived from _Cenellipsis_ simply by
this equatorial constriction. But it can also be derived from _Cyphanta_ by
loss of the medullary shell.

Subgenus 1. _Artiscium_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial rods or
spines.

1. _Artiscus paniscus_, n. sp.

Pores of the shell regular, hexagonal, four times as broad as the bars; ten
to twelve pores on the half meridian, eight to nine on the half equator.
Surface smooth. (The network of this species is quite the same as that of
the proximal cameræ of _Panartus diploconus_, Pl. 40, fig. 1.)

_Dimensions._--Main axis of the shell 0.12, equatorial axis 0.07; meshes
0.013, bars 0.003.

_Habitat._--Pacific, central area, Station 271, depth 2425.

2. _Artiscus facetus_, n. sp.

Pores of the shell regular, circular, with prominent hexagonal frame, twice
as broad as the bars; twenty-two to twenty-four pores on the half meridian,
fourteen to fifteen on the half equator. Surface a little thorny. (The
shell of this species exhibits nearly the same formation as that of
_Cyphonium virgineum_, Pl. 39, fig. 12, but without enclosed medullary
shells.)

_Dimensions._--Main axis of the shell 0.15, equatorial axis 0.09; meshes
0.01, bars 0.005.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

3. _Artiscus paniculus_, n. sp.

Pores of the shell regular, circular, without hexagonal frame, as broad as
the bars; fourteen to sixteen pores on the half meridian, ten to eleven on
the half equator. Surface quite smooth.

_Dimensions._--Main axis of the shell 0.13, equatorial axis 0.06; meshes
0.01, bars 0.01.

_Habitat._--Fossil in the Tertiary rocks of Barbados (Haeckel).

{356}4. _Artiscus panarius_, n. sp.

Pores of the shell irregular, roundish, twice to four times as broad as the
bars; sixteen to eighteen pores on the half meridian, twelve to thirteen on
the half equator. Surface smooth.

_Dimensions._--Main axis of the shell 0.11, equatorial axis 0.06; meshes
0.005 to 0.02, bars 0.001 to 0.004.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

Subgenus 2. _Artidium_, Haeckel, 1881, Prodromus, p. 462.

_Definition._--Surface of the shell covered with radial rods or spines.

5. _Artiscus elegans_, n. sp.

Pores of the shell regular, circular, with hexagonal frames, twice as broad
as the bars; fourteen pores on the half meridian, eight to nine on the half
equator. From every corner of the hexagonal frames (between every three
pores) starts a thin, three-sided pyramidal spine, twice as large as a
pore.

_Dimensions._--Main axis 0.13, equatorial axis 0.08; meshes 0.007, bars
0.004; spines 0.015 long.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

6. _Artiscus nodosus_, n. sp. (Pl. 39, fig. 9).

Pores of the shell subregular, circular, without hexagonal frame, three
times as broad as the bars; sixteen to eighteen on the half meridian, ten
to twelve on the half equator. Irregularly scattered on the whole surface a
variable number (twenty-five to thirty in all) of stout short radial spines
or rather blunt rods; the length and thickness of these is the same, and
equals the size of two to three meshes; its form resembles a truncated
six-sided pyramid.

_Dimensions._--Main axis 0.11, equatorial axis 0.08; meshes 0.01, bars
0.003; length and thickness of the radial sticks 0.02.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

7. _Artiscus hystrix_, n. sp.

Pores of the shell irregular, roundish, of very unequal size and dissimilar
form, twice to eight times as broad as the bars; ten to fifteen on the half
meridian, seven to nine on the half equator. Irregularly scattered on the
whole surface a large number of thin conical spines, about as large as the
meshes, partly directed radially, partly obliquely. (The shell of this
species resembles very much that of _Cyphonium ceratospyris_ (p. 366) =
_Didymocyrtis ceratospyris_, Monogr. d. Radiol., 1862, Taf. xxii. fig. 14,
but is without the enclosed inner shells.)

{357}_Dimensions._--Main axis 0.12, equatorial axis 0.07; meshes 0.004 to
0.02, bars 0.002 to 0.005; spines 0.01 to 0.02.

_Habitat._--Atlantic, Eastern Tropical part, Station 346, surface.

Genus 153. _Stylartus_,[192] Haeckel, 1881, Prodromus, p. 462.

_Definition._--#Artiscida# with two strong, solid, polar spines, or two
bunches of polar spines, opposite on the two poles of the main axis.

The genus _Stylartus_ differs from _Artiscus_ by the production of two
large opposite spines in the main axis, starting from both poles of it;
sometimes every spine is surrounded by a group of smaller radial spines.
The genus is nearly allied to _Ellipsoxiphus_ (p. 295), and differs from it
only in the equatorial stricture of the ellipsoidal shell. But it may also
be derived from the similar _Cyphinus_ (Pl. 39, fig. 14) by loss of the
medullary shell.

Subgenus 1. _Stylartella_, Haeckel.

_Definition._--On each pole of the main axis only one single large spine.

1. _Stylartus bipolaris_, n. sp. (Pl. 48, fig. 5).

Shell thick walled, rough; both its chambers nearly spherical, with
irregular, roundish pores, twice to four times as broad as the bars; eight
to ten on the half equator of each chamber. Polar spines very strong,
conical, as long as the greatest breadth.

_Dimensions._--Length of the shell (without spines) 0.18, greatest breadth
0.13; length of the polar spines 0.13, basal breadth 0.03.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

2. _Stylartus bicuspis_, n. sp.

Shell thin walled, smooth, with regular, circular pores, twice as broad as
the bars; twelve to fourteen on the half equator of each chamber. Polar
spines very stout, straight, three-sided pyramidal, half as long as the
shell.

_Dimensions._--Length of the shell 0.16, greatest breadth 0.12; length of
the polar spines 0.08, basal thickness 0.025.

_Habitat._--Pacific, central area. Station 266, depth 2750 fathoms.

Subgenus 2. _Stylartura_, Haeckel.

_Definition._--On each pole of the main axis a bunch of several spines.

{358}3. _Stylartus palatus_, n. sp.

Shell thick walled, thorny, with irregular, roundish pores, twice to three
times as broad as the bars; fifteen to eighteen on the half equator of each
chamber. On each pole of the main axis a large conical spine, surrounded by
a group of ten to fifteen smaller spines, one-third to two-thirds as long
as the breadth of the deep equatorial stricture.

_Dimensions._--Length of the shell 0.14, greatest breadth 0.09; length of
the polar spines 0.03 to 0.06, basal breadth 0.005 to 0.015.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

4. _Stylartus penicillus_, n. sp.

Shell thin walled, smooth, with irregular, roundish pores, three to four
times as broad as the bars; ten to twelve on the half equator of each
chamber. On each pole of the main axis a brush-like bunch of twenty to
thirty thin conical radial spines, half as long as the shell.

_Dimensions._--Length of the shell 0.15, greatest breadth 0.1; length of
the polar spines 0.08, basal breadth 0.005.

_Habitat._--Equatorial Atlantic, Station 347, surface.

Genus 154. _Cannartus_,[193] Haeckel, 1881, Prodromus, p. 462.

_Definition._--#Artiscida# with two hollow polar tubes (fenestrated
cylindrical or conical tubuli, opposite on the two poles of the main axis).

The genus _Cannartus_ differs from _Artiscus_ by the production of two
opposite hollow tubes on both poles of the main axis. The cavity of the
conical or cylindrical fenestrated tubes communicates freely with that of
the shell; the network in both is the same. The distal end of the tubes is
nearly always broken off, sometimes closed, with a conical apex.
_Cannartus_ can be derived either from _Pipettella_ by a transverse
equatorial constriction, or from _Cannartiscus_ by the loss of the
medullary shell, or from _Artiscus_ by the production of the polar tubes.

1. _Cannartus violina_, n. sp. (Pl. 39, fig. 10).

Pores of the shell subregular, circular, twice to three times as broad as
the bars; eighteen to twenty on the half meridian, fourteen to sixteen on
the half equator. Shell-wall in the "tropical zone" of both halves
thickened. Polar tubes nearly cylindrical, about as long as the main axis,
distal ends broken off.

_Dimensions._--Main axis 0.14, equatorial axis 0.09; meshes 0.008, bars
0.003; length of the polar tubes 0.12, breadth 0.02 to 0.03.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

{359}2. _Cannartus bitubulus_, n. sp.

Pores of the shell regular, circular, hexagonally framed, of the same
breadth as the bars; fourteen to sixteen on the half meridian, ten to
twelve on the half equator. Polar tubes cylindrical, longer than the main
axis, sulcated, distal ends broken off. (Similar to _Pipetta tuba_, Pl. 39,
fig. 7, but with equatorial stricture and without medullary shell.)

_Dimensions._--Main axis 0.16, equatorial axis 0.12; meshes and bars 0.006;
length of the polar tubes 0.2, breadth 0.025.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

3. _Cannartus biscottus_, n. sp.

Pores of the shell irregular, roundish, twice to three times as broad as
the bars; ten to twelve on the half meridian, six to eight on the half
equator. Polar tubes conical, shorter than the main axis, with closed
apex. (Similar to _Cannartidium bicinctum_, Pl. 39, fig. 18, but without
enclosed medullary shell.)

_Dimensions._--Main axis 0.11, equatorial axis 0.07; meshes 0.005 to 0.01,
bars 0.003; length of the polar tubes 0.08, breadth on the base 0.03.

_Habitat._--Equatorial Atlantic, Station 348, depth 2450 fathoms.

Family XV. #CYPHINIDA#, Haeckel (Pl. 39, figs. 11-19).

_Cyphinida_, Haeckel, 1881, Prodromus, p. 462.

_Definition._--#Prunoidea# with ellipsoidal twin-shell, divided by an
equatorial stricture into two communicating hemiellipsoidal or
hemispherical chambers; this external twin-shell (cortical shell) is either
simple or double, and encloses one or more internal concentric shells
(medullary shells). Central capsule ellipsoidal, commonly with an
equatorial constriction.

The family #Cyphinida# have the same characteristic twin-form of the
cortical shell as the Artiscida, but differ from them in the presence of a
simple or double internal medullary shell, connected with the cortical
shell by radial beams. The fenestrated shell is therefore composed of two
or more concentric shells as in the Druppulida, but differs from these in
the ring-like equatorial constriction.

_The Medullary Shell_, in the middle of the central capsule, is either
single or double, composed of two concentric shells. As in the Druppulida,
the form of the medullary shells is either spherical (Pl. 39, fig. 12_a_)
or lenticular, compressed from both poles of the main axis (Pl. 39, fig.
18_a_); sometimes the inner medullary shell is spherical, the outer
lenticular.

_The Radial Beams_, which connect the medullary shell with the equatorial
constriction of the cortical shell, lie either all in the equatorial plane
(Pl. 39, fig. 11) or near it on both sides (Pl. 39, figs. 12_a_, 18_a_).

{360}The _Cortical Shell_ is commonly simple (Pl. 39, figs. 12, 18),
sometimes composed of two concentric shells (Pl. 39, fig. 13), rarely of
three. As in the Artiscida, also in the Cyphinida, from both poles of the
main axis solid spines or hollow fenestrated tubes are often developed (Pl.
39, figs. 14, 16-18).

_The Central Capsule_ of the Cyphinida (Pl. 39, fig. 13) is of the same
form as in the Artiscida, generally ellipsoidal, but with an equatorial
ring-like stricture, which divides it into two equal halves. It encloses
the simple or double medullary shell, and is perforated by the radial beams
starting from this. From the inner surface of the surrounding cortical
shell it is separated by a thicker or thinner jelly-layer, the calymma.
(Compare also Taf. xxii. fig. 14 of my Monograph, 1862.)

_Synopsis of the Genera of Cyphinida._

-------------------------------------------------------------------------
Cortical shell without peculiar spines or hollow fenestrated tubes on
both poles of the main axis.
-------------------------------------------------------------------------
Cortical twin- {Medullary shell
shell simple { simple, 155. _Cyphanta_.
{
{Medullary shell
{ double, 156. _Cyphonium_.

Cortical twin- {Cortical shell
shell double { double, 157. _Cypassis_.
or triple; {
medullary {Cortical shell
shell double. { triple, 158. _Cyphocolpus_.
-------------------------------------------------------------------------
Cortical shell simple, with peculiar spines or hollow fenestrated tubes
on both poles of the main axis.
-------------------------------------------------------------------------
Two opposite {Medullary shell
polar spines { simple, 159. _Cyphinus_.
(or bunches {
of spines). {Medullary shell
{ double, 160. _Cyphinidium_.

Two opposite {Medullary shell
hollow { simple, 161. _Cannartiscus_.
fenestrated {
polar tubes. {Medullary shell
{ double, 162. _Cannartidium_.

Genus 155. _Cyphanta_,[194] n. gen.

_Definition._--#Cyphinida# with simple cortical shell and simple medullary
shell, without polar spines or tubes.

The genus _Cyphanta_ is the most simple of all Cyphinida, and can be
regarded as the common ancestral form of this family. It may be derived
phylogenetically from _Druppula_ by a ring-like constriction in the
equatorial plane of the ellipsoidal cortical shell, or from _Artiscus_ by
secondary formation of a central (spherical or ellipsoidal) medullary
shell.

Subgenus 1. _Cyphantella_, Haeckel.

_Definition._--Surface of the cortical shell smooth, without spines or
thorns.

{361}1. _Cyphanta colpodes_, n. sp.

Cortical shell thin walled, with smooth surface; its pores regular,
hexagonal, three to four times as broad as the bars; ten to twelve on the
half meridian, seven to eight on the half equator. Medullary shell
spherical, its diameter one-third of the equatorial axis of the cortical
shell. (The cortical shell of this species is nearly identical with the
central half of the cortical shell of _Panartus diploconus_, Pl. 40, fig.
1.)

_Dimensions._--Main axis of the cortical shell 0.12, equatorial axis 0.07;
meshes 0.013; bars 0.003; diameter of the medullary shell 0.025.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

2. _Cyphanta circopora_, n. sp.

Cortical shell thick walled, with smooth surface; pores subregular,
circular, twice as broad as the bars; eighteen to twenty on the half
meridian, twelve to fourteen on the half equator. Medullary shell
spherical, its diameter one-half of the equatorial axis of the cortical
shell. (The shell of this species is very similar to that of _Cannartiscus
amphiconiscus_, Pl. 39, fig. 19, but without the polar tubes of that
species.)

_Dimensions._--Main axis of the cortical shell 0.13, equatorial axis 0.08;
meshes 0.007, bars 0.003; diameter of the medullary shell 0.04.

_Habitat._--Pacific, central area. Station 266, depth 2750 fathoms.

3. _Cyphanta lævis_, Haeckel.

_Ommatospyris lævis_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 318.

Cortical shell thin walled, with smooth surface; pores of it irregular,
roundish, of very different size (some very large in the tropical circles
of both hemispheres). Pores twice to six times as broad as the bars; ten to
twelve on the half meridian, six to eight on the half equator. Medullary
shell spheroidal, compressed, its main axis somewhat shorter than its
equatorial axis, which attains half the length of that of the cortical
shell. (This species may be perhaps identical with _Ommatospyris lævis_ of
Ehrenberg, the diagnosis of which is insufficient and figure not given.)

_Dimensions._--Main axis of the cortical shell 0.1, equatorial axis 0.06;
meshes 0.005 to 0.02, bars 0.002 to 0.04; diameter of the medullary shell
0.03.

_Habitat._--Philippine Sea (Ehrenberg), Station 213, depths 2650 and 3300
fathoms.

4. _Cyphanta arachnoides_, n. sp.

Cortical shell very delicate and thin walled, cobweb-like, with smooth
surface; pores irregular, polygonal (mostly pentagonal or hexagonal), eight
to ten times as broad as the thread-like bars; ten to twelve on the half
meridian, six to eight on the half equator. Medullary shell spheroidal,
compressed, its main axis shorter than its equatorial axis, which is about
one-third that of the cortical shell. (This species is very similar to the
middle part of the shell of _Ommatocampe profundissima_, Ehrenberg, 1872,
Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. viii. fig. 6.)

{362}_Dimensions._--Main axis of the cortical shell 0.11, equatorial axis
0.06; meshes 0.01 to 0.02, bars 0.002; diameter of the medullary shell
0.02.

_Habitat._--Atlantic, Canary Islands (Lanzerote), surface.

Subgenus 2. _Cyphantissa_, Haeckel.

_Definition._--Surface of the cortical shell spiny, everywhere scattered
with numerous thorns or spines.

5. _Cyphanta hispida_, n. sp.

Cortical shell thick walled, with spiny surface; pores regular, circular,
with hexagonal frame, twice as broad as the bars; twelve to fourteen on the
half meridian, eight to ten on the half equator. From the corners of the
hexagonal frames (between every three pores) arise short, straight, conical
radial spines, somewhat longer than the breadth of the pores. Medullary
shell subspherical, its diameter about one-third of the equatorial axis of
the cortical shell. (The appearance of the cortical shell is the same as
that of the middle part of _Peripanartus amphiconus_, Pl. 40, fig. 5.)

_Dimensions._--Main axis of the cortical shell 0.12, equatorial axis 0.07;
meshes 0.01, bars 0.005; length of the spines 0.012; diameter of the
medullary shell 0.02.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

6. _Cyphanta hystrix_, n. sp.

Cortical shell thin walled, with thorny surface; pores irregular, roundish,
of very unequal size, twice to six times as broad as the bars; twelve to
eighteen on the half meridian, eight to thirteen on the half equator.
Between the pores irregularly scattered, bristle-like, thin spines about
the same size, partly straight, partly oblique, rising from the surface.
Medullary shell lenticular, compressed, its equatorial axis one and a half
times the length of the main axis, and about half that of the cortical
shell.

_Dimensions._--Main axis of the cortical shell 0.15, equatorial axis 0.09;
meshes 0.004 to 0.02, bars 0.003 to 0.006; length of the spines 0.02;
diameter of the medullary shell 0.04.

_Habitat._--South Atlantic, Station 318, surface.

Genus 156. _Cyphonium_,[195] n. gen.

_Definition._--#Cyphinida# with simple cortical shell and double medullary
shell, without polar spines or tubes.

The genus _Cyphonium_ contains a number of very common species, among which
are the earliest known forms of this family, partly described by Ehrenberg
as Ommatospyris {363}(which genus contains also a number of other
#Prunoidea#), partly by me (1862) as _Didymocyrtis_. Both names are
inadequate, as allusions to quite different families of Nassellaria, but
may be retained as significations of subgeneric divisions. _Cyphonium_
differs from _Cyphanta_ by the double medullary shell, which is either
spherical or lenticular.

Subgenus 1. _Ommatospyris_, Ehrenberg (_partim_).

_Definition._--Surface of the cortical shell smooth, without thorns or
spines.

1. _Cyphonium coscinoides_, n. sp.

_Ommatospyris coscinoides_, Haeckel, 1881, Prodromus, p. 462.

Cortical shell thin walled, with smooth surface; pores regular, hexagonal,
three to four times as broad as the bars; five to six on the half meridian,
eight to nine on the half equator of each chamber. Medullary shells both
spherical. (The cortical shell of this species is nearly the same as that
of _Cyphanta colpodes_, and as the middle part of the cortical shell of
_Panartus diploconus_, Pl. 40, fig. 1.)

_Dimensions._--Main axis of the cortical shell 0.11, equatorial axis 0.06;
greatest breadth of the chambers 0.08; pores 0.013, bars 0.003.

_Habitat._--North Pacific, Station 244, surface.

2. _Cyphonium virgineum_, n. sp. (Pl. 39, figs. 12, 12_a_).

_Ommatospyris virginea_, Haeckel, 1881, Prodromus et Atlas (pl. xxxix.
fig. 12_a_).

Cortical shell thick walled, with smooth surface (sometimes a little rough
with very small thorns); pores subregular, circular, with hexagonal
elevated frames, about as broad as the bars; ten to twelve on the half
meridian, sixteen to eighteen on the half equator of each chamber.
Medullary shells (fig. 12_a_) both spherical. (Sometimes, as in the figured
specimen, the shell is somewhat irregular, an individual abnormality.)

_Dimensions._--Main axis of the cortical shell 0.13 to 0.15; equatorial
axis of the structure 0.08 to 0.09, of each chamber 0.1 to 0.12, pores and
bars 0.005 to 0.007.

_Habitat._--Pacific, western tropical part, Station 225, depth 4475
fathoms.

3. _Cyphonium ethmarium_, n. sp.

_Ommatospyris ethmaria_, Haeckel, 1881, Prodromus, p. 462.

Cortical shell thin walled, with quite smooth surface; pores subregular,
circular (without hexagonal frame), twice as broad as the bars; nine to ten
on the half meridian of each chamber, sixteen to eighteen on its half
equator. Medullary shells both spherical. (This species resembles the
proximal internal chambers of _Peripanartus atractus_, Pl. 40, fig. 7.)

_Dimensions._--Main axis 0.13, equatorial axis 0.07; greatest breadth 0.09;
pores 0.006, bars 0.003.

_Habitat._--Equatorial Atlantic, Station 347, surface.

{364}4. _Cyphonium trinacrium_, n. sp.

Cortical shell thick walled, with rough surface; pores subregular, circular
(without hexagonal frame), not broader than the bars; five to six on the
half meridian of each chamber, nine to ten on its half equator. Medullary
shells both compressed, lenticular. This species resembles _Cypassis
entomocora_ vel _Ommatocampe trinacria_, Stöhr, 1880, _loc. cit._, p. 90,
Taf. ii. fig. 1, but has not its external mantle. It may be the ancestral
form of it (both in an ontogenetic and phylogenetic sense).

_Dimensions._--Main axis 0.11, equatorial axis 0.07; greatest breadth 0.08;
pores 0.005, bars 0.005.

_Habitat._--Fossil in Tertiary rocks of Sicily, Caltanisetta, Haeckel
(Grotte, Stöhr?).

5. _Cyphonium diattus_, n. sp.

Cortical shell thin walled, with quite smooth surface; pores irregular,
polygonal, mostly pentagonal or hexagonal, three to six times as broad as
the bars; six to seven on the half meridian of the chamber, ten to twelve
on its half equator. Medullary shells both spherical. (Resembles _Cyphonium
profundum_, Ehrenberg, 1872, _loc. cit._, Taf. x. fig. 5, but does not
possess the spines of the surface.)

_Dimensions._--Main axis 0.13, equatorial axis 0.06; greatest breadth 0.08;
pores 0.005 to 0.012, bars 0.002 to 0.004.

_Habitat._--Indian Ocean, western part (Zanzibar), Pullen, depth 2200
fathoms.

6. _Cyphonium mammarium_, n. sp.

Cortical shell thick walled, with smooth surface (sometimes a little
rough); pores irregular, roundish, twice to three times as broad as the
bars; eleven to twelve on the half meridian of each chamber, sixteen to
nineteen on its half equator. Internal medullary shell spherical, external
lenticular compressed, sometimes both spherical or both compressed.
(Resembles the internal cortical twin-shell of _Cyphocolpus virginis_, Pl.
40, fig. 11.)

_Dimensions._--Main axis 0.14, equatorial axis 0.08, greatest breadth 0.09;
pores 0.004 to 0.006, bars 0.002.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Subgenus 2. _Ommatocyrtis_, Haeckel.

_Definition._--Surface of the cortical shell thorny or spiny.

7. _Cyphonium hexagonium_, n. sp.

_Didymocyrtis hexagonia_, Haeckel, 1881, Prodromus.

Cortical shell thin walled, with spiny surface; pores regular or
subregular, hexagonal, twice to three times as broad as the bars; five to
six on the half meridian, nine to ten on the half equator {365}of each
chamber. Radial spines between them conical, about as long as the pores.
Medullary shells both spherical.

_Dimensions._--Main axis 0.14, equatorial axis 0.08; greatest breadth of
each chamber 0.1; pores 0.012, bars 0.004.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

8. _Cyphonium facettarium_, n. sp.

_Didymocyrtis facettaria_ Haeckel, 1881, Prodromus.

Cortical shell thick walled, with spiny surface; pores subregular,
circular, with elevated hexagonal frames, twice as broad as the bars; eight
to nine on the half meridian, fourteen to sixteen on the half equator of
each chamber. In the corners of the hexagons (between every three meshes)
arise radial spines, about twice as long as the pores. Both medullary
shells spheroidal compressed. (This species is nearly identical with the
internal cortical twin-shell of _Peripanartus amphiconiscus_, Pl. 40, fig.
5.)

_Dimensions._--Main axis 0.12, equatorial axis 0.07; greatest breadth of
both chambers 0.09; pores 0.01, bars 0.005, spines 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

9. _Cyphonium cribellum_, n. sp. (Pl. 39, fig. 11).

Cortical shell thick walled, with spiny surface; pores regular, circular,
about as broad as the bars; seven to eight on the half meridian of each
chamber, thirteen to fifteen on its half equator. Radial spines between
them conical, twice as long as the pores. Medullary shells both lenticular.

_Dimensions._--Main axis 0.11, equatorial axis 0.06; greatest breadth 0.08;
pores and bars 0.004.

_Habitat._--North Atlantic, Station 353, surface.

10. _Cyphonium profundum_, Haeckel.

_Ommatospyris profunda_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 297, Taf. viii. fig. 5, Taf. x. fig. 5.

Cortical shell thin walled, with thorny surface; pores irregular,
polygonal, three to six times as broad as the thin bars; five to six on the
half meridian of each chamber, ten to twelve on its half equator. Thorns
of the surface short. Medullary shells both spherical.

_Dimensions._--Main axis 0.1, equatorial axis 0.06; greatest breadth 0.07;
pores 0.005 to 0.012, bars 0.002, spines 0.005.

_Habitat._--Pacific, tropical zone; Philippine Sea, depth 3300 fathoms,
Ehrenberg; Stations 266 to 274, depth 2350 to 2925 fathoms.

{366}11. _Cyphonium ceratospyris_, Haeckel.

_Didymocyrtis ceratospyris_, Haeckel, 1862, Monogr. d. Radiol., p. 445,
pl. xxii. figs. 14-16.

_Haliomma didymocyrtis_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 816.

Cortical shell thin walled, with spiny surface; pores irregular, roundish,
three to nine times as broad as the bars; five to six on the half meridian
of each chamber, eight to nine on its half equator. Spines of the surface
partly very short, partly as long as the largest pores, either radially or
obliquely inserted. Medullary shells both spherical. (Compare the detailed
description and figures also of the soft body in my Monograph, _loc. cit._)

_Dimensions._--Main axis of the cortical shell 0.12, equatorial axis (in
the stricture) 0.06; greatest breadth 0.08; pores 0.03 to 0.2, bars 0.002
to 0.006, spines 0.05 to 0.015.

_Habitat._--Mediterranean (Messina), Canary Islands (Lanzerote), Haeckel.

Genus 157. _Cypassis_,[196] n. gen.

_Definition._--#Cyphinida# with double cortical shell and double medullary
shell, without polar spines or tubes.

The genus _Cypassis_ differs from _Cyphonium_ by duplication of the
cortical twin-shell; from the outer surface of the simple twin-shell arise
numerous radial spines, which become connected by anastomosing transverse
branches, and in this manner form an outer envelope or mantle. _Cypassis_
may also be regarded as a _Cromyodruppa_, the double cortical shell of
which is constricted in the equatorial plane.

Subgenus 1. _Didymospyris_, Haeckel (1881).

_Definition._--Surface of the shell smooth, without thorns or spines.

1. _Cypassis palliata_, n. sp.

Inner cortical shell thin walled, with regular, circular,
hexagonally-framed pores, twice as broad as the bars; seven to eight on the
half meridian of each chamber, ten to twelve on its half equator. Outer
cortical shell very thin, with smooth surface, and very small, irregular,
roundish pores, twice to five times smaller than those of the inner shell.
Distance between the two cortical shells equals the diameter of the outer
medullary shell, which, like the inner, is spherical. (The network of this
species resembles that of _Peripanartus amphiconiscus_, Pl. 40, fig. 5.)

_Dimensions._--Main axis of the external cortical shell 0.18, of the
internal 0.12; greatest breadth (in the equator of each chamber) of the
former 0.13, of the latter 0.09; pores of the outer 0.002 to 0.005, of the
inner shell 0.01, bars 0.002 to 0.005.

_Habitat._--South Pacific, Station 288, surface.

{367}2. _Cypassis eucolpos_, n. sp.

Inner cortical shell thick walled, with regular, circular pores, twice as
broad as the bars (without hexagonal frames); nine to ten on the half
meridian of each chamber, fifteen to seventeen on its half equator. Outer
cortical shell very thin, with smooth surface, and very small, irregular,
roundish pores, twice to three times smaller than those of the inner shell.
Distance between the two cortical shells about one and a half times the
diameter of the inner medullary shell, which, like the outer, is spherical.
(The inner cortical shell resembles that of _Cannartiscus amphiconiscus_,
Pl. 39, fig. 19, but without polar tubes.)

_Dimensions._--Main axis of the external cortical shell 0.2, of the
internal 0.15; greatest breadth of the former 0.16, of the latter 0.12;
pores of the outer 0.002, of the inner 0.006, bars 0.003.

_Habitat._--South Atlantic, Station 319, surface.

Subgenus 2. _Didymocyrtis_, Haeckel (1862).

_Definition._--Surface of the cortical shell thorny or spiny.

3. _Cypassis entomocora_, Haeckel.

_Astromma entomocora_, Ehrenberg, 1847, Mikrogeol., Taf. xxii. fig. 32.

_Haliomma didymum_, Ehrenberg, 1844, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 83.

? _Haliomma amphisiphon_, Ehrenberg, 1844, Monatsber. d. k. Akad. d.
Wiss. Berlin, p. 267.

_Ommatospyris entomocora_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, pp. 66, 74.

? _Ommatocampe trinacria_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 90,
Taf. ii. fig. 1.

_Didymospyris entomocora_, Haeckel, 1881, Prodromus.

_Didymocyrtis entomocora_, Haeckel, 1862, Monogr. d. Radiol., p. 445.

Inner cortical shell thick walled, with regular, circular pores, twice as
broad as the bars (without hexagonal frames); eight to nine on the half
meridian of each chamber, ten to eleven on its half equator. Outer cortical
shell thin walled, with regular, circular pores like those of the inner.
Distance between the two cortical shells equals about twice the diameter of
one pore (or the short axis of the lenticular double medullary shell).
Surface covered with short conical spines (in the figure of Ehrenberg
broken off).

_Dimensions._--Main axis of the external cortical shell 0.2, of the
internal 0.15; greatest breadth of the former 0.13, of the latter 0.1;
pores of the outer 0.005 to 0.01, of the inner 0.01, bars 0.004; length of
the surface spines 0.005 to 0.01.

_Habitat._--Fossil in the Tertiary rocks of Barbados, Bermuda, and Sicily
(Caltanisetta); also living in the greatest depth of the Pacific, Stations
268, 225, depth 2900 to 4475.

4. _Cypassis puella_, n. sp. (Pl. 39, fig. 13).

_Didymospyris colpodes_, Haeckel, 1881, Prodromus et Atlas (pl. xxxix.,
fig. 13).

Internal cortical shell thin walled, with irregular, circular pores of very
different size, once to four times as broad as the bars; fourteen to
fifteen on the half meridian of each chamber, eighteen {368}to twenty on
its half equator. Outer cortical shell thin walled, with irregular,
circular pores, only one-third to one-half as large as those of the inner.
Distance between the two shells equals the largest diameter of the double
lenticular medullary shell. Surface of both cortical shells covered with
short conical spines (not longer than the largest pores).

_Dimensions._--Main axis of the external cortical shell 0.2, of the
internal 0.15, of the central capsule 0.1; greatest breadth of the first
0.14, of the second 0.1, of the third 0.07; pores of the outer shell 0.001
to 0.005, of the inner 0.002 to 0.01, bars 0.002 to 0.003; length of the
surface spines 0.005 to 0.01.

_Habitat._--Pacific, central area; Stations 266 to 274, surface; Atlantic,
Canary Islands (Haeckel), Station 354, surface.

5. _Cypassis halicora_, n. sp.

Internal cortical shell thin walled, with irregular, roundish pores, once
to three times as broad as the bars; ten to eleven on half meridian of each
chamber, fourteen to sixteen on its half equator. Outer cortical shell thin
walled, with very delicate network, spindle-like, with conical
prolongations at both poles. Distance between the two shells larger than
the greatest diameter of the double lenticular medullary shell. Surface of
both cortical shells covered with innumerable small thorns. (Resembles
closely _Cyphocolpus virginis_, Pl. 40, fig. 11, but without the third
shell.)

_Dimensions._--Main axis of the external cortical shell 0.21, of the
internal 0.14, greatest breadth of the former 0.13, of the later 0.09;
pores 0.003 to 0.009; bars of the outer shell 0.001, of the inner 0.004.

_Habitat._--Indian Ocean, Ceylon (Haeckel), surface.

Genus 158. _Cyphocolpus_,[197] n. gen.

_Definition._--#Cyphinida# with triple cortical shell and double medullary
shell, without polar spines or tubes.

The genus _Cyphocolpus_ differs from both foregoing genera by the increased
number of the cortical twin-shells. Whilst these are simple in _Cyphonium_,
double in _Cypassis_, they are triple in _Cyphocolpus_, composed of three
concentric envelopes. The three genera named represent a phylogenetic
series, which is repeated in the ontogenetic development of _Cyphocolpus_.

1. _Cyphocolpus didymus_, n. sp.

Inner cortical shell with circular, subregular pores, twice as broad as the
bars; five to six on the half meridian of each chamber, eight to ten on its
half equator. Middle cortical shell also with subregular, circular pores of
the same size. Outer cortical shell with smooth surface, with more
irregular, roundish pores of very different size. The distance between
every two shells equals the diameter of the inner medullary shell, which,
like the outer, is spheroidal.

{369}_Dimensions._--Main axis of the outer cortical shell 0.3, of the
middle 0.22, of the inner 0.15; greatest breadth of the inner cortical
shell 0.1, its pores 0.01, bars 0.005.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

2. _Cyphocolpus virginis_, n. sp. (Pl. 40, fig. 11).

_Zygartus virginis_, Haeckel, 1881, Prodromus et Atlas (pl. xl. fig. 11).

Inner cortical shell with irregular, roundish pores, twice to three times
as broad as the bars; eleven to twelve on the half meridian of each
chamber, sixteen to eighteen on its half equator. Middle cortical shell
with very delicate network, its irregular, polygonal pores of the same size
as those of the inner, but separated by extremely thin bars. Outer cortical
shell with spiny surface, and with more irregular, roundish pores of
different size. The distance between each two shells is variable, and
attains more than the diameter of the outer medullary shell, which, like
the inner, is spheroidal, slightly compressed.

_Dimensions._--Main axis of the outer cortical shell 0.26, of the middle
0.22, of the inner 0.14; greatest breadth of the inner cortical shell 0.09,
its pores 0.004 to 0.006, bars 0.002.

_Habitat._--Pacific, central area, Station 271, surface.

Genus 159. _Cyphinus_,[198] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Cyphinida# with simple cortical shell and simple medullary
shell, with two opposite polar spines (or bunches of polar spines) on the
poles of the main axis.

The genus _Cyphinus_ differs from _Cyphanta_ by the development of two
opposite spines (or bunches of spines) on both poles of the main axis. It
simulates therefore the bipolar formation of _Stylartus_, and differs from
this in the possession of a medullary shell.

Subgenus 1. _Cyphinoma_, Haeckel.

_Definition._--On both poles of the main axis only one single large spine.

1. _Cyphinus amphacanthus_, n. sp.

Cortical shell with smooth surface, and circular, regular pores, twice as
broad as the bars; five to six pores on the half meridian of each chamber,
ten to twelve on its half equator. On both poles of the main axis, one
single, very strong, conical spine, about half as long as this axis, and
half as broad at the base as the spherical medullary shell.

_Dimensions._--Main axis of the cortical shell (without spines) 0.2,
greatest breadth (in the equator of each chamber) 0.1; pores 0.006, bars
0.003; length of the polar spines 0.1, basal thickness of them 0.02.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{370}2. _Cyphinus dixiphus_, n. sp.

Cortical shell with rough surface, and irregular, roundish pores, twice to
four times as broad as the bars; eight to nine pores on the half meridian
of each chamber, twelve to fourteen on its half equator. On both poles of
the main axis, one single, strong, conical spine, with prominent edges at
the base, and nearly as broad as the spherical medullary shell, nearly as
long as the main axis.

_Dimensions._--Main axis (without spines) 0.18, greatest breadth 0.13;
pores 0.003 to 0.008, bars 0.002; length of the polar spines 0.16, basal
thickness 0.03.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Subgenus 2. _Cyphinura_, Haeckel.

_Definition._--On both poles of the main axis a bunch or circle of several
spines.

3. _Cyphinus amphilophus_, n. sp. (Pl. 39, fig. 14).

_Cyphinidium amphilophus_, Haeckel, 1882, Atlas (pl. xxxix. fig. 14).

Cortical shell thick walled, with rough surface, and subregular, circular
pores of different size, twice to five times as broad as the bars; nine to
ten pores on the half meridian of each chamber, twelve to fourteen on its
half equator. On both poles of the main axis a bunch of six to eight strong
conical spines, the largest of which are about as long as the greatest
breadth of the shell; their basal part is furrowed, and nearly half as
thick as the spherical medullary shell.

_Dimensions._--Main axis of the cortical shell (without spines) 0.18 to
0.2, greatest breadth 0.1 to 0.12; pores 0.004 to 0.01, bars 0.002; length
of the polar spines 0.04 to 0.12, basal thickness 0.02 to 0.04.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

4. _Cyphinus penicillatus_, Haeckel.

_Ommatospyris penicillata_, Ehrenberg, 1872, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 318; Abhandl. d. k. Akad. d. Wiss. Berlin, Taf.
viii. fig. 4.

Cortical shell thin walled, with rough surface, and irregular, polygonal
pores, six to eight times as broad as the bars; five to six pores on the
half meridian of each chamber, nine to ten on its half equator. Around both
poles of the main axis a polar circle of eight to ten divergent radial
spines, scarcely thicker than the bars, and about as long as the diameter
of the spherical medullary shell. (In the figure of Ehrenberg the network
is too thin and the spines too short.)

_Dimensions._--Main axis (without spines) 0.1, greatest breadth 0.09; pores
0.008 to 0.014, bars 0.001 to 0.002; length of the spines 0.02 to 0.04.

_Habitat._--Pacific, Philippine Sea, 3300 fathoms, Ehrenberg; Station 206,
depth 2100 fathoms.

{371}Genus 160. _Cyphinidium_,[199] n. gen.

_Definition._--#Cyphinida# with simple cortical shell and double medullary
shell, with two opposite polar spines (or bunches of polar spines) on the
poles of the main axis.

The genus _Cyphinidium_ differs from _Cyphinus_ only in the duplication of
the medullary shell, which is composed of two concentric spheres or
somewhat compressed lenticular spheroids. It exhibits therefore the same
relation to _Cyphinus_ that _Stylatractus_ in the Druppulida bears to
_Lithatractus_. Possibly the two former genera are derived from the two
latter by an annular constriction in the equatorial plane.

Subgenus 1. _Cyphinidoma_, Haeckel.

_Definition._--On both poles of the cortical shell one single polar spine
or a bunch of polar spines.

1. _Cyphinidium amphistylium_, n. sp.

Cortical shell thick walled, with smooth surface, and subregular, circular
pores, twice to three times as broad as the bars; eight to ten on the half
meridian, sixteen to eighteen on the half equator of each chamber. Both
medullary shells spherical. The cortical shell resembles very much that of
_Cannartiscus amphiconiscus_ (Pl. 39, fig. 19), but exhibits instead of the
hollow polar tubes two strong solid polar spines of conical form, half as
long as the main axis, and as broad at the base as the inner medullary
shell.

_Dimensions._--Main axis of the cortical shell (without polar spines) 0.14,
greatest breadth 0.1; pores 0.008, bars 0.004; length of the polar spines
0.08, basal thickness 0.02.

_Habitat._--North Pacific, off Japan, Station 241, depth 2300 fathoms.

2. _Cyphinidium apicatum_, Haeckel.

_Ommatospyris apicata_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 317.

Cortical shell thin walled, with thorny surface, and irregular, polygonal
pores, three to five times as broad as the thin bars; eight to nine on the
half meridian, thirteen to fifteen on the half equator of each chamber. On
both poles a bunch of four to eight strong conical spines, the central of
which (in the main axis) is much larger than the others, and twice as long
as the diameter of the outer spheroidal medullary shell; size and number of
the polar spines is very variable (commonly five to six).

{372}_Dimensions._--Main axis (without spines) 0.12, greatest breadth 0.08;
pores 0.005 to 0.01, bars 0.002; length of the polar spines 0.02 to 0.06,
basal thickness 0.01.

_Habitat._--Northern Pacific, Californian Sea, depth 2500 fathoms,
Ehrenberg; Station 253, depth 3125 fathoms.

Subgenus 2. _Cyphinidura_, Haeckel.

_Definition._--On both poles of the cortical shell, a circle of divergent
polar spines.

3. _Cyphinidium coronatum_, n. sp.

Cortical shell with thorny surface, and irregular, roundish pores, twice to
four times as broad as the bars; seven to eight on the half meridian,
eleven to thirteen on the half equator of each chamber. Around both poles
of the main axis occurs a circle of ten to twelve strong, radially
divergent spines, about as long as the diameter of the outer medullary
shell, which, like the inner, is spheroidal. (This species is identical
with the younger developmental stage of _Panicium coronatum_, Pl. 40, fig.
4, before the formation of two distal chambers.)

_Dimensions._--Main axis 0.12, greatest breadth 0.1; pores 0.008 to 0.016,
bars 0.004; length of the polar spines 0.02, thickness 0.005.

_Habitat._--Pacific, central area, Stations 270 to 274, depth 2350 to 2925
fathoms.

Genus 161. _Cannartiscus_,[200] n. gen.

_Definition._--#Cyphinida# with simple cortical shell and simple medullary
shell, with two hollow fenestrated polar tubes, opposite on both poles of
the main axis.

The genus _Cannartiscus_ differs from _Cyphanta_ in the development of two
hollow fenestrated tubes, opposite on both poles of the main axis. It
simulates therefore the characteristic formation of _Cannartus_, and
differs from this Artiscid in the possession of a medullary shell.

1. _Cannartiscus amphiconiscus_, n. sp. (Pl. 39, fig. 19).

_Cannartidium amphiconiscus_, Haeckel, 1882, Atlas (pl. xxxix. fig. 19).

Cortical shell thick walled, smooth, with regular, circular pores, twice as
broad as the bars; nine to ten on the half meridian, sixteen to eighteen on
the half equator of each chamber. Polar tubes conical, about as long as one
single chamber, on the base somewhat broader than the simple spherical
medullary shell. Pores of the tubes only one-third as great as those of the
chambers.

_Dimensions._--Main axis of the cortical shell (without tubes) 0.14,
greatest breadth 0.1; {373}pores 0.008, bars 0.004. Length of the polar
tubes 0.07, basal breadth 0.04; pores 0.003, bars 0.002.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

2. _Cannartiscus amphicylindrus_, n. sp.

Cortical shell thick walled, rough, with subregular, circular pores, twice
to four times as broad as the bars; six to seven on the half meridian, ten
to twelve on the half equator of each chamber. Polar tubes cylindrical, on
the distal end open (broken off?) nearly as long as the main axis, somewhat
narrower than the spherical medullary shell. Pores of the tubes much
smaller than those of the chambers.

_Dimensions._--Main axis (without tubes) 0.17, greatest breadth 0.12; pores
0.006 to 0.012, bars 0.003. Length of the polar tubes 0.15, breadth of them
0.03; pores 0.003, bars 0.002.

_Habitat._--Pacific, central area, Station 268, 2900 fathoms; the same form
occurs fossil in the rocks of Barbados.

Genus 162. _Cannartidium_,[201] n. gen.

_Definition._--#Cyphinida# with simple cortical shell and double medullary
shell, with two hollow fenestrated polar tubes, opposite on both poles of
the main axis.

The genus _Cannartidium_ differs from _Cannartiscus_ and _Cannartus_ in the
duplication of the medullary shell, which is composed of two concentric
spheres or compressed lenticular spheroids. The three genera named form
therefore one morphological series, with identical cortical shell, and only
differing in the absence or presence of a simple or double medullary shell.

Subgenus 1. _Cannartidella_, Haeckel.

_Definition._--Surface of the cortical shell smooth or a little rough, but
without spines or fenestrated protuberances.

1. _Cannartidium amphiconicum_, n. sp.

Cortical shell thick walled, smooth with regular circular pores twice as
broad as the bars; eight to nine on the half meridian, fifteen to sixteen
on the half equator of each chamber. Polar tubes conical, with smaller
pores, tapering towards the closed apex, about as long as one single
chamber, at the base one-third as broad as the equatorial constriction.
Both concentric medullary shells spherical. (Nearly identical with
_Cannartiscus amphiconiscus_, Pl. 39, fig. 19, but differs in the
possession of the double medullary shell.)

{374}_Dimensions._--Main axis of the cortical shell (without tubes) 0.15,
greatest breadth 0.11; pores 0.009, bars 0.004; length of the polar tubes
0.08, basal breadth 0.04.

_Habitat._--Pacific, western tropical part, Station 225, depth 4475
fathoms.

2. _Cannartidium amphicanna_, Haeckel.

"Curious twin Polycystin," Bury, 1862, Polycystins of Barbados, pl. xx.
fig. 4.

Cortical shell thick walled, rough, with regular, circular pores, not
broader than the bars; six to seven on the half meridian, ten to twelve on
the half equator of each chamber. Polar tubes cylindrical, at the distal
end open (broken off?), nearly as long as the main axis, about one-third as
broad as the equatorial constriction. Both concentric medullary shells
spherical.

_Dimensions._--Main axis 0.14, greatest breadth 0.11; pores and bars 0.005;
length of the polar tubes 0.13, breadth of them 0.03.

_Habitat._--Fossil in the Barbados deposits.

3. _Cannartidium amphisiphon_, Haeckel.

? _Haliomma amphisiphon_, Ehrenberg, 1844, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 267.

Cortical shell thick walled, rough, with irregular, roundish pores, twice
to five times as broad as the bars; five to six on the half meridian, nine
to ten on the half equator of each chamber. Polar tubes conical, with
smaller pores, about as long as one single chamber, half as broad at the
base as the equatorial constriction. (This deep-sea form is probably
identical with that fossil species which Ehrenberg, in 1844, described as
_Haliomma amphisiphon_, and which he afterwards, in 1875, erroneously
identified with his _Astromma entomocora_ (= _Cypassis entomocora_). But
the figure given of the latter is quite different from the given diagnosis
of the former.)

_Dimensions._--Main axis of the cortical shell (without tubes) 0.13,
greatest breadth 0.1; pores 0.006 to 0.12, bars 0.003; length of the polar
tubes 0.07, basal breadth of them 0.04.

_Habitat._--Tropical Atlantic, Antilles, Station 24, depth 390 fathoms;
fossil in the Tertiary rocks of Bermuda.

4. _Cannartidium bicinctum_, n. sp. (Pl. 39, fig. 18).

Cortical shell thick walled, with a thickened ring-shaped protuberance in
the greatest breadth of both chambers. Pores subregular, circular, twice to
three times as broad as the bars; seven to eight on the half meridian,
thirteen to fifteen on the half equator of each chamber. Polar tubes
conical, with smaller pores, somewhat shorter than the greatest breath,
nearly as thick at the base as the outer spheroidal medullary shell (fig.
18_a_).

_Dimensions._--Main axis (without tubes) 0.12, greatest breadth 0.08; pores
0.003 to 0.006, bars 0.002; length of the polar tubes 0.08, basal thickness
0.03.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

{375}Subgenus 2. _Cannartidissa_, Haeckel.

_Definition._--Surface of the cortical shell with conical fenestrated
protuberances in the greatest breadth of both chambers.

5. _Cannartidium mammiferum_, n. sp. (Pl. 39, fig. 16).

Cortical shell-thin walled, having a circle of six to eight conical
protuberances in the greatest breadth of both chambers, each of which
exhibits a central apical pore, surrounded by a circle of six to eight
oblique larger pores. Between the protuberances occur nine to ten pores on
the half meridian of each chamber, circular, subregular, twice to three
times as broad as the bars. Polar tubes cylindrical, conical at the closed
end, with smaller pores, about as long as the greatest breadth of the
chambers, as broad as the spherical outer medullary shell.

_Dimensions._--Main axis (without tubes) 0.13, greatest breadth (including
the protuberances) 0.1; pores 0.005 to 0.008, bars 0.003; length of the
polar tubes 0.09, basal thickness 0.02.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

6. _Cannartidium mastophorum_, n. sp. (Pl. 39, fig. 17).

Cortical shell thin walled, of the same form and structure as in the
foregoing species, differs from this mainly in the form of the polar tubes,
which are not cylindrical but conical, tapering gradually from the base
towards the closed apex. Besides this, the conical protuberances (six in
the greatest periphery of each chamber) are more regular and acute, with
smaller and more numerous pores. Both medullary shells are here also
spherical.

_Dimensions._--Main axis (without tubes) 0.14, greatest breadth (including
the protuberances) 0.13; pores 0.006 to 0.009, bars 0.004; length of the
polar tubes 0.07, basal thickness 0.03.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Family XVI. #PANARTIDA#, Haeckel (Pl. 40, figs. 1-9).

_Definition._--#Prunoidea# with a four-jointed cortical shell, the external
shell being divided by three parallel transverse constrictions into four
chambers, in the centre enclosing two internal concentric shells (medullary
shells). Central capsule cylindrical, commonly four-jointed (with three
transverse annular constrictions).

The family #Panartida# (Pl. 40, figs. 1-9) is characterised by its
four-jointed cortical shell, constantly composed of four chambers, lying
one behind another in the elongated main axis. They are separated by three
ring-like constrictions lying in three parallel transverse planes; the
middle of these is the equatorial plane. In the centre of this latter lies
constantly a double medullary shell.

The Panartida must be derived from the Cyphinida by further increase in the
longitudinal or main axis, and by repetition of the equatorial constriction
in two transverse planes parallel to it and at an equal distance from it.

{376}_The Medullary Shell_ is always double in this family, composed of two
small concentric shells lying in the middle of the central capsule. They
are either spherical or lenticular, compressed towards both poles of the
main axis; sometimes the inner is spherical, the outer lenticular. From the
equatorial circumference of the latter starts a number of radial beams,
which perforate the central capsule, and are inserted into the equatorial
constriction of the cortical shell. Commonly all radial beams lie in the
equatorial plane, sometimes also a part of them on both sides of it.

_The Cortical Shell_ is constantly composed of four completely latticed
chambers, which are only separated by the three annular constrictions. In
many species all four chambers have the same form and size (Pl. 40, fig.
3), but in the greater number of species both proximal (or central)
chambers are different in size, shape, or structure from both distal (or
polar) chambers. The cortical shell is either simple or double, composed of
two concentric four-jointed shells; rarely it is triple or multiple,
composed of three or more shells fitting one inside the other. Very often
the outer cortical shell is incomplete, and only developed around the two
proximal chambers of the complete inner cortical shell, both distal
chambers of the latter remaining simple.

On both poles of the main axis often are developed solid spines
(_Panicium_, Pl. 40, fig. 4) or hollow fenestrated tubes (_Panarium_, Pl.
40, fig. 9), as in the foregoing families. Besides this, the surface of the
cortical shell may be armed with spines.

_The Central Capsule_ of the Panartida is always cylindrical, on both poles
hemispherical; commonly it is more or less distinctly four-jointed, with
three ring-like constrictions corresponding to those of the enclosing
cortical shell. From the inner surface of the latter it is separated by a
thinner or thicker jelly-mantle.

_Synopsis of the Genera of Panartida._

On both poles of the main { Cortical shell simple, 163. _Panartus_.
axis neither peculiar {
polar spines nor { Cortical shell double or
fenestrated tubes. { triple, 164. _Peripanartus_.

On both poles of the main { Cortical shell simple, 165. _Panicium_.
axis a large apical {
spine or a group of { Cortical shell double or
polar spines. { triple, 166. _Peripanicium_.

On both poles of the main { Cortical shell simple, 167. _Panarium_.
axis a conical or {
cylindrical fenestrated { Cortical shell double or
tube. { triple, 168. _Peripanarium_.

Genus 163. _Panartus_,[202] n. gen.

_Definition._--#Panartida# with simple cortical shell and double medullary
shell, without polar spines or tubes on both poles of the main axis.

{377}The genus _Panartus_ represents the common ancestral form of all
Panartida, as all other genera of this subfamily must be derived from it
and are only further developmental stages, from an ontogenetic as well as
from a phylogenetic point of view. The cortical shell of _Panartus_ is
constantly composed of four fenestrated chambers jointed to one another in
the main axis; both proximal chambers are separated from one another by the
equatorial ring-like constriction, in the centre of which lies the double
medullary shell; both distal chambers are separated from the former by two
other annular constrictions (in planes parallel to the equatorial plane).
All four chambers may exhibit the same (kidney-shaped) form and structure
(in the subgenera _Panartella_ and _Panartoma_); or the proximal chamber
may differ more or less in shape and size from the distal (in the subgenera
_Panartissa_ and _Panartura_). The outer surface of the cortical shell is
sometimes smooth (as in _Panartella_ and _Panartissa_), at other times
spiny or thorny (as in _Panartoma_ and _Panartura_). The double medullary
shell is sometimes spherical, or commonly compressed at both poles and
spheroidal or lenticular.

Subgenus 1. _Panartella_, Haeckel.

_Definition._--Surface of the cortical shell smooth, without spines or
thorns. All its four chambers exhibit nearly the same form and structure.

1. _Panartus tetraplus_, n. sp.

All four chambers of the cortical shell of the same form, size, and
structure, kidney-shaped, twice as broad as long. Pores rather regular,
hexagonal, all nearly of the same size and form, four times as broad as the
thin bars; five to six pores on the half meridian, nine to ten on the half
equator of each chamber. Surface of the cortical shell smooth, its main
axis three times as long as its equatorial axis (in the median
constriction). Both concentric medullary shells spherical; the equatorial
axis of the outer half as long as that of the cortical shell. (All four
chambers of this species have the same appearance as the two proximal
chambers of _Panartus diploconus_, Pl. 40, fig. 1.)

_Dimensions._--Main axis of the cortical shell 0.21, equatorial axis 0.07;
breadth of every chamber 0.09; meshes 0.013, bars 0.03.

_Habitat._--Central area of the Pacific, Stations 270 to 274, depth 2350 to
2925 fathoms.

2. _Panartus tetracolus_, n. sp.

All four chambers of the cortical shell of the same form, size, and
structure, kidney-shaped, twice as broad as long. Pores regular, circular,
with hexagonal frame, twice as broad as the bars; six to seven pores on the
half meridian, ten to twelve on the half equator of each chamber. Surface
of the cortical shell smooth; its main axis three times as long as the
equatorial axis. Both concentric medullary shells spheroidal, somewhat
compressed at both poles; the equatorial axis {378}of the outer almost
equals half that of the cortical shell. (All four chambers of this species
exhibit the structure of the two proximal chambers of _Peripanartus
amphiconus_, Pl. 40, fig. 5.)

_Dimensions._--Main axis of the cortical shell 0.24, equatorial axis 0.08;
breadth of every chamber 0.01; meshes 0.01, bars 0.005.

_Habitat._--Equatorial Atlantic, Station 348, depth 2450 fathoms.

3. _Panartus tetrameres_, n. sp.

All four chambers of the cortical shell of the same form, size, and
structure, kidney-shaped, twice as broad as long. Pores rather regular,
circular, without hexagonal frame, about as broad as the bars; five to six
pores on the half meridian, ten to twelve on the half equator of each
chamber. Surface of the cortical shell smooth, its main axis three times as
long as the equatorial axis. Both concentric medullary shells spheroidal,
compressed; the outer half as broad as the constriction. (This species is
very much like _Ommatocampe increscens_, Stöhr, 1880, _loc. cit._, Taf. ii.
fig. 2, and may be considered as the ancestral form of it.)

_Dimensions._--Main axis of the cortical shell 0.2, equatorial axis 0.065;
breadth of every chamber 0.08; meshes and bars 0.005.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Caltanisetta, Haeckel.

4. _Panartus tetraphalangus_, n. sp.

All four chambers of the cortical shell of the same form, size, and
structure, kidney-shaped, twice as broad as long. Pores irregular,
polygonal (mostly pentagonal or hexagonal), four to eight times as broad as
the thin bars; four to seven on the half meridian, eight to fifteen on the
half equator of each chamber. Surface of the cortical shell smooth, its
main axis three times as long as its equatorial axis. Both concentric
medullary shells compressed, the outer, half as broad as the constriction.

_Dimensions._--Main axis of the cortical shell 0.26, equatorial axis 0.09;
breadth of every chamber 0.12; meshes 0.003 to 0.02, bars 0.001 to 0.003.

_Habitat._--Fossil in the rocks of Barbados, Haeckel.

5. _Panartus tetrathalamus_, n. sp. (Pl. 40, fig. 3).

All four chambers of the cortical shell of the same form, size, and
structure, kidney-shaped, twice as broad as long. Pores irregular,
roundish, twice to three times as broad as the bars; a circle of larger
pores at the base of both distal chambers; five to six pores on the half
meridian, ten to twelve on the half equator of each chamber. Surface of the
cortical shell smooth; its main axis two and a half times as long as the
equatorial axis. Both medullary shells spheroidal, compressed, the outer
one-third as broad as the constriction. (In the specimen figured spines
begin to arise from both distal chambers; this forms a transition to
_Panartus quadrijugus_.)

_Dimensions._--Main axis of the cortical shell 0.18, equatorial axis 0.07;
breadth of every chamber 0.09; meshes 0.003 to 0.01, bars 0.002 to 0.004.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, in various
depths.

{379}Subgenus 2. _Panartissa_.

_Definition._--Surface of the cortical shell smooth, without spines or
thorns. Both its proximal chambers differ in form and structure from the
two distal chambers.

6. _Panartus diploconus_, n. sp. (Pl. 40, fig. 1).

Both proximal chambers nearly ellipsoidal, one and a half times as broad as
long, with regular hexagonal pores, four times as broad as the bars; five
to six pores on the half meridian, nine to ten on the half equator of each
chamber. Both distal chambers conical, with a circle of ten to twelve large
square pores at the base; the other pores very small and numerous,
roundish, sixteen to twenty in the height of each cone, and the same number
on the half periphery of its base. Both concentric medullary shells
compressed, the outer, half as broad as the constriction of the cortical
shell, the surface of which is quite smooth.

_Dimensions._--Total length of the shell (or main axis) 0.34; greatest
breadth of each chamber 0.09; pores of the proximal chambers 0.013, bars
0.003; pores and bars of the distal chambers 0.003; large basal square
pores 0.02.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

7. _Panartus amphiconus_, n. sp.

Both proximal chambers nearly kidney-shaped, one and a half times as broad
as long, with regular, circular pores, with hexagonal frames, twice as
broad as the bars; six to seven pores on the half meridian, ten to twelve
on the half equator of each chamber. Both distal chambers conical, with a
circle of ten to twelve very large pores at the base; the other pores very
small, roundish, ten to fifteen in the height of each cone, twenty to
thirty on the half basal periphery. Both concentric medullary shells
compressed, the outer one-third as broad as the constriction of the
cortical shell, whose surface is smooth. (This species resembles
_Peripanartus amphiconus_, Pl. 40, fig. 5, but is without the surface
spines and the outer envelope.)

_Dimensions._--Total length of the shell 0.28; greatest breadth of the
proximal chambers 0.1, of the distal chambers 0.14; pores of the former
0.008, of the latter 0.004; bars of both 0.003 to 0.005.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

8. _Panartus fusiformis_, n. sp.

Both proximal chambers nearly ellipsoidal, one and a half times as long as
broad, with irregular, roundish pores, twice to three times as broad as the
bars; eight to ten pores on the half meridian, twelve to fifteen on the
half equator of each chamber. Both distal chambers hemiellipsoidal or
egg-shaped, somewhat higher and narrower than the proximal chambers, with a
circle of larger square pores at their base; the other pores roundish and
very irregular. Both medullary shells spheroidal, one-third as broad as the
constriction of the cortical shell, whose surface is quite smooth. (This
{380}species is similar to _Peripanartus atractus_, Pl. 40, fig. 7, but is
without the superficial spines and the equatorial girdle-like envelope of
the latter.)

_Dimensions._--Total length of the shell 0.25 to 0.3; greatest breadth (in
the equator of the proximal chambers) 0.08 to 0.1; pores of the proximal
chambers 0.008 to 0.012, pores of the distal chambers half their size, bars
0.002 to 0.006.

_Habitat._--North Pacific, Station 237, surface.

9. _Panartus profundissimus_, Haeckel.

_Ommatocampe profundissima_, Ehrenberg, 1872, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 297, Taf. viii. fig. 6.

Both proximal chambers nearly ellipsoidal, with irregular, polygonal pores,
six to eight times as broad as the bars; four to five pores on the half
meridian, eight to nine on the half equator of each chamber. Both distal
chambers hemispherical, with a very delicate irregular network. Medullary
shells spherical; surface of the shell smooth.

_Dimensions._--Total length of the shell 0.15; greatest breadth in the
equator of the proximal chambers 0.08; pores of the proximal chambers
0.015, bars 0.002.

_Habitat._--Philippine Sea, depth 3300 fathoms, Ehrenberg; Station 213,
depth 2050 fathoms.

Subgenus 3. _Panartoma_, Haeckel.

_Definition._--Surface of the cortical shell thorny, covered with scattered
spines. All its four chambers exhibit the same form and structure.

10. _Panartus quadriceps_, n. sp.

All four chambers of the cortical shell of the same form, size, and
structure, kidney-shaped, twice as broad as long. Pores rather regular,
circular, with hexagonal frames, twice to three times as broad as the bars;
six to seven pores on the half meridian, ten to twelve on the half equator
of each chamber. Surface spiny; from the corners of the hexagonal frames
(between every three pores) arise short radial spines. Both medullary
shells spherical, the outer one-third as broad as the constriction of the
cortical shell. (All the four chambers of this species have the same form
as the two proximal chambers of _Peripanartus amphiconus_, Pl. 40, fig. 5.)

_Dimensions._--Main axis of the cortical shell 0.24, equatorial axis 0.08;
meshes 0.01, bars 0.003 to 0.005; length of the spines 0.01 to 0.02.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

11. _Panartus quadrijugus_, n. sp.

All four chambers of the cortical shell of the same form and size,
kidney-shaped, twice as broad as long. Pores irregular, roundish, twice to
three times as broad as the bars; a circle of larger pores at the base of
both distal chambers; six to eight pores on the half meridian, twelve to
{381}fourteen on the half equator of each chamber. Surface spiny,
everywhere covered with small irregular thorns. (This species differs from
the smooth _Panartus tetrathalamus_, Pl. 40, fig. 3, almost entirely by the
thorny surface.)

_Dimensions._--Main axis 0.2, equatorial axis 0.08; breadth of every
chamber 0.1; meshes 0.005 to 0.01, bars 0.003, spines 0.01 to 0.02.

_Habitat._--Cosmopolitan; on the surface of the Atlantic, Indian, and
Pacific Oceans.

12. _Panartus quadrigeminus_, n. sp.

All four chambers of the cortical shell nearly of the same size and form,
kidney-shaped, twice as broad as long. Pores irregular, polygonal (mostly
pentagonal or hexagonal), six to eight times as broad as the thin bars;
five to seven on the half meridian, twelve to sixteen on the half equator
of each chamber. Surface of the cortical shell spiny, covered with numerous
irregularly scattered, often oblique, bristle-like thorns.

_Dimensions._--Main axis 0.27, equatorial axis 0.1; breadth of each chamber
0.13; meshes 0.01 to 0.02, bars 0.001 to 0.005.

_Habitat._--Cosmopolitan; surface of the Atlantic and Pacific, many
Stations.

Subgenus 4. _Panartura_, Haeckel.

_Definition._--Surface of the cortical shell thorny, covered with scattered
spines. Both its proximal chambers differ in form or structure from the two
distal chambers.

13. _Panartus spinosus_, n. sp.

Both proximal chambers nearly ellipsoidal, with regular, circular pores
enclosed by hexagonal frames. Both distal chambers conical, with a circle
of ten to twelve large square pores at the base; the other pores small,
roundish. The cortical shell of this species is quite the same as that of
_Panartus amphiconus_ (Pl. 40, fig. 5), but differs in the absence of the
external envelope (or the second cortical shell), and is covered with
short, conical spines on the whole surface.

_Dimensions._--Total length of the cortical shell 0.25, greatest breadth
(on the base of the distal chambers) 0.12; meshes of the proximal chambers
0.008, of the distal chambers 0.004; bars 0.002 to 0.04, spines 0.01 to
0.02.

_Habitat._--North Pacific, Station 240, surface.

14. _Panartus setosus_, Haeckel.

_Ommatocampe setosa_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 297, Taf. viii. fig. 7.

Both proximal chambers kidney-shaped, broader than the two hemispherical
distal chambers. Meshes of the cortical shell irregular, polygonal, or
subregular hexagonal; at the base of each {382}distal chamber a circle of
large square meshes. The breadth of the pores is eight to twelve times that
of the bars between them. The whole surface is covered with small
bristle-like spines.

_Dimensions._--Total length of the cortical shell 0.14, greatest breadth
(in the equator of each proximal chamber) 0.07; meshes 0.008 to 0.015, bars
0.002, spines 0.01.

_Habitat._--Pacific (Philippine and Californian Sea) 3300 and 2300 fathoms,
Ehrenberg; Station 200, depth 250 fathoms; Station 253, depth 3125 fathoms.

15. _Panartus pluteus_, n. sp. (Pl. 40, fig. 2).

Both proximal chambers kidney-shaped, with irregular, roundish pores, twice
to four times as broad as the bars; their surface everywhere covered with
short conical thorns. From both polar circles arise ten to twelve radial
rods, which bear a fenestrated cap, and form thus either the beginning of a
second, external, cortical shell, or (if remaining thus) two imperfect
distal chambers. The spherical segment, which forms their surface, is
concentric with the proximal chambers, is perforated by the same irregular,
roundish pores, and covered with numerous bristle-like spines.

_Dimensions._--Main axis of the cortical shell 0.17, equatorial axis (in
the constriction) 0.06; distance between the proximal and distal chambers
0.03; pores 0.003 to 0.01, bars 0.003 to 0.005.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Genus 164. _Peripanartus_,[203] n. gen.

_Definition._--#Panartida# with double cortical shell and double medullary
shell, without polar spines or tubes on both poles of the main axis.

The genus _Peripanartus_ differs from _Panartus_ only in the development of
an outer reticulated envelop, which mantle-like surrounds the shell and
represents a second or external cortical shell. This mantle or veil
envelops either the whole internal cortical shell or only both proximal
chambers. _Peripanartus_ exhibits the same relation to _Panartus_ as in the
foregoing ancestral family _Cypassis_ bears to _Cyphonium_.

Subgenus 1. _Peripanartula_, Haeckel.

_Definition._--Surface of the outer cortical shell smooth, without spines
or thorns.

1. _Peripanartus palliatus_, n. sp.

Internal cortical shell with four kidney-shaped chambers of equal size and
similar form, identical with the cortical shell of _Panartus tetrathalamus_
and _Panartus quadrijugus_ (Pl. 40, fig. 3), with irregular, roundish
pores, twice to three times as broad as the bars. It differs from these two
species {383}only by the external cortical shell enveloping the whole
internal one, in the same manner as in _Peripanartus cylindrus_ (Pl. 40,
fig. 6). The two shells are connected by numerous radial rods. The surface
of the outer cortical shell is quite smooth, its irregular, roundish pores
about half as large as those of the inner cortical shell.

_Dimensions._--Main axis of the external cortical shell 0.27, of the
internal 0.2; greatest breadth of the former (in the equator of each
chamber) 0.13, of the latter 0.09; pores of the external cortical shell
0.002 to 0.004, of the internal 0.004 to 0.008; bars of the former 0.002,
of the latter 0.003.

_Habitat._--Pacific, central area, Stations 271, 272, depth 2425 to 2600
fathoms.

2. _Peripanartus lævigatus_, n. sp.

Internal cortical shell with four unequal chambers, both proximal
kidney-shaped (with subregular polygonal pores, twice to three times as
broad as the bars), both distal nearly hemispherical, with subregular,
roundish pores of half that size. External cortical shell envelops only the
proximal chambers, and appears as the direct continuation of the internal
shell of the distal chambers, with the same small roundish pores. The outer
surface is quite smooth. This species has nearly the same structure as
_Peripanicium amphicorona_ (Pl. 40, fig. 8), but differs from it by the
absence of the polar spine-circles, and by the perfect smoothness of the
surface.

_Dimensions._--Main axis of the shell 0.25, greatest breadth (in the
equator of the proximal chambers) 0.15; equatorial stricture of the
external shell 0.10, of the internal 0.07; pores of the internal shell of
the proximal chambers 0.013, of the external 0.006; bars 0.003.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

3. _Peripanartus amphiconus_, n. sp. (Pl. 40, fig. 5).

Inner cortical shell with four very unequal chambers; both proximal
chambers kidney-shaped (with regular, circular, hexagonally framed pores,
twice as broad as the bars); both distal chambers conical, with a circle of
large square pores at the base, the other pores very small, roundish.
Outer cortical shell very thin, quite smooth, with irregular, roundish
pores, envelops in younger specimens only the distal chambers. But the
radial spines on the surface of the internal cortical shell of the proximal
chambers indicate that these will also be afterwards enclosed by the
growing of the external shell. This species exhibits the progressive
(ontogenetic as well as phylogenetic) development of _Panartus amphiconus_.

_Dimensions._--Main axis of the outer cortical shell 0.3, its greatest
breadth (at the prominent distal ends of the radial rods between the large
square pores) 0.15; pores 0.003 to 0.001, bars 0.005; main axis of the
inner cortical shell 0.26, its greatest breadth 0.13; pores of its distal
chambers 0.003, of its proximal chambers 0.01; large square pores 0.03;
bars 0.004.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

{384}Subgenus 2. _Peripanartium_, Haeckel.

_Definition._--Surface of the outer cortical shell covered with spines or
thorns.

4. _Peripanartus atractus_, n. sp. (Pl. 40, fig. 7).

Inner cortical shell with four very unequal chambers; both proximal
chambers spheroidal (with subregular, roundish pores, twice to three times
as broad as the bars), both distal chambers nearly conical, with a circle
of eight to ten very large square pores at the base; the other pores very
small, roundish. Outer cortical shell envelops only the proximal chambers
and the basal half of the distal chambers, appearing as the direct
continuation of the microporous covering of their apical half. The whole
external cortical shell is inflated in the equatorial zone, and hence
assumes an irregular, spindle-like appearance. Its whole surface is rough
with innumerable very short thorns.

_Dimensions._--Main axis 0.28, equatorial axis 0.17; pores of the external
cortical shell 0.002 to 0.004, bars 0.002; main axis of the proximal
chambers of the internal cortical shell 0.07, equatorial axis 0.09; pores
0.007, bars 0.004.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

5. _Peripanartus cylindrus_, n. sp. (Pl. 40, fig. 6).

Internal cortical shell with four unequal chambers; both proximal chambers
kidney-shaped (with regular, circular, hexagonally framed pores of the same
breadth as the bars), both distal chambers nearly hemispherical, with a
circle of eight to ten very large square pores at their base; the other
pores very small, roundish. External cortical shell envelops the whole
internal like a perfect hollow cylinder, which is closed at both poles by a
hemispherical cap. The whole surface of this cylindrical mantle is
perforated by innumerable very small roundish pores, and covered with small
bristle-like spines.

_Dimensions._--Main axis of the cylinder 0.3, equatorial axis 0.12; pores
and bars 0.006; spines of its surface 0.012 to 0.15; main axis of the
internal cortical shell 0.2; equatorial axis (in the constriction) 0.06;
pores and bars of its proximal chambers 0.007, of its distal chambers
0.003; large square pores at their base 0.02.

_Habitat._--Pacific, central area, Stations 270 to 274, depth 2350 to 2925
fathoms.

Genus 165. _Panicium_,[204] n. gen.

_Definition._--#Panartida# with simple cortical shell and double medullary
shell, with two opposite apical spines on both poles of the main axis, or
with a group of polar spines.

The genus _Panicium_ comprises those forms of _Panartus_ which develop a
peculiar armature on both poles of the cortical shell. This may be only a
single strong spine {385}on each pole, lying in the main axis (subgenus
_Panicidium_); or a bunch or a circle of polar spines diverging radially
(subgenus _Panartidium_); in the latter case the polar spines can either
form a bunch on the pole itself, or a crown of thorns around it.

Subgenus 1. _Panicidium_, Haeckel.

_Definition._--On both poles of the main axis only a single large spine.

1. _Panicium amphacanthum_, n. sp.

All four chambers of the cortical shell nearly of the same size and form,
kidney-shaped, with subregular, circular pores, twice as broad as the bars;
five to six pores on the half meridian, ten to eleven on the half equator
of each chamber. Surface covered with small spines. On both poles of the
main axis is a very strong, conical, apical spine, half as long as the main
axis, on the base one-fourth as broad as the equatorial axis.

_Dimensions._--Main axis (without polar spines) 0.22, equatorial axis 0.06;
pores 0.01, bars 0.005; length of the polar spines 0.12, basal thickness
0.015.

_Habitat._--Antarctic, Station 152, depth 1260 fathoms.

2. _Panicium amphistylus_, n. sp.

Both proximal chambers kidney-shaped, both distal chambers nearly
hemispherical, somewhat smaller. Pores irregular, roundish, twice to three
times as broad as the bars; four to six on the half meridian, eight to ten
on the half equator of each chamber; a circle of larger square pores on the
base of each distal chamber. Surface thorny. On both poles of the main axis
one strong, pyramidal, apical spine, one-third as long as the main axis, on
the base one-fourth as broad as the equatorial constriction. (Similar to
_Panarium tubularium_, Pl. 40, fig. 9, but without tubes.)

_Dimensions._--Main axis (without polar spines) 0.2, equatorial axis 0.05;
pores 0.005 to 0.01, bars 0.003; length of the polar spines 0.07, basal
thickness 0.012.

_Habitat._--Antarctic, Station 156, depth 1975 fathoms.

Subgenus 2. _Panartidium_, Haeckel.

_Definition._--On both poles of the main axis a bunch or a circle of
aggregated spines.

3. _Panicium scoparium_, n. sp.

Both proximal chambers kidney-shaped, both distal chambers nearly
hemispherical, somewhat smaller. Pores irregular, polygonal, twice to five
times as broad as the bars; six to eight on the half meridian, twelve to
fourteen on the half equator of each chamber. Surface spiny, covered with
thin, {386}bristle-like spines. On both poles of the main axis a bunch of
six to eight aggregated larger conical spines, about as high as a single
chamber.

_Dimensions._--Main axis (without polar spines) 0.2, equatorial axis 0.06;
pores 0.005 to 0.015, bars 0.003; length of the polar spines 0.05, basal
thickness 0.005.

_Habitat._--Pacific, central area, Stations 270 to 274, depth 2350 to 2925
fathoms.

4. _Panicium coronatum_, n. sp. (Pl. 40, fig. 4).

Both proximal chambers kidney-shaped, covered with short stout spines.
Pores irregular, roundish, twice to four times as broad as the bars; seven
to eight on the half meridian, eleven to thirteen on the half equator of
each chamber. Both distal chambers cap-like, separated from the former by a
circle of ten to twelve very large square pores; the other pores very
small, roundish. The circumpolar area is smooth, surrounded by a circle of
ten to twelve very stout, conical, radial spines, which arise from the
distal ends of the bars separating the large square meshes.

_Dimensions._--Main axis (without spines) 0.22, equatorial axis 0.08;
greatest breadth in the equator of the chambers 0.1; pores of the proximal
chambers 0.007 to 0.015, bars 0.004; square pores of the distal chambers
0.02 to 0.03, small pores 0.005, bars 0.004; length of the radial spines of
the polar circles 0.07, basal thickness 0.007.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 166. _Peripanicium_,[205] n. gen.

_Definition._--#Panartida# with double cortical shell and double medullary
shell, on both poles of the main axis with two opposite apical spines, or
with a bunch or circle of polar spines.

The genus _Peripanicium_ differs from _Panicium_ only in the development of
an outer reticulated envelope, which mantle-like surrounds the shell, and
represents a second or external cortical shell. As in _Panicium_, its
ancestral form, so also in _Peripanicium_, there can be distinguished two
subgenera:--_Peripanicea_, where only a single large spine arises from each
pole (lying in the main axis), and _Peripanicula_, where a variable number
of spines is to be found, either diverging bunch-like from the pole itself,
or surrounding it as a crown-like polar circle.

Subgenus 1. _Peripanicea_, Haeckel.

_Definition._--On both poles of the main axis only a single large spine.

1. _Peripanicium amphixiphus_, n. sp.

Internal cortical shell with four unequal chambers, with thorny surface.
Both proximal chambers kidney-shaped (with subregular hexagonal pores,
three times as broad as the bars). {387}Both distal chambers nearly
hemispherical (with a circle of ten to twelve very large square pores at
their base, the other pores very small). From both poles arises a single,
strong, conical spine, half as long as the main axis. The internal cortical
shell is very much like that of _Panarium tubularium_ (Pl. 40, fig. 9), but
differs by the solid polar spines (instead of the hollow tubules) and by
the formation of an external, cylindrical, very thin, cortical shell, which
envelops the whole internal, and is connected with it by numerous beams.
The surface of the external shell is quite smooth, and perforated by
innumerable very small roundish pores.

_Dimensions._--Main axis of the external cortical shell 0.25, of the
internal 0.19; transverse axis of the former 0.13, of the latter 0.08;
pores of the proximal chambers of the internal shell 0.01, bars 0.003;
pores of external cortical shell 0.002, bars 0.001; length of both polar
spines 0.12, basal thickness of them 0.02.

_Habitat._--South Atlantic, near Tristan da Cunha, Station 333, depth 2025
fathoms.

Subgenus 2. _Peripanicula_, Haeckel.

_Definition._--On both poles of the main axis a bunch or a circle of
aggregated polar spines.

2. _Peripanicium amphicorona_, n. sp. (Pl. 40, fig. 8).

_Peripanartus amphicorona_, Haeckel, 1881, Prodromus et Atlas (pl. xl.
fig. 8).

Internal cortical shell with four very unequal chambers. Both proximal
chambers kidney-shaped, with subregular, polygonal pores, three times as
broad as the bars. Both distal chambers double cone-shaped; the proximal
cone of these formed by a circle of ten to twelve radial beams, which
separate the same number of large square meshes; the distal cone with five
to six circles of very small roundish, irregular pores. The continuation of
the thin porous lamella of this distal cone forms the external cortical
shell, whose surface is a little rough with innumerable very small thorns.
On both poles is a large circular opening (nearly as broad as the
equatorial constriction of the inner shell), surrounded by a delicate crown
of thorns. The spines of this crown equal the diameter of the polar
opening; they are very thin and bristle-like, connected by a few bridges
and diverge outwards.

_Dimensions._--Main axis (without the crowns of polar thorns) 0.26,
greatest breadth (in the equator of the proximal chambers) 0.15; equatorial
constriction of the external shell 0.12, of the internal 0.07; pores of the
proximal chambers of the internal cortical shell 0.01, of the external
0.005, bars 0.003; diameter of the circular polar opening 0.05; length of
the polar spines 0.05.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

3. _Peripanicium coronarium_, n. sp.

Internal cortical shell with four unequal chambers. Both proximal chambers
kidney-shaped, with irregular, roundish pores, twice to four times as broad
as the bars. Both distal chambers cap-like, with a basal circle of ten to
twelve large square meshes, the other pores very small. External
{388}cortical shell very thin, with smooth surface, and very small,
irregular, roundish pores; it envelops the whole internal shell at a
constant distance, which equals the breadth of the square meshes. From the
distal ends of the radial beams between the square meshes arise on every
polar circle ten to twelve strong spines, as direct prolongations of those
beams. These form two regular, polar crowns of thorns. The inner part of
the thorns (between both shells) has only one-third to one-fourth the
length of the outer free part. (This species represents a further
development of _Panicium coronatum_, Pl. 40, fig. 4, by secondary formation
of an external mantle, like that of _Peripanartus atractus_, Pl. 40, fig.
7.)

_Dimensions._--Main axis 0.27, greatest breadth 0.15; pores of the internal
proximal chambers 0.008 to 0.016, bars 0.004; square meshes of the distal
chambers 0.03; pores of the outer cortical shell 0.003; length of the
spines of the polar circles 0.1 to 0.15.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

Genus 167. _Panarium_,[206] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Panartida# with simple cortical shell and double medullary
shell, with two hollow fenestrated tubes, opposite on the poles of the main
axis.

The genus _Panarium_ differs from _Panartus_ by two hollow latticed tubes,
which start from both poles of the main axis and lie in it. It repeats
therefore in this family the same peculiar and remarkable formation, which
we find in _Pipettella_ among the Ellipsida, in _Pipetta_ among the
Druppulida, in _Cannartidium_ among the Cyphinida, in _Cannartus_ among the
Artiscida, &c.

Subgenus 1. _Panarelium_, Haeckel.

_Definition._--Surface of the shell smooth, without spines or thorns.

1. _Panarium facettarium_, n. sp.

Surface of the cortical shell smooth. All the four chambers nearly of the
same size and form, kidney-shaped, about twice as broad as long. Pores of
these subregular, circular, with hexagonal frames, about as broad as the
bars; nine to ten on the half meridian, twelve to fourteen on the half
equator of each chamber. Polar tubuli nearly cylindrical, longer than half
the main axis of the cortical shell, about one-fourth as broad as the
equatorial constriction. Pores of the tubuli of the same shape as those of
the chambers, but only half as large. (This species is like _Pipetta tuba_,
Pl. 39, fig. 7, but distinguished by three parallel transverse
constrictions.)

_Dimensions._--Total length of the shell (without tubuli) 0.26; breadth of
each chamber 0.12; pores 0.01, bars 0.01; length of the tubuli 0.15,
breadth of them 0.03.

_Habitat._--Pacific central area, Station 268, depth 2900 fathoms.

{389}2. _Panarium pipettarium_, n. sp.

Surface of the cortical shell smooth. Both proximal chambers nearly
kidney-shaped; both distal chambers somewhat smaller, hemispherical. Pores
of all four chambers subregular, circular, without hexagonal frames, twice
as broad as the bars; six to seven pores on the half meridian, ten to
eleven on the half equator of each chamber. Polar tubuli slender, conical,
nearly half as long as the main axis of the shell, one-third as broad at
the base as the equatorial constriction. Pores of the tubuli of the same
shape as those of the chambers, but only half as large. (This species
resembles somewhat _Cannartiscus amphiconiscus_, Pl. 39, fig. 19, but with
double the number of chambers.)

_Dimensions._--Total length of the shell (without tubuli) 0.25, greatest
breadth 0.1; pores 0.008, bars 0.004; length of the polar tubuli 0.11,
basal breadth 0.03.

_Habitat._--Pacific, western tropical part, Station 225, depth 4475
fathoms.

Subgenus 2. _Panaromium_, Haeckel.

_Definition._--Surface of the shell spiny, covered with short thorns.

3. _Panarium annularium_, n. sp.

Surface of the cortical shell thorny, everywhere covered with small
bristle-like spines. All four chambers nearly of the same size and form,
kidney-shaped. Their pores subregular, circular, three to four times as
broad as the bars; seven to eight on the half meridian, ten to eleven on
the half equator of each chamber. Polar tubuli conical, nearly half as long
as the main axis, half as broad at the base as the equatorial constriction.
Pores of the tubuli like those of the chambers, but only half as broad.

_Dimensions._--Total length of the shell (without tubuli) 0.16; breadth of
each chamber 0.07; pores 0.006 to 0.008, bars 0.002; length of the polar
tubuli 0.07, basal breadth 0.03.

_Habitat._--Northern Pacific, Station 253, depth 3125 fathoms.

4. _Panarium artophorum_, n. sp.

Surface of the shell thorny, with scattered small spines. Both proximal
chambers nearly kidney-shaped; both distal hemispherical, somewhat smaller.
Pores of all chambers irregular, roundish, little broader than the bars;
four to five on the half meridian, six to eight on the half equator of each
chamber. Polar tubuli cylindrical, longer than the half main axis, only
one-fourth as broad the equatorial constriction. Pores of the tubuli only
one-third to one-fourth as broad as those of the chambers.

_Dimensions._--Total length of the shell (without tubuli) 0.22, greatest
breadth (in the equator of the proximal chambers) 0.09; pores and bars
0.008 to 0.012; length of the polar tubuli 0.14, breadth of them 0.02.

_Habitat._--Southern Pacific, Station 289, depth 2550 fathoms.

{390}5. _Panarium tubularium_, n. sp. (Pl. 40, fig. 9).

Surface of the cortical shell thorny, covered with small spines. Both
proximal chambers nearly kidney-shaped, with irregular, polygonal pores,
three to four times as broad as the bars; five to six pores on the half
meridian, nine to ten on the half equator of each chamber. Both distal
chambers nearly hemispherical, with a circle of ten to twelve large square
pores at their base; the other pores much smaller, irregular, roundish.
Polar tubuli prismatic, with prominent edges, nearly half as long as the
main axis, only one-fourth as broad as the equatorial constriction. Pores
of the tubuli very small, in longitudinal series between the edges.

_Dimensions._--Total length of the shell (without tubuli) 0.18 to 0.2,
greatest breadth (in the equator of the proximal chambers) 0.07 to 0.09;
pores 0.003 to 0.01, bars 0.002 to 0.004; length of the polar tubuli 0.06
to 0.09, breadth of them 0.015 to 0.02.

_Habitat._--Pacific, central area, Stations 270 to 272, depth 2425 to 2925
fathoms.

Genus 168. _Peripanarium_,[207] n. gen.

_Definition._--#Panartida# with double cortical shell and double medullary
shell, with two hollow fenestrated tubes, opposite on the poles of the main
axis.

The genus _Peripanarium_ differs from _Panarium_ only in the development of
an outer reticulated envelop, which mantle-like surrounds the shell and
represents a second or external cortical shell. It bears therefore to
_Panarium_ the same relation as _Peripanartus_ to _Panartus_, &c.

1. _Peripanarium cenoconicum_, n. sp.

Internal cortical shell with four kidney-shaped chambers of nearly the same
form and size; their pores irregular, roundish, twice to four times as
broad as the bars. External cortical shell cylindrical, hemispherical at
both poles, with irregular, very small, roundish pores, only one-third to
one-half as large as those of the internal. From both poles of the
internal cortical shell arises a hollow conical fenestrated tube, which
perforates the external. The outer free part of this slender cone equals
in length the inner part or the distance between the two cortical shells,
which is about equal to the length of one chamber. External surface covered
with small thorns. (This species has the same appearance as _Desmartus
larvalis_, Pl. 40, fig. 12, but has only four chambers instead of six.)

_Dimensions._--Main axis (without cones) 0.23, greatest breadth 0.11; pores
of the inner cortical shell 0.004 to 0.009, of the outer 0.002 to 0.003;
bars of the former 0.002, of the latter 0.001; total length of the polar
cones 0.05, basal thickness of them 0.03.

_Habitat._--West Tropical Pacific, Station 225, depth 4475 fathoms.

{391}2. _Peripanarium cenocylindricum_, n. sp.

Internal cortical shell with four unequal chambers, the two proximal
kidney-shaped, the two distal conical, and somewhat smaller; pores
irregular, polygonal-roundish, twice to five times as broad as the bars.
External cortical shell nearly spindle-shaped, in the equatorial zone
inflated, conical, tapering towards both poles, its pores very small,
roundish, its surface covered with small thorns. The shell of this species
resembles very much _Peripanartus atractus_ (Pl. 40, fig. 7), but differs
by the hollow cylindrical polar tubes, which arise from both poles of the
inner cortical shell and attain more than half the length of the main axis.
They are open at their ends (broken away), about as broad as the outer
medullary shell, and perforated by irregular, roundish pores.

_Dimensions._--Main axis (without polar tubes) 0.28, greatest breadth (in
the equator) 0.16; pores of the inner cortical shell 0.004 to 0.01, of the
outer 0.002 to 0.005; bars of the former 0.004, of the latter 0.002; total
length of the polar tubes 0.16, breadth of them 0.03.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

Family XVII. #ZYGARTIDA#, Haeckel (Pl. 40, figs. 10-13).

_Zygartida_, Haeckel, 1881, Prodromus, p.462 (_sensu restricto_).

_Definition._--#Prunoidea# with annulated cortical shell, the external
shell being divided by five or more parallel transverse constrictions into
six or more chambers, enclosing in the centre two internal concentric
shells (medullary shells). Central capsule cylindrical, commonly annulated
(corresponding to the transverse annular constrictions of the cortical
shell).

The family #Zygartida#, the seventh and last of the #Prunoidea#, comprises,
in the sense here restricted, all those #Prunoidea# in which the cortical
shell is annulated and composed of six to twenty or more (at least six)
cameræ, lying one behind another in the elongated main axis, and separated
by five or more annular constrictions. When I constituted this family in
1881 (_loc. cit._), I had given to it a much wider extent, embracing all
those #Prunoidea# which exhibit annular constrictions of the cortical
shell; as the number of these, in consequence of further researches, is
much increased, it seems now more convenient to restrict the family to the
extent here given.

No doubt the Zygartida must be derived from the Panartida by progressive
growth of the cortical shell in the main axis and accompanying increase of
the number of its chambers. Whilst this number in the Panartida is
constantly restricted to four, in the Zygartida it amounts to ten, twenty,
or more, and is in the lowest case six (Pl. 40, fig. 10). The maximum
number is variable in the different species, but of course also different
in the various degrees of individual development. Each Zygartid is at the
beginning of its growth a Cyphinid, later a Panartid. The number of the
annular constrictions separating the chambers and lying in parallel
transverse planes is at least {392}five, but may amount to nineteen, to
twenty-one or more; they lie constantly in pairs on both sides of the
equatorial constriction (Pl. 40, figs. 12, 13).

The cortical shell may either remain simple (_Ommatocampe_, Pl. 40, fig.
10), or become double (_Desmocampe_, Pl. 40, fig. 12), or sometimes triple
(_Zygocampe_, Pl. 40, fig. 13). In the latter cases the outer (secondary
and tertiary) cortical shells are commonly incomplete, and only developed
around the proximal chambers of the complete first (primary) cortical
shell, its distal chambers remaining simple.

_The Medullary Shell_ is constantly double, as in the Panartida; its form
is either spherical or lenticular, compressed in the direction of the main
axis. It is always connected with the equatorial constriction of the
cortical shell by a number of radial beams, lying either in the equatorial
plane or on each side of it (Pl. 40, figs. 10-13).

_The Central Capsule_ of the Zygartida is constantly cylindrical; its
increasing growth on both poles of the axis corresponds to that of the
including cortical shell. Commonly (but not always) its cylindrical surface
is annulated, with five or more transverse strictures, corresponding to
those of the cortical shell. From the inner surface of the latter it is
separated by a jelly-mantle, the calymma.

_Synopsis of the Genera of Zygartida._

I. Ommacampida. { Without polar tubes, 169. _Ommatocampe_.
1. Cortical shell {
simple. { With two hollow fenestrated
(Medullary shell { tubes, on the poles of
double.) { the axis, 170. _Ommatartus_.

II. Desmocampida. { Without polar tubes, 171. _Desmocampe_.
2. Cortical shell {
double. { With two hollow fenestrated
(Medullary shell { tubes, on the poles of
double.) { the axis, 172. _Desmartus_.

III. Zygocampida. { Without polar tubes, 173. _Zygocampe_.
3. Cortical shell {
triple. { With two hollow fenestrated
(Medullary shell { tubes, on the poles of
double.) { the axis, 174. _Zygartus_.

Genus 169. _Ommatocampe_,[208] Ehrenberg, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 832.

_Definition._--#Zygartida# with simple cortical shell and double medullary
shell, without polar tubes.

The genus _Ommatocampe_ was founded by Ehrenberg in 1860 for one of his
"_Haliommatina_," with the following diagnosis:--"Shell rod-like, long,
articulate, with nucleus, without spines, with four or more joints." The
species figured by him, _Ommatocampe {393}polyarthra_, exhibits five pairs
of chambers on both sides of the meridian plane of the cortical shell, the
centre of which includes a double medullary shell. It represents the most
simple form of all Zygartida, and may be derived phylogenetically from
_Cyphonium_ simply by multiplication of the chambers of the cortical shell,
growing on both poles of the main axis.

Subgenus 1. _Ommatocampium_.

_Definition._--Surface of the cortical shell smooth or rough, without
thorns or spines, also without polar spines.

1. _Ommatocampe polyarthra_, Ehrenberg.

_Ommatocampe polyarthra_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 279, Taf. vi. fig. 9.

Cortical shell with smooth surface, composed of six to ten chambers of the
same size and form. Every chamber kidney-shaped, about twice as broad as
long, with three transverse rows of circular, subregular pores, twice to
three times as broad as the bars. Both medullary shells spherical.

_Dimensions._--Length of the six-chambered cortical shell 0.15; greatest
breadth of each chamber 0.04; pores 0.005 to 0.007, bars 0.002 to 0.003.

_Habitat._--Californian Sea, depth 2600 fathoms, Ehrenberg; Pacific,
central area, Station 268, depth 2900 fathoms; fossil in the Tertiary rocks
of Barbados, Haeckel.

2. _Ommatocampe increscens_, Stöhr.

_Ommatocampe increscens_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 90,
Taf. ii. fig. 2, _a_, _b_.

Cortical shell with rough surface, composed of six to ten chambers of
nearly the same size and form; the breadth of the chambers a little
increasing towards both poles. Every chamber kidney-shaped, the proximal
chambers twice as broad as long, with four or five transverse rows of
pores; the distal chambers three to four times as broad as long, with two
or three transverse rows of pores. Form of the pores irregular, roundish,
bars between them in the distal part smaller, in the proximal part larger
than the pores. Both medullary shells compressed lenticular.

_Dimensions._--Length of the six-chambered cortical shell 0.22; greatest
breadth of the chambers 0.09 to 0.1; pores and bars 0.003 to 0.006.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Stöhr.

3. _Ommatocampe annulata_, n. sp.

Cortical shell with smooth surface, composed of six to twelve chambers of
nearly the same size and form. Each chamber kidney-shaped, twice as broad
as long, with four to five transverse rows of {394}irregular, roundish
pores, once to five times as broad as the bars. On the base of each distal
chamber (beginning in the second or third pair) a circle of twelve to
sixteen larger square pores. Both medullary shells compressed lenticular.
(Resembles the inner cortical shell of _Desmartus larvalis_, Pl. 40, fig.
12, but is without the polar tubes and the outer envelope of this species.)

_Dimensions._--Length of the six-chambered cortical shell 0.23; greatest
breadth of each chamber 0.07; pores 0.002 to 0.01, bars 0.002.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, in various
depths.

Subgenus 2. _Ommatocampula_, Haeckel.

_Definition._--Surface of the cortical shell thorny or spiny, but without
peculiar polar spines and without regular coronals of spines.

4. _Ommatocampe erucæformis_, n. sp.

Cortical shell with spiny surface, composed of six to twelve chambers of
nearly the same size and form, the distal chambers a little smaller. The
form and structure of the cortical shell is nearly the same as in the
foregoing species, only the pores are somewhat larger, and the whole
surface is covered with bristle-like radial spines, about half as long as
the breadth of one chamber. Both medullary shells compressed and
lenticular.

_Dimensions._--Length of the six-chambered cortical shell 0.24; greatest
breadth of each chamber 0.08; pores 0.005 to 0.012, bars 0.002.

_Habitat._--North Atlantic, Canary Islands, Station 353, depth 2965
fathoms.

5. _Ommatocampe nereis_, n. sp. (Pl. 40, fig. 10).

Cortical shell with spiny surface, composed of six (or more) chambers of
different size and structure. Both proximal chambers kidney-shaped, with
subregular, circular, hexagonally framed pores, twice as broad as the bars.
All following chambers cap-like, with much smaller, irregular, roundish
pores, at the base of every chamber a circle of ten to twelve large square
pores. Spines of the surface short, irregularly scattered. Both medullary
shells spherical. (All the observed specimens possessed only six chambers.)

_Dimensions._--Length of the six-chambered cortical shell 0.25; greatest
breadth of each chamber 0.08; pores of both proximal chambers 0.01, bars
0.005; basal pores of the other chambers 0.02; pores of their distal caps
0.003 to 0.006, bars 0.003.

_Habitat._--Pacific, central area, Stations 271 to 274, depth 2350 to 2750
fathoms.

Subgenus 3. _Ommatocorona_, Haeckel.

_Definition._--Surface of the cortical shell spiny, on every chamber a
regular circle or coronal of radial spines.

{395}6. _Ommatocampe chætopodum_, n. sp.

Cortical shell with spiny surface, composed of six (or more) kidney-shaped
chambers of different size and form. Both proximal chambers kidney-shaped,
with subregular, circular, hexagonally framed pores, about as broad as the
bars. All the following chambers hemispherical, with irregular, roundish
pores; at the base of every chamber a circle of ten to twelve larger square
pores. The beams between these latter are prolonged into free radial
spines, twice as thick as the bars. Therefore every chamber is surrounded
by a circle of radial spines, like those of _Panicium coronatum_ (Pl. 40,
fig. 4). Both medullary shells lenticular.

_Dimensions._--Length of the six-chambered cortical shell 0.3, greatest
breadth 0.08, pores and bars of both proximal chambers 0.01; pores of the
other chambers 0.002 to 0.008; square pores 0.02, bars 0.005; length of the
coronal spines 0.03, thickness 0.01.

_Habitat._--Indian Ocean, Madagascar, Rabbe.

Subgenus 4. _Ommatacantha_, Haeckel.

_Definition._--Surface of the shell smooth or spiny, on the poles of the
main axis occur two strong opposite polar spines.

7. _Ommatocampe amphilonche_, n. sp.

Cortical shell composed of six kidney-shaped chambers of nearly the same
size and structure; every chamber twice as broad as long, with four to five
transverse rows of irregular, roundish pores, once to four times as broad
as the bars. Both medullary shells lenticular. Surface of the cortical
shell covered with bristle-like spines. On both poles of the main axis one
larger, strong, conical spine, about half as long as this axis, and on the
base as broad as the inner medullary shell. (Resembles on the whole
_Desmartus larvalis_, Pl. 40, fig. 12, but without external mantle, and
with two solid polar spines instead of the polar tubes.)

_Dimensions._--Length of the six-chambered cortical shell 0.24; greatest
breadth of each chamber 0.08; pores 0.002 to 0.08, bars 0.002; length of
the polar spines 0.12, basal thickness 0.02.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Genus 170. _Ommatartus_,[209] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Zygartida# with simple cortical shell and double medullary
shell, with two hollow fenestrated tubes, opposite on both poles of the
main axis.

The genus _Ommatartus_ differs from _Ommatocampe_ by the development of two
hollow fenestrated tubes on both poles of the main axis, and bears
therefore the same relation to it as _Cannartidium_ to _Cyphonium_. The
former two genera differ from the two latter by the augmentation of the
chambers of the cortical shell.

{396}1. _Ommatartus amphicanna_, n. sp.

Cortical shell with spiny surface, composed of six chambers of equal size
and kidney-shaped; every chamber with four to five transverse rows of
irregular, roundish pores, twice to three times as broad as the bars. Both
medullary shells lenticular. Polar tubes conical, about as long as one
chamber, with smaller pores. Both medullary shells spheroidal, more or less
compressed. (Resembles _Desmartus larvalis_, Pl. 40, fig. 12, but is
without the external shell.)

_Dimensions._--Length of the six-chambered cortical shell 0.24; greatest
breadth of each chamber 0.07; pores 0.008 to 0.012, bars 0.004; length of
the polar tubes 0.04 to 0.05, basal thickness 0.02.

_Habitat._--Pacific, central area, Stations 270 to 274, depth 2350 to 2925
fathoms.

2. _Ommatartus amphisiphon_, n. sp.

Cortical shell with spiny surface, composed of six or eight chambers of
different size and form. Both proximal chambers kidney-shaped, each with
five to six transverse rows of irregular, polygonal pores, three to four
times as broad as the bars. The other (four to six) chambers somewhat
smaller, more hemispherical, with smaller, irregular pores, but on the base
of each a circle of ten to twelve larger square pores. Polar tubuli
prismatic, about half as long as the main axis, with prominent edges and
longitudinal rows of smaller pores between them. Both medullary shells
lenticular. (Resembles closely _Panarium tubularium_, Pl. 40, fig. 9, but
with six to eight chambers instead of four, and with longer tubuli.)

_Dimensions._--Length of the six-chambered cortical shell 0.26; greatest
breadth of the proximal chambers 0.09, of the distal chambers 0.07; pores
of the former 0.01, of the latter 0.004; large square pores 0.013, bars
0.003; length of the polar tubes 0.1 to 0.12, thickness 0.02.

_Habitat._--South Pacific, Station 297, depth 1775 fathoms.

3. _Ommatartus amphobolus_, n. sp.

Cortical shell with spiny surface, composed of six chambers of different
size and form. Both proximal chambers kidney-shaped, with four to five
transverse rows of subregular, circular pores, twice as broad as the bars.
Both middle chambers cap-like, with a basal circle of ten to twelve larger
square pores, and small roundish pores on the cap. Both distal chambers
smaller, conical, with very small roundish pores. Polar tubuli conical,
nearly half as long as the main axis, also with very small pores. Both
medullary shells lenticular.

_Dimensions._--Length of the six-chambered shell 0.28, greatest breadth
0.08; pores of the proximal chambers 0.01, bars 0.005; pores of the distal
chambers and the polar tubes 0.002 to 0.004, bars 0.002; length of the
polar tubes 0.13, basal thickness 0.03.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

Genus 171. _Desmocampe_,[210] n. gen.

_Definition._--#Zygartida# with double cortical shell and double medullary
shell, without polar tubes.

{397}The genus _Desmocampe_ differs from _Ommatocampe_ in the duplication
of the jointed cortical shell; the radial spines, which start from the
surface of the inner cortical shell, are connected one with another by
transverse communicating branches which form an outer envelop around it;
but this reticulated mantle is commonly not quite perfect and more or less
irregular.

1. _Desmocampe catenula_, n. sp.

Inner cortical shell with six to eight chambers of the same size and form.
Every chamber kidney-shaped, with three to four transverse rows of
circular, subregular pores, twice as broad as the bars. Outer cortical
shell cylindrical, hemispherical at both poles, with smooth surface and
irregular, roundish pores of very different size. Distance between the two
cortical shells equals the diameter of the outer medullary shell, which,
like the inner, is spherical. (Resembles Ommatocampe polyarthra, Ehrenberg,
1872, _loc. cit._, Taf. vi, fig. 9, but differs in the external mantle.)

_Dimensions._--Length of the six-chambered inner cortical shell 0.16, of
the outer 0.21; greatest breadth of each chamber of the former 0.05, of the
latter 0.09; pores of the inner 0.006, bars 0.003; pores of the outer 0.002
to 0.008, bars 0.002.

_Habitat._--North Pacific, Station 237, off Japan, surface.

2. _Desmocampe tænioides_, n. sp.

Inner cortical shell with six to ten chambers of nearly the same size and
form; the distal chambers somewhat smaller. Every chamber kidney-shaped,
with four to five transverse rows of irregular roundish pores, twice to
three times as broad as the bars. Outer cortical shell cylindrical,
hemispherical at both poles, with spiny surface; its pores like those of
the inner, but the bars between them much thinner. Distance between the two
cortical shells equals the diameter of the outer medullary shell. Both
medullary shells lenticular. (Resembles _Desmartus larvalis_, Pl. 40, fig.
12, but is without polar tubes.)

_Dimensions._--Length of the six-chambered inner cortical shell 0.25, of
the outer 0.3; greatest breadth of the former 0.07, of the latter 0.11;
pores 0.005 to 0.012; bars of the inner shell 0.004, of the outer 0.001.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Desmocampe aphrodite_, n. sp.

Inner cortical shell with six to ten chambers of different size and
structure. Both proximal chambers kidney-shaped, with five to six
transverse rows of subregular, circular, hexagonally-framed pores, twice as
broad as the bars. All following chambers cap-like, with much smaller,
irregular, roundish pores, on the base of each a circle of ten to twelve
large square pores. Outer cortical shell cylindrical, on both poles
hemispherical, with spiny surface and very delicate network of small
polygonal pores. Both medullary shells spherical. (The inner cortical shell
of this species resembles that of _Ommatocampe nereis_, Pl. 40, fig. 10;
the outer that of _Cyphocolpus virginis_, Pl. 40, fig. 11.)

{398}_Dimensions._--Length of the six-chambered internal cortical shell
0.25, of the outer 0.3; greatest breadth of the former 0.08, of the latter
0.12; pores of the proximal chambers of the inner shell 0.01, bars 0.005;
pores of the distal chambers 0.003 to 0.005, square pores 0.02, bars 0.003;
pores of the external cortical shell 0.002 to 0.006, bars 0.001.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

4. _Desmocampe atractus_, n. sp.

Inner cortical shell with six chambers of very different size and
structure. Both proximal chambers kidney-shaped, with seven to eight
transverse rows of subregular, roundish pores, twice to three times as
broad as the bars. Both middle chambers cap-like, on the base with a circle
of eight to ten very large square pores, on the distal cap with small
irregular, roundish pores. Both distal chambers conical, also with small
irregular, roundish pores. Outer cortical shell spindle-shaped, inflated in
the equatorial zone, tapering conically towards both poles, with very
delicate network of small roundish pores and thin bars. Surface covered
with innumerable very small spines. Both medullary shells lenticular.
(Resembles very much _Peripanartus atractus_, Pl. 40, fig. 7, but differs
in the number of the chambers and their proportion to the outer mantle,
which envelops spindle-like all six chambers.)

_Dimensions._--Length of the six-chambered internal cortical shell 0.27, of
the outer 0.32; greatest breadth of the former 0.09, of the latter 0.15;
pores of the inner cortical shell (on an average)--proximal chambers 0.008,
middle chambers 0.02, distal chambers 0.004, bars 0.004; pores of the outer
cortical shell 0.002 to 0.004, bars 0.002.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

Genus 172. _Desmartus_,[211] n. gen.

_Definition._--#Zygartida# with double cortical shell and double medullary
shell, with two hollow fenestrated tubes, opposite on both poles of the
main axis.

The genus _Desmartus_ differs from _Desmocampe_ by the development of two
hollow fenestrated tubes, opposite on both poles of the main axis, and
bears therefore the same relation to it as _Ommatartus_ to _Ommatocampe_.
Both the former genera can be produced from the two latter by duplication
of the cortical shell.

1. _Desmartus larvalis_, n. sp. (Pl. 40, fig. 12).

_Zygartus larvalis_, Haeckel, 1881, Prodromus et Atlas (pl. xl, fig. 12).

Inner cortical shell composed of six kidney-shaped chambers of nearly the
same size and structure; every chamber twice as broad as long, with four to
five transverse rows of irregular, roundish pores, twice to five times as
broad as the bars; the basal pores of the distal chambers {399}somewhat
larger and more square. Outer cortical shell cylindrical, hemispherical at
both poles, with irregular, polygonal pores, on an average twice as large
as those of the inner, but the bars between them much thinner. Outer
surface spiny. Both medullary shells lenticular. Polar tubes conical, a
little longer than one internal chamber, as broad at the base as the inner
medullary shell. Sometimes the tubes exhibit prominent edges (as in the
lower spine of fig. 12); the pores of these are very small, and roundish.

_Dimensions._--Length of the six-chambered internal cortical shell 0.23, of
the external 0.3; greatest breadth of the former 0.07, of the latter 0.11;
pores of the inner shell 0.004 to 0.01, of the outer 0.01 to 0.02; bars of
the former 0.002, of the latter 0.001; length of the polar tubes 0.05,
basal thickness 0.02.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

2. _Desmartus tubulatus_, n. sp.

Inner cortical shell composed of six to ten kidney-shaped chambers,
tapering in size towards both poles, every chamber with six to seven
transverse rows of irregular, roundish pores, twice to six times broader
than the bars. Outer cortical shell spindle-shaped, in the equatorial zone
inflated, tapering towards both poles, its network similar to the inner,
only more delicate. Polar tubes conical, twice as long as an inner chamber,
as broad at the base as the outer medullary shell. Both medullary shells
lenticular.

_Dimensions._--Length of the six-chambered internal cortical shell 0.25, of
the external 0.32; greatest breadth of the former 0.08, of the latter 0.12;
pores of the inner shell 0.002 to 0.012, of the outer 0.003 to 0.01; bars
of the former 0.002, of the latter 0.001; length of the polar tubes 0.07,
basal thickness 0.03.

_Habitat._--North Pacific, Station 256, depth 2950 fathoms.

Genus 173. _Zygocampe_,[212] n. gen.

_Definition._--#Zygartida# with triple (or multiple) cortical shell and
double medullary shell, without polar tubes.

The genus _Zygocampe_ differs from _Desmocampe_ and _Ommatocampe_ by the
multiplication of the cortical shell, which is composed of three or more
concentric envelopes. The three mentioned genera form therefore a
phylogenetic series, produced by the concentric increase on the outside of
the jointed cortical shell. Commonly the second cortical shell is not as
complete as the first (or innermost), and the third (or outermost) is yet
more incomplete. Rarely the number of the concentric cortical shells
surpasses three.

1. _Zygocampe pupula_, n. sp.

Inner cortical shell with six to twelve chambers of nearly the same size
and form. Every chamber kidney-shaped, with four to five transverse rows of
circular, subregular pores, twice as {400}broad as the bars. Middle
cortical shell cylindrical, hemispherical at both poles, with subregular
circular pores somewhat smaller than those of the inner; the bars also
thinner. Outer cortical shell of the same form as the middle, but with very
delicate network, and quite irregular, roundish pores and very thin bars.
Surface quite smooth. Both medullary shells lenticular. (May be regarded in
a phylogenetic as well as an ontogenetic sense, as the further
developmental form of _Desmocampe catenula_ and _Ommatocampe polyarthra_.)

_Dimensions._--Length of the six-chambered inner cortical shell 0.17, of
the middle 0.22, of the outer 0.27; greatest breadth of the first 0.05, of
the second 0.09, of the third 0.12; pores of the inner cortical shell
0.005, of the middle 0.004, of the outer 0.003 to 0.012; bars of the first
0.003, of the second 0.002, of the third 0.001.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Zygocampe corasium_, n. sp.

Inner cortical shell with six to eight chambers of different size and form.
Both proximal chambers kidney-shaped, with five to six transverse rows of
subregular, circular, hexagonally framed pores, twice as broad as the bars.
All following chambers cap-like, the distal somewhat smaller; their pores
much smaller, irregular, roundish; only at the base of each chamber a
circle of ten to twelve large square pores. Middle cortical shell
cylindrical, in the equatorial zone a little constricted, hemispherical at
both poles, with irregular delicate network of roundish, polygonal meshes.
Outer cortical shell of the same form as the middle, but with a very
delicate and quite irregular network of polygonal meshes. Commonly this
outer mantle is incomplete, and sometimes interwoven in a spongy manner
with the middle (or also with the inner). Surface covered with many
irregular, thin, bristle-like spines. Both medullary shells lenticular.
(May be considered in a phylogenetic and ontogenetic sense as a further
developmental stage of _Desmocampe aphrodite_ and _Ommatocampe nereis_, Pl.
40, fig. 10.)

_Dimensions._--Length of the six-chambered inner cortical shell 0.24, of
the middle 0.3, of the outer 0.36; greatest breadth of the first 0.08, of
the second 0.12, of the third 0.16, pores of the first (on an average)
0.005 to 0.02, of the second 0.003 to 0.015, of the third 0.01 to 0.05;
bars corresponding 0.005 or 0.003 or 0.001.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

3. _Zygocampe chrysalidium_, n. sp. (Pl. 40, fig. 13).

Inner cortical shell with six to eighteen chambers of different size and
form. Both proximal chambers kidney-shaped, with six to seven transverse
rows of subregular, circular pores, three to four times as broad as the
bars. All following chambers with more irregular pores, with a circle of
ten to twelve larger square pores at the base. Middle cortical shell with
smaller, irregular, roundish pores. Outer cortical shell with larger
polygonal, quite irregular pores. Both outer shells appear commonly
incomplete or somewhat irregularly developed, and sometimes in a spongy
manner interwoven with one another. Surface covered with irregular,
bristle-like spines. Both medullary shells spherical. The breadth of the
chambers decreases gradually towards both poles, so that the whole shell
assumes a spindle form. Sometimes on both poles is developed a short
conical {401}polar tube (in fig. 13 only on the upper pole), and so this
species is transformed into _Zygartus chrysalis_.

_Habitat._--Pacific, central area, Stations 272 to 274, depth 2350 to 2750
fathoms.

Genus 174. _Zygartus_,[213] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Zygartida# with triple (or multiple) cortical shell and
double medullary shell, with two hollow fenestrated tubes, opposite on both
poles of the main axis.

The genus _Zygartus_ differs from Zygocampe by the development of two
hollow fenestrated tubes, on the two opposite poles of the main axis. It
exhibits therefore to the latter the same relation that _Desmartus_ does to
_Desmocampe_ and _Ommatartus_ to _Ommatocampe_; it differs from these by
the multiplication of the cortical shell, which is composed of at least
three concentric envelopes.

1. _Zygartus doliolum_, n. sp.

Inner cortical shell composed of six (or more) kidney-shaped chambers, all
nearly of the same size and structure, every chamber with four to five
transverse rows of irregular, roundish pores, twice to four times as broad
as the bars; the basal pores scarcely larger than the apical pores. Middle
cortical shell nearly of the same structure as the inner, only the pores
larger and the bars thinner. Outer cortical shell cylindrical,
hemispherical at both poles, its network very delicate, with large
irregular, polygonal pores, and very thin bars between them; its surface
covered with thin bristle-like spines. Both medullary shells lenticular.
Polar tube cylindrical, with conical apex, and with very small pores; its
length equals the breadth of two internal chambers, its breadth that of the
inner medullary shell. (This species appears to be a further developed form
of _Ommatocampe annulata_ and _Desmartus larvalis_, Pl. 40, fig. 12.)

_Dimensions._--Length of the six-chambered inner cortical shell 0.25, of
the middle 0.3, of the outer 0.35; greatest breadth of the first 0.07, of
the second 0.11, of the third 0.15; pores of the inner shell 0.005 to 0.01,
of the middle 0.003 to 0.007, of the outer 0.008 to 0.02; bars of the first
0.002, of the second and third 0.001; length of the polar tubes 0.15, basal
thickness 0.02.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

2. _Zygartus chrysalis_, n. sp. (Pl. 40, fig. 13).

Inner cortical shell composed of six to twenty (commonly eight to twelve)
chambers of different size and form. Both proximal chambers nearly
hemispherical, with spiny surface and subregular, {402}circular pores,
three to four times as broad as the bars. All following chambers cap-like,
tapering towards both poles, with more irregular, roundish pores, with a
circle of ten to twelve larger square pores at the base. Middle cortical
shell with smaller roundish pores. Outer cortical shell with larger
polygonal, quite irregular pores. Both medullary shells lenticular. Polar
tubes conical or cylindrical with conical apex, of very variable length,
sometimes not longer than one internal chamber, at other times twice to
four times as long (in the figured specimen not fully developed, as also a
part of the shells). Differs from _Zygocampe chrysalis_ only by the
production of polar tubes.

_Dimensions._--Length of the six-chambered inner cortical shell 0.25, of
the middle 0.3, of the outer 0.35; greatest breadth of the first 0.07, of
the second 0.11, of the third 0.14; other measures the same as in
_Zygocampe chrysalis_; length of the tubes 0.05 to 0.12 or more.

_Habitat._--Pacific, central area, Stations 270 to 274, depth 2350 to 2925
fathoms.

----

Suborder V. DISCOIDEA, Haeckel (Pls. 31-38, 41-48).

_Discida_ vel _Discoidea_, Haeckel, 1862, Monogr. d. Radiol., pp. 56,
476.
_Discoida_, _Discoidea_, _Discida_, Haeckel, 1878, Protistenreich, p.
103.

_Definition._--SPUMELLARIA with discoidal or lenticular central capsule
(often with radial prolongations, rarely allomorphic); with discoidal or
lenticular fenestrated siliceous shell (often with radial spines or
fenestrated arms, rarely allomorphic). Growth reduced or diminished in the
direction of one dimensive axis.

The section #Discoidea# comprises those SPUMELLARIA in which the
fenestrated shell is more or less discoidal or lenticular, flattened or
compressed in the direction of one axis. The geometric fundamental form of
the latticed shell, which in the #Sphæroidea# is a sphere, here becomes a
flat disk, like a medal, or a biconvex lens, sometimes also a biconcave
lens. The #Discoidea# can be derived from the #Sphæroidea# by shortening of
one axis. This shortened vertical axis is the main axis of the disk; both
its poles are constantly equal. Perpendicular to this axis is the
equatorial plane of the disk by which it becomes divided into equal halves.
In the simplest forms of #Discoidea# all axes of this horizontal equatorial
plane (all "equatorial axes" or "cross axes") are equal; in the most of the
genera and species these cross axes are different, so that rays of stronger
growth ("perradii") alternate with rays of weaker growth ("interradii").
The number of these cross axes distinguishable is commonly two to four,
rarely more. In the direction of these are developed either radial marginal
spines or spongy arms.

The order #Discoidea# was founded in my Monograph (1862, p. 476) as the
family "Discida" (Radiolaria with flat discoidal or biconvex lenticular
shell), comprising the {403}"_Calodictya_ and _Lithocyclidina_" of
Ehrenberg and a great part of his "_Haliommatina_." As three different
subfamilies of that family I separated the Coccodiscida (with five genera),
the Trematodiscida (with seven genera), and the Discospirida (with two
genera; Monogr. d. Radiol., p. 485). A fourth group of #Discoidea# was
constituted by the Spongodiscida (with eight genera, including the
_Spongocyclida_), which at that time I united with the Spongurida, because
of their spongy structure (_loc. cit._, p. 452).

As the number of fossil #Discoidea# found in the Tertiary rocks of Barbados
and of the Mediterranean shores (Sicily and Greece) is comparatively very
large, we find even in the first system of Polycystina of Ehrenberg (1847),
not less than twelve genera distinguished, viz., six _Calodictya_, two
_Haliommatina_, and four _Lithocyclidina_ (Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, 1847, p. 53). The whole number of Radiolarian genera
distinguished in that first system was forty-four. The diagnoses of them
given by Ehrenberg were as usual very insufficient. The characters of the
three families given by him were the following:--_Calodictya_--"Testarum
intus spongiosarum et nucleo destitutarum orbes; _Haliommatina_--Testæ
subglobosæ nucleus radiatus; _Lithocyclidina_--Testarum disci in media
parte nucleati margine celluloso." In the latest work of Ehrenberg (1875,
p. 157) the same system was repeated, but some new genera added; and
thirty-eight different species, appertaining to the #Discoidea#, were
figured in the same work (Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, Tafs.
xx.-xxx.).

Richard Hertwig, 1879, in his excellent work, Der Organismus der
Radiolarien (pp. 57-68), gave a detailed description of the skeleton of
some #Discoidea#, and arrived at the conclusion that this whole family had
a spirally constructed skeleton, and should therefore be derived from the
Lithelida. But this conclusion is certainly erroneous, and in my opinion
the whole explanation of that spiral structure, and of its signification in
the development of #Discoidea#, is the weakest part of that otherwise very
important work.

In my Prodromus (1881, p. 456) I gave a provisional system of the #Discida#
or #Discoidea# from the immense quantity of new material collected by the
Challenger, and could distinguish not less than eighty-four genera. This
number is from subsequent research only augmented by seven, so that in the
following pages ninety-one genera with five hundred and one species are
described. In the Prodromus I had disposed them in four different families,
which number is now increased to six. These six families can be again
disposed in two main groups or sections, the #Phacodiscaria# and the
#Cyclodiscaria#, each section with three families.

The #Phacodiscaria# are characterised by the possession of a typical
"phacoid shell," and contain the three families Cenodiscida, Phacodiscida,
and Coccodiscida. On the other hand, the #Cyclodiscaria# are distinguished
by the absence of such a "phacoid shell," and contain the three families
Porodiscida, Pylodiscida, and Spongodiscida. Both sections exhibit an
analogous development.

{404}The #Cenodiscida# (Pl. 48, figs. 1-3) open the series of the
#Discoidea# as their simplest forms; a discoidal or lenticular simple
lattice-shell encloses a central capsule of the same form, and is separated
from it by the calymma or jelly-veil. The common ancestral form of this
family is _Cenodiscus_, without radial marginal spines; it can be derived
from _Cenosphæra_ in the most simple way, by flattening in one axis. If on
the equatorial margin of the lens a peculiar solid girdle be developed, we
obtain _Zonodiscus_; in all other genera of the Cenodiscida radial spines
are developed on the margin. As the simple lenticular cortical shell of
_Cenodiscus_, in which the central capsule is enclosed, is most
characteristic not only of this family, but also of the two following
families, we call it the phacoid shell (that is, a lenticular extracapsular
or cortical lattice-shell).

The #Phacodiscida# (Pls. 31-35), the second family, have the same
extracapsular "phacoid shell" as the Cenodiscida, but differ from these by
the possession of one or two intracapsular concentric medullary shells,
which are connected with the former by radial beams, perforating the
lenticular central capsule. The radial beams are commonly numerous, and
arranged in two opposite bunches around the shortened main axis. But often
also besides these occur other longer radial beams, situated in the
equatorial plane; the number of these is commonly four, and they form a
regular rectangular cross, lying opposite in pairs in two equatorial
diameters, perpendicular one to another. In the simplest genera of this
family (the _Sethodiscida_) the equatorial margin of the phacoid shell is
simple or surrounded by a solid smooth girdle; in all other genera are
developed on the margin solid radial spines lying in the equatorial plane,
either regularly disposed in a somewhat constant number (two to eight,
_Heliosestrida_), or irregularly disposed, in a larger and more variable
number (ten to twenty or more, _Heliodiscida_).

The #Coccodiscida# (Pls. 36-38) form a third family of the #Discoidea#,
directly associated with the Phacodiscida; both have the same
characteristic extracapsular "phacoid shell," which is connected by radial
beams with a simple or double, intracapsular medullary shell. But whilst in
the foregoing family the equatorial margin of the phacoid shell is simple
or only armed with radial spines, in the Coccodiscida it is surrounded by
peculiar concentric chambered girdles, or rings, which resemble those of
the following family, the Porodiscida. Each of these "chambered girdles" is
composed of a circular ring in the equatorial plane, a variable number of
radial beams dividing it into incomplete chambers, and two porous
cover-plates or "sieve-plates," covering the upper and lower face of the
disk. These sieve-plates may be regarded as incomplete lenticular cortical
shells, which are only developed in the peripheral part of the disk, whilst
their central part is represented by the only complete cortical shell, the
"phacoid shell." The number of these concentric chamber-girdles amounts to
from one to ten or more. The margin of the disk is either simple
(Lithocyclida) or armed with radial spines (Stylocyclida), or provided with
two to five chambered radial arms (Astracturida); the structure of the arms
is the same as that of the girdles.

{405}The #Porodiscida# (Pls. 41-47), the largest family of all #Discoidea#,
begins the series of #Cyclodiscaria#, or those #Discoidea# in which there
is no "phacoid shell," but a small simple central chamber surrounded by a
number of small latticed chambers of nearly the same size and form. In the
Porodiscida these chambers are arranged in complete circular concentric
rings or spiral convolutions; in the small family of Pylodiscida the
central chamber is surrounded by three radial arm-chambers separated by
three open spaces; in the third family of #Cyclodiscaria#, the
Spongodiscida, all the chambers are arranged more or less irregularly, and
the whole disk becomes spongy; also the surface of the disk is spongy,
whilst in both former families it is covered by two regular even porous
plates or "sieve-plates." Probably all #Cyclodiscaria# can be derived
phylogenetically from _Archidiscus_, a very small and simple lenticular
disk, which is composed of a small spherical latticed central chamber and
of a single concentric chambered ring or girdle; the margin of this ring is
connected with the central chamber by a variable number of radial beams.
This _Archidiscus_ can be derived either from _Saturnalis_ (Pl. 13, fig.
16.) by the development of lattice-work between the equatorial ring and
both polar faces of the concentric central chamber, or from _Sethodiscus_
(Pl. 33, figs. 1-3) by flattening of the lenticular shell, so that the
enclosed inner medullary shell (the central chamber) meets the outer
phacoid shell at both poles.

The Porodiscida are commonly flat or biconvex (rarely biconcave) disks, the
central chamber of which is surrounded not by a single, but by a variable
number (commonly three to six) of concentric chambered girdles or rings;
they arise from _Archidiscus_ by apposition of new concentric chambered
rings around the first ring, all lying in the equatorial plane. Afterwards
the disk often becomes thickened by apposition of concentric chamber-rings
on both flat sides also, so that two to four or more layers are stratified
one over the other. The circular concentric rings often become interrupted,
or spirally convoluted (wholly or partially); also the chambers sometimes
become irregularly crowded. But in all cases both surfaces of the disk
(upper and lower) continue to be porous plates or sieve-plates, at least in
the centre, but they never become spongy.

The margin of the disk exhibits in the Porodiscida a great variety of
different forms, serving for distinction of subfamilies and of genera. In
the Trematodiscida the margin remains quite simple, as in the Archidiscida,
or is only surrounded by a hyaline equatorial girdle. In the Ommatodiscida
it is distinguished by one or two peculiar oscula, surrounded by a corona
of spines. The Stylodictyida are distinguished by a number of solid radial
spines, and the Euchitonida by a number of chambered, or spongy, radial
arms, arising from the margin of the disk and lying in the equatorial
plane. The variety of these radial marginal appendages is in the
Porodiscida much greater than in the Coccodiscida.

The #Pylodiscida# (Pl. 48, figs. 12-20) represent a new, small, but very
remarkable family of #Discoidea#, all triradial, and distinguished by the
peculiar formation of {406}large open spaces in the latticed discoidal
shell, which reappear in a similar shape among the #Larcoidea# in the
Pylonida (_Tetrapyle_, &c.). We get the best understanding of this peculiar
formation if we return to _Archidiscus_, probably the common ancestral form
of all #Cyclodiscaria#, of the Porodiscida as well as of the Pylodiscida
and Spongodiscida. In some species of _Archidiscus_ (_Archidiscus
hexoniscus_, _Archidiscus pyloniscus_, &c.) the small lenticular shell is
composed of a spherical latticed central chamber and of a concentric
equatorial girdle composed of six such chambers, either all six equal, or
alternately larger and smaller. This latter form is nearly identical with
_Triodiscus_, and if we imagine the lattice-work of only three
ring-chambers complete, whilst that of the three alternating chambers is
reduced to the marginal ring, we get _Triopyle_, by loss of this ring
_Triolene_ (a disk composed of four simple lattice-chambers, lying in one
plane, three radial around one central spherule). The genera mentioned form
together the subfamily of Triopylida. In the second subfamily, Hexapylida,
the same formation is doubled; here three double arm-chambers are separated
by three double spaces (two in each radius). Also here the three distal
spaces may be either quite open (_Pylolena_), or half closed by the
marginal girdle (_Hexapyle_), or quite loosely latticed (_Pylodiscus_). If
the margin of this latter form become surrounded by a perfect chambered
equatorial girdle, we get _Discozonium_, and if this acquire a peculiar
marginal ostium (surrounded by a corona of spines) we arrive at
_Discopyle_. These two latter genera form the third subfamily, the
Discopylida. The eight genera of Pylodiscida represent therefore a
continuous phylogenetic series.

The #Spongodiscida# are the sixth and last family of the #Discoidea#,
differing from the five other families in the irregular, spongy structure
of the discoidal skeleton; both surfaces of the flat disk (upper and lower)
are here principally covered with a rough, spongy framework, whilst in the
five other families they are covered by the flat and smooth porous plates
or sieve-plates. Nevertheless there is no sharp boundary between the
Spongodiscida and the closely allied Porodiscida. In these latter also the
discoidal shell becomes often more or less spongy (mainly in the peripheral
part, _e.g._, in _Myelastrum_, Pl. 47); but at least the central part of
the disk here remains constantly covered by sieve-plates. The massive
skeleton of the Spongodiscida is either of perfectly irregular structure,
only composed of innumerable fine branched siliceous threads, interwoven in
all possible directions; or only the outer part of the disk is composed of
such spongy framework, whilst the central part is more or less distinctly
composed of concentric chambered rings, as in the Porodiscida. These latter
forms indeed exhibit an immediate transition to this family, and were
formerly (in 1862) separated by me as Spongocyclida. Also the polymorphous
shape of the disk margin in the Spongodiscida is quite analogous to that of
the Porodiscida. Whilst in the first subfamily, the Spongophacida, the
margin is quite simple; in the second, the Spongotrochida, it is armed with
solid radial spines; and in the third, the Spongobrachida, it is provided
with two, three, {407}or four spongy, radial arms--the former as well as
the latter lying in the equatorial plane of the disk, either regularly or
irregularly disposed.

_The Equatorial Margin_ of the lenticular disk exhibits in all six families
of #Discoidea# similar characters, mainly serving for the distinction of
subfamilies and genera. In the most primitive genera of all six families
the margin is simple, without radial prolongations (spines or arms); it is
quite simple in _Cenodiscus_, _Sethodiscus_, _Phacodiscus_, _Lithocyclia_,
_Coccodiscus_, _Archidiscus_, _Porodiscus_, _Pylodiscus_, and
_Spongodiscus_. In some genera the simple margin of the lenticular disk is
bordered and surrounded by a thin, hyaline, equatorial girdle of silex,
either quite solid or slightly porous (_Zonodiscus_, _Periphæna_,
_Perizona_, _Perichlamydium_, and _Spongophacus_).

A quite peculiar and remarkable character of few genera is the development
of one or two oscula, larger marginal openings, which are surrounded by a
corona of spines, and probable are fit for the issue a peculiar bunch of
pseudopodia or of a "sarcode flagellum." Such oscula occur only in two
families of #Cyclodiscaria#; in the Porodiscida and Pylodiscida; in the
former _Ommatodiscus_, in the latter _Discopyle_ (Pl. 48, figs. 19, 20) is
distinguished by a single marginal osculum; besides this, in the former
occurs _Stomatodiscus_, with two such oscula, opposite on the poles of one
axis (Pl. 48, fig. 8). These oscula may be compared with the similar polar
formations in some Ellipsida (_Lithomespilus_) and in many Cyrtoidea; but
they do not prove a nearer affinity with the latter, and are only
analogous, not homologous.

_Radial Spines_ occur on the margin in the equatorial plane of the
#Discoidea# in the greatest variety of number, form, size, and disposition.
If the number be low (between two and eight) they are commonly regularly
disposed; if the number be larger (ten to twenty or more) their disposition
becomes commonly more or less irregular. The regular disposition is of
great promorphological importance, as indicating the axes in which the
growth is preponderant, and introduces other peculiar radial formations.
Regarding these axes we can generally distinguish two groups, _Artiacantha_
with a paired number (two, four, eight), and _Perissacantha_ with odd
numbers (usually three). The section of _Artiacantha_ could be divided into
three following groups:--A. Stylodiscida, with two radial spines only,
lying opposite on both poles of one equatorial axis (the "first cross
axis")--_Stylodiscus_, _Sethostylus_, _Stylocyclia_, _Xiphodictya_,
_Spongolonche_ (Pl. 31, figs. 9-12; Pl. 38, fig. 1; Pl. 42, figs. 10-12,
&c.); B. Staurodiscida, with four radial spines, lying opposite in pairs on
the poles of two crossed equatorial axes, perpendicular to one another
(first and second cross axes)--_Crucidiscus_, _Sethostaurus_,
_Staurocyclia_, _Staurodictya_, _Spongostaurus_ (Pl. 31, figs. 1-8; Pl. 37,
figs. 1-4; Pl. 42, figs. 1-6; Pl. 48, fig. 2, &c.); C. Octostylida, with
eight radial spines, opposite in pairs in four axes, which are crossed at
angles of 45°--_Heliosestrum_, _Astrosestrum_, &c. (Pl. 32, figs. 4, 5; Pl.
34, figs. 3, 6); in this latter case sometimes the radial symmetry is the
same as in many Medusæ, four larger (perradial) {408}spines alternating
with four smaller (interradial), indicating radii of first and second
order. The section of _Perissacantha_ is much smaller, and commonly
represented only by triradial forms, with three spines at equal distances
(120°)--_Triactis_, _Tripocyclia_, _Tripodictya_, _Spongotripus_, &c. (Pl.
33, fig. 6; Pl. 37, fig. 5; Pl. 42, figs. 7-9).

_Radial Arms_ on the margin of the disk appear in similar variety of
number, form, and disposition as the radial spines; but the number is here
commonly limited to from two to four, rarely five to six. The arms are
absent in the families Cenodiscida and Phacodiscida; in the four other
families they return under similar forms. These arms are direct
prolongations of the disk, and exhibit the same structure, so that they may
be regarded both as centrifugal productions of certain radii, and also
inversely as peripheral parts of a disk, the interjacent radii of which are
reduced. The regular disposition and shape of the arms, an important
character for the distinction of genera and species, is repeated in a quite
analogous manner in the four above mentioned families, so that we can
distinguish the following groups--A. Amphibrachida, with two radial arms,
opposite on the poles of one equatorial axis (the first cross
axis)--_Diplactura_, _Amphibrachium_, _Spongobrachium_ (Pl. 38, figs. 3-5;
Pl. 44, figs. 6-11); B. Triobrachida, with three radial arms; the most
important group (with all Pylodiscida); either all three arms are equal and
disposed at equal distances (_Trigonactura_, _Dictyastrum_,
_Rhopalodictyum_, Pl. 38, figs. 6-9; Pl. 43, figs. 5, 13, 16; Pl. 48, figs.
12-19), or a single odd arm differs in size and position, and is often
larger than both the opposite paired arms (_Rhopalastrum_, _Euchitonia_,
Pl. 43, figs. 6, 10, 15, &c.); C. Tetrabrachida, with four radial arms,
opposite in pairs in two crossed axes, commonly perpendicular one to
another, _Stauractura_, _Stauralastrum_, _Spongaster_, &c. (Pls. 46, 47).

The arms are commonly simple, undivided, but sometimes also forked or
branched (Pl. 43, figs. 15, 16; Pl. 47). Their basal parts are either free,
separately inserted into the margin of the circular central disk, or they
are connected by a "patagium," a peculiar connecticulum, like a
web-membrane, which is composed of a chambered, commonly more or less
spongy framework, different in texture from the lattice-work of the arms
(Pl. 38, figs. 8, 9; Pl. 43, figs. 9-16; Pl. 46). Sometimes the patagium
overgrows the whole shell. A peculiar modification of it appears in
_Stephanastrum_ (Pl. 44, fig. 1), where only the distal parts of the arms
are connected by the ring-shaped patagium, whilst the basal parts are free;
therefore open gates rest between them, like those of the Pylodiscida (Pl.
48, figs. 12-20).

The _Central Capsule_ of the #Discoidea# is constantly discoidal, more or
less lenticular; in some cases more biconvex, with vaulted faces and thin
margin; in others more medal-shaped, with flat faces and thick margin. In
the Cenodiscida alone the capsule lies freely inside the simple phacoid
shell, and is separated from it by the jelly-veil. In the other five
families the capsule encloses the central parts of the skeleton, and is
enveloped by the superficial parts of it, whilst its membrane is perforated
by radial beams {409}connecting the latter with the former. In the
Phacodiscida and Coccodiscida the capsule encloses the simple or double
medullary shell, but is itself enclosed by the cortical phacoid shell. In
all #Cyclodiscaria# (in the Porodiscida, Pylodiscida, and Spongodiscida)
the capsule fills out the greatest part of the chambered or spongy
skeleton, and is only protected by the superficial parts of it, in the
Porodiscida and Pylodiscida by the covering sieve-plates, in the
Spongodiscida by the spongy cortical substance of the shell. The growth of
the capsule corresponds to that of the including shell, gradually
increasing on the margin in the equatorial plane. Whilst in the greater
number of #Discoidea# its form continues circular, in many forms provided
with radial arms it enters into the arms and assumes their form. The
protoplasm of the capsule is commonly coloured by brown or red pigment, and
often contains many oil-globules. The nucleus is originally enclosed by the
medullary shell or the central chamber, and with increasing size enters
into the surrounding parts; in the #Cyclodiscaria# it often fills out the
internal concentric rings. The extracapsular jelly or the calymma is
commonly thick, and envelops the greater part or the whole body.

_Synopsis of the Families of the_ #Discoidea#.

-------------------------------------------------------------------------
I. Section #Phacodiscaria#.
Discoidea with external phacoid shell (or lenticular latticed cortical
shell).
-------------------------------------------------------------------------
Phacoid shell simple, without enclosed 1. CENODISCIDA.
medullary shell

Phacoid shell with { Margin without
simple or { chambered girdles, 2. PHACODISCIDA.
double {
enclosed { Margin surrounded by
medullary shell. { by chambered
{ girdles. 3. COCCODISCIDA.
-------------------------------------------------------------------------
II. Section #Cyclodiscaria#.
Discoidea without external phacoid shell (no lenticular latticed
cortical shell).
-------------------------------------------------------------------------
Surface of the { Concentric rings around
shell covered { the central chamber
by convex or { complete (without
even porous { open spaces), 4. PORODISCIDA.
sieve-plates {
(not spongy). { Concentric rings around
{ the central chamber
{ interrupted by three
{ open spaces, 5. PYLODISCIDA.

Surface of the shell spongy, not covered
by peculiar porous sieve-plates, 6. SPONGODISCIDA.

Family XVIII. #CENODISCIDA#, n. fam. (Pl. 31, fig. 11; Pl. 48, figs. 1-3).

_Definition._--#Discoidea# with simple extracapsular phacoid shell (or
lenticular latticed cortical shell), without medullary shell and without
chambered equatorial girdles.

The new family #Cenodiscida# opens the long series #Discoidea#, as their
most simple and primitive form. The circular lenticular central capsule is
enclosed by a {410}simple latticed shell of the same form, only separated
from it by a thinner or thicker jelly-veil. The lenticular or discoidal
fenestrated shell is therefore an extracapsular or "cortical shell,"
without an enclosed medullary shell.

The few genera of the Cenodiscida differ only in the shape of the
equatorial margin of the lenticular disk. In the first subfamily,
Zonodiscida, the margin is either quite simple (_Cenodiscus_) or surrounded
by a smooth, solid equatorial girdle (_Zonodiscus_). In the second
subfamily, Trochodiscida, the margin is armed with solid radial spines,
lying in the equatorial plane. According to the number and disposition of
these marginal spines, we distinguish _Stylodiscus_ (with two spines,
opposite in one equatorial axis), _Crucidiscus_ (with four spines, opposite
in pairs in two equatorial axes, perpendicular one to another),
_Theodiscus_ (with three marginal spines), and _Trochodiscus_ (with
numerous, commonly twenty to thirty, irregularly disposed spines). The
spines are constantly simple, not branched; sometimes more conical or
cylindrical, at other times more angular or pyramidal.

The two convex faces of the lenticular shell are constantly of similar
shape, commonly smooth, sometimes more or less thorny, or armed with
bristle-shaped radial spines. The pores are commonly more or less regular,
circular, and disposed in series, which are occasionally more radial, at
other times more concentric. If the wall of the hollow lens be rather
thick, the difference in the shape of the central and peripheral pores is
often striking. The central pores perforating the thick wall
perpendicularly are short cylindrical tubes; the marginal pores perforating
it in an oblique direction are longer conical tubes. The bars between the
central pores are often somewhat smaller.

_The Central Capsule_ of the Cenodiscida is in all cases a perfect,
circular, biconvex lens, the equatorial diameter of which is commonly
between two-thirds and three-fourths of the enclosing lattice-shell. The
interval between the two is filled up by the jelly-veil, or the hyaline
"calymma," which is perforated by the numerous pseudopodia that pass
through the shell-pores.

As the Cenodiscida possess the most simple shell-form of all #Discoidea#,
we may regard _Cenodiscus_ as the common ancestral form of this large
section, in the same manner as _Cenosphæra_ is the ancestral form of
#Sphæroidea#, _Cenellipsis_ of the Prunoidea, Cenolarcus of the Larcoidea.
But it is also possible that a part of Cenodiscida (or all?) arises from
the Phacodiscida by reduction and loss of the medullary shell. For in some
cases we find arising from the inside of the shell centripetal radial
beams, which end at a certain equal distance from the hollow centre (Pl.
31, fig. 11; Pl. 38, fig. 2). Cenodiscus itself can be derived either from
_Cenosphæra_ by compression of the spheroidal shell in one axis, or from
_Sethodiscus_ by loss of the intracapsular medullary shell, or from
_Actidiscus_ (the lenticular Actissa) by formation of a cortical shell
around the lenticular central capsule.

{411}_Synopsis of the Genera of the Cenodiscida._

I. Subfamily {Margin quite simple,
Zonodiscida. { without equatorial girdle, 175. _Cenodiscus_.
Margin of the disk {
without radial {Margin surrounded by a solid
spines. { siliceous equatorial girdle, 176. _Zonodiscus_.

II. Subfamily {Two spines opposite in one
Trochodiscida. { equatorial axis, 177. _Stylodiscus_.
Margin of the disk {
armed with radial {Three spines on the margin of
spines (lying in { the disk, 178. _Theodiscus_.
the equatorial {
plane). {Four spines opposite in pairs
{ in two perpendicularly
{ crossed equatorial axes, 179. _Crucidiscus_.
{
{Ten to twenty or more radial
{ spines (variable in number
{ and commonly irregular
{ in disposition), 180. _Trochodiscus_.

Subfamily 1. ZONODISCIDA, Haeckel.

_Definition._--#Cenodiscida# without radial spines on the margin of the
disk.

Genus 175. _Cenodiscus_,[214] n. gen.

_Definition._--#Cenodiscida# with simple margin of the circular disk,
without surrounding equatorial girdle and without radial spines.

The genus _Cenodiscus_ is the most simple and primitive form of all
#Discoidea#, and represents possibly the common ancestral form of this
order. The latticed shell is a simple biconvex lens, and encloses a smaller
central capsule of the same form, separated from it by the jelly-veil.
_Cenodiscus_ can be derived phylogenetically either from _Cenosphæra_ by
lenticular flattening of a simple latticed sphere, or directly from
_Actissa_ by formation of a lenticular fenestrated shell around the
lentiform central capsule. Possibly also some forms of _Cenodiscus_ can be
derived from _Sethodiscus_ by reduction and loss of the medullary shell.

1. _Cenodiscus phacoides_, n. sp. (Pl. 48, figs. 1, 1_a_).

Disk with smooth surface, without radial ribs or spines. Margin of the
lenticular biconvex disk thin, simple. Pores regular, circular; fifteen to
sixteen on the radius of the disk. (Very similar to _Sethodiscus
phacoides_, but without medullary shell.)

_Dimensions._--Diameter of the disk 0.2, of the pores 0.005.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{412}2. _Cenodiscus rotula_, n. sp.

Disk with smooth surface, without medial ribs or spines. Margin of the disk
blunt, very thick, rounded; both faces little convex. Pores regular,
circular; thirteen to fourteen on the radius of the disk. (Similar to
_Phacodiscus rotula_, Pl. 35, fig. 7, but without medullary shell.)

_Dimensions._--Diameter of the disk 0.16, of the pores 0.006.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

3. _Cenodiscus lenticula_, n. sp.

Disk with thorny surface, scattered with small, conical spines. Margin of
the lenticular biconvex disk thin. Pores irregular, roundish; ten to eleven
on the radius of the disk.

_Dimensions._--Diameter of the disk 0.15, of the pores 0.008.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Genus 176. _Zonodiscus_,[215] n. gen.

_Definition._--#Cenodiscida# with surrounding solid equatorial girdle on
the margin of the lenticular disk, without radial spines.

The genus _Zonodiscus_ differs from the preceding _Cenodiscus_ only in the
development of a solid siliceous girdle around the keen margin of the
lenticular disk. This form can also be derived from _Periphæna_ or
_Perizona_ by reduction and loss of the medullary shell. The same girdle
formation returns not only in both these Phacodiscida (Pl. 32, fig. 7; Pl.
33, fig. 4), but also in the Porodiscid _Perichlamydium_.

1. _Zonodiscus saturnalis_, n. sp.

Disk with smooth surface, without radial spines. Pores regular, circular,
fifteen to sixteen on the radius of the disk, in its distal half arranged
in fifty to sixty radial series, which are separated by prominent radial
crests or ribs. The crests are prolonged into the proximal half of the thin
solid equatorial girdle, which is one-third as broad as the radius of the
disk.

_Dimensions._--Diameter of the disk 0.25, of the pores 0.05.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

Subfamily 2. TROCHODISCIDA, Haeckel.

_Definition._--#Cenodiscida# with radial spines on the margin of the disk,
disposed in the equatorial plane.

Genus 177. _Stylodiscus_,[216] n. gen.

_Definition._--#Cenodiscida# with two radial spines on the margin of the
disk, opposite in one equatorial axis.

{413}The genus _Stylodiscus_ opens the series of the Trochodiscida or of
those Cenodiscida in which the thin margin of the hollow lenticular disk is
armed with a number of solid radial spines, situated in its equatorial
plane. _Stylodiscus_ is at the same time the most simple form of the
Stylodiscida, or of the numerous #Discoidea# (belonging to different
families) in which the disk bears only two spines, opposite in one
equatorial axis. Hitherto only two species of this genus have been
observed, but they seem to represent two different subgenera.

Subgenus 1. _Stylentodiscus_, Haeckel.

_Definition._--Internal cavity of the shell with centripetal axial rods.

1. _Stylodiscus endostylus_, n. sp. (Pl. 31, fig. 11).

_Sethostylus endostylus_, Haeckel, 1881, Prodromus et Atlas (pl. xxxi,
fig. 11).

Disk with smooth surface and dentated margin. Pores irregular, roundish;
ten to twelve on the radius of the disk. Marginal teeth conical, short,
irregular. Both opposite polar spines cylindrical, longer than the diameter
of the disk, and as broad as one larger pore. On the inside of the hollow
disk both spines are prolonged into two strong centripetal axial rods,
which do not reach the centre. Also a number of smaller centripetal axial
rods surrounds the central cavity, so that an original medullary shell
(_Sethostylus_) seems to have been lost (comp. above, p. 410).

_Dimensions._--Diameter of the disk 0.25, of the pores 0.005 to 0.02;
length of the polar spines 0.3 and more, thickness 0.012.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Subgenus 2. _Stylexodiscus_, Haeckel.

_Definition._--Internal cavity of the shell without axial rods.

2. _Stylodiscus amphistylus_, n. sp.

Disk with smooth surface and smooth margin. Pores regular, circular; eight
to nine on the radius of the disk. Both opposite polar spines pyramidal,
sulcated, about as long as the radius of the disk, and three times as long
as broad at the base. Inner cavity of the disk simple, without centripetal
rods.

_Dimensions._--Diameter of the disk 0.15, of the pores 0.007; length of the
polar spines 0.08, basal breadth 0.03.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

{414}Genus 178. _Theodiscus_,[217] n. gen.

_Definition._--#Cenodiscida# with three radial spines on the margin of the
disk, placed in its equatorial plane.

The genus _Theodiscus_ is the most simple form of those very numerous
#Discoidea# in which three rays are developed on the margin of the disk; a
symbol of the Christian dogma of the Trinity. Commonly the three angles
between the three spines are equal, more rarely one angle is larger than
the two others which are equal. The shell of some species is nearly
spherical (formerly separated by me as _Theosphæra_), whilst in others it
is a flattened biconvex lens.

Subgenus 1. _Theodiscoma_, Haeckel.

_Definition._--Angles between the three radial spines equal (triangle
equilateral).

1. _Theodiscus divinus_, n. sp.

Disk nearly spherical, with smooth surface. Pores regular, circular,
hexagonally framed; seven to eight on the radius. Three angles between the
spines equal. Spines prismatic, straight, twice as broad as one pore, eight
to eleven times as long as the shell diameter.

_Dimensions._--Diameter of the disk 0.1, of the pores 0.006; length of the
spines 0.8 to 1.1, breadth 0.01.

_Habitat._--Central Pacific, Station 265, surface.

2. _Theodiscus christianus_, n. sp.

Disk nearly spherical, with smooth surface. Pores regular, circular,
prolonged into short cylindrical, prominent tubuli, half as high as broad;
eight to nine on the radius. Three angles between the spines equal. Spines
prismatic, straight, twice as broad as one pore, twelve to fifteen times as
long as the shell diameter.

_Dimensions._--Diameter of the disk 0.08, of the pores 0.005; length of the
spines 1.1 to 1.2, breadth 0.01.

_Habitat._--Central Pacific, Station 271, surface.

3. _Theodiscus trinitatis_, n. sp.

Disk a rather flattened, biconvex lens, about twice as broad as thick.
Pores regular, circular, not prolonged into tubuli; five to six on the
radius. Three angles between the spines equal. Spines pyramidal, three
times as broad as one pore, about twice as long as the shell diameter.

{415}_Dimensions._--Diameter of the disk 0.06, of the pores 0.007; length
of the spine 0.12, basal breadth 0.02.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

Subgenus 2. _Theodiscura_, Haeckel.

_Definition._--Angles between the three radial spines unequal, two paired
angles equal, larger or smaller than the odd angle (triangle isosceles).

4. _Theodiscus vanitatis_, n. sp.

Disk nearly spherical, with smooth surface. Pores irregular, roundish;
eight to ten on the radius. Three angles between the spines unequal; one
odd angle larger than both others. Spines pyramidal, of unequal size; one
odd spine larger, both others smaller than the shell diameter.

_Dimensions._--Diameter of the disk 0.12, pores 0.004 to 0.006; length of
the odd spine 0.2, of the paired spines 0.1.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

5. _Theodiscus nirvana_, n. sp.

Disk a flat biconvex lens, about twice as broad as thick. Pores regular,
circular; ten to twelve on the radius. Three angles between the spines
unequal; one odd angle smaller than both others. Spines prismatic, very
long and thin, twice as broad as one pore, ten to twelve times as long as
the shell diameter.

_Dimensions._--Diameter of the disk 0.1, pores 0.005; length of the spines
1 to 1.2 or more, breadth 0.01.

_Habitat._--Indian Ocean, Ceylon, surface, Haeckel.

Genus 179. _Crucidiscus_,[218] n. gen.

_Definition._--#Cenodiscida# with four radial spines on the margin of the
disk, crossed in the equatorial plane.

The genus _Crucidiscus_ is the most simple form of the Staurodiscida, or of
the numerous #Discoidea# (belonging to different families) in which the
margin of the disk bears four radial spines, lying in the equatorial plane,
and crossed at right angles. Whilst commonly the internal shell-cavity of
_Crucidiscus_ is quite simple, in one case it bears four centripetal axial
rods, as inner prolongations of the outer radial cross-spines, perhaps
indications of a lost medullary shell (comp. p. 410).

{416}Subgenus 1. _Staurentodiscus_, Haeckel.

_Definition._--Internal cavity of the shell with centripetal axial rods.

1. _Crucidiscus endostaurus_, n. sp. (Pl. 48, fig. 2).

Disk with smooth surface and smooth simple margin. Pores regular, circular;
thirteen to fourteen on the radius of the disk. Four crossed spines
conical, strong, longer than the radius of the disk, on the inside
prolonged into four thinner centripetal axial rods, which do not reach the
centre. In the middle part of the disk also some other short axial rods
arise from the inside, not reaching the centre (as in _Stylodiscus
endostylus_, Pl. 31, fig. 11).

_Dimensions._--Diameter of the disk 0.16, of the pores 0.004; length of the
spines 0.2, breadth 0.014.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

Subgenus 2. _Staurexodiscus_, Haeckel.

_Definition._--Internal cavity of the shell without centripetal axial rods.

2. _Crucidiscus cuspidatus_, n. sp.

Disk with smooth surface and simple smooth margin. Pores regular, circular;
seven to eight on the radius of the disk. Four crossed spines pyramidal,
with prominent edges, somewhat shorter than the radius of the disk, twice
as broad at their thick base as one pore. No internal axial rods in the
shell cavity.

_Dimensions._--Diameter of the disk 0.12, of the pores 0.005; length of the
spines 0.05, basal breadth 0.025.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Crucidiscus echinatus_, n. sp.

Disk with thorny surface and thorny margin. Pores regular, circular, six to
seven on the radius. Four crossed spines pyramidal (four sided?), about as
long as the radius of the disk, as broad at their base as one pore. No
internal axial rods. Spines of the surface short, conical.

_Dimensions._--Diameter of the disk 0.14, of the pores 0.008; length of the
spines 0.08, basal breadth 0.02.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

4. _Crucidiscus cruciatus_, n. sp.

Disk with smooth surface and broad dentated margin. Pores regular,
circular; eleven to twelve on the radius. Four crossed spines pyramidal,
sulcated, about as long as the radius of the disk, as {417}broad at their
base as the radiated margin, which bears fifty to sixty triangular pointed
teeth of irregular size. (Similar to _Sethostaurus cruciatus_, Pl. 31, fig.
5, but without internal axial rods and medullary shell.)

_Dimensions._--Diameter of the disk 0.2, of the pores 0.007; length of the
spines 0.1, basal breadth 0.02.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

Genus 180. _Trochodiscus_,[219] n. gen.

_Definition._--#Cenodiscida# with numerous (ten to twenty or more) radial
spines on the margin of the disk, situated in its equatorial plane.

The genus _Trochodiscus_ bears on the thin margin of the latticed disk a
variable number of radial spines (commonly ten to twenty, sometimes fifty
to eighty or more). Their size and disposition are commonly more or less
irregular. For the most part all the marginal spines lie in the equatorial
plane; but sometimes part of them are crowded into two or four parallel
girdles. Several species of this genus are very similar to some species of
_Heliodiscus_, and only differ in the absence of the medullary shell,
perhaps in consequence of the phylogenetic loss of it; if this be the case,
the former are to be derived from the latter (compare p. 410).

Subgenus 1. _Trochodisculus_, Haeckel.

_Definition._--Bases of the marginal spines free, not connected by a solid
equatorial girdle.

1. _Trochodiscus cenophacus_, n. sp.

Disk with smooth surface. Pores regular, circular; eleven to twelve on the
radius. Marginal spines sixteen to twenty, conical, of irregular size and
disposition, about three times as long as broad, and as long as the radius
of the disk.

_Dimensions._--Diameter of the disk 0.15, of the pores 0.006; length of the
spines 0.06 to 0.09, basal breadth 0.02 to 0.03.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

2. _Trochodiscus medusinus_, n. sp.

Disk with smooth surface. Pores regular, circular; nine to ten on the
radius. Marginal spines ten to twelve, equilateral triangular, deeply
sulcated, about half as long and broad as the radius of the disk. (Very
similar to _Heliosestrum medusinum_, Pl. 34, fig. 6, but without enclosed
medullary shell. Compare p. 410.)

_Dimensions._--Diameter of the disk 0.2, of the pores 0.01; length of the
spines 0.05, basal breadth 0.05.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

{418}3. _Trochodiscus odontotrochus_, n. sp.

Disk with smooth surface, in the distal part radially sulcated. Pores
irregular, roundish; twelve to fourteen on the radius. Marginal spines very
short and numerous (thirty to forty), conical, scarcely one-fourth as long
as the radius of the disk, which resembles a wheel with marginal teeth.

_Dimensions._--Diameter of the disk 0.18, of the pores 0.004; length of the
spines 0.02, basal breadth 0.01.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

4. _Trochodiscus solaris_, n. sp.

Disk with smooth surface. Pores subregular, circular; eight to nine on the
radius. Marginal spines very numerous (sixty to eighty), in two to four
girdles densely crowded together, the largest as long as the diameter of
the disk, bent and conical. (Similar to _Heliodiscus solaster_, Pl. 34,
fig. 4, but without medullary shell.)

_Dimensions._--Diameter of the disk 0.15, of the pores 0.008; length of the
spines 0.05 to 0.15, basal breadth 0.01.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

5. _Trochodiscus echiniscus_, n. sp.

Disk with spiny surface. Pores irregular, roundish; ten to eleven on the
radius. Marginal spines conical, very numerous (forty to fifty), irregular
in size and disposition, the largest half as long as the radius of the
disk. (Similar to _Heliodiscus echiniscus_, Pl. 34, fig. 5, but without
medullary shell.)

_Dimensions._--Diameter of the disk 0.18, of the pores 0.005 to 0.015;
length of the spines 0.02 to 0.04, basal breadth 0.01.

_Habitat._--North Atlantic, Færöe Channel, John Murray.

Subgenus 2. _Pristodiscus_, Haeckel.

_Definition._--Bases of the marginal spines connected by a solid siliceous
equatorial girdle.

6. _Trochodiscus stellaris_, n. sp. (Pl. 48, fig. 3).

Disk with smooth surface. Pores subregular, circular; seven to eight on the
radius. Marginal spines twelve to sixteen, triangular, of subregular size
and disposition, about half as long as the radius of the disk, connected at
their broad base by a solid equatorial girdle of half the breadth; girdle
and spines radially striped.

_Dimensions._--Diameter of the disk 0.25, of the pores 0.015; length of the
spines 0.04 to 0.06, basal breadth 0.02 to 0.04.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

{419}7. _Trochodiscus cingillum_, n. sp.

Disk with smooth surface, in the distal part radially sulcated. Pores
regular, circular; twelve to thirteen on the radius. Marginal spines twenty
to twenty-four, triangular, of equal size and at regular distances, only
one-fourth as long as the radius of the disk, connected at their broad base
by a solid equatorial girdle of the double breadth. (Similar to
_Heliodiscus cingillum_, Pl. 33, fig. 7, but without medullary shell.)

_Dimensions._--Diameter of the disk 0.22, of the pores 0.004; length of the
spines 0.02, basal breadth 0.02

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Family XIX. #PHACODISCIDA#, Haeckel (Pls. 31-35).

_Phacodiscida_, Haeckel, 1881, Prodromus, p. 456.

_Definition._--#Discoidea# with simple extracapsular phacoid shell (or
lenticular latticed cortical shell), connected by radial beams with an
intracapsular, simple or double, concentric medullary shell, without
chambered equatorial girdles.

The family #Phacodiscida# comprises a large number of splendid forms (about
a hundred species), which agree with the preceding Cenodiscida in the
possession of the characteristic extracapsular "phacoid shell," but differ
from them in having one or two intracapsular "medullary shells"; these
concentric spherical medullary shells are connected with the lenticular
cortical shell or phacoid shell by means of radial beams perforating the
central capsule. The Phacodiscida bear therefore the same relation to the
Cenodiscida that the Disphærida and Trisphærida do to the Monosphærida.

Formerly several species belonging to the family were described by
Ehrenberg and Johannes Müller, but not distinguished from the Sphæroidea,
genus _Haliomma_ (_e.g._, _Haliomma sol_ et _Haliomma humboldtii_ of the
former, _Haliomma amphidiscus_ of the latter). For these oldest known
species I constituted in 1862 my genus _Heliodiscus_ (Monogr. d. Radiol.,
p. 436). Some other genera were afterwards (1875) figured by Ehrenberg as
_Periphæna_ and _Chilomma_. The rich material of the Challenger revealed
this family as very polymorphic and widely distributed, so that in my
Prodromus (1881, p. 457) I could enumerate eighteen different genera of
Phacodiscida. This number is here reduced to fifteen, uniting several of
them into one genus as "subgenera."

_The Medullary Shell_ of the Phacodiscida, or the intracapsular latticed
shell, is either simple and spherical, or double, composed of two
concentric spheres, which are united by a variable number of radial beams.
We could distinguish therefore as two subfamilies the Carpodiscida (with
simple medullary shell) and the Thecodiscida (with double concentric
medullary shell); the former corresponding to the Carposphærida (or
_Dyosphæria_), the latter to the Thecosphærida (or _Triosphæria_). But as
this difference seems not to be so important as the different shape of the
disk margin, we prefer this latter as a character {420}for the separation
of subfamilies. The form of the medullary shells is commonly quite
spherical (as also in the above mentioned #Sphæroidea#); sometimes it is a
little compressed in the same direction as the enclosing lenticular phacoid
shell. The diameter of the latter is commonly three to four times as large
as the diameter of the medullary shell; if this be double (in the
Thecodiscida) then the diameter of the outer medullary shell is commonly
three to four times as large as that of the inner. The radial beams
connecting the two seem to be very variable in number and disposition
(compare Pl. 31, fig. 8; Pl. 32, figs. 3, 4_a_, 7, 8_a_; Pl. 33, figs. 2,
3; Pl. 35, figs. 4, 8, 9, &c.).

_The Radial Beams_, which connect the medullary shell with the cortical or
"phacoid shell," and which pierce the membrane of the lenticular central
capsule, are commonly aggregated into two polar bunches around the
shortened axis of the disk (compare Pl. 31, fig. 8; Pl. 32, figs. 3, 8; Pl.
33, figs. 2, 3, &c.). Their number seems to be usually between ten and
thirty. More rarely piercing radial beams lie also in the equatorial plane,
and then commonly as inner prolongations of the outer marginal spines; so
we find two opposite in one axis, in _Heliostylus_ (Pl. 34, figs. 1, 2), or
four opposite in pairs in two crossed axes, in _Phacostaurus_ (Pl. 31,
figs. 1, 2, 7). Often the thickness of the beams increases from the centre
towards the periphery.

_The Phacoid Shell_, or the lenticular extracapsular cortical shell,
exhibits in the Phacodiscida quite the same general character as in the
Cenodiscida, above described (p. 410). Commonly the equatorial diameter of
the lens is twice to three times as large as the vertical diameter or the
shortened "main axis." The convexity of both opposite faces is either quite
even to the sharp margin (Pl. 31, fig. 2; Pl. 33, figs. 2, 5), or the
central part of the lens is more strongly vaulted, and often the margin is
thickened or truncated (Pl. 31, fig. 10; Pl. 35, fig. 7). The surface of
the lens is commonly smooth, but sometimes also covered with bristle-shaped
radial spines (Pl. 34, figs. 3, 5; Pl. 32, figs. 2, 3); rarely these spines
are prolonged and branched (Pl. 35, figs. 3, 5). The pores of the phacoid
shell are usually quite regular, circular, and regularly arranged, either
in more concentric or in more radial rows; the latter are sometimes
separated by radial crests arising towards the margin (Pl. 35, fig. 6). If
the wall of the phacoid shell be much thickened, the pores in its central
part are shorter and cylindrical, in its marginal part longer and conical
(Pl. 31, fig. 7; Pl. 32, fig. 1).

_The Margin of the Lens_ of the Phacodiscida is very polymorphic, and
serves mainly for the separation of genera. In the first subfamily,
Sethodiscida, it is either quite simple (Pl. 35, figs. 6-8) or surrounded
by a thin solid equatorial girdle, the basal part of which is often
radially striped (Pl. 32, figs. 7, 8). In the second subfamily,
Heliosestrida, we find on the margin a small number of radial spines in the
equatorial plane regularly disposed, either two opposite in one axis (Pl.
31, figs. 9-12) or four opposite in pairs in two crossed axes (Pl. 31,
figs. 1-8), or eight opposite in pairs in four axes, crossed at angles of
45° (Pl. 34, figs. 3, 6); in the latter case we can sometimes distinguish
{421}(as in many Medusæ) four larger perradial spines alternating with four
smaller interradial spines. Often in one and the same species occur
abnormalities in number and disposition of the radial spines, three or five
spines instead of four, or also seven or nine spines instead of eight;
often both halves of the disk become asymmetrical. If the number of the
marginal spines exceed eight to ten, they commonly become very variable in
size and irregular in disposition; these variations characterise the third
subfamily, Heliodiscida. Commonly also here all spines lie in the
equatorial plane; but sometimes they become crowded in several parallel
circles on both sides of the margin (Pl. 32, fig. 1). The form of the
marginal spines is commonly conical or flattened triangular, often also
pyramidal or deeply furrowed (Pl. 31, figs. 6-9). Very rarely the spines
are fenestrated (Pl. 35, fig. 1); only in one genus (_Heliodrymus_) they
are all or partly branched (Pl. 33, fig. 9; Pl. 35, figs. 3, 5).

The peculiar development of the phacoid shell has been already described by
J. Müller (compare my Monograph, 1862, pp. 156, 438).

_The Central Capsule_ of the Phacodiscida is everywhere circular,
lenticular, envelops the medullary shell, and is enclosed by the phacoid
shell, perforated by the radial beams, which connect the latter to the
former.

_Synopsis of the Genera of the Phacodiscida._

-------------------------------------------------------------------------
I. Subfamily Sethodiscida.
Margin of the disk without radial spines.
-------------------------------------------------------------------------
Margin simple, without {Medullary shell simple, 181. _Sethodiscus_.
equatorial girdle. {
{Medullary shell double, 182. _Phacodiscus_.

Margin surrounded by a {Medullary shell simple, 183. _Periphæna_.
hyaline equatorial girdle. {
{Medullary shell double, 184. _Perizona_.
-------------------------------------------------------------------------
II. Subfamily Heliosestrida.
Margin of the disk with two to eight solid radial spines, usually
quite regularly disposed. (Number usually constant.)
-------------------------------------------------------------------------
Two radial spines {Medullary shell simple, 185. _Sethostylus_.
(opposite in one axis). {
{Medullary shell double, 186. _Phacostylus_.

Three radial spines. Medullary shell simple, 187. _Triactiscus_.

Four radial spines (in {Medullary shell simple, 188. _Sethostaurus_.
cross form). {
{Medullary shell double, 189. _Phacostaurus_.

Six radial spines. Medullary shell simple, 190. _Distriactis_.

Eight radial spines. {Medullary shell simple, 191. _Heliosestrum_.
{
{Medullary shell double, 192. _Astrosestrum_.
-------------------------------------------------------------------------
III. Subfamily Heliodiscida.
Margin of the disk with numerous (ten to twenty or more) radial
spines, usually irregularly disposed. (Number variable.)
-------------------------------------------------------------------------
Radial spines all simple, {Medullary shell simple, 193. _Heliodiscus_.
not branched. {
{Medullary shell double, 194. _Astrophacus_.

Radial spines all or }Medullary shell simple, 195. _Heliodrymus_.
partly branched. }

{422}Subfamily 1. SETHODISCIDA, Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# without radial spines on the margin of the
disk.

Genus 181. _Sethodiscus_,[220] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with simple medullary shell and simple margin
of the circular disk, without surrounding equatorial girdle and without
radial spines.

The genus _Sethodiscus_ is the most simple and primitive form of all
Phacodiscida, and may be regarded as the common ancestral form of this
family. The simple spherical medullary shell is connected by a variable
number of radial beams with the lenticular or discoidal cortical shell (or
"phacoid shell"). The margin of this latter is quite simple, circular,
without solid equatorial girdle or radial spines. From the nearly allied
genus _Carposphæra_ of the #Sphæroidea#, its probable ancestral form,
_Sethodiscus_ can be derived simply by lenticular compression of the
spheroidal cortical shell.

Subgenus 1. _Sethodiscinus_, Haeckel.

_Definition._--Surface of the disk smooth, without radial ribs or spines.

1. _Sethodiscus phacoides_, n. sp.

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; fourteen to fifteen on the radius of the disk.
(Very similar to _Periphæna cincta_, Pl. 33, fig. 4, but without the girdle
of the margin.)

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.07, of
the pores 0.005.

_Habitat._--Pacific, central area, Stations 270 to 274, in various depths.

2. _Sethodiscus macroporus_, n. sp.

Disk with smooth surface, twice as broad as the medullary shell. Pores
regular, circular, very large; five to six on the radius of the disk.
(Remarkable for the extraordinary size of the pores, which reaches half the
radius of the medullary shell.)

_Dimensions._--Diameter of the disk 0.1, of the medullary shell 0.05, of
the pores 0.012.

_Habitat._--North Atlantic, Gulf Stream, Færöe Channel, John Murray.

3. _Sethodiscus microporus_, n. sp.

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular, very small; twenty-two to twenty-four on the radius of
the disk. (The small pores are scarcely half as broad as the thick bars
between them.)

{423}_Dimensions._--Diameter of the disk 0.25, of the medullary shell 0.06,
of the pores 0.002.

_Habitat._--South Pacific, Station 288, surface.

4. _Sethodiscus lenticula_, n. sp. (Pl. 33, figs. 1, 2).

Disk with smooth surface, four times as broad as the medullary shell. Pores
irregular, polygonal; eight to nine on the radius of the disk. (The pores
of the medullary shell, fig. 2, are also irregular, polygonal, or
roundish.)

_Dimensions._--Diameter of the disk 0.17, of the medullary shell 0.04, of
the pores 0.01.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

5. _Sethodiscus macrococcus_, n. sp. (Pl. 33, fig. 3).

Disk with smooth surface, two and a half times as broad as the medullary
shell. Pores irregular, polygonal; eleven to twelve on the radius of the
disk. (The pores of the medullary shell, fig. 3, are regular, circular,
with elevated hexagonal frames between them. The figured specimen is a
young one, both halves of the biconvex disk being not yet united in the
equatorial plane.)

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.065, of
the pores 0.01.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

6. _Sethodiscus micrococcus_, n. sp.

Disk with smooth surface, five times as broad as the medullary shell. Pores
irregular, roundish, very small; twenty-six to twenty-eight on the radius
of the disk. (The pores of the small medullary shell are regularly
circular, of the same size as those of the disk.)

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.04, of
the pores 0.002.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

Subgenus 2. _Sethodisculus_, Haeckel.

_Definition._--Surface of the disk with elevated radial ribs or spines.

7. _Sethodiscus radiatus_, Haeckel.

_Haliomma radians_, Ehrenberg, 1854, Mikrogeol., Taf. xix. fig. 50.

_Haliomma radiatum_, Ehrenberg, 1854, Mikrogeol., Taf. xxi. fig. 54.

Disk with radiated surface, four times as broad as the medullary shell.
Pores regularly circular; nine to ten on the radius of the disk, arranged
in radial series (about forty), which are separated by smooth crests, not
prominent on the smooth margin.

_Dimensions._--Diameter of the disk 0.12, of the medullary shell 0.03, of
the pores 0.003.

_Habitat._--Fossil in Tertiary deposits of the Mediterranean, Greece,
Sicily, Oran, &c.

{424}8. _Sethodiscus echinatus_, Haeckel.

_Haliomma echinatum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxvii. fig. 2.

Disk with radiated thorny surface, three times as broad as the medullary
shell. Pores regular, circular; eleven to twelve on the radius of the disk,
arranged in radial series (about sixty), which are separated by thorny
crests, prominent a little on the dentated margin.

_Dimensions._--Diameter of the disk 0.18, of the medullary shell 0.06, of
the pores 0.01.

_Habitat._--Fossil in the rocks of Barbados.

9. _Sethodiscus cristatus_, n. sp.

Disk with radiated surface, four times as broad as the medullary shell.
Pores regularly circular; sixteen to seventeen on the radius of the disk,
arranged in radial series (about eighty), which are separated in the distal
half by smooth elevated crests, strongly prominent on the dentated margin.
(Very similar to _Phacodiscus cristatus_, Pl. 35, fig. 6, but with simple
medullary shell.)

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05, of
the pores 0.005.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Genus 182. _Phacodiscus_,[221] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with double medullary shell and simple margin
of the circular disk, without surrounding equatorial girdle and without
radial spines.

The genus _Phacodiscus_ differs from the ancestral genus _Sethodiscus_ only
in the duplication of the medullary shell, and has therefore the same
relation to it that _Thecosphæra_ in the #Sphæroidea# exhibits to
_Carposphæra_. Both former discoidal genera differ from the two latter
spheroidal by the lenticular flattening of the cortical shell.

Subgenus 1. _Phacodiscinus_, Haeckel.

_Definition._--Surface of the disk smooth, without radial ribs or spines.

1. _Phacodiscus rotula_, n. sp. (Pl. 35, fig. 7).

Disk with smooth surface, four and a half times as broad as the outer and
fourteen times as broad as the inner medullary shell. Pores regularly
circular; sixteen to eighteen on the radius of the disk. Margin of the lens
very thick, truncated, nearly as broad as the outer medullary shell.

_Dimensions._--Diameter of the disk 0.21, of the outer medullary shell
0.045, of the inner 0.015; pores 0.008.

_Habitat._--North Pacific, Station 224, depth 1850 fathoms.

{425}2. _Phacodiscus lentiformis_, n. sp. (Pl. 35, fig. 8).

Disk with smooth surface, three and a half times as broad as the outer, and
ten times as broad as the inner medullary shell. Pores regularly circular;
twelve to thirteen on the radius of the disk. Margin of the disk sharp, as
in a biconvex lens.

_Dimensions._--Diameter of the disk 0.25, of the outer medullary shell
0.07, of the inner 0.025; pores 0.01.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms; fossil in
Barbados.

3. _Phacodiscus grandis_, n. sp.

Disk with thorny surface, five times as broad as the outer, and fifteen
times as broad as the inner medullary shell. Pores regularly circular;
twenty-two to twenty-four on the radius of the disk. Margin of the disk
thick, rounded.

_Dimensions._--Diameter of the disk 0.45, of the outer medullary shell
0.09, of the inner 0.03; pores 0.01.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Subgenus 2. _Phacodisculus_, Haeckel.

_Definition._--Surface of the disk with elevated radial ribs or spines.

4. _Phacodiscus echiniscus_, n. sp.

Disk with spiny surface, four times as broad as the outer, and ten times as
broad as the inner medullary shell. Pores irregularly roundish; fourteen to
fifteen on the radius of the disk. Margin of the disk sharp, as in a
biconvex lens.

_Dimensions._--Diameter of the disk 0.3, of the outer medullary shell 0.75,
of the inner 0.03; pores 0.007.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

5. _Phacodiscus clypeus_, n. sp. (Pl. 35, figs. 6, 9).

Disk with radiated surface, four times as broad as the outer, and twelve
times as broad as the inner medullary shell. Pores regularly circular;
sixteen to eighteen on the radius of the disk; in the distal half disposed
in sixty to seventy radial series, which are separated by prominent crests;
the sharp margin therefore a little jagged.

_Dimensions._--Diameter of the disk 0.2, of the outer medullary shell 0.05,
of the inner 0.017; pores 0.006.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

{426}Genus 183. _Periphæna_,[222] Ehrenberg, 1873 Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 246.

_Definition._--#Phacodiscida# with simple medullary shell, without radial
spines, but with a solid equatorial girdle around the margin of the
lenticular disk.

The genus _Periphæna_, founded by Ehrenberg in 1873 for the fossil
_Periphæna decora_ of Barbados, differs from its ancestral form
_Sethodiscus_ in the development of a very thin siliceous solid girdle
around the margin of the lenticular disk; this girdle lies in the
equatorial plane of the shell, and reappears in similar form in
_Perichlamydium_ among the Porodiscida, in _Spongophacus_ among the
Spongodiscida, and in _Zonodiscus_ among the Cenodiscida.

1. _Periphæna cincta_, n. sp. (Pl. 33, fig. 4).

Disk with smooth surface, four times as broad as the medullary shell. Pores
regularly circular; fourteen to sixteen on the radius of the disk. Girdle
of the margin about half as broad as the radius of the medullary shell, in
the distal half structureless, in the proximal half with seventy to eighty
short radial ribs.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05, of
the pores 0.005.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

2. _Periphæna decora_, Ehrenberg.

_Periphæna decora_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 80, Taf. xxviii. fig. 6.

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; twenty to twenty-two on the radius of the disk,
disposed in radial series. Girdle of the margin nearly as broad as the
radius of the medullary shell, in the distal half structureless, in the
proximal half with eighty to ninety short radial ribs.

_Dimensions._--Diameter of the disk 0.25, of the medullary shell 0.08, of
the pores 0.06.

_Habitat._--Fossil in Barbados.

3. _Periphæna statoblastus_, n. sp.

Disk with smooth surface, five times as broad as the medullary shell. Pores
regularly circular; twenty-four to twenty-six on the radius of the disk,
disposed in radial series, those in the distal half of the disk being
separated by piercing radial beams. Girdle of the margin twice as broad as
the radius of the medullary shell, in the whole breadth with one hundred
and twenty to one hundred and thirty piercing radial ribs, the
prolongations of the beams.

_Dimensions._--Diameter of the disk 0.3, of the medullary shell 0.06, of
the pores 0.04.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

{427}Genus 184. _Perizona_,[223] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with double medullary shell, without radial
spines, but with a solid equatorial girdle around the margin of the
lenticular disk.

The genus _Perizona_ differs from its ancestral form _Phacodiscus_ in the
development of a thin solid siliceous girdle, lying in the equatorial
plane around the margin of the disk. The two genera bear the same relation
to each other that _Periphæna_ bears to _Sethodiscus_. But in the two
latter genera the spherical medullary shell is simple, in the two former
double.

1. _Perizona scutella_, n. sp. (Pl. 32, fig. 7).

Disk with smooth surface, in the distal third radiated, four times as broad
as the outer, and ten times is broad as the inner medullary shell. Pores
subregular, circular; thirteen to fourteen on the radius of the disk, in
the marginal part separated by eighty to ninety prominent radial crests,
which are prolonged into the proximal half of the solid girdle. Breadth of
the girdle equal to that of the inner medullary shell.

_Dimensions._--Diameter of the disk 0.25, of the outer medullary shell
0.06, of the inner 0.025; pores 0.007.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

2. _Perizona pterygota_, n. sp. (Pl. 32, figs. 8, 8_a_).

Disk with smooth surface, six times as broad as the outer, and fifteen
times as broad as the inner medullary shell. Pores regularly circular;
eighteen to twenty on the radius of the disk. Margin much thickened and
truncated, abruptly separated in the equatorial circumference from the
peripheral sharp margin of the solid girdle; breadth of the girdle equal to
that of the outer medullary shell.

_Dimensions._--Diameter of the disk 0.25, of the outer medullary shell
0.04, of the inner 0.015; pores 0.006.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Subfamily 2. HELIOSESTRIDA, Haeckel.

_Definition._--#Phacodiscida# with a constant number of radial spines on
the margin of the disk (two, three, four, six, or eight), which are
commonly regularly disposed (sometimes more or less irregularly).

{428}Genus 185. _Sethostylus_,[224] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with simple medullary shell and with two
radial spines on the margin of the disk, opposite in the equatorial axis.

The genus _Sethostylus_ opens the series of the Heliosestrida or of those
Phacodiscida in which a constant number of radial spines (two to eight) is
more or less regularly disposed on the margin of the disk. All these
marginal spines lie in the equatorial plane of the lens, and have in the
same species a rather constant number and similar size, a certain form and
disposition. Sometimes they incline more or less to irregular variations.
In _Sethostylus_ there are only two spines, opposite in the equatorial
diameter of the lens; it corresponds to _Stylocyclia_ among the
Coccodiscida, and to _Xiphodictya_ among the Porodiscida.

Subgenus 1. _Sethostylium_, Haeckel.

_Definition._--Margin of the disk simple, smooth, without equatorial girdle
and without a corona of marginal spines.

1. _Sethostylus distyliscus_, n. sp. (Pl. 31, fig. 9).

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; nine to ten on the radius of the disk, in the
outer rows cylindrical. Margin of the disk simple, smooth, thin, without
peculiar equatorial girdle, radially striped by the prominent beams of the
peripheral series of pores. Both marginal spines of equal size, pyramidal,
sulcated, about as long as the radius of the disk and as broad at the base
as the medullary shell.

_Dimensions._--Diameter of the disk 0.13 to 0.16, of the medullary shell
0.04 to 0.05; length of both opposite spines 0.06 to 0.08, basal breadth
0.03 to 0.04.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

2. _Sethostylus dicylindrus_, n. sp. (Pl. 31, fig. 10).

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; sixteen to eighteen on the radius of the disk. Margin of
the disk simple, smooth, thick, without peculiar equatorial girdle. Both
marginal spines cylindrical, about as long as the diameter of the disk,
about twice as broad as a single pore. (Walls of the disk in the central
part twice to three times as thick as in the peripheral part.)

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of both spines 0.2 to 0.3, breadth 0.008.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

{429}Subgenus 2. _Heliostylus_, Haeckel, 1881, Prodromus, p. 457.

_Definition._--Margin of the disk with a solid equatorial girdle or a
corona of radial spines.

3. _Sethostylus dentatus_, n. sp. (Pl. 34, fig. 1).

_Heliostylus dentatus_, Haeckel, 1881, Prodromus et Atlas (pl. xxxiv.
fig. 1).

Disk with smooth surface, six times as broad as the medullary shell. Pores
regular, circular; eighteen to twenty on the radius of the disk. Margin of
the disk with a broad solid equatorial girdle, which bears fifty to sixty
strong conical teeth on the periphery; teeth about as long as the diameter
of the medullary shell. Both opposite marginal spines cylindro-conical,
about as long as the diameter of the disk, and as broad as the girdle.

_Dimensions._--Diameter of the disk 0.3, of the medullary shell 0.05;
length of both main spines 0.2 to 0.3, breadth 0.02.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

4. _Sethostylus serratus_, n. sp. (Pl. 34, fig. 2).

_Heliostylus serratus_, Haeckel, 1881, Prodromus et Atlas (pl. xxxiv.
fig. 2).

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; fourteen to sixteen on the radius of the disk. Margin
with a broad solid equatorial girdle, which bears forty to fifty strong
conical teeth on the periphery; teeth about as long as the radius of the
medullary shell. Both opposite marginal spines spindle-shaped, about as
long as the diameter of the disk, and as broad as the girdle.

_Dimensions._--Diameter of the disk 0.3, of the medullary shell 0.075;
length of both main spines 0.2 to 0.3, breadth 0.03.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

5. _Sethostylus hastatus_, n. sp.

Disk with smooth surface, two and a half times as broad as the medullary
shell. Pores irregular, roundish; ten to twelve on the radius of the disk.
Margin with a broad solid equatorial girdle, which bears on the periphery
six conical teeth, three on each half between the two main spines, which
are cylindrical, longer than the diameter of the disk, and half as broad as
the girdle. (This species can be derived from _Heliosestrum_, two opposite
spines of the eight marginal spines being much stronger developed than the
other six.)

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.06;
length of both main spines 0.2, breadth 0.01.

_Habitat._--North Atlantic, Station 354, surface.

{430}6. _Sethostylus spicatus_, n. sp.

Disk with spiny surface, four times as broad as the medullary shell. Pores
irregular roundish, with many unequal conical spines between them. Margin
with an equatorial girdle, composed of three to four concentric series of
conical radial spines, about as long as the radius of the medullary shell.
Both main spines very large, cylindrical, longer than the diameter of the
disk, and as broad as the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the main spines 0.3 or more, breadth 0.025.

_Habitat._--Fossil in the rocks of Barbados, Haeckel; living in the depths
of the Equatorial Atlantic, Station 348, depth (2450) fathoms.

Genus 186. _Phacostylus_,[225] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with double medullary shell and with two
radial spines on the margin of the disk, opposite in the equatorial axis.

The genus _Phacostylus_ differs from the foregoing _Sethostylus_ by the
duplication of the medullary shell, and bears the same relation to it as
_Phacodiscus_ does to _Sethodiscus_, or as _Amphicyclia_ in the
Coccodiscida does to _Stylocyclia_.

Subgenus 1. _Phacostylium_, Haeckel.

_Definition._--Margin of the disk simple, smooth, without equatorial
girdle, and without a corona of marginal spines.

1. _Phacostylus amphistylus_, n. sp. (Pl. 31, fig. 12).

Disk with smooth surface, four times as broad as the outer, and twelve
times as broad as the inner medullary shell. Pores regular, circular;
twelve to fourteen on the radius of the disk. Margin of the disk simple,
smooth, without spines and equatorial girdle. Both polar spines conical,
longer than the diameter of the disk, furrowed at the base, and twice as
broad as the inner medullary shell.

_Dimensions._--Diameter of the disk 0.16 to 0.18, of the outer medullary
shell 0.045, of the inner 0.015; length of the polar spines 0.2, basal
breadth 0.03.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Phacostylus amphixiphus_, n. sp.

Disk with smooth surface, three times as broad as the outer, and eight
times as broad as the inner medullary shell. Pores irregular, roundish; ten
to eleven on the radius. Margin of the disk {431}simple, smooth, without
spines and equatorial girdle. Both polar spines sword-shaped triangular,
two-edged, about as long as the radius of the disk, and as broad at the
base as the inner medullary shell.

_Dimensions._--Diameter of the disk 0.15, of the outer medullary shell
0.05, of the inner 0.02; length of the polar spines 0.08, basal breadth
0.02.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

Subgenus 2. _Astrostylus_, Haeckel.

_Definition._--Margin of the disk with a solid equatorial girdle or a
corona of radial spines.

3. _Phacostylus amphipyramis_, n. sp.

Disk with spiny surface, four and a half times as broad as the outer, and
fourteen times as broad as the inner medullary shell. Pores irregular,
roundish; eight to ten on the radius. Margin of the disk with a corona of
irregular, radial spines. Both opposite polar spines pyramidal, twice as
long as broad, and nearly as long as the radius of the disk.

_Dimensions._--Diameter of the disk 0.22, of the outer medullary shell
0.05, of the inner 0.016; length of the polar spines 0.1, basal breadth
0.05.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

4. _Phacostylus caudatus_, n. sp. (Pl. 32, fig. 6).

_Astrosestrum caudatum_, Haeckel, 1881, Prodromus et Atlas (pl. xxxii.
fig. 6).

Disk with smooth surface, two and a half times as broad as the outer, and
six times as broad as the inner medullary shell. Pores regular, circular;
six to seven on the radius of the disk. Margin with a solid equatorial
girdle, and irregularly bordered with eight to ten conical spines; two
opposite of these are much longer than the others. (This species can be
derived from _Astrosestrum_, two opposite marginal spines being much more
strongly developed than the six to eight others.)

_Dimensions._--Diameter of disk 0.12, of the outer medullary shell 0.05, of
the inner 0.02; length of the polar spines 0.1 to 0.25, basal breadth
0.025.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

5. _Phacostylus maximus_, n. sp.

Disk with smooth surface, five times as broad as the outer, and ten times
as broad as the inner medullary shell. Pores regular, circular; twenty to
twenty-two on the radius. Margin with a solid equatorial girdle, bearing on
the periphery one hundred to one hundred and twenty plain teeth, and
two very large polar spines, which are cylindrical, longer than the
diameter of the disk, and as broad at the furrowed base as the radius
of the outer medullary shell. (Similar to _Sethostylus dentatus_,
Pl. 34, fig. 1, but much larger, and with a double medullary shell.)
{432}_Dimensions._--Diameter of the disk 0.4, of the outer medullary shell
0.08, of the inner 0.04; length of the polar spines 0.5, basal breadth
0.04.

_Habitat._--Fossil in the rocks of Barbados.

Genus 187. _Triactiscus_,[226] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with simple medullary shell, and with three
radial spines on the margin of the disk, placed in the equatorial plane.

The genus _Triactiscus_ exhibits on the margin of the lenticular phacoid
shell three radial solid spines, the distance between which is sometimes
equal, at other times unequal. Either all the three spines are of equal
size and similar form, or one odd arm is smaller than the two paired arms.
The triradial form of this genus is repeated in _Tripodocyclia_ among the
Coccodiscida, and in _Tripodictya_ among the Porodiscida. The medullary
shell of _Triactiscus_ is simple.

1. _Triactiscus tripyramis_, n. sp. (Pl. 33, fig. 6).

Disk with smooth surface and smooth margin, three times as broad as the
medullary shell. Pores regular, circular; twelve to thirteen on the radius
of the disk. Three radial spines of nearly equal size and equidistant, or
one odd spine a little smaller, and the opposite angle (between the paired
spines) also smaller. Spines pyramidal, with broad prominent edges, one to
one and a half times as long as the radius of the disk, as broad at the
base as the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.15; of the medullary shell 0.05,
pores 0.006.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Triactiscus tricuspis_, n. sp. (Pl. 33, fig. 5).

Disk with smooth surface and smooth margin, five times as broad as the
medullary shell. Pores irregular, roundish; ten to eleven on the radius of
the disk. Three radial spines of unequal size and at unequal distances,
very short, conical, not longer than the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.03;
pores 0.004 to 0.008.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

3. _Triactiscus tripodiscus_, Haeckel.

_Haliomma triactis_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 236, Taf. xxviii. fig, 4.

Disk with thorny surface and spiny margin, three times as broad as the
medullary shell. Pores regular, circular; eight to nine on the radius of
the disk. Three radial spines of different {433}size and at unequal
distances, one odd spine as long as the radius, both paired spines as long
as the diameter of the disk; the odd angle between the latter is smaller.
Spines pyramidal, with broad edges.

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.05,
pores 0.008.

_Habitat._--Fossil in the rocks of Barbados.

Genus 188. _Sethostaurus_,[227] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with simple medullary shell and with four
radial spines on the margin of the disk, crossed in the equatorial plane.

The genus _Sethostaurus_ exhibits four marginal spines, which form commonly
a more or less regular cross in the equatorial plane. Sometimes the size
and disposition of the four spines become more or less different, and also
the angles between them vary; the regular rectangular cross passes over
into a bilateral or irregular form. The medullary shell is simple. The same
cross-form of the disk is seen in _Staurocyclia_ among the Coccodiscida,
and in _Staurodictya_ among the Porodiscida.

Subgenus 1. _Sethostaurium_, Haeckel.

_Definition._--Margin of the disk without a solid equatorial girdle or a
corona of spines.

1. _Sethostaurus orthostaurus_, n. sp. (Pl. 31, figs. 1, 2).

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; thirteen to fourteen on the radius of the disk.
Margin simple, smooth, without girdle. Four crossed spines equal,
pommel-shaped, angular, contracted at the base, scarcely as long as the
diameter of the medullary shell.

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.05;
length of the crossed spines 0.04, breadth 0.02.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Sethostaurus conostaurus_, n. sp. (Pl. 34, figs. 7, 8).

Disk with spiny surface, three times as broad as the medullary shell. Pores
irregular, roundish; seven to eight on the radius. Margin simple, smooth,
without girdle. Four crossed spines subregular, equal, conical, about as
long as the diameter and as broad at the base as the radius of the
medullary shell. (Fig. 7 exhibits the normal form, fig. 8 an abnormality
with five spines.)

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.06;
length of the crossed spines 0.05, basal breadth 0.03.

_Habitat._--South Atlantic, Station 333, surface.

{434}3. _Sethostaurus rhombostaurus_, n. sp. (Pl. 31, fig. 4).

Disk with smooth surface, two and a half times as broad as the medullary
shell. Pores regular, circular; ten to eleven on the radius. Margin simple,
smooth, without girdle. Two opposite spines longer than the diameter of the
disk, four times as long as the two others, which only equal its radius.
Spines sword-shaped, angular.

_Dimensions._--Diameter of the disk 0.14, of the medullary shell 0.06;
length of the major spines 0.2, of the minor 0.05, breadth 0.025.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

4. _Sethostaurus recurvatus_, n. sp. (Pl. 31, fig. 3).

Disk with smooth surface, three times as broad as the medullary shell.
Pores irregular, roundish; eight to nine on the radius. Margin simple,
smooth, without girdle. Four spines cylindrical, irregularly curved, of
different sizes; one single very large, much longer than the three others;
two opposite lateral spines recurved, hook-shaped.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.05;
length of the major spine 0.32, of the opposite spine 0.16, of both lateral
spines 0.08.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Subgenus 2. _Heliostaurus_, Haeckel 1881, Prodromus, p. 457.

_Definition._--Margin of the disk with a solid equatorial girdle or a
corona of spines.

5. _Sethostaurus cruciatus_, n. sp. (Pl. 31, fig. 5).

_Heliostaurus cruciatus_, Haeckel, 1881, Prodromus et Atlas (pl. xxxi.
fig. 5).

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; eleven to twelve on the radius. Margin with a
solid, radially striped girdle, which bears on the periphery forty to fifty
triangular pointed teeth of unequal length. Four crossed spines of equal
size and similar form, pyramidal, sulcated, about as long as the radius of
the disk, as broad at the base as the girdle.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.06;
length of the crossed spines 0.1, basal breadth 0.02.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

6. _Sethostaurus coronatus_, n. sp.

Disk with spiny surface, twice as broad as the medullary shell. Pores
irregular, roundish; nine to ten on the radius. Margin of the disk with a
solid broad girdle, bearing on the periphery a corona of thirty to forty
flat tongue-shaped teeth of different length. Four crossed spines
{435}prismatic with prominent edges, about as long as the diameter of the
disk, and as broad as the girdle.

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.08,
length of the crossed spines 0.18, breadth 0.02.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

7. _Sethostaurus gigas_, n. sp.

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; twenty to twenty-two on the radius. Margin of the disk
with a narrow solid girdle, bearing on the periphery one hundred to one
hundred and twenty slender flat pointed teeth. Four crossed spines equal,
conical, about as long as the radius of the disk, as broad at their
furrowed base as the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.4, of the medullary shell 0.1; length
of the crossed spines 0.5, basal breadth 0.05.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 189. _Phacostaurus_,[228] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with double medullary shell and with four
radial spines on the margin of the disk, crossed in the equatorial plane.

The genus _Phacostaurus_ differs from the foregoing _Sethostaurus_ by the
duplication of the medullary shell, and bears to it the same relation as
_Phacodiscus_ does to _Sethodiscus_. Also in this genus the cross of the
shell is commonly regular, rectangular, sometimes more or less irregular.

Subgenus 1. _Phacostaurium_, Haeckel.

_Definition._--Margin of the disk without a solid equatorial girdle or a
corona of spines.

1. _Phacostaurus oceanidum_, n. sp. (Pl. 31, fig. 6).

Disk with smooth surface, three times as broad as the outer and eight times
as broad as the inner medullary shell. Pores regular, circular; eleven to
twelve on the diameter of the disk. Margin smooth. Four crossed spines
pyramidal, deeply sulcate, about as long as the diameter of the outer
medullary shell, and two-thirds as broad at the base.

_Dimensions._--Diameter of the disk 0.15, of the outer medullary shell
0.05, of the inner 0.02; length of the spines 0.05, basal breadth 0.03.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

{436}Subgenus 2. _Astrostaurus_, Haeckel, 1881, Prodromus, p. 457.

_Definition._--Margin of the disk with a solid equatorial girdle, or a
corona of spines.

2. _Phacostaurus quadrigatus_, n. sp.

Disk with smooth surface, four times as broad as the outer and ten times as
broad as the inner medullary shell. Pores irregular, roundish; fifteen to
sixteen on the diameter of the disk. Margin with a broad solid equatorial
girdle, from which arise four crossed spines, conical, as long as the
radius of the disk.

_Dimensions._--Diameter of the disk 0.2, of the outer medullary shell 0.05,
of the inner 0.02; length of the spines 0.1, basal breadth 0.03.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

3. _Phacostaurus magnificus_, n. sp. (Pl. 31, figs. 7, 8).

_Astrostaurus magnificus_, Haeckel, 1881, Prodromus et Atlas (pl. xxxi.
figs.7, 8).

Disk with smooth surface, three times as broad as the outer and nine times
as broad as the inner medullary shell. Pores regular, circular; thirteen to
fourteen on the diameter of the disk. Margin with a corona of fifty to
sixty conical spines of unequal length. Four spines of the cross very large
pyramidal, deeply sulcated, longer than the radius of the disk, and as
broad at the base as the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.22, of the outer medullary shell
0.07, of the inner 0.025; length of the four spines 0.16, basal breadth
0.04.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

Genus 190. _Distriactis_,[229] n. gen.

_Definition._--#Phacodiscida# with simple medullary shell and with six
radial spines on the margin of the disk placed in the equatorial plane.

The genus _Distriactis_ exhibits on the margin of the lenticular phacoid
shell six radial solid spines. Their distance is commonly equal, rarely
unequal. Either all six spines are of equal size and similar form, or three
larger (perradial) alternating with three smaller (interradial), so that
_Distriactis_ appears to be derived from _Triactiscus_ by development of
three secondary between three primary spines.

1. _Distriactis liriantha_, n. sp.

Disk smooth, four times as broad as the medullary shell, with regular,
circular pores (about ten on the radius). Six marginal spines of equal size
and at equal distances, triangular, sulcate, half as long as the radius.
(Similar to _Heliosestrum medusinum_, Pl. 34, fig. 6.)

{437}_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the spines 0.05, basal breadth 0.04.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

2. _Distriactis corallantha_, n. sp.

Disk thorny, three times as broad as the medullary shell, with regular,
circular pores, about eight on the radius. Six marginal spines of equal
size and equidistant, conical, nearly as long as the diameter of the disk,
half as broad at the base as the medullary shell.

_Dimensions._--Diameter of the disk 0.12, of the medullary shell 0.04;
length of the spines 0.1, breadth 0.02.

_Habitat._--South Pacific, Station 300, surface.

3. _Distriactis alterna_, n. sp.

Disk smooth, five times as broad as the medullary shell, about twelve pores
on the radius. Six marginal spines pyramidal, at equal distances, but
unequal alternating size; three larger (perradial) spines as long as the
shell radius, three smaller between them (interradial) half as long and
broad.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.03;
length of the three major spines 0.08, of the three minor 0.05.

_Habitat._--North Atlantic, Station 353, surface.

4. _Distriactis palmantha_, n. sp.

Disk rough, four times as broad as the medullary shell, about ten pores on
the radius. Six marginal spines conical, at equal distances, but unequal
alternating size; three larger (perradial) spines as long as the shell
diameter, three smaller between them (interradial) as long as the radius.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the three major spines 0.2, of the three minor 0.1,

_Habitat._--South Atlantic, Station 325, surface.

5. _Distriactis amphithecta_, n. sp.

Disk thorny, three times as broad as the medullary shell. Six marginal
spines conical, of very different size; one single (anterior) much larger
than the five others, the opposite (posterior) much smaller. The other four
spines are paired, the anterior pair larger than the posterior.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.05;
length of the longest spine 0.2, of the shortest 0.04, of the anterior pair
0.1, of the posterior 0.07.

_Habitat._--Indian Ocean, Cocos Islands, surface, Rabbe.

{438}Genus 191. _Heliosestrum_,[230] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with simple medullary shell and with eight
radial spines on the margin of the disk (more or less regularly disposed,
sometimes seven or nine).

The genus _Heliosestrum_ (with simple medullary shell) and the following
_Astrosestrum_ (with double medullary shell) contain those Phacodiscida in
which the margin of the lenticular disk bears eight radial spines, commonly
more or less regularly disposed in the equatorial plane, so that the equal
angles between them amount to 45°. There are, however, many exceptions to
this regular eight-rayed form, either the angles between the eight spines
becoming unequal or the number of the spines amounting to seven or nine
(sometimes also six or ten) in one and the same species. But the regular
form is as prevalent, and also in the abnormal forms indicated by the
position of the spines, that we separate the genus _Heliosestrum_ from
_Heliodiscus_. Both genera are rich in common species, and in some of the
former four perradial (larger) spines alternate regularly with four
interradial (smaller) spines.

Subgenus 1. _Heliosestantha_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines free, not connected by an equatorial girdle.

1. _Heliosestrum medusinum_, n. sp. (Pl. 34, fig. 6).

_Heliodiscus medusinus_, Haeckel, 1881, Atlas (pl. xxxiv. fig. 6).

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; about ten on the radius of the disk. Eight marginal
spines regularly distributed (sometimes seven or nine, more or less
irregular), angular, nearly pyramidal (with equilateral triangular
outline), about as long and broad as the diameter of the medullary shell,
without connecting equatorial girdle. From the broad base of each spine run
eight to ten deep furrows convergent to its apex.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the radial spines 0.05, basal breadth 0.05.

_Habitat._--North Pacific, Station 266, surface.

2. _Heliosestrum octastrum_, n. sp.

Disk with smooth surface, three times as broad as the medullary shell.
Pores irregular, roundish; eight to nine on the radius. Eight marginal
spines regularly distributed (sometimes {439}seven or nine, more or less
irregular), conical, about as long as the diameter of the disk, without a
connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.05;
length of the radial spines 0.16, basal breadth 0.01.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

3. _Heliosestrum solarium_, Haeckel.

_Haliomma sol_ (_partim_), Ehrenberg, 1875 (_non_ 1844), Abhandl. d. k.
Akad. d. Wiss. Berlin, p. 74, Taf. xxviii. fig. 1.

Surface of the disk smooth; its diameter six times as large as that of the
medullary shell. Pores regular, circular; nine to ten on the radius. Eight
marginal spines regularly distributed (sometimes seven or nine, more or
less irregular), conical, about as long as the diameter of the medullary
shell, without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.18, of the medullary shell 0.03;
length of the radial spines 0.03, basal breadth 0.01.

_Habitat._--Fossil in the Tertiary rocks of Barbados.

4. _Heliosestrum liriope_, n. sp.

Surface of the disk smooth; its diameter four times as large as that of the
medullary shell. Pores subregular, circular; seven to eight on the radius.
Eight marginal spines regularly distributed, conical, compressed,
alternating longer and shorter; the longer equal to the diameter of the
disk, the shorter to the radius.

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.4;
length of the major radial spines 0.15, of the minor 0.09.

_Habitat._--North Pacific, Station 236, surface.

5. _Heliosestrum quadrigeminum_, n. sp.

Surface of the disk smooth; its diameter four times as large as that of the
medullary shell. Pores irregular, roundish; six to seven on the radius.
Eight marginal spines triangular, flat, regularly disposed, alternating
longer and shorter, the major as long as the radius of the shell, and half
as broad at the base, with three to four deep furrows on both sides,
without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.04;
length of the larger radial spines 0.08, of the smaller 0.04, basal breadth
0.04.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

6. _Heliosestrum contiguum_, Haeckel.

_Haliomma contiguum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxvii. fig. 5.

Disk with smooth surface, four times as broad as the medullary shell.
Pores irregular, roundish, touching, with very thin bars; nine to ten on
the radius. Eight marginal spines (often {440}seven or nine) conical, half
as long as the radius, twice as long as broad, without a connecting
equatorial girdle.

_Dimensions._--Diameter of the disk 0.17, of the medullary shell 0.04;
length of the radial spines 0.04, basal breadth 0.02.

_Habitat._--Fossil in Barbados.

7. _Heliosestrum irregulare_, n. sp.

Disk with smooth surface, four times as broad as the medullary shell. Pores
irregular, roundish; nine to eleven on the radius. Eight conical radial
spines of irregular size and distribution, often seven or nine, about as
long as the radius of the disk (in some cases longer, in others shorter).
No connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the radial spines 0.1, basal breadth 0.02.

_Habitat._--Central Pacific, Stations 270 to 272, depth 2425 to 2925
fathoms.

Subgenus 2. _Heliosestilla_, Haeckel.

_Definition._--Surface of the disk armed with radial spines. Bases of the
marginal spines free, without a connecting equatorial girdle.

8. _Heliosestrum octonum_, n. sp. (Pl. 34, fig. 3).

Disk with spiny surface, three times as broad as the medullary shell. Pores
circular, of very different sizes; eight to ten on the radius. Eight
conical marginal spines of equal size and equidistant, as long as the
radius of the disk, and one-third as broad at the base as the medullary
shell. Numerous spines on the surface bristle-shaped, on the margin half as
long as the eight main spines, in the central part shorter.

_Dimensions._--Diameter of the disk 0.18, of the medullary shell 0.06;
length of the marginal spines 0.1, basal breadth 0.02.

_Habitat._--Central Pacific, Station 271, surface.

9. _Heliosestrum ægineta_, n. sp.

Disk with spiny surface, four times as broad as the medullary shell. Pores
irregular, roundish, of different sizes; twelve to fourteen on the radius.
Eight pyramidal marginal spines regularly disposed, four larger (perradial)
alternating with four smaller (interradial); the former as long as the
radius of the disk, the latter half as long.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the marginal spines 0.05 to 0.1, basal breadth 0.03.

_Habitat._--South Atlantic, Station 330, surface.

{441}Subgenus 3. _Heliosestomma_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines connected by a solid equatorial girdle.

10. _Heliosestrum octangulum_, n. sp.

Disk with smooth surface, octagonal, twice as broad as the medullary shell.
Pores regular, circular; nine to ten on the radius. Equatorial girdle
narrow, radially striped, connecting the points of the eight short,
regularly disposed, marginal spines in such a manner that the whole shell
forms a regular octagon with rectilinear sides.

_Dimensions._--Diameter of the disk 0.11 to 0.12, of the medullary shell
0.05 to 0.06; length of the sides of the octagon 0.06.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

11. _Heliosestrum octogonium_, n. sp.

Disk with smooth surface, octagonal, four times as broad as the medullary
shell. Pores irregular, roundish; eight to nine on the radius. Equatorial
girdle broad, smooth, connecting the points of the eight short, triangular,
flat marginal spines in such a manner that the whole shell forms a
subregular octagon with concave, nearly equal sides. (Sometimes seven or
nine spines instead of eight are developed.)

_Dimensions._--Diameter of the disk 0.16 to 0.18, of the medullary shell
0.04 to 0.05; length of the radial spines 0.02 to 0.04, basal breadth 0.03.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

12. _Heliosestrum craspedotum_, n. sp.

_Haliomma humboldti_, var., Bury, 1862, Polycystins of Barbados, pl.
viii. fig. 4 (below).

Disk with smooth surface, three times as broad as the medullary shell.
Pores subregular, circular; ten to twelve on the radius. Eight marginal
spines (sometimes seven or nine) triangular, deeply sulcated, half as long
and one-third as broad as the radius of the disk, connected by a narrow,
radially striped equatorial girdle.

_Dimensions._--Diameter of the disk 0.18, of the medullary shell 0.06;
length of the radial spines 0.05, basal breadth 0.03.

_Habitat._--Central Pacific, Stations 271 to 274, at various depths, also
fossil in Barbados.

Genus 192. _Astrosestrum_,[231] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with double medullary shell and with eight
radial spines on the margin of the disk (more or less regularly disposed,
sometimes seven or nine).

{442}The genus _Astrosestrum_ differs from the foregoing _Heliosestrum_ by
the duplication of the medullary shell. The eight marginal spines in the
majority of individuals are regularly formed and disposed, of equal size
and equidistant. But there are frequent exceptions to this rule, either the
angles between the eight spines being more or less different, or the number
amounting to seven or nine, instead of eight. Here also in some species
four larger (perradial) spines alternate regularly with four smaller
(interradial spines), after the same law of symmetry, which is common in
the Medusæ.

Subgenus 1. _Astrosestantha_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines free, not connected by an equatorial girdle.

1. _Astrosestrum ephyra_, n. sp. (Pl. 32, figs. 4, 4_a_).

Disk with smooth surface, three times as broad as the outer and nine times
as broad as the inner medullary shell. Pores subregular, circular; seven to
eight on the radius of the disk. Eight marginal spines (sometimes seven or
nine) more or less irregularly disposed, of variable size, commonly as long
as the radius of the disk, twice as long as broad, pyramidal, sulcate,
without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.12, of outer medullary shell 0.04, of
the inner 0.014; length of the marginal spines 0.05 to 0.07, basal breadth
0.03.

_Habitat._--Central Pacific, Stations 270 to 274, depths 2350 to 2925
fathoms.

2. _Astrosestrum nauphanta_, n. sp. (Pl. 32, fig. 5).

Disk with smooth surface, two and a half times as broad as the outer and
five times as broad as the inner medullary shell. Pores regular, circular;
eight to nine on the radius of the disk. Eight marginal spines (often seven
or nine) more or less regularly disposed, of equal size, half as long as
the radius of the disk, and quite as broad at the base, compressed
triangular, sulcate, without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.15, of the outer medullary shell
0.06, of the inner 0.03; length of the marginal spines 0.04, basal breadth
0.04.

_Habitat._--Pacific, central area, Stations 266 to 268, depth 2700 to 2900
fathoms.

3. _Astrosestrum octacanthum_, Haeckel.

_Haliomma octacanthum_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 295, Taf. viii. fig. 11.

Disk with smooth surface, twice as broad as the outer and six times as
broad as the inner medullary shell. Pores regular, circular; six to seven
on the radius of the disk. Eight marginal spines {443}regularly disposed,
of variable size, the longest as long as the radius, pyramidal, not broader
at the base than one pore, without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.12, of the outer medullary shell
0.06, inner 0.02; length of the marginal spines 0.03 to 0.06, basal breadth
0.01.

_Habitat._--Western Pacific, Philippine Sea, 3300 fathoms, Ehrenberg;
Station 225, depth 4475 fathoms.

4. _Astrosestrum acraspedum_, n. sp.

Disk with smooth surface, three times as broad as the outer and six times
as broad as the inner medullary shell. Pores regular, circular; nine to ten
on the radius of the disk. Eight marginal spines regularly disposed, of
alternating size; four major (perradial) spines as long as the diameter of
the disk, four minor (interradial) half as long; spines cylindrical, not
broader at the base than one pore, without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.2, of the outer medullary shell 0.07,
of the inner 0.035; length of the larger spines 0.2, of the smaller 0.1,
basal breadth 0.01.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Subgenus 2. _Astrosestilla_, Haeckel.

_Definition._--Surface of the disk covered with radial spines. Bases of the
marginal spines free, not connected by an equatorial girdle.

5. _Astrosestrum acanthastrum_, n. sp.

Disk with spiny surface, three times as broad as the outer and nine times
as broad as the inner medullary shell. Pores subregular, circular; ten to
eleven on the radius. Eight marginal spines (sometimes seven or nine)
regularly disposed, triangular, about half as long as the radius of the
disk, and one-third as broad at the base, without a connecting equatorial
girdle.

_Dimensions._--Diameter of the disk 0.18, of the outer medullary shell
0.06, of the inner 0.02; length of the radial spines 0.05, basal breadth
0.03.

_Habitat._--North Atlantic, Gulf Stream, Færöe Channel, surface, John
Murray.

6. _Astrosestrum echinastrum_, n. sp.

Disk with spiny surface, twice as broad as the outer and four times as
broad as the inner medullary shell. Pores irregular, roundish; eight to
nine on the radius. Eight marginal spines more or less regularly disposed,
pyramidal, nearly as long as the radius of the disk, and not broader at the
base than one large pore, without a connecting equatorial girdle.

_Dimensions._--Diameter of the disk 0.15, of the outer medullary shell
0.07, of the inner 0.04; length of the radial spines 0.07, basal breadth
0.01.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Caltanisetta, Teuscher.

{444}Subgenus 3. _Astrosestomma_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines connected by a solid equatorial girdle.

7. _Astrosestrum pelagia_, n. sp.

Disk with smooth surface, three times as broad as the outer and eight times
as broad as the inner medullary shell. Pores regular, circular; eight to
nine on the radius. Eight marginal spines conical, about as long as the
radius of the disk, of nearly equal length, one-third as broad at the base,
connected by a radially striped equatorial girdle.

_Dimensions._--Diameter of the disk 0.18, of the outer medullary shell
0.06, of the inner 0.023; length of the radial spines 0.09, basal breadth
0.03.

_Habitat._--Indian Ocean, Ceylon, Haeckel, surface.

8. _Astrosestrum floscula_, n. sp.

Disk with smooth surface, two and a half times as broad as the outer and
five times as broad as the inner medullary shell. Pores irregular,
roundish; ten to eleven on the radius. Eight marginal spines triangular,
plain, scarcely half as long as the radius of the disk, nearly as broad at
the base, connected by an even equatorial girdle.

_Dimensions._--Diameter of the disk 0.2, of the outer medullary shell 0.08,
of the inner 0.04; length of the radial spines 0.09, basal breadth 0.07.

_Habitat._--South Atlantic, Station 325, surface.

Subfamily 3. HELIODISCIDA, Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with a variable number of radial spines on
the margin of the disk (ten to twenty or more), which are commonly more or
less irregular (sometimes regularly formed and disposed).

Genus 193. _Heliodiscus_,[232] Haeckel, 1862, Monogr. d. Radiol., p. 436.

_Definition._--#Phacodiscida# with simple medullary shell and with numerous
(ten to twenty or more) simple radial spines on the margin of the disk
(commonly with a variable number and an irregular disposition of the
undivided spines).

The genus _Heliodiscus_, the most common and polymorphic of all
#Phacodiscida#, was founded by me in 1862 as the first known type of this
family (_loc. cit._). I use {445}here the diagnosis of this genus in a
restricted sense, including only the species, in which the number of
marginal spines amounts to ten, twenty, or more (sometimes fifty to eighty,
rarely more than one hundred). The number of spines is in the different
species variable, and their disposition commonly more or less irregular,
whilst in the preceding genera (with two, three, four, six, or eight
marginal spines) their distance and form are commonly regular.

Subgenus 1. _Heliodiscetta_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines free, without a connecting equatorial girdle.

1. _Heliodiscus asteriscus_, n. sp. (Pl. 33, fig. 8).

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; ten to twelve on the radius of the disk. Marginal
spines fifteen to twenty, conical, often double contoured, of variable size
and disposition, the largest as long as the radius of the disk, as broad at
the base as one pore.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.05;
length of the radial spines 0.04 to 0.07, basal breadth 0.01.

_Habitat._--Cosmopolitan; Mediterranean (Corfu), Indian Ocean (Madagascar),
Atlantic and Pacific, many Stations, surface and various depths.

2. _Heliodiscus trochiscus_, n. sp. (Pl. 34, figs. 10, 13).

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; fourteen to sixteen on the radius. Marginal spines
twelve to sixteen, conical, of variable size and disposition, the largest
as long as the diameter of the medullary shell, twice as broad as one pore.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the radial spines 0.01 to 0.025, basal breadth 0.005.

_Habitat._--Central Pacific, Stations 272 to 274, surface.

3. _Heliodiscus trigonodon_, n. sp.

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; eighteen to twenty on the radius. Marginal spines
sixteen to twenty, equilateral triangular, flat, smooth, about as long and
broad as the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the radial spines 0.03, basal breadth 0.03.

_Habitat._--Central Pacific, Stations 266 to 274, depths 2350 to 2925
fathoms.

{446}4. _Heliodiscus glyphodon_, n. sp. (Pl. 35, fig. 2).

_Heliosestrum glyphodon_, Haeckel, 1881, Prodromus et Atlas (pl. xxxv.
fig. 2).

Disk with smooth surface, four times as broad as the medullary shell. Pores
subregular, circular; seven to eight on the radius. Marginal spines ten to
twelve, equilateral triangular, flat, deeply furrowed, twice as long as
broad, and as long as the radius of the disk.

_Dimensions._--Diameter of the disk 0.14, of the medullary shell 0.035;
length of the radial spines 0.07, basal breadth 0.035.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

5. _Heliodiscus helianthus_, Haeckel.

_Haliomma helianthus_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxvii. fig. 1.

Disk with smooth surface, five times as broad as the medullary shell. Pores
regular, oblong, disposed regularly in sixty to eighty radial series;
fourteen to sixteen on the radius. Marginal spines sixty to eighty,
conical, about as long as the diameter of the medullary shell, twice as
broad as one pore.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.04;
length of the radial spines 0.04, basal breadth 0.005.

_Habitat._--Fossil in Barbados.

6. _Heliodiscus sol_, Haeckel.

_Haliomma sol_, Ehrenberg, 1854, Mikrogeol., Taf. xix. fig. 52.

_Heliodiscus sol_, Haeckel, 1862, Monogr. d. Radiol., p. 438.

Disk with smooth surface, four times as broad as the medullary shell. Pores
irregular, roundish; eight to ten on the radius. Marginal spines twenty to
thirty, conical, the largest as long as the radius of the medullary shell,
their bases widely distant. (The species from Barbados figured by
Ehrenberg, 1875, as _Haliomma sol_, appertains to _Heliosestrum solare_.)

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the radial spines 0.02 to 0.025, basal breadth 0.01.

_Habitat._--Fossil in Tertiary rocks of Greece (Ægina), Ehrenberg.

7 _Heliodiscus siculus_, Stöhr.

_Heliodiscus siculus_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 89, Taf.
i. fig. 14.

Disk with smooth surface, three times as broad as the medullary shell.
Pores irregular roundish; seven to eight on the radius. Marginal spines
forty to fifty, conical, very irregular, the largest as long as the radius
of the medullary shell, their bases coming in contact.

_Dimensions._--Diameter of the disk 0.13 to 0.15, of the medullary shell
0.04 to 0.05; length of the radial spines 0.02 to 0.03, basal breadth 0.002
to 0.006.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte (Stöhr),
Caltanisetta (Teuscher).

{447}8. _Heliodiscus polymorphus_, n. sp. (Pl. 34, figs. 11, 12).

Disk with smooth surface, four times as broad as the medullary shell. Pores
irregular, roundish; ten to twelve on the radius. Marginal spines ten to
fifteen, pyramidal, angular, very irregular and variable in size and
distribution, the largest nearly as long as the diameter of the disk, as
broad at the base as the radius of the medullary shell.

_Dimensions._--Diameter of the disk 0.15 to 0.2, of the medullary shell
0.04 to 0.05; length of the radial spines 0.06 to 0.18, basal breadth 0.02
to 0.03.

_Habitat._--Central Pacific, Stations 265 to 268, depths 2700 to 2900
fathoms.

9. _Heliodiscus solaster_, n. sp. (Pl. 34, fig. 4).

Disk with smooth surface, three times as broad as the medullary shell.
Pores subregular, roundish; nine to ten on the radius. Marginal spines
fifty to sixty, cylindro-conical, flexuose, very variable in size, the
largest nearly as long as the diameter of the disk. The spines lie not only
in the equatorial plane (as usual), but also in two to four crowded girdles
on both sides of it.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.05;
length of the radial spines 0.05 to 0.15, basal breadth 0.008.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

Subgenus 2. _Heliodiscilla_, Haeckel.

_Definition._--Surface of the disk covered with radial spines. Bases of
the marginal spines free, without a connecting equatorial girdle.

10. _Heliodiscus phacodiscus_, Haeckel.

_Heliodiscus phacodiscus_, Haeckel, 1862, Monogr. d. Radiol., p. 437,
Taf. xvii. figs. 5-7.

_Haliomma phacodiscus_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 815.

Disk with spiny surface, three times as broad as the medullary shell. Pores
regular, circular; six to eight on the radius. Marginal spines twelve to
sixteen, conical, as long as the radius of the disk, and one-third as broad
as the diameter of the medullary shell.

_Dimensions._--Diameter of the disk 0.12 to 0.16, of the medullary shell
0.04 to 0.05; length of the marginal spines 0.06 to 0.08, basal breadth
0.01 to 0.02.

_Habitat._--Mediterranean (Messina), Canary Islands (Lanzerote).

11. _Heliodiscus amphidiscus_, Haeckel.

_Heliodiscus amphidiscus_, Haeckel, 1862, Monogr. d. Radiol., p. 437.

_Haliomma amphidiscus_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 154, Taf. ii. figs. 3-7.

Disk with spiny surface, three times as broad as the medullary shell.
Pores regular, circular; eight to ten on the radius. Marginal spines twelve
to fifteen, bristle-shaped, not larger than the surface spines, about as
long as the radius of the medullary shell.

{448}_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.05;
length of the marginal spines 0.03, basal breadth 0.002.

_Habitat._--Mediterranean (south shore of France), J. Müller.

12. _Heliodiscus echiniscus_, n. sp. (Pl. 34, fig. 5).

Disk with spiny surface, three times as broad as the medullary shell. Pores
irregular, roundish or polygonal; eight to ten on the radius. Marginal
spines thirty to fifty, pyramidal, angular, of very variable size, number,
and disposition, gradually passing into the surface-spines; the largest
nearly as long as the diameter of the medullary shell.

_Dimensions._--Diameter of the disk 0.14, of the medullary shell 0.045;
length of the radial spines 0.02 to 0.04, basal breadth 0.006.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

13. _Heliodiscus pertusus_, n. sp. (Pl. 35, fig. 1).

_Heliosestrum pertusum_, Haeckel, 1881, Prodromus et Atlas (pl. xxxv.
fig. 1).

Disk with spiny surface, three times as broad as the medullary shell. Pores
irregular, circular, hexagonally framed; eight to ten on the radius.
Marginal spines ten to thirty, very variable in size and disposition;
commonly eight to twelve larger spines, which are pyramidal, about as long
as the radius of the disk and perforated by two to four irregular, longish
pores. Between these fenestrated large spines are commonly ten to twenty
smaller conical spines, gradually passing into those of the surface.

_Dimensions._--Diameter of the disk 0.13 to 0.16, of the medullary shell
0.04 to 0.05; length of the marginal spines 0.04 to 0.08, basal breadth
0.01 to 0.015.

_Habitat._--North Pacific, Stations 241 to 244, depths 2300 to 2900
fathoms.

Subgenus 3. _Heliodiscomma_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines connected by a solid equatorial girdle.

14. _Heliodiscus cingillum_, n. sp. (Pl. 33, fig. 7).

Disk with smooth surface, five times as broad as the medullary shell. Pores
regular, circular; twelve to fourteen on the radius. Equatorial girdle
about as broad as the medullary shell, in the proximal half radially
striped, on the margin with twenty to twenty-four short, flat, triangular
spines, which are shorter than the breadth of the girdle.

_Dimensions._--Diameter of the disk 0.25, of the medullary shell 0.05;
length of the marginal spines 0.02, basal breadth 0.02.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

{449}15. _Heliodiscus humboldti_, Haeckel.

_Heliodiscus humboldti_, Haeckel, 1862, Monogr. d. Radiol., p. 438.

_Haliomma humboldti_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvi. fig. 27;
Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, Taf. xxvii. fig. 3.

_Haliomma humboldti_, Bury, 1862, Polycystins of Barbados, pl. viii. fig.
3 (at left).

Disk with smooth surface, four times as broad as the medullary shell. Pores
regular, circular; eighteen to twenty on the radius. Equatorial girdle half
as broad as the medullary shell, in the proximal half radially striped, on
the margin with sixteen to twenty short, flat, triangular teeth, which are
longer than the breadth of the girdle.

_Dimensions._--Diameter of the disk 0.12 to 0.2, of the medullary shell
0.03 to 0.05; length of the marginal spines 0.02 to 0.04, basal breadth
0.02.

_Habitat._--Fossil in Barbados (common and very variable).

16. _Heliodiscus marginatus_, n. sp. (Pl. 34, fig. 9).

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular; nine to ten on the radius. Equatorial girdle
one-fourth to one-half as broad as the medullary shell, in the whole
breadth radially striped, on the margin with twelve to eighteen very short
and broad, triangular, marginal spines.

_Dimensions._--Diameter of the disk 0.15 to 0.2, of the medullary shell
0.05 to 0.06; length of the marginal spines 0.01 to 0.02, basal breadth
0.02.

_Habitat._--Central Pacific, Stations 265 to 268, depth 2900 fathoms.

17. _Heliodiscus sulcatus_, n. sp.

Disk with smooth surface, four times as broad as the medullary shell. Pores
subregular, circular; twelve to fifteen on the radius. Equatorial girdle as
broad as the medullary shell, in the whole breadth radially striped on the
margin with ten to fifteen triangular, deeply sulcated teeth, about as long
and broad as the medullary shell. (Similar to _Heliodiscus glyphodon_, Pl.
35, fig. 2, but with broad sulcated girdle and shorter, more numerous
spines.)

_Dimensions._--Diameter of the disk 0.15 to 0.2, of the medullary shell
0.04 to 0.05; length of the radial spines 0.05 to 0.06, basal breadth 0.04
to 0.05.

_Habitat._--Atlantic and Pacific, tropical part, many Stations, surface.

18. _Heliodiscus umbonatus_, Haeckel.

_Haliomma umbonatum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 74, Taf. xxvii. fig. 4.

Disk with smooth surface, three times as broad as the medullary shell.
Pores irregular, roundish; eight to ten on the radius. Equatorial girdle
half as broad as the medullary shell, hyaline, not radially striped, on the
margin with ten to twenty triangular, irregular, smooth teeth, very
variable in size and disposition.

{450}_Dimensions._--Diameter of the disk 0.15 to 0.2, of the medullary
shell 0.05 to 0.06; length of the marginal spines 0.02 to 0.08, basal
breadth 0.01 to 0.03.

_Habitat._--Cosmopolitan; Atlantic, Pacific, in various depths; also fossil
in Barbados and Sicily.

Subgenus 4. _Heliodiscura_, Haeckel.

_Definition._--Surface of the disk covered with radial spines. Bases of the
marginal spines connected by a solid equatorial girdle.

19. _Heliodiscus apollinis_, n. sp.

Disk with spiny or bristly surface, three times as broad as the medullary
shell. Pores regular, circular; eleven to twelve on the radius. Equatorial
girdle narrow, on the margin with sixteen to twenty broad, flat, triangular
teeth, which are half as long and one-fourth as broad as the medullary
shell. (Very similar to _Astrophacus apollinis_, Pl. 32, fig. 2, but with
simple medullary shell.)

_Dimensions._--Diameter of the disk 0.18, of the medullary shell 0.06;
length of the marginal spines 0.03, basal breadth 0.015.

_Habitat._--Mediterranean (Corfu), Haeckel, surface.

20. _Heliodiscus zoroaster_, n. sp.

Disk with spiny surface, four times as broad as the medullary shell. Pores
subregular, circular; fourteen to sixteen on the radius. Equatorial girdle
broad, radially striped, on the margin with ten to twelve pyramidal, deeply
sulcated radial spines, which are nearly as long as the radius of the disk,
and one-fourth as broad at the base.

_Dimensions._--Diameter of the disk 0.24, of the medullary shell 0.06;
length of the marginal spines 0.11, basal breadth 0.03.

_Habitat._--Indian Ocean, between Aden and Ceylon, Haeckel, surface.

Genus 194. _Heliodrymus_,[233] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with simple medullary shell and with numerous
(ten to twenty or more) branched radial spines on the margin of the disk
(commonly with a variable number and an irregular disposition of the
ramified spines).

The genus _Heliodrymus_ differs from the nearly allied _Heliodiscus_ by the
ramification of the marginal spines, a character hitherto observed in no
other genus of Phacodiscida. The branching is more or less irregular,
either a simple bifurcation or a repeated fissure; the spines and their
branches are commonly more or less flexuose. {451}We can distinguish two
subgenera: in _Heliocladus_ the surface of the disk is smooth, in
_Heliodendrum_ covered with bristle-shaped radial spines, which are either
simple or also branched, sometimes longer than the thick marginal spines.

Subgenus 1. _Heliocladus_, Haeckel, 1881, Prodromus, p. 457.

_Definition._--Surface of the disk smooth, without radial spines.

1. _Heliodrymus dendrocyclus_, n. sp. (Pl. 33, fig. 9).

_Heliocladus dendrocyclus_, Haeckel, 1881, Prodromus et Atlas (pl.
xxxiii. fig. 9).

Disk with smooth surface, three times as broad as the medullary shell.
Pores regular, circular, hexagonally framed; eight to nine on the radius.
Marginal spines sixteen to twenty, cylindrical, very strong, flexuose,
irregularly branched, nearly as long as the diameter of the disk. Between
these main spines, each of which bears two to six irregular branches, are
scattered on the margin numerous smaller simple spines.

_Dimensions._--Diameter of the disk 0.16, of the medullary shell 0.05;
length of the main spines 0.1 to 0.14, breadth 0.01 to 0.02.

_Habitat._--Central Pacific, Station 271, surface.

2. _Heliodrymus furcatus_, n. sp.

Disk with smooth surface, four times as broad as the medullary shell. Pores
irregular, roundish; ten to twelve on the radius. Marginal spines twenty to
twenty-five, cylindrical, flexuose, forked, about as long as the radius of
the disk; fork-branches irregular, of unequal size. Some smaller simple
spines are scattered between the forked ones.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.04;
length of the radial spines 0.07 to 0.09, breadth 0.01.

_Habitat._--Central Pacific, Station 265, surface.

3. _Heliodrymus grottensis_, Haeckel.

_Heliodiscus grottensis_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 89,
Taf. i. fig. 13.

Disk with smooth surface, two and a half times as broad as the medullary
shell. Pores irregular, roundish; eight to nine on the radius. Marginal
spines twenty to thirty, conical, very irregular in form, size, and
disposition; the smaller simple, the larger irregularly branched and half
as long as the radius of the disk.

_Dimensions._--Diameter of the disk 0.17, of the medullary shell 0.07;
length of the marginal spines 0.02 to 0.04, basal breadth 0.01 to 0.02.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

{452}Subgenus 2. _Heliodendrum_, Haeckel.

_Definition._--Surface of the disk armed with simple or branched radial
spines.

4. _Heliodrymus setosus_, n. sp.

Disk with spiny surface, four times as broad as the medullary shell. Pores
regular, circular; twelve to thirteen on the radius. Marginal spines ten to
twelve, cylindrical, irregularly branched, each with two to eight flexuose
branches of different sizes; the largest as long as the diameter of the
disk. Spines of the surface bristle-shaped, half as long, not branched.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.05;
length of the marginal spines 0.12 to 0.18, breadth 0.025.

_Habitat._--North Pacific, Station 254, surface.

5. _Heliodrymus ramosus_, n. sp. (Pl. 35, figs. 3, 4).

Disk with spiny surface, three times as broad as the medullary shell. Pores
regular, circular, hexagonally framed; eleven to twelve on the radius.
Marginal spines sixteen to twenty, cylindrical, about as long as the
radius, irregularly forked or branched, with unequal flexuose branches.
Spines of the surface nearly as long, bristle-shaped, also irregularly
branched.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.05;
length of the spines 0.06 to 0.08, basal breadth 0.01 to 0.015.

_Habitat._--South Pacific, Station 288, surface.

6. _Heliodrymus viminalis_, n. sp. (Pl. 35, fig. 5).

Disk with spiny surface, two and a half times as broad as the medullary
shell. Pores irregular, roundish; ten to twelve on the radius. Marginal
spines fifteen to twenty, cylindro-conical, strong, partly simple, partly
forked, about as long as the diameter of the disk. Spines of the surface
very numerous, bristle-shaped, longer than the marginal spines, and more
branched.

_Dimensions._--Diameter of the disk 0.15, of the medullary shell 0.06;
length of the marginal spines 0.11 to 0.14, basal breadth 0.01 to 0.015;
length of the surface spines 0.2 to 0.03.

_Habitat._--Central Pacific, Station 271, surface.

Genus 195. _Astrophacus_,[234] Haeckel, 1881, Prodromus, p. 457.

_Definition._--#Phacodiscida# with double medullary shell and with numerous
(ten to twenty or more) simple radial spines on the margin of the disk
(commonly with a variable number and an irregular disposition of the
undivided spines).

The genus _Astrophacus_ differs from the similar _Heliodiscus_ in the
duplication of the medullary shell. The number and disposition of the
radial spines of the margin {453}(commonly between ten and twenty) is also
here variable in one and the same species. The greater number of observed
species of _Astrophacus_ resemble in a very remarkable manner the
corresponding species of _Heliodiscus_, and differ only in the double
medullary shell.

Subgenus 1. _Astrophacetta_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines free, without a connecting equatorial girdle.

1. _Astrophacus asteriscus_, n. sp.

Disk with smooth surface, three times as broad as the outer and eight times
as broad as the inner medullary shell. Pores regular, circular; twelve to
fourteen on the radius. Marginal spines fifteen to twenty, of variable size
and disposition; the largest as long as the radius of the disk, as broad at
the base as one pore. (Very similar to _Heliodiscus asteriscus_, Pl. 33,
fig. 8, but differing in the double medullary shell.)

_Dimensions._--Diameter of the disk 0.2, of the outer medullary shell 0.07,
of the inner 0.025; length of the marginal spines 0.05 to 0.1, basal
breadth 0.01.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

2. _Astrophacus trochiscus_, n. sp. (Pl. 34, fig. 14).

Disk with smooth surface, three times as broad as the outer and seven times
as broad as the inner medullary shell. Pores irregular, roundish; twelve to
thirteen on the radius. Marginal spines sixteen to twenty, conical, of
irregular variable size and disposition; the largest as long as the inner
medullary shell. (Differs from _Heliodiscus trochiscus_ in the double
medullary shell.)

_Dimensions._--Diameter of the disk 0.22, of the outer medullary shell
0.08, of the inner 0.03.

_Habitat._--North Atlantic, Station 354, surface.

3. _Astrophacus solaris_, n. sp. (Pl. 32, fig. 1).

Disk with smooth surface, three times as broad as the outer and seven times
as broad as the inner medullary shell. Pores subregular, roundish; twelve
to fourteen on the radius. Marginal spines one hundred to one hundred and
twenty, conical, flexuose, of irregular size and form; the largest
one-third as long as the diameter of the disk. The spines lie not only in
the equatorial plane (as is usual) but also in two to four crowded girdles
on both sides of it. (Very similar to _Heliodiscus solaster_, Pl. 34, fig.
4, but of double the size, with double the number of spines and with a
double medullary shell.)

_Dimensions._--Diameter of the disk 0.3, of the outer medullary shell 0.11,
of the inner 0.045; length of the spines 0.03 to 0.1, basal breadth 0.01 to
0.02.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe, surface.

{454}Subgenus 2. _Astrophacilla_, Haeckel.

_Definition._--Surface of the disk covered with radial spines. Bases of the
marginal spines free, without a connecting equatorial girdle.

4. _Astrophacus phacodiscus_, n. sp. (Pl. 32, fig. 3).

Disk with spiny surface, two and a half times as broad as the outer and
seven times as broad as the inner medullary shell. Pores subregular,
circular; ten to twelve on the radius. Marginal spines twelve to sixteen,
conical, stout, nearly as long as the radius of the disk, and as broad at
the base as the inner medullary shell. The numerous bristle-shaped spines
of the surface are scarcely half as long. (Similar to _Heliodiscus
phacodiscus_, Haeckel, Monogr. d. Radiol., Taf. xvii. figs. 5-7, but
differing in the double medullary shell.)

_Dimensions._--Diameter of the disk 0.18, of the outer medullary shell
0.07, of the inner 0.025; length of the marginal spines 0.08 to 0.09, basal
breadth 0.03.

_Habitat._--South Pacific, Station 300, surface.

Subgenus 3. _Astrophacomma_, Haeckel.

_Definition._--Surface of the disk smooth, without radial spines. Bases of
the marginal spines connected by a solid equatorial girdle. (Perhaps =
_Chilomma_(?) Ehrenberg, 1847, Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, p. 54.)

5. _Astrophacus cingillum_, n. sp.

_Haliomma humboldti_, var., Bury, 1862, Polycystins of Barbados, pl.
viii. fig. 3 (at right).

Disk with smooth surface, three times as broad as the outer and eight times
as broad as the inner medullary shell. Pores regular, circular; eleven to
thirteen on the radius. Equatorial girdle about as broad as the inner
medullary shell, in the proximal half radially striped, on the margin with
eighteen to twenty-four short, flat, triangular spines, about as long as
the breadth of the girdle. (Similar to _Heliodiscus cingillum_, but with
double medullary shell.)

_Dimensions._--Diameter of the disk 0.2, of the outer medullary shell 0.07,
of the inner 0.025; length of the spines 0.02, basal breadth 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms; also
fossil in Barbados.

6. _Astrophacus saturnus_, Haeckel.

? _Chilomma saturnus_, Ehrenberg, 1861, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 297; 1872, Abhandl. d. k. Akad. d. Wiss. Berlin, p. 286,
Taf. ii. fig. 5.

? _Chilomma saturnus_, Haeckel, 1862, Monogr. d. Radiol., p. 447.

Disk with smooth surface, two and a half times as broad as the outer, six
times as broad as the inner medullary shell. Pores large, irregular,
roundish; five to seven on the radius (?). Equatorial girdle very broad,
radially striped, nearly as broad as the outer medullary shell, perforated
by {455}twenty to thirty (or more?) radial spines. (The position of this
species, and the identity of _Chilomma_ with _Astrophacomma_, remains
doubtful, as the imperfect figure given by Ehrenberg of _Chilomma
saturnus_, the only species of the genus, is in contradiction with his
vague description, as is very often the case.)

_Dimensions._--Diameter of the disk 0.12 (with girdle 0.22) of the outer
medullary shell 0.05, of the inner 0.02.

_Habitat._--Arctic Ocean (Greenland, depth 1000 fathoms), Ehrenberg.

Subgenus 4. _Astrophacura_, Haeckel.

_Definition._--Surface of the disk covered with radial spines. Bases of the
marginal spines connected by a solid equatorial girdle.

7. _Astrophacus apollinis_, n. sp. (Pl. 32, fig. 2).

Disk with spiny surface, three times as broad as the outer, eight times as
broad as the inner medullary shell. Pores regular, circular; eleven to
twelve on the radius of the disk. Equatorial girdle narrow, smooth, on the
margin with twelve to sixteen broad, flat, triangular spines, of the same
length as the numerous bristle-shaped spines of the surface, which reach
half the radius of the disk. (Very similar to _Heliodiscus apollinis_, but
differing in the double medullary shell.)

_Dimensions._--Diameter of the disk 0.24, of the outer medullary shell
0.08, of the inner 0.03; length of the radial spines 0.06, basal breadth
0.03.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Family XX. #COCCODISCIDA#, Haeckel (Pls. 36-38).

_Coccodiscida_, Haeckel, 1862, Monogr. d. Radiol., p. 485.
_Coccodiscida_, Haeckel, 1881, Prodromus, p. 458.
_Lithocyclidina_, Ehrenberg, 1847, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 214 (_partim_).

_Definition._--#Discoidea# with extracapsular phacoid shell (or lenticular
latticed cortical shell), connected by radial beams with an intracapsular,
simple or double, concentric medullary shell, and surrounded by one or more
concentric chambered equatorial girdles on the margin.

The family #Coccodiscida# was founded by me in 1862 for those #Discoidea#
which agree with the Phacodiscida in the formation of the lenticular
"phacoid shell" (including a simple or double medullary shell), but differ
from them in the development of peculiar concentric chambered rings or
girdles around the equatorial margin of the disk, similar to those of the
Porodiscida.

The Coccodiscida represent a polymorphic family, in which we here
distinguish sixteen genera with fifty-seven species; it comprises the
greater part of those {456}#Discoidea# which Ehrenberg united in his group
Lithocyclidina (1875, represented by four genera and eight species);
several of these, however, appertain to quite different families, as his
_Astromma entomocora_, _Lithocyclia amphitrites_, &c. His knowledge of the
structure was very imperfect. The peculiar differentiation of the genera
and species exhibits the greatest analogy to that of the following family,
Porodiscida, though the structure of the central disk in both families is
quite different.

_The Phacoid Shell_, or the circular, lenticular cortical shell exhibits in
the Coccodiscida quite the same structure and composition as in the
Phacodiscida, described above (p. 420), so that there can be no doubt as to
the phylogenetic origin of the former from the latter. Quite in the same
way in both families, the lenticular "phacoid shell" is connected by
numerous, short, radial beams with the intracapsular, simple or double,
medullary shell; and also here these beams are commonly disposed in two
groups around the poles of the shortened main axis of the lens, so that
their distal ends are implanted in both circumpolar areas (Pl. 36, figs.
2-6; Pl. 37, figs. 3, 7; Pl. 38, figs. 2, 7). The medullary shell is
commonly simple, spherical, sometimes a little lenticularly compressed;
more rarely it is double, composed of two concentric lattice-shells, which
are connected by radial beams; in this case either both concentric
medullary shells are spherical, or the inner is spherical, and the outer
lenticular, very rarely the inner is lenticular also. In average size and
structure they agree perfectly with those of the Phacodiscida.

_The Chamber Girdles_ or "chambered rings" around the equatorial margin of
the disk, which constitute the only difference between the Coccodiscida and
the Phacodiscida, seem to exhibit a considerable degree of difference of
structure in the numerous species of this family; but I regret that I
cannot explain them here satisfactorily. The study of these structures is
extremely difficult because of the thickness and darkness of the massive
opaque shells; to get a perfect knowledge of them, it is indispensable to
compare slides made in different directions (horizontal slides through the
equatorial and parallel planes, vertical slides through radial and parallel
planes, oblique slides in different directions). But this requires a long
time and a most careful study of the slides, which are very difficult to
get in satisfactory condition. Therefore the following remarks can have
only a provisional value.

In all Coccodiscida we can distinguish on the equatorial chamber-girdle of
the lens-margin (even on superficial inspection) three different elements
of structure, viz.:--(A) concentric circular rings in the equatorial plane;
(B) numerous radial beams piercing the former and dividing them into
imperfect chambers; (C) porous plates or sieve-plates on both convex faces
of the disk. The probable morphological significance of these three
elements is the following:--Each ring or girdle corresponds to an outer
lenticular cortical shell, which is only developed on the marginal part,
whilst its central part is represented by the phacoid shell. Therefore the
radial beams (separating the imperfect chambers) are the same as in the
concentric Polysphærida, and the {457}sieve-plates of the surface are the
porous walls of the cortical shell itself. The correctness of this
explanation seems to be proved by such forms as figured in Pl. 38, figs. 2,
4, where the whole surface of the phacoid shell is covered by a concentric
chamber-work, as a central continuation of the marginal concentric rings.
If we imagine a system of perfect concentric lenticular phacoid shells,
compressed strongly from both poles of the shortened main axis, we get the
same figure.

Rarely one single girdle only is developed on the equatorial margin of the
lenticular disk (Pl. 37, figs. 2, 3, 5). Commonly the number of concentric
girdles amounts to three to six, often to ten to twelve or more. Some of
these largest Coccodiscida reach a considerable size. Commonly all girdles
are of the same breadth, which is about equal to the radius or to the
diameter of the inner medullary shell. Rarely the first (or innermost)
girdle differs by its greater breadth from the succeeding ones (Pl. 36,
fig. 8).

Only in few Coccodiscida the girdle-building remains restricted to the
equatorial planes, so that all the chambers lie in it. Commonly on both
sides of this plane become developed several layers, and often the number
of these (three to six or more) increases towards the periphery; in other
cases not their number, but their height increases. Therefore very often
the margin of the discoidal shell is much thickened, as thick as the centre
of the lenticular phacoid shell (or even more); whilst between the latter
and the former (on the proximal girdles) the disk is considerably thinner
(Pl. 36, figs. 2, 4; Pl. 37, figs. 7, 8; Pl. 38, figs. 2, 4). The
stratified layers communicate by large openings between their chambers. The
radial beams are commonly more or less regular and piercing, but also
frequently irregular and interrupted; often their number increases towards
the margin by intercalation of new beams.

_The Pores_ of the sieve-plates, which cover both sides of the chambered
disk, appear on the margin of the phacoid shell as direct continuations of
the pores of the latter, and sometimes they are so regularly disposed that
one single circular pore is situated on the surface of each chamber (Pl.
36, fig. 7; Pl. 37, fig. 1). But commonly the pores are of variable size
and number, two to three on each chamber, and often quite irregularly
scattered.

_The Margin_ of the chambered disk exhibits many differences, which afford
characters for the distinction of genera. In the first subfamily, the
Lithocyclida, the margin is quite simple without radial appendages. In the
second subfamily, the Stylocyclida, it is armed with solid radial spines
lying in the equatorial plane, and often regularly disposed in the same
manner as in the Phacodiscida (compare above, p. 421). In the third
subfamily, the Astracturida, the margin bears two or more (commonly three
or four) chambered arms, also situated in the plane of the disk, and of the
same structure as the circular chambered girdles (Pl. 38). In some cases
even the whole system of chamber-girdles is represented only by the radial
arms, which are inserted immediately on the margin of the phacoid shell. We
may regard therefore these formations as imperfect chambered disks, which
are developed only in the direction of certain rays {458}(perradii), and
reduced in the direction of the alternating rays (interradii). In some
Astracturida the chambered arms exhibit a structure different from the more
irregular chamber-work between them, so that we can distinguish the latter,
connecting the arms like a web-membrane, as a peculiar "patagium" (as in
many Porodiscida) (Pl. 38, figs. 8, 9). Often the distal ends of the
chambered arms are armed with a radial spine (Pl. 38, figs. 5, 6, 9).

_The Central Capsule_ of the Coccodiscida is originally always of the same
form and shape as in their ancestral group, the Phacodiscida; a circular
lenticular disk, which envelops the simple or double medullary shell and is
enclosed by the cortical phacoid shell. But whilst in the Phacodiscida the
phacoid shell envelops the central capsule perfectly, in the Coccodiscida
it envelops only the capsule from the two flat sides (by the upper and
lower sieve-plates); the marginal part of the lenticular capsule overgrows
the margin of the phacoid shell by peripheral extension, and fills out the
chambered cavity of the concentric girdles. In the Lithocyclida and the
Stylocyclida, where there are no chambered arms, the central capsule
remains a simple circular lens or disk; in the Astracturida, where
chambered radial arms surround the margin of the circular central disk, the
capsule enters also into these arms and fills out the greatest part of
their chambered cavities.

_Synopsis of the Genera of Coccodiscida_.

-------------------------------------------------------------------------
I. Subfamily Lithocyclida.
Margin of the disk simple, circular, without radial appendages.
-------------------------------------------------------------------------
Neither radial spines nor {Medullary shell simple, 196. _Lithocyclia_.
chambered arms on the {
circular margin. {Medullary shell double, 197. _Coccodiscus_.
-------------------------------------------------------------------------
II. Subfamily Stylocyclida.
Margin of the disk armed with solid radial spines.
-------------------------------------------------------------------------
Two opposite spines. {Medullary shell simple, 198. _Stylocyclia_.
{
{Medullary shell double, 199. _Amphicyclia_.

Three radial spines. Medullary shell simple, 200. _Trigonocyclia_.

Four crossed spines. Medullary shell simple, 201. _Staurocyclia_.

Five to ten or more {Medullary shell simple, 202. _Astrocyclia_.
radial spines. {
{Medullary shell double, 203. _Coccocyclia_.
-------------------------------------------------------------------------
III. Subfamily Astracturida.
Margin of the disk with two to five or more (commonly three or four)
hollow radial chambered arms (with or without a connecting
patagium). (Medullary shell commonly simple.)
-------------------------------------------------------------------------
Two arms, opposite in {Without patagium, 204. _Diplactura_.
one axis. {
{With patagium, 205. _Amphiactura_.

Three radial arms (at {Without patagium, 206. _Trigonactura_.
equal distances). {
{With patagium, 207. _Hymenactura_.

Four arms (in two {Without patagium, 208. _Astractura_.
crossed diameters). {
{With patagium, 209. _Stauractura_.

Five radial arms (at {Without patagium, 210. _Pentactura_.
variable distances). {
{With patagium, 211. _Echinactura_.

{459}Subfamily 1. LITHOCYCLIDA, Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with simple circular disk, without any radial
appendages of the margin (either solid radial spines or chambered arms).

Genus 196. _Lithocyclia_,[235] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Coccodiscida# with simple circular margin of the disk,
without radial appendages. Medullary shell simple.

The genus _Lithocyclia_ is the most simple form of Coccodiscida, and
represents the common ancestral form of this family, from which all other
genera of it can be derived. The lenticular, biconvex disk is quite simple,
composed of a variable number of concentric, circular, chambered rings,
which are pierced by radial beams, and which surround the circular
lenticular cortical shell or "phacoid shell." The latter contains a simple
spherical medullary shell in its centre, and is connected with it by radial
beams. The margin of the disk is circular, quite simple, without radial
spines or chambered arms.

1. _Lithocyclia cingulata_, n. sp.

Phacoid shell (or lenticular porous cortical shell) three times as broad as
the spherical enclosed medullary shell, surrounded by one single chambered
girdle or ring (with about forty chambers of equal size, separated by
radial beams). Margin of the disk circular, smooth. Pores of the convex
covering plates regular, circular; eight on the radius of the phacoid
shell, two on the breadth of the ring.

_Dimensions._--Diameter of the disk (with one ring) 0.13, of the phacoid
shell 0.1, of the medullary shell 0.033.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

2. _Lithocyclia lenticula_, n. sp. (Pl. 36, figs. 3, 4).

Phacoid shell two and a half times as broad as the enclosed medullary
shell, surrounded by three chambered girdles of equal size, which are
divided by piercing radial beams each into about fifty chambers. Margin of
the disk thorny. Pores irregular, roundish; seven on the radius of the
phacoid shell, two on the breadth of each girdle.

_Dimensions._--Diameter of the disk (with three girdles) 0.2, of the
phacoid shell 0.11, of the medullary shell 0.045.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

{460}3. _Lithocyclia ocellus_, Ehrenberg.

_Lithocyclia ocellus_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvi. fig. 30;
Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, Taf. xxix. fig. 3.

Phacoid shell three times as broad as the medullary shell, surrounded by
numerous (seven to eleven) chambered rings, which are divided by piercing
radial beams each into sixty to ninety chambers. Margin of the disk smooth.
Pores regular, circular; nine on the radius of the phacoid shell, one
single pore on each chamber.

_Dimensions._--Diameter of the disk (with eleven rings) 0.22, of the
phacoid shell 0.1, of the medullary shell 0.035.

_Habitat._--Fossil in the rocks of Barbados.

4. _Lithocyclia monococcus_, n. sp.

_Stephanopyxis dubiosa_ (?), Bury, 1862, Polycystins of Barbados, pl.
xiii. figs. 1, 2.

Phacoid shell four times as broad as the medullary shell, surrounded by
numerous (five to eight) chambered rings, which are divided by piercing
radial beams each into fifty to seventy chambers. Margin of the disk
thickened, thorny. Pores regular, circular; seven on the radius of the
phacoid shell, one single pore on each chamber.

_Dimensions._--Diameter of the disk (with eight rings) 0.18, of the phacoid
shell 0.12, of the medullary shell 0.03.

_Habitat._--Pacific, central area, Station 267, depth 2700 fathoms; also
fossil in the rocks of Barbados.

5. _Lithocyclia heteropora_, n. sp.

Phacoid shell two and a third times as broad as the medullary shell,
surrounded by five to nine chambered rings, which are divided by piercing
radial beams each into fifty to seventy chambers. Margin of the disk
smooth. Pores very different in the inner and outer part of the surface; in
the phacoid shell larger, regular, circular, eight on its radius, in the
chambered periphery very small and irregular, somewhat spongy.

_Dimensions._--Diameter of the disk (with nine rings) 0.2, of the phacoid
shell 0.13, of the medullary shell 0.055.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

Genus 197. _Coccodiscus_,[236] Haeckel, 1862, Monogr. d. Radiol., p. 485.

_Definition._--#Coccodiscida# with simple circular margin of the disk,
without radial appendages. Medullary shell double.

The genus _Coccodiscus_ has quite the same form and structure as the
preceding _Lithocyclia_, and differs from it only in the double medullary
shell, composed of two concentric latticed spheres; sometimes the inner
medullary shell is spherical, the outer {461}lenticular; the latter is
connected with the lenticular phacoid shell (or cortical shell) by radial
beams.

1. _Coccodiscus lamarckii_, n. sp. (Pl. 36, fig. 1).

Phacoid shell (or lenticular porous cortical shell) very thick walled (as
thick as one ring), three times as broad as the outer and seven times as
broad as the inner medullary shell, surrounded by two to three chambered
rings of equal breadth, each of which is divided by piercing radial beams
into thirty-six to forty square chambers. Margin of the disk circular,
smooth. Pores regular, circular, of equal size; ten on the radius of the
phacoid shell, two on the breadth of each chamber.

_Dimensions._--Diameter of the disk (with two rings) 0.2, of the phacoid
shell 0.14, outer medullary shell 0.05, inner 0.02.

_Habitat._--Western Tropical Pacific, Station 220, depth 1100 fathoms.

2. _Coccodiscus darwinii_, Haeckel.

_Coccodiscus darwinii_, Haeckel, 1862, Monogr. d. Radiol., p. 486, Taf.
xxviii. figs. 11, 12.

Phacoid shell three times as broad as the outer and nine times as broad as
the inner medullary shell, surrounded by five to eight chambered rings of
equal breadth, each of which is divided by forty piercing radial beams into
forty square chambers of equal size. Margin of the disk smooth, circular.
Pores irregular, roundish, of unequal size; eleven on the radius of the
phacoid shell, one to two on each chamber.

_Dimensions._--Diameter of the disk (with eight rings) 0.32, of the phacoid
shell 0.11, outer medullary shell 0.036, inner 0.012.

_Habitat._--Mediterranean, Messina, surface.

3. _Coccodiscus goethei_, n. sp. (Pl. 36, fig. 2).

Phacoid shell two and a half times as broad as the outer and five times as
broad as the inner medullary shell, surrounded by three to seven chambered
rings of equal breadth, each of which is divided by piercing radial beams
into sixty to eighty chambers. Margin of the disk thorny. Pores regular,
circular; eight on the radius of the phacoid shell, a single one on the
breadth of each chamber.

_Dimensions._--Diameter of the disk (with seven rings) 0.25, of the phacoid
shell 0.1, of the medullary shell 0.04, inner 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Subfamily 2. STYLOCYCLIDA, Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with solid radial spines on the margin of the
circular disk, situated in its equatorial plane (without chambered arms).

{462}Genus 198. _Stylocyclia_,[237] Ehrenberg, 1847, Monatsber. d. k.
preuss. Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Coccodiscida# with two opposite solid radial spines on the
margin of the circular disk. Medullary shell simple.

The genus _Stylocyclia_ opens the series of the Stylocyclida or of those
Coccodiscida in which the margin of the chambered disk is armed with solid
radial spines, situated in its equatorial plane, but without chambered
arms. _Stylocyclia_ is the most simple form of this subfamily, and bears
only two marginal spines, opposite in one equatorial axis of the disk. The
medullary shell is simple. This genus corresponds to _Xiphodictya_ in the
family Porodiscida. The genus was previously known only by one single
species described by Ehrenberg.

1. _Stylocyclia dimidiata_, Ehrenberg.

_Stylocyclia dimidiata_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxix. fig. 4.

Phacoid shell three times as broad as the medullary shell, surrounded by
five to eight chambered rings of equal breadth, divided by eighty to ninety
piercing radial beams into square chambers. Pores subregular, circular; six
to seven on the radius of the phacoid shell, a single one on each chamber.
Both opposite marginal spines strong, club-shaped, their thickness
decreasing from the margin towards the centre of the disk. (The figure of
Ehrenberg is very incomplete.)

_Dimensions._--Diameter of the disk (with eight rings) 0.25, of the phacoid
shell 0.11, of the medullary shell 0.035.

_Habitat._--Fossil in the Tertiary rocks of Barbados; living in the depths
of the Equatorial Atlantic, Station 348, depth (2450) fathoms.

2. _Stylocyclia prionacantha_, n. sp. (Pl. 37, fig. 6).

Phacoid shell two and a half times as broad as the medullary shell,
surrounded by five to six chambered rings, which are divided by fifty to
sixty piercing radial beams into square chambers. Pores in the thick-walled
phacoid shell regularly circular, increasing in size from the centre; eight
to nine on its radius. Pores on the surface of the chambered girdle
smaller, very irregular, two to three on each ring. Both marginal spines
longer than the diameter of the disk, with broad serrated edges.

_Dimensions._--Diameter of the disk (with five rings) 0.23, of the phacoid
shell 0.12, of the medullary 0.05.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

{463}3. _Stylocyclia excavata_, n. sp. (Pl. 37, fig. 8).

Phacoid shell four times as broad as the medullary shell, surrounded by
four chambered rings, which are divided by twenty to thirty radial beams
into broad chambers. The height of the rings increases strongly from the
centre, so that the fourth ring is two and a half times as high as the
first. Pores irregular, roundish; five to six on the radius of the phacoid
shell, one to two on the breadth of each ring. Margin of the disk smooth.
Both marginal spines thin and long, cylindrical, arising from the medullary
shell, longer than the diameter of the disk.

_Dimensions._--Diameter of the disk (with four rings) 0.25, of the phacoid
shell 0.1, of the medullary shell 0.025.

_Habitat._--Fossil in the rocks of Barbados. Indian Ocean, Cocos Islands,
Rabbe.

4. _Stylocyclia amphacantha_, n. sp.

Phacoid shell twice as broad as the medullary shell, surrounded by five
chambered rings, divided by piercing beams each into forty to fifty
chambers. Pores regular, circular; eight to nine on the radius of the
phacoid shell, one on each chamber. Margin of the disk thorny. Both
marginal spines strong, conical, as long as its radius.

_Dimensions._--Diameter of the disk (with five rings) 0.24, of the phacoid
shell 0.1, of the medullary shell 0.05.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Genus 199. _Amphicyclia_,[238] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with two opposite solid radial spines on the
margin of the circular disk. Medullary shell double.

The genus _Amphicyclia_ has the same form and structure as _Stylocyclia_,
and differs from it only in the double concentric medullary shell. It bears
therefore to the latter the same relation that _Coccodiscus_ does to
_Lithocyclia_.

1. _Amphicyclia chronometra_, n. sp. (Pl. 38, fig. 1).

Phacoid shell three times as broad as the outer and nine times as broad as
the inner medullary shell, surrounded by two broad chambered rings, which
are divided into irregular chambers by fifty to sixty radial beams of
different distance. Margin of the disk thorny, lacerated. Pores irregular,
roundish; twelve to sixteen on the radius of the phacoid shell, two to
three on the breadth of each ring. The two opposite marginal spines strong,
prismatic, with prominent edges, about as long as the diameter of the disk.

_Dimensions._--Diameter of the disk (with two rings) 0.24, of the phacoid
shell 0.15, outer medullary shell 0.05, inner 0.017.

_Habitat._--Pacific, central area, Stations 265 to 268, depths 2700 to 2900
fathoms.

{464}2. _Amphicyclia amphistyla_, n. sp. (Pl. 37, fig. 7).

_Stylocyclia amphistyla_, Haeckel, 1879, MS. et Atlas (pl. xxxvii. fig.
7).

Phacoid shell thin walled, two and a half times as broad as the outer and
seven times as broad as the inner medullary shell, divided by eighty to
ninety radial beams into irregular chambers, which are stratified in four
to five floors. Pores regular, circular; eight to nine on the radius of
the phacoid shell, two on the breadth of each chamber. Both marginal
spines cylindrical.

_Dimensions._--Diameter of the disk (with seven rings) 0.27, of the phacoid
shell 0.1, outer medullary shell 0.04, inner 0.014.

_Habitat._--North Atlantic, Canary Islands, Station 354, surface.

3. _Amphicyclia pachydiscus_, n. sp. (Pl. 38, fig. 2).

Phacoid shell very thick, spongy, twice as broad as the outer and five
times as broad as the inner medullary shell, divided by fifty to seventy
radial beams into subregular chambers, which are stratified in five to six
floors. Pores irregular, roundish; ten to twelve on the radius of the
phacoid shell, two to three on the breadth of each chamber. The two
opposite marginal spines quadrangular prismatic, very long, as broad as the
radius of the inner medullary shell. Fig. 2 exhibits a vertical section
(slide) through the centre.

_Dimensions._--Diameter of the disk (with five rings) 0.22, of the phacoid
shell 0.1, outer medullary shell 0.05, inner 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms; fossil in
Barbados.

Genus 200. _Trigonocyclia_,[239] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with three solid radial spines on the margin
of the circular disk. Medullary shell simple.

The genus _Trigonocyclia_ (only known by one single species, once observed)
bears on the margin of the chambered disk three solid radial spines, at
equal distances one from another. It corresponds to the genus
_Tripodictya_ amongst the Porodiscida.

1. _Trigonocyclia triangularis_, n. sp. (Pl. 37, fig. 5).

Phacoid shell three times as broad as the medullary shell, connected with
it by six equidistant radial beams, three of which are prolonged into
marginal spines. In the equatorial plane only one single chambered ring,
divided by forty-four radial beams into narrow chambers. Pores large,
irregular, roundish; five on the radius of the phacoid shell, two on each
chamber. Surface and margin of the disk spiny. Three strong conical
marginal spines, divergent at equal angles, as long as the radius of the
disk.

_Dimensions._--Diameter of the disk 0.16, of the phacoid shell 0.12, of the
medullary shell 0.04.

_Habitat._--Indian Ocean, Zanzibar, depth 2200 fathoms, Pullen.

{465}Genus 201. _Staurocyclia_,[240] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with four solid radial spines on the margin
of the disk, crossed in two equatorial diameters perpendicular one to
another. Medullary shell simple.

The genus _Staurocyclia_ is characterised by four radial spines on the
margin of the disk, which are opposite in pairs and situated in two
equatorial diameters, commonly perpendicular one to another. They form
therefore a rectangular cross (rarely more or less irregular). The genus
corresponds exactly to _Staurodictya_ in the family Porodiscida. The
medullary shell is simple.

1. _Staurocyclia cruciata_, n. sp. (Pl. 37, fig. 1).

Phacoid shell two and a half times as broad as the medullary shell,
surrounded by six to eight regular rings, which are divided by fifty to
seventy piercing beams into square chambers. Pores regular, circular; six
on the radius of the phacoid shell, one single pore on each chamber. Four
crossed radial spines, arising from the medullary shell by thin bases,
prolonged over the dentated margin as four strong quadrangular swords, as
long as the radius of the disk.

_Dimensions._--Diameter of the disk (with eight rings) 0.23, of the phacoid
shell 0.09, of the medullary shell 0.035.

_Habitat_--Pacific, central area, Station 267, depth 2700 fathoms.

2. _Staurocyclia serrata_, n. sp.

? _Haliomma_ sp., Bury, 1862, Polycystins of Barbados, pl. xxii. fig. 2.

Phacoid shell twice as broad as the medullary shell, surrounded by three to
four spongy rings, which are divided by thirty to forty radial beams into
irregular chambers. Pores in the phacoid shell regular, circular, five on
its radius; in the chambered spongy girdle much smaller and irregular.
Margin dentated, with four crossed, very large, quadrangular spines, as
long as the radius of the disk, with wing-shaped, serrated edges.

_Dimensions._--Diameter of the disk (with four rings) 0.2, of the phacoid
shell 0.1, of the medullary shell 0.05.

_Habitat._--Fossil in the rocks of Barbados.

3. _Staurocyclia phacostaurus_, n. sp. (Pl. 37, figs. 2, 3).

_Phacostaurus pyramidalis_, Haeckel, 1879, MS. (as a separate genus).

Phacoid shell two and a quarter times as broad as the medullary shell,
connected with it by twelve radial beams (four equatorial and eight
divergent) and surrounded by one single ring, which is divided by forty
radial beams into regular chambers. Pores regular, honeycomb-like in the
{466}phacoid shell, seven on its radius; smaller on the equatorial ring,
three on its breadth. Four marginal spines pyramidal four-sided, as long
and as broad at the base as the ring.

_Dimensions._--Diameter of the disk (with one ring) 0.14, of the phacoid
shell 0.11, of the medullary shell 0.048.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

4. _Staurocyclia magniducis_, n. sp. (Pl. 37, fig. 4).

_Coccostaurus magniducis_, Haeckel, 1881, MS. et Atlas (pl. xxxvii. fig.
4).

Phacoid shell twice as broad as the medullary shell, connected with it by
numerous radial beams and surrounded by eight chambered rings, which are
divided by one hundred to one hundred and twenty piercing radial beams into
small chambers. Pores subregular, circular; ten on the radius of the
phacoid shell, two on the breadth of each chamber. Margin of the disk armed
with numerous bristle-shaped radial spines, as long as the breadth of the
chambered girdle. Four very large crossed spines, nearly as long as the
diameter of the disk, quadrangular, with four dentated edges; at the
club-shaped distal end thorny, as broad as the medullary shell and three
times as broad as at the narrow base. I name this splendid species in the
honour of H.R.H. the Grand Duke of Saxe-Weimar, Carl Alexander, the
magnanimous protector of arts and sciences, the rector magnificentissimus
of the University of Jena.

_Dimensions._--Diameter of the disk (with eight rings) 0.27, of the phacoid
shell 0.11, of the medullary shell 0.05; length of the four crossed
club-spines 0.2, basal breadth 0.02, distal breadth 0.06.

_Habitat._--Indian Ocean, Ceylon, Belligemma, surface (Haeckel).

Genus 202. _Astrocyclia_,[241] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with numerous (five or more, commonly thirty
to sixty) solid radial spines on the margin of the circular disk. Medullary
shell simple.

The genus _Astrocyclia_ exhibits on the margin of the circular chambered
disk a large but variable number of solid radial spines, commonly between
thirty and sixty. They are the external prolongations of the inner piercing
radial beams, which divide the concentric rings of the disk into chambers.
All the spines lie in the equatorial plane of the disk. The genus
corresponds to _Stylodictya_ in the family Porodiscida.

1. _Astrocyclia solaster_, n. sp. (Pl. 36, fig. 7).

Phacoid shell two and a half times as broad as the medullary shell,
surrounded by four to six regular rings of equal breadth, which are divided
by thirty to forty piercing radial beams into broad {467}chambers. Pores
regular, circular; seven on the radius of the phacoid shell, one single
pore on each chamber (the size increasing from the centre). Margin with
thirty to forty angular, broad, dentate spines, the prolongations of the
inner radial beams.

_Dimensions._--Diameter of the disk (with six rings) 0.22, of the phacoid
shell 0.1, of the medullary shell 0.04.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

2. _Astrocyclia stella_, Haeckel.

_Lithocyclia stella_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 78, Taf. xxix. fig. 2.

Phacoid shell three times as broad as the medullary shell, surrounded by
four to eight rings of increasing breadth from the centre, the outer of
which are more or less spongy. Each ring is divided by forty to fifty
piercing radial beams into chambers of increasing size. Pores regular,
circular; a single one on each chamber, ten on the radius of the phacoid
shell. On the margin are scattered ten to twenty thick and short spines,
four-sided pyramidal, about as long and as broad at the base as the
diameter of the medullary shell.

_Dimensions._--Diameter of the disk (with eight rings) 0.25, of the phacoid
shell 0.09, of the medullary shell 0.03.

_Habitat._--Fossil in the rocks of Barbados.

3. _Astrocyclia rotula_, n. sp.

Phacoid shell four times as broad as the medullary shell, surrounded by one
single ring, which is divided by forty radial beams into equal chambers.
Pores regular, circular; two on the breadth of the ring, six on the radius
of the phacoid shell. Margin with forty short, conical spines, the
prolongations of the inner radial beams, twice as long as the breadth of
the ring.

_Dimensions._--Diameter of the disk 0.13, of the phacoid shell 0.1, of the
medullary shell 0.025.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

4. _Astrocyclia arachnia_, n. sp.

Phacoid shell twice as broad as the medullary shell, surrounded by four to
six rings of equal breadth, which are divided by twelve to twenty radial
beams into broad chambers. Pores regular, circular; two on the breadth of
each ring, eight on the radius of the phacoid shell. Margin with twelve to
twenty very long and thin, bristle-shaped, radial spines, the prolongations
of the inner beams, longer than the diameter of the disk. (Resembles
_Stylodictya arachnia_, but differs in the possession of a phacoid shell.)

_Dimensions._--Diameter of the disk (with six rings) 0.22, of the phacoid
shell 0.09, of the medullary shell 0.045.

_Habitat._--North Atlantic, Canary Islands.

{468}5. _Astrocyclia heterocycla_, n. sp. (Pl. 36, fig. 8). Phacoid shell
two and a half times as broad as the medullary shell, surrounded by three
rings of unequal breadth, the first ring as broad as the second and third
together. They are divided by fifty to sixty piercing radial beams into
chambers, which are square in the second and third rings, and half as large
as in the first. The radial beams are alternately thicker and thinner,
prolonged into short marginal spines. Pores subregular, circular; eight on
the radius of the phacoid shell, two on the breadth of the first ring, a
single one on each chamber of the second and third rings.

_Dimensions._--Diameter of the disk (with three rings) 0.2, of the phacoid
shell 0.12, of the medullary shell 0.05.

_Habitat._--Equatorial Atlantic, Station 348, depth (2450) fathoms; also
fossil in Barbados.

Genus 203. _Coccocyclia_,[242] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with numerous (five or more) solid radial
spines on the margin of the circular disk. Medullary shell double.

The genus _Coccocyclia_ has the same form and structure as the foregoing
_Astrocyclia_, and differs from it only in the double concentric medullary
shell. It bears therefore to the latter the same relation as _Coccodiscus_
to _Lithocyclia_.

1. _Coccocyclia liriantha_, n. sp.

Phacoid shell three times as broad as the outer and ten times as broad as
the inner medullary shell, connected with both by six equidistant piercing
radial beams. The margin of the phacoid shell is surrounded by one or two
equatorial rings, each as broad as the outer medullary shell, and divided
by thirty to forty radial beams into narrow chambers. Pores subregular,
circular, about nine to ten on the radius of the phacoid shell, three to
four on the breadth of each ring. Margin of the disk ciliated, with six
regularly disposed radial spines as prolongations of the inner six piercing
beams. Each spine cylindrical, delicately crenulated, about as broad as the
inner medullary shell, and once to twice as long as the diameter of the
whole disk, elegantly undulated, with a conical terminal point.

_Dimensions._--Diameter of the disk (with two rings) 0.32, of the phacoid
shell 0.2, of the outer medullary shell 0.06, of the inner 0.02.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

2. _Coccocyclia heliantha_, n. sp. (Pl. 36, figs. 5, 6).

Phacoid shell two and a half times as broad as the outer and six times as
broad as the inner medullary shell, surrounded by six to eight rings of
nearly equal breadth, the outer somewhat smaller. They are divided by fifty
to sixty piercing radial beams into irregular chambers. Pores
{469}irregular, roundish; ten on the radius of the phacoid shell, a single
one on each chamber of the equatorial girdle. Margin of the disk with
numerous strong conical radial spines of different breadths, arranged in
several circles, the strongest in the equatorial plane, about as broad as
one ring and twice as long. Surface thorny.

_Dimensions._--Diameter of the disk (with seven rings) 0.3, of the phacoid
shell 0.12, of the outer medullary shell 0.05, of the inner 0.02.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms; also
fossil in Barbados.

Subfamily 3. ASTRACTURIDA, Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with two or more (commonly three or four)
chambered radial arms on the margin of the disk, situated in its equatorial
plane (with or without a connecting patagium between the arms).

Genus 204. _Diplactura_,[243] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with two opposite chambered arms on the
margin of the circular disk, without a connecting patagium.

The genus _Diplactura_ is the simplest form of the Astracturida, or of
those Coccodiscida in which the margin of the disk bears chambered arms,
situated in its equatorial plane. This subfamily corresponds to the
Euchitonida among the Porodiscida, to the Spongobrachida among the
Spongodiscida. In _Diplactura_ there are only two free arms, opposite in
one equatorial diameter (corresponding to _Amphibrachium_ among the former,
to _Spongolena_ among the latter).

Subgenus 1. _Diplacturium_, Haeckel.

_Definition._--Arms blunt at the distal end, without radial spines.

1. _Diplactura diplobrachia_, n. sp.

Phacoid shell twice as broad as the medullary shell, with six pores on its
radius, without a perfect chambered ring around it. Arms club-shaped, one
and a half times as long as the diameter of the phacoid shell, at the
distal end rounded, blunt, as broad as the latter, at the base only half as
broad.

_Dimensions._--Diameter of the phacoid shell 0.08, of the medullary shell
0.04; length of the arms 0.12, basal breadth 0.04, distal breadth 0.08.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

{470}Subgenus 2. _Diplactinium_, Haeckel.

_Definition._--Arms on the distal end armed with a radial terminal spine.

2. _Diplactura diploconus_, n. sp. (Pl. 38, fig. 5).

_Amphiactura diploconus_, Haeckel, 1877, MS. et Atlas (pl. xxxviii. fig.
5).

Phacoid shell three times as broad as the medullary shell, with eight pores
on its radius, surrounded by a single chambered ring. Arms club-shaped,
twice as long as the diameter of the phacoid shell, at the distal end as
broad as the latter, at the base only half as broad. Both poles of the
common axis of the arms are armed with a strong spindle-shaped terminal
spine.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.03; length of the arms (without terminal spines) 0.17, basal breadth
0.05, distal breadth 0.09.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

Genus 205. _Amphiactura_,[244] Haeckel, 1881, Prodromus, p. 458.

_Definition._--#Coccodiscida# with two opposite chambered arms on the
margin of the circular disk, connected by a spongy patagium.

The genus _Amphiactura_ differs from the foregoing _Diplactura_ in the
development of a patagium between the arms, and therefore bears the same
relation to it as _Amphymenium_ in the Porodiscida does to _Amphibrachium_,
or _Spongobrachium_ in the Spongodiscida does to _Spongolene_. In this and
in the following #Discoidea# provided with a patagium, this connecting web
constantly exhibits a different texture of its framework, which is
sometimes more regularly chambered, at other times more irregularly spongy.

1. _Amphiactura amphibrachia_, n. sp. (Pl. 38, figs. 3, 4).

Phacoid shell three times as broad as the medullary shell, with eight pores
on its radius. Arms nearly equilateral triangular, twice as long as the
diameter of the phacoid shell, at the truncated distal end as broad as the
latter, at the base only one-third as broad. Patagium a circular lenticular
disk, enveloping only the basal third of the arms, with three to four
concentric circular rings, divided into chambers by about forty radial
beams, which are prolonged beyond the margin of the patagium into radial
spines. The vertical section (fig. 4) shows that the chambers of each arm
(eleven to twelve transverse rows in the radius) are disposed in two
layers.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.03; length of the arms 0.18, basal breadth 0.03, distal breadth 0.09.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

{471}Genus 206. _Trigonactura_,[245] Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Coccodiscida# with three chambered arms on the margin of
the circular or triangular disk, without a connecting patagium.

The genus _Trigonactura_ exhibits three radial arms, which in all known
species are separated by three equal angles. The terminal points of the
arm-axes are the corners of an equilateral triangle. It corresponds
therefore to _Dictyastrum_ among the Porodiscida.

Subgenus 1. _Trigonacturium_, Haeckel.

_Definition._--Distal end of the arms blunt or truncated, without a
terminal spine.

1. _Trigonactura pythagoræ_, Haeckel.

_Astromma pythagoræ_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 66, Taf. xxx. fig. 2.

Phacoid shell circular, twice as broad as the medullary shell, with five
pores on its radius, without a completely surrounding chambered girdle.
Arms nearly square, at the truncated distal end as broad as long, and
scarcely broader than at the base, two-thirds as long as the diameter of
the central disk.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length of the arms 0.07, distal breadth 0.07.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, in various depths;
also fossil in Tertiary rocks of Barbados and Nicobar.

2. _Trigonactura rhopalastrella_, n. sp.

_Stephanastrum_ sp., Bury, 1862, Polycystins of Barbados, pl. xv. fig. 7.

Phacoid shell twice as broad as the medullary shell, with five pores on its
radius, without a perfect chambered ring around it. Arms club-shaped, at
the blunt distal end rounded, as long as the diameter of the phacoid shell;
their basal breadth is one-fourth, their distal breadth one-half of its
length.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length of the arms 0.1, basal breadth 0.025, distal breadth 0.05.

_Habitat._--Pacific, central area, Stations 270 to 274, depths 2400 to 2800
fathoms; also fossil in Tertiary rocks of Barbados.

{472}3. _Trigonactura trigonobrachia_, n. sp.

Phacoid shell two and a half times as broad as the medullary shell, with
six pores on its radius, without a perfect chambered girdle. Arms nearly
equilateral triangular, twice as long as the diameter of the phacoid shell,
at the truncated distal end as broad as its diameter and three times as
broad as at the narrow base. (Resembles _Hymenactura hexagona_, but without
a patagium.)

_Dimensions._--Diameter of the phacoid shell 0.11, of the medullary shell
0.045; length of the arms 0.2, basal breadth 0.035, distal breadth 0.1.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

4. _Trigonactura lanceolata_, n. sp.

Phacoid shell three times as broad as the medullary shell, with eight pores
on its radius, surrounded by one perfect chambered ring. Arms lanceolate,
three times as long as the diameter of the phacoid shell, and four times as
long as broad in the middle part, at both ends very narrow, blunt.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.03; length of the arms 0.25, breadth in the midst 0.06.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Subgenus 2. _Trigonactinium_, Haeckel.

_Definition._--Distal end of the arms armed with a radial spine.

5. _Trigonactura triacantha_, n. sp. (Pl. 38, figs. 6, 7).

Phacoid shell two and a half times as broad as the medullary shell, with
eight pores on its radius, surrounded by one perfect chambered ring. Arms
nearly lanceolate, in the middle part one and a half times as broad as at
both ends, twice as long as the phacoid shell, at the distal end with a
strong pyramidal terminal spine. Through the spongy framework of each arm
shine sixteen to eighteen transverse bars (as septa of the joints) and six
to eight longitudinal piercing beams.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.038; length of the arms 0.2, greatest breadth 0.08.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

6. _Trigonactura trigonodiscus_, n. sp.

? _Astromma_ sp., Bury, 1862, Polycystins of Barbados, Taf. xv. fig. 2.

Phacoid shell triangular, twice as broad as the spherical medullary shell,
with six to seven pores on its radius, surrounded by one chambered ring.
Arms equilateral triangular, as long as {473}the diameter of the central
disk and at the base half as broad, slowly decreasing in breadth towards
the distal end, which is armed with a strong pyramidal spine.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length of the arms 0.12, basal breadth 0.07.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms; also
fossil in Barbados.

7. _Trigonactura trixiphos_, n. sp.

Phacoid shell circular, twice as broad as the medullary shell, without a
completely surrounding chambered ring, with four pores on its radius. Arms
club-shaped, about as long as the diameter of the central disk, at the base
half as broad as long, at the rounded distal end broader, and armed with a
strong pyramidal terminal spine, nearly as long as the arm itself.

_Dimensions._--Diameter of the phacoid shell 0.08, of the medullary shell
0.04; length of the arms (without terminal spine) 0.08, basal breadth 0.04,
distal breadth 0.06.

_Habitat._--Fossil in the rocks of Barbados.

Genus 207. _Hymenactura_,[246] Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Coccodiscida# with three chambered arms on the margin of
the circular or triangular disk, connected by a spongy patagium.

The genus _Hymenactura_ differs from the foregoing _Trigonactura_ in the
spongy patagium between the arms, and bears therefore to it the same
relation that among the Porodiscida _Hymeniastrum_ does to the simpler
_Dictyastrum_. The oldest known species of this genus is _Hymenactura
pythagoræ_, described by Ehrenberg as _Hymeniastrum pythagoræ_, but
differing from this in the structure of the central disk.

Subgenus 1. _Hymenacturium_, Haeckel.

_Definition._--Distal end of the arms blunt or truncated, without terminal
spines.

1. _Hymenactura archimedis_, n. sp. (Pl. 38, fig. 8).

Phacoid shell three times as broad as the medullary shell, with eight pores
on its radius. Arms nearly trapezoidal, somewhat longer than the diameter
of the central disk, at the truncated distal end nearly as broad, at the
base only half as broad. In each arm eleven to twelve transverse rows of
square chambers, each of which exhibits on the surface one large pore.
Patagium enveloping the basal half of the arms, with four to five convex
rows of chambers, forming together a circular concentric disk.

_Dimensions._--Diameter of the phacoid shell 0.11, of the medullary shell
0.035; length of the arms 0.12, basal breadth 0.05, distal breadth 0.1.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

{474}2. _Hymenactura pythagoræ_, Haeckel.

_Hymeniastrum pythagoræ_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvi. fig.
31; Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, p. 76, Taf. xxx. fig. 5.

Phacoid shell three times as broad as the medullary shell, with six pores
on its radius. Arms nearly trapezoidal, somewhat longer than the diameter
of the central disk, at the truncated distal end nearly as broad, at the
base two-thirds as broad. In each arm eight to nine transverse rows of
square chambers, each chamber with one pore on the surface. Patagium
enveloping the basal half of the arms, with four to five rectilinear
parallel rows of chambers, forming together a regular triangle. Differs
from the preceding species mainly in the quite different structure of the
patagium, from the true _Hymeniastrum pythagoræ_ in the phacoid shell of
the central disk, which encloses only one simple medullary shell.

_Dimensions._--Diameter of the phacoid shell 0.11, of the medullary shell
0.04; length of the arms 0.13, basal breadth 0.07, distal breadth 0.12.

_Habitat._--Fossil in the rocks of Barbados.

3. _Hymenactura trigona_, n. sp.

_Astromma_ sp., Bury, 1862, Polycystins of Barbados, pl. xv. fig. 1.

Phacoid shell twice as broad as the medullary shell, with four to five
pores on its radius. Arms slender, lanceolate, three times as long as the
diameter of the phacoid shell and five times as long as broad in the middle
part; distal ends blunt. Patagium enveloping only the basal half of the
arms, forming a regular triangle with concave sides.

_Dimensions._--Diameter of the phacoid shell 0.08, of the medullary shell
0.035; length of the arms 0.2, greatest breadth 0.04.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms; fossil in the
rocks of Barbados.

4. _Hymenactura hexagona_.

_Hymeniastrum_ sp., Bury, 1862, Polycystins of Barbados, pl. xv. fig. 3.

Phacoid shell twice as broad as the medullary shell, with four to five
pores on its radius. Arms nearly equilateral triangular, three times as
long as the diameter of the phacoid shell, at the truncated distal end
nearly three-fourths as broad as long, at the narrow base only one-fourth
as broad. Patagium complete, enveloping the arms completely, so that the
whole body represents a regular hexagonal disk.

_Dimensions._--Diameter of the phacoid shell 0.08, of the medullary shell
0.04; length of the arms 0.18, basal breadth 0.04, distal breadth 0.16.

_Habitat._--Fossil in the rocks of Barbados and Nicobar.

{475}Subgenus 2. _Hymenactinium_, Haeckel.

_Definition._--Distal ends of the arms armed with terminal spines.

5. _Hymenactura ptolemæi_, n. sp.

_Astromma_ sp., Bury, 1862, Polycystins of Barbados, pl. xv. figs. 5, 6.

Phacoid shell twice as broad as the medullary shell, with six to seven
pores on its radius. Arms nearly square, about as large as the phacoid
shell, at the truncated distal end little broader than at the base, and
armed with a strong pyramidal terminal spine. Patagium incomplete,
enveloping the basal half of the arms.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length and greatest breadth of the arms 0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms; fossil
in the rocks of Barbados.

6. _Hymenactura copernici_, n. sp. (Pl. 38, fig. 9).

Phacoid shell three times as broad as the medullary shell, with six pores
on its radius. Arms lanceolate, nearly twice as long as the phacoid shell,
in the middle part twice as broad as the medullary shell, with a strong
conical terminal spine at the distal end. In each arm about ten transverse
rows of chambers. Patagium enveloping the basal half of the arms, with four
convex parallel rows of chambers.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.035; length of the arms 0.17, greatest breadth 0.07.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Genus 208. _Astractura_,[247] Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Coccodiscida# with four chambered arms on the margin of the
circular or quadrangular disk, crossed in two equatorial diameters, without
a connecting patagium.

The genus _Astractura_ has the form of a regular cross, four radial arms
being opposite in two equatorial diameters perpendicular one to another. In
the Porodiscida the same form is repeated by _Stauralastrum_, in the
Spongodiscida by _Spongasteriscus_. The oldest known species of the genus
is _Astromma aristotelis_ of Ehrenberg, in which genus this author
confounded triradial and four-radial forms.

{476}Subgenus 1. _Astracturium_, Haeckel.

_Definition._--Distal end of the arms blunt or truncated, without terminal
spines.

1. _Astractura ordinata_, n. sp.

? _Astromma_ sp., Bury, 1862, Polycystins of Barbados, pl. xiv. fig. 3.

Phacoid shell three times as broad as the medullary shell, with six pores
on its radius, without a completely developed chambered ring. Arms
trapezoidal, somewhat longer than the radius of the disk, at the truncated
distal end as broad as long, at the base one-third smaller.

_Dimensions._--Diameter of the phacoid shell 0.08, of the medullary shell
0.03; length of the arms 0.05, distal breadth 0.05, basal breadth 0.035.

_Habitat._--Tropical Atlantic, Station 348, depth 2450 fathoms; also fossil
in Barbados.

2. _Astractura clavigera_, n. sp.

_Astromma aristotelis_, Bury, 1862, Polycystins of Barbados, pl. iv. fig.
2.

Phacoid shell twice as broad as the medullary shell, with ten pores on its
radius, surrounded by one perfect chambered ring. Arms club-shaped, about
as long as the radius of the central disk, at the rounded distal end
two-thirds, at the base one-third as broad as long.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length of the arms 0.06, basal breadth 0.02, distal breadth 0.04.

_Habitat._--Pacific, central area, Stations 270 to 274, depths 2350 to 2925
fathoms; fossil in the rocks of Barbados.

Subgenus 2. _Astractinium_, Haeckel.

_Definition._--Distal end of the arms furnished with a radial spine.

3. _Astractura aristotelis_, Haeckel.

_Astromma aristotelis_, Ehrenberg, 1856, Microgeol., Taf. xxxvi. fig. 32;
Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, p. 66, Taf. xxx. fig. 4.

_Astromma aristotelis_, Haeckel, 1862, Monogr. d. Radiol., p. 489.

? _Astromma aristotelis_, Bury, 1862, Polycystins of Barbados, pl. xiv.
fig. 4.

Phacoid shell three times as broad as the medullary shell, with six to
eight pores on its radius, surrounded by one perfect chambered ring. Arms
nearly trapezoidal, about as long as the diameter of the central disk, at
the base half as broad, at the truncated distal end nearly as broad as
long, and armed with a large pyramidal terminal spine. Each arm with about
eight transverse and longitudinal rows of chambers.

{477}_Dimensions._--Diameter of the phacoid shell 0.11, of the medullary
shell 0.04; length of the arms 0.1, basal breadth 0.05, distal breadth
0.09.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, at various depths;
fossil in the Tertiary rocks of Barbados and Nicobar.

4. _Astractura democriti_, n. sp.

_Astromma aristotelis_, var., Ehrenberg, 1875, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 66, Taf, xxx. fig. 3.

? _Astromma aristotelis_, var., Bury, 1862, Polycystins of Barbados, pl.
xiv. fig. 4.

Phacoid shell twice as broad as the medullary shell, with five to seven
pores on its radius, without perfect chambered ring. Arms about twice as
long as broad, longer than the diameter of the central disk, at the
truncated distal end a little broader than at the base, and armed with a
strong, pyramidal, terminal spine.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length of the arms 0.15, basal breadth 0.04, distal breadth 0.08.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms; fossil
in Barbados.

5. _Astractura hippocratis_, n. sp.

Phacoid shell three times as broad as the medullary shell, with seven pores
on its radius, surrounded by two to three perfect chambered rings. Arms
nearly square, scarcely as long and nearly as broad as the diameter of the
central disk, at the rounded distal end armed with a very strong,
pyramidal, terminal spine, longer than the arm itself, and at the base as
broad as the medullary shell.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.03; length of the arms 0.08, breadth 0.1.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Genus 209. _Stauractura_,[248] Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Coccodiscida# with four chambered arms on the margin of the
circular or quadrangular disk, crossed in two equatorial diameters,
connected by a spongy patagium.

The genus _Stauractura_ differs from the foregoing in the patagium between
the arms, and bears therefore the same relation to it as _Histiastrum_ in
the Porodiscida does to _Stauralastrum_. All known species of this genus
form a regular square, if we connect the end points of the arm axes by
lines.

Subgenus 1. _Stauracturium_, Haeckel.

_Definition._--Distal end of the arms blunt or truncated, without terminal
spines.

{478}1. _Stauractura octogona_, n. sp.

Phacoid shell twice as broad as the medullary shell, with eight pores in
its radius. Arms nearly square, little larger than the phacoid shell, with
broad truncated distal ends. The corners of the latter are so connected by
the complete patagium, that the whole shell forms a regular octagon.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.05; length of the arms 0.12, distal breadth 0.11.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Stauractura tetragona_, n. sp.

Phacoid shell three times as broad as the medullary shell, with eleven
pores on its radius. Arms club-shaped, nearly as long as the diameter of
the phacoid shell, and one and a half times as long as broad at the distal
part; at the distal end rounded, blunt, without terminal spine. The
terminal points of the arms are so connected by a thin, complete patagium,
that the whole shell becomes a regular square.

_Dimensions._--Diameter of the phacoid shell 0.13, of the medullary shell
0.045; length of the arms 0.12, greatest breadth 0.08.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Subgenus 2. _Stauractinium_, Haeckel.

_Definition._--Distal end of the arms furnished with a radial spine.

3. _Stauractura medusina_, n. sp.

Phacoid shell two and a half times as broad as the medullary shell, with
nine pores on its radius. Arms club-shaped, one and a half times as long as
the diameter of the phacoid shell, and in the outer third nearly as broad
as the latter, at the base scarcely one-third as broad; their truncated
distal end armed with a strong pyramidal terminal spine. Patagium
incomplete, enveloping only the basal half of the arms, with three
rectilinear parallel rows of chambers, forming a square.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.04; length of the arms 0.14, greatest breadth 0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

4. _Stauractura quadrata_, n. sp.

Phacoid shell twice as broad as the medullary shell, with seven pores on
the radius. Arms nearly square, about as large as the phacoid shell, with a
strong pyramidal terminal spine at the distal end. Patagium complete,
between every two arms triangular, with rectilinear parallel {479}rows of
chambers; it connects the end points of the arm radius in such a manner
that the whole shell becomes a regular square.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.045; length of the arms 0.1, greatest breadth 0.08.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Genus 210. _Pentactura_,[249] Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Coccodiscida# with five chambered arms on the margin of the
circular or pentagonal disk, without a connecting patagium.

The genus _Pentactura_ possesses five free radial arms, and resembles
_Pentalastrum_ among the Porodiscida. The distance of the five arms seems
to be sometimes equal, at other times different in one and the same
species.

1. _Pentactura pentactis_, Haeckel.

_Astromma pentactis_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 66, Taf. xxx. fig. 1.

Phacoid shell two and a half times as broad as the medullary shell, with
eight pores on its radius. Arms regularly or irregularly disposed, nearly
square, about half as large as the phacoid shell, at the truncated distal
end little broader than at the base. In the specimen figured, and very
imperfectly described by Ehrenberg, (_loc. cit._), the arms were
asymmetrically disposed, two opposite in one axis of the disk, two others
on one side of this axis, the fifth opposite to these on the other side.
Another specimen which I found in the Barbados rocks had five arms of equal
size, regularly disposed, at equal intervals.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.04; length and breadth of the arms 0.05 to 0.06.

_Habitat._--Fossil in the rocks of Barbados.

2. _Pentactura astropecten_, n. sp.

Phacoid shell three times as broad as the medullary shell, with ten pores
on its radius. Arms regularly disposed, club-shaped, nearly twice as long
as the diameter of the phacoid shell, and equal to it in breadth at the
rounded distal end. The latter is armed with a short conical spine, and is
twice as broad as the base.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.03; length of the arms 0.17, basal breadth 0.05, distal breadth 0.08.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

{480}Genus 211. _Echinactura_,[250] n. gen.

_Definition._--#Coccodiscida# with five chambered arms on the margin of the
circular or pentagonal disk, connected by a spongy patagium.

The genus _Echinactura_ differs from the foregoing _Pentactura_ in the
patagium between the arms, and bears therefore the same relation to it as
_Pentinastrum_ in the Porodiscida does to _Pentalastrum_.

1. _Echinactura culcita_, n. sp.

Phacoid shell pentagonal, three times as broad as the medullary shell, with
nine pores on its radius. Arms egg-shaped, about as long as the diameter of
the phacoid shell, and in the distal part half as broad. The blunt ends of
the arms are so connected by the complete patagium that the whole shell
becomes a regular pentagonal disk.

_Dimensions._--Diameter of the phacoid shell 0.1, of the medullary shell
0.035; length of the arms 0.11, basal breadth 0.04, distal breadth 0.06.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Echinactura asteriscus_, n. sp.

Phacoid shell twice as broad as the medullary shell, with seven pores on
its radius. Arms lanceolate, twice as long as the diameter of the phacoid
shell, in the middle part nearly as broad as the latter, at the distal end
with a strong conical terminal spine. Patagium incomplete, enveloping only
the basal half of the arms.

_Dimensions._--Diameter of the phacoid shell 0.09, of the medullary shell
0.045; length of the arms 0.2, greatest breadth 0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Echinactura goniaster_, n. sp.

Phacoid shell three times as broad as the medullary shell, with nine pores
on its radius. Arms club-shaped, one and a half times as long as the
diameter of the phacoid shell, in the distal part nearly as broad as the
latter, in the basal part one-third as broad; at the end a strong pyramidal
spine. Patagium nearly complete, with concave dentated margin between each
pair of arms, therefore the pentagonal disk of the whole shell with concave
sides.

_Dimensions._--Diameter of the phacoid shell 0.12, of the medullary shell
0.04; length of the arms (without terminal spine) 0.18, basal breadth 0.04,
distal breadth 0.1.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

{481}Family XXI. #PORODISCIDA#, Haeckel (Pls. 41-47).

_Porodiscida_, Haeckel, 1881, Prodromus, p. 459.
_Trematodiscida_ et _Discospirida_, Haeckel, 1862, Monogr. d. Radiol.,
pp. 485, 491, 513.
_Calodictya_, Ehrenberg, 1847, Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, p. 53 (_partim_).

_Definition._--#Discoidea# without phacoid shell, with flat discoidal
shell, in which a simple spherical central chamber is surrounded by
concentric chambered rings (each ring divided by radial beams into
imperfect chambers). Surface of the disk on the two flat sides covered by
a porous sieve-plate.

The family #Porodiscida# is by far the largest and richest in different and
common forms among all #Discoidea#; already in my Monograph (1862) nine
genera and twenty-eight species have been enumerated. Their number is here
increased to more than thirty genera and two hundred species. Many of these
species appertain to the most common and widely distributed SPUMELLARIA,
both living and fossil. But the study of their structure is not easy, and
requires (as in the foregoing Coccodiscida) not only careful examination of
the facial views of the disk, but also of the marginal view and of slides
and sections through different planes.

In my Monograph (1862, pp. 485, 491, 513) I had constituted for these
#Discoidea# two different families, the Trematodiscida and Discospirida;
but the comparative study of a far greater number of different types in the
Challenger collection has since convinced me that those two families are
but little different, and united by transitional forms within one and the
same genus, so that they must be united as Porodiscida. Of the group, which
Ehrenberg formerly had called "Calodictya," many genera appertain to the
Porodiscida, whilst many others are true Spongodiscida.

The Porodiscida represent the first and the most important family of the
#Cyclodiscaria#, or of those #Discoidea# which are devoid of the peculiar
extracapsular lenticular "phacoid shell," characteristic of the three
preceding families (united therefore as #Phacodiscaria#). Probably all
#Cyclodiscaria# can be derived from _Archidiscus_, from a morphological as
well as a phylogenetic point of view. _Archidiscus_ seems to be the common
ancestral form not only of the Porodiscida, but also of the nearly allied
Pylodiscida and Spongodiscida. This important _Archidiscus_ (Pl. 48, figs.
9-11) is a small lenticular circular disk, in which a simple latticed
spherical central chamber is surrounded by one single concentric ring,
connected with it by a variable number of radial beams in the equatorial
plane. From this typical _Archidiscus_, as from their "architype," all
other #Cyclodiscaria# may be derived; the Porodiscida by regular apposition
of new concentric chambered rings on the margin, the Spongodiscida by
irregular apposition of a spongy framework, the Pylodiscida by a peculiar
interrupted, concentric, triradial growth, three radial arm-chambers
alternating with three open gates or holes, so that already the first
chambered ring is not complete.

{482}_Archidiscus_ (Pl. 48, figs. 9-11) is not only the common phylogenetic
ancestral form of all #Cyclodiscaria#, but also the common ontogenetic
original form of all Porodiscida, or at least of the greater part of them.
The numerous species of _Archidiscus_, which are distinguished in the
sequel, are at the same time the embryonic forms of different Porodiscida,
corresponding to the "biogenetic main law of development." The small shell
of _Archidiscus_ is sometimes completely lenticular, circular, at other
times more or less polygonal; commonly on the biconvex centre much thicker
than on the margin, but sometimes also of nearly equal thickness (like a
medal or a short cylinder). The latticed central chamber of it is probably
in the majority of species spherical, but in some more or less compressed,
lenticular; the number of small pores on its surface is probably commonly
between ten and twenty (four to eight usually being visible on each
hemisphere). The number of radial beams, which connect it with the
equatorial ring, varies commonly from four to eight; but sometimes only two
or three are to be found, in other cases nine to ten or more. The regular
disposition of these beams (in certain equatorial axes of the disk) is
probably of great importance, as determining the later development of
characteristic radial appendages of the margin in the more highly developed
Porodiscida. The equatorial ring itself, forming the margin of the
lenticular disk, is either a simple solid ring or a broader latticed
girdle; in the latter case it merges slowly into the opposite sieve-plates
of the two flat disk sides, or the porous "cover-plates," covering its
parallel or convex surfaces. These latter can be regarded as direct
peripheral continuations of the polar regions of the spherical central
chamber. The ring-chambers, surrounding the latter in a single circle, are
commonly of nearly the same breadth, but often also of different irregular
size. Their number varies between two and ten or more, but commonly between
four and eight; each ring-chamber is covered on the upper and lower side by
the sieve-plate, bounded on the inner (proximal) side by the wall of the
central chamber, on the outer (distal) side by the marginal ring, on both
lateral sides by the contiguous neighbouring ring-chambers.

The important question as to the phylogenetic origin of _Archidiscus_ can
be answered in a twofold way. The most simple form of _Archidiscus_
(_Archidiscus dioniscus_) can be derived immediately from the
Stylosphærida, _Saturnalis_ (Pl. 13, fig. 16), only by the development of
lattice-work between the equatorial ring and the two polar faces of the
concentric central chamber (on the surface of the biconvex jelly-mantle).
But on the other hand _Archidiscus_ may also be derived from the simplest
Phacodiscida, _Sethodiscus_ (Pl. 33, figs. 1-3), by the stronger
compression of the biconvex lenticular shell, so that the enclosed
medullary shell on the two poles runs together with the lenticular phacoid
shell, of which only the peripheral part remains free, and thus forms the
chambered ring; this latter explanation seems the more natural in many
cases, as often in the Porodiscida the central chamber is enclosed in one
or two concentric spherical or lenticular lattice-shells.

{483}The second subfamily of the Porodiscida are the Trematodiscida, which
are derived from the preceding Archidiscida by concentric growth in the
equatorial plane. In the same way in which the simple chambered ring of
_Archidiscus_ is connected by radial beams with the central chamber, so in
_Porodiscus_ a variable number of concentric rings is connected with that
first ring. The number of these concentric rings varies between two and ten
or more, but commonly amounts to between three and five. The radial beams
connecting them are either piercing or interrupted; their number increases
gradually from the centre towards the periphery. The chambers between them
are sometimes more regular, at other times more irregular in size and form.
Their upper and lower wall is formed by the two covering "sieve-plates,"
or the porous cover-plates, which are continued from the central disk to
the margin. If these two sieve-plates continue being parallel, the disk
becomes medal-shaped or a short cylinder; if the two plates become more or
less concavely vaulted one against the other, the disk becomes biconvex
lenticular, the middle part thicker than the marginal part. Rarely the
contrary is the case, the margin thicker than the centre, and then the disk
biconcave.

In my Monograph I had separated as two different subfamilies the true
Trematodiscida (with circular concentric rings) from the Discospirida (with
spirally convoluted rings). But the enormous mass of specimens, which I
afterwards examined in the Challenger collection, has convinced me that
this separation was not natural. For in one and the same genus of most
nearly allied forms we find on one hand quite regular concentric circular
forms (_Trematodiscus_), on the other hand spirally convoluted forms
(_Discospira_), and connecting between them such forms as are in the
central part concentric, in the marginal part spiral (_Perispira_)--or
conversely, these in the centre spiral, on the margin concentric
(_Centrospira_)--and frequently also more or less irregular forms with
interrupted rings (_Atactodiscus_); therefore, all those genera (Prodromus,
1881, Nos. 448-452) have only the value of subgenera of _Porodiscus_. But a
distinct genus is _Perichlamydium_, in which the two sieve-plates run on
the margin of the lens and form a broad hyaline porous or solid girdle.
More important is the distinction of the Ommatodiscida, in which the margin
of the disk exhibits one larger osculum, armed with a corona of spines
(_Ommatodiscus_), or two oscula, opposite on the poles of one axis
(_Stomatodiscus_). Whilst in many Porodiscida all chambers of the
concentric rings lie in one and the same (equatorial) plane, in many others
with further growth they become stratified in floors, and the whole disk is
therefore composed of two to four or more parallel disks, each with a
system of concentric chambered rings or girdles, quite as in the majority
of the Coccodiscida (p. 457). Often the central part of the lenticular
disk becomes thickened by apposition of such floors or strata, whilst the
marginal part remains simple, with one single stratum. The communication
between the chambers of the different strata seems to be the same as in the
similar Coccodiscida.

Also the margin of the disk exhibits in the Porodiscida the same
characteristic {484}differences as in the foregoing family. In the
subfamily of Stylodictyida it bears a certain number of solid radial
spines, often regularly disposed (as in the Stylocyclida). In the subfamily
of Euchitonida the margin is distinguished by the possession of two to six
or more (commonly three or four) chambered arms, also situated in the plane
of the disk, and of the same structure (sometimes more or less irregular,
spongy). These arms are very variable in size, form, and structure, exhibit
the same peculiarities as in the Astracturida, and are sometimes free, at
other times connected by a "patagium" or an interbrachial spongy skeleton
of different structure, like a web membrane (compare above, p. 458). In
some genera the arms become forked or branched on the distal end.
Sometimes their distal end bears a terminal radial spine.

_The Central Capsule_ of the Porodiscida assumes generally the form of the
including shell, with or without arms, but is constantly somewhat smaller,
as it remains enclosed by the sieve-plates of the disk surface. Often the
capsule is filled with many coloured oil-globules, disposed regularly in
the chamber rows. The nucleus of it is enclosed by the central chamber, and
in many cases by this and the innermost concentric rings.

_Synopsis of the Genera of the Porodiscida._

-------------------------------------------------------------------------
I. Section of the Porodiscida--Archidiscaria.
Central chamber of the disk surrounded only by one single chambered
ring.
--------------------
1. Subfamily Archidiscida.
Only one single chambered girdle surrounds the central chamber; margin
smooth or spiny.
-------------------------------------------------------------------------
Margin smooth, without spines, 212. _Archidiscus_.

Margin armed with radial spines, 213. _Axodiscus_.

II. Section of the Porodiscida--Astrodiscaria.

Central chamber of the disk surrounded by two or more (commonly three to
six) concentric chambered rings or radiated girdles.
-------------------------------------------------------------------------
II. Section of the Porodiscida--Astrodiscaria.
Central chamber of the disk surrounded by two or more (commonly three
to six) concentric chambered rings or radiated girdles.
--------------------
2. Subfamily Trematodiscida.
Margin of the disk quite simple, without radial appendages (spines or
chambered arms), without peculiar oscula.
-------------------------------------------------------------------------
Margin simple, without an equatorial girdle, 214. _Porodiscus_.

Margin with a thin porous equatorial girdle, 215. _Perichlamydium_.
-------------------------------------------------------------------------
3. Subfamily Ommatodiscida. Margin of the disk without chambered arms but
distinguished by one or two large oscula, or wide openings armed with a
crown of spines.
-------------------------------------------------------------------------
Disk with a single marginal osculum, 216. _Ommatodiscus_.

Disk with two opposite marginal oscula, 217. _Stomatodiscus_.
-------------------------------------------------------------------------
4. Subfamily Stylodictyida.
Margin of the disk without peculiar oscular openings and without
chambered arms, but armed with solid radial spines.
-------------------------------------------------------------------------
Radial spines of the { Two opposite spines, 218. _Xiphodictya_.
margin two, three, or{
four, usually quite { Three equidistant
regularly disposed. { spines, 219. _Tripodictya_.
{
{ Four crossed spines, 220. _Staurodictya_.

Radial spines of the { Margin simple, without
margin five or more, { an equatorial girdle, 221. _Stylodictya_.
commonly irregularly {
disposed (generally {Margin with a thin porous
ten to twelve). { equatorial girdle, 222. _Stylochlamydium_.
-------------------------------------------------------------------------
5. Subfamily Euchitonida.{485}
Margin of the disk with radial chambered arms (or hollow chambered
appendages), on the distal end of the arms with or without solid
radial spines.
-------------------------------------------------------------------------
{ Two arms { Without a patagium, 223. _Amphibrachium_.
{ simple, {
Two arms, { undivided. { With a patagium, 224. _Amphymenium_.
opposite {
in one { Two arms { Without a patagium, 225. _Amphirrhopalum_.
main axis.{ forked (or {
{ one simple, {
{ other { With a patagium, 226. _Amphicraspedum_.
{ forked). {

{ { Regular, 227. _Dictyastrum_.
{ { Without a {
Three { Three arms { patagium. { Bilateral, 228. _Rhopalastrum_.
arms { simple, {
(either { undivided. {
regular, { { { Regular, 229. _Hymeniastrum_.
equal, or { { With a {
bilateral,{ { patagium. { Bilateral, 230. _Euchitonia_.
with { { {
unequal {
arms). { { Without a
{ { Either { patagium, 231. _Chitonastrum_.
{ Three arms {regular or {
{ forked. {bilateral. { With a
{ { { patagium, 232. _Trigonastrum_.

Four arms { { { Regular, 233. _Stauralastrum_.
(either { { Without a {
regular, { Four arms { patagium. { Bilateral, 234. _Hagiastrum_.
with four { simple, {
equal { undivided. { { Regular, 235. _Histiastrum_.
crossed { { With a {
arms, or { { patagium. { Bilateral, 236. _Tessarastrum_.
bilateral,{ {
with { { With a terminal
paired { { patagial girdle, 237. _Stephanastrum_
arms). {
{ { Regular, 238. _Dicranastrum_.
{ Four arms { Without a {
{ forked. { patagium. { Bilateral, 239. _Myelastrum_.

{ Five arms { Without a patagium, 240. _Pentalastrum_.
{ simple, {
Five arms { undivided. { With a patagium, 241. _Pentinastrum_.
(equal or {
unequal). { Five arms { Without a patagium, 242. _Pentophiastrum_.
{ forked. {

Six arms { Six arms { Without a patagium, 243. _Hexalastrum_.
(equal or { simple, {
unequal). { undivided. { With a patagium, 244. _Hexinastrum_.

Subfamily 1. ARCHIDISCIDA, Haeckel.

_Definition._--#Porodiscida# with a simple spherical or lenticular latticed
central chamber, surrounded by a single concentric latticed ring, which is
divided by radial beams into two to six or more radial chambers.

{486}Genus 212. _Archidiscus_,[251] n. gen.

_Definition._--#Porodiscida# with a simple central chamber, surrounded by a
single concentric ring, which is divided by radial beams into two to six or
more radial chambers, without radial spines on the margin.

The genus _Archidiscus_ begins the long and polymorphous series of the
#Cyclodiscaria# or of those #Discoidea# which do not possess the peculiar
"phacoid shell" characteristic of the three preceding families, united as
"#Phacodiscaria#." As already mentioned above, both these groups are
probably of independent origin, derived from the #Sphæroidea# in different
ways (compare pp. 402, 405, &c.). Among all #Cyclodiscaria# _Archidiscus_
is the most simple, and probably the common ancestral form, from which the
other genera may be derived.

Subgenus 1. _Dioniscus_, Haeckel.

_Definition._--Ring with two chambers, separated by two radial beams.

1. _Archidiscus dioniscus_, n. sp.

Ring circular, connected with the central chamber by two radial beams,
opposite in one axis, therefore two equal semicircular ring chambers. (This
primitive form has an interesting reference to _Saturnalis_, Pl. 13, fig.
16, and differs from it only in the lattice-work covering both faces of the
lenticular disk, the margin of which forms the ring.)

_Dimensions._--Diameter of the ring 0.05, of the central chamber 0.016.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

2. _Archidiscus dithalamus_, n. sp.

Ring roundish, connected with the central chamber by two radial beams, not
opposite in one axis; both semicircular ring chambers more or less unequal,
one of them larger than the other, and sometimes much more prominent. (If
this prominence increase, we can regard it as the beginning of spiral
convolutions, _Discospira_.)

_Dimensions._--Diameter of the ring 0.06, of the central chamber 0.014.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

Subgenus 2. _Trioniscus_, Haeckel.

_Definition._--Ring with three chambers, separated by three radial beams.

{487}3. _Archidiscus trioniscus_, n. sp.

Ring triangular, roundish, equilateral, connected with the central chamber
by three radial beams at equal distances (120°); therefore three equal ring
chambers. (Resembles the central part of the disk of _Tripodictya
trigonaria_, Pl. 42, fig. 8, and may be the ancestral form of it.)

_Dimensions._--Diameter of the ring 0.04, of the central chamber 0.014.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

4. _Archidiscus trithalamus_, n. sp.

Ring irregular, roundish, connected with the central chamber by three
radial beams at unequal distances; therefore three ring chambers of
different size. (If these differences be important, they introduce a spiral
convolution in the further development of the _Porodiscus_ arising from
it.)

_Dimensions._--Diameter of the ring 0.06, of the central chamber 0.016.

_Habitat._--South Pacific, Station 288, surface.

Subgenus 3. _Tetroniscus_, Haeckel.

_Definition._--Ring with four chambers, separated by four radial beams.

5. _Archidiscus stauroniscus_, n. sp. (Pl. 48, figs. 9, 9_a_).

Ring regular, square, connected with the central chamber by four radial
beams at equal distances, opposite in pairs in two axes perpendicular one
to another; therefore four equal ring chambers (or congruent quadrants of
the ring). Resembles the central part of the disk of _Staurodictya medusa_,
&c. (Pl. 42, figs. 1-3); also of _Staurodictya gracilis_, Ehrenberg, 1875
(Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. xxiii. fig. 3).

_Dimensions._--Diameter of the ring 0.05, of the central chamber 0.016.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

6. _Archidiscus tetroniscus_, n. sp.

Ring elliptical, of rhomboidal fundamental form, connected with the central
chamber by four radial beams, halving the sides of the rhombus and opposite
in pairs in two axes which are not perpendicular one to another; therefore
four ring chambers in pairs different, two opposite equal and larger than
the two others.

_Dimensions._--Diameter of the ring 0.05 to 0.07, of the central chamber
0.016.

_Habitat._--Central Pacific, Station 263, depth 2650 fathoms.

7. _Archidiscus tetrathalamus_, n. sp.

Ring irregular, quadrangular, connected with the central chamber by four
radial beams of increasing unequal length; therefore all four ring chambers
of different size, gradually increasing {488}in the following quadrants of
the ring. Important as an ancestral type of such spiral and semi-spiral
forms as _Staurodictya cruciata_ (Pl. 42, figs. 4, 12, &c.) and
_Stylodictya clavata_, _Stylodictya stellata_, &c., of Ehrenberg, 1875
(Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. xxiii. figs. 2, 7, 8, 9).

_Dimensions._--Diameter of the ring 0.05 to 0.08, of the central chamber
0.06.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Subgenus 4. _Pentoniscus_, Haeckel.

_Definition._--Ring with five chambers, separated by five radial beams.

8. _Archidiscus pentoniscus_, n. sp.

Ring pentagonal or nearly circular, regular, connected with the central
chamber by five radial beams of equal length and at equal distances (72°);
therefore all five chambers of the ring of equal size and similar form.
(Resembles the central part of the disk of _Pentinastrum asteriscus_, Pl.
44, fig. 2.)

_Dimensions._--Diameter of the ring 0.05, of the central chamber 0.015.

_Habitat._--North Pacific, Station 244, surface.

Subgenus 5. _Hexoniscus_, Haeckel.

_Definition._--Ring with six chambers, separated by six radial beams.

9. _Archidiscus hexoniscus_, n. sp. (Pl. 48, figs. 10, 10_a_).

Ring regular, hexagonal, or nearly circular, connected with the central
chamber by six radial beams of equal length and at equal distances (60°);
therefore all six chambers of the same size and form. (Resembles the
central part of the disk of _Hexinastrum geryonidum_, Pl. 44, fig. 4.)

_Dimensions._--Diameter of the ring 0.06, of the central chamber 0.018.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

10. _Archidiscus pyloniscus_, n. sp. (Pl. 48, figs. 11, 11_a_).

Ring triangular, connected with the central chamber by six radial beams at
alternating distances; therefore three larger chambers (of looser network)
alternate with three smaller chambers (of denser network); pores of the
former twice to three times as large as those of the latter. This species
is of peculiar importance, as an immediate transitional form to the
Pylodiscida. If these three larger ring chambers lose their few
lattice-beams and so became open gates, we get _Triolene_ or _Triopyle_,
the original forms of the Pylodiscida.

_Dimensions._--Diameter of the ring 0.05, of the central chamber 0.015.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

{489}11. _Archidiscus hexathalamus_, n. sp.

Ring irregular, roundish, or hexagonal, connected with the central chamber
by six radial beams of unequal increasing length; therefore all six ring
chambers of gradually increasing size (beginning a spiral convolution,
original form of some _Discospira_).

_Dimensions._--Diameter of the ring 0.05 to 0.07, of the central chamber
0.014.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

Subgenus 6. _Circoniscus_, Haeckel.

_Definition._--Ring with seven or more chambers, separated by seven or more
radial beams.

12. _Archidiscus octoniscus_, n. sp.

Ring circular or regular octagonal, connected with the central chamber by
eight equidistant radial beams; therefore eight ring chambers of equal
size. (Compare the central part of the disk of _Porodiscus quadrigatus_,
Pl. 41, fig. 3.)

_Dimensions._--Diameter of the ring 0.04, of the central chamber 0.014.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

13. _Archidiscus polythalamus_, n. sp.

Ring circular, connected with the central chamber by nine to ten or more
radial beams at nearly equal distances. Ring chambers nine to ten, more or
less equal, sometimes also eleven to twelve, more different. (This species
is very variable, and may perhaps be divided into a number of different
"transformistic species.")

_Dimensions._--Diameter of the ring 0.04 to 0.06, of the central chamber
0.015.

_Habitat._--Central Pacific, Stations 265 to 274, depths 2350 to 2925
fathoms.

Genus 213. _Axodiscus_,[252] n. sp.

_Definition._--#Porodiscida# with a simple central chamber, surrounded by
one single concentric ring, which is divided by radial beams into two to
six or more radial chambers; margin of the disk armed with radial spines.

The genus _Axodiscus_ differs from the preceding _Archidiscus_ only in the
shape of the margin of the small lenticular disk, which is armed with a
variable number of radial spines, indicating certain axes or radii of the
shell. If these marginal spines at certain equal distances from the margin
branch and their distal ends become united by these branches forming a
concentric second ring, the genus passes into _Porodiscus_. The different
number and disposition of the marginal spines are probably very important,
{490}as determining the later development of two to four or more radii in
the different genera of Porodiscida.

1. _Axodiscus stylophorus_, n. sp.

Ring circular, with two equal semicircular chambers, connected with the
central chamber by two opposite radial beams, which are prolonged outside
into two strong conical spines.

_Dimensions._--Diameter of the ring 0.05, of the central chamber 0.016.

_Habitat._--Central Pacific, Station 266, depth 2750 fathoms.

2. _Axodiscus triradiatus_, n. sp.

Ring regular, triangular, equilateral, connected with the central chamber
by three equidistant radial beams, which are prolonged outside into three
short conical spines. (Differs from _Archidiscus trioniscus_ in the
possession of marginal spines.)

_Dimensions._--Diameter of the ring 0.04, of the central chamber 0.013.

_Habitat._--Central Pacific, Station 267, depth 2700 fathoms.

3. _Axodiscus staurophorus_, n. sp.

Ring regular, square, connected with the central chamber by four radial
beams, which lie opposite in pairs in two perpendicularly crossed axes, and
are prolonged outside into four delicate cylindrical spines.

_Dimensions._--Diameter of the ring 0.045, of the central chamber 0.015.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

4. _Axodiscus hexagonus_, n. sp.

Ring regular, hexagonal, connected with the central chamber by six radial
beams at equal distances, which are prolonged outside into six strong
pyramidal spines.

_Dimensions._--Diameter of the ring 0.06, of the central chamber 0.018.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

5. _Axodiscus trigonus_, n. sp.

Ring regular, triangular, equilateral, connected with the central chamber
by six radial beams of alternating length and distance; therefore three
smaller (perradial) chambers (with denser and darker network) alternating
with three larger (interradial) chambers (with looser and finer network).
On the margin three strong conical radial spines, arising in the radius of
the larger chambers. (Differs from _Archidiscus pyloniscus_, Pl. 48, fig.
11, mainly in the strong angular spines on the three corners of the
triangular shell.)

_Dimensions._--Diameter of the ring 0.05, of the central chamber 0.014.

_Habitat._--South Pacific, Station 296, depth 1825 fathoms.

{491}6. _Axodiscus octogonus_, n. sp.

Ring circular or nearly octagonal, connected with the central chamber by
eight radial beams at nearly equal distances, which are prolonged outside
into eight short conical spines. (Differs from _Archidiscus octoniscus_ in
the marginal prolongation of the eight beams.)

_Dimensions._--Diameter of the ring 0.04, of the central chamber 0.013.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

7. _Axodiscus spinosus_, n. sp.

Ring circular, connected with the central chamber by ten to twelve radial
beams at nearly equal distances, which are prolonged outside into short
conical spines of variable length. (May be regarded as an aculeate variety
of _Archidiscus polythalamus_.)

_Dimensions._--Diameter of the ring 0.04 to 0.05, of the central chamber
0.014.

_Habitat._--Central Pacific, Stations 265 to 274, depths 2350 to 2950
fathoms.

Subfamily 2. TREMATODISCIDA, Haeckel, 1862, Monogr. d. Radiol., p. 491
(_sensu emendato et restricto_).

_Definition._--#Porodiscida# without radial appendages of the disk (solid
spines or chambered arms on the margin), and without peculiar oscula on the
margin of the disk, which is composed of two to four or more concentric
rings.

Genus 214. _Porodiscus_,[253] Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Porodiscida# with simple circular disk, composed of several
rings (without radial appendages or peculiar oscula on the margin of the
disk).

The genus _Porodiscus_ is, next to its ancestral form, _Archidiscus_, the
simplest and most primitive form of the Porodiscida, from which all other
genera of this family can be derived. The disk is quite simple, without any
marginal appendages, composed of a variable number of rings, commonly of
circular form, sometimes more or less polygonal, elliptical, or irregular.
In my Monograph (1862, pp. 491, 513) I had separated the species, here
united in _Porodiscus_, into two different genera: _Trematodiscus_ with
concentric rings, and _Discospira_ with spiral rings. But the extended
study of these very common forms in a great number of specimens in the
Challenger collection has convinced me that the separation of those two
genera cannot be maintained. In one and the same locality, where one single
characteristic disk-form is very common, we find intermingled quite regular
disks with only concentric, circular rings (_Trematodiscus_), and other
disks with one single perfect spiral ring (_Discospira_); and between
{492}these a smaller number of specimens, in which the rings of the disk
are partly concentric, partly spiral; either the rings of the central part
of the disk are concentric, the outer spiral (_Perispira_), or the
proportion is inverse (_Centrospira_); and sometimes the whole disposition
of the concentric and spiral rings is irregular, and the rings often
interrupted (_Atactodiscus_). Therefore it appears more natural to give to
all these different forms only the value of subgenera of _Porodiscus_, as I
have already proposed in my Prodromus (1881, p. 459). Even the numerous
species of _Porodiscus_ (mainly characterised by the equal or different
breadth of the rings, and by the number, form, and size of the connecting
radial beams and of the superficial pores) are for the most part very
variable and hard to distinguish, as all those characters are not constant.
_Porodiscus_ is a quite "transformistic genus."

Subgenus 1. _Trematodiscus_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 841.

_Definition._--All rings of the disk concentric (commonly circular, rarely
a little elliptical or polygonal).

1. _Porodiscus orbiculatus_, Haeckel.

_Trematodiscus orbiculatus_, Haeckel, 1862, Monogr. d. Radiol., p. 492,
Taf. xxix. fig. 1.

_Trematodiscus orbiculatus_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 108.

All rings of the disk circular, concentric, of equal breadth, connected by
numerous alternating radial beams. Chambers differing little in size, about
as large as the central chamber. Pores regular, circular, two to two and a
half on the breadth of each ring.

_Dimensions._--Diameter of the disk (with nine rings) 0.18; breadth of each
ring 0.01; pores 0.003.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface; also fossil in Tertiary rocks of Barbados, Sicily, and Nicobar.

2. _Porodiscus concentricus_, Haeckel.

_Trematodiscus concentricus_, Haeckel, 1862, Monogr. d. Radiol., p. 493.

_Trematodiscus concentricus_, Stöhr, 1880, Palæontogr., vol. xxvi. p.
108.

_Flustrella concentrica_, Ehrenberg, 1838, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 132; _Ibid._, 1875, p. 72, Taf. xxii. fig. 13.

? _Flustrella concentrica_, Ehrenberg, 1854, Mikrogeol., Taf. xix. fig.
61, Taf. xx. fig. 42, Taf. xxi. fig. 51, Taf. xxxvi. fig. 29.

All rings of the disk circular, concentric, of equal breadth, connected by
numerous piercing radial beams. Chambers different in size, increasing
from the centre towards the periphery. Pores regular, circular, one and
half to two on the breadth of each ring.

_Dimensions._--Diameter of the disk (with eight rings) 0.16; breadth of
each ring 0.01; pores 0.003.

_Habitat._--Fossil in many Tertiary rocks--Barbados, Sicily, Greece, &c.

_Porodiscus haeckelii_ = _Trematodiscus haeckelii_, Bütschli, 1882, L. N.
41, Taf. xxiv. figs. 5_a_, 5_b_.

{493}3. _Porodiscus flustrella_, n. sp. (Pl. 41, fig. 1).

_Trematodiscus flustrella_, Haeckel, 1866, MS. Canar. Ins.

All rings of the disk circular, concentric, of equal breadth, connected by
numerous piercing radial beams. Chambers different in size, increasing from
the centre towards the periphery. Pores very irregular, polygonal, or
roundish, one to three on the breadth of each ring.

_Dimensions._--Diameter of the disk (with six rings) 0.12; breadth of each
ring 0.01; pores 0.002 to 0.006.

_Habitat._--Cosmopolitan; Atlantic, Pacific, &c., many Stations, surface.

4. _Porodiscus sorites_, Haeckel.

_Trematodiscus sorites_, Haeckel, 1862, Monogr. d. Radiol., p. 492, Taf.
xxix. fig. 2.

All rings of the disk circular, concentric, of equal breadth, connected by
numerous alternating radial beams. Chambers different in size, increasing
from the centre. Pores regular, circular, one to one and a half on the
breadth of each ring.

_Dimensions._--Diameter of the disk (with six rings) 0.12; breadth of each
ring 0.01; pores 0.006.

_Habitat._--Mediterranean (Messina), surface.

5. _Porodiscus macroporus_, Haeckel.

_Trematodiscus macroporus_, Haeckel, 1879, MS.

_Flustrella macropora_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 160.

All rings of the disk circular, concentric, of equal breadth, connected by
numerous piercing radial beams. Chambers of different breadth, increasing
from the centre. Pores regular, very large, nearly square, one single pore
on the breadth of each ring.

_Dimensions._--Diameter of the disk (with five rings) 0.11; breadth of each
ring 0.01; pores 0.005.

_Habitat._--Fossil in the Tertiary rocks of Barbados.

6. _Porodiscus microporus_, Haeckel.

_Trematodiscus microporus_, Stöhr, 1880, Palæontogr., vol xxvi. p. 108,
Taf. iv. fig. 17.

All rings of the disk concentric, either circular or a little elliptical;
the innermost rings of the same breadth as the central chamber, the third
ring much broader. Radial beams between them alternating; chambers of
different size. Pores very small, subregular, everywhere of the same size,
four to five pores on the breadth of each ring.

_Dimensions._--Diameter of the disk (with three rings) 0.15; breadth of the
inner rings 0.02, of the outer 0.03; pores 0.0016.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

{494}7. _Porodiscus ellipticus_, Haeckel.

_Trematodiscus ellipticus_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 108,
Taf. iv. fig. 16.

All rings of the disk concentric, either circular or a little elliptical,
connected by eight piercing radial beams; central chamber elliptical, of
the same breadth as the first ring, broader than the following rings, the
breadth of which decreases towards the periphery. Pores irregular, in the
inner rings twice as broad as in the outer (third) ring, four to five on
the breadth of each ring.

_Dimensions._--Diameter of the disk (with three rings) 0.18; breadth of the
inner rings 0.03, of the outer 0.02; pores 0.003 to 0.006.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

8. _Porodiscus heterocyclus_, Haeckel.

_Trematodiscus heterocyclus_, Haeckel, 1862, Monogr. d. Radiol., p. 493,
Taf. xxix. fig. 3.

? _Flustrella cyclia_, Harting, 1863, Fauna Banda-Zee, p. 11, pl. i. fig.
19.

_Trematodiscus heterocyclus_, Stöhr, 1880, Palæontogr., vol. xxvi. p.
108.

All rings of the disk concentric, circular, connected by numerous radial
beams, which are partly piercing, partly alternating. The breadth of the
rings increases gradually from the centre towards the periphery,
corresponding also to the size of the pores; two to three circular pores on
the breadth of each ring.

_Dimensions._--Diameter of the disk (with six rings) 0.17; breadth of the
second ring 0.007, of the sixth 0.02; pores 0.002 to 0.006.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface; also
fossil in Tertiary rocks of Barbados and the Mediterranean.

9. _Porodiscus quadrigatus_, n. sp. (Pl. 41, fig. 3).

All rings of the disk concentric, of nearly equal breadth, connected by
four interradial beams, perpendicular one to another. The first ring
(surrounding the central chamber) with eight chambers, the second only with
four (alternating with the four radial beams). Size of the four chambers of
each ring increases much towards the periphery. Pores regular, circular,
about two on the breadth of each ring.

_Dimensions._--Diameter of the disk (with six rings) 0.18; breadth of each
ring 0.05; pores 0.004.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Subgenus 2. _Perispira_, Haeckel, 1881, Prodromus, p. 459.

_Definition._--The inner rings of the disk concentric (commonly circular);
the outer rings spirally convoluted.

{495}10. _Porodiscus perispira_, n. sp. (Pl. 41, fig. 2).

_Perispira perforata_, Haeckel, 1881, Prodromus.

Inner rings of the disk (two or three) concentric, outer rings (three or
four) spirally convoluted, spiral line simple. All rings nearly of the same
breadth, connected by alternating irregular radial beams. Pores irregular,
two to three on the breadth of each ring.

_Dimensions._--Diameter of the disk (with six rings) 0.17; breadth of each
ring 0.016; pores 0.003 to 0.005.

_Habitat._--Pacific, central area, Stations 272 to 274, surface.

11. _Porodiscus radiatus_, n. sp.

_Perispira radiata_, Haeckel, 1881, Prodromus.

Inner rings of the disk (four or five) concentric, outer rings (three or
four) spirally convoluted, spiral line simple. All rings connected by
piercing radial beams (eight in the inner half, sixteen in the outer half).
Breadth of the rings and of the pores increasing from the centre towards
the periphery, three to four pores on the breadth of each ring.

_Dimensions._--Diameter of the disk (with eight rings) 0.18; breadth of the
second ring 0.006, of the eighth ring 0.02; pores 0.002 to 0.006.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Subgenus 3. _Centrospira_, Haeckel, 1881, Prodromus, p. 459.

_Definition._--The inner rings of the disk spirally convoluted, the outer
rings concentric (commonly circular).

12. _Porodiscus centrospira_, n. sp. (Pl. 41, fig. 6).

_Centrospira perispongidium_, Haeckel, 1881, Prodromus.

Inner rings of the disk (two or three) spirally convoluted (with simple or
double spiral line), outer rings (three or four) concentric, subcircular.
All rings nearly of the same breadth, connected by alternating radial
beams. Pores subregular, two to three on the breadth of each ring.
Lattice-work in the periphery of the disk a little spongy (as in Pl. 41,
fig. 11).

_Dimensions._--Diameter of the disk (with five rings) 0.15; breadth of each
ring 0.015; pores 0.003.

_Habitat._--Pacific, central area, Station 267, depth 2700 fathoms.

Subgenus 4. _Discospira_, Haeckel, 1862, Monogr. d. Radiol., p. 513.

_Definition._--All rings of the disk spirally convoluted, forming parts of
a simple or double spiral turning.

{496}13. _Porodiscus helicoides_, Haeckel.

_Discospira helicoides_, Haeckel, 1862, Monogr. d. Radiol., p. 514, Taf.
xxix. fig. 7.

_Trematodiscus helicoides_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 841.

All rings of the disk spirally convoluted around the central chamber;
spiral line regular, simple. All rings nearly of the same breadth,
connected by numerous alternating radial beams. Chambers little different
in size, little longer than broad. Pores of equal size, regular, two on the
breadth of each ring.

_Dimensions._--Diameter of the disk (with ten rings) 0.2; breadth of each
ring 0.01; pores 0.0025.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific; also fossil in
the Tertiary rocks of the Mediterranean, Sicily, Oran, &c.

14. _Porodiscus spiralis_, Haeckel.

_Flustrella spiralis_, Ehrenberg, 1840, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 210; Mikrogeol., 1854, Taf. xix. fig. 62.

_Discospira spiralis_, Haeckel, 1862, Monogr. d. Radiol., p. 513.

All rings of the disk spirally convoluted around the central chamber;
spiral line regular, simple. All rings nearly of the same breadth,
connected by numerous (twelve to sixteen) piercing radial beams. Chambers
increasing in size from the centre towards the periphery. Pores irregular,
of different size, one and a half to two on the breadth of each ring.

_Dimensions._--Diameter of the disk (with eight rings) 0.16; breadth of
each ring 0.01; pores 0.004.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific; also
fossil in the Tertiary rocks of Barbados, Sicily, Greece, &c.

15. _Porodiscus operculina_, Haeckel.

_Discospira operculina_, Haeckel, 1862, Monogr. d. Radiol., p. 514, Taf.
xxix. fig. 8.

All rings of the disk spirally convoluted around the central chamber,
spiral line regular, simple. Breadth of the rings and length of the
chambers increasing in size from the centre towards the periphery. Most
part of chambers twice as long as broad. Pores irregular, of very different
sizes, two to three on the breadth of each ring.

_Dimensions._--Diameter of the disk (with six rings) 0.16; breadth of the
rings 0.012 to 0.016; pores 0.002 to 0.006.

_Habitat._--Mediterranean (Messina), surface.

16. _Porodiscus bilix_, Haeckel.

_Discospira bilix_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 113, Taf. vi.
fig. 3.

All rings of the disk spirally convoluted around the large elliptical
central chamber. Spiral convolutions of the inner rings simple, of the
outer rings double; the latter twice as broad as the {497}former. Pores
regular, circular, in the inner rings one pore on the breadth, in the outer
rings two to three pores. Radial beams piercing, numerous, on the margin
prominent.

_Dimensions._--Diameter of the disk (with six rings) 0.14; breadth of the
inner rings 0.007, of the outer 0.013; pores 0.0017.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Stöhr.

17. _Porodiscus bispiralis_, Haeckel.

_Stylodictya bispiralis_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 160. Taf. xxiv. fig. 1.

All rings of the disk spirally convoluted, in a perfect double spiral, with
increasing breadth of the rings; the fourth ring twice as broad as the
first. Pores regular, circular; in the inner rings one pore on the breadth,
in the outer rings two to three pores. Radial beams piercing, numerous,
prominent on the margin.

_Dimensions._--Diameter of the disk (with four rings) 0.1; breadth of the
inner rings 0.006, of the outer 0.012; pores 0.0015.

_Habitat._--Fossil in the rocks of Barbados; living in the depths of the
Central Pacific, Station 268, depth 2900 fathoms.

18. _Porodiscus duplex_, Haeckel.

_Discospira duplex_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 114, Taf.
vi. fig. 4.

All rings of the disk spirally convoluted, in a perfect double spiral, with
increasing breadth of the broad rings. Pores regular, circular; in the
inner rings two to three, in the outer four to five on the breadth. Radial
beams interrupted. Margin of the rings thorny.

_Dimensions._--Diameter of the disk (with three rings) 0.15; breadth of the
rings 0.02 to 0.04; pores 0.007.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

19. _Porodiscus semispiralis_, n. sp. (Pl. 41, fig. 4).

All rings of the disk spirally convoluted, divided by four radial,
perpendicularly crossed and zigzag shaped beams into four quadrants; the
quarter rings of each quadrant halving the rings of each adjacent quarter.
Half spiral line often irregular or partly interrupted. Breadth of all
rings nearly equal. Pores irregular, roundish, two on the breadth of each
ring. Margin of the disk dentated.

_Dimensions._--Diameter of the disk (with nine rings) 0.22; breadth of each
ring 0.012; pores 0.002 to 0.006.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Subgenus 5. _Atactodiscus_, Haeckel, 1881, Prodromus, p. 459.

_Definition._--Rings of the disk more or less irregular, partly concentric,
partly spirally convoluted, often interrupted.

{498}20. _Porodiscus deformis_, Haeckel.

_Discospira deformis_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 114, Taf.
vi. fig. 6.

Rings of the disk irregular, partly concentric, partly spiral, often
interrupted, increasing in breadth from the centre. Radial beams not
piercing; pores regular, circular, two to six on the breadth of the
different rings. (Very variable and irregular, sometimes more spiral, at
other times more concentric, but always with equal pores.)

_Dimensions._--Diameter of the disk (with four rings) 0.18; breadth of the
first ring 0.014, of the second 0.03, of the fourth 0.06; pores 0.005.

_Habitat._--Fossil in the Tertiary rocks of Barbados and Sicily; living in
the Central Pacific, Stations 266 to 268, depths 2700 to 2900 fathoms.

21. _Porodiscus irregularis_, n. sp. (Pl. 41, fig. 7).

_Atactodiscus irregularis_, Haeckel, 1881, Prodromus, p. 459.

_Perispongidium irregulare_, Haeckel, 1878, MS. et Atlas (pl. xli. fig.
7).

Rings of the disk irregular, partly concentric, partly spiral, often
interrupted, with nearly equal breadth. Radial beams not piercing; pores
irregular, roundish, two to four on the breadth of each ring; network in
the periphery of the disk spongy. (Very variable and irregular, sometimes
more spiral, at other times more concentric; disk in the peripheral part
often more or less spongy. Differs from the preceding by the equal breadth
of the rings and the different size of the pores.)

_Dimensions._--Diameter of the disk (with six rings) 0.15; breadth of each
ring 0.011; pores 0.002 to 0.004.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface; also
fossil in Barbados.

Genus 215. _Perichlamydium_,[254] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Porodiscida# with a simple circular disk (without radial
spines and chambered arms), surrounded on the margin by a thin porous (but
not chambered) equatorial girdle.

The genus _Perichlamydium_ differs from _Porodiscus_ only in the
development of a thin, porous, equatorial girdle, which surrounds the
circular margin of the chambered disk. This girdle lies in the equatorial
plane of the lenticular disk, and represents a very delicate siliceous
plate, perforated by numerous small pores. Sometimes the proximal part of
the girdle is ribbed by thin radial beams, the distal prolongations of the
radial rods of the central disk. If these ribs reach the margin of the
girdle and are prominent over it, _Perichlamydium_ passes over into
_Stylochlamydium_.

{499}1. _Perichlamydium praetextum_, Ehrenberg.

_Perichlamydium praetextum_, Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 43; Mikrogeol., 1854, Taf. xxii. fig. 21 (non
20).

_Perichlamydium praetextum_, Haeckel, 1862, Monogr. d. Radiol., p. 495.

All rings of the disk (three to four) concentric, circular, of equal
breadth, with interrupted (not piercing) radial beams. Equatorial girdle
without radial beams, nearly as broad as the disk; its circular pores of
the same size as those of the disk; about two pores on the breadth of each
ring.

_Dimensions._--Diameter of the disk (without the girdle, with four rings)
0.11; breadth of each ring 0.012; breadth of the girdle 0.06 to 0.1; pores
0.004.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, surface; also fossil
in the Tertiary rocks of Barbados and Sicily.

2. _Perichlamydium saturnus_, n. sp. (Pl. 41, fig. 5).

All rings of the disk (three to four) concentric, circular (sometimes
partly concentric, circular, partly spiral, or irregular), with interrupted
(not piercing) radial beams. Equatorial girdle without radial beams, about
half as broad as the disk; its circular pores very small, scarcely half as
large as those of the disk; about three pores on the breadth of each ring.
(Very variable in the ring-form, differs from the preceding in the small
pores of the girdle.)

_Dimensions._--Diameter of the disk (without the girdle, with three rings)
0.11; breadth of each ring 0.02, pores 0.004, breadth of the girdle 0.05,
pores 0.002.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

3. _Perichlamydium accrescens_, Haeckel.

_Discospira accrescens_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 114,
Taf. vi. fig. 5.

All rings of the disk (six to seven) not concentric, convoluted in a simple
spiral, of nearly equal breadth, with interrupted (not piercing) radial
beams. Equatorial girdle in the proximal part with numerous radial beams,
which do not reach its margin; its pores half as large as those of the
disk, where one to two pores arise on the breadth of each ring. (The girdle
becomes twice to three times as broad as in the figure of Stöhr.)

_Dimensions._--Diameter of the disk (with seven rings, without the girdle)
0.13; breadth of each ring 0.007 to 0.01; pores 0.0036; breadth of the
girdle 0.05, pores of it 0.0017.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte (Stöhr),
Caltanisetta (Haeckel); living in the Central Pacific, Station 266, depth
2750 fathoms.

4. _Perichlamydium spirale_, Ehrenberg.

_Perichlamydium spirale_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 80, Taf. xxii. fig. 12.

All rings of the disk (three to four) not concentric, convoluted in a
simple spiral, of increasing breadth from the centre; connected by numerous
piercing radial beams. Equatorial girdle about {500}half as broad as the
disk, without radial beams; its pores of the same size as those of the
disk, regular, circular; three pores on the breadth of the first ring, six
pores of the fourth ring.

_Dimensions._--Diameter of the disk (with four rings, without the girdle)
0.12; breadth of the first ring 0.01, of the fourth 0.02; breadth of the
girdle 0.05; pores 0.003.

_Habitat._--Fossil in the rocks of Barbados.

Subfamily 3. OMMATODISCIDA, Stöhr, 1880, Palæontographica, vol. xxvi. p.
115.

_Definition._--#Porodiscida# without radial appendages of the
concentrically annulated disk, but distinguished by one single or two
opposite large marginal oscula, or wide openings on the margin of the disk,
armed with a coronet of spines.

Genus 216. _Ommatodiscus_,[255] Stöhr, 1880, Palæontographica, vol. xxvi.
p. 115.

_Definition._--Porodiscida without chambered arms and radial spines on the
margin of the circular or elliptical disk, but with one large marginal
osculum or opening surrounded by a coronet of spines.

The genus _Ommatodiscus_, together with the following _Stomatodiscus_, form
the peculiar small subfamily of Ommatodiscida, founded by Stöhr in 1880
(_loc. cit._). These remarkable Porodiscida, very nearly allied to
_Porodiscus_, are distinguished by one or two large openings on the margin
of the disk, and these "marginal oscula" are constantly armed with a
coronet of spines (comparable to the osculum coronatum of _Sycon_ in the
Calcispongiæ). Probably in the living Ommatodiscida the osculum is the door
from which a "sarcode-flagellum" issues (comp. above, p. 407). Perhaps this
osculum is comparable to the peculiar coronet of spines which is developed
on one pole of the shell axis in some Ellipsida (_Lithomespilus_,
_Lithapium_). The internal structure of the disk is commonly more or less
obscure, as the lenticular disk is much thickened, sometimes nearly
ellipsoidal. It is possible that the Ommatodiscida are more nearly related
to the Lithelida than to the Porodiscida; but there is no indication of an
internal trizonal medullary shell. Also the apparent resemblance to the
Cyrtida is of no morphological value; both groups are of quite independent
phylogenetic origin.

Subgenus 1. _Ommatodiscinus_, Haeckel.

_Definition._--Disk circular.

1. _Ommatodiscus decipiens_, Stöhr.

_Ommatodiscus decipiens_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 115,
Taf. vi. figs. 8, 8_a._

Disk circular, with two chambered rings around the spherical central
chamber, of equal breadth. Chambers twice as high as broad. Pores very
small, one-third as broad as the bars {501}between them, two on the breadth
of each ring. Osculum of the same breadth as the central chamber,
surrounded by numerous very short teeth.

_Dimensions._--Diameter of the disk 0.12; breadth of the central chamber
and of each ring 0.03; pores 0.0015.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

2. _Ommatodiscus stöhrii_, n. sp.

Disk circular with three chambered rings around the spherical central
chamber, of equal breadth. Chambers broader than high. Pores of the same
breadth as the bars between them, four on the breadth of each ring. Osculum
of the same breadth as the central chamber, surrounded by a corona of ten
to twenty thin, bristle-shaped teeth, as long as its diameter.

_Dimensions._--Diameter of the disk 0.2; breadth of the central chamber and
of each ring 0.03; pores 0.004.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

3. _Ommatodiscus circularis_, n. sp.

Disk circular, with four circular chambered rings around the spherical
central chamber, of increasing breadth; the fourth ring twice as broad as
the second. Chambers about as broad as high. Pores large, twice as broad as
the bars, one to two on the breadth of each ring. Osculum twice as broad as
the central chamber, surrounded by a coronal of strong conical teeth, twice
as long as its diameter.

_Dimensions._--Diameter of the disk 0.18; breadth of the central chamber
and inner rings 0.015, of the outer rings 0.03; pores 0.01.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Subgenus 2. _Ommatodisculus_, Haeckel.

_Definition._--Disk elliptical.

4. _Ommatodiscus haeckelii_, Stöhr.

_Ommatodiscus haeckelii_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 115,
Taf. vi. figs. 7, 7_a._

Disk elliptical (6 : 7), with four chambered rings around the elliptical
central chamber, of equal breadth. Chambers about as high as broad. Pores
small, half as broad as the bars, two on the breadth of each ring. Osculum
three times as broad as the central chamber, surrounded by a crown of
strong conical teeth.

_Dimensions._--Length of the disk 0.18, breadth 0.16; breadth of each ring
and of the central chamber 0.02; pores 0.003.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte (Stöhr),
Caltanisetta (Haeckel).

{502}5. _Ommatodiscus lævigatus_, Stöhr.

_Ommatodiscus lævigatus_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 116,
Taf. vi. figs. 9, 9_a._

Disk elliptical (3 : 4), with three chambered rings around the circular
central chamber, the third ring half as broad as the second. Chambers twice
as high as broad. Pores very small, one-third as broad as the bars. Osculum
twice as broad as the central chamber, armed with a crown of short conical
teeth.

_Dimensions._--Length of the disk 0.15, breadth 0.11; breadth of inner
rings 0.02, of the outer 0.01; pores 0.0017.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

6. _Ommatodiscus fragilis_, Stöhr.

_Ommatodiscus fragilis_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 116,
Taf. vi. figs. 10, 10_a._

Disk elliptical (4 : 5), with five chambered rings around the elliptical
central chamber, the fifth ring twice as broad as each of the others.
Chambers about as high as broad. Pores very small, one-fifth as broad as
the bars. Osculum three times as broad as the central chamber, surrounded
by a coronet of short teeth.

_Dimensions._--Length of the disk 0.17, breadth 0.13; breadth of the inner
rings 0.01, of the outer 0.02; pores 0.001.

_Habitat._--Fossil in Tertiary rocks of Sicily and Barbados; living in
depths of the Tropical Atlantic and Pacific, Station 353, depth 2965
fathoms, Station 265, depth 2900 fathoms, &c.

Genus 217. _Stomatodiscus_,[256] n. gen.

_Definition._--#Porodiscida# without chambered arms and radial spines on
the margin of the circular or elliptical disk, but with two large,
opposite, marginal oscula, or openings surrounded by a coronet of spines.

The genus _Stomatodiscus_ has the same characteristic structure as the
foregoing _Ommatodiscus_, and differs from it only in the duplication of
the large marginal openings. Whilst in the latter there is only one such
marginal osculum, here we find on the margin of the disk two oscula,
opposite on the poles of one equatorial axis.

1. _Stomatodiscus amphistomus_, n. sp.

Disk circular, with three concentric rings of equal breadth around the
central chamber. Pores irregular, roundish, about two on the breadth of
each ring. Surface of the lenticular shell spiny. On two opposite points of
the margin a large osculum, three to four times as broad as the central
chamber, armed with a coronet of strong pyramidal spines of different
length, the longest equal to the radius of the disk.

{503}_Dimensions._--Diameter of the disk (with three rings) 0.12; breadth
of each ring 0.016; pores 0.004.

_Habitat._--South Pacific, Station 302, depth 1450 fathoms.

2. _Stomatodiscus osculatus_, n. sp. (Pl. 48, fig. 8).

Disk elliptical, nearly twice as long as broad, with three concentric rings
around the elliptical central chamber, one piercing radial beam in the main
axis, the other beams interrupted. Surface of the shell with small
scattered thorns. Pores very irregular, roundish, partly aggregated in
groups of four to eight smaller porules. On both poles of the main axis a
large elliptical marginal osculum, about as large as the central chamber,
armed with a coronet of short conical spines.

_Dimensions._--Length of the disk (with three rings) 0.18, breadth 0.1;
pores 0.001 to 0.006.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Subfamily 4. STYLODICTYIDA, Haeckel, 1881, Prodromus, p. 459.

_Definition._--#Porodiscida# with solid radial spines on the margin of the
concentrically annulated disk, situated in the equatorial plane of the disk
(without chambered arms and marginal oscula).

Genus 218. _Xiphodictya_,[257] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with two opposite, solid, radial spines on the
margin of the circular or elliptical disk.

The genus _Xiphodictya_ opens the series of the Stylodictyida or of those
Porodiscida in which the margin of the chambered disk is armed with solid
radial spines, all situated in the equatorial plane of the disk.
_Xiphodictya_ exhibits the minimum number of spines, two being opposite on
the poles of one equatorial axis of the disk. It repeats, therefore, in
this family the same amphistylic formation as _Sethostylus_ in the
Phacodiscida and _Stylocyclia_ in the Coccodiscida.

Subgenus 1. _Xiphodictyon_, Haeckel.

_Definition._--All rings of the disk concentric, circular.

1. _Xiphodictya amphibelonia_, n. sp. (Pl. 42, fig. 10).

All rings of the disk concentric, circular, of equal breadth. Pores
irregular, roundish, one and a half to two on the breadth of each ring.
Margin of the disk thorny, of the same thickness as {504}the central part
of the medal-shaped or cylindrical disk. Two opposite radial spines very
long and thin, cylindrical, twice to three times as long as the diameter of
the disk, only half as thick as the breadth of one ring.

_Dimensions._--Diameter of the disk (with five rings) 0.17; breadth of each
ring 0.014; pores 0.003 to 0.012.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

2. _Xiphodictya amphirrhopalia_, n. sp. (Pl. 42, fig. 11).

All rings of the disk concentric, circular, with increasing breadth from
the centre; the fifth ring twice as broad as the second. Central chamber
very large. Margin of the lenticular disk thorny, much thinner than the
central part. Pores irregular, roundish, two to three on the breadth of
each ring. Two opposite radial spines club-shaped, about as long as the
diameter of the disk, in the outer spindle-shaped part three to four times
as broad as at the base.

_Dimensions._--Diameter of the disk (with six rings) 0.17; breadth of the
second ring 0.008, of the fifth ring 0.016.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms; also
fossil in the Tertiary rocks of Sicily (Caltanisetta).

Subgenus 2. _Xiphospira_, Haeckel.

_Definition._--All rings of the disk or a part of them not concentric,
spirally convoluted; sometimes irregular or interrupted.

3. _Xiphodictya staurospira_, n. sp. (Pl. 42, fig. 12).

All rings of the disk not concentric, half spiral. Four radial beams,
crossed perpendicularly and zigzag-shaped, divide each ring into four
quadrants; the quarter-ring of each quadrant halves the two adjacent rings.
All rings of equal breadth. Pores irregular, roundish, two on the breadth
of each ring. Two opposite radial spines pyramidal, somewhat shorter than
the diameter of the disk.

_Dimensions._--Diameter of the disk (with six rings) 0.16; breadth of each
ring: 0.014; pores 0.003 to 0.007.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

4. _Xiphodictya heliospira_, n. sp.

All rings of the disk not concentric, convoluted in a simple, regular,
spiral line; all nearly of equal breadth. Pores subregular, circular, two
on the breadth of each ring. Two opposite radial spines conical, about as
long as the radius of the disk. Margin of the disk thorny. (Differs from
_Stylodictya heliospira_, Pl. 41, fig. 8, mainly by the two strong,
opposite, radial spines.)

{505}_Dimensions._--Diameter of the disk (with six rings) 0.14; breadth of
each ring 0.01 to 0.012; pores 0.002 to 0.006.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Genus 219. _Tripodictya_,[258] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with three solid, equidistant, radial spines
on the margin of the circular or triangular disk.

The genus _Tripodictya_ exhibits three radial spines on the margin of the
disk, divergent at equal angles; rarely in some specimens the angles differ
more or less. Perhaps this genus bears a near relation to the Dictyastrida
(or to the Euchitonida with three chambered arms--_Dictyastrum_,
_Euchitonia_, &c.).

1. _Tripodictya trigonaria_, n. sp. (Pl. 42, fig. 8).

All rings of the disk concentric, triangular, with three equal convex
sides, all of the same breadth; first and second rings with three simple
chambers, third and fourth rings with six chambers, fifth ring with twelve
chambers. Central chamber also equilateral triangular, from its three
corners arise three piercing perradial beams; from the second ring arise
three interradial beams, alternate with the latter; from the fourth ring
between these and the former arise six adradial beams. Pores subregular,
two on the breadth of each ring. Three marginal spines pyramidal, as long
as the radius of the disk and as broad as one ring.

_Dimensions._--Diameter of the disk (with five rings) 0.11; breadth of each
ring 0.011; pores 0.004.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

2. _Tripodictya triacantha_, n. sp. (Pl. 42, fig. 7).

All rings of the disk concentric, circular, of the some breadth, connected
by very numerous irregular radial beams, the number of which increases
towards the periphery. Pores irregular, roundish, two on the breadth of
each ring. Three marginal spines spindle-shaped, as long as the radius of
the disk, and in the middle part as broad as one ring.

_Dimensions._--Diameter of the disk (with eight rings) 0.2; breadth of each
ring 0.012; pores 0.004.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

3. _Tripodictya tribelonia_, n. sp. (Pl. 42, fig. 9).

All rings of the disk, or a part of them, not concentric, spirally
convoluted, of equal breadth, connected by numerous irregular, interrupted
radial beams, the number of which increases from the centre. (In one
marginal view of the disk, Pl. 42, fig. 9, the disk seemed to be composed
of seven {506}parallel chambered plates in the central part, and five
similar plates in the peripheral part.) Pores irregular, roundish, three on
the breadth of each ring. Three marginal spines long and thin, cylindrical
(as in _Staurodictya cruciata_, Pl. 42, fig, 4), longer than the diameter
of the disk, and half as thick as one ring.

_Dimensions._--Diameter of the disk (with nine rings) 0.18; breadth of each
ring 0.01; pores 0.002 to 0.004.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

Genus 220. _Staurodictya_,[259] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with four solid radial spines (commonly
crossed at right angles) on the margin of the circular or quadrangular
disk.

The genus _Staurodictya_ is characterised by four marginal spines, lying
opposite in pairs in two crossed equatorial diameters of the disk,
perpendicular one to another. Sometimes the regular rectangular position of
the spines becomes more or less irregular. Commonly the marginal spines are
the extreme prolongations of four internal crossed radial beams, which are
either rectilinear or zigzag-shaped. In the latter case the concentric
disposition of the circular rings becomes more or less spiral. Perhaps
_Staurodictya_ has a near relation to the Stauralastrida (or to the
Euchitonida with four crossed chambered arms--_Stauralastrum_,
_Histiastrum_, &c.).

Subgenus 1. _Staurodictyon_, Haeckel.

_Definition._--All rings of the disk concentric (commonly circular,
sometimes with four incisions, produced by two crossed constrictions, or
nearly square).

1. _Staurodictya medusa_, n. sp. (Pl. 42, fig. 3).

All rings of the disk concentric, of nearly equal breadth, divided into
four quadrants by two perradial constrictions (perpendicular one to
another). Pores irregular, roundish, two to three on the breadth of each
ring. Four marginal spines strong, pyramidal, nearly as long as the radius
of the disk. Margin between them denticulated.

_Dimensions._--Diameter of the disk (with four rings) 0.13; breadth of each
ring 0.015; pores 0.002 to 0.01.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

2. _Staurodictya ciliata_, n. sp. (Pl. 42, fig. 2).

All rings of the disk concentric, of nearly equal breadth, circular or
roundish, without perradial constrictions. Pores subregular, circular,
three to four on the breadth of each ring. Four marginal {507}spines short,
pyramidal, about as long as the breadth of one ring. Margin between them
ciliated, with thin radial bristles.

_Dimensions._--Diameter of the disk (with four rings) 0.13; breadth of each
ring 0.015; pores 0.0025.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

3. _Staurodictya elegans_, n. sp. (Pl. 42, fig. 1).

All rings of the disk concentric, circular, or roundish, of increasing
breadth towards the margin; the fifth ring twice as broad as the first.
Pores irregular, roundish, two to three on the breadth of each ring. Four
marginal spines very large, conical, with a thin pedicle at the base,
cancellated by ten to twelve deep furrows, about as long as the radius of
the disk and three times as long as broad at the base (above the pedicle).
Margin between them ciliated, with numerous short radial spines.

_Dimensions._--Diameter of the disk (with six rings) 0.14; breadth of the
inner rings 0.006, of the outer 0.012; pores 0.002 to 0.008.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

4. _Staurodictya quadrispina_, Haeckel.

_Stylodictya quadrispina_, Haeckel, 1862, Monogr. d. Radiol., p. 496,
Taf. xxix. fig. 4.

All rings of the disk concentric, circular, of increasing breadth towards
the margin; the fourth ring twice as broad as the second. Pores irregular,
roundish, two to three on the breadth of each ring. Four marginal spines
short and thin, conical, about as long as the breadth of one ring. Margin
between them smooth.

_Dimensions._--Diameter of the disk (with six rings) 0.12; breadth of the
inner rings 0.006, of the outer rings 0.012; pores 0.001 to 0.006.

_Habitat._--Mediterranean, Atlantic (Canary Islands), surface.

Subgenus 2. _Staurospira_, Haeckel.

_Definition._--All rings of the disk or a part of them not concentric,
spirally convoluted; spiral line simple or double, sometimes half or
irregular, interrupted.

5. _Staurodictya cruciata_, n. sp. (Pl. 42, figs. 4, 5).

All rings of the disk nearly of the same breadth, not concentric,
half-spiral, interrupted by four zigzag-shaped radial beams crossed in two
diameters perpendicular one to another. The quarter ring of each quadrant
halving both neighbouring quarters. Pores irregular, roundish, two on the
breadth of each ring. Pores of the outermost (eighth) ring much smaller
than the others. Four marginal spines cylindrical or nearly spindle-shaped,
thick, about as long as the radius of the disk. Margin between them smooth.

_Dimensions._--Diameter of the disk (with eight rings) 0.2; breadth of each
ring 0.011; pores 0.004.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

{508}6. _Staurodictya splendens_, Haeckel.

_Stylodictya splendens_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiii. fig. 9.

All rings of the disk nearly of the same breadth, not concentric,
half-spiral, interrupted by four zigzag-shaped perradial beams, crossed in
two diameters. The quarter ring of each quadrant halving both neighbouring
quarters. Pores regular, circular, only one single on the breadth of each
ring. Four marginal spines conical or spear-shaped, about half as long as
the radius of the disk. Margin between them ciliated, with short
bristle-shaped radial spines.

_Dimensions._--Diameter of the disk (with eight rings) 0.2; breadth of each
ring 0.012; pores 0.004.

_Habitat._--Fossil in the rocks of Barbados.

7. _Staurodictya grandis_, n. sp.

All rings of the disk (twelve to sixteen) of the same breadth, not
concentric, irregular, partly spiral, interrupted by irregular turnings,
and by ramified radial beams, which divide each ring into numerous square
chambers. Pores subregular, circular, only one single on the breadth of
each ring (and on each chamber). Four marginal spines short and stout,
conical, twice as long as broad at the base, four to five times as long as
the breadth of one ring. Margin between them dentated.

_Dimensions._--Diameter of the disk (with sixteen rings) 0.3; breadth of
each ring 0.01; pores 0.005.

_Habitat._--Pacific, central area, Stations 265 to 268, depth 2900 fathoms;
also fossil in the Tertiary rocks of Barbados.

8. _Staurodictya ocellata_, Haeckel.

_Stylodictya ocellata_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiii. fig. 7.

All rings of the disk irregular, not concentric, half-spiral, interrupted
by four zigzag-shaped, crossed, radial beams; the quarter ring of each
quadrant halving both neighbouring quarters. Breadth of the rings
increasing towards the periphery; the fourth ring twice as broad as the
second. Pores irregular, with increasing size from the centre, three on the
breadth of each ring. Four marginal spines thick and long, cylindrical;
margin between them smooth.

_Dimensions._--Diameter of the disk (with four rings) 0.1; breadth of the
inner rings 0.01, of the outer 0.02; pores 0.002 to 0.006.

_Habitat._--Fossil in the rocks of Barbados.

_Stylodictya haeckelii_, Zittel, 1876, L. N. 29, p. 85, Taf. ii. fig. 9, is
a fossil Cretaceous species, related to _Stylodictya multispina_.

{509}Genus 221. _Stylodictya_,[260] Ehrenberg, 1847, Monatsber. d. k.
preuss. Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Porodiscida# with numerous (five or more, commonly eight to
twelve) solid radial spines, regularly or irregularly disposed on the
margin of the circular or polygonal disk; margin simple, without a porous
equatorial girdle.

The genus _Stylodictya_ comprises the majority of this subfamily, in which
the number of the marginal spines exceeds four. Commonly we find eight to
twelve spines, more or less regularly disposed (four perradial and four
interradial, or four perradial and eight adradial); but often also the
number and disposition become irregular (sometimes very large). In my
Monograph (1862 pp. 495, 515) I had separated the concentric disks with
closed circular rings (as true _Stylodictya_, s. str.) from the spiral
disks with convoluted rings (_Stylospira_). But I retain these two groups
here only as two subgenera, as intermediate forms between them are very
common, and often a part of the disk concentric, a part spiral (compare
above, p. 492).

Subgenus 1. _Stylodictyon_, Haeckel, 1862, Monogr. d. Radiol., p. 495.

_Definition._--All rings of the disk concentric, commonly circular (rarely
a little elliptical or polygonal).

1. _Stylodictya gracilis_, Ehrenberg.

_Stylodictya gracilis_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvi. fig. 28.

_Stylodictya gracilis_, Ehrenberg, 1873, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 257; Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, Taf.
xxiii. fig. 3.

_Stylodictya gracilis_, Haeckel, 1862, Monogr. d. Radiol., p. 499.

All rings of the disk concentric, circular, of equal breadth (the first
ring sometimes, but not constantly, four-lobed). Pores regular, circular,
small, three on the breadth of each ring. Four perradial beams (crossed in
two perpendicular diameters) beginning from the circular central chamber,
four interradial beams from the first or second ring (sometimes others
between them). Beams prolonged into eight to twelve (or more) marginal
spines, bristle-shaped, as long as the radius of the shell.

_Dimensions._--Diameter of the disk (with four rings) 0.12; breadth of each
ring 0.013; pores 0.0025.

_Habitat._--Fossil in Tertiary rocks of Barbados and Nicobar; living in the
depths of the Pacific and Atlantic.

{510}2. _Stylodictya multispina_, Haeckel.

_Stylodictya multispina_, Haeckel, 1862, Monogr. d. Radiol., p. 496, Taf.
xxix. fig. 5.

_Stylodictya forbesii_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 160, Taf. xxiii. fig. 6.

All rings of the disk concentric, circular, of equal breadth. Pores
regular, circular, two and a half to three on the breadth of each ring.
Eight to twelve radial beams beginning from the central chamber, others
from the inner rings. Commonly from the third or fourth ring arise
twenty-four to thirty (sometimes forty or more) piercing beams, which are
prolonged at the margin into bristle-shaped radial spines, as long as the
breadth of two to four rings.

_Dimensions._--Diameter of the disk (with seven rings) 0.2; breadth of each
ring 0.013; pores 0.004.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific, surface
and various depths.

3. _Stylodictya hastata_, Ehrenberg.

_Stylodictya hastata_, Ehrenberg, 1873, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 257; Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, Taf.
xxiii. fig. 5.

_Stylodictya gracilis_, Bury, 1862, Polycystins of Barbados, pl. ii. fig.
1.

All rings of the disk concentric, circular, of equal breadth. Pores
regular, circular, very small, two on the breadth of each ring. Eight to
twelve marginal spines very large, spear-shaped, sulcated, pyramidal,
nearly as long as the diameter of the disk, with a thin pedicle at the
base, above this as thick as the breadth of one ring.

_Dimensions._--Diameter of the disk (with five rings) 0.13; breadth of each
ring 0.012; pores 0.003.

_Habitat._--Fossil in the rocks of Barbados.

4. _Stylodictya stellata_, Bailey.

_Stylodictya stellata_, Bailey, 1856, Amer. Journ., vol. xxii. p. 6, pl.
i. fig. 20.

_Stylodictya stellata_, Haeckel, 1862, Monogr. d. Radiol., p. 499.

All rings of the disk concentric, circular, of equal breadth. Pores
regular, circular, very small, two on the breadth of each ring. Marginal
spines fourteen (probably variable in number, twelve to sixteen), very
thick and short, conical; their length seems to be equal to their basal
breadth and to the breadth of one ring. Related to _Stylochlamydium_?

_Dimensions._--Diameter of the disk (with five rings) 0.11; breadth of each
ring 0.01; pores 0.003.

_Habitat._--North Pacific, depths of the Kamtschatka Sea, Bailey; Station
241, depth 2300 fathoms.

5. _Stylodictya arachnia_, Haeckel.

_Stylodictya arachnia_, Haeckel, 1862, Monogr. d. Radiol., p. 497.

_Stylocyclia arachnia_, J. Müller, 1856, p. 492; Abhandl. d. k. Akad. d.
Wiss. Berlin, 1858, p. 41, Taf. i. figs. 8, 9.

All rings of the disk concentric, circular, or polygonal; their breadth
increases from the centre towards the periphery, so that the fourth or
fifth ring is twice as broad as the second. Pores {511}subregular,
circular, two on the breadth of each ring. Radial beams partly piercing.
Marginal spines eight to sixteen (commonly twelve), bristle-shaped, very
thin, once to three times as long as the diameter of the disk. (On the
numerous varieties of this common species compare my Monograph, 1862, p.
498.)

_Dimensions._--Diameter of the disk (with ten rings) 0.22; breadth of the
inner rings 0.004 to 0.008, of the outer 0.012 to 0.015; pores 0.003 to
0.005.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

6. _Stylodictya solmaris_, n. sp.

All rings of the disk concentric, polygonal, with eight to sixteen rounded
corners; their breadth increases from the centre; eighth ring twice as
broad as the second. Pores subregular circular, large, only one pore on the
breadth of each ring. Twenty to thirty marginal spines, bristle-shaped,
undulating, about as long as (or longer than) the diameter of the disk.

_Dimensions._--Diameter of the disk (with eight rings) 0.2; breadth of the
inner rings 0.005, of the outer 0.012; pores 0.004 to 0.008.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

7. _Stylodictya octogonia_, n. sp.

All rings of the disk concentric, increasing in breadth from the centre.
The outer rings (five to eight) regular, octogonal, twice as broad as the
circular inner rings. Pores subregular, circular, two to three on the
breadth of each ring. Eight piercing perradial spines (alternating with the
eight corners of the octogonal rings) bristle-shaped, longer than the
diameter of the disk.

_Dimensions._--Diameter of the disk (with eight rings) 0.24; breadth of the
inner rings 0.006, of the outer 0.014; pores 0.005 to 0.01.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

Subgenus 2. _Stylodictula_, Haeckel.

_Definition._--Rings of the disk partly concentric, completely annular;
partly spiral or interrupted, often irregular.

8. _Stylodictya perispira_, n. sp.

Inner rings of the disk (two to four) concentric, circular, or roundish,
outer rings forming a simple spiral, breadth a little increasing from the
centre. Pores subregular, circular, about two on the breadth of each ring.
Marginal spines bristle-shaped, ten to fifteen, about as long as the
diameter of the disk.

_Dimensions._--Diameter of the disk (with eight rings) 0.2; breadth of each
ring 0.013; pores 0.004.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

{512}9. _Stylodictya centrospira_, n. sp. (Pl. 41, fig. 9).

Inner rings of the disk (three or four) convoluted in a simple or double
spiral, often interrupted or irregular; outer rings (two to three)
concentric, circular, or roundish. Breadth of the rings variable,
irregular. Pores irregular, of very different sizes, in the outer
concentric part twice to four times as large as in the inner spiral part.
Radial beams partly interrupted, partly piercing. Marginal spines fifty to
eighty, very variable in size and number, commonly fifteen to twenty strong
conical spines, twice to three times as long as the ring-breadth, and
numerous (thirty to sixty) smaller spines. Very variable.

_Dimensions._--Diameter of the disk (with seven rings) 0.2; breadth of the
rings 0.01 to 0.02; pores 0.002 to 0.01.

_Habitat._--Pacific, central area, Stations 263 to 274, depth 2350 to 2925
fathoms.

10. _Stylodictya setigera_, Ehrenberg.

_Stylodictya setigera_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiii. fig. 4.

Inner rings of the disk (two to three) convoluted in a simple or double
spiral, outer rings (two to three) concentric, circular, or roundish.
Breadth of the rings nearly equal. Pores regular, circular, three on the
breadth of each ring. Marginal spines twenty to forty, bristle-shaped,
about twice as long as the ring-breadth. (The specimen figured by Ehrenberg
was a young one; in older specimens I found the inner spiral disk
surrounded by two to three concentric circular rings.)

_Dimensions._--Diameter of the disk (with six rings) 0.16; breadth of the
rings 0.12; pores 0.003.

_Habitat._--Fossil in the rocks of Barbados; also living in the depths of
the Central Pacific, Station 266, depth 2750 fathoms.

Subgenus 3. _Stylospira_, Haeckel, 1862, Monogr. d. Radiol., p. 515.

_Definition._--All rings of the disk convoluted in a simple or double
spiral (sometimes also in quartered half-spirals).

11. _Stylodictya heliospira_, n. sp. (Pl. 41, fig. 8).

_Stylospira heliospira_, Haeckel, 1879, MS.

All rings of the disk convoluted in a simple regular spiral, increasing in
breadth from the centre towards the periphery; the sixth ring twice as
broad as the second. Pores irregular, roundish, two on the breadth of each
ring. Marginal spines numerous, thirty to forty, bristle-shaped, nearly
half as long as the radius of the disk.

_Dimensions._--Diameter of the disk (with seven rings) 0.16; breadth of the
inner rings 0.008, of the outer 0.016; pores 0.002 to 0.006.

_Habitat._--Mediterranean, Portofino near Genoa, surface; also fossil in
the Tertiary rocks of Sicily; Caltanisetta, Haeckel.

{513}12. _Stylodictya hertwigii_, Haeckel.

_Stylospira arachnia_, R. Hertwig, 1879, Organismus d. Radiol., p. 59,
Taf. vi. fig. 8.

All rings of the disk convoluted in a simple regular spiral, with
increasing breadth from the centre; the fifth ring twice as broad as the
second. Pores regular, circular, two on the breadth of each ring. Twelve
piercing radial beams and some others interrupted, prolonged into twelve to
twenty radial marginal spines, bristle-shaped, about as long as the
diameter of the disk.

_Dimensions._--Diameter of the disk (with five rings) 0.15; breadth of the
second ring 0.01, of the fifth 0.02; pores 0.006.

_Habitat._--Mediterranean (Messina), R. Hertwig.

13. _Stylodictya dujardinii_, Haeckel.

_Stylospira dujardinii_, Haeckel, 1862, Monogr. d. Radiol., p. 515, Taf.
xxix. figs. 9, 10.

All rings of the disk convoluted in a simple regular spiral, of nearly
equal breadth. Pores regular, circular, two on the breadth of each ring.
Very numerous (twenty to thirty or more) piercing radial beams, prolonged
into bristle-shaped marginal spines, about as long as the diameter of the
disk.

_Dimensions._--Diameter of the disk (with six rings) 0.12; breadth of each
ring 0.01; pores 0.004.

_Habitat._--Mediterranean (Messina), Haeckel.

14. _Stylodictya echinastrum_, Ehrenberg.

_Stylodictya echinastrum_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiii. fig. 1.

All rings of the disk convoluted in a double spiral, of nearly equal
breadth. Pores irregular, roundish, two to three on the breadth of each
ring. Marginal spines numerous, twenty to thirty, of very different size,
the largest conical, strong, about as long as the diameter of the disk, and
on the base as broad as one ring.

_Dimensions._--Diameter of the disk (with five rings) 0.12; breadth of each
ring 0.01; pores 0.002 to 0.006.

_Habitat._--Fossil in the rocks of Barbados.

15. _Stylodictya clavata_, Ehrenberg.

_Stylodictya clavata_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 84, Taf. xxiii. fig. 2.

All rings of the disk convoluted in a half spiral, of nearly equal breadth;
each ring by four zigzag beams (crossed in two perpendicular diameters)
divided into four equal quarters; the spiral line of each quarter ring
halving both neighbouring quarters. Pores regular, circular, two on the
breadth of each ring. Eight marginal spines short, conical, with thin
pedicle; four perradial (as {514}prolongations of the four internal beams)
alternating with four interradial spines arising from the margin of the
disk.

_Dimensions._--Diameter of the disk (with five rings) 0.13; breadth of each
ring 0.014; pores 0.003.

_Habitat._--Fossil in the rocks of Barbados.

Genus 222. _Stylochlamydium_,[261] Haeckel, 1881, Prodromus, p. 460.

The genus _Stylochlamydium_ is intermediate between _Perichlamydium_ (with
which it was formerly united) and _Stylodictya_. It deals with the former
in the peculiar equatorial girdle, with the latter in the radial spines of
the disk margin, which pierce the girdle. To both these genera it shows
slow transitions, and can hardly be subjected to a sharp definition.

Subgenus 1. _Stylochlamys_, Haeckel.

_Definition._--All rings of the disk concentric, circular (or somewhat
polygonal).

1. _Stylochlamydium asteriscus_, n. sp. (Pl. 41, fig. 10).

_Perichlamydium asteriscus_, Haeckel, 1879, MS.

All rings of the disk concentric, circular, or polygonal, with increasing
breadth from the centre; the fifth ring twice as broad as the second. Pores
irregular, roundish, three to four on the breadth of each ring; in the
rings two to six times as large as in the equatorial girdle, which is half
as broad or two-thirds as broad as the radius of the disk. Twelve
bristle-shaped radial spines are connected by the girdle near to the
points; four crossed spines arising from the central chamber; two others
between these in each quadrant arising from the first ring.

_Dimensions._--Diameter of the disk (with five rings) 0.15, with the girdle
0.25; breadth of the second ring 0.01, of the fifth ring 0.02; pores 0.001
to 0.005.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Stylochlamydium limbatum_, Haeckel.

_Perichlamydium limbatum_, Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 43; Mikrogeol., 1854, Taf. xxii. fig. 20.

_Perichlamydium limbatum_, Haeckel, 1862, Monogr. d. Radiol., p. 494.

All rings of the disk concentric, circular, of equal breadth. Pores
regular, circular, two on the breadth of each ring; twice to three times as
large as the fine pores of the equatorial girdle, which is {515}about half
as broad as the radius of the disk. Twelve (or eleven) bristle-shaped
radial spines are connected by the girdle near to the points, irregularly
disposed.

_Dimensions._--Diameter of the disk (with five rings) 0.12, with the girdle
0.2, breadth of each ring 0.01; pores 0.001 to 0.003.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Caltanisetta, Grotte.

3. _Stylochlamydium venustum_, Haeckel.

_Perichlamydium venustum_, Bailey, 1856, Amer. Journ., vol. xxii. p. 5,
pl. i. figs. 16, 17.

_Perichlamydium venustum_, Haeckel, 1862, Monogr. d. Radiol., p. 495.

All rings of the disk concentric, circular, of equal breadth. Pores
regular, circular, everywhere of nearly equal size, three on the breadth of
each ring, a little smaller in the equatorial girdle, which is about as
broad as the radius of the disk. Twenty to twenty-four bristle-shaped
radial spines, irregularly disposed, proceed with their free points over
the margin.

_Dimensions._--Diameter of the disk (with four rings) 0.1, with the girdle
0.3; breadth of each ring 0.011; pores 0.002.

_Habitat._--North Pacific, Kamtschatka, Bailey.

4. _Stylochlamydium æquale_, Haeckel.

_Perichlamydium æquale_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 109, pl.
v. fig. 2.

All rings of the disk concentric, circular, with increasing breadth from
the centre; the sixth ring twice as broad as the second. Pores regular,
circular, everywhere of equal size; on the breadth of the inner rings one,
of the outer two, of the girdle three pores. Girdle only one-fourth as
broad as the radius of the disk. About twenty bristle-shaped radial spines,
irregularly disposed, are connected by the girdle near to the points.

_Dimensions._--Diameter of the disk (with six rings) 0.11, with the girdle
0.17; breadth of the second ring 0.006, of the sixth 0.013; pores 0.006.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

Subgenus 2. _Stylochlamyum_, Haeckel.

_Definition._--Rings of the disk all (or in part) not concentric, spirally
convoluted or irregular.

5. _Stylochlamydium perispirale_, Haeckel.

_Perichlamydium limbatum_, var. Stöhr, 1880, Palæontogr., vol. xxvi. p.
109, Taf. v. fig. 1.

Inner rings of the disk concentric, circular, outer rings convoluted
spirally, all rings of equal breadth. Pores regular, circular, two on the
breadth of each ring, twice as large as in the equatorial {516}girdle,
which is about half as broad as the radius of the disk. Twelve to sixteen
bristle-shaped radial spines, irregularly disposed, are connected by the
girdle near to the points.

_Dimensions._--Diameter of the disk (with six rings) 0.12, with the girdle
0.2; breadth of each ring 0.011; pores in the central disk 0.004, in the
girdle 0.002.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

6. _Stylochlamydium spongiosum_, Haeckel.

_Perichlamydium spongiosum_, Stöhr, 1880, Palæontogr., vol. xxvi, p. 109,
Taf. v. fig. 3.

Rings of the disk partly concentric, partly spiral, more or less irregular
and often interrupted, with increasing breadth from the centre. Central
part of the disk more or less spongy and obscure. Equatorial girdle half as
broad as the radius of the chambered disk, with smaller pores than the
latter, pierced by twenty to thirty thin, bristle-shaped radial beams,
which proceed over the margin of the disk.

_Dimensions._--Diameter of the disk (with ten rings) 0.2, with the girdle
0.3; breadth of the rings 0.005 to 0.015; pores 0.001 to 0.005.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms; also
fossil in the Tertiary rocks of Barbados and Sicily.

Subfamily 5. EUCHITONIDA, Haeckel.

_Definition._--#Porodiscida# with two or more (commonly three or four)
radial chambered or spongy arms on the margin of the concentrically
annulated disk, situated in its equatorial plane (with or without a
connecting patagium between the arms).

Genus 223. _Amphibrachium_,[262] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with two simple, undivided, chambered arms,
opposite in one axis, without a patagium.

The genus _Amphibrachium_ opens the long series of the Euchitonida, or of
those Porodiscida which bear on the margin of the circular central disk a
certain number of chambered arms, composed of a series of chambers which
are separated by transverse septa. The first group or tribe of this
subfamily is formed by the Amphibrachida, in which the disk bears only two
arms opposite on the poles of one axis. The simplest form of these is
_Amphibrachium_, in which both arms are simple, equal, and without a
patagium or spongy connecticulum.

{517}Subgenus 1. _Amphibrachella_, Haeckel.

_Definition._--Both arms equal, of the same form and size, blunt at the
distal end, without a terminal spine.

1. _Amphibrachium sponguroides_, n. sp.

Both opposite arms of the same form and size, nearly cylindrical, three
times as long as broad, with six to eight transverse septa or joints, at
the distal end rounded, blunt, without a terminal spine.

_Dimensions._--Radius of each arm 0.22, breadth 0.065.

_Habitat._--Antarctic Ocean, Station 154, surface.

2. _Amphibrachium lanceolatum_, n. sp.

Both arms equal, lanceolate, in the middle part three times as broad as at
the two ends, two and a half times as long as broad, with eight to nine
transverse septa, at the distal end blunt, without a terminal spine.

_Dimensions._--Radius of each arm 0.3, greatest breadth 0.09.

_Habitat._--Pacific, central area, Station 267, depth 2700 fathoms.

3. _Amphibrachium dilatatum_, n. sp. (Pl. 44, fig. 6).

Both arms equal, trapezoidal, somewhat broader than long, on the convex
distal end three times as broad as on the narrow base, with five to six
transverse septa, without a terminal spine. Central disk large, somewhat
irregular, with three to four rings, twice as broad as the base of the
arms.

_Dimensions._--Radius of each arm 0.32, basal breadth 0.15, terminal
breadth 0.44.

_Habitat._--South Atlantic, off Patagonia, Station 319, surface.

Subgenus 2. _Amphibrachidium_, Haeckel.

_Definition._--Both arms equal, of the same size and form, with terminal
spines.

4. _Amphibrachium amphilonche_, n. sp.

Both arms equal, lanceolate, in the middle part four times as broad as at
the two ends, twice as long as broad; on the distal end of each arm is a
long conical spine.

_Dimensions._--Radius of each arm (without terminal spine) 0.2, breadth
0.08.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{518}5. _Amphibrachium capitatum_, n. sp.

Both arms equal, club-shaped, three times as long as broad, in the outer
distal half thickened, three times as broad as at the narrow base; on the
distal end of each arm a strong, angular, terminal spine. (The form of the
arms like that of _Stephanastrum capitatum_, Pl. 44, fig. 1.)

_Dimensions._--Radius of each arm 0.22, basal breadth 0.02, terminal
breadth 0.06.

_Habitat._--Equatorial Atlantic, Station 347, surface.

6. _Amphibrachium armatum_, n. sp.

Both arms equal, four times as long as broad, in the thickened distal part
twice as broad as at the base, thorny, with twenty to thirty larger spines
on the distal end, and a very large pyramidal spine in the longitudinal
axis. (The form of the arms like that of the odd arm in _Euchitonia
carcinus_, Pl. 43, fig. 10.)

_Dimensions._--Radius of each arm 0.24, basal breadth 0.03, distal breadth
0.06.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

Subgenus 3. _Amphibrachoma_, Haeckel.

_Definition._--Both arms of different size or form, without terminal
spines.

7. _Amphibrachium indicum_, n. sp.

Both arms club-shaped, but very different in size and form; larger arm
three times as long and twice as broad as the smaller arm; the larger with
nine joints slowly increasing in size, the smaller with four joints,
rapidly increasing; the terminal joint three times as broad as the basal.
Distal end blunt, rounded, without spines.

_Dimensions._--Radius of the larger arm 0.24, of the smaller 0.08; distal
breadth of the former 0.06, of the latter 0.03; basal breadth 0.015.

_Habitat._--Indian Ocean, Ceylon, Haeckel, surface.

Subgenus 4. _Amphibrachura_, Haeckel.

_Definition._--Both arms of different size or form, with terminal spines.

8. _Amphibrachium clavula_, n. sp.

Both arms different in size and form; larger arm club-shaped, four times as
long as broad, at the distal end three times as broad as at the base, and
twice as long as the smaller arm, which resembles a stalked knob, with thin
basal peduncle and spherical distal part. Ends of the two arms {519}thorny
(with numerous smaller, and three to five larger spines); one very large
conical terminal spine on each pole of the main axis.

_Dimensions._--Radius of the larger arm 0.3, of the smaller 0.15; distal
breadth of the former 0.06, of the latter 0.04; basal breath 0.02.

_Habitat._--South Atlantic, Station 333, surface.

Genus 224. _Amphymenium_,[263] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with two simple, undivided, chambered arms,
opposite in one axis, connected by a patagium.

The genus _Amphymenium_ differs from the preceding _Amphibrachium_, its
ancestral form, by development of a patagium or connecticulum between both
arms. This forms a latticed or more spongy envelop, which surrounds either
the middle part of the shell, or the whole shell with exception of the
distal ends of both arms. If the envelop become very spongy, the shell may
be confounded with the cylindrical Ellipside _Spongocore_ (nearly allied to
_Spongurus_); possibly also _Ommatogramma_ of Ehrenberg belongs to this
genus.

Subgenus 1. _Ommatogramma_, Ehrenberg (?).

_Definition._--Both opposite arms of the same size and form, blunt, without
terminal spines.

1. _Amphymenium pupula_, n. sp. (Pl. 44, fig. 8).

Both arms equal, twice as long as broad, three-jointed; the terminal joint
egg-shaped, as large as both other joints together; distal end rounded,
blunt. Patagium nearly complete, enveloping the arms with exception of the
distal end. Perimeter nearly spindle-shaped.

_Dimensions._--Radius of each arm 0.17, greatest breadth 0.06; transverse
breadth of the patagium 0.13.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

2. _Amphymenium naviculare_, Haeckel.

? _Ommatogramma navicularis_, Ehrenberg, 1872, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 317; Abhandl. d. k. Akad. d. Wiss. Berlin, Taf.
vi. fig. 7.

Both arms equal, three times as long as broad, spongy, not jointed; distal
end a little club-shaped, blunt. Patagium nearly complete, enveloping the
arms with exception of the distal end. Perimeter nearly lanceolate. The
imperfect diagnosis and figure of Ehrenberg make it doubtful {520}whether
this species belongs to the Porodiscida (_Amphymenium_) or to the
Spongodiscida (_Spongobrachium_) or perhaps to the Spongurida
(_Spongocore_).

_Dimensions._--Radius of each arm 0.1, greatest breadth 0.03; transverse
breadth of the patagium 0.05.

_Habitat._--North Pacific, Californian Sea, depth 2600 fathoms, Ehrenberg.

3. _Amphymenium zygartus_, n. sp. (Pl. 44, fig. 7).

Both arms equal, four times as long as broad, with seven to eight joints,
slowly decreasing in size towards the blunt end. Patagium incomplete,
protecting only the middle part of the shell on both sides; on each side
two parallel lattice-plates, connected by transverse radial beams,
perpendicular to the surface. Perimeter nearly rectilinear. (Resembles much
certain forms of _Zygartus_, Pl. 40, but is a true Discoid, no Prunoid.)
Compare also Pl. 45, fig. 8.

_Dimensions._--Radius of each arm 0.2, greatest breadth 0.05; transverse
breadth of the patagium 0.11.

_Habitat._--Pacific, central area, Station 271, surface.

4. _Amphymenium monstrosum_, n. sp. (Pl. 44, fig. 11).

Both arms equal, little longer than broad, with six to seven convex joints.
The axis of both arms is not common and straight, as in all other species
of this genus, but broken, therefore the incomplete patagium, which
envelops only two to three joints of the arms, is on one side convex, on
the other side concave; it is formed by a simple lattice-plate, connected
with the arms by numerous radial beams. This anomalous form, seen only
once, may perhaps be a monstrosity of _Euchitonia_.

_Dimensions._--Radius of each arm 0.12, greatest breadth 0.06; transverse
breadth of the patagium 0.13.

_Habitat._--South Pacific, Station 290, surface.

Subgenus 2. _Ommathymenium_, Haeckel.

_Definition._--Both opposite arms of the same size and form, armed at the
distal end with terminal spines.

5. _Amphymenium amphistylium_, n. sp. (Pl. 44, fig. 9).

Both arms equal, three times as long as broad, thickened towards the
truncated distal end, and armed with a strong pyramidal terminal spine.
Each arm with seven joints, separated by convex, transverse septa, and
halved by a radial beam lying in the longitudinal axis. Patagium
incomplete, cylindrical, enveloping only the middle part of the shell.

_Dimensions._--Radius of each arm 0.18, greatest breadth 0.06; transverse
breadth of the patagium 0.08.

_Habitat._--North Atlantic, Station 354, surface.

{521}6. _Amphymenium fusiforme_, n. sp.

Both arms equal, lanceolate, three times as long as broad in the width,
with seven to eight joints. Distal end pointed, armed with a strong conical
terminal spine. Patagium complete, enveloping the whole shell with
exception of the terminal spines. Whole form spindle-shaped.

_Dimensions._--Radius of each arm 0.2, greatest breadth 0.07; transverse
breadth of the patagium 0.15.

_Habitat._--South Atlantic, Station 330, surface.

Genus 225. _Amphirrhopalum_,[264] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with two chambered arms, opposite in one axis,
without a patagium; one arm or both forked at the distal end.

The genus _Amphirrhopalum_ differs from _Amphibrachium_, its ancestral
form, by bifurcation of the distal ends of the arms, which may affect
either both arms, or only one of them.

Subgenus 1. _Amphirrhopalium_, Haeckel.

_Definition._--Both opposite arms of similar size and form, with blunt
branches, without terminal spines.

1. _Amphirrhopalum ximorphum_, n. sp.

Both arms equal, in the proximal half simple, in the distal half forked,
with six to seven transverse septa; distal end of each branch blunt,
without terminal spine, somewhat broader than the base of the whole arm.
Axis of the branches concavely curved. (Resembles _Amphicraspedum
maclaganium_, Pl. 45, fig. 11, but wants the patagium.)

_Dimensions._--Radius of the arms 0.18, basal breadth 0.065; terminal
breadth of each branch 0.075.

_Habitat._--North Pacific, off Japan, Station 240, surface.

2. _Amphirrhopalum amphidicranum_, n. sp.

Both arms equal, in the proximal half simple, in the distal half forked,
with irregular septa; distal end of each arm blunt, without a terminal
spine, smaller than the basal breadth of the arm. Axis of the branches
straight. (Resembles _Dicranastrum furcatum_, Pl. 47, fig. 2, but without
lateral arms.)

_Dimensions._--Radius of each arm 0.15, basal breadth 0.03; terminal
breadth of each branch 0.02.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

{522}Subgenus 2. _Amphirrhopella_, Haeckel.

_Definition._--Both opposite arms of similar size and form, with terminal
spines.

3. _Amphirrhopalum bigeminum_, n. sp.

Both arms equal, in the proximal larger half simple, in the distal smaller
half forked; each branch triangular, with a strong conical terminal spine.
Axis of the branches straight. (Resembles _Dicranastrum cornutum_, Pl. 45,
fig. 2, but without lateral arms.)

_Dimensions._--Radius of the arms (without spines) 0.15, basal breadth
0.03; breadth of the bifurcation 0.08.

_Habitat._--South Pacific, Station 284, surface.

4. _Amphirrhopalum echinatum_, n. sp. (Pl. 45, fig. 10).

Both arms equal, in the proximal smaller half simple, nearly square, in the
distal larger half forked; the branches thorny, armed at the end with
numerous spines, one larger on the terminal pole of the concavely curved
arm-axis.

_Dimensions._--Radius of the arms (without spines) 0.15, basal breadth
0.05; breadth of branches 0.03.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Subgenus 3. _Amphirrhopoma_, Haeckel.

_Definition._--Both opposite arms of different size or form, without
terminal spines.

5. _Amphirrhopalum ypsilon_, n. sp.

Both arms very different. Larger arm simple, egg-shaped, twice as long as
broad; smaller arm in the basal half simple, nearly square, in the distal
half forked; both branches egg-shaped, blunt. (Resembles _Amphicraspedum
wyvilleanum_, Pl. 45, fig. 12, but wants the patagium.)

_Dimensions._--Radius of the larger simple arm 0.18, breadth 0.09; radius
of the smaller forked arm 0.15, breadth of its branches 0.05.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

Genus 226. _Amphicraspedum_,[265] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with two chambered arms, opposite in one axis,
connected by a lateral patagium; one arm or both forked at the distal end.

The genus _Amphicraspedum_ exhibits the same bifurcation of the arms as
does _Amphirrhopalum_, but differs from this ancestral form in the
development of a patagium, an external connecticulum between the arms,
which envelops the disk totally or partially.

{523}Subgenus 1. _Amphicraspedon_, Haeckel.

_Definition._--Both arms of equal size and form, without terminal spines of
the branches.

1. _Amphicraspedum maclaganium_, n. sp. (Pl. 45, fig. 11).

Both arms equal, in the proximal half simple, in the distal half forked,
with six to seven transverse septa; distal end of each branch rounded,
blunt, somewhat broader than the base of the whole arm. Divergent axes of
both branches concavely curved. Patagium incomplete, with elliptical
perimeter, enveloping only the middle part of the shell. I call this
interesting species in honour of Miss Nellie Maclagan, the learned
translator of several zoological papers from German into English.

_Dimensions._--Radius of each arm 0.25, basal breadth 0.07; distal breadth
of each branch 0.08; equatorial breadth of the patagium 0.25.

_Habitat._--North Atlantic, off Halifax, Station 50, surface.

Subgenus 2. _Amphicraspedina_, Haeckel.

_Definition._--Both arms of different size or form, without terminal spines
on the branches.

2. _Amphicraspedum wyvilleanum_, n. sp. (Pl. 45, fig. 12).

Both arms different. Larger arm simple, egg-shaped, with eleven convex
joints, one and a half times as long as broad; smaller arm in the basal
half simple, triangular, with six cap-like joints, in the distal half
forked; both branches egg-shaped, with five joints and blunt ends. Patagium
nearly complete with four to five concave chamber-rows. Called in honour of
Sir C. Wyville Thomson.

_Dimensions._--Radius of the larger simple arm 0.18, breadth 0.08; radius
of the smaller forked arm 0.16; breadth of the branches 0.05; transverse
breadth of the patagium 0.2.

_Habitat._--South Atlantic, Station 333, surface.

Subgenus 3. _Amphicraspedula_, Haeckel.

_Definition._--Both arms of different size or form, with terminal spines of
the branches.

3. _Amphicraspedum murrayanum_, n. sp. (Pl. 44, fig. 10).

_Amphymenium murrayanum_, Haeckel, 1879, MS. et Atlas (pl. xliv. fig.
10).

Both arms different in size; the larger one and a half times as long and
broad as the smaller. Both arms triangular, forked at the broader distal
end, with two very strong, conical, divergent, {524}straight terminal
spines. Patagium broad, incomplete, with circular perimeter. Called in
honour of my friend Dr. John Murray.

_Dimensions._--Radius of the larger arm (including the spines) 0.24, of the
smaller 0.16; distance of the terminal points of the former 0.18, of the
latter 0.09; diameter of the patagium 0.2.

_Habitat._--North Atlantic, Færöe Channel, Gulf Stream, surface, John
Murray.

Genus 227. _Dictyastrum_,[266] Ehrenberg, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 830.

_Definition._--#Porodiscida# with three simple, undivided, chambered arms,
without a patagium; triangular shell regular, with three equal arms and
three equal angles.

The genus _Dictyastrum_ is the simplest form of the Trigonastrida, or of
the Porodiscida, in which the margin of the central disk is furnished with
three chambered arms. In _Dictyastrum_ these are quite simple and regular,
without a patagium, separated by equal angles, so that the whole shell
represents a regular, equilateral triangle, if we connect the distal points
of the arms by lines. The genus _Dictyastrum_, founded by Ehrenberg in
1860, differs from his _Rhopalodictyum_--after his own diagnosis--only by
an insignificant difference in the form of the simple arms, which is
scarcely a specific character. I therefore apply this name here in the
above amended sense, seeing that the only figured species of Ehrenberg
(_Dictyastrum angulatum_) occurs in two different, but externally very
similar forms: one of these is a true Porodiscid (_Dictyastrum_) with two
porous covering-plates and concentric rings; the other is a true
Spongodiscid (_Rhopalodictyum_) with quite spongy, irregular network, and
is probably identical with the _Rhopalodictyum truncatum_ of Ehrenberg.

Subgenus 1. _Dictyastrella_, Haeckel.

_Definition._--Arms with blunt ends, without terminal spines.

1. _Dictyastrum angulatum_, Ehrenberg.

_Dictyastrum angulatum_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 289, Taf. viii. fig. 18.

Arms nearly square with straight edges, towards the truncated end a little
broader, about the same diameter as the triangular central disk. The figure
of Ehrenberg seems to represent a Spongodiscid (_Rhopalodictyum
angulatum_), but in the same locality (Philippine Sea) occurs also a true
_Dictyastrum_ of quite the same form, but with three to four concentric
rings of the central disk, and with jointed arms.

{525}_Dimensions._--Radius of each arm (length from the centre to the
distal end) 0.13; breadth of the truncated end 0.1.

_Habitat._--Tropical Pacific, Philippine Sea, Station 200, depth 250
fathoms.

2. _Dictyastrum bandaicum_, Haeckel.

_Rhopalastrum bandaicum_, Harting, 1863, Mikr. Fauna Banda-Zee, p. 16,
Taf. iii. fig. 45.

Arms nearly square, with convex edges, in the middle a little broader than
at both ends, about half the diameter of the central disk. Differs from the
nearly allied preceding species by the half size of the arms and the convex
edges.

_Dimensions._--Radius of each arm 0.12, its greatest breadth 0.07.

_Habitat._--Tropical Pacific, Banda Sea, Harting.

3. _Dictyastrum hexagonum_, n. sp. (Pl. 43, fig. 7).

_Rhopalastrum hexagonum_, Haeckel, 1880, Atlas (pl. xliii. fig. 1).

Arms nearly triangular, one and a third times as broad at the distal end as
long, and three times as broad as at the base. Central disk about the same
diameter. In each arm six simple broad chambers. If we connect the six
corners of the truncated distal ends by straight lines, we get a regular
hexagon.

_Dimensions._--Radius of each arm 0.17, basal breadth 0.06, terminal
breadth 0.17.

_Habitat._--Equatorial Atlantic, Station 347, surface.

4. _Dictyastrum trirrhopalum_, n. sp.

Arms club-shaped, five times as long as broad at the base, at the thickened
end three times as broad as at the base. Diameter of the central disk
equals half the length of the arms. (Similar to _Rhopalastrum malleus_, Pl.
43, fig. 1, but with three equal angles and much smaller disk.)

_Dimensions._--Radius of each arm 0.25, basal breadth 0.04, distal breadth
0.12.

_Habitat._--Pacific, central area, Station 273, surface.

Subgenus 2. _Dictyastromma_, Haeckel.

_Definition._--Arms on the distal end provided with terminal spines.

5. _Dictyastrum trispinosum_, n. sp. (Pl. 43, fig. 5).

_Rhopalastrum trispinosum_, Haeckel, 1881, Prodromus et Atlas (pl. xliii.
fig. 5).

Arms trapezoid, at the rounded distal end twice as broad as at the base,
with a strong and short, conical, terminal spine. Diameter of the circular
central disk about equal to the length and the greatest breadth of the
arms.

_Dimensions._--Radius of each arm 0.15, basal breadth 0.06, distal breadth
0.11.

_Habitat._--Equatorial Atlantic, Station 347, surface.

{526}6. _Dictyastrum triactis_, Ehrenberg.

_Dictyastrum triactis_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 306.

Arms rectilinear, four times as long as broad, with parallel edges, pointed
at the distal end, with a short terminal spine. Diameter of the circular
central disk equal to double the breadth of the arms.

_Dimensions._--Radius of each arm 0.2, breadth 0.04.

_Habitat._--Pacific, Philippine Sea, Station 206, depth 2100 fathoms.

7. _Dictyastrum aculeatum_, n. sp.

Arms lanceolate, three times as long as broad, twice as broad in the middle
as at either end, with thorny surface and numerous conical terminal spines,
one very large in the radius. Central disk triangular, about as broad as
the arms. (Resembles _Rhopalastrum arcticum_, Pl. 43, fig. 6, but differs
by the equal angles and the triangular disk.)

_Dimensions._--Radius of each arm 0.2, breadth 0.06.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 228. _Rhopalastrum_,[267] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Porodiscida# with three simple undivided, chambered arms,
without a patagium; triangular shell bilateral, one odd arm opposite to the
odd angle between two paired arms.

The genus _Rhopalastrum_, founded by Ehrenberg (1847) with a very
insufficient diagnosis, is here retained for those Trigonastrida that agree
in the generic characters with the only species figured by him, viz.,
_Rhopalastrum lagenosum_ (compare my Monograph, 1862, p. 500). It
comprises, therefore, such Euchitonida as agree with the preceding
_Dictyastrum_ in the simple form of the three arms and the absence of a
patagium, but differ from it in the different size of the three angles, and
often also in the divergent form and size of the three arms; one odd arm is
opposite to the odd angle between the two paired arms.

Subgenus 1. _Rhopalastrella_, Haeckel.

_Definition._--Arms with blunt ends, without terminal spines.

1. _Rhopalastrum truncatum_, Haeckel.

_Rhopalastrum truncatum_, Haeckel, 1862, Monogr. d. Radiol., p. 500, Taf.
xxix. fig. 6.

Distance of both paired arms about half as large as their distance from the
odd arm. All three arms nearly of the same form and size, very short and
broad; their breadth nearly equals that of {527}the roundish central disk,
whilst their length reaches only one-fourth of it. End of the arms convex
rounded, without spines.

_Dimensions._--Radius of each arm 0.1, breadth 0.15.

_Habitat._--Mediterranean (Messina), Atlantic (Canary Islands), surface,
Haeckel.

2. _Rhopalastrum pistillum_, Stöhr.

_Rhopalastrum pistillum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 110,
Taf. v. fig. 4.

Distance between the paired arms about two-thirds as large as their
distance from the odd arm. All three arms nearly of the same form and size,
about three times as long as the diameter of the central disk, at the base
one-third as broad as at the convex rounded end, without spines. Stöhr has
only observed a fragment with one arm; some perfect specimens, which I
found in the Caltanisetta-rock, exhibited nearly the same form as
_Rhopalastrum malleus_ (Pl. 43, fig. 1), but differ from this by the
smaller disk, the broader arms, and the smaller angle between the paired
arms.

_Dimensions._--Radius of all three arms 0.2; basal breadth of each arm
0.045, terminal breadth 0.13.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte (Stöhr),
Caltanisetta (Haeckel).

3. _Rhopalastrum malleus_, n. sp. (Pl. 43, fig. 1).

Distance between the paired arms one and a third times as large as their
distance from the odd arm. All three arms nearly of the same form and size,
hammer-shaped, three times as broad at the truncated distal end as at the
base. Central disk broader than the arms.

_Dimensions._--Radius of each arm 0.25, basal breadth 0.05, distal breadth
0.15.

_Habitat._--South Atlantic, Station 325, surface.

4. _Rhopalastrum lagenosum_, Ehrenberg.

_Rhopalastrum lagenosum_, Ehrenberg, 1847, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 43; Mikrogeol., 1854, Taf. xxii. fig. 22.

_Rhopalastrum lagenosum_, Haeckel, 1862, Monogr. d. Radiol., p. 501.

_Flustrella bilobata_, Ehrenberg, 1844, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 81.

_Haliomma lagena_, Ehrenberg, 1840, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 200.

Distance between the paired arms one and a half times as great as their
distance from the odd arm. All three arms nearly of the same form and size,
about one and a half times as long as the diameter of the central disk, at
the base half as broad as at the convex rounded end, without spines.
Ehrenberg has only observed a fragment with two paired arms; some perfect
specimens with three arms, observed by me, differed from the nearly allied
_Rhopalastrum pistillum_ (from the same locality) by the larger disk, the
broader arms, and the larger unpaired angle.

_Dimensions._--Radius of all three arms 0.2, basal breadth 0.05, terminal
breadth 0.09.

_Habitat._--Fossil in Tertiary rocks of Sicily, Caltanisetta.

{528}5. _Rhopalastrum ypsilinum_, n. sp. (Pl. 43, fig. 2).

Distance between the paired arms half as large as their distance from the
odd arm. All three arms at the concavely curved distal end three times as
broad as at the narrow base. Odd arm twice as long and broad as the paired
arms. Central disk smaller than the latter. No terminal spines.

_Dimensions._--Radius of the odd arm 0.32, of the paired arms 0.2; terminal
breadth of the former 0.2, of the latter 0.12.

_Habitat._--Indian Ocean, Cocos Islands, Rabbe, surface.

6. _Rhopalastrum clavatum_, n. sp.

Distance between the paired arms half as large as their distance from the
odd arm. All three arms club-shaped, at the thickened, nearly spherical,
distal end three times as broad as at the narrow base. Odd arm nearly twice
as long and broad as the paired arms. Central disk equal to the distal knob
of the latter. No terminal spines.

_Dimensions._--Radius of the odd arm 0.3, of the paired arms 0.18; distal
breadth of the former 0.16, of the latter 0.08.

_Habitat._--South Atlantic, Station 332, surface.

7. _Rhopalastrum irregulare_, n. sp. (Pl. 43, fig. 8).

Distance between all three arm-points different. All three arms
cylindrical, nearly of the same length, but of different form, irregularly
curved, about four times as long as broad, with blunt ends.

_Dimensions._--Length of each arm about 0.2, breadth 0.05.

_Habitat._--Antarctic Sea, Station 157, depth 1950 fathoms.

Subgenus 2. _Rhopalastromma_, Haeckel.

_Definition._--Arms on the distal end provided with one or more terminal
spines.

8. _Rhopalastrum martellum_, n. sp.

Distance between all three arm-points nearly the same; but the odd arm is
one and a half times as large as both paired arms, and is perpendicular to
the common axis of the latter, therefore the shell has the form of a
hammer. Each arm is twice as broad at the distal end as at the base, and
armed with a conical terminal spine; the latter is vertical in the odd arm,
horizontal in the paired arms.

_Dimensions._--Radius of the odd arm 0.3, of the paired arms 0.15; distal
breadth of the former 0.12, of the latter 0.06.

_Habitat._--North Atlantic, Station 354, surface.

{529}9. _Rhopalastrum triceros_, n. sp. (Pl. 43, fig. 4).

Distance between the paired arms about two-thirds as large as their
distance from the odd arm. All three arms club-shaped, three times as broad
at the thickened distal part as at the base, and armed with one single,
conical, terminal spine. Odd arm of the same breadth, but twice as long as
the paired arms.

_Dimensions._--Radius of the odd arm 0.35, of the paired arms 0.2; basal
breadth 0.04, distal breadth 0.12.

_Habitat._--Pacific, central area, Station 274, surface.

10. _Rhopalastrum hexaceros_, n. sp. (Pl. 43, fig. 3).

Distance between the paired arms equals four-fifths of their distance from
the odd arm. All three arms nearly of the same size, about square, a little
broader at the truncated distal end, which is armed at both corners with a
strong, conical, radial spine.

_Dimensions._--Radius of each arm 0.2, basal breadth 0.1, distal breadth
0.12.

_Habitat._--Indian Ocean, Ceylon, Belligemma, Haeckel, surface.

11. _Rhopalastrum arcticum_, n. sp. (Pl. 43, fig. 6).

Distance between the paired arms half as large as their distance from the
odd arm, which is a little larger. All three arms of the same form,
lanceolate, twice to three times as long as broad, twice as broad in the
middle as at either end. Each arm with twelve to fourteen transverse septa,
at the distal end with a bunch of conical spines, and one single, very
large, pyramidal, terminal spine.

_Dimensions._--Radius of each arm (without spine) 0.17, greatest breadth of
it 0.05 to 0.06.

_Habitat._--Arctic Ocean, lat. 83° 19' N., North Polar expedition of the
"Alert."

Genus 229. _Hymeniastrum_,[268] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Porodiscida# with three simple, undivided, chambered arms,
connected by a patagium; triangular shell regular, with three equal arms
and three equal angles.

The genus _Hymeniastrum_ was founded by Ehrenberg (1847) with a very
incomplete diagnosis, and hitherto known only by one single species,
figured by him as _Hymeniastrum pythagoræ_ (Mikrogeol., 1854, Taf. xxxvi.
fig. 31). This form occurs in two different states, externally quite
identical; in one state the central disk (as figured, _loc. cit._), is a
simple lens or hollow disk, containing a medullary shell or "central
chamber"; in the other state the central disk is composed of two concentric
rings surrounding the "central chamber." We retain here the name
_Hymeniastrum_ for this latter state, {530}expressed in the diagnosis given
above, and call the former state (the Coccodiscid) _Hymenactura_ (compare
above, p. 473). One practical advantage, obtained in this way, is that all
genera of #Discoidea# ending with "_-astrum_" belong to the Porodiscida.
_Hymeniastrum_ differs from _Dictyastrum_ by the possession of a patagium,
and from _Euchitonia_ by the equal size of the angles and the arms.

Subgenus 1. _Hymenastrella_, Haeckel.

_Definition._--Arms with blunt ends, without terminal spines.

1. _Hymeniastrum leydigii_, Haeckel.

_Euchitonia leydigii_, Haeckel, 1862, Monogr. d. Radiol., p. 510, Taf.
xxxi. figs. 4, 5.

_Hymeniastrum leydigii_, Haeckel, 1881, Prodromus, p. 460.

_Histiastrum trinacrium_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 843.

Arms twice as long as broad, two-thirds as broad at the base as at the
blunt, nearly truncated distal end. Patagium nearly complete, perfectly
filling out the interbrachial spaces, with six to seven regular, concave
chamber-rows; only the truncate terminal faces of the arms free.

_Dimensions._--Radius of the arms 0.15 to 0.18, breadth on their base 0.04
to 0.05, on their broadest distal end 0.06 to 0.07.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface. Very
common and variable.

2. _Hymeniastrum köllikeri_, Haeckel.

_Euchitonia köllikeri_, Haeckel, 1862, Monogr. d. Radiol., p. 511, Taf.
xxxi. figs. 6, 7.

_Hymeniastrum köllikeri_, Haeckel, 1881, Prodromus, p. 460.

Arms nearly lanceolate, twice as long as broad, half as broad at the base
as at the broadest middle part, pointed at the distal end. Patagium
complete, enveloping the whole triangular disk, with irregular
chamber-rows.

_Dimensions._--Radius of the arms 0.2, breadth on their base 0.04, on the
broadest part 0.08.

_Habitat._--Mediterranean (Messina), surface.

3. _Hymeniastrum gümbelii_, Haeckel.

_Stylactis gümbelii_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 112, Taf.
vi. fig. 1.

Arms twice as long as broad, nearly twice as broad in the circular distal
half as in the square proximal half. Patagium incomplete, circular, with
eight to nine convex chamber-rows, enveloping only the proximal square
halves of the arms; the lenticular distal halves remain free.

_Dimensions._--Radius of the arms 0.14; breadth at the base 0.04, at the
distal lenticular part 0.07.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Stöhr.

{531}4. _Hymeniastrum euclidis_, n. sp. (Pl. 43, fig. 13).

_Euchitonia euclidis_, Haeckel, 1881, Prodromus, p. 460 et Atlas (pl.
xliii. fig. 13).

Arms one and a half times as long as broad, club-shaped, three times as
broad in the oval distal part as in the narrow square proximal part.
Patagium complete, enveloping the whole triangular disk, and also the
convex ends of the arms, with eight to nine convex chamber-rows.

_Dimensions._--Radius of the arms 0.18; breadth at the base 0.025, at the
broadest distal part 0.07.

_Habitat._--Pacific, central area, Stations 270 to 274, surface.

5. _Hymeniastrum pythagoræ_, Ehrenberg.

_Hymeniastrum pythagoræ_, Ehrenberg, 1854 (_partim_), Mikrogeol., Taf.
xxxvi. fig. 31.

Arms nearly as broad as long, two-thirds as broad at the base as at the
truncated distal end. Patagium incomplete, enveloping only the basal half
of the arms, with four to five rectilinear parallel chamber-rows. (This
form has the greatest resemblance to the figure given by Ehrenberg, _loc.
cit._, but differs by the central disk, which is composed of two concentric
rings surrounding the small central chamber; compare above _Hymenactura
pythagoræ_, p. 474.)

_Dimensions._--Radius of the arms 0.18; breadth at the base 0.08, at the
truncated end 0.12.

_Habitat._--Equatorial Atlantic, Station 347, depth 2250 fathoms.

6. _Hymeniastrum archimedis_, n. sp.

Arms nearly triangular, at the truncated, slightly convex end three times
as broad as at the narrow base and one and a third times as broad as long;
each arm with six simple chambers. Patagium incomplete, enveloping only the
basal half of the arms with three to four convex chamber-rows. (Differs
from _Rhopalastrum hexagonum_, Pl. 43, fig. 7, only by the patagium.)

_Dimensions._--Radius of the arms 0.2; breadth at the base 0.07, at the
truncated end 0.2.

_Habitat._--Pacific, central area, Station 274, surface.

Subgenus 2. _Hymenastromma_, Haeckel.

_Definition._--Arms at the distal end provided with radial spines.

7. _Hymeniastrum ternarium_, Haeckel.

_Histiastrum ternarium_, Ehrenberg, 1875, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 76, Taf. xxiv. fig. 2.

Arms two and a half times as long as broad, three times as broad at the
truncated end as at the narrow base, with a strong, conical, radial spine
at the end. Patagium incomplete, enveloping only the basal half of the
arms, with three to four convex chamber-rows.

_Dimensions._--Radius of the arms (without terminal spine) 0.02; breadth at
the base 0.025, at the broadest terminal part 0.08.

_Habitat._--Fossil in the rocks of Barbados.

{532}8. _Hymeniastrum trigonarium_, n. sp.

Arms one and a half times as long as broad, a little broader at the rounded
end than at the base, with three strong conical radial spines, one larger
(in the radius of each arm) and two smaller on both sides of this. Patagium
complete, with four to five rectilinear parallel chamber-rows, enveloping
the whole arms (with exception of the terminal spines) and forming a
perfect equilateral triangle.

_Dimensions._--Radius of the arms (without terminal spines) 0.18; breadth
at the base 0.1, at the distal part 0.12.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

Genus 230. _Euchitonia_,[269] Ehrenberg, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 831 (_sensu emendato_).

_Definition._--#Porodiscida# with three simple, undivided, chambered arms,
connected by a patagium; triangular shell bilateral, one odd arm opposite
to the odd angle between the two paired arms.

The genus _Euchitonia_, quite insufficiently characterised by Ehrenberg,
was founded by him (1860) for one single species, afterwards (1872)
described and figured as _Euchitonia furcata_. Retaining this species
correctly as the type of this genus, I give to it here the above diagnosis.
In my Monograph (1862, p. 503) I described seven Mediterranean species of
_Euchitonia_. Three of these have in common the characters according to the
present diagnosis: _Euchitonia mülleri_, _Euchitonia virchowii_,
_Euchitonia beckmanni_; two others appertain (on account of the regular,
not bilateral form) to _Hymeniastrum_, and two others (on account of the
forked, not simple arms) to _Trigonastrum_. Afterwards (1880) three true
fossil species of _Euchitonia_ were described by Stöhr (_Euchitonia
cruciata_, _Euchitonia zittelii_, and _Euchitonia acuta_). Some species of
this genus are cosmopolitan, and appertain to the most common and
everywhere represented #Discoidea#.

Subgenus 1. _Stylactis_, Ehrenberg, 1872 (_loc. cit._).

_Definition._--Arms with blunt ends, without terminal spines.

1. _Euchitonia furcata_, Ehrenberg.

_Euchitonia furcata_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 308; Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. vi. iii.
fig. 6.

Distance between the paired arms about half as large as their distance from
the odd arm. All three arms nearly of the same size and form, about twice
as long as broad, one and a half times at the {533}blunt convex end as
broad as at the base. Patagium incomplete, with concave chamber-rows and
irregular network, only enveloping the basal half of the arms.

_Dimensions._--Radius of all three arms 0.15, basal breadth of each arm
0.04, terminal breadth 0.06.

_Habitat._--North Pacific, Californian Sea, depth 2600 fathoms, Ehrenberg.

2. _Euchitonia mülleri_, Haeckel.

_Euchitonia mülleri_, Haeckel, 1862, Monogr. d. Radiol., p. 508, Taf.
xxx. figs. 5-10.

_Euchitonia mülleri_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 110, Taf.
v. fig. 5.

Distance between the paired arms about two-thirds as large as their
distance from the odd arm, which is somewhat larger. Length of the arms
equals two and a half times the breadth of the blunt convex end or five
times the breadth of the base. Patagium with concave chamber-rows, nearly
complete, enveloping the arms with exception of the terminal face. (This
common species is very variable; compare my Monograph.)

_Dimensions._--Radius of all three arms 0.16 to 0.22, basal breadth 0.03 to
0.05, terminal breadth 0.06 to 0.08.

_Habitat._--Cosmopolitan; one of the most common #Discoidea# in all seas,
on the surface as well as at different depths; also fossil in the Tertiary
rocks of Barbados and Sicily.

3. _Euchitonia triangulum_, Haeckel.

_Stylactis triangulum_, Ehrenberg, 1872, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 320; Abhandl. d. k. Akad. d. Wiss. Berlin, Taf. viii.
fig. 9.

_Stylactis triangulum_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 113, Taf.
vi. fig. 2.

Distance between the paired arms about two-thirds as large as their
distance from the odd arm. All three arms nearly of the same size and form,
about one and a half times as long as broad, nearly as broad at the base as
at the blunt rounded end. Patagium incomplete, with convex chamber-rows,
enveloping about two-thirds of the arms.

_Dimensions._--Radius of all three arms 0.15, basal breadth 0.05, terminal
breadth 0.06.

_Habitat._--Pacific and Atlantic, Stations 253, 272, 354, surface; also
fossil in the Tertiary rocks of Barbados and Sicily.

4. _Euchitonia cruciata_, Stöhr.

_Euchitonia cruciata_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 111, Taf.
v. fig. 7.

Distance between the paired arms about one and a half times as large as
their distance from the odd arm, which is a little larger. Length of the
arm nearly equals twice the breadth, which is the same at the base and at
the rounded blunt ends. Patagium incomplete, with concave chamber-rows,
enveloping only the base of the arms, and forms between them three other
smaller arms; therefore the whole shell forms six angles with six
alternating arms.

_Dimensions._--Radius of all three arms 0.15, basal breadth 0.05.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

{534}5. _Euchitonia lanceolata_, n. sp. (Pl. 43, fig. 9).

Distance between the paired arms about one and a third times as large as
their distance from the odd arm. All three arms nearly of the same size and
form, three times as long as broad, lanceolate, much broader in the middle
part than at both ends; distal end cuspidate, but not spiny. Patagium
incomplete, with convex chamber-rows, envelops about two-thirds of the
arms.

_Dimensions._--Radius of all three arms 0.18, greatest breadth (in the
width) 0.06.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

6. _Euchitonia zittelii_, Haeckel.

_Stylactis zittelii_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 112, Taf.
v. fig. 8.

Distance between the paired arms about one-fifth as large as their distance
from the odd arm. All three arms nearly of the same size and form, in the
proximal half thinner and nearly square, in the distal half thicker and
circular; the latter half twice to three times as broad as the former.
Patagium incomplete, with convex chamber-rows, envelops only the proximal
square half of the arms.

_Dimensions._--Radius of all three arms 0.15, basal breadth 0.03, terminal
breadth 0.08.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte, Stöhr.

7. _Euchitonia stöhrii_, n. sp. (Pl. 43, fig. 12).

Distance between the paired arms about four-fifths as large as their
distance from the odd arm, which is one and a half times as long as the
former. Arms about three times as long as broad, twice as broad in the
distal half as in the proximal half, with rounded blunt ends. Patagium
complete, with concave chamber-rows, enveloping the whole shell, also the
ends of the arms.

_Dimensions._--Radius of the paired arms 0.2, of the odd arm 0.25; basal
breadth 0.03, distal breadth 0.06.

_Habitat._--Fossil in the rocks of Barbados and of Nicobar Islands,
Haeckel. Indian Ocean, Cocos Islands (Rabbe). Also in the Central Pacific,
Station 267.

8. _Euchitonia beckmannii_, Haeckel.

_Euchitonia beckmannii_, Haeckel, 1862, Monogr. d. Radiol., p. 505, Taf.
xxxi. fig. 1.

Distance between the paired arms scarcely half as large as their distance
from the odd arm, which is somewhat larger. Each arm with six simple broad
chambers (without radial septa), the terminal chambers semilunar, convex,
blunt, four times as broad as the first (basal) chamber. Patagium
incomplete, with convex chamber-rows, enveloping the arms with exception of
the broad blunt terminal face.

_Dimensions._--Radius of the paired arms 0.15, of the odd arm 0.18; basal
breadth 0.02 to 0.03, terminal breadth 0.1 to 0.15.

_Habitat._--Mediterranean (Messina), Haeckel, surface.

{535}9. _Euchitonia virchowii_, Haeckel.

_Euchitonia virchowii_, Haeckel, 1862, Monogr. d. Radiol., p. 503, Taf.
xxx. figs. 1-4.

_Histiastrum fasciatum_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 842.

Distance between the paired arms about half as large as their distance from
the odd arm, which is somewhat larger. Each arm with six broad chambers,
bisected by a radial septum; the terminal chamber convex, blunt, twice as
broad as the basal chamber. Patagium incomplete, with concave chamber-rows,
enveloping the arms with exception of the broad blunt terminal face.

_Dimensions._--Radius of the paired arms 0.15, of the odd arm 0.16; basal
breadth 0.05, terminal breadth 0.1 to 0.12.

_Habitat._--Mediterranean (Messina), Atlantic (Canary Islands).

Subgenus 2. _Pteractis_, Ehrenberg, 1872 (_loc. cit._).

_Definition._--Arms provided with radial spines at the distal end.

10. _Euchitonia elegans_, Haeckel.

_Pteractis elegans_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 299, Taf. viii. fig. 3.

Distance between the paired arms half as large as their distance from the
odd arm. This latter is straight, while both the former are concavely
curved towards the middle line. Arms five times as long as broad, at the
distal end pointed and armed with a short conical terminal spine. Patagium
nearly complete, enveloping four-fifths of the arms, with four to five
concave chamber-rows.

_Dimensions._--Radius of the arms 0.2, breadth of them 0.03.

_Habitat._--Tropical Pacific, Philippine Sea, depth 3300 fathoms
(Ehrenberg).

11. _Euchitonia carcinus_, n. sp. (Pl. 43, fig. 10).

Distance between the paired arms scarcely one-fourth as great as their
distance from the odd arm. This latter is straight, twice as long as the
former, which are concavely curved towards the middle line. The odd arm is
three times as broad at the distal end as at the narrow base. The end of
each arm is furnished with a strong triangular radial spine and a group of
smaller spines. Patagium incomplete, with two to three concave
chamber-rows, enveloping only the basal half of the arms.

_Dimensions._--Radius of the paired arms 0.15, breadth 0.03; radius of the
odd arm 0.3, breadth on its base 0.02, on its distal end 0.07.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

{536}12. _Euchitonia acuta_, Stöhr.

_Euchitonia acuta_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 111, Taf. v.
fig. 6.

Distance between the paired arms two-thirds as great as their distance from
the odd arm. All three arms nearly of the same size and form, two and a
half times as long as broad, nearly lanceolate, at their broadest part one
and a half times as broad as at their base. Patagium complete, enveloping
the whole triangular disk, with five to six concave chamber-rows. In the
figure of Stöhr the ends of the arms are simply pointed, while I find in
the same fossil form a short terminal conical spine.

_Dimensions._--Radius of all three arms 0.14; breadth at the base 0.04, at
the broadest part 0.06.

_Habitat._--Fossil in Tertiary rocks of Sicily, Grotte (Stöhr),
Caltanisetta (Haeckel).

13. _Euchitonia ypsiloides_, Haeckel.

_Histiastrum ypsiloides_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 843.

Distance between the paired arms two-thirds as great as their distance from
the odd arm, which is somewhat larger. Length of the arms equals five times
the breadth of the narrow base, which is half that of the distal end; this
latter is armed with three to five short conical spines. Patagium complete,
with six to seven concave chamber-rows, enveloping the whole arms with the
exception of the terminal spines. (Differs from _Euchitonia mülleri_ almost
solely by the possession of terminal spines.)

_Dimensions._--Radius of the arms 0.18 to 0.2; breadth at the base 0.04, at
the distal end 0.08.

_Habitat._--Mediterranean (Messina), Atlantic (Canary Islands), surface.

14. _Euchitonia echinata_, n. sp. (Pl. 43, fig. 11).

Distance between paired arms three-fourths as great as their distance from
the odd arm, which is somewhat larger. Arms one and a half times as long as
broad, somewhat constricted in the middle part, armed at the rounded ends
with numerous (thirty to forty) strong, conical spines. Patagium complete,
with four or five rectilinear parallel chamber-rows, enveloping the whole
of the arms with the exception of the spiny ends.

_Dimensions._--Radius of the arms (without spines) 0.2, breadth 0.06 to
0.09.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

Genus 231. _Chitonastrum_,[270] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with three forked, chambered arms, without a
patagium. (Arms and angles between them either equal or unequal.)

The genus _Chitonastrum_ differs from its ancestral form, _Dictyastrum_, by
the bifurcation of the distal ends of the arms. The few species of this
genus are partly {537}regular (like _Dictyastrum_), partly bilateral (like
_Rhopalastrum_). If the number of species increases much, these two
subgenera may be separated into two genera: _Chitonastrella_ corresponding
to the former, _Chitonastromma_ to the latter.

Subgenus 1. _Chitonastrella_, Haeckel.

_Definition._--All three arms of the same size and form, equidistant;
fundamental form of the shell therefore an equilateral triangle.

1. _Chitonastrum triglochin_, n. sp.

All three arms equal and equidistant. Each arm has the form of an isosceles
triangle, twice as high as broad; the truncated apex of the triangle is
inserted into the large central disk, whilst its distal base (four times as
broad) is divided by a deep incision (half as long as the arm). Each arm
with ten to twelve joints, simple in its basal half, double in its distal
half. Axes of the six branches straight. (Resembles _Trigonastrum
regulare_, Pl. 43, fig. 16, but differs in the absence of a patagium.)

_Dimensions._--Radius of each arm 0.24, greatest breadth 0.11, basal
breadth 0.03.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

2. _Chitonastrum bathybium_, n. sp.

All three arms equal and equidistant, in the basal two-thirds simple,
rectilinear, three times as long as broad, in the distal third forked, both
branches equal, straight, blunt, half as broad as the basal part.

_Dimensions._--Radius of each arm 0.18, basal breadth 0.04, breadth of the
branches 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Subgenus 2. _Chitonastromma_, Haeckel.

_Definition._--One odd arm different in size or form from the two other
arms, which are paired; distance between them different; fundamental form
of the shell therefore a bilateral, isosceles triangle.

3. _Chitonastrum jugatum_, n. sp. (Pl. 43, fig. 14).

_Dictyastrum jugatum_, Haeckel, 1881, Prodromus et Atlas (pl. xliii. fig.
14).

Arms very different; odd arm club-shaped, twice as long as broad, at the
blunt distal end twice as broad as at the base; its axis is perpendicular
to the common axis of both paired arms, which are only two-thirds as long,
not so broad, and in the distal half divided into two branches; the
anterior branch is straight, nearly horizontal, the posterior shorter and
curved backwards.

{538}_Dimensions._--Radius of the odd arm 0.2, of the paired arms 0.15;
distal breadth of the former 0.08, basal breadth 0.04; breadth of the
paired arms 0.04.

_Habitat._--South Atlantic, Station 325, surface.

4. _Chitonastrum dicranoides_, n. sp.

All three arms in the basal half simple, nearly square, in the distal half
forked; branches straight, blunt. Odd arm twice as large as the paired
arms; angle between the latter larger than the angles between them and the
odd arm. (The form of the arms resembles _Dicranastrum furcatum_, Pl. 47,
fig. 2.)

_Dimensions._--Radius of the odd arm 0.24, of the paired arms 0.12; basal
breadth 0.06.

_Habitat._--North Atlantic, Station 353, surface.

5. _Chitonastrum lyra_, n. sp. (Pl. 43, fig. 15).

_Dictyastrum lyra_, Haeckel, 1881, Prodromus et Atlas (pl. xliii. fig.
15).

All three arms forked and nearly of the same size, but different in form
and position. The distance between the branches of the two paired arms is
only one-fourth of the distance between them and the odd arm. Each arm in
the basal two-thirds is simple, with eleven to twelve transverse septa, in
the distal third forked, each branch with four to five transverse septa.
The branches of each arm are curved convexly one to another, ending
obtusely. The axis of the simple proximal part is straight in the odd arm,
in the paired arms curved concavely towards the middle line. In the figured
specimen, which I observed living in Portofino (in September 1880), the
central chamber of the central disk and the first surrounding ring were
filled with the nucleus of the cell; both external rings were filled (like
all chambers of the arms) with pink oil-globules of the red central
capsule. From the mantle, enveloping the shell, radiated innumerable fine
pseudopodia (much too short in the figure), and between the two paired arms
a long "sarcode-flagellum."

_Dimensions._--Radius of each arm 0.16; greatest breadth of the odd arm
0.04; basal breadth of the paired arms 0.02; distance of both branches of
each arm 0.08.

_Habitat._--Mediterranean, Portofino, near Genoa, Haeckel.

Genus 232. _Trigonastrum_,[271] n. gen.

_Definition._--#Porodiscida# with three forked, chambered arms, connected
by a patagium. (Arms and angles between them either equal or unequal.)

The genus _Trigonastrum_ differs from the preceding _Chitonastrum_, its
ancestral form, in the development of a patagium between the arms. It bears
therefore to the latter the same relation that _Euchitonia_ does to
_Rhopalastrum_.

{539}Subgenus 1. _Trigonastrella_, Haeckel.

_Definition._--All three arms of the same size and form, equidistant;
fundamental form of the shell therefore a regular, equilateral triangle.

1. _Trigonastrum regulare_, n. sp. (Pl. 43, fig. 16).

_Chitonastrum regulare_, Haeckel, 1881, Prodromus et Atlas (pl. xliii.
fig. 16).

All three arms equal and equidistant. Each arm has the form of an isosceles
triangle, twice as high as broad, the truncated apex of which is inserted
into the large circular central disk, whilst its distal base (four times as
broad) is divided by a deep incision (half as long as the arm). Each arm
with ten to twelve joints. Patagium between the arms nearly complete,
spongy. (Differs from _Chitonastrum triglochin_ mainly in the possession of
a patagium.)

_Dimensions._--Radius of each arm 0.24, greatest breadth 0.11, basal
breadth 0.03; length of the sides of the regular triangle 0.45.

_Habitat._--Pacific, central area, Station 274, surface.

Subgenus 2. _Trigonastromma_, Haeckel.

_Definition._--One odd arm different in form or size from the other two
arms, which are paired; distance between them different; fundamental form
of the shell therefore an isosceles triangle.

2. _Trigonastrum krohnii_, Haeckel.

_Euchitonia krohnii_, Haeckel, 1862, Monogr. d. Radiol., p. 507.

All three arms different; distance between the two paired arms smaller than
their distance from the odd arm. Odd arm with six joints. Each paired arm
with seven joints, increasing in breadth towards the distal end. Odd arm
and one paired arm forked at the end, the other paired arm simple. Patagium
nearly complete. (The asymmetry in this form may perhaps be an individual
anomaly, as also in _Myelastrum anomalum_, Pl. 47, fig. 9.)

_Dimensions._--Radius of the arms about 0.13 to 0.16, breadth 0.05 to 0.08.

_Habitat._--North Atlantic, Funchal, Madeira, Krohn, surface.

3. _Trigonastrum gegenbauri_, Haeckel.

_Euchitonia gegenbauri_, Haeckel, 1862, Monogr. d. Radiol., p. 506, Taf.
xxxi. figs. 2, 3.

Arms different; distance between the paired arms larger than their distance
from the odd arm, which is one-third shorter. Odd arm egg-shaped, simple,
with seven joints, undivided. Both paired arms equal, with ten joints, in
the distal third forked. Patagium nearly complete.

_Dimensions._--Radius of the odd arm 0.14, greatest breadth 0.08; radius of
the paired arms 0.2.

_Habitat._--Mediterranean (Messina), Haeckel, surface.

{540}Genus 233. _Stauralastrum_,[272] n. gen.

_Definition._--#Porodiscida# with four simple, undivided, chambered arms,
without a patagium; quadrangular shell a regular cross, with four equal
arms placed at right angles.

The genus _Stauralastrum_ is the most simple form of the Tessarastrida, or
of those Porodiscida in which the margin of the central disk is armed with
four chambered arms. In _Stauralastrum_ these four arms are quite simple
and equal, without a patagium, separated by four right angles, so that the
whole shell represents a regular rectangular cross. If we connect the
distal points of the arms by lines, we get a complete square. (In my
Prodromus, 1881, the species of this genus were united with _Hagiastrum_,
which genus I now retain for the simple bilateral Tessarastrida.)

Subgenus 1. _Stauralastrella_, Haeckel.

_Definition._--Ends of the arms blunt, without terminal spines.

1. _Stauralastrum cruciforme_, n. sp. (Pl. 45, fig. 6).

Arms very thin, nearly linear, four to five times as long as broad, of
equal breadth at the base and at the truncated distal end; their breadth
equals one-third of the radius of the central disk. Edges of the arms
parallel.

_Dimensions._--Radius of each arm 0.8, breadth 0.016.

_Habitat._--South Pacific, Station 293, surface.

2. _Stauralastrum lanceolatum_, n. sp.

Arms lanceolate, three times as long as broad, in their middle part three
times as broad as at both ends; their greatest breadth nearly equals the
diameter of the central disk. (The arms have the same form as in
_Euchitonia lanceolata_, Pl. 43, fig. 9.) Edges of the arms convex.

_Dimensions._--Radius of each arm 0.3, greatest breadth (in the middle
part) 0.08.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

3. _Stauralastrum ordo_, n. sp.

Arms trapezoid, about as long as broad, twice as broad at their truncated
distal end as at the base; their basal breadth equals the radius of the
central disk, which exhibits two to three rings. (The arms have nearly the
same form as those in _Hagiastrum mosis_, Pl. 45, fig. 3.) Edges of the
arms rectilinear, divergent towards the ends.

_Dimensions._--Radius of each arm 0.12, basal breadth 0.04, distal breadth
0.08.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

{541}4. _Stauralastrum clavigerum_, n. sp.

Arms twice as long as broad, in their distal half lenticular, nearly
circular, twice as broad as in their square proximal half; their distal
breadth equals the diameter of the central disk, which exhibits three to
four rings. Edges of the arms concave.

_Dimensions._--Radius of each arm 0.2, basal breadth 0.04, distal breadth
0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

5. _Stauralastrum dilatatum_, n. sp.

Arms of nearly equal length and breadth, at their convex distal end three
times as broad as at their narrow base; their distal breadth three times as
large as the radius of the central disk, which exhibits three to four
rings. (Resembles _Histiastrum quadrigatum_, Pl. 46, fig. 3, but has no
patagium.) Edges of the arms concave.

_Dimensions._--Radius of each arm 0.15, basal breadth 0.04, terminal
breadth 0.12.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

Subgenus 2. _Stauralastromma_, Haeckel.

_Definition._--Ends of the arms with one or more terminal spines.

6. _Stauralastrum rhopalophorum_, n. sp. (Pl. 45, fig. 1).

_Hagiastrum rhopalophorum_, Haeckel, 1881, Prodromus, p. 460.

Arms cylindrical, eight times as long as broad at their base, at their
distal end club-shaped, three times as broad as at their base; their distal
breadth twice as large as the diameter of the central disk, which exhibits
two to three rings. Surface thorny, with larger spines towards the end, and
one radial, very strong, angular terminal spine. Edges of the arms
parallel.

_Dimensions._--Radius of each arm (without terminal spine) 0.32, basal
breadth 0.03, terminal breadth 0.1.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

7. _Stauralastrum antiquum_, n. sp.

_Rhopalastrum_ sp. Bury, 1862, Polycystins of Barbados, pl. xiv. fig. 5.

Arms six times as long as broad at their base, in their distal half nearly
spherical, three times as broad as in their cylindrical basal half; their
distal breadth nearly equals the diameter of the central disk which
exhibits three to four rings. On the end of each arm one strong, angular,
terminal spine. (Differs from the preceding species by larger central disk
and stouter arms, also by less developed spines.) Edges of the arms
parallel.

_Dimensions._--Radius of each arm 0.25, basal breadth 0.04, terminal
breadth 0.12.

_Habitat._--Fossil in the Barbados rocks; and living in the depths of the
Central Pacific, Station 266, depth 2750 fathoms.

{542}8. _Stauralastrum staurolonche_, n. sp.

Arms four times as long as broad at their base, gradually increasing
towards their truncated end, which is one and a half times as broad as
their base; their distal breadth equals the radius of the central disk,
which exhibits four to five rings. At the end of each arm is a very strong
conical terminal spine. (Resembles _Histiastrum quaternarium_, _Abhandl. k.
Akad. Wiss. Berlin_, 1875, Taf. xxiv. fig. 3, but has no patagium.) Edges
of the arms rectilinear, divergent.

_Dimensions._--Radius of each arm 0.25, basal breadth 0.045, distal breadth
0.07.

_Habitat._--Fossil in the Barbados rocks; and living in the depth of the
Equatorial Atlantic, Station 348, depth (2450) fathoms.

9. _Stauralastrum horridum_, n. sp.

Arms three times as long as broad at their base, gradually increasing
towards their rounded end, which is twice as broad as their base, their
distal breadth equals the diameter of the central disk, which exhibits four
to five rings. Surface thorny, at the distal end of each arm is a group of
twenty to twenty-five smaller and five to six larger, straight, conical
spines. Edges of the arms rectilinear, divergent.

_Dimensions._--Radius of each arm 0.15, basal breadth 0.05, distal breadth
0.1.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

Genus 234. _Hagiastrum_,[273] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with four simple, undivided, chambered arms,
without a patagium; quadrangular shell bilateral, two opposite arms of the
main axis (or principal arms) different from the two others (or lateral
arms).

The genus _Hagiastrum_, as here defined, was formerly united by me with the
foregoing _Stauralastrum_, but differs from it by the bilateral or
symmetrical form. Whilst in the latter all four arms and the four angles
between them are equal, they are here differentiated into pairs.

Subgenus 1. _Hagiastrella_, Haeckel.

_Definition._--Both longitudinal arms of equal size and form.

1. _Hagiastrum buddhae_, n. sp. (Pl. 45, fig. 5).

Cross rectangular. Both longitudinal arms of equal size, twice as long as
the transverse arms; all arms smooth, club-shaped, twice as broad at their
globose distal part as at their base, each with three large conical
terminal spines.

{543}_Dimensions._--Radius of the principal arms 0.4, of the lateral arms
0.2; basal breadth 0.06, distal breadth 0.12.

_Habitat._--Indian Ocean, Belligemma, Ceylon, surface, Haeckel.

2. _Hagiastrum bramae_, n. sp.

Cross rectangular. Both longitudinal arms of equal size, one and a half
times as long as the transverse arms; all arms thorny, club-shaped, at
their pear-shaped distal part three times as broad as at their base,
provided with numerous conical spines, one larger terminal spine at their
distal point. (Resembles _Stauralastrum rhopalophorum_, Pl. 45, fig. 1, but
is distinguished by the different size of the arm-pairs, and by the
stronger spines.)

_Dimensions._--Radius of the principal arms 0.3, of the lateral arms 0.2;
basal breadth 0.04, distal breadth 0.12.

_Habitat._--Indian Ocean, off Maldive Islands, surface, Haeckel.

Subgenus 2. _Hagiastromma_, Haeckel.

_Definition._--The two longitudinal arms different in size or form.

3. _Hagiastrum mosis_, n. sp. (Pl. 45, fig. 3).

Cross rectangular. All four arms nearly isosceles, triangular, at their
narrow base half as broad as at their truncated, concavely fluted, distal
end. The posterior principal arm with twelve to thirteen joints, twice as
long as the anterior arm, which has six to seven joints and is one and a
half times as long as the two lateral arms (with four to five joints).

_Dimensions._--Radius of the posterior arm 0.3, of the anterior 0.15, of
the lateral arms 0.1; basal breadth 0.05, terminal breadth 0.1.

_Habitat._--Mediterranean (Smyrna), surface, Haeckel.

4. _Hagiastrum mohammedis_, n. sp.

Cross with unequal angles, the anterior little smaller than the posterior.
All four arms club-shaped, thorny, three times as broad at their globose
distal end as at their narrow base, and furnished with ten to twelve
conical spines. Posterior principal arm twice as long as the anterior, and
four times as long as the rudimentary lateral arms.

_Dimensions._--Radius of the posterior arm 0.4, of the anterior 0.2, of the
lateral arms 0.1; basal breadth 0.02 to 0.03, distal breadth 0.06 to 0.1.

_Habitat._--Philippine Sea, Samboangan, Station 200, surface.

5. _Hagiastrum christi_, n. sp.

Cross with unequal angles, the anterior somewhat smaller than the
posterior. All four arms of similar form, lanceolate, in their middle twice
as broad as at either obtuse end. The posterior {544}principal arm with
twelve joints, one and a half times as long as the anterior (with nine
joints) and twice as long as the two lateral arms (each with six joints).
The form and structure of the arms in this species are nearly the same as
in _Tessarastrum straussi_ (Pl. 45, fig. 8); but the arms are broader in
the middle, and are not connected by a patagium.

_Dimensions._--Radius of the principal posterior arm 0.2, of the anterior
0.15, of each lateral arm 0.1; greatest breadth (in the width) 0.01, basal
breadth 0.03.

_Habitat._--North Atlantic, Færöe Channel, Gulf Stream, surface, John
Murray.

Genus 235. _Histiastrum_,[274] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Porodiscida# with four simple, undivided, chambered arms,
connected by a patagium; square shell a regular cross, with four equal arms
and four right angles between them.

The genus _Histiastrum_, quite insufficiently characterised by Ehrenberg
(1847), was afterwards (1875) illustrated by the figures of two different
fossil species. One of these, _Histiastrum ternarium_, with three arms,
belongs to _Hymeniastrum_; the other, _Histiastrum quaternarium_, is here
retained as the true, typical representative species of the genus. It
differs from its ancestral form _Stauralastrum_, by the possession of a
patagium, from _Tessarastrum_ by the regular square form of the shell.

Subgenus 1. _Histiastrella_, Haeckel.

_Definition._--Distal ends of the arms blunt, without terminal spines.

1. _Histiastrum quadrigatum_, n. sp. (Pl. 46, fig. 3).

Arms at their distal end nearly as broad as long, and four times as broad
as at their narrow base; their lateral edges concave, their terminal edge
convex, without spines. Each arm is divided by seven to eight convex
transverse septa into eight to nine simple, broad chambers. Central disk
with three to four rings, about as broad as the fifth chamber. Patagium
complete, connecting all the lateral edges of the arms.

_Dimensions._--Radius of each arm 0.15, basal breadth 0.03, terminal
breadth 0.12.

_Habitat._--Equatorial Atlantic, Station 347, surface.

2. _Histiastrum excisum_, n. sp.

Arms four times as long as broad at their base, and twice as broad at their
rounded blunt distal end as at their base; their lateral edges rectilinear,
divergent. Central disk with three to four rings, {545}somewhat broader
than their distal end. Patagium incomplete, connecting only the basal half
of the arms, with three to four concave chamber-rows, on the margin
concave. (May be regarded as _Euchitonia mülleri_, with four arms.)

_Dimensions._--Radius of each arm 0.2, basal breadth 0.05, distal breadth
0.1.

_Habitat._--Atlantic, surface, Canary Islands.

3. _Histiastrum velatum_, n. sp. (Pl. 46, fig. 4).

_Dictyastrum velatum_, Haeckel, 1879, Atlas (pl. xlvi. fig. 4).

Arms pear-shaped, rapidly increasing from their narrow base, nearly
circular, little longer than broad; each with eight to nine transverse
chamber-rows; their lateral edges at their base concave, at their end
circular. Central disk with three to four rings, somewhat smaller than one
arm. Patagium complete, with six to seven radial beams, filling out
perfectly the intervals between the arms. A peculiar girdle of finer
network and equal breadth surrounds the whole equatorial periphery of the
disk, and gives it the appearance of a square with rounded corners.

_Dimensions._--Radius of each arm 0.2, basal breadth 0.03, distal breadth
0.12.

_Habitat._--South Atlantic, Station 330, surface.

Subgenus 2. _Histiastromma_, Haeckel.

_Definition._--Distal ends of the arms spiny, furnished with one or more
terminal spines.

4. _Histiastrum quaternarium_, Ehrenberg.

_Histiastrum quaternarium_, Ehrenberg, 1875, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 74, Taf. xxiv. figs. 3, 4.

Arms six times as long as broad at their base, with rectilinear, little
divergent edges; at their truncated distal end a little broader, with one
single, very strong, conical, terminal spine. Central disk with four to
five rings, somewhat broader than the arm. Patagium incomplete, enveloping
only the basal half of the arms.

_Dimensions._--Radius of each arm 0.18, basal breadth 0.03, distal breadth
0.05.

_Habitat._--Fossil in the rocks of Barbados.

5. _Histiastrum gladiatum_, n. sp.

_Astromma_ sp., Bury, 1862, Polycystins of Barbados, pl. v. fig. 1.

Arms triangular, eight times as long as broad at their base; at their
distal end three times as broad as at their base, with rectilinear,
divergent edges; their truncated end with a large conical, terminal spine.
Central disk with four to five rings, broader than the arms. Patagium
incomplete, enveloping only the basal half of the arms.

{546}_Dimensions._--Radius of each arm 0.12, basal breadth 0.013, distal
breadth 0.04.

_Habitat._--Fossil in the rocks of Barbados; and living in the depth of the
Central Pacific, Station 268, depth 2900 fathoms.

6. _Histiastrum boseanum_, n. sp. (Pl. 46, fig. 1).

Arms linear, twelve times as long as broad at their base (at their distal
end twice as broad as at their base), with rectilinear, parallel edges. The
club-shaped end thickened, dentate, with two lateral rows of strong teeth
in the equatorial plane, and with one very stout, angular, terminal spine.
Central disk with two to three rings, broader than the arms. Patagium
incomplete, with four to five concave chamber-rows, enveloping only the
basal half of the arms. I call this splendid species in honour of Dr. Graf
Bose, the great friend of nature and patron of the University of Jena.

_Dimensions._--Radius of each arm 0.25, basal breadth 0.02, distal breadth
0.05.

_Habitat._--Indian Ocean, Madagascar, Rabbe.

7. _Histiastrum coronatum_, n. sp.

_Stephanastrum_ sp., Bury, 1862, Polycystins of Barbados, pl. iv. fig. 1.

Arms linear, eight times as long as broad, with rectilinear parallel edges;
the thickened, nearly spherical, distal end three times as broad, with five
strong conical spines, one middle (perradial) larger and two smaller on
each side of it. Central disk with two to three rings, of the same breadth
as the terminal knot of the arms. Patagium incomplete, square, enveloping
the arms, with the exception of the knot.

_Dimensions._--Radius of each arm 0.16, basal breadth 0.02, terminal
breadth 0.06.

_Habitat._--Fossil in the rocks of Barbados.

8. _Histiastrum circulare_, n. sp.

_Stephanastrum_ sp., Bury, 1862, Polycystins of Barbados, pl. xxiii. fig.
1.

Arms linear in their inner half, egg-shaped and three times as broad in
their outer half, with ten to twelve strong terminal spines, the middle
(perradial) larger, in all three times as long as broad. Central disk with
three to four rings, broader than their distal knobs. Patagium nearly
complete, circular, enveloping the whole arms, with exception of the
outermost end.

_Dimensions._--Radius of each arm 0.22, basal breadth 0.02, terminal
breadth 0.06.

_Habitat._--Fossil in the rocks of Barbados; and living in the depths of
the Central Pacific, Station 268, depth 2900 fathoms.

9. _Histiastrum pentadiscus_, n. sp. (Pl. 46, fig. 2).

Arms in their inner half linear, twice as long as broad, in their outer
half circular, three times as broad, with the same structure as the central
disk, exhibiting three concentric rings around one {547}central chamber.
Patagium complete, spongy, with radiating beams, enveloping the whole disk,
with the exception of the outermost end of the arms, which is armed with
twelve to sixteen strong conical spines, the middle (perradial) spine much
larger.

_Dimensions._--Radius of each arm (without terminal spine) 0.18, basal
breadth 0.025, terminal breadth 0.08.

_Habitat._--South Atlantic, Station 333, surface.

Genus 236. _Tessarastrum_,[275] n. gen.

_Definition._--#Porodiscida# with four simple, undivided, chambered arms,
connected by a patagium; quadrangular shell bilateral, two opposite arms of
the main axis (or principal arms) different from the two others (or lateral
arms).

The genus _Tessarastrum_, formerly united by me with _Histiastrum_, differs
from the latter in its bilateral or symmetrical form, and bears therefore
the same relation to it that _Hagiastrum_ does to _Stauralastrum_.

Subgenus 1. _Tessarastrella_, Haeckel.

_Definition._--Both longitudinal arms of equal size and form.

1. _Tessarastrum straussii_, n. sp. (Pl. 45, fig. 8).

_Histiastrum straussii_, Haeckel, 1881, Prodromus et Atlas (pl. xlv. fig.
8).

Cross not rectangular. Both principal arms of equal size and form, four
times as long as broad, and twice as long as the broader lateral arms; the
former with ten to eleven, the latter with five to six joints, separated by
convex transverse septa. Distal ends of the arms blunt. Axes of the smaller
arms not perpendicular to that of the larger arms; therefore the anterior
angles between them smaller than the posterior angles. Patagium between the
arms incomplete. I call this remarkable species after the great German
philosopher David Strauss.

_Dimensions._--Radius of the principal arms 0.24, of the lateral arms 0.12;
greatest breadth (in the middle) of the former 0.05, of the latter 0.06.

_Habitat._--North Pacific, _Hyalonema_-ground, March 5, 1875.

2. _Tessarastrum spinozæ_, n. sp.

Cross rectangular. Both principal arms of equal size and form, ten times as
long as broad, and twice as long as the lateral arms, which are only five
times as long as broad. All arms linear, at their distal end club-shaped,
and armed with twenty to thirty very strong angular spines. Patagium
incomplete, enveloping only the basal half of the arms. (Resembles
_Histiastrum boseanum_, {548}Pl. 46, fig. 1, but differs in the broader
arms and the unequal size of both pairs.) I call this species after the
great monistic philosopher Benedictus Spinoza.

_Dimensions._--Radius of the principal arms 0.3, of the lateral arms 0.16;
basal breadth 0.025, distal breadth 0.05.

_Habitat._--South Atlantic, Station 333, surface.

3. _Tessarastrum brunonis_, n. sp. (Pl. 45, fig. 9).

_Histiastrum brunonis_, Haeckel, 1881, Prodromus et Atlas (pl. xlv. fig.
9).

Cross rectangular. Both principal arms of equal size and form, three times
as long as broad, each with ten joints, three times as long as the lateral
arms, which are nearly square, with four joints. All arms rounded, at their
truncated end little broader than at their base. No spines. Patagium
complete, envelops the whole shell, and is composed of two parallel
lattice-lamellæ on each side of the flat disk, which are connected by very
fine perpendicular bars. This is shown clearly in fig. 9, Pl. 45, where the
disk is seen from the edge. I call this species after the great Italian
philosopher Giordano Bruno.

_Dimensions._--Radius of the principal arms 0.22, of the lateral arms 0.12;
basal breadth 0.04, distal breadth 0.05.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

Subgenus 2. _Tessarastromma_, Haeckel.

_Definition._--The two principal arms of different size or form.

4. _Tessarastrum democriti_, n. sp. (Pl. 45, fig. 7).

_Histiastrum democriti_, Haeckel, 1881, Prodromus et Atlas (pl. xlv. fig.
7).

Cross not rectangular; the two anterior angles smaller than the two
posterior. All four arms club-shaped, twice as broad at their rounded
obtuse distal end as at their base, of unequal length. Posterior principal
arm one and a fourth times as long as the posterior, and one and two-thirds
as long as the lateral arms. Patagium incomplete, enveloping only the basal
half of the arms. I call this species after the great Greek philosopher
Democritus.

_Dimensions._--Radius of the posterior arm 0.3, of the anterior 0.25, of
each lateral arm 0.22; basal breadth 0.05, distal breadth 0.1.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 237. _Stephanastrum_,[276] Ehrenberg, 1847, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 54.

_Definition._--#Porodiscida# with four simple, undivided, chambered arms,
connected on the distal ends by a spongy, square or rhomboidal, patagial
girdle (or a patagium with four large, interbrachial openings). Shell
either regular or bilateral (with equal or unequal arms).

{549}The genus _Stephanastrum_, founded (1847) by Ehrenberg for the very
peculiar _Stephanastrum rhombus_, differs from the nearly allied foregoing
genera in the imperfect development of the peculiar patagium, connecting
only the distal ends of the four arms, while it is absent at their base.
Two new species, different from _Stephanastrum rhombus_ by the regular
square form, were found in the Challenger collection.

Subgenus 1. _Stephanastrella_, Haeckel.

_Definition._--All four arms of the cross have the same size.

1. _Stephanastrum quadratum_, n. sp. (Pl. 46, fig. 5).

All four arms of the same size, six times as long as broad at their base,
ending with a strong, short, four-sided pyramidal spine. In the outer half
of each arm are two opposite lateral spongy wings, which form an
equilateral triangle, and from union of the bases of the four triangles
arises the peculiar patagium, which forms a square with four large
interbrachial openings.

_Dimensions._--Radius of each arm 0.25, basal breadth 0.035; length of the
sides of the square patagium 0.3.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

2. _Stephanastrum capitatum,_ n. sp. (Pl. 44, fig. 1).

All four arms of the same size, five times as long as broad at their base,
at their distal end with a spongy, nearly spherical capitulum of twice
their breadth, provided with a very strong, angular, pyramidal, terminal
spine (half as long as the arm). All four arms connected by a square
patagium, arising immediately below the capitula, and perforated by four
large interbrachial openings.

_Dimensions._--Radius of each arm (without the terminal spine) 0.25, basal
breadth 0.05; length of the sides of the square patagium 0.3.

_Habitat._--Pacific, central area. Station 265, depth 2900 fathoms.

Subgenus 2. _Stephanastromma_, Haeckel.

_Definition._--Two opposite arms of the cross larger than the two others.

3. _Stephanastrum rhombus,_ Ehrenberg.

_Stephanastrum rhombus_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvi. fig.
33; Abhandl. d. k. Akad. d. Wiss. Berlin, 1875, Taf. xxv. fig. 1.

Two arms of the longitudinal axis one and a third times as long as two arms
of the transverse axis. All four arms linear, about eight times as long as
broad, at their distal end somewhat {550}thickened, club-shaped, with a
pyramidal terminal spine. The ends of the arms are connected by a
riband-shaped, straight, spongy patagium of the same breadth as the arms.
Between the rhomboidal patagium and the arms remain four large rectangular
triangles as interbrachial openings.

_Dimensions._--Radius of the longer arms 0.2, of the shorter 0.15; basal
breadth 0.02; length of the sides of the rhombic patagium 0.25.

_Habitat._--Fossil in the rocks of Barbados.

Genus 238. _Dicranastrum_,[277] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with four forked, spongy, or chambered arms,
without a patagium; shell regular (not bilateral), with four equal arms
crossed at right angles.

The genus _Dicranastrum_ comprises a number of very remarkable, hitherto
unknown, Euchitonida, which are rather common in the Pacific (mainly on the
surface), and characterised by the bifurcation of the cross-arms of the
regular square shell. It bears therefore to its probable ancestral form,
_Stauralastrum_, the same relation that in the triradiate Euchitonida
_Chitonastrum_ does to _Dictyastrum_. The arms are commonly of very
delicate structure, more or less spongy.

Subgenus 1. _Dicranaster_, Haeckel.

_Definition._--Each cross-arm with two simple branches.

1. _Dicranastrum furcatum_, n. sp. (Pl. 47, fig. 2).

Arms simply forked, three times as long as broad at their base, with eight
blunt ends of the fork-branches. The simple proximal half of each arm about
the same size as each branch of the dichotomous distal part, twice as long
as broad. Edges of the arms ragged.

_Dimensions._--Radius of each arm 0.38, basal breadth 0.12; breadth of the
forked part 0.3.

_Habitat._--Pacific, central area, Station 271, surface.

2. _Dicranastrum dichotomum_, n. sp.

Arms simply forked, four times as long as broad at their base; each arm
with two blunt branches. The simple proximal part of each arm is three
times as long and twice as broad as each branch of the dichotomous distal
part. Ends of the arms blunt, truncated.

_Dimensions._--Radius of each arm 0.35, basal breadth 0.08; breadth of the
forked part 0.2.

_Habitat._--South Pacific, Station 281, surface.

{551}3. _Dicranastrum cornutum_, n. sp. (Pl. 45, fig. 2).

_Hagiastrum cornutum_, Haeckel, 1879, Atlas (pl. xlv. fig. 2).

Four arms simply forked, four times as long as broad; each arm with two
triangular diverging branches ending in strong conical spines. The simple
basal part of each arm about of the same length as each branch of the
distal part, twice as long as broad. Distance between the terminal spines
of each arm nearly as great as its length.

_Dimensions._--Radius of the arm 0.24, basal breadth 0.05; breadth across
the bifurcation 0.12.

_Habitat._--South Atlantic, Station 325, surface.

4. _Dicranastrum antilope_, n. sp.

Four arms simply forked, three times as long as broad; each arm with two
lanceolate diverging branches, ending in strong angular spines. The simple
basal part of each arm is twice as long as each branch of the distal part.
Distance of the two terminal spines of each arm scarcely half as great as
its length.

_Dimensions._--Radius of the arm 0.32, basal breadth 0.11; breadth of the
branches 0.07.

_Habitat._--Tropical Atlantic (off Ascension), Station 342, surface.

Subgenus 2. _Tricranastrum_, Haeckel, 1881, Prodromus, p. 460.

_Definition._--Each cross arm with three terminal branches, one middle
(perradial) and two lateral (adradial) branches.

5. _Dicranastrum wyvillei_, n. sp. (Pl. 47, fig. 3).

_Tricranastrum wyvillei_, Haeckel, 1879, Natürl. Schöpfungsgesch., p.
705, Taf. xvi. fig. 5.

Arms trifid, one and a half times as long as broad; each arm with three
blunt terminal branches of equal size. The simple basal part of each arm
twice as long as the trifid distal part. (The central capsule depicted in
fig. 3, Pl. 47, has the same form as the skeleton, and is only a little
smaller.)

_Dimensions._--Radius of the arm 0.4, basal breadth 0.12, greatest breadth
(in the distal part) 0.24.

_Habitat._--Pacific, central area, Station 271, surface.

6. _Dicranastrum tricuspis_, n. sp.

Arms trifid, twice as long as broad; each arm with three pointed terminal
branches, ending in strong conical spines, the middle branch somewhat
larger than the other two. The simple basal part of each arm three times as
long as the trifid distal part.

_Dimensions._--Radius of the arm 0.3, basal breadth 0.06, greatest breadth
(in their distal part) 0.15.

_Habitat._--North Pacific, Station 244, surface.

{552}7. _Dicranastrum trifarium_, n. sp.

Arms trifid, three times as long as broad at their base; each arm with
three pointed terminal branches, ending in strong sulcate spines; the
middle branch twice as large as the two others. The simple basal part of
each arm two and a half times as long as the trifid distal part.

_Dimensions._--Radius of the arm 0.35, basal breadth 0.03, greatest breadth
(in the distal part) 0.05.

_Habitat._--South Pacific, Station 298, surface.

Subgenus 3. _Tetracranastrum_, Haeckel.

_Definition._--Each cross-arm with four terminal branches, the two
fork-branches being again bifurcated.

8. _Dicranastrum bifurcatum_, n. sp. (Pl. 47, figs. 1, 1_a_).

Arms doubly forked or quadripartite, six times as long as broad at their
base; each arm in its proximal half simple, three times as long as broad;
in its distal half doubly forked; the secondary branches with blunt,
roundish ends, nearly as large as the primary branches. Central disk (fig.
1_a_) with three concentric rings around the central chamber; from its
periphery radiate thin radial beams in the spongy framework of the delicate
arms. The central capsule has the same form as the skeleton, and is only a
little smaller.

_Dimensions._--Radius of the arm 0.45, basal breadth 0.08; breadth of the
terminal branches 0.03.

_Habitat._--North Pacific, east of Japan, Station 241, surface.

Genus 239. _Myelastrum_,[278] Haeckel, 1881, Prodromus, p. 460.

_Definition._--#Porodiscida# with four forked, spongy, or chambered arms,
without a patagium; shell bilateral, with paired different arms; two equal
anterior arms of different shape from the two equal posterior arms.

The genus _Myelastrum_ differs from the foregoing _Dicranastrum_, the
ancestral form, by the twofold differentiation of the four arms. Whilst in
the latter all four arms are equal, separated by equal angles, here the two
anterior arms are constantly different from the two posterior. The lateral
angles between the two arm-pairs are equal, while the posterior and the
anterior angle (between the two arms of each pair) are more or less
different. The shell assumes, therefore, a very characteristic bilateral
form, similar to the "quadricorn cross" of the grey central substance in
the transverse section of the human medulla spinalis. Though the spongy
shell is commonly a most delicate and {553}thin disk it nevertheless
reaches unusual dimensions, its diameter in some species being more than a
millimetre.

Subgenus 1. _Myelastrella_, Haeckel.

_Definition._--Posterior arms simple, undivided; anterior arms lobated or
cleft, with one or more incisions at the distal end.

1. _Myelastrum medullare_, n. sp. (Pl. 47, fig. 13).

Anterior arms bifid, nearly square, with a shallow incision at their broad
truncated end. Posterior arms somewhat smaller, nearly triangular, with
simple blunt ends. Sagittal constriction three-fourths as large as the
transverse one. Surface smooth.

_Dimensions._--Radius of the anterior arms 0.36, of the posterior 0.3;
longitudinal constriction 0.24, transverse 0.36.

_Habitat._--Pacific, central area, Station 271, surface.

2. _Myelastrum spinale_, n. sp.

Anterior arms bifid, twice as long as broad, with a deep incision at their
truncated end. Posterior arms slender, half as large, with simple blunt
ends. Sagittal constriction one and a half times as large as the
transverse. Surface spiny.

_Dimensions._--Radius of the anterior arms 0.4, of the posterior 0.2;
longitudinal constriction 0.3, transverse 0.2.

_Habitat._--North Pacific, Station 248, surface.

3. _Myelastrum heteropterum_, n. sp. (Pl. 47, fig. 8).

Anterior arms trifid, about as long as broad, with two incisions at their
broad truncated end. Posterior arms about half as large, simple, with
rounded blunt ends. Sagittal constriction two-thirds as large as the
transverse. Surface bristly.

_Dimensions._--Radius of the anterior arms 0.65, of the posterior 0.035;
longitudinal constriction 0.4, transverse 0.6.

_Habitat._--South Pacific, Station 291, surface.

Subgenus 2. _Myelastromma_, Haeckel.

_Definition._--All four arms (posterior as well as anterior) lobated or
cleft.

4. _Myelastrum octocorne_, n. sp. (Pl. 47, fig. 12).

All four arms with a deep incision in their distal half; anterior arms
broader, but shorter than the posterior; in the anterior arms the two lobes
are of the same size, in the posterior arms the {554}median lobe is longer
than the lateral lobe. Sagittal constriction little smaller than the
transverse. Surface of the disk rough.

_Dimensions._--Radius of the anterior arms 0.45, of the posterior 0.62;
longitudinal constriction 0.25, transverse 0.3.

_Habitat._--Pacific, central area, Station 271, surface.

5. _Myelastrum farfalla_, n. sp. (Pl. 47, fig. 10).

All four arms with a shallow incision at their distal end. Anterior arms
broader, but shorter than the posterior; in each arm the anterior lobe is
shorter than the posterior. Sagittal constriction smaller than the
transverse. Surface smooth.

_Dimensions._--Radius of the anterior arms 0.4, of the posterior 0.6;
longitudinal constriction 0.5, transversal stricture 0.6.

_Habitat._--Pacific, central area, Station 274, surface.

6. _Myelastrum papilio_, n. sp. (Pl. 47, fig. 6).

All four arms with a shallow incision at their distal end. Anterior arms
triangular, of the same breadth, but of the double length of the square
posterior arms. Sagittal and transverse constrictions of the same length.
Surface ciliated. (Resembles a butterfly.)

_Dimensions._--Radius of the anterior arms 0.7, of the posterior 0.4;
longitudinal and transverse constrictions 0.4.

_Habitat._--North Pacific, near Japan, Station 241, surface.

7. _Myelastrum decaceros_, n. sp. (Pl. 47, fig. 7).

Anterior arms bifid, with one shallow incision at their distal end.
Posterior arms of the same length, but much broader, trifid, with two
incisions (one larger and one smaller). Sagittal and transverse
constrictions of the same length. Surface spiny.

_Dimensions._--Radius of all four arms 0.4; longitudinal and transverse
constrictions 0.35.

_Habitat._--Pacific, central area, Station 272, surface.

8. _Myelastrum dodecaceros_, n. sp. (Pl. 47, figs. 11, 11_a_).

Anterior arms trifid, with two shallow incisions at their distal end.
Posterior arms about one-third larger, also trifid, with one deeper
anterior and one shallower posterior incision. Sagittal constriction much
longer than the transverse. Surface ciliated, covered with numerous thin
and long radial bristles (fig. 11_a_).

_Dimensions._--Radius of the anterior arms 0.5, of the posterior 0.7;
longitudinal constriction 0.5, transverse 0.35.

_Habitat._--Pacific, central area, Station 270, surface.

{555}9. _Myelastrum ciliatum_, n. sp.

Anterior arms trifid, with two shallow incisions. Posterior arms somewhat
smaller, also trifid, with two deeper incisions. Sagittal constriction a
little larger than the transverse. Margin of the disk ciliated, with radial
bristle-shaped spines, as prolongations of the inner radial beams, arising
from the central disk.

_Dimensions._--Radius of the anterior arms 0.6, of the posterior 0.5;
longitudinal constriction 0.4, transverse 0.35.

_Habitat._--South Pacific, Station 288, surface.

10. _Myelastrum lobatum_, n. sp.

Anterior arms somewhat broader but shorter than the posterior arms. Each
arm four-lobed, with three terminal shallow incisions of nearly equal size.
Sagittal constriction smaller than the transverse. Surface of the disk
bristly.

_Dimensions._--Radius of the anterior arms 0.4, of the posterior 0.5;
longitudinal constriction 0.3, transverse 0.35.

_Habitat._--North Pacific, Station 253, surface.

11. _Myelastrum rotula_, n. sp.

Anterior arms little broader than the posterior, but of the same length.
Each arm four-lobed, with three terminal deep incisions of equal size.
Sagittal and transverse constrictions equal. The whole disk nearly
circular, resembles a wheel with sixteen spokes. Surface smooth.

_Dimensions._--Radius of all four arms 0.5; longitudinal and transverse
constrictions 0.3.

_Habitat._--Pacific, central area, Station 274, surface.

12. _Myelastrum giganteum_, n. sp.

Anterior arms somewhat broader and shorter than the posterior. Each arm
four-lobed, with three shallow terminal incisions, the middle incision
twice as deep as the two laterals. Sagittal constriction a little larger
than the transverse. Margin ciliated, with radial bristle-shaped spines as
prolongations of the inner radial beams, proceeding from the central disk
(as in _Myelastrum dodecaceros_, Pl. 47, figs. 11, 11_a_).

_Dimensions._--Radius of the anterior arms 0.6, of the posterior 0.8;
longitudinal constriction 0.6, transverse 0.5.

_Habitat._--Pacific, central area, Station 271, surface.

{556}13. _Myelastrum anomalum_, n. sp. (Pl. 47, fig. 9).

All four arms of different size and form; anterior arms broader, posterior
longer; one anterior arm trifid, the three other arms bifid; length of the
branches unequal; all four angles between the arms unequal. (This anomalous
form, seen only once, may be an individual abnormality.)

_Dimensions._--Radius of the arms 0.4 to 0.7; constrictions 0.3.

_Habitat._--North Pacific, Station 237, surface.

Genus 240. _Pentalastrum_,[279] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Porodiscida# with five simple, undivided, chambered arms,
without a patagium.

The genus _Pentalastrum_ opens the small series of Euchitonida, in which
the shell is not provided with three or four arms, as usual, but with five.
All forms of this little group are rare. Some species resemble in their
external form and in the articulation of their arms certain forms of
Asterida. In _Pentalastrum_, the most simple genus, the five arms are
simple, not forked, and without a patagium. It can be derived from
_Dictyastrum_ or _Stauralastrum_ by increase of the number of arms.

Subgenus 1. _Pentalastrella_, Haeckel.

_Definition._--All five arms equal, with equal angles between them. Shell a
regular pentagon.

1. _Pentalastrum asteracanthion_, n. sp.

All five arms equal, club-shaped, at their thickened obtuse end three times
as broad as at their base, twice as long as broad. Angles between the arms
equal.

_Dimensions._--Radius of each arm 0.2, basal breadth 0.03, distal breadth
0.08.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Pentalastrum astropecten_, n. sp.

All five arms equal, with five to six distinct, simple joints, the basal
joint two-thirds as broad as the terminal joint, which bears a strong
conical spine. Angles between the arms equal. (Resembles _Pentinastrum
asteriscus_, Pl. 44, fig. 2, but has no patagium.)

_Dimensions._--Radius of each arm 0.14, basal breadth 0.024, distal breadth
0.036.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

{557}Subgenus 2. _Pentalastromma_, Haeckel.

_Definition._--Arms of different sizes, one odd arm larger than the two
others; the opposite odd angle generally different from the four other
angles.

3. _Pentalastrum ophidiaster_, n. sp. (Pl. 44, fig. 3).

Arms nearly triangular, at their obtuse truncated distal end twice as broad
as at their base. Four arms equal, with five joints each; the fifth arm
twice as long, with seven joints. Angles between the arms nearly equal; the
odd angle a little larger.

_Dimensions._--Radius of the larger odd arm 0.25, of the four smaller arms
0.15; basal breadth 0.035, distal breadth 0.07.

_Habitat._--Pacific, central area, Station 274, surface.

4. _Pentalastrum cometa_, n. sp.

Arms nearly cylindrical, at their obtuse truncated distal end one and a
half times as broad as at their base. Posterior odd arm very large, with
eleven joints, about three times as long as the two lateral arms (with five
joints each) and four times as long as the two anterior arms (with three
joints each). Angles between the paired arms different; the two lateral
angles smaller than the two posterior, and these smaller than the odd
anterior angle.

_Dimensions._--Radius of the odd posterior arm 0.5, of the lateral arms
0.25, of the anterior arms 0.18; basal breadth 0.05, distal breadth 0.08.

_Habitat._--South Pacific, Station 288, surface.

Genus 241. _Pentinastrum_,[280] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Porodiscida# with five simple, undivided, chambered arms,
connected by a patagium.

The genus _Pentinastrum_ differs from the foregoing _Pentalastrum_ only in
the development of a patagium or connecticulum between the arms, and bears
therefore the same relation to it that _Histiastrum_ does to
_Stauralastrum_, or _Hymeniastrum_ to _Dictyastrum_.

1. _Pentinastrum asteriscus_ , n. sp. (Pl. 44, fig. 2).

All arms equal, twice as long as broad, at their base two-thirds as broad
as at their truncated distal end, which bears a strong, pyramidal, terminal
spine. Each arm is divided by five transverse septa into six joints or
chambers, and each of these by a radial beam into a pair of chambers. The
five radial beams arise from the innermost chamber of the central disk, and
end in the five terminal {558}spines. The diameter of the central disk is
larger than the length of the arms. The angles between the arms are equal
and filled up by an incomplete patagium, so that the whole disk forms a
regular pentagon with five concave sides.

_Dimensions._--Radius of each arm (without terminal spine) 0.14; breadth at
their base 0.02, at their terminal joint 0.03; radius of the central disk
0.06.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Pentinastrum goniaster_, n. sp.

? _Stephanastrum_ sp., Bury, 1862, Polycystins of Barbados, pl. xx. fig.
1.

All five arms equal, four times as long as broad, club-shaped, at their
globose distal end twice as broad as at their base, and armed with a strong
conical terminal spine. Diameter of the central disk equals only one-third
of the length of the arms. The articulation of the spongy arms is somewhat
obscure. Patagium complete, totally fills up the interbrachial spaces, so
that the whole disk forms a regular pentagon with five rectilinear sides,
except that the terminal spines project at the corners.

_Dimensions._--Radius of each arm 0.18, basal breadth 0.02, distal breadth
0.04; radius of the central disk 0.05.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms; also fossil in
the rocks of Barbados.

Genus 242. _Pentophiastrum_,[281] n. gen.

_Definition._--#Porodiscida# with five forked chambered arms, without a
patagium.

The genus _Pentophiastrum_ differs from the two preceding genera by the
bifurcation of the five arms, and can be derived either from _Pentalastrum_
by the ramification of the distal ends of the arms, or from the similar
_Myelastrum_ by the increase in the number of arms.

1. _Pentophiastrum dicranastrum_, n. sp.

All five arms equal, with equal angles between them. Each arm in the basal
half simple, in the distal half forked; both branches of it equal, with
obtuse ends. (This regular species resembles _Dicranastrum furcatum_, Pl.
47, fig. 2, but with five rays instead of four; also the form of the arms
is more slender and the edges smooth.)

_Dimensions._--Radius of each arm 0.25, breadth 0.05.

_Habitat._--Equatorial Atlantic, Station 347, depth 2250 fathoms.

Subgenus _Pentophiastromma_, Haeckel.

_Definition._--Arms of different size, one odd arm opposite to the angle
between both arm-pairs.

{559}2. _Pentophiastrum caudatum_, n. sp. (Pl. 47, fig. 5).

Arms in pairs different; four arms in the basal half simple, in the distal
half forked; the fifth (posterior) odd arm simple, undivided, cylindrical;
the anterior pair a little smaller than the posterior; the neighboring
branches of the two pairs on each side larger than the two others. Axes of
the arms and their branches straight.

_Dimensions._--Radius of the arms about 0.5, breadth 0.12.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

3. _Pentophiastrum forcipatum_, n. sp. (Pl. 47, fig. 4).

Arms in pairs different, all in the basal two thirds simple, in the distal
third forked. Only in the posterior (odd) arm both branches are equal, in
the four others unequal. The common axis of the posterior lateral pair is
horizontal, perpendicular to the median line; the axes of the anterior pair
are pincer-like, concavely curved towards the median line or principal
axis.

_Dimensions._--Radius of the arms about 0.5, breadth 0.14.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Genus 243. _Hexalastrum_,[282] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Porodiscida# with six simple chambered arms, without a
patagium.

The genus _Hexalastrum_, together with the following _Hexinastrum_,
encloses those _Euchitonida_ in which the number of the chambered arms
surrounding the central disk amounts to six. This is the highest number of
these articulated marginal appendages which is reached in any #Discoidea#.
Formerly (1881) in my Prodromus, p. 459, I supposed that the same number
was reached also by one Coccodiscid, and called this genus _Hexactura_.
Afterwards I was convinced that this form was also a _Hexalastrum_.

1. _Hexalastrum palmanthum_, n. sp.

All six arms equal, with equal angles between them. Each arm club-shaped,
three times as long as broad, twice as broad at the thickened distal end as
at the base, without a terminal spine.

_Dimensions._--Radius of each arm 0.2, basal breadth 0.02, distal breadth
0.06.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

2. _Hexalastrum crinanthum_, n. sp.

All six arms equal, with equal angles between them. Each arm club-shaped,
four times as long as broad, at the egg-shaped distal end three times as
broad as in the linear basal part, provided {560}with numerous short
conical spines and one longer terminal spine. (Resembles _Stauralastrum
rhopalophorum_, Pl. 45, fig. 1, but with six rays instead of four.)

_Dimensions._--Radius of each arm 0.3, basal breadth 0.03, distal breadth
0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

Subgenus _Hexalastromma_, Haeckel.

_Definition._--Arms more or less different in size or form; shell
bilateral.

3. _Hexalastrum orchidaceum_, n. sp. (Pl. 44, fig. 5).

_Hexactura orchidacea_, Haeckel, 1881, Prodromus, p. 459.

Arms different in length, so that two unequal opposite odd arms determine
the main axis, and the four other arms lie on both sides of this as two
different pairs. The proportion of their relative length is the
following:--anterior lateral arms five, anterior odd arm six; posterior
lateral arms seven, posterior odd arm eight. Each arm is club-shaped, two
to three times as long as broad, and divided into six to eight joints by
five to seven transverse septa; its distal end is armed with a terminal
spine and twice as broad as its base.

_Dimensions._--Radius of the posterior odd arm 0.4, of the anterior odd arm
0.3; of the posterior lateral pair 0.35, of the anterior lateral pair 0.25;
basal breadth 0.08, distal breadth 0.16.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Genus 244. _Hexinastrum_,[283] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Porodiscida# with six simple, undivided, chambered arms,
connected by a patagium.

The genus _Hexinastrum_ differs from its ancestral form _Hexalastrum_ by
the development of a patagium between the arms. The only observed species
is regular.

1. _Hexinastrum geryonidum_, n. sp. (Pl. 44, fig. 4).

_Hexalastrum geryonidum_, Haeckel, 1879, Atlas (pl. xliv. fig. 4).

Disk quite regular with six radii; all six arms of the same size and form,
at their broad, convexly rounded, smooth end five times as broad as at
their narrow base, and little longer than broad. Each arm is divided by
eight transverse septa into nine simple joints or chambers of the same
height; the breadth of the distal chambers increases rapidly. The regular,
hexagonal, central disk exhibits four concentric rings around the central
chamber. Patagium between the arms incomplete, with concavely fluted edge.

_Dimensions._--Radius of each arm 0.15, basal breadth 0.016, distal breadth
0.08; radius of the central disk 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

{561}Family XXII. #PYLODISCIDA#, n. fam. (Pl. 48, figs. 12-20).

_Definition._--#Discoidea# without phacoid shell, with flat discoidal
shell, in which a simple spherical central chamber is surrounded by one or
two concentric triradial girdles; each girdle with three gates, separated
by three simple arm-chambers. Surface of the disk with three open or
latticed gates on each flat side.

The family #Pylodiscida# represents a new small but interesting group of
#Discoidea#, which exhibits rather complex affinities to different groups
of #Sphærellaria#. In my Prodromus (1881, p. 464) I had enumerated only two
genera of this family, _Triopyle_ and _Hexapyle_, and had united them with
_Tetrapyle_ and allied genera in the family Pylonida. Indeed, the
resemblance of skeletal structure in the two groups is very great. The most
simple forms of both groups exhibit a simple spherical latticed central
chamber, which is surrounded by few latticed chambers of similar size and
form, separated by open gates. But in the Pylonida these chambers are
opposite in pairs, and form together a complete lattice-girdle around the
central chamber, whereas in the Pylodiscida the chambers are not opposite
in pairs in one axis, and form therefore only latticed half girdles, which
arise from the central chamber like radial arms, and may perhaps better be
called "arm-chambers"; their number is constantly three. The free open
spaces between these three arm-chambers form three gates, comparable to the
two or four gates of _Amphipyle_, _Tetrapyle_, &c., and become afterwards
closed by lattice-work in a similar way in both groups. A more important
difference between them is indicated by the further mode of growth. The
Pylonida build new girdles in all three dimensive planes (alternating in
the transverse, lateral and sagittal planes); their geometric fundamental
form is therefore the "lentellipsis" or the "triaxial ellipsoid." The
Pylodiscida, however, grow only at the periphery of the discoidal shell in
one single plane (the equatorial plane); their fundamental form is
therefore the biconvex lens or the flat disk (a shortened cylinder). This
important difference is my deciding motive, in separating the latter from
the former and in regarding the Pylodiscida as true #Discoidea#, the more
so as they can easily be derived from _Archidiscus_, the fundamental and
ancestral form of the Porodiscida.

One single form of _Archidiscus_ seems to be of peculiar importance in this
relation, viz., _Archidiscus hexoniscus_ (Pl. 48, fig. 10). In this species
the simple central chamber is surrounded by a latticed ring or girdle,
composed of six equal chambers of the same size and form, all lying in the
same plane with the central chamber. In a nearly allied species, viz.,
_Archidiscus pyloniscus_, the six ring-chambers are different, three
smaller (with denser network) alternating with three larger (of looser
network); if we imagine the network of the latter reduced to a marginal bar
we get _Triopyle_, and if also this bar disappear by reduction we get
_Triolena_, the most simple form of the Pylodiscida. {562}Even _Archidiscus
pyloniscus_ may possibly represent the same form among the Porodiscida as
_Triodiscus lenticula_ among the Pylodiscida; this important form indicates
clearly the close affinity of the two families.

If we take the latter, nearly identical form as the common starting point
of both families of Cyclodiscaria, then probably _Triopyle_ and _Triolene_
must be regarded as retrograde forms, derived from _Triodiscus_ in the one
case, from _Archidiscus_ in the other, by reduction of three interradial
arm-chambers, whilst three perradial only remain. But it is also possible
that the most simple form, _Triolene_, originated independently from some
_Cenosphæra_, three simple radial chambers, like the latticed central
chamber, being derived from the latter by apposition in three equidistant
radii, whilst three other radii between them remained free. In this case
the other genera of Pylodiscida are derived from their ancestral form
_Triolene_.

Adopting this latter view, we find that all eight genera of Pylodiscida,
here distinguished, may be regarded as following members of a continuous
series. If the three simple arm-chambers of _Triolene_, surrounding the
equal central chamber, become united at their distal ends by a concentric
equatorial ring, then originates _Triopyle_; and this graduates into
_Triodiscus_ by fenestration of the three open gates between the three
latticed arms. Whilst these three genera form together the subfamily
Triopylida, a second family, Hexapylida, is composed of three other
analogous genera, in which the same process of development becomes
repeated.

_Pylolena_, the most simple form of Hexapylida, arises from _Triodiscus_ by
the development of three new arm-chambers (of the second order) which are
apposed at the distal end of the three primary arm-chambers (of the first
order) in the same radius. If the distal ends of these three secondary
arm-chambers become united by a concentric latticed ring or girdle, we get
_Hexapyle_ (with six open gates, two in each radius), and if its six gates
become afterwards closed by loose lattice-work, we arrive at _Pylodiscus_
(a repetition of _Triodiscus_).

A third subfamily, Discopylida, is formed by the building of a chambered
equatorial girdle around the margin of _Pylodiscus_. This girdle has quite
the same structure as the similar chambered rings or girdles of the
Porodiscida and Coccodiscida. Between the two sieve-plates of the disk
surface is enclosed a variable number (twelve to twenty-four or more) of
chambers, imperfectly separated by radial beams, which connect the margin
of the _Pylodiscus_-shell with an outer peripheral concentric ring. In
_Discozonium_ this marginal ring is perfect, whilst in _Discopyle_ it is
interrupted by a peculiar large opening, a "marginal osculum" surrounded by
a corona of spines, quite the same remarkable formation which we
encountered in _Ommatodiscus_ among the Porodiscida.

All Pylodiscida are therefore triradial (with three perradial arms and
three interradial gates between them), and many of them have a great
resemblance to certain triradial Porodiscida and Spongodiscida, perhaps not
only a morphological resemblance, but also {563}a true phylogenetic
relation. But it is remarkable that we do not find further forms of
development in this family, by multiplication either of the arm-chambers
(further growth in the three perradii) or of the concentric chambered rings
(in the periphery of the disk margin).

The central capsule of the Pylodiscida is constantly flat, discoidal, and
enclosed between the two sieve-plates of the surface. Its form is either
circular or triangular.

_Synopsis of the Genera of the Pylodiscida._

-------------------------------------------------------------------------
I. Subfamily Triopylida.
Three gates between three simple arm-chambers.
-----------------------------------------------------------------------
Three gates open, without a barring equatorial
girdle, 245. _Triolena_.

Three gates barred by a { Gate-faces simple, 246. _Triopyle_.
latticed equatorial {
girdle. { Gate-faces latticed, 247. _Triodiscus_.
-------------------------------------------------------------------------
II. Subfamily Hexapylida.
Six gates between three double arm-chambers (three inner and three
outer); no chambered marginal girdle.
-------------------------------------------------------------------------
Three outer gates open, without a barring equatorial
girdle, 248. _Pylolena_.

{ Both faces of the outer
Three outer gates barred { gates simple, 249. _Hexapyle_.
by a latticed (second) {
equatorial girdle. { Both faces of the outer
{ gates latticed, 250. _Pylodiscus_.
-------------------------------------------------------------------------
III. Subfamily Discopylida.
Six gates between three double arm-chambers.
------------------------------------------------------------------------
{ No peculiar osculum on
In the equatorial plane { the margin of the disc, 251. _Discozonium_.
on the margin of the {
Pylodiscus-shell is a { One pecuiar osculum (with
chambered equatorial { a corona of spines) on
girdle. { the margin of the disc, 252. _Discopyle_.

Subfamily 1. TRIOPYLIDA, Haeckel.

_Definition._--#Pylodiscida# with a simple, spherical or lenticular,
central chamber, surrounded by three simple arm-chambers, which are
separated by three notches or gates.

Genus 245. _Triolena_,[284] n. gen.

_Definition._--#Pylodiscida# with a simple, spherical or lenticular,
central chamber, surrounded by three simple arm-chambers. Notches between
the three arms open.

The genus _Triolena_ is the most simple form of all Pylodiscida, and must
be regarded as their common ancestral form, from an ontogenetic as well as
a phylogenetic point of view. The small shell is composed of a simple,
spherical or lenticular, latticed, central chamber, and of three simple,
surrounding equal arms, which are also simple latticed chambers, lie in the
equatorial plane, and are separated by three equal angles or open gates.

{564}1. _Triolena primordialis_, n. sp. (Pl. 48, fig. 12).

Arm-chambers trapezoid, nearly square, of the same size as the circular,
lenticular, primordial, central chamber. Surface of the disk smooth.

_Dimensions._--Diameter of the shell 0.045, of the central chamber 0.015,
of each arm 0.015.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Triolena tribelone_, n. sp.

Arm-chambers lanceolate, of the same breadth as and twice the length of the
triangular, central chamber; at the pointed end of each arm is a conical
terminal spine (in the equatorial plane). Surface of the disk thorny.

_Dimensions._--Diameter of the shell 0.05, of the central chamber 0.015;
length of the arms 0.02, breadth 0.016.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

3. _Triolena trispinosa_, n. sp.

Arm-chambers ovate, in the basal half nearly as broad as the hexagonal
central chamber, at the pointed distal end with a strong conical radial
spine of double the length. Surface rough.

_Dimensions._--Diameter of the shell 0.055, of the central chamber 0.02.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

4. _Triolena hexabelone_, n. sp.

Arm-chambers nearly triangular, at the base half as broad as the hexagonal
central chamber, at the truncate distal end one and a half times as broad,
and armed with two radial conical spines (in the equatorial plane). Surface
of the disk smooth.

_Dimensions._--Diameter of the shell 0.06, of the central chamber 0.02.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

5. _Triolena trigonalis_, n. sp.

Arm-chambers nearly triangular, at the base half as broad as the circular
central chamber, at the concave lunulate distal end twice as broad, and
armed with four conical radial spines (two on each side of the equatorial
plane). Surface thorny.

_Dimensions._--Diameter of the shell 0.05, of the central chamber 0.015.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

{565}Genus 246. _Triopyle_,[285] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylodiscida# with a simple, spherical or lenticular,
central chamber, surrounded by three simple arm-chambers. Notches between
the three arms transformed into gates by a connecting equatorial girdle.

The genus _Triopyle_ differs from the preceding _Triolene_ in the
development of a simple ring or latticed equatorial girdle, which connects
the distal ends of the three arm-chambers, and transforms the open notches
between them into three gates. The ring may be circular, triangular, or
hexagonal.

1. _Triopyle circulus_, n. sp.

Disk circular, three times as broad as the hexagonal central chamber. Three
arm-chambers trapezoidal, at the convex distal end as broad, at the base
half as broad as the three circular gates between them. Surface smooth. No
marginal spines on the girdle.

_Dimensions._--Diameter of the disk 0.045, of the gates 0.015.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

2. _Triopyle hexagona_, n. sp. (Pl. 48, fig. 13).

Disk hexagonal, three times as broad as the circular central chamber. Three
arm-chambers trapezoidal, at the truncated distal end as broad, at the base
half as broad as the three triangular gates between them. Surface smooth.
No marginal spines.

_Dimensions._--Diameter of the disk 0.05, of the gates 0.02.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

3. _Triopyle trigona_, n. sp.

Disk triangular, four times as broad as the circular central chamber. Three
arm-chambers nearly triangular, at the narrow base half as broad, at the
distal end twice as broad as the circular or roundish gates. Surface
smooth. On the margin (in the equatorial plane) three strong pyramidal
spines (at the end of the arms).

_Dimensions._--Diameter of the disk 0.04, of the gates 0.015.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

4. _Triopyle cordigera_, n. sp.

Disk hexagonal, four times as broad as the hexagonal central chamber. Three
arm-chambers triangular, at the base half as broad as at the truncated
distal end, about the same size as the {566}three heart-shaped gates. On
the six corners of the margin (which forms a regular hexagon) are six
pyramidal radial spines, as prolongations of the arm-edges.

_Dimensions._--Diameter of the disk 0.05, of the gates 0.02.

_Habitat._--Indian Ocean, Ceylon, surface, Haeckel.

5. _Triopyle renigera_, n. sp.

Disk hexagonal, five times as broad as the circular central chamber. Three
arm-chambers trapezoidal, at the base one-third, at the distal end
two-thirds as broad as the three kidney-shaped gates. On the six corners of
the margin (which forms an irregular hexagon) six conical radial spines, as
prolongations of the arm-edges.

_Dimensions._--Diameter of the disk 0.06, of the gates 0.02.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

6. _Triopyle spinigera_, n. sp.

Disk roundish, triangular, four times as broad as the triangular central
chamber, which is armed with three radial spines between the arms.
Arm-chambers club-shaped, at the narrow base one-fourth, at the distal end
half as broad as the square gates. On the margin twelve large conical
spines, two opposite on each face of the distal end of each arm-edge. Three
smaller radial spines on the three corners of the girdle (in the same
interradial meridian planes as the three spines of the central chamber).
Compare _Triodiscus spinosus_.

_Dimensions._--Diameter of the disk 0.05, of the gates 0.02.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

Genus 247. _Triodiscus_,[286] n. sp.

_Definition._--#Pylodiscida# with a simple, spherical or lenticular,
central chamber, surrounded by three simple arm-chambers. Notches between
the three arms closed by lattice-work and by an equatorial girdle.

The genus _Triodiscus_ differs from the preceding _Triopyle_ in the
development of loose lattice-work on both sides of the discoidal shell.
This network closes the gates and transforms the whole shell into a
fenestrated lens. The singular species of _Triodiscus_ correspond to
certain species of _Triopyle_.

1. _Triodiscus lenticula_, n. sp.

Disk circular, lenticular, three times as broad as the central chamber.
Three arm-chambers trapezoidal, of the same size and form as the three
gates between them, which are closed by a loose {567}delicate network
(differs from _Triopyle circulus_ by the production of the two convex
latticed plates, which envelop the whole lens). Margin smooth.

_Dimensions._--Diameter of the disk 0.045, of the gates 0.015.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

2. _Triodiscus trigonus_, n. sp.

Disk triangular, four times as broad as the central chamber. Three
arm-chambers at the base half as broad, at the distal end twice as broad as
the roundish gates. Surface smooth. On the three corners of the margin (in
the arm-radius) three strong spines. (Differs from _Triopyle trigona_ only
in the loose framework closing the gates.)

_Dimensions._--Diameter of the disk 0.04, of the gates 0.015.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

3. _Triodiscus spinosus_, n. sp. (Pl. 48, fig. 14).

Disk subcircular, four times as broad as the triangular central chamber.
Three arm-chambers club-shaped, at the base one-third, at the distal end
two-thirds as broad as the semicircular gates. Surface thorny. On the
margin fifteen larger radial spines, three on the corners of the disk (in
the radius of the gates), twelve on the two faces of the arms ends (two
opposite on the edge of each end.) (Differs from _Triopyle spinigera_
mainly by the delicate hexagonal network closing the gates.)

_Dimensions._--Diameter of the disk 0.05, of the gates 0.02.

_Habitat._--North Atlantic, Canary Islands (Lanzerote, Haeckel).

Subfamily 2. HEXAPYLIDA, Haeckel.

_Definition._--#Pylodiscida# with triopyle-shaped medullary shell,
surrounded by three distal arm-chambers, which are separated by three open
notches or gates.

Genus 248. _Pylolena_,[287] n. gen.

_Definition._--#Pylodiscida# with triopyle-shaped medullary shell,
surrounded by three distal arm-chambers. Notches between the three arms
open.

The genus _Pylolena_ opens the series of the Hexapylida, or of those
Pylodiscida in which the centre of the shell is formed by a tri-radiated
medullary shell like _Triopyle_. In the equatorial plane of this
triopyle-shaped disk are developed on its margin three distal arm chambers,
as prolongations of the three arms of _Triopyle_, but much larger. In
_Pylolene_ the three angles or notches between the distal arms remain open,
repeating the form of _Triolene_.

{568}1. _Pylolena inermis_, n. sp.

Arms club-shaped, at the base half as broad, at the rounded distal end as
broad as the triopyle-shaped medullary shell. Surface and margin smooth.

_Dimensions._--Diameter of the cortical shell 0.15, of the medullary shell
0.05.

_Habitat._--Equatorial Atlantic, Station 347, surface; Central Pacific,
Station 272, surface.

2. _Pylolena armata_, n. sp. (Pl. 48, fig. 15).

Arms trapezoid, at the base half as broad, at the distal end twice as broad
as the triopyle-shaped medullary shell. Surface and margin thorny. Twelve
strong conical spines at the distal end of the three arms, two opposite on
both faces of the arm-edges.

_Dimensions._--Diameter of the cortical shell 0.2, of the medullary shell
0.06.

_Habitat._--South Atlantic, Station 325, surface.

Genus 249. _Hexapyle_,[288] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylodiscida# with triopyle-shaped medullary shell,
surrounded by three distal arm-chambers. Notches between the three arms
transformed into gates by a connecting equatorial girdle.

The genus _Hexapyle_ differs from the preceding _Pylolene_ in the
development of an equatorial ring or latticed girdle, which connects the
free extremities of the three distal ends and transforms the open notches
between them into gates. It simulates therefore the formation of
_Triopyle_, from which it differs by duplication of the arm-joints and of
the gates (in each radius occur one proximal and one distal gate).

1. _Hexapyle triangula_, n. sp.

Cortical shell triangular, with three rounded corners, three times as broad
as the triangular triopyle-shaped medullary shell. Surface smooth or rough,
but not spiny. Three arms two-thirds as broad as the three egg-shaped gates
of each side.

_Dimensions._--Diameter of the cortical shell (or length of one side of the
triangle) 0.15, of the medullary shell 0.05; breadth of the gates 0.06, of
the bridges between them 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Hexapyle sexangula_, n. sp.

Cortical shell hexagonal, with six equal sides, four times as broad as the
triangular triopyle-shaped medullary shell. Surface rough, but not spiny.
Three arms (on their smallest part) half as broad as the three triangular
gates (on their broadest part).

{569}_Dimensions._--Diameter of the cortical shell 0.16, of the medullary
shell 0.04; breadth of the gates 0.06, of the bridges 0.03.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

3. _Hexapyle circularis_, n. sp.

Cortical shell circular, four times as broad as the triangular,
Triopyle-shaped medullary shell. Surface smooth. Three arms half as broad
as the three kidney-shaped gates on each side of the shell.

_Dimensions._--Diameter of the cortical shell 0.2, of the medullary shell
0.05; breadth of the gates 0.08, of the bridges between them 0.04.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

4. _Hexapyle triacantha_, n. sp.

Cortical shell triangular, thorny, with three pointed corners, prolonged
into three strong conical radial spines; its diameter four times as great
as that of the medullary shell. Three arms of the same breadth as the three
egg-shaped gates of each side.

_Dimensions._--Diameter of the cortical shell (or length of one side of the
triangle, without spines) 0.2, of the medullary shell 0.05; breadth of the
gates and of the bridges between them 0.06.

_Habitat._--Indian Ocean, surface; Madagascar, Rabbe.

5. _Hexapyle hexacantha_, n. sp.

Cortical shell hexagonal, thorny, three times as broad as the medullary
shell. Six stronger conical radial spines on the six corners at equal
distances, lying in the equatorial plane of the disk. Three half girdles
half as broad as the three triangular gates of each side.

_Dimensions._--Diameter of the cortical shell 0.15, of the medullary shell
0.05; breadth of the gates 0.05, of the bridges 0.025.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

6. _Hexapyle dodecantha_, n. sp. (Pl. 48, fig. 16).

Cortical shell triangular, thorny, with rounded corners, three and a half
times as broad as the medullary shell. Twelve stronger radial spines, six
opposite in pairs on each side of the discoidal shell, as prolongations of
the lateral edges of the triangular gates, which are about the same breadth
as the half girdles between them.

_Dimensions._--Diameter of the cortical shell 0.21, of the medullary shell
0.06; breadth of the gates and of the bridges 0.06.

_Habitat._--Pacific, central area, Station 270, surface.

{570}7. _Hexapyle polyacantha_, n. sp.

Cortical shell circular, thorny, with a circle of numerous conical radial
spines in the periphery; its diameter nearly three times as great as that
of the medullary shell. Three half-girdles half as broad as the three
kidney-shaped gates on each side of the discoidal shell.

_Dimensions._--Diameter of the cortical shell 0.11, of the medullary shell
0.04; breadth of the gates 0.04, of the bridges between them 0.02.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 250. _Pylodiscus_,[289] n. gen.

_Definition._--#Pylodiscida# with _Triopyle_-shaped medullary shell,
surrounded by three distal arm-chambers. Notches between the three arms
closed by lattice-work and by an equatorial girdle.

The genus _Pylodiscus_ differs from _Hexapyle_ in the development of two
convex plates of lattice-work, which close the six open gates and transform
the disk into a biconvex fenestrated lens. _Pylodiscus_ simulates therefore
the characteristic form of _Triodiscus_; but whilst in the latter we find
only three arm-joints and three simple gates, their number is doubled in
the former.

1. _Pylodiscus triangularis_, n. sp. (Pl. 48, fig. 17).

Cortical shell triangular, with rounded corners, three times as broad as
the triangular, _Triopyle_-shaped medullary shell. Three arms about as
broad as the three egg-shaped gates between them. (Differs from _Hexapyle
triangula_ only by the delicate network of both triangular convex covering
plates, which close the open gates.) Six larger and many smaller spines on
the margin of the disk.

_Dimensions._--Diameter of the disk 0.15, of the gates 0.05.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

2. _Pylodiscus sexangularis_, n. sp.

Cortical shell hexangular, equilateral, four times as broad as the
circular, _Triopyle_-shaped medullary shell. Surface spiny. Three arms
nearly of the same form and size as the three triangular gates between
them. (Similar to _Hexapyle sexangularis_, but differing in the irregular
loose network closing the gates.)

_Dimensions._--Diameter of the disk 0.18, of the gates 0.06.

_Habitat._--North Pacific, south of Japan, Station 237, surface

{571}3. _Pylodiscus cardiopylus_, n. sp.

Cortical shell hexangular, equilateral, five times as broad as the
triangular, _Triopyle_-shaped medullary shell. Surface smooth. Three arms
triangular, at the distal end as broad as the three heart-shaped gates. On
the margin six strong pyramidal spines in the equatorial plane (adradial,
as prolongations of the lateral arm-edges).

_Dimensions._--Diameter of the disk 0.15, of the gates 0.05.

_Habitat._--Indian Ocean, Sunda Strait, Rabbe.

4. _Pylodiscus nephropylus_, n. sp.

Cortical shell circular, three times as broad as the circular,
_Triopyle_-shaped medullary shell. Surface spiny. Three arms trapezoidal,
half as broad as the three kidney-shaped gates. On the margin twelve
stronger, conical, radial spines, opposite in pairs on both sides of the
disk, disposed in six pairs at the distal end of the arms.

_Dimensions._--Diameter of the disk 0.2, of the medullary shell 0.06.

_Habitat._--Central Pacific, Station 268, depth 2900 fathoms.

Subfamily 3. DISCOPYLIDA, Haeckel.

_Definition._--#Pylodiscida# with _Triopyle_-shaped medullary shell and
_Pylodiscus_-shaped cortical shell, which is surrounded by a marginal
chambered equatorial girdle.

Genus 251. _Discozonium_,[290] n. gen.

_Definition._--#Pylodiscida# with _Triopyle_-shaped medullary shell and
_Pylodiscus_-shaped cortical shell, which is surrounded by an equatorial
chambered girdle. No peculiar osculum on the margin of the disk.

The genus _Discozonium_ and the following _Discopyle_ make up together the
small group of the Discopylida, or those Pylodiscida in which a discoidal
shell like _Pylodiscus_ is surrounded by a marginal equatorial girdle; this
girdle is divided into twelve to twenty-four or more chambers by radial
beams, which are the external prolongations of the radial marginal spines
of _Pylodiscus_. The latter genus bears therefore to _Discozonium_ the same
relation as _Sethodiscus_ in the other #Discoidea# does to _Lithocyclia_,
or _Phacodiscus_ to _Coccodiscus_.

1. _Discozonium cyclonium_, n. sp.

Disk circular, lenticular, with smooth margin, three times as broad as the
triangular, _Triopyle_-shaped medullary shell. Three gates of the cortical
shell kidney-shaped, twice as broad as the three {572}arms. Chambered
equatorial girdle with twenty to twenty-four subregular chambers (similar
to _Discopyle osculata_, Pl. 48, fig. 19, but without any marginal
osculum).

_Dimensions._--Diameter of the disk 0.2, of the cortical shell 0.14, of the
medullary shell 0.07.

_Habitat._--Central Pacific, Station 265, depth 2900 fathoms.

2. _Discozonium trigonium_, n. sp.

Disk triangular, with thorny margin, four times as broad as the triangular,
_Triopyle_-shaped medullary shell. Three gates of the cortical shell
egg-shaped, scarcely as broad as the three arms. Chambered equatorial
girdle with twenty-four to thirty irregular chambers.

_Dimensions._--Diameter of the disk 0.24, of the cortical shell 0.2, of the
medullary shell 0.06.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

3. _Discozonium hexagonium_, n. sp. (Pl. 48, fig. 18).

Disk hexagonal, with spiny margin, four times as broad as the triangular,
_Triopyle_-shaped medullary shell. Three gates of the cortical shell
kidney-shaped, one and a half times as broad as the arms. Chambered
equatorial girdle with twelve large regular chambers; the radial beams
between them are prolonged into twelve strong pyramidal marginal spines
(three perradial on the ends of the arms, three interradial on the radii of
the gates, six adradial between the former and latter).

_Dimensions._--Diameter of the disk 0.2, of the cortical shell 0.12, of the
medullary shell 0.05.

_Habitat._--Central Pacific, Station 271, depth 2425 fathoms.

Genus 252. _Discopyle_,[291] n. gen.

_Definition._--#Pylodiscida# with _Triopyle_-shaped medullary shell and
_Pylodiscus_-shaped cortical shell, which is surrounded by an equatorial
chambered girdle. One peculiar osculum, surrounded by a corona of spines,
on the margin of the disk.

The genus _Discopyle_ differs from the preceding genus _Discozonium_ in the
development of a peculiar marginal osculum, and bears therefore to it the
same relation as, in the Porodiscida, _Ommatodiscus_ does to _Porodiscus_
(compare above, p. 500). This peculiar osculum is here also surrounded by a
corona of spines, and serves probably for the exit or outlet of a bunch of
pseudopodia or a "sarcode-flagellum." Only two species of _Discopyle_ have
been observed, which represent perhaps better two different genera; in one
species the disk is circular, in the other elliptical. In this latter the
osculum lies on one pole of the main axis.

{573}1. _Discopyle osculata_, n. sp. (Pl. 48, fig. 19).

Disk circular with spiny margin, three times as broad as the triangular,
_Triopyle_-shaped medullary shell. Three gates of the cortical shell
kidney-shaped, on the inside with an interradial spine, twice as broad as
the three pentagonal arms. Chambered equatorial girdle with twenty-four
subregular chambers, in the radius of one odd gate with a large marginal
osculum, which is as broad as the medullary shell, and surrounded by a
dense corona of twenty to thirty strong conical spines.

_Dimensions._--Diameter of the disk 0.15, of the cortical shell 0.1, of the
medullary shell 0.05, of the marginal osculum 0.06.

_Habitat._--Central Pacific, Station 272, depth 2600 fathoms.

2. _Discopyle elliptica_, n. sp. (Pl. 48, fig. 20).

Disk elliptical, four-fifths as broad as long, with spiny margin, three
times as broad as the triangular, _Triopyle_-shaped medullary shell. Three
gates of the cortical shell roundish, on the inside with an interradial
spine, little broader than the quadrangular arms. Chambered equatorial
girdle with twenty to thirty irregular chambers, on one pole of the main
axis with a large marginal osculum, which is one-third as broad as the
length of the main axis, and armed with a corona of twenty to thirty short
conical spines. The osculum does not correspond to a certain radius.

_Dimensions._--Diameter of the disk 0.15, of the cortical shell 0.08, of
the medullary shell 0.04, of the marginal osculum 0.05.

_Habitat._--Central Pacific, Station 267, depth 2700 fathoms.

Family XXIII. #SPONGODISCIDA#, Haeckel (Pl. 41, fig. 11).

_Spongodiscida et Spongocyclida_, Haeckel, 1862, Monogr. d. Radiol.,
pp. 452, 460, 469.
_Spongodiscida_, Haeckel, 1881, Prodromus, p. 461.
_Calodictya_, Ehrenberg, 1847, Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin, p. 53 (_partim_).

_Definition._--#Discoidea# without a phacoid shell, with a flat discoidal
shell, in which a simple spherical central chamber is surrounded by an
irregular spongy framework (sometimes with concentric rings around the
central chamber). Surface of the disk quite spongy, without porous
sieve-plates.

The family #Spongodiscida# is the sixth and last family of the #Discoidea#,
and bears to the other families of this group the same relation as the
family Spongosphærida does to the other #Sphæroidea#, or the Spongurida to
the other #Prunoidea#. Its characteristic structure consists in the
irregular spongy framework of the disk, and mainly in the rough, irregular
shape of its spongy surface, which is never covered with porous plates
(neither phacoid shell nor corresponding sieve-plates), as in all other
#Discoidea#. Of course a little spongy structure occurs also in many
Porodiscida {574}and even in some Coccodiscida (principally in the
peripheral part of the disk or its chambered arms); but both flat (or
convex) surfaces of the disk (at least in the central part) remain here
constantly as simple lattice-plates, whilst in all Spongodiscida the whole
surface of the disk is spongy.

When I constituted the family Spongodiscida in my Monograph (1862, pp. 452,
460) I had separated from them the Spongocyclida, exhibiting in the central
part of the disk a more or less distinct concentric arrangement of the
spongy chambers, whilst in the former the delicate spongy framework is
quite irregular, composed of branched siliceous threads, connected and
interwoven in all directions. But in all Spongocyclida the whole surface of
the spongy disk is quite as irregularly rough and deprived of smooth
sieve-plates as in all true Spongodiscida, and the more or less concentric
structure of the central part of the disk in the former (very variable and
often scarcely able to be recognised) seems not sufficient to separate both
groups; even the single genera cannot be sufficiently separated by this
character. I now therefore give up entirely the group of Spongocyclida (as
already done in my Prodromus, 1881). Nevertheless the concentric annular
structure in the dark central part of some Spongodiscida is very
interesting as transition to the Porodiscida; it indicates already that the
former are derived from the latter. Even the single genera in both
families are corresponding.

In the new system of "Polycystina," which Ehrenberg gave, 1875 (Abhandl. d.
k. Akad. d. Wiss. Berlin, p. 157), are enumerated under the _Calodictya_
four genera "with spongy disk," viz., _Spongodiscus_, _Rhopalodictyum_,
_Dictyocoryne_, _Spongaster_. Indeed these four genera, which I here
retain, are true Spongodiscida, and must be separated from the other
_Calodictya_, the greater part of which are Porodiscida. The number of
species of true Spongodiscida now amounts to sixty-seven, which I dispose
in thirteen genera.

The whole family may be divided into three subfamilies. The first of these
are the Spongophacida (corresponding to the Trematodiscida among the
Porodiscida), in which the circular margin of the spongy disk bears no
radial appendages; either the margin is quite simple, spongy
(_Spongodiscus_), or surrounded by a hyaline, solid, or porous equatorial
girdle (_Spongophacus_). The disk is either more lenticular (biconvex) or
more flat discoidal (a shortened cylinder), rarely a little biconcave
(thicker at the margin than in the centre). The spongy framework of the
solid disk is either quite irregular (_Spongodisculus_), or in the central
part with concentric circular rings (_Spongocyclia_), or in the central
part spirally convoluted (_Spongospira_).

The second subfamily, Spongotrochida, corresponds to the Stylodictyida
(among the #Porodiscida#), and is distinguished by solid radial spines on
the margin of the disk, disposed in the equatorial plane either irregularly
or regularly (after the same order as in the other families of
#Discoidea#).

The third subfamily, Spongobrachida, correspond perfectly to the
Euchitonida {575}(among the Porodiscida), bearing on the disk-margin two,
three, or four spongy arms, commonly disposed regularly in the equatorial
plane. Here also occurs the peculiar formation of a "patagium," or of an
interbrachial spongy framework different from that of the arms, which
connects the arms like a web-membrane in the equatorial plane.

The spongy framework exhibits in all these Spongodiscida no remarkable
differences, being everywhere composed of fine branched solid siliceous
threads, interwoven in all directions, with irregular meshes of very
different size.

The central capsule of all Spongodiscida is filled up with the same spongy
framework which covers also both its sides. It grows according to the
enveloping skeleton, but remains constantly smaller. The form of the
central capsule is circular (lenticular or discoidal) in the Spongophacida
and Spongotrochida, whilst in the Spongobrachida it enters into the radial
spongy arms, developed from the margin of the spongy disk.

_Synopsis of the Genera of the Spongodiscida._

{Spongy disk with simple
I. Subfamily { margin (without peculiar
Spongophacida. { equatorialgirdle), 253. _Spongodiscus_.
Spongy disk {
without radial {Spongy disk with a peculiar
appendages. { (solid or porous) equatorial
{ girdle, 254. _Spongophacus_.

{ {Two opposite
{ { spines, 255. _Spongolonche_.
II. Subfamily {Few (two, {
Spongotrochida. { three, or {Three marginal
Spongy disk with { four) radial { spines, 256. _Spongotripus_.
solid radial { spines {
spines on the { regularly {Four crossed
margin (in the { disposed. { spines, 257. _Spongostaurus_.
equatorial {
plane). { {Spines only
{Numerous (five { on the
{ to ten or { margin
{ more) radial { (equatorial), 258. _Stylotrochus_.
{ spines, {
{ often {Spines on both
{ irregularly { sides of the
{ disposed. { disk, 259. _Spongotrochus_.

{ {Without a
{Two arms, { patagium, 260. _Spongolena_.
{ opposite in {
{ one axis. {With a
{ { patagium, 261. _Spongobrachium_.
III. Subfamily {
Spongobrachida. { {Without
Spongy disk with { { a patagium, 262. _Rhopalodictyum_.
spongy radial {Three arms on {
arms on the { the margin. {With a
margin (in the { { patagium, 263. _Dictyocoryne_.
equatorial {
plane). { {Without
{ { a patagium, 264. _Spongasteriscus_.
{Four arms in {
{ cross form. {With a
{ { patagium, 265. _Spongaster_.

Subfamily 1. SPONGOPHACIDA, Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with a simple circular disk, without radial
appendages on the margin (neither solid spines nor chambered arms).

{576}Genus 253. _Spongodiscus_,[292] Ehrenberg, 1854, Monatsber. d. k.
preuss. Akad. d. Wiss. Berlin, p. 237.

_Definition._--#Spongodiscida# with a simple circular disk, without radial
appendages and without an equatorial girdle on the margin.

The genus _Spongodiscus_ represents the most simple and primitive form of
the Spongodiscida, or of those #Discoidea# in which the central disk is
more or less spongy, composed of an irregular fine framework. In my
Monograph (1862, pp. 452, 460, 469) I had separated the true _Spongodiscus_
(first described by Ehrenberg, _loc. cit._) and the _Spongocyclia_; the
former being characterised by the irregular spongy framework of the whole
disk, whilst in the latter this framework includes in the central part some
concentric circular rings (approaching _Porodiscus_). In the same way
afterwards Stöhr (1880, _loc. cit._) separated the genus _Spongospira_ as
spongy disks, which include in the central part some spiral convolutions.
But as these differences are rather inconstant and not sharply discernible,
I think it now better to regard these three forms as subgenera of
_Spongodiscus_. All three have the common simple circular disk, without any
marginal appendages.

Subgenus 1. _Spongodisculus_, Haeckel.

_Definition._--Spongy framework of the disk quite irregular, without
concentric rings or spiral convolutions.

1. _Spongodiscus mediterraneus_, Haeckel.

_Spongodiscus mediterraneus_, Haeckel, 1862, Monogr. d. Radiol., p. 461,
Taf. xii. figs. 14, 15.

Spongy disk plain on both sides, with quite an irregular framework, without
concentric rings and without radial piercing beams. Texture everywhere
uniform; meshes eight to ten times as broad as the bars.

_Dimensions._--Diameter of the disk 0.08 to 0.24, of the meshes 0.005 to
0.008.

_Habitat._--Mediterranean (Messina), Haeckel; also fossil in the Tertiary
rocks of Sicily (Grotte), Stöhr.

2. _Spongodiscus radiatus_, n. sp.

Spongy disk plain on both sides, with quite irregular framework, without
concentric rings, but with numerous piercing radial beams which are not
prolonged into marginal spines. Texture everywhere uniform; meshes once and
a half to twice as broad as the bars.

_Dimensions._--Diameter of the disk 0.12 to 0.15, of the meshes 0.002 to
0.003.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

{577}3. _Spongodiscus resurgens_, Ehrenberg.

_Spongodiscus resurgens_, Ehrenberg, 1854, Mikrogeol., Taf. xxxvB. B.
iv., fig. 16.

_Spongodiscus resurgens_, Stöhr, 1880, Palæontogr., xxvi. p. 117, Taf.
vi. fig. 11.

Spongy disk lenticular, biconvex, in the darker centre much thicker than
towards the thin periphery, with an irregular framework, without concentric
rings, but with numerous piercing radial beams. Texture equal; meshes three
to four times as broad as the bars.

_Dimensions._--Diameter of the disk 0.1 to 0.3, of the meshes 0.003 to
0.006.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface; also
fossil in the Tertiary rocks of Sicily and Barbados.

4. _Spongodiscus favus_, Ehrenberg.

_Spongodiscus favus_, Ehrenberg, 1861, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 301.

Spongy disk lenticular, biconvex, in the darker centre much thicker than
towards the periphery, with an irregular framework, without concentric
rings. Texture different, in the outer half with numerous radial beams and
loose network, the meshes of which are three to four times as large as in
the darker and denser framework of the centre.

_Dimensions._--Diameter of the disk 0.2, of the outer meshes 0.008 to
0.012, of the inner meshes 0.002 to 0.003.

_Habitat._--North Atlantic, Greenland, Færöe Channel (John Murray),
surface.

5. _Spongodiscus biconcavus_, n. sp.

Spongy disk biconcave, in the peripheral, ring-like, thickened part twice
as thick as in the hollowed central part, with an irregular framework,
without concentric rings. Texture different, in the outer half looser than
in the inner, darker part; meshes of the outer part five to six times, of
the inner twice to three times, as broad as the bars.

_Dimensions._--Diameter of the disk 0.25, of the outer meshes 0.01 to
0.012, of the inner 0.004 to 0.006.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Subgenus 2. _Spongocyclia_, Haeckel, 1862, Monogr. d. Radiol., p. 469.

_Definition._--Spongy framework of the disk in the inner part with
concentric, circular rings, in the outer part irregular.

{578}6. _Spongodiscus cycloides_, Haeckel.

_Spongodiscus cycloides_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 843.

_Spongocyclia cycloides_, Haeckel, 1862, Monogr. d. Radiol., p. 469, Taf.
xxviii. fig. 1.

Spongy disk on both sides plain, in the central part with five to ten
concentric, circular rings, in the peripheral part quite irregularly and
densely spongy. Meshes twice to four times as broad as the bars.

_Dimensions._--Diameter of the disk 0.1 to 0.2, of the meshes 0.003 to
0.006.

_Habitat._--Mediterranean (Messina), North Atlantic (Canary Islands).

7. _Spongodiscus spongocyclia_, Haeckel.

_Spongocyclia triangularis_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 119,
Taf. vii. fig. 5.

Spongy disk lenticular, in the thicker central part with eleven to twelve
circular, concentric rings, in the thinner, peripheral zone irregularly
spongy. Meshes twice to four times as broad as the bars. (The triangular
form in the specimen figured by Stöhr is accidental, produced by the broken
margin.)

_Dimensions._--Diameter of the disk 0.2 to 0.3, of the meshes 0.006 to
0.008.

_Habitat._--Fossil in Tertiary rocks of Barbados (Haeckel) and Sicily
(Stöhr).

Subgenus 3. _Spongospira_, Stöhr, 1880, Palæontogr., vol xxvi. p. 120.

_Definition._--Spongy framework of the disk in the inner part with spiral
convolutions, in the outer part irregular.

8. _Spongodiscus florealis_, Haeckel.

_Spongospira florealis_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 120,
Taf. vii. fig. 6.

Spongy disk lenticular, in the thicker central part with five to six spiral
convolutions, in the outer peripheral zone irregularly spongy. No radial
beams piercing the framework. Meshes three to four times as broad as the
bars.

_Dimensions._--Diameter of the disk 0.26, of the meshes 0.006 to 0.008.

_Habitat._--Fossil in Tertiary rocks of Sicily (Grotte), Stöhr.

9. _Spongodiscus spiralis_, n. sp.

_Spongospira spiralis_, Haeckel, 1881, Prodromus.

Spongy disk on both sides plain, with twelve to sixteen spiral convolutions
in the central part, with irregular, spongy framework in the outer part,
pierced by numerous interrupted radial beams. Meshes five to six times as
broad as the bars.

_Dimensions._--Diameter of the disk 0.2 to 0.3, of the meshes 0.01 to
0.012.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

{579}Genus 254. _Spongophacus_,[293] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with a simple circular disk, without radial
appendages, on the margin surrounded by a thin porous or solid equatorial
girdle.

The genus _Spongophacus_, represented hitherto only by a single but
interesting species, differs from _Spongodiscus_ by the peculiar girdle
which surrounds the margin of the disk in the equatorial plane. It
simulates the same formation as _Perichlamydium_ in the Porodiscida, and
resembles also _Periphæna_, &c., in the Phacodiscida.

1. _Spongophacus periphæna_, n. sp.

Spongy disk lenticular, with an irregular, dense framework, in the centre
darker than in the peripheral part. Margin of the disk very thin,
surrounded by a broad, circular girdle, lying in the equatorial plane,
about as broad as the half radius of the spongy disk. The inner part of the
girdle is perforated by numerous irregular, small pores, which pass over
gradually into the spongy meshes; the outer part is quite homogeneous,
solid, transparent, with an extremely thin margin.

_Dimensions._--Diameter of the disk 0.2; breadth of the girdle 0.05.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Subfamily 2. SPONGOTROCHIDA, Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with a circular disk, the margin of which is
armed with solid radial spines, situated in the equatorial plane (rarely
also on both sides of the disk with radial spines).

Genus 255. _Spongolonche_,[294] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with two solid marginal spines, opposite in
one equatorial diameter of the disk.

The genus _Spongolonche_ opens the series of the Spongotrochida, or of
those Spongodiscida in which the margin of the disk is armed with solid
radial spines, situated in the equatorial plane. _Spongolonche_ possesses
only two such spines, opposite in one equatorial diameter of the disk; it
corresponds therefore to _Stylocyclia_ among the Coccodiscida, to
_Xiphodictya_ among the Porodiscida.

{580}1. _Spongolonche conostyla_, n. sp..

Spongy disk circular, with irregular framework, without concentric rings.
Both opposite radial spines conical, about as long as the radius of the
disk, and four times as long as broad at the base. Margin of the disk
nearly smooth.

_Dimensions._--Diameter of the disk 0.16; length of the radial spines 0.18,
basal thickness 0.045.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

2. _Spongolonche amphistyla_, n. sp.

Spongy disk circular, with four to six concentric rings in the inner part,
with quite an irregular framework in the outer part. Both opposite radial
spines cylindrical, twice to three times as long as the diameter of the
disk, at the base about as broad as two meshes of the framework. Margin of
the disk ciliated.

_Dimensions._--Diameter of the disk 0.2; length of the radial spines 0.4 to
0.6, basal thickness 0.01.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 256. _Spongotripus_,[295] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with three solid radial spines on the margin
of the circular or triangular disk.

The genus _Spongotripus_ is characterised by three marginal spines, which
are commonly regularly disposed, more rarely in a bilateral or an irregular
manner. It corresponds to _Trigonocyclia_ among the Coccodiscida, to
_Tripodictya_ among the Porodiscida.

Subgenus 1. _Spongotripodiscus_, Haeckel.

_Definition._--Radial spines of equal size and distance; triangle regular.

1. _Spongotripus regularis_, n. sp.

Spongy disk circular; three radial spines on its margin of equal size and
equidistant, strong, conical, about as long as the diameter of the disk,
and five times as long as broad at the base.

_Dimensions._--Diameter of the disk 0.15; length of the spines 0.16, basal
breadth 0.03.

_Habitat._--Pacific, central area, Station 272, surface.

{581}2. _Spongotripus neumayri_, Haeckel.

_Spongechinus neumayri_, Dunikowski, 1882, Denskchr. d. k. Akad. d. Wiss.
Wien, xlv. p. 28, Taf. v. fig. 59.

Spongy disk circular, nearly spherical; three radial spines on its margin
of equal size and equidistant, pyramidal, nearly as long as the diameter of
the disk, and three times as long as broad at the base.

_Dimensions._--Diameter of the disk 0.15 to 0.2; length of the spines 0.11,
basal breadth 0.04.

_Habitat._--Fossil in the Jura of the Alps and in Tertiary rocks of
Barbados.

3. _Spongotripus strepsiceros_, n. sp.

Spongy disk circular; three radial spines on its margin of equal size and
equidistant, angular, twice as long as the diameter of the disk, spirally
twisted like the horns of Antilope strepsiceros.

_Dimensions._--Diameter of the disk 0.15; length of the spines 0.3, basal
breadth 0.02.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Subgenus 2. _Spongotripodium_, Haeckel.

_Definition._--Radial spines of different size or at different distances;
triangle either isosceles or irregular.

4. _Spongotripus ypsilon_, n. sp.

Spongy disk triangular, isosceles, with convex sides; three spines angular,
of different size and at different distances; the odd spine straight and
twice as long as the two paired spines, which are more approximated and
curved concavely one to the other, like y.

_Dimensions._--Diameter of the disk 0.12; length of the odd spine 0.2, of
the paired spines 0.1, basal breadth 0.04.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

5. _Spongotripus irregularis_, n. sp.

Spongy disk circular; three spines conical, irregularly curved, all three
of different size, and at irregular distances; once to three times as long
as the diameter of the disk; margin of the disk thorny.

_Dimensions._--Diameter of the disk 0.1; length of the spines 0.1 to 0.3,
basal breadth 0.05.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

{582}Genus 257. _Spongostaurus_,[296] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with four solid radial spines on the margin
of the circular or square disk, commonly crossed in two equatorial
diameters perpendicular one to another.

The genus _Staurodictya_ exhibits four marginal spines in cross form.
Commonly the cross is regular and rectangular, the four spines being
opposite in pairs in two perpendicular diameters; sometimes more or less
irregular. The genus repeats _Staurodictya_ among the Porodiscida,
_Staurocyclia_ among the Coccodiscida.

1. _Spongostaurus cruciatus_, n. sp.

Spongy disk circular; four radial spines on its margin opposite in two
crossed equatorial diameters, perpendicular one to another, conical, about
as long as the diameter of the disk.

_Dimensions._--Diameter of the disk 0.16; length of the spines 0.18, basal
breadth 0.04.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

2. _Spongostaurus serratus_, n. sp.

Spongy disk circular; four crossed radial spines very large, twice to three
times as long as the diameter of the central disk, serrated on both edges,
with two rows of strong conical perpendicular teeth (similar to the saw of
_Pristis antiquorum_).

_Dimensions._--Diameter of the disk 0.1; length of the spines 0.2 to 0.3,
breadth 0.02.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

3. _Spongostaurus hastatus_, n. sp.

Spongy disk square, with concave sides; four crossed radial spines
cylindrical, on the broader distal end spear-shaped or nearly lanceolate,
one and a half times as long as the diameter of the disk.

_Dimensions._--Diameter of the disk 0.13; length of the spines 0.2, basal
breadth 0.01, distal breadth 0.03.

_Habitat._--Tropical Atlantic, Station 347, surface.

4. _Spongostaurus quadratus_, n. sp.

Spongy disk square, with rectilinear sides; four crossed radial spines
pyramidal, arising from the corners of the square, about as long as its
half diagonal.

_Dimensions._--Diameter of the disk 0.16; length of the spines 0.08, basal
breadth 0.02.

_Habitat._--Pacific, central area, Station 271, surface.

{583}Genus 258. _Stylotrochus_,[297] Haeckel, 1862, Monogr. d. Radiol., p.
463.

_Definition._--#Spongodiscida# with numerous solid radial spines on the
margin of the disk (five to ten or more), all situated in the equatorial
plane.

The genus _Stylotrochus_ comprises those Spongodiscida in which the margin
of the circular disk bears numerous radial spines. All these spines lie in
the same equatorial plane, whilst in the following genus they are disposed
over the whole surface of the disk. _Stylotrochus_ corresponds to
_Astrocyclia_ among the Coccodiscida, to _Stylodictya_ among the
Porodiscida. The spongy framework of the disk is either quite irregular
(_Stylotrochiscus_), or includes in the middle part some concentric
circular rings (_Stylospongia_).

Subgenus 1. _Stylotrochiscus_, Haeckel.

_Definition._--Spongy framework of the whole disk irregular, without
concentric circular rings or spiral convolutions.

1. _Stylotrochus arachnius_, Haeckel.

_Spongotrochus arachnius_, Haeckel, 1862, Monogr. d. Radiol., p. 464.

Spongy framework of the whole disk irregular. Eight to twelve marginal
spines very long and thin, bristle-shaped, twice to four times as long as
the diameter of the disk. (Very similar to the common _Stylodictya
arachnia_, but without concentric circular rings and sieve-plates, with
quite irregular network of fine bars.)

_Dimensions._--Diameter of the disk 0.12 to 0.15; length of the radial
spines 0.2 to 0.6, basal breadth 0.001.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface; also fossil in Tertiary rocks of Barbados and the Mediterranean.

2. _Stylotrochus craticulatus_, Haeckel.

_Spongotrochus craticulatus_, Stöhr, 1880, Palæontogr., vol. xxvi. p.
118, Taf. vi. fig. 12.

? _Spongodiscus aculeatus_, Ehrenberg, 1854, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 246.

Spongy framework of the whole disk irregular. Sixteen to twenty short
marginal spines (twice to four times as long as the diameter of one mesh of
the framework), free prolongations of internal radial beams which arise
from the darker centre of the disk. (The interruption of the disk-margin on
one point of its circumference, figured by Stöhr as osculum or
"Mündungs-Oeffnung," is probably an accidental abnormality; I did not find
it in other specimens.)

{584}_Dimensions._--Diameter of the disk 0.2 to 0.25; length of the radial
spines 0.005 to 0.018, basal breadth 0.001 to 0.003.

_Habitat._--Pacific, central area, Stations 266 to 268, surface; fossil in
Barbados and Sicily.

3. _Stylotrochus helianthus_, n. sp.

Spongy framework of the whole disk irregular. Thirty to fifty very large,
conical radial spines, about as long as the diameter of the disk, and at
the base two to four times as broad as one mesh of the framework (without
internal prolongations).

_Dimensions._--Diameter of the disk 0.15 to 0.2; length of the radial
spines 0.16 to 0.24, basal breadth 0.008 to 0.016.

_Habitat._--Pacific, central area, Station 272, surface.

4. _Stylotrochus rhabdostylus_, Haeckel.

_Spongosphæra rhabdostyla_, Ehrenberg, 1875, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 256, Taf. xxvi. figs. 1, 2.

Spongy framework of the whole disk irregular. Four very large marginal
primary spines crossed in two equatorial diameters, perpendicular one to
another, and between them numerous (twelve to twenty or more) smaller
secondary spines. The latter arise from the margin, whilst the former
pierce the disk and are nearly united in its centre. All the spines are
cylindrical, the smaller as broad as one mesh, the larger three to six
times as broad.

_Dimensions._--Diameter of the disk 0.2; length of the four main spines
0.15 to 0.3, breadth 0.01 to 0.02; length of the accessory spines 0.05 to
0.1, breadth 0.005.

_Habitat._--Fossil in the rocks of Barbados.

5. _Stylotrochus heteracanthus_, Haeckel.

_Spongotrochus heteracanthus_, Haeckel, 1862, Monogr. d. Radiol., p. 464.

Spongy framework of the whole disk irregular. Ten very long needle-shaped
marginal spines (symmetrically distributed), and between them numerous very
fine, shorter, accessory, bristle-shaped spines.

_Dimensions._--Diameter of the disk 0.16; length of the main spines 0.16,
basal breadth 0.002; length of the secondary spines 0.03.

_Habitat._--Mediterranean (Messina), surface.

Subgenus 2. _Stylospongia_, Haeckel, 1862, Monogr. d. Radiol., p. 473.

_Definition._--Spongy framework of the disk in the inner part with
concentric rings or spiral convolutions, in the outer part quite irregular.

{585}6. _Stylotrochus huxleyi_, Haeckel.

_Stylospongia huxleyi_, Haeckel, 1862, Monogr. d. Radiol., p. 473, Taf.
xxviii. fig. 7.

Spongy framework of the disk in the inner part with five concentric rings,
in the outer part quite irregular. Ten marginal spines, conical at the
base, about as long as the radius of the disk, without inner piercing
prolongations.

_Dimensions._--Diameter of the disk 0.12; length of the radial spines 0.06,
basal breadth 0.003.

_Habitat._--Mediterranean (Messina), Haeckel.

7. _Stylotrochus geddesii_, n. sp. (Pl. 41, fig. 11).

_Stylospongidium geddesii_, Haeckel, 1881, Atlas (pl. xli. fig. 11).

Spongy framework of the disk in the inner part with four to eight
concentric rings (or partially spiral convolutions), in the outer part
quite irregular. Thirty to fifty pyramidal marginal spines of variable
size, one-fourth to one-half as long as the radius of the disk, outer
prolongations of inner piercing radial beams, which arise from various
concentric rings. I call this interesting species, which is intermediate
between _Stylodictya_ and _Stylotrochus_, in honour of the morphologist Mr.
Patrick Geddes of Edinburgh.

_Dimensions._--Diameter of the disk 0.15 to 0.25; length of the radial
spines 0.03 to 0.06, basal breadth 0.004 to 0.01.

_Habitat._--Pacific, central area, Stations 270 to 274, in 2350 to 2925
fathoms.

Genus 259. _Spongotrochus_,[298] Haeckel, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 844.

_Definition._--#Spongodiscida# with numerous solid radial spines (five to
ten or more), which are scattered over the whole surface and the margin of
the disk, or regularly disposed on both sides of it.

The genus _Spongotrochus_ differs from the foregoing and nearly allied
genus by the distribution of the numerous radial spines. These are not
confined to the margin of the disk, but also scattered on its whole
surface, and sometimes symmetrically disposed on both its sides in a
regular manner. Also in this genus the spongy framework is sometimes quite
irregular (_Spongotrochiscus_), at other times in the middle part with
enclosed concentric rings (_Stylospongidium_).

Subgenus 1. _Spongotrochiscus_, Haeckel, 1862, Monogr. d. Radiol., p. 463.

_Definition._--Spongy framework of the whole disk irregular, without
concentric rings or spiral convolutions.

{586}1. _Spongotrochus brevispinus_, Haeckel.

_Spongotrochus brevispinus_, Haeckel, 1862, Monogr. d. Radiol., p. 462,
Taf. xxvii. figs. 4, 5.

Spongy framework of the whole disk irregular, everywhere equal. The whole
surface of the disk covered with numerous short, needle-shaped, radial
spines, about as long as the thickness of the disk, which is one-fifth of
its diameter.

_Dimensions._--Diameter of the disk 0.18; length of the radial spines 0.03
to 0.04, basal breadth 0.002.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

2. _Spongotrochus multispinus_, n. sp.

Spongy framework of the whole disk irregular, in the centre darker. The
whole surface of the disk covered with numerous conical radial spines,
about as long as the radius of the disk.

_Dimensions._--Diameter of the disk 0.22; length of the radial spines 0.1,
basal breadth 0.008.

_Habitat._--Pacific, central area, Stations 271 to 274, surface.

3. _Spongotrochus longispinus_, Haeckel.

_Spongotrochus longispinus_, Haeckel, 1862, Monogr. d. Radiol., p. 463,
Taf xxvii. figs. 2, 3.

Spongy framework of the whole disk irregular, everywhere equal. Surface
thorny. Twenty long, needle-shaped, radial spines, about as long as the
diameter of the disk, symmetrically distributed in pairs on both flat sides
of the disk, so that the opposite pairs lie in five equidistant, meridian
planes (compare the figures).

_Dimensions._--Diameter of the disk 0.2; length of the twenty radial spines
0.2, basal breadth 0.001.

_Habitat._--Mediterranean (Messina), surface.

Subgenus 2. _Stylospongidium_, Haeckel, 1881, Prodromus, p. 460.

_Definition._--Spongy framework of the disk in the inner part with
concentric rings or spiral convolutions, in the outer part quite irregular.

4. _Spongotrochus scutella_, n. sp.

_Spongotrochus ehrenbergii_, Bütschli, 1882, L. N. 41, Taf. xxvi. figs.
1_a_, 1_b_.

Spongy framework of the disk in the inner part with four to six concentric
rings, in the outer part quite irregular. The whole surface of the disk
covered with bristle-shaped radial spines, about half as long as the radius
of the disk.

_Dimensions._--Diameter of the disk 0.24; length of the radial spines 0.06,
basal breadth 0.02.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

{587}5. _Spongotrochus parma_, n. sp.

Spongy framework of the disk in the inner part with five to eight spiral
convolutions (or partly concentric rings), in the outer part quite
irregular. Thirty to forty long, bristle-shaped, radial spines, about as
long as the diameter of the disk, disposed on both flat sides of the disk,
but not on the margin.

_Dimensions._--Diameter of the disk 0.16; length of the radial spines 0.18,
basal breadth 0.003.

_Habitat._--South Atlantic, Station 325, surface.

Subfamily 3. SPONGOBRACHIDA, Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with two or more (commonly three or four)
spongy radial arms on the margin of the disk, situated in its equatorial
plane (with or without a connecting patagium between the arms).

Genus 260. _Spongolena_,[299] n. gen.

_Definition._--#Spongodiscida# with two opposite spongy arms on the margin
of the disk, without a connecting patagium.

The genus _Spongolena_ opens the series of the Spongobrachida, or of the
Spongodiscida provided with radial spongy arms on the margin of the disk.
_Spongolena_ is the most simple form of this subfamily, and bears only two
simple opposite arms, without a connecting patagium. It corresponds to
_Amphibrachium_ (Porodiscida) and to _Diplactura_ (Coccodiscida). Also
there is no patagium. _Spongolena_ may easily be confounded with
_Spongurus_ (compare my Prodromus, 1881, p. 461); but in the true
_Spongurus_ (an ellipsoid) the transverse section is circular, in
_Spongolena_ elliptical.

1. _Spongolena rhopalura_, n. sp.

Arms club-shaped, three times as long as broad, at the rounded distal end
twice as broad as at the base, and twice as long as the diameter of the
circular central disk. Surface nearly smooth.

_Dimensions._--Radius of the arms (or the distance from the centre to the
distal arm-end) 0.16, distal breadth 0.05, basal breadth 0.03.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

2. _Spongolena spongura_, n. sp.

Arms nearly cylindrical, twice as long as broad, and a little longer than
the diameter of the elliptical central disk. Surface thorny, some longer
bristle-shaped spines on the distal end of the {588}arms. (Very similar to
the ellipsoid _Spongurus cylindricus_, Monogr. d. Radiol., p. 465, Taf.
xxvii. fig. 1, but differs in the compressed lenticular (not ellipsoidal)
form of the central disk; the transverse section of the arm is elliptical,
not circular.)

_Dimensions._--Radius of the arms 0.1, breadth 0.04.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

3. _Spongolena cypselura_, n. sp.

Arms nearly triangular, not longer than broad, about half as large as the
elliptical central disk, at the broader distal end with two very large,
widely divergent lateral spines, and between them several smaller, like the
tail of a swallow. Surface thorny.

_Dimensions._--Radius of the arms 0.2, distal breadth (without spines)
0.15, basal breadth 0.07.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Genus 261. _Spongobrachium_,[300] Haeckel, 1881, Prodromus, p. 461.

_Definition._--#Spongodiscida# with two opposite spongy arms on the margin
of the disk, connected by a spongy patagium of different texture.

The genus _Spongobrachium_ differs from the foregoing only in the loose
spongy patagium, which envelops both opposite spongy arms. It corresponds
to _Amphymenium_ among the Porodiscida and to _Amphiactura_ among the
Coccodiscida.

1. _Spongobrachium ellipticum_, Haeckel.

_Spongocyclia elliptica_, Haeckel, 1862, Monogr. d. Radiol., p. 470, Taf.
xxviii. fig. 2.

_Spongodiscus ellipticus_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 844.

Arms nearly square, scarcely as long and broad as the radius of the
circular central disk, at the broader distal end truncated. Patagium
complete, enveloping the whole disk with the arms, and forming a larger
elliptical disk of looser framework. (In my Monograph, 1862, _loc. cit._, I
had not distinguished the opposite darker arms, opposite in the longer axis
of the elliptical disk, from the enveloping looser framework of the
patagium. In larger specimens of the Challenger collection this distinction
is very evident.)

_Dimensions._--Radius of the arms 0.12, breadth 0.05; major axis of the
elliptical patagium 0.24, minor 0.16.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

2. _Spongobrachium lanceolatum_, n. sp.

Arms club-shaped, twice as long as broad, at the distal end pointed, five
times as long as the radius of the circular central disk. Patagium
complete, enveloping the whole disk with the arms, {589}and forming a
larger lanceolate disk of looser framework. (Similar in form to
_Amphymenium pupula_, Pl. 44, fig. 8, but with an irregular spongy
framework and pointed ends.)

_Dimensions._--Radius of the arms 0.16, breadth 0.05; major axis of the
lanceolate patagium 0.32, minor axis 0.12.

_Habitat._--Pacific, central area, Station 270, surface.

Genus 262. _Rhopalodictyum_,[301] Ehrenberg, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 830.

_Definition._--#Spongodiscida# with three spongy arms on the margin of the
circular or triangular disk, without a connecting patagium.

The genus _Rhopalodictyum_ comprises those very common forms of
Spongodiscida in which the margin of the disk is provided with three
simple, free, spongy arms. It corresponds to _Dictyastrum_ and
_Rhopalastrum_ among the Porodiscida, to _Trigonactura_ among the
Coccodiscida. The typical species of this genus is _Rhopalodictyum
abyssorum_ of Ehrenberg, the only species figured by him. His diagnosis of
the genus was very insufficient, and agreed with that of his _Dictyastrum_
(compare above the improved diagnosis of this genus, p. 526, and my
Monograph, p. 466).

Subgenus 1. _Rhopalodictya_, Haeckel.

_Definition._--Triangular shell regular, with three arms of equal size and
equidistant.

1. _Rhopalodictyum abyssorum_, Ehrenberg.

_Rhopalodictyum abyssorum_, Ehrenberg, 1872, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 299, Taf. viii. fig. 17.

Arms of equal size and equidistant, club-shaped, about as long as the
diameter of the triangular central disk, and a little longer than the
breadth of the pear-shaped rounded distal end. Surface rough.

_Dimensions._--Radius of the arms 0.11, basal breadth 0.03, distal breadth
0.06.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific; tropical zone,
surface.

2. _Rhopalodictyum truncatum_, Ehrenberg.

_Rhopalodictyum truncatum_, Ehrenberg, 1861, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 301.

? _Dictyastrum angulatum_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 289, Taf. viii. fig. 18.

Arms of equal size and equidistant, nearly square, with straight edges,
towards the truncated end a little broader, about of the same diameter as
the central triangular disk. This species is indicated {590}by Ehrenberg
only by the short diagnosis, "Radiis stellæ tribus apice truncatis." It is
probably identical with his figure of _Dictyastrum angulatum_ (_loc.
cit._). This latter name I have retained for the similar Porodiscid (above,
p. 526), mainly because the genera _Dictyastrum_ and _Rhopalodictyum_,
according to the insufficient diagnosis of Ehrenberg, seem to be identical.
(Compare my Monograph, 1862, p. 466.)

_Dimensions._--Radius of each arm 0.13, basal breadth 0.09, distal breadth
0.1.

_Habitat._--Tropical Pacific, Philippine Sea, Station 200, depth 250
fathoms; Atlantic (Mexican Gulf Stream).

3. _Rhopalodictyum subacutum_, Ehrenberg.

_Rhopalodictyum subacutum_, Ehrenberg, 1861, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 301.

Arms of equal size and equidistant, club-shaped, three times as long as the
diameter of the central disk, which equals the breadth of the thickened
distal end; the latter is armed with a strong pyramidal terminal spine.
(The diagnosis of Ehrenberg is "Radiis stellæ tribus, apice cuneatis
subacutis.")

_Dimensions._--Radius of the arms 0.2, basal breadth 0.04, distal breadth
0.07.

_Habitat._--North Atlantic, surface and various depths; Greenland; Mexican
Gulf Stream; Færöe Channel (John Murray).

4. _Rhopalodictyum bifidum_, n. sp.

Arms of equal size and equidistant, in the distal half forked, twice as
long as the diameter of the central disk; both fork branches half as broad
as the simple basal part, truncated at the distal end.

_Dimensions._--Radius of the arms 0.16, basal breadth 0.08, distal breadth
0.04.

_Habitat._--North Pacific, Station 237, surface.

Subgenus 2. _Triactinosphæra_, Dunikowski, 1882, Denkschr. d. k. Akad. d.
Wiss. Wien, vol. 45, ii. p. 192.

_Definition._--Triangular shell bilateral or irregular, with three arms of
different size or distance.

5. _Rhopalodictyum zittelii_, Haeckel.

_Triactinosphæra zittelii_, Dunikowski, 1882, Denkschr. d. k. Akad. d.
Wiss, Wien, vol. 45, ii. p. 192.

Arms of different size and at unequal distances, one odd arm being a little
shorter than the two paired arms; the odd angle between the latter is
larger than the paired angles between them and the {591}former. Arms
club-shaped, their basal semi-cylindrical, half as broad as the spherical
distal half, which bears a strong, conical, terminal spine. The discoverer
of this remarkable, very old, Liassic species, Dunikowski, supposes that it
is not a true Discoid, from the absence of a central disk; in my opinion
the central disk (nearly spherical) has the same relation to the arms as in
many other #Discoidea#, their equatorial plane is the same.

_Dimensions._--Radius of the odd arm 0.18, of the paired arms 0.24; basal
breadth 0.08, distal breadth 0.16.

_Habitat._--Fossil in the Alpine Lias, Schafberg, near Salzburg,
Dunikowski.

6. _Rhopalodictyum elongatum_, n. sp.

Arms of different size and at unequal distances, one odd arm being twice as
long as both paired arms; the odd angle between the latter much larger than
both equal paired angles. Arms nearly cylindrical, little flattened, three
to six times as long as broad, with rounded blunt distal end.

_Dimensions._--Radius of the arms 0.15 to 0.3, breadth 0.05.

_Habitat._--Pacific, central area, Station 274, surface.

7. _Rhopalodictyum curvatum_, n. sp.

Arms of different size and at unequal distances, irregular, more or less
curved, nearly cylindrical, five to ten times as long as broad, with
rounded blunt distal end. (There were observed only two specimens of this
remarkable irregular species; in one specimen all three arms were simple,
in the other one odd arm forked. The length of the arms and the size of the
angles between them seems to be very different and variable.)

_Dimensions._--Radius of the arms 0.3 to 0.6 to 1.15, breadth 0.06 to 0.12.

_Habitat._--Pacific, central area, Station 271, surface.

Genus 263. _Dictyocoryne_,[302] Ehrenberg, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 830.

_Definition._--#Spongodiscida# with three spongy arms on the margin of the
circular or triangular disk, connected by a spongy patagium of different
texture.

The genus _Dictyocoryne_ differs from the foregoing _Rhopalodictyum_ only
in the patagium, connecting the three spongy arms, and bears to it the same
relation as in the Porodiscida _Euchitonia_ does to _Rhopalastrum_. The
typical species of this genus is _Dictyocoryne profunda_, the only species
figured by Ehrenberg. After his {592}insufficient diagnosis _Dictyocoryne_
was identical with his _Spongaster_ (compare my Monograph, p. 467); but as
the type of this latter genus (_Spongaster tetras_) figured by him, 1872,
had four crossed arms, we retain this genus here separate.

Subgenus 1. _Dictyocorynula_, Haeckel.

_Definition._--Triangular shell regular, with three arms of equal size and
equal distance.

1. _Dictyocoryne profunda_, Ehrenberg.

_Dictyocoryne profunda_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 307, Taf. vii. fig. 23.

Arms of equal size and equidistant, club-shaped, in the outer circular half
three times as broad as in the inner cylindrical half, and much larger than
the small triangular central disk. Patagium complete, forming an
equilateral triangle with rounded corners.

_Dimensions._--Radius of the arms 0.14, basal breadth 0.025, distal breadth
0.07.

_Habitat._--Pacific, Philippine Sea, depth 3300 fathoms, Ehrenberg; Station
198, depth 2150 fathoms; Station 274, depth 2750 fathoms.

2. _Dictyocoryne tetradiscus_, n. sp.

Arms of equal size and equidistant, club-shaped, their outer circular half
is quite as large as the circular central disk, and is connected with it by
the thin cylindrical inner half, which is scarcely one-fourth as broad.
Patagium complete, forming an equilateral triangle with rounded corners and
convex sides.

_Dimensions._--Radius of the arms 0.16, basal breadth 0.02, distal breadth
0.08.

_Habitat._--Pacific, central area, Stations 271 to 274, in 2350 to 2750
fathoms.

3. _Dictyocoryne euchitonia_, Haeckel.

_Dictyocoryne euchitonia_, Haeckel, 1862, Monogr. d. Radiol., p. 468.

Arms of equal size and equidistant, lanceolate, twice as long as broad in
the distal part, and three times as long as the small circular central
disk. Patagium complete, forming an equilateral triangle with pointed
corners. (Very similar to _Euchitonia köllikeri_, Monograph, 1862, p. 511,
Taf. xxxi. fig. 6, but quite spongy, not concentric.)

_Dimensions._--Radius of the arms 0.15, basal breadth 0.03, distal breadth
0.06.

_Habitat._--Mediterranean (Messina), Haeckel.

{593}4. _Dictyocoryne trigona_, n. sp.

Arms of equal size and equidistant, lanceolate, three times as long as
broad in the middle part, and as the diameter of the triangular central
disk; their distal end armed with a strong conical radial spine. Patagium
complete, forming an equilateral triangle with pointed corners.

_Dimensions._--Radius of the arms 0.2, basal breadth 0.02, distal breadth
0.06.

_Habitat._--North Atlantic, Canary Islands, surface.

Subgenus 2. _Dictyocorynium_, Haeckel.

_Definition._--Triangular shell bilateral or irregular, with three arms of
different sizes or at different distances.

5. _Dictyocoryne charybdaea_, Haeckel.

_Spongocyclia charybdaea_, Haeckel, 1862, Monogr. d. Radiol., p. 472,
Taf. xxviii. figs. 5, 6.

_Spongodiscus charybdaeus_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 844.

Arms at different distances, nearly equilateral triangular, scarcely half
as long as the radius of the large circular central disk. Both paired arms
touching at their bases, separated by a great distance from the opposite
odd arm. Patagium complete, nearly pentagonal. (The illustration in my
Monograph, in the coloured plate xxviii., is better than my description, as
I had not exactly separated the arms from the patagium.)

_Dimensions._--Radius of the arms 0.18, basal breadth 0.08.

_Habitat._--Mediterranean (Messina), surface.

6. _Dictyocoryne pentagona_, Stöhr.

_Dictyocoryne pentagona_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 118,
Taf. vii. fig. 2.

Arms at different distances, club-shaped, little longer than broad at their
rounded distal end. Both paired arms smaller and more approximate than the
opposite odd arm. Patagium complete, very large, enveloping the whole
shell, and forming a pentagon with rectilinear base and transverse
constriction, and with five rounded corners.

_Dimensions._--Radius of the arms 0.1 to 0.15, middle breadth 0.04.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Stöhr.

7. _Dictyocoryne agrigentina_, Stöhr.

_Dictyocoryne agrigentina_, Stöhr, 1880, Palæontogr., vol. xxvi. p. 118,
Taf. vii. fig. 1.

Arms at different distances, club-shaped, about twice as long as broad, and
smaller than the large triangular central disk. Both paired arms smaller
and more approximate than the opposite odd {594}arm. Patagium incomplete,
enveloping only the basal half of the arms as a circular spongy disk of
loose framework.

_Dimensions._--Radius of the arms 0.18, basal breadth 0.05, distal breadth
0.08.

_Habitat._--Fossil in the Tertiary rocks of Sicily, Grotte, Stöhr.

8. _Dictyocoryne echinata_, n. sp.

Arms unequal and at different distances, club-shaped, in the distal half
twice as broad as in the basal half, three times as long as the small
triangular central disk, at their distal end armed with one larger and
several smaller conical spines. Both paired arms with touching bases, and
only half as large as the doubly remote odd arm. Patagium incomplete,
enveloping only the basal half of the arms, and forming a circular disk of
looser framework. Surface thorny.

_Dimensions._--Radius of the arms 0.15 to 0.2, basal breadth 0.03, distal
breadth 0.06.

_Habitat._--South Atlantic, Station 325, surface.

Genus 264. _Spongasteriscus_,[303] Haeckel, 1862, Monogr. d. Radiol., p.
474 (_sensu restricto_).

_Definition._--SPONGODISCIDA with four spongy arms on the margin of the
circular or quadrangular disk, crossed in two equatorial diameters, without
a connecting patagium.

The genus _Spongasteriscus_ (in the restricted definition here stated)
exhibits on the margin of the disk four spongy arms, which form either a
regular or a bilateral cross. It corresponds to _Stauralastrum_ and
_Myelastrum_ among the Porodiscida, to _Astractura_ among the Coccodiscida.

Subgenus 1. _Spongasteriscinus_, Haeckel.

_Definition._--Cross formed by the four arms regular, rectangular, with
equal-sized and equidistant arms.

1. _Spongasteriscus ovatus_, n. sp.

Arms at equal distances, forming a regular, rectangular cross, egg-shaped,
with a broader rounded distal end, one and a third times as long as broad,
and three times as long as the radius of the central disk; in the latter
three to four concentric rings.

_Dimensions._--Radius of the arms 0.1, greatest breadth 0.06.

_Habitat._--Western Tropical Pacific, Station 225.

{595}2. _Spongasteriscus clavatus_, n. sp.

Arms at equal distances, forming a regular, rectangular cross, club-shaped,
about as long as the diameter of the central disk, and at their rounded
distal end one and a third times as long as broad, at their narrow base
only one third as broad. In the centre five to six concentric rings.

_Dimensions._--Radius of the arms 0.13, distal breadth 0.06, basal breadth
0.02.

_Habitat._--Pacific, central area. Station 271, surface.

3. _Spongasteriscus mucronatus_, n. sp.

Arms at equal distances, forming a regular, rectangular cross, club-shaped,
three times as long as the radius of the central disk, in the distal half
nearly circular, three to four times as broad as at the narrow base. The
rounded distal end armed with a strong pyramidal spine. In the centre no
concentric rings. (Similar to _Stauralastrum rhopalophorum_, Pl. 45, fig.
1, but quite spongy.)

_Dimensions._--Radius of the arms 0.12, basal breadth 0.03, distal breadth
0.02.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

4. _Spongasteriscus furcatus_, n. sp.

Arms at equal distances, forming a regular, rectangular cross, in the
distal third forked; both fork branches half as long as the basal undivided
part of the arm, which is twice as long as broad. Distal ends of the eight
branches blunt, rounded. In the central disk no concentric rings.

_Dimensions._--Radius of the arms 0.12, basal breadth 0.03; distal breadth
of the branches 0.02.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

5. _Spongasteriscus armatus_, n. sp.

Arms at equal distances, forming a rectangular, regular cross, in the
distal half forked; both fork branches of the same length as the basal
undivided part of the arm, which is nearly square. Distal ends of the eight
branches armed with a strong pyramidal spine. In the central disk no
concentric rings. (Similar to _Dicranastrum cornutum_, Pl. 45, fig. 2, but
quite spongy.)

_Dimensions._--Radius of the arms 0.18, basal breadth 0.03, distal breadth
0.015.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Subgenus 2. _Spongasterisculus_, Haeckel.

_Definition._--Cross formed by the four arms, bilateral or irregular, with
the arms at different distances.

{596}6. _Spongasteriscus quadricornis_, Haeckel.

_Spongasteriscus quadricornis_, Haeckel, 1862, Monogr. d. Radiol., p.
474, Taf. xxviii. figs. 8-10.

_Spongodiscus quadricornis_, Haeckel, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 844.

Arms at different distances, forming a bilateral or irregular cross,
grouped in two opposite pairs; their form equilateral triangular; their
length smaller than the radius of the large circular central disk, which
exhibits in the interior eight to sixteen concentric rings.

_Dimensions._--Radius of the arms 0.2, of the central disk 0.13; basal
breadth of the arms 0.08.

_Habitat._--Mediterranean (Messina), Haeckel, surface.

7. _Spongasteriscus tetraceros_, Haeckel.

_Spongasteriscus tetraceros_, Haeckel, 1862, Monogr. d. Radiol., p. 475.

Arms at different distances, forming a bilateral or irregular cross,
grouped in two opposite pairs; their form isosceles triangular; their
length larger than the radius of the large elliptical central disk, which
exhibits in the interior six to twelve concentric rings.

_Dimensions._--Radius of the arms 0.16, of the central disk 0.1; basal
breadth of the arms 0.06.

_Habitat._--Mediterranean (Messina), North Atlantic (Canary Islands),
surface.

8. _Spongasteriscus myelastrum_, n. sp.

Arms at different distances, forming a bilateral or irregular cross,
grouped in two opposite pairs; the arms of one pair broader and shorter
than the arms of the other pair. Each arm in its basal half simple, in the
distal half forked; ends of the fork branches blunt. In the central disk no
concentric rings. (Similar to _Myelastrum octocorne_, Pl. 47, fig. 12, but
quite spongy.)

_Dimensions._--Radius of the arms 0.2, basal breadth 0.05, distal breadth
0.02.

_Habitat._--North Pacific, Station 236, surface.

Genus 265. _Spongaster_,[304] Ehrenberg, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 833.

_Definition._--#Spongodiscida# with four spongy arms on the margin of the
circular or quadrangular disk, connected by a spongy patagium of different
texture.

The genus _Spongaster_ differs from the foregoing _Spongasteriscus_ in the
patagium connecting the spongy arms, and bears therefore to it the same
relation as, in the Porodiscida, _Histiastrum_ does to _Stauralastrum_, or,
in the Coccodiscida, _Stauractura_ does to _Astractura_. The typical
specimen, figured by Ehrenberg (_Spongaster tetras_), exhibits a regular,
square disk, as also some other species. In a certain number of other
species (formerly united by me with _Spongocyclia_) the quadrangular disk
is bilateral.

{597}Subgenus 1. _Spongastrella_, Haeckel.

_Definition._--Cross formed by the four arms regular, rectangular, with the
arms of equal size and equidistant.

1. _Spongaster tetras_, Ehrenberg.

_Spongaster tetras_, Ehrenberg, 1872, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 299, Taf. vi. (iii.) fig. 8.

_Dictyocoryne tetras_, Haeckel, 1862, Monogr. d. Radiol., p. 469.

Arms at equal distances, forming a rectangular, regular Myelastrum,
papiliocross, club-shaped, about twice as long as the diameter of the
square central disk and eight times as long as broad at their base. Distal
ends rounded, perfectly enclosed by the complete patagium, which forms a
regular square, with slightly concave sides.

_Dimensions._--Radius of the arms 0.12, distal breadth 0.03, basal breadth
0.015; length of the square side 0.2.

_Habitat._--Cosmopolitan; Atlantic, Indian, Pacific, surface and in various
depths.

2. _Spongaster quadratus_, n. sp.

Arms at equal distances, forming a regular, rectangular cross, club-shaped,
about four times as long as the diameter of the central circular disk, and
five times as long as broad at the base. Basal third of the arms square;
distal two thirds triangular, three times as broad, with a truncated distal
end. Patagium complete, perfectly enveloping the arms, and bordered by an
elegant, radially striated, broad edge, forming a regular square. (Similar
to _Histiastrum quadratum_, Pl. 46, fig. 4, but quite spongy.)

_Dimensions._--Radius of the arms 0.15, distal breadth 0.06, basal breadth
0.02; length of the square side 0.25.

_Habitat._--Pacific, central area, Stations 270 to 274, depths 2350 to 2925
fathoms.

3. _Spongaster cruciatus_, n. sp.

Arms at equal distances, forming a rectangular, regular cross, lanceolate,
three times as long as broad and as the diameter of the central circular
disk. Ends of the arms provided with a short conical spine. Patagium
incomplete, enveloping only the basal half of the arms, forming a regular
square with concave sides.

_Dimensions._--Radius of the arms 0.18, greatest breadth 0.05; length of
the square side 0.2.

_Habitat._--South Pacific, Station 288, surface.

{598}4. _Spongaster pentacyclus_, n. sp.

Arms at equal distances, forming a rectangular, regular cross, of the same
size and form as the circular central disk, so that the dark interior part
of the shell is composed of five equal circular disks, situated in a
quincuncial manner. The clearer complete patagium, enveloping the whole
cross perfectly, forms a regular square with rounded edges.

_Dimensions._--Radius of the arms 0.2; diameter of each of the five
circular disks 0.01; length of the square side 0.35.

_Habitat._--West Indies, Cuba, surface (Thomson).

Subgenus 2. _Spongastromma_, Haeckel.

_Definition._--Cross formed by the four arms bilateral or irregular, with
the arms at different distances.

5. _Spongaster orthogonus_, Haeckel.

_Spongocyclia orthogona_, Haeckel, 1862, Monogr. d. Radiol., p. 471, Taf.
xxviii. fig. 3.

_Spongodiscus orthogonus_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 844.

Arms at different distances, forming a bilateral cross, grouped in two
opposite pairs of equal size and similar form. Arms club-shaped, about as
long as the diameter of the central circular disk, enveloped perfectly by
the complete patagium, which forms a regular rectangle, the longer side of
which is one and a half times as long as the shorter side. (The arms in my
figure are not distinctly enough marked.)

_Dimensions._--Radius of the arms 0.08, breadth 0.02; length of the larger
side of the rectangle 0.15, of the smaller 0.1.

_Habitat._--Mediterranean (Messina), surface.

6. _Spongaster scyllaeus_, Haeckel.

_Spongocyclia scyllaea_, Haeckel, 1862, Monogr. d. Radiol., p. 471, Taf.
xxviii. fig. 4.

_Spongodiscus scyllaeus_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 844.

Arms at different distances, forming a bilateral cross, grouped in two
opposite pairs of different size and form, one pair smaller and less
divergent than the other. Arms club-shaped, little longer than the radius
of the central circular disk, enveloped perfectly by the complete patagium,
which forms a trapezium; the convergent longer sides of the latter are one
and a half times as long as the larger, and twice as long as the smaller
parallel side. (The arms are in my figure, _loc. cit._, not distinctly
enough marked.)

_Dimensions._--Radius of the arms 0.12 to 0.14, breadth 0.02; length of
both convergent sides 0.24, of the larger parallel side 0.18, of the
smaller 0.12.

_Habitat._--Pacific, central area, Station 272, surface; Mediterranean
(Messina).

{599}----

Suborder VI. LARCOIDEA, Haeckel, 1883 (Pls. 9, 10, 49, 50).

_Definition_.--SPUMELLARIA with lentelliptical central capsule (rarely
somewhat modified or allomorphic), with a lentelliptical fenestrated
siliceous shell (often modified or allomorphic, and sometimes quite
irregular). Growth different in the three unequal dimensive axes,
perpendicular one to another. The typical Lentellipsis is characterised by
three elliptical dimensive planes of different sizes, perpendicular one to
another.

The section #Larcoidea#, the fourth and last of the #Sphærellaria#,
comprises all those forms of this group in which the fenestrated shell
originally is lentelliptical, characterised by different growth in three
different axes, perpendicular one to another, all three equal on both
poles. The geometrical fundamental form of the shell is therefore a
lentellipsis or a triaxial ellipsoid; and this typical form is preserved
completely in the majority of #Larcoidea# in the pure geometrical form of
the central capsule.

The three dimensive axes, which determine the typical form of #Larcoidea#,
are commonly differentiated in such a way, that the first, the longitudinal
or principal axis, is the longest; both its poles, oral and aboral (or
anterior and posterior) are equal. The second, the lateral or transverse
axis, is commonly less than the first, greater than the third axis; both
its poles are the equal lateral poles (right and left not differentiated).
The third dimensive axis, the equatorial or sagittal axis, is commonly the
shortest; both its equal poles are the sagittal poles (dorsal and ventral
poles not different). The relative size of the three dimensive axes in the
human body exhibits similar relations.

The three dimensive planes of the #Larcoidea#, the sagittal, lateral, and
transverse planes, are elliptical, all three of different sizes. The first
plane, the median or sagittal plane, is commonly as regards size between
the two others; its major axis is the principal, its minor the sagittal
axis; it separates the right half of the body from the left. The second
plane or lateral plane, is commonly larger than the two others; its major
axis the principal, its minor the transverse axis; it separates the dorsal
half of the body from the ventral. The third plane, the equatorial or zonal
plane, is commonly less than the two others; its major axis the lateral,
its minor the sagittal axis; it separates the two principal halves of the
body, the oral and aboral halves.

In my Monograph (1862) only very few forms of #Larcoidea# are described,
_Tetrapyle_ and _Lithelius_ (the latter representing a peculiar family,
Lithelida). In my Prodromus (1881, pp. 463, 464) I disposed all observed
forms of #Larcoidea# in two different families, the Pylonida and Lithelida.
The rich materials of the Challenger collection have since offered an
astonishing number of new and interesting forms of this section, so that I
can enumerate here fifty-one genera and two hundred and sixty-five species.
I dispose them here in four subsections and nine families. Three of these
have regular lentelliptical shells, which are not articulate, and
{600}without annular constrictions (Larcarida, Larnacida, Pylonida); these
form the subsection Pylolarcida. Two other families (Tholonida and
Zonarida) are distinguished by annular constrictions, which divide the
regular lentelliptical shell into a number of dome-shaped chambers or
cupolas; we call these Thololarcida. A third group, Spirolarcida, comprises
the #Larcoidea# with spiral growth; the two families of Lithelida and
Streblemida. Finally a fourth group the Sorolarcida is formed by the
#Larcoidea# with irregular shells, also two families, the Phorticida and
Soreumida.

The first family of #Larcoidea#, the Larcarida, contains the most simple
forms, beginning with _Cenolarcus_, a quite simple lentelliptical latticed
shell. In _Coccolarcus_ we find already two concentric shells, connected by
radial beams, an inner medullary and an outer cortical shell. In
_Spongolarcus_ the lentelliptical shell becomes spongy.

The second family, Larnacida, is very similar to the Larcarida, and seems
to diverge only by the different mode of connection between the two
concentric lentelliptical shells. But in truth this slight difference is of
great morphological importance, as it depends on a quite different and
peculiar mode of growth. In the foregoing Larcarida (_Coccolarcus_, &c.),
the concentric shells originate in the same manner as in the concentric
#Prunoidea# and #Sphæroidea#, by radial beams, which arise from the surface
of the inner (medullary) shell and become connected by a network to form
the outer (cortical) shell. Here, in the Larnacida, a quite similar shell
originates in a quite different way, first arrived at in the Pylonida
(_Trizonium_). Both concentric shells become here connected by peculiar
lattice girdles, which are developed in the perimeter of the three
elliptical dimensive planes. Firstly, on both sides of a simple, spherical,
or lentelliptical central chamber, arise two lateral wings (on the poles of
the transverse axis), and build around the former a transverse girdle. This
is crossed by a larger lateral girdle, the minor axis of which is the major
of the former, and perpendicular to both girdles is yet developed a third,
the sagittal girdle. If the open fissures or "gates" between these three
girdles become closed by network, we obtain _Larnacilla_, the probable
ancestral form of all Larnacida.

Whilst in _Larnacilla_ and _Larnacidium_ this typical trizonal
lentelliptical shell constitutes by itself alone the whole skeleton, in the
other Larnacida it becomes overgrown by outer envelops, and so becomes
enclosed in the interior of the central capsule as a "_Larnacilla_-shaped
medullary shell." If the enclosing external envelops be simply latticed, we
get the subfamily Larnacalpida; if they be spongy, we get the
Larnacospongida.

The third family, Pylonida, is the most important of all #Larcoidea#, as
not only the largest and most interesting number of species belong to it,
but also many other genera (far the greater part of all #Larcoidea#) may be
derived from it. The peculiar character of the Pylonida is determined by
the imperfect fenestration of the lentelliptical shell growing in the three
dimensive axes in a quite different manner. Each elliptical dimensive plane
becomes circumscribed by an elliptical latticed girdle (or fenestrated
{601}ring), and between these three girdles (perpendicular one to another)
remain wide open fissures of the shell or "gates" (_Pylæ_). The beginning
of the shell-building is the same as in _Larnacilla_, the most simple form
of Larnacida. From a quite simple medullary shell, a spherical,
subspherical, ellipsoidal, or lentelliptical central chamber, arise two
latticed wings, opposite on the poles of the transverse axis
(_Monozonium_). Both wings are short and wide hollow fenestrated tubes, the
axes of which are parallel to the principal axis. Therefore they form
together with the central chamber an elliptical transverse girdle. This
first girdle becomes crossed by a second lateral girdle; from both poles of
the transverse axis arise latticed wings, which unite on the poles of the
principal axis, therefore the minor axis of this second larger ring is the
major axis of the first smaller ring (_Dizonium_). Between the two crossed
rings remain four wide open gates. Now follows the development of a third
sagittal girdle, arising from both poles of the principal axis and
overgrowing the four gates. But as this third girdle is larger than the
second, four other larger gates arise between the two (in planes
perpendicular to the former four gates). Now we have the characteristic and
most important trizonal shell (_Trizonium_), composed of three elliptical
lattice-girdles of different size, perpendicular one to another, and
enclosing a simple central chamber. If the four gates of this _Trizonium_
become closed by lattice-work, it passes over into _Larnacilla_, the most
important ancestral form of the Larnacida.

This most significant "trizonal shell," either incompletely latticed in
_Trizonium_ (with four open gates), or completely latticed by fenestration
of the four gates, in _Larnacilla_, is to be found in far the greater part
of all #Larcoidea#, representing the medullary shell, which is overgrown by
an outer cortical shell. In many #Larcoidea#, in which this
"_Larnacilla_-shell" is absent, it is perhaps lost by phylogenetic
reduction, or retrograde metamorphosis.

The same process of triple girdle-building, by which the typical
_Trizonium_-shell or _Larnacilla_-shell is produced (Haplozonaria), is
repeated once or twice in the larger forms of Pylonida. The first system of
three girdles (perpendicular one to another) becomes overgrown by a second
system of the same formulation in the Diplozonaria, and this becomes
overgrown by a third system in the Triplozonaria; in the highest genus of
this group, _Pylozonium_, we find not less than nine girdles (three
systems, each of three girdles). Till now only one genus of the whole
polymorphous family was well known, _Tetrapyle_ (with five girdles, three
of the medullary, two of the cortical shell). If the gates between the
girdles remain open, all these forms must be regarded as Pylonida; if the
gates afterwards become closed by a network, they pass over into other
families.

The fourth family of the _Larcoidea_ is the Tholonida, distinguished by the
polythalamous shell being composed of a certain number of roundish or
hemispherical chambers (domes or cupolas), which surround a primordial
central chamber in quite regular disposition, lying opposite in pairs on
the poles of the three dimensive axes. {602}If we imagine that each "wing"
(or open half-girdle) of the Pylonida becomes closed by a lattice-work, and
so transformed into a hemispherical or roundish cupola, we obtain the
characteristic shell of the Tholonida. Indeed every girdle of the former
corresponds to a pair of opposite domes of the latter. The axis of each
pair of domes is one of the three dimensive axes.

The primordial chamber of the Tholonida (or the central chamber, around
which all cupolas are regularly disposed) is either a simple lentelliptical
lattice-shell, like _Cenolarcus_, or it is a trizonal shell (with an
enclosed concentric medullary shell), like _Larnacilla_. As in both cases
the building and the disposition of the cupolas around it are quite the
same, we can suppose that the whole family of Tholonida may have been
derived originally from _Larnacilla_ (or _Trizonium_), and that the
Cenotholida (with a simple central chamber) are sprung from the
Coccotholida (with a _Larnacilla_-shaped central chamber) by reduction and
loss of the original medullary shell.

The family Tholonida can be divided into three subfamilies according to the
disposition of the cupola-pairs in one, two, or three dimensive axes. In
the Cubotholida lie two cupolas on the poles of the transverse axis of the
central chamber (corresponding to _Amphipyle_); in the Staurotholida we
find four cupolas crosswise disposed, on the poles of the transverse and
principal axes (corresponding to _Tetrapyle_); in the Cubotholida are at
least six cupolas, on the poles of all three dimensive axes (corresponding
to _Tholonium_). In all three cases the number of cupolas may be augmented
by the secondary apposition of other chambers or domes in the same
disposition. Sometimes also the whole cortical shell becomes enclosed by an
external veil or mantle of delicate network. The lentelliptical (or often
nearly cubical) central chamber becomes often reduced, so that its sides
are incompletely latticed or widely opened; in some Cubotholida only the
twelve edges of the eight cornered cubical central chamber remain; its six
sides are quite open and only over-vaulted by the six hemispherical
cupolas. From the opposite points of the latter (in the deep annular
constrictions between them) often arise radial spines, and these lie
commonly in diagonal planes, separating the dome-pairs.

A similar dome-building or a composition of the polythalamous shell by
pairs of cupolas we find also in the next (fifth) family, the Zonarida. But
here the true cause of the peculiar dome-structure is quite different, not
an apposition of new chambers, but the constriction of a cortical
shell-like _Larnacalpis_ by two or more constrictions. These constrictions
lie in dimensive planes (or in planes parallel to these), and therefore the
cupolas are (all or partly) in diagonal planes, a condition quite opposite
to that found in the Tholonida. One of the annular constrictions is
constantly in the sagittal plane (separating the right and left halves of
the shell). The number of the constrictions in the few genera is two,
three, and four, and therefore the number of the cupolas four, six, or
eight. As this cortical shell constantly encloses a trizonal medullary
shell (or _Larnacilla_-shell), we cannot doubt that the Zonarida must be
derived from the Larnacida.

{603}Whilst in all the foregoing five families of #Larcoidea# the
shell-form is regular and their geometrical fundamental form is a
lentellipsis (or a triaxial ellipsoid, with three unequal isopolar
dimensive axes), in the four remaining families of this suborder the shell
becomes bilateral or irregular (with the poles of the axes unequal). In two
of these families (Lithelida and Streblemida) the growth of the shell
becomes spiral, in the last two families (Soreumida and Phorticida) quite
irregular. But as in all four families we encounter the typical trizonal
medullary shell (or _Larnacilla_-shell), we are convinced that they must be
derived (wholly or partially) from the Larnacida.

The Lithelida (the sixth family) are #Larcoidea# with spiral growth and
bilateral form (like _Nautilus_); therefore the spiral line lies in one
plane and this spiral plane divides the whole shell into two symmetrical
halves (right and left). The axis of the spiral (around which the shell
winds) is a straight line, one of the three dimensive axes. In the greater
part of Lithelida (in the Larcospirida) the primordial of central chamber
of the polythalamous shell is a trizonal medullary shell or
_Larnacilla_-shell, and the growth of the first spiral turning begins as
the development of the first (transverse) cortical girdle of _Amphipyle_;
but as one wing (or lateral half) of this girdle grows more rapidly than
the other, it overgrows the latter and begins the spiral winding; if the
other wing follow and overgrow the first, the spiral becomes double. Each
of the three dimensive girdles (of the Pylonida) may begin the spiral
winding. There can be no doubt that all these Lithelida (the Larcospirida)
must be derived from the Pylonida, by unequal growth of the two halves of
one girdle. Perhaps from these may also derived the other part of this
family, the Spiremida (_Spirema_ and _Lithelius_); in these the primordial
chamber of the spiral shell is simple, and may be derived by reduction of
the original _Larnacilla_-shell. But it is also possible that the Spiremida
proceed directly from the Larcarida, and that their ancestors did not
possess a _Larnacilla_-shell.

The Streblemida (the seventh family) are #Larcoidea# with spiral growth and
asymmetrical form of the polythalamous shell (like _Helix_ or
_Turrilites_); therefore the spiral line is twisted like a winding stair,
and the spiral face is curved and divides the shell into two unequal
halves. The Streblemida have the same likeness and relation to the
turbinoid Foraminifera (_Rotalia_, _Globigerina_, &c.) as the Lithelida to
the nautiloid Foraminifera (_Polystomella_, _Nummulina_, &c.). As in these
calcareous Rhizopods also the peculiar growth of the siliceous Streblemida
begins from a primordial chamber to which a variable number of roundish
chambers (of increasing size) is apposed. But the building of these
chambers and of their septa is by no means so regular and complete as in
the greater number of turbinoid Foraminifera. As in a part of this family
the primordial chamber is _Larnacilla_-shell, these also may be derived
from the Larnacida, but the other part (with simple central chamber) is
perhaps produced directly from the Larcarida.

The eighth family, Soreumida, is perhaps derived from the Streblemida by
the loss of the spiral growth. The polythalamous shell is similar to the
latter, but the chambers are {604}aggregated without any order, like the
Acervulinida among the Foraminifera. In some cases also here the primordial
chamber is a trizonal _Larnacilla_-shell, in other cases it is a simple,
subspherical or lentelliptical shell.

The last family, the Phorticida, is formed of irregular #Larcoidea#, in
which a lentelliptical trizonal _Larnacilla_-shell (as an inner medullary
shell) is enveloped by an irregular, latticed, or spongy cortical shell.
They can be regarded as abnormalities or irregular deformities of Larnacida
or Pylonida.

The central capsule of the #Larcoidea# is originally lentelliptical and
preserves this form, the "triaxial ellipsoid," in the greater number of
genera. In some groups it follows the prevalent growth of the shell in the
direction of one of the three dimensive axes, and becomes prolonged in this
way. In many chambered forms (particularly Tholonida and Zonarida) the
growing central capsule gets constricted, corresponding to the
constrictions of the shell. In the Soreumida and Phorticida its form often
becomes irregular. But in general for the greater number of #Larcoidea# the
lentelliptical form of their central capsule is quite characteristic.

_Synopsis of the Families of_ #Larcoidea#.

-------------------------------------------------------------------------
#Larcoidea# with a regular or symmetrical shell, the growth of which is
determined by the three dimensive axes. (Both poles of each axis are
equal.)
-------------------------------------------------------------------------
{Medullary shell absent or
Cortical shell completely { simple (spherical or
latticed, without external { lentelliptical), 1. LARCARIDA.
gates (or interzonal {
fissures), without annular {Medullary shell trizonal or
constrictions and domes. { _Larnacilla_-shaped
{ (composed of three
{ dimensive girdles), 2. LARNACIDA.

Cortical shell incompletely latticed, with two to four or
more symmetrically disposed gates or fissures remaining
between latticed dimensive girdles, 3. PYLONIDA.

Cortical shell completely }Constrictions of the
latticed, without external } cortical shell in
gates (or interzonal } diagonal planes; domes
fissures), with two or } in dimensive axes, 4. THOLONIDA.
more annular constrictions,}
which separate three to }Constrictions of the
six or more dome-shaped } cortical shell in
protuberances. } dimensive planes; domes
} in diagonal axes, 5. ZONARIDA.
-------------------------------------------------------------------------
#Larcoidea# with a symmetrical or irregular shell, either with
spiral growth or with quite irregular growth. (Both poles of one axis
are different.)
-------------------------------------------------------------------------
{Spiral cortical shell bilateral
{ (with plane spiral), 6. LITHELIDA.
Cortical shell with {
spiral growth. {Spiral cortical shell
{ asymmetrical (with ascending
{ spiral), 7. STREBLEMIDA.

{Cortical shell simple, with
{ one single chamber, 8. PHORTICIDA.
Cortical shell with quite {
irregular growth. {Cortical shell composed of
{ a number of heaped up
{ or aggregated chambers, 9. SOREUMIDA.

{605}Family XXIV. #LARCARIDA#, Haeckel, 1883 (Pl. 50, figs. 1, 2).

_Definition._--#Larcoidea# with a regular, completely latticed,
lentelliptical cortical shell, without open gates and annular
constrictions; medullary shell absent or simple (not trizonal), connected
with the cortical shell by radial beams.

The family #Larcarida# opens the long series of #Larcoidea# as the most
simple group of this suborder. It commences with _Cenolarcus_, a quite
simple lentelliptical latticed shell, which is characterised by three
unequal isopolar dimensive axes, perpendicular one to another. The major of
these three axes is the longitudinal or principal, the middle is the
lateral or transverse, and the minor is the equatorial or sagittal axis (as
in the human body). Among the three dimensive planes, which are determined
by pairs of these axes, the lateral plane is the largest (halved by the
crossed principal and lateral axes). The intermediate is the sagittal plane
or median plane (halved by the crossed principal and sagittal axes). The
smallest is the equatorial plane or transverse plane (halved by the crossed
lateral and sagittal axes). Therefore the shell has all the characters of
the true _Lentellipsis_ or of the "triaxial ellipsoid," and its axes agree
with the three axes of the "rhombic crystalline system."

In the three subfamilies of Larcarida this lentelliptical shell assumes a
different shape: in the Cenolarcida it remains simple, in the Spongolarcida
it becomes spongy (sometimes quite filled out with a spongy framework), in
the Coccolarcida it is composed of two or more concentric lentelliptical
shells (at least an inner medullary and one outer cortical shell). These
shells are simply connected by radial beams, and not, as in the Larnacida,
by latticed wings (or half girdles).

The network of the Larcarida shell is sometimes regular, commonly irregular
(as in the greater number of #Larcoidea#). The surface of the shell is
sometimes smooth or thorny, at other times covered with radial spines.
These are often symmetrically disposed, either on the poles of the
dimensive axes or in crossed diagonal planes.

The central capsule is a true "lentellipsis" in a geometrical sense; it is
halved by three elliptical dimensive planes of different sizes,
perpendicular one to another. In the Cenolarcida the central capsule lies
freely inside the simple (cortical) shell, only separated from it by the
jelly-mantle. In the Coccolarcida it contains the medullary shell, and is
enclosed by the simple or double cortical shell, perforated by the radial
beams connecting the two shells. The spongy shell of the Spongolarcida
exhibits a different relation to the central capsule: in _Spongolarcus_ the
latter lies freely in the internal cavity of the spongy shell; in
_Stypolarcus_, where this cavity is quite filled with a spongy network, the
central capsule also contains a part of it.

The morphological and phylogenetic relations of the Larcarida to the other
families of SPUMELLARIA admit of a different explanation. As this family
contains the {606}most simple forms of all #Larcoidea#, we can regard the
_Cenolarcus_ as the common ancestral form of this group, having originated
from _Actilarcus_ (or the lentelliptical _Actissa_) by the building of a
simple lentelliptical lattice-shell. But it is also possible that a part of
the Larcarida (or all?) descend from Larnacida by reduction or loss of the
original _Larnacilla_-shell (compare _Cenolarcus triaxonius_, p. 607).

_Synopsis of the Genera of the Larcarida._

-------------------------------------------------------------------------
I. Subfamily Cenolarcida.
Shell simple, latticed (lentelliptical cortical shell).
-------------------------------------------------------------------------
Without radial spines, 266. _Cenolarcus_.

With radial spines, 267. _Larcarium_.
-------------------------------------------------------------------------
II. Subfamily Coccolarcida.
Shell composed of two or more concentric latticed shells (inner
medullary and outer cortical).
-------------------------------------------------------------------------
Without radial spines, 268. _Coccolarcus_.

With radial spines, 269. _Larcidium_.
-------------------------------------------------------------------------
III. Subfamily Spongolarcida.
Shell spongy, partly or whole composed of a spongy framework.
-------------------------------------------------------------------------
With an internal cavity, 270. _Spongolarcus_.

Without an internal cavity, 271. _Stypolarcus_.

Subfamily 1. CENOLARCIDA, Haeckel.

_Definition._--#Larcarida# with a simple, lentelliptical latticed shell
(cortical shell without a medullary shell).

Genus 266. _Cenolarcus_,[305] n. gen.

_Definition._--#Larcarida# with a simple, lentelliptical latticed shell,
without a medullary shell, without radial spines.

The genus _Cenolarcus_ begins the group of #Larcoidea# as the most simple
form of this suborder. It corresponds to _Cenosphæra_ among the
#Sphæroidea#, to _Cenodiscus_ among the #Discoidea#, to _Cenellipsis_ among
the #Prunoidea#. The simple latticed shell is distinguished from that of
the three other genera by its typical lentelliptical form, a triaxial
ellipsoid with three dimensive axes of unequal length. Probably
_Cenolarcus_ is the original ancestral form of the #Larcoidea#, derived
from _Actilarcus_ (the lentelliptical _Actissa_) by the formation of a
simple fenestrated shell around the lentelliptical central capsule. But
possibly also some species of _Cenolarcus_ may be derived from
_Coccolarcus_ or _Larnacilla_ by reduction and loss of the medullary shell
(compare _Cenolarcus triaxonius_, n. sp.).

{607}1. _Cenolarcus primordialis_, n. sp. (Pl. 50, figs. 7, 7_a_, 7_b_).

Network of the shell regular, with circular, hexagonally framed pores,
twice as broad as the elevated bars; about ten pores on the half meridian,
eight on the half equator. Surface a little rough. Proportion of the three
dimensive axes = 2 : 2.5 : 3.

_Dimensions._--Principal axis (or length) 0.12, transverse axis (or
breadth) 0.1, sagittal axis (or thickness) 0.08; pores 0.01, bars 0.005.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

2. _Cenolarcus dimensivus_, n. sp.

Network of the shell regular, with circular pores (without hexagonal
frames), three times as broad as the thin bars; about twelve pores on the
half meridian, nine on the half equator. Surface thorny. Proportion of the
three dimensive axes = 3 : 4 : 5.

_Dimensions._--Principal axis 0.15, transverse axis 0.12, sagittal axis
0.09; pores 0.012, bars 0.004.

_Habitat._--North Pacific, Station 244, surface.

3. _Cenolarcus triaxonius_, n. sp.

Network of the shell regular, with circular, hexagonally framed pores, four
times as broad as the thin bars; about twelve pores on the half meridian,
nine on the equator. Surface smooth. From the inner surface of the shell
arise six very thin radial beams, opposite in pairs in the three dimensive
axes; all six beams end freely in a little knob, at an equal distance from
the centre; therefore this remarkable species seems to have lost a
medullary shell (descending from _Coccolarcus_ or _Larnacilla_?).
Proportion of the three dimensive axes = 2 : 2.5 : 3.

_Dimensions._--Principal axis 0.13, transverse axis 0.11, sagittal axis
0.09; pores 0.012, bars 0.03.

_Habitat._--Tropical Atlantic, Station 338, depth 1990 fathoms.

4. _Cenolarcus lentellipticus_, n. sp.

Network of the shell regular, with circular pores of the same breadth as
the thick bars; about eighteen pores on the half meridian, fourteen on the
half equator. Surface smooth. Proportion of the three dimensive axes =
2 : 3 : 4.

_Dimensions._--Principal axis 0.16, transverse axis 0.12, sagittal axis
0.08; pores and bars 0.004.

_Habitat._--Western Tropical Pacific, Station 224, surface.

5. _Cenolarcus minimus_, n. sp.

Network of the shell subregular, with very small circular pores of the same
breadth as the bars; only four pores on the half meridian, three on the
half equator. Surface smooth. Proportion of the three dimensive axes =
3 : 4 : 5.

{608}_Dimensions._--Principal axis 0.05, transverse 0.04, sagittal axis
0.03; pores and bars 0.006.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Genus 267. _Larcarium_,[306] n. gen.

_Definition._--#Larcarida# with a simple, lentelliptical latticed shell,
without a medullary shell; surface covered with radial spines.

The genus _Larcarium_ differs from _Cenolarcus_ only in the possession of
radial spines on the surface of the simple fenestrated lentelliptical
shell. These spines are commonly disposed symmetrically, opposite in pairs,
either on the poles of the three dimensive axes, or on the poles of certain
diagonal axes. Larcarium differs from the similar genera _Larcidium_,
_Larnacidium_, and _Larnacantha_ by the absence of any medullary shell.

1. _Larcarium amphistylum_, n. sp.

Shell thorny, with two large conical spines, opposite on both poles of the
principal axis, somewhat longer than it. Network of the shell regular, with
circular, hexagonally framed pores, twice as broad as the bars; about
twelve pores on the half meridian, ten on the half equator. Proportion of
the three dimensive axes = 3 : 4 : 5.

_Dimensions._--Principal axis (or length) 0.15, transverse axis (or
breadth) 0.12, sagittal axis (or thickness) 0.09; pores 0.01, bars 0.005.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

2. _Larcarium staurostylum_, n. sp.

Shell smooth, with four short three-sided pyramidal spines of equal length,
opposite in pairs on the poles of the principal and lateral axes. Network
of the shell regular, with circular pores of the same breadth as the bars;
about eight pores on the half meridian, six on the half equator. Proportion
of the three dimensive axes = 2 : 2.5 : 3.

_Dimensions._--Length of the shell 0.12, breadth 0.1, thickness 0.08; pores
and bars 0.006.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

3. _Larcarium hexastylum_, n. sp.

Shell smooth, with six short conical spines of equal length (= the shortest
axis of the shell), opposite in pairs on the poles of the three dimensive
axes. Network of the shell subregular, with {609}circular pores three times
as broad as the bars; about eleven pores on the half meridian, nine on the
half equator. Proportion of the three dimensive axes = 3 : 4 : 5.

_Dimensions._--Length of the shell 0.1, breadth 0.08, thickness 0.06, pores
0.006; bars 0.002.

_Habitat._--North Pacific, Station 244, surface.

4. _Larcarium axostylum_, n. sp.

Shell thorny, with six strong conical radial spines, opposite in pairs on
the poles of the three dimensive axes. All three pairs are of different
sizes, the length of each spine corresponding nearly to the size of the
shell-axis, of which it is the prolongation. Network of the shell
subregular, with circular pores twice as broad as the bars; ten on the half
meridian, eight on the half equator. Proportion of the three dimensive axes
= 2 : 3 : 4.

_Dimensions._--Length of the shell 0.12, breadth 0.09, thickness 0.06;
pores 0.01, bars 0.005.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

5. _Larcarium octostylum_, n. sp.

Shell thorny, with eight thin cylindrical radial spines, opposite in pairs
in two crossed diagonal planes. Network of the shell irregular, with
roundish pores, twice to four times as broad as the bars; nine to eleven on
the half meridian, five to seven on the half equator. Proportion of the
three dimensive axes = 3 : 4 : 5.

_Dimensions._--Length of the shell 0.11, breadth 0.09, thickness 0.07;
pores 0.006 to 0.012, bars 0.003.

_Habitat._--Pacific, central area, Station 267, surface.

6. _Larcarium polystylum_, n. sp.

Shell thorny, with numerous (twenty to thirty) stronger conical radial
spines, about as long as the shortest axis of the shell. Network of the
shell irregular, with roundish pores, twice to four times as broad as the
bars; seven to eight on the half meridian, five to six on the half equator.
Proportion of the three dimensive axes = 1 : 2 : 3.

_Dimensions._--Length of the shell 0.09, breadth 0.06, thickness 0.03;
pores 0.006 to 0.012, bars 0.003.

_Habitat._--South Atlantic, Station 323, surface.

7. _Larcarium chætostylum_, n. sp.

Shell bristly, with very numerous (sixty to eighty or more) very thin,
bristle-like, radial spines, somewhat longer than the longest axis of the
shell. Network irregular, with very small roundish pores, about the same
size as the bars; sixteen to eighteen on the half meridian, thirteen to
fifteen on the half equator. Proportion of the three dimensive axes =
1 : 1.5 : 2.

{610}_Dimensions._--Length of the shell (without spines) 0.13, breadth 0.1,
thickness 0.07; pores and bars 0.004 to 0.006.

_Habitat._--Pacific, central area, Station 273, surface.

Subfamily 2. COCCOLARCIDA, Haeckel.

_Definition._--#Larcarida# with encased lentelliptical shell, composed of
two or more concentric lentelliptical latticed shells, which are united by
radial beams (at least one inner medullary shell and one outer cortical
shell).

Genus 268. _Coccolarcus_,[307] n. gen.

_Definition._--#Larcarida# with two concentric latticed shells, an outer
lentelliptical cortical shell, and an inner (spherical or lentelliptical)
medullary shell, both connected by radial beams. Surface without radial
spines.

The genus _Coccolarcus_ differs from _Cenolarcus_ by the possession of an
internal medullary shell. This is quite simple, either spherical or
lentelliptical, and connected with the outer cortical shell by a number of
radial beams. In the similar _Larnacilla_ this connection is effected by
four internal latticed lamellæ (the half lateral wings of the transverse
girdle); therefore we find here four internal gates (on the poles of the
principal axis), absent in _Coccolarcus_.

1. _Coccolarcus lentellipsis_, n. sp.

Cortical shell with smooth surface and regular network; pores circular,
twice as broad as the bars; about thirteen on the half meridian, eleven on
the half equator. Proportion of the three dimensive axes = 3 : 4 : 5.
Medullary shell spherical, half as broad as the transverse radius.

_Dimensions._--Principal axis (or length) of the cortical shell 0.15,
transverse axis (or breadth) 0.12, sagittal axis (or thickness) 0.09; pores
0.01, bars 0.005; diameter of the medullary shell 0.03.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Coccolarcus platellipsis_, n. sp.

Cortical shell with thorny surface and irregular network; pores roundish,
twice to three times as broad as the bars; sixteen to eighteen on the half
meridian, ten to twelve on the half equator. Proportion of the three
dimensive axes = 1 : 2 : 3. Medullary shell lentelliptical, one-third as
large as the cortical shell.

{611}_Dimensions._--Length of the cortical shell 0.17, breadth 0.11,
thickness 0.06; pores 0.008 to 0.012, bars 0.004; medullary shell 0.03 to
0.06.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

Genus 269. _Larcidium_,[308] n. gen.

The genus _Larcidium_ differs from the foregoing _Coccolarcus_ only in the
possession of radial spines, and bears to it the same relation that
_Larcarium_ does to _Cenolarcus_. The spines are commonly symmetrically
disposed, opposite in pairs in the dimensive axes, sometimes also in
diagonal axes.

1. _Larcidium dissacanthum_, n. sp.

Cortical shell with thorny surface and regular network; pores circular,
hexagonally framed, three times as broad as the bars; about thirteen on the
half meridian, eleven on the half equator. Proportion of the three
dimensive axes = 2 : 3 : 5. Medullary shell spherical, one-fourth as broad
as the cortical, connected with it by two thin beams, lying in the
principal axis, and prolonged on its poles into two strong conical spines,
somewhat longer than the greatest axis.

_Dimensions._--Length of the cortical shell (or principal axis) 0.15,
breadth 0.09, thickness 0.07; pores 0.015, bars 0.005; medullary shell
0.025.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

2. _Larcidium hexacanthum_, n. sp.

Cortical shell with smooth surface and regular network; pores circular,
twice as broad as the bars; about eleven on the half meridian, nine on the
half equator. Proportion of the three axes = 2 : 2.5 : 3. Medullary shell
spherical, one-third as broad as the cortical. On the surface six strong,
three-sided pyramidal spines, all about as long as the breadth of the
cortical shell.

_Dimensions._--Length of the cortical shell 0.12, breadth 0.1, thickness
0.08; pores 0.01, bars 0.005; medullary shell 0.035.

_Habitat._--Pacific, central area. Station 266, surface.

3. _Larcidium axacanthum_, n. sp.

Cortical shell with rough surface and irregular network; pores roundish,
twice to four times as broad as the bars; about fifteen to nineteen on the
half meridian, twelve to fourteen on the half {612}equator. Proportion of
the three axes = 2 : 3 : 4. Medullary shell lentelliptical, of the same
form and structure as the cortical, but only one-third as large, connected
with it by six thin radial beams, lying in pairs in the three dimensive
axes; on the outside they are prolonged into six strong conical radial
spines, which are in pairs of different size (as in _Larcarium axostylum_);
the length of each spine nearly equals the axis of the cortical shell, of
which it is the prolongation.

_Dimensions._--Length of the cortical shell (and the principal spines)
0.18, breadth of it (and length of the lateral spines) 0.135, thickness of
it (and length of the sagittal spines) 0.09; pores 0.005 to 0.013, bars
0.003.

_Habitat._--Pacific, central area, Station 271, surface.

4. _Larcidium octacanthum_, n. sp.

Cortical shell thorny, with irregular network; pores roundish, once to four
times as broad as the bars; about thirteen to fifteen on the half meridian,
eleven to thirteen on the half equator. Proportion of the three axes =
1 : 2.5 : 4. Medullary shell lentelliptical, one-fifth as large as the
cortical, connected with it by eight radial beams, which are situated in
two crossed diagonal planes (opposite in pairs), and are prolonged on the
surface into eight long and thin cylindrical radial spines similar to
_Tetrapyle octacantha_.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.11, thickness
0.045; pores 0.003 to 0.012, bars 0.003; medullary shell 0.02 to 0.03.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

5. _Larcidium dodecanthum_, n. sp. (Pl. 50, figs. 8, 8_a_).

Cortical shell rough, with irregular network; pores roundish, twice to four
times as broad as the bars; about twelve to fourteen on the half meridian,
eight to ten on the half equator. Proportion of the three axes = 2 : 3 : 4.
Medullary shell lentelliptical, one-third as large as the cortical,
connected with it by twelve thin radial beams, which are prolonged outside
into twelve strong conical radial spines, about half as long as the breadth
of the shell.

_Dimensions._--Length of the cortical shell 0.14, breadth 0.1, thickness
0.07; pores 0.008 to 0.015, bars 0.004; medullary shell 0.04 to 0.05.

_Habitat._--South Pacific, Station 288, surface.

6. _Larcidium polyacanthum_, n. sp.

Cortical shell spiny, with irregular network; pores roundish, once to twice
as broad as the bars; about ten to twelve on the half meridian, six to
eight on the half equator. Proportion of the three axes = 1 : 2 : 3.
Medullary shell lentelliptical, half as large as the cortical shell,
connected with it by numerous (twenty to twenty-five) radial beams, which
are prolonged outside into three-sided pyramidal spines, about as long as
the breadth of the shell.

_Dimensions._--Length of the cortical shell 0.18, breadth 0.12, thickness
0.06; pores 0.01 to 0.015, bars 0.008; medullary shell 0.03 to 0.09.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{613}Subfamily 3. SPONGOLARCIDA, Haeckel.

_Definition._--#Larcarida# with spongy lentelliptical shell (with or
without enclosed medullary shell).

Genus 270. _Spongolarcus_,[309] n. gen.

_Definition._--#Larcarida# with lentelliptical spongy shell, containing a
central cavity of the same form, without medullary shell (without radial
spines).

The genus _Spongolarcus_ differs from _Cenolarcus_ (its probable ancestral
form) only in the development of spongy framework forming the wall of the
hollow lentelliptical shell. It corresponds, therefore, to _Plegmosphæra_
among the #Sphæroidea#, and to _Spongellipsis_ among the #Prunoidea#. From
these two similar spongy #Sphærellaria# it differs in its characteristic
lentelliptical form, with three dimensive axes of unequal length.

1. _Spongolarcus lentellipsis_, n. sp.

Spongy network of the shell very loose, its meshes fifteen to twenty times
as broad as the bars. Surface of the shell nearly smooth; diameter of its
internal cavity twice as large as the thickness of its wall. Proportion of
the three dimensive axes = 2 : 3 : 4.

_Dimensions._--Length 0.16, breadth 0.12, height 0.08; thickness of the
spongy wall 0.05.

_Habitat._--Pacific, central area, Station 274, surface.

2. _Spongolarcus triaxonius_, n. sp.

Spongy network of the shell rather loose, its meshes twelve to sixteen
times as broad as the bars. Surface of the shell rough; diameter of its
internal cavity about eight times as large as the thickness of its wall.
Proportion of the three dimensive axes = 2 : 2.5 : 3.

_Dimensions._--Length 0.25, breadth 0.2, height 0.16; thickness of the
spongy wall 0.025.

_Habitat._--North Pacific, Station 253, surface.

3. _Spongolarcus dimensivus_, n. sp.

Spongy network of the shell dense, its meshes four to eight times as broad
as the bars. Surface of the shell thorny; diameter of its internal cavity
about fifteen times as large as the thickness of its wall. Proportion of
the three dimensive axes = 3 : 4 : 5.

_Dimensions._--Length 0.2, breadth 0.16, height 0.12; thickness of the
spongy wall 0.01.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

{614}4. _Spongolarcus amphicentria_, Haeckel.

? _Amphicentria salpa_, Ehrenberg, 1861, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 296; Abhandl. d. k. Akad. d. Wiss. Berlin, 1872, Taf.
ii. fig. 18.

? _Spongurus salpa_, Haeckel, 1862, Monogr. d. Radiol., p. 466.

Spongy network of the shell compact, its meshes about the same breadth as
the bars. Surface of the shell spiny, with some larger spines around the
poles of the axis; diameter of the internal cavity about six times as large
as the thickness of its wall. Proportion of the three dimensive axes =
1 : 2 : 3. (Perhaps this _Spongolarcus_ is identical with _Amphicentria
salpa_, very imperfectly described and figured by Ehrenberg, _loc. cit._?)

_Dimensions._--Length 0.14, breadth 0.09, height 0.05; thickness of the
spongy wall 0.015.

_Habitat._--North Atlantic; off Greenland, 1000 fathoms, Ehrenberg; Station
64, depth (2700) fathoms.

Genus 271. _Stypolarcus_,[310] n. gen.

_Definition._--#Larcarida# with lentelliptical spongy shell, composed of
compact spongy framework, without central cavity and medullary shell
(without radial spines).

The genus _Stypolarcus_ differs from _Spongolarcus_ in the absence of any
central cavity. This is quite filled up by spongy framework, which forms
the whole mass of the lentelliptical body. _Stypolarcus_ bears therefore
the same relation to _Spongolarcus_ that _Styptosphæra_ does to
_Plegmosphæra_.

1. _Stypolarcus spongiosus_, n. sp.

Lentelliptical shell composed in the whole mass of loose, spongy framework
of similar texture, with irregular meshes, about ten to twenty times as
broad as the thin bars. Surface rough, without radial spines. Proportion of
the three axes = 3 : 4 : 5.

_Dimensions._--Length 0.2, breadth 0.16, height 0.12.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Family XXV. #LARNACIDA#, Haeckel, 1883 (Pl. 50, figs. 3-8).

_Definition._--#Larcoidea# with a regular, completely latticed,
lentelliptical cortical shell, without open gates and annular
constrictions; either this cortical shell or the enclosed medullary shell
is trizonal, composed of three elliptical, latticed, dimensive girdles of
different sizes, perpendicular one to another.

{615}The family #Larnacida# immediately follows the Larcarida as the next
simple group of all #Larcoidea#; some genera of both groups (such as
_Larnacalpis_ and _Coccolarcus_, or _Larnacantha_ and _Larcidium_) may
easily be confounded from their being so much alike. In both the
lentelliptical shell is composed of two concentric shells, an inner
(medullary) and an outer (cortical) shell. But the connection between these
shells and the construction of the inner shell is quite different in the
two groups. Whilst in the Larcarida the medullary shell is connected with
the cortical shell simply by radial beams, here in the Larnacida this
connection is effected by two latticed lamellæ, which are the lateral wings
of a transverse girdle. Therefore we encounter here for the first time that
peculiar mode of growth which characterises the greater part of the
#Larcoidea#, but particularly the Pylonida. But whilst in the Pylonida
between the three crossed lattice-girdles, remain four open gates, here in
the Larnacida the gates become closed by lattice-work; the lentelliptical
cortical shell becomes perfect.

The most simple genus of Larnacida, and no doubt the common ancestral form
of this whole family, is _Larnacilla_ (Pl. 50, figs. 1, 1_a_, 1_b_). The
most important shell of this typical genus is composed of a simple
lentelliptical medullary shell and of three elliptical latticed girdles
surrounding it, perpendicular one to another. These three "dimensive
girdles" lie in the perimeter of the three dimensive planes, the minor (and
first) in the equatorial plane, the second (and major) in the lateral
plane, the third (and intermediate) in the sagittal plane. Therefore we
have before us the same "trizonal shell" as in the important genus
_Trizonium_ among the Pylonida. But whilst in _Trizonium_, as in all
Pylonida, the four gates between the girdles remain open, here in
_Larnacilla_ they become perfectly closed by lattice-work.

The formation of the typical "_Larnacilla_-shell" begins with a simple,
spherical or lentelliptical lattice-shell, from both sides of which arise
two latticed "lateral wings" opposite on the poles of the transverse axis.
These two wings are comparable to the lateral chambers of _Tholartus_
(among the Tholonida), but differ by two large openings. Each wing is a
short cylindrical tube with latticed wall, open at both ends; the axis of
the tube (going through the centre of the open ends) is parallel to the
principal axis of the whole shell (and of the central chamber). Therefore
both wings form together a transverse ring, the middle of which encloses
the central chamber. The distal parts of both wings grow towards the poles
of the principal axis; if they became united here, the second (lateral)
girdle would be complete. Between it and the first girdle four open gates
remain ("_Tetrapyle_"); but these become overgrown by the third or sagittal
girdle, and at last the gates between this and the two other girdles become
closed by lattice-work. This perfect fenestration of the trizonal cortical
shell, and the complete closing of the gates between the girdles by
network, is the only difference between _Trizonium_ and _Larnacilla_.

In _Larnacilla_ and in the nearly allied _Larnacidium_ (only differing by
radial spines {616}on the surface) the "trizonal shell" is an external or
"cortical shell," enclosing the central capsule, the interior of which only
contains the simple central chamber and the jointed proximal parts of both
lateral wings. In the other genera of Larnacida (by proceeding growth) this
trizonal _Larnacilla_-shell becomes enclosed by the growing central capsule
and is now only a "medullary shell," whilst on the outside of the central
capsule in the same manner is developed an outer cortical shell
(_Larnacalpis_, _Larnacantha_); and perhaps the same process may be
repeated. But sometimes also this cortical shell becomes doubled by a
simple envelop of network (_Larnacoma_). In the Larnacospongida the
cortical shell is composed of a spongy framework (corresponding to the
Spongolarcida in the foregoing family).

The lattice-work of the Larnacida is commonly irregular (as in most other
#Larcoidea#), and its pores have little signification for the different
species. On the outer surface often arise radial spines, symmetrically
disposed either in dimensive planes or in diagonal planes.

The central capsule is constantly a true lentellipsis or a "triaxial
ellipsoid," characterised by three halving, elliptical dimensive planes,
perpendicular one to another. It bears a different relation to the skeleton
in the two subfamilies of Larnacida. In the first subfamily, the
Larnacillida (_Larnacilla_, _Larnacidium_), the central capsule encloses
only the simple spherical or lentelliptical central chamber ("simple
medullary shell"), and is enveloped by the trizonal cortical shell. In the
other subfamily, the Larnacalpida (_Larnacalpis_, _Larnacospongus_, &c.),
this trizonal "_Larnacilla_-shell" becomes enclosed by the overgrowing
central capsule, which now becomes enveloped by an external, latticed or
spongy, lentelliptical "cortical shell."

_Synopsis of the Genera of Larnacida._

-------------------------------------------------------------------------
I. Subfamily Larnacillida.
Medullary shell simple, spherical or subspherical. Cortical shell
lentelliptical, trizonal; between them four gates.
-------------------------------------------------------------------------
Shell without radial spines, 272. _Larnacilla_.

Shell with radial spines, 273. _Larnacidium_.
-------------------------------------------------------------------------
II. Subfamily Larnacalpida.
Medullary shell _Larnacilla_-shaped, double; the inner spherical or
subspherical, the outer lentelliptical and trizonal.
-------------------------------------------------------------------------
{ { Without radial
{ Cortical { spines, 274. _Larnacalpis_.
{ shell {
Cortical shell simple { simple. { With radial
or double, but not { { spines, 275. _Larnacantha_.
spongy. {
{ Cortical shell double,
{ without radial spines, 276. _Larnacoma_.

{ Shell without radial
Cortical shell simple { spines, 277. _Larnacospongus_.
or double, wholly or {
partly spongy. { Shell with radial spines, 278. _Larnacostupa_.

{617}Subfamily 1. LARNACILLIDA, Haeckel.

_Definition._--#Larnacida# with a simple, spherical or lentelliptical,
medullary shell, connected by the lateral wings of a latticed transverse
girdle with the simple lentelliptical trizonal cortical shell; the central
capsule encloses the former and is enveloped by the latter.

Genus 272. _Larnacilla_,[311] n. gen.

_Definition._--#Larnacida# with a simple lentelliptical cortical shell,
connected by the lateral wings of a latticed transverse girdle with the
simple, spherical or lentelliptical, medullary shell. Surface without
radial spines.

The genus _Larnacilla_ represents the most simple form of Larnacida, and at
the same time the most important common ancestral form, from which the
greater number of #Larcoidea# may be derived, viz., all those genera which
possess the characteristic "_Larnacilla_-shaped medullary shell." This
typical form of medullary shell may be derived from the genus _Trizonium_
among the Pylonida by the closing of the four open gates of this genus. The
free opening of these four gates becomes overgrown and closed by
lattice-work, developed from the free edges of the three crossed girdles,
and thus finally all three girdles are united in the form of a simple
lentelliptical shell (Pl. 50, figs. 1, 1_a_, 1_b_). Seen from the sagittal
poles (or from the poles of the shortest axis, fig. 1), the shell exhibits
on both sides of the small spherical medullary shell the two lateral wings
of the transverse girdle from the face; seen from the lateral poles (or
from the poles of the transverse axis, fig. 1_a_), one of these wings
appears in the optical section as an oblong ring, which seemingly encloses
the concentric medullary shell, and on both sides is grown together with
the sagittal girdle; seen from the principal poles (or from the poles of
the longitudinal axis, fig. 1_b_), both wings exhibit their elliptical
opening (at the right and left from the central medullary shell). The two
concentric shells are only connected by the two lateral tube-like wings of
the transverse girdle; the lateral and the sagittal girdles have no
connection with the medullary shell. The latter is sometimes spherical, at
other times lentelliptical.

1. _Larnacilla typus_, n. sp. (Pl. 50, fig. 1, 1_a_, 1_b_).

Cortical shell with smooth surface and with subregular network; pores twice
as broad as the bars; about twelve pores on the half meridian, ten on the
half equator. Proportion of the three dimensive axes = 2 : 3 : 4. Internal
four gates (between transverse and lateral girdles) roundish-triangular,
little broader than high. Medullary shell spherical, scarcely one-third as
broad as the lentelliptical cortical shell.

{618}_Dimensions._--Length of the cortical shell (or principal axis) 0.13,
breadth of it (or transverse axis) 0.1, height of it (or sagittal axis)
0.07; pores 0.006, bars 0.003; medullary shell (diameter) 0.03.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

2. _Larnacilla prometor_, n. sp.

Cortical shell with rough surface and regular network; pores three times as
broad as the bars; about ten on the half meridian, eight on the half
equator. Proportion of the three axes = 1 : 2 : 3. Internal four gates
(between transverse and lateral girdles) kidney-shaped, twice as broad as
high. Medullary shell lentelliptical, of the same form as the external
cortical shell, but only one third as large.

_Dimensions._--Length of the cortical shell 0.15, breadth 0.1, height 0.05,
pores 0.015, bars 0.005; medullary shell 0.02 to 0.05.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

3. _Larnacilla subglobosa_, n. sp.

Cortical shell nearly spherical, with thorny surface and irregular network;
pores roundish, twice to four times as broad as the bars; twelve to sixteen
in the half circumference. Proportion of the three axes very little
different = 1.3 : 1.4 : 1.5. Internal four gates elliptical, one and a half
times as broad as high. Medullary shell spherical, one-fourth as broad as
the cortical shell.

_Dimensions._--Length of the cortical shell 0.15, breadth 0.14, height
0.13; pores 0.006 to 0.012, bars 0.003; medullary shell 0.035.

_Habitat._--Pacific, central area. Station 266, depth 2750 fathoms.

4. _Larnacilla medullaris_, n. sp.

Cortical shell lentelliptical, with smooth surface and irregular network;
pores roundish, very small, scarcely as broad as the bars; about eight to
nine on the half meridian, six to seven on the half equator. Proportion of
the three axes = 3 : 4 : 6. Internal four gates elliptical. Medullary shell
spherical, scarcely one-third as broad as the cortical shell. (This small
species may be only the medullary shell of some other Larcoid, the cortical
shell of which is not yet formed.)

_Dimensions._--Length of the cortical shell 0.06, breadth 0.04, height
0.03; pores and bars about 0.004; medullary shell 0.013.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Genus 273. _Larnacidium_,[312] n. gen.

{619}The genus _Larnacidium_ has the same shell-formation as the foregoing
_Larnacilla_, and differs from it only in the possession of radial spines
on the surface, which in all known species exhibit a symmetrical
disposition. From the nearly allied genus _Larcidium_ it differs in the
characteristic mode of connection between the two shells, owing to the
different kind of growth. In _Larcidium_ this connection is effected only
by radial beams, whereas in _Larnacidium_ (as in all Larnacida) by two
lateral latticed tubes, the wings of the primary transverse girdle.

1. _Larnacidium staurobelonium_, n. sp.

Cortical shell smooth, with four strong conical, radial spines in the
lateral plane, opposite in pairs, two on the poles of the principal and two
on the poles of the transverse axis. Pores subregular, circular, twice as
broad as the bars; about twelve on the half meridian. Proportion of the
three axes = 2 : 3 : 4. Medullary shell spherical, one-third as broad as
the cortical shell.

_Dimensions._--Length of the cortical shell 0.12, breadth 0.09, height
0.06; pores 0.008, bars 0.04; medullary shell 0.03.

_Habitat._--North Pacific Station 244, depth 2900 fathoms.

2. _Larnacidium hexabelonium_, n. sp.

Cortical shell thorny, with six strong, three-sided pyramidal, radial
spines, lying opposite in pairs on the poles of the three dimensive axes.
Pores subregular, circular, three times as broad as the bars; about
fourteen on the half meridian. Proportion of the three axes = 2 : 2.5 : 3.
Medullary shell spherical, one-third as broad as the cortical shell.

_Dimensions._--Length of the cortical shell 0.14, breadth 0.11, height
0.08; pores 0.01, bars 0.003; medullary shell 0.04.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

3. _Larnacidium polybelonium_, n. sp.

Cortical shell very spiny, with numerous (twenty to thirty or more) larger
thin radial spines, about as long as the shell. Pores irregular, twice to
five times as broad as the bars; about sixteen on the half meridian.
Proportion of the three axes = 2 : 2.5 : 3. Medullary shell
lentelliptical, half as large as the cortical shell.

_Dimensions._--Length of the cortical shell 0.11, breadth 0.09, height
0.07; pores 0.004 to 0.01, bars 0.002; length of the medullary shell 0.06,
breadth 0.05, height 0.04.

_Habitat._--Western Tropical Pacific, Station 225. depth 4475 fathoms.

Subfamily 2. LARNACALPIDA, Haeckel.

_Definition._--#Larnacida# with a double, trizonal, _Larnacilla_-shaped
medullary shell, enclosed in the central capsule, and enveloped by a simple
or double, latticed or spongy, lentelliptical, cortical shell.

{620}Genus 274. _Larnacalpis_,[313] n. gen.

_Definition._--#Larnacida# with a simple lentelliptical cortical shell,
without radial spines. Medullary shell double, _Larnacilla_-shaped.

The genus _Larnacalpis_ represents the most simple form of the sub-family
Larnacalpida, and is very important as the common original form of all
those #Larcoidea# in which a double _Larnacilla_-shaped medullary shell is
surrounded by a simple, perfectly closed, latticed, lentelliptical cortical
shell. Therefore the same typical, trizonal, lentelliptical shell, which in
_Larnacilla_ represents the external envelop (or cortical shell) of the
central capsule, here in _Larnacalpis_ becomes enclosed as an internal
nucleus (or medullary shell) in the interior of the central capsule, and
this latter becomes overgrown by a new lentelliptical cortical shell. The
connection between the two shells of _Larnacalpis_ is either effected by a
number of radial beams (e.g., in _Larnacalpis triaxonia_ by six beams
situated in the three dimensive axes), or by two lateral, latticed,
tube-like wings, which are repetitions of the smaller lateral wings
connecting its external shell with the internal medullary shell (as in
_Larnacalpis lentellipsis_). The latter species may be regarded as a
_Pylonium_ with a completely latticed shell.

1. _Larnacalpis lentellipsis_, n. sp. (Pl. 50, figs. 2, 2_a_, 2_b_).

Cortical shell with thorny surface and irregular network; pores roundish,
twice to four times as broad as the bars; about sixteen on the half
meridian, twelve on the half equator. Proportion of the three dimensive
axes = 2 : 3 : 4. Medullary shell one-third as large as the cortical, with
four elliptical internal gates, connected with it by two opposite beams in
the principal axis and by two latticed wings in the transverse axis;
therefore between the two shells are four large kidney-shaped gates, halved
by the polar beams (as in _Octopyle_).

_Dimensions._--Length of the cortical shell (or principal axis) 0.14,
breadth (or transverse axis) 0.11, height (or sagittal axis) 0.07; pores
0.01 to 0.02, bars 0.005; length of the medullary shell 0.05, breadth 0.04,
height 0.03.

_Habitat._--Pacific, central area, Station 273, depth 2600 fathoms.

2. _Larnacalpis phacodiscus_, n. sp.

Cortical shell with thorny surface and regular network; pores circular,
twice as broad as the bars; about ten on the half meridian, eight on the
half equator. Proportion of the three axes = 2 : 2.5 : 3. Medullary shell
half as large as the cortical, with four kidney-shaped internal gates.

_Dimensions._--Length of the cortical shell 0.13, breadth 0.11, height
0.09; pores 0.012, bars 0.006; length of the medullary shell 0.06, breadth
0.045, height 0.03.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

{621}3. _Larnacalpis macrococcus_, n. sp.

Cortical shell with spiny surface and regular network; pores circular,
small, of the same breadth as the bars; about twenty-two on the half
meridian, nineteen on the half equator. Proportion of the three axes =
2 : 3 : 4. Medullary shell two-thirds as large as the cortical, with four
wide internal semicircular gates.

_Dimensions._--Length of the cortical shell 0.1, breadth 0.075, height
0.05; pores and bars 0.003; length of the medullary shell 0.066, breadth
0.05, height 0.032.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

4. _Larnacalpis subsphærica_, n. sp.

Cortical shell with rough surface and irregular network; roundish pores
twice to five times as broad as the bars; about twenty-four on the half
meridian, twenty-one on the half equator. Proportion of the three axes =
1.2 : 1.3 : 1.4. Medullary shell half as large as the cortical, with four
elliptical internal gates.

_Dimensions._--Length of the cortical shell 0.14, breadth 0.13, height
0.12; pores 0.004 to 0.01, bars 0.002; length of the medullary shell 0.08,
breadth 0.07, height 0.06.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

5. _Larnacalpis triaxonia_, n. sp. (Pl. 50, fig. 3).

Cortical shell with smooth surface and peculiar network, composed of four
meridian rows of larger pores (five large elliptical pores on each half
meridian, the largest in the equator) and numerous small irregular pores
between them. Proportion of the three axes = 2 : 3 : 4. Medullary shell
with four semicircular internal gates, about one-fourth as large as the
cortical, connected with it by six thin radial beams, opposite in pairs in
the three dimensive axes.

_Dimensions._--Length of the cortical shell 0.14, breadth 0.1, height 0.07;
large pores 0.03, small pores 0.002 to 0.01, bars 0.002 to 0.01; length of
the medullary shell 0.04, breadth 0.03, height 0.02.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

Genus 275. _Larnacantha_,[314] n. gen.

_Definition._--#Larnacida# with a simple lentelliptical cortical shell,
armed with symmetrically disposed radial spines. Medullary shell double,
_Larnacilla_-shaped.

The genus _Larnacantha_ has the same characteristic shell-formation as the
foregoing _Larnacalpis_, and differs from it only in the possession of
radial spines, which are symmetrically distributed on the surface in a
definite order. Commonly these spines are external prolongations of the
internal radial beams, which connect the double _Larnacilla_-shaped
medullary shell with the simple lentelliptical cortical shell.

{622}1. _Larnacantha dissacantha_, n. sp.

Cortical shell smooth, with two large cylindrical spines, opposite on the
poles of the principal axis, and somewhat longer than it. Pores regular,
circular, three times as broad as the bars; about eleven on the half
meridian. Proportion of the three axes = 2 : 3 : 4. Medullary shell
one-third as large as the cortical shell.

_Dimensions._--Length of the cortical shell (principal axis) 0.13, breadth
(transverse axis) 0.1, height (sagittal axis) 0.07; pores 0.009, bars
0.003; length of the _Larnacilla_-shaped medullary shell 0.045.

_Habitat._--North Pacific, Station 256, depth 2950 fathoms.

2. _Larnacantha stauracantha_, n. sp.

Cortical shell smooth, with four large conical spines in the lateral plane,
two larger opposite on the poles of the principal, two smaller on those of
the transverse axis. Pores regular, circular, twice as broad as the bars;
about seventeen on the half meridian. Proportion of the three axes =
3 : 3.5 : 4. Medullary shell one-third as large as the cortical shell.

_Dimensions._--Length of the cortical shell 0.15, breadth 0.13, height
0.11; pores 0.008, bars 0.004; length of the medullary shell 0.05.

_Habitat._--Pacific, central area, Station 274, surface.

3. _Larnacantha quadricornis_, n. sp.

Cortical shell spiny, with four strong, horn-like curved spines in the
lateral plane, opposite in pairs on the poles of the crossed diagonal axes.
Pores irregular, roundish, twice to four times as broad as the bars; about
fourteen on the half meridian. Proportion of the three axes = 2 : 3 : 4.
Medullary shell one-third as large as the cortical shell.

_Dimensions._--Length of the cortical shell 0.14, breadth 0.11, height
0.07; pores 0.006 to 0.012, bars 0.003; length of the medullary shell 0.05.

_Habitat._--Pacific, central area, Station 270, surface.

4. _Larnacantha hexacantha_, n. sp. (Pl. 50, fig. 4).

Cortical shell thorny, with six strong conical radial spines in the lateral
plane, two opposite on the poles of the principal axis, four others
opposite in pairs on the poles of the two crossed diagonal axes. Pores with
peculiar distribution; twelve large elliptical pores (nearly of the size of
the medullary shell) symmetrically disposed in four crossed meridians
(between the sagittal and the lateral meridians), separated by bands of
smaller irregular pores. Proportion of the three axes = 3 : 4 : 5.
Medullary shell hexagonal, one-third as large as the cortical shell.

_Dimensions._--Length of the cortical shell 0.14, breadth 0.11, height
0.08; large pores 0.04, small pores 0.003 to 0.01, bars 0.004; length of
the medullary shell 0.05.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

{623}5. _Larnacantha bicruciata_, n. sp. (Pl. 50, fig. 5).

Cortical shell thorny, in the lateral plane with eight strong conical
radial spines, alternating with eight smaller spines; four of the eight
stronger spines opposite on the poles of the principal and transverse axes
(in the figure 5, by mistake, not represented large enough), four others
between those, opposite on the poles of the two crossed diagonal axes.
Pores with a peculiar disposition; on both flat sides of the lentellipsis
an elliptical ring of eight large elliptical pores (alternating with the
eight stronger radial spines), separated by bands of smaller irregular
pores. Proportion of the three axes = 2 : 3 : 4. Medullary shell nearly
half as long as the cortical shell.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.13, height
0.08; large pores 0.03, small pores 0.003 to 0.01, bars 0.006; length of
the medullary shell 0.07.

_Habitat._--Indian Ocean, surface, Madagascar (Rabbe).

6. _Larnacantha octacantha_, n. sp.

Cortical shell thorny, with eight long and thin, cylindrical, radial
spines, lying opposite in pairs in two crossed diagonal planes. Pores
irregular, roundish, twice to five times as broad as the bars; about
sixteen on the half meridian. Proportion of the three axes = 1 : 1-1/3 : 2.
Medullary shell scarcely one-fourth as long as the cortical shell. (This
species resembles closely the common _Tetrapyle octacantha_, from which it
seems to be developed by a complete over-growing of the four gates, which
become closed by a network connecting the free edges of the transverse and
lateral girdles.)

_Dimensions._--Length of the cortical shell 0.18, breadth 0.13, height 0.1;
pores 0.005 to 0.015, bars 0.003; length of the medullary shell 0.04.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

7. _Larnacantha cladacantha_, n. sp.

Cortical shell very spiny, with eight longer ramified spines, lying
opposite in pairs in two crossed diagonal planes; each spine with two to
six irregular, lateral branches. Pores irregular, roundish, twice to three
times as broad as the bars; about twelve on the half meridian. Proportion
of the three axes = 5 : 6 : 7. Medullary shell nearly half as long as the
cortical shell. (Differs from the foregoing by the branching spines end the
larger medullary shell.)

_Dimensions._--Length of the cortical shell 0.15, breadth 0.13, height
0.11; pores 0.01 to 0.015, bars 0.005; length of the medullary shell 0.07.

_Habitat._--Indian Ocean, surface, Cocos Islands (Rabbe).

8. _Larnacantha prismatica_, n. sp. (Pl. 50, fig. 6).

Cortical shell smooth, four-sided prismatic, with eight short, parallel,
three-sided pyramidal spines; these lie opposite in pairs in four parallel
longitudinal lines, as prolongations of the four {624}lateral edges of the
prism, and arise from its eight corners. Pores regular, circular, three
times as broad as the bars; about fourteen on the half meridian. Proportion
of the three axes = 2 : 3 : 4. Medullary shell half as long as the cortical
shell.

_Dimensions._--Length of the cortical shell 0.11, breadth 0.07, height
0.05; pores 0.006, bars 0.002; length of the medullary shell 0.06.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

9. _Larnacantha decacantha_, n. sp.

Cortical shell thorny, with ten short and stout, conical, radial spines,
two opposite on the poles of the principal axis (as prolongations of inner
axial beams), eight others opposite in pairs in two crossed diagonal
planes. Pores irregular, roundish, twice to four times as broad as the
bars; about sixteen on the half meridian. Proportion of the three axes =
3 : 3.75 : 4. Medullary shell about one-third as long as the cortical
shell.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.15, height
0.12; pores 0.008 to 0.016, bars 0.004; length of the medullary shell 0.06.

_Habitat._--North Atlantic, Station 354, surface.

10. _Larnacantha dodecantha_, n. sp.

Cortical shell nearly smooth, but with twelve strong conical radial spines;
four in the lateral plane opposite in pairs (two on the poles of the
principal, and two on the poles of the transverse axis); eight others
opposite in pairs in two crossed diagonal planes. Pores with a peculiar
disposition: twelve large elliptical pores in two crossed meridian planes
(alternating with the twelve spines), separated by bands of irregular small
pores. Proportion of the three axes = 1 : 1.5 : 2. Medullary shell
hexagonal, one-third as long as the cortical shell.

_Dimensions._--Length of the cortical shell 0.15, breadth 0.11, height
0.08; large pores 0.03, small pores 0.005 to 0.01, bars 0.003; length of
the medullary shell 0.05.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

11. _Larnacantha drymacantha_, n. sp.

Cortical shell very spiny, on the whole surface covered with a forest of
numerous (thirty to fifty or more) large branched spines, about the length
of the shell; each spine with three to nine lateral branches, simple or
forked (very similar to _Cromyodrymus abietinus_, Pl. 30, fig. 6). Pores
very irregular, roundish. Proportion of the three axes = 2 : 2.5 : 3.
Medullary shell half as long as the cortical shell.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.13, height 0.1;
pores 0.005 to 0.015, bars 0.03; length of the medullary shell 0.08.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{625}Genus 276. _Larnacoma_,[315] n. gen.

_Definition._--#Larnacida# with double lentelliptical cortical shell,
without radial spines. Medullary shell double, _Larnacilla_-shaped.

The genus _Larnacoma_ has originated from the nearly allied _Larnacalpis_
by duplication of the cortical shell. Whilst in both genera the connection
between the _Larnacilla_-shaped medullary shell and the primary cortical
shell is the same, many short radial beams arise from the surface of the
latter in _Larnacoma_, which at constant equal distances from it unite by a
network forming the secondary or outer cortical shell. It differs from the
similar Druppulida (_Cromyodruppa_) by the sagittal flattening of the
lentelliptical shell and the _Larnacilla_-form of the double medullary
shell.

1. _Larnacoma lentellipticum_, n. sp.

Shell with smooth surface and elliptical perimeter, one and a third times
as long as broad. All four shells lentelliptical. Distance between the two
cortical shells about twice as great as the distance of the inner cortical
and outer medullary shell. Network of both outer shells irregular, with
large roundish pores, twice to six times as broad as the bars.

_Dimensions._--Length (or principal axis) of the first (innermost) shell
0.03, second 0.08, third 0.16, fourth (outermost) 0.27, breadth (or
transverse axis) corresponding--(A) 0.02, (B) 0.05, (C) 0.11, (D) 0.2.

_Habitat._--South Atlantic, Station 323, depth 1900 fathoms.

2. _Larnacoma quadruplex_, n. sp.

Shell with thorny surface and elliptical perimeter, one and a fifth times
as long as broad. All four shells lentelliptical. Distance between the two
cortical shells somewhat smaller than the distance between the inner
cortical and outer medullary shell. Network of both outer shells irregular,
with large roundish pores, twice to ten times as broad as the bars.

_Dimensions._--Length of the first shell 0.02, second 0.06, third 0.15,
fourth 0.24; breadth corresponding--(A) 0.016, (B) 0.04, (C) 0.11, (D) 0.2.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

3. _Larnacoma hexagonium_, n. sp.

Shell with thorny surface and hexagonal perimeter, as long as broad. All
four shells hexagonal, connected by six piercing radial beams (two in the
principal axis, four others in two crossed diagonals). Distance between the
two cortical shells somewhat greater than the distance between the inner
cortical and outer medullary shell. Network of both outer shells
subregular, with small circular pores, twice as broad as the bars.

{626}_Dimensions._--Length of the first shell 0.02, second 0.05, third
0.09, fourth 0.16; breadth corresponding--(A) 0.015, (B) 0.035, (C) 0.065,
(D) 0.12.

_Habitat._--South Atlantic, west of Tristan da Cunha, Station 332, depth
2200 fathoms.

Genus 277. _Larnacospongus_,[316] n. gen.

_Definition._--#Larnacida# with spongy lentelliptical cortical shell,
without radial spines. Medullary shell double, _Larnacilla_-shaped.

The genus _Larnacospongus_ differs from the nearly allied genera
_Larnacalpis_ and _Larnacoma_ by the spongy texture of the lentelliptical
cortical shell whilst the enclosed medullary shell in both genera is the
same trizonal _Larnacilla_-shell. Therefore _Larnacospongus_ (and the
following nearly related _Larnacostupa_) can be derived directly by
development of a spongy envelop either from _Larnacilla_ and _Larnacalpis_,
or from _Trizonium_ and _Amphipyle_. But some species of these spongy
genera appear to be derived rather from _Tetrapyle_ or _Pylonium_, perhaps
also from _Cubotholus_. Their phylogenetic origin may be explained in
different ways.

1. _Larnacospongus larnacillifer_, n. sp.

Cortical shell lentelliptical, one and a half times as long as broad, with
rough surface and rather loose spongy framework, directly enclosing a
trizonal _Larnacilla_-shell of the same form, but of only one-third its
size.

_Dimensions._--Length of the whole shell 0.17, breadth 0.12; length of the
medullary shell 0.06, breadth 0.04.

_Habitat._--South Atlantic, east coast of Patagonia, Station 319, surface.

2. _Larnacospongus tetrapylifer_, n. sp.

Cortical shell lentelliptical, one and a third times as long as broad, with
thorny surface; composed of an outer envelop of loose spongy framework and
an inner lattice-shell with four kidney-shaped gates, like _Tetrapyle;_ the
latter encloses a trizonal medullary shell of one-fourth its size.

_Dimensions._--Length of the whole shell 0.22, breadth 0.16; length of the
medullary shell 0.045, breadth 0.035.

_Habitat._--South Atlantic, west of Tristan da Cunha, Station 332, surface.

{627}Genus 278. _Larnacostupa_,[317] n. gen.

_Definition._--#Larnacida# with spongy lentelliptical cortical shell, with
radial spines on the surface. Medullary shell double, _Larnacilla_-shaped.

The genus _Larnacostupa_ differs from the preceding Larnacospongus only in
the possession of radial spines, covering either the whole surface
irregularly or disposed in a certain symmetrical order.

1. _Larnacostupa octacantha_, n. sp.

Cortical shell lentelliptical, with thorny surface, and rather dense,
irregular, spongy framework, which arises from an inner latticed cortical
shell, like that of _Tetrapyle_ or _Pylonium_. This latter is twice as
large as the enclosed _Larnacilla_-shell. Eight long and thin, cylindrical,
radial spines, opposite in pairs in two diagonal planes. (Seems to be the
common _Tetrapyle octacantha_, enveloped by an outer spongy framework
mantle.)

_Dimensions._--Length of the whole shell (without spines) 0.22, breadth
0.16; length of the medullary shell 0.06, breadth 0.04.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

2. _Larnacostupa spinosa_, n. sp.

Cortical shell lentelliptical, about one and a half times as long as broad,
with very lax and irregular spongy framework, arising from a nearly
quadrangular lattice-shell like that of _Tetrapyle_; this latter encloses a
_Larnacilla_-shell of half its size. Whole surface covered with thin
bristle-like radial spines, of about the length of the shell.

_Dimensions._--Length of the whole shell (without spines) 0.18, breadth
0.12; length of the medullary shell 0.05, breadth 0.035.

_Habitat._--Antarctic Ocean, off Kerguelen, Station 150, surface.

3. _Larnacostupa dendrophora_, n. sp.

Cortical shell nearly spherical, scarcely longer than broad, with lax,
irregular spongy framework, arising from a lentelliptical trizonal
medullary shell (like _Larnacilla_). Whole surface covered with thin
arborescent radial spines, about half as long as the shell, each spine with
three to six irregular branches.

_Dimensions._--Length of the whole shell (without spines) 0.15, breadth
0.13; length of the medullary shell 0.07, breadth 0.04.

_Habitat._--Antarctic Ocean, Station 154, surface.

{628}Family XXVI. #PYLONIDA#, Haeckel, 1881 (Pl. 9).

_Pylonida_, Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Larcoidea# with regular, incompletely latticed cortical
shell, distinguished by two to four or more symmetrically disposed gates or
large fissures remaining between one to three latticed dimensive girdles
(perpendicular one to another). One, two, or three concentric systems of
such girdles (each system with three girdles) may be developed.

The family #Pylonida# is the most important and interesting among all the
#Larcoidea#, not only because it is much richer in different and peculiar
forms than the other families of this section, but also because it has
direct and very complex relations to all the other families of #Larcoidea#.
It is even possible that the Pylonida represent the original ancestral
group of the whole section, and that the apparently simpler group of the
Larcarida must be derived from the former by retrogressive metamorphosis.

Till the year 1881 the family Pylonida, which here now exhibits ten genera
with eighty-six species, was only represented by one single species,
accurately described and extensively illustrated by Johannes Müller in
1858, the well known and widely distributed cosmopolitan _Tetrapyle
octacantha_ (Abhandl. d. k. Akad. d. Wiss. Berlin, p. 33, Taf. iii.). A
slight modification of it was afterwards described by Ehrenberg as
_Schizomma quadrilobum_ (Abhandl. d. k. Akad. d. Wiss. Berlin, 1872, Taf.
ii. fig. 12). A more accurate description of it, with a good explanation of
its characteristic growth, was given in 1879 by Richard Hertwig in his
Organismus der Radiolarien (pp. 52-54, Taf. iv. figs. 7, 8; Taf. vi. figs.
2, 5). In my Prodromus (1881, p. 463) I constituted for a large number of
allied species, detected in the Challenger collection, the special family
Pylonida, and distinguished among it twelve different genera. However, I
think it now better to restrict the definition of the family as given in
the above definition, and to remove from it a number of genera formerly
with it united, as the genera _Triopyle_ and _Hexapyle_, appertaining to
the #Discoidea#.

The characteristic type of all true Pylonida is clearly demonstrated by
their peculiar mode of growth, the consequence of which is the imperfect
lattice-work of the fenestrated larcoid shell. This remarkable growth is
effected by the development of elliptical latticed girdles (or rings),
which enclose a quite simple, spherical, subspherical, or lentelliptical
primordial shell. The girdles lie in three different planes, perpendicular
to one another, and are of different sizes; each girdle being somewhat
larger than the foregoing and somewhat smaller than the following girdle.
Between these latticed girdles remain on the surface of the shell large
openings or "gates," which are not closed by network, and it is just the
symmetrical disposition and form of these open "gates," separated and
enclosed by the fenestrated girdles, which give to the Pylonida their
characteristic appearance.

{629}To understand clearly this peculiar constitution of the Pylonida-shell
by a system of alternating girdles, developing one after the other it is
indispensable to pay careful attention to the three different elliptical
dimensive planes, which characterise all #Larcoidea#, and to the three
different dimensive axes, which bisect those planes. The girdle which first
develops around simple primordial shell or central chamber is the
transverse girdle, lying in the equatorial plane; then comes, secondly, the
lateral girdle, lying in the lateral plane; and thirdly follows the
sagittal girdle, lying in the sagittal or median plane. The three simplest
genera of the Pylonida--_Monozonium_, _Dizonium_, _Trizonium_--represent
these three different stages, with one, two, or three girdles. These three
genera constitute the first subfamily, Haplozonaria (with one single system
of girdles); all three girdles lie in the surface of a simple
lentelliptical cortical shell.

From this first subfamily the other two subfamilies of Pylonida must be
derived, by repetition of the same characteristic process of growth. In the
Diplozonaria a second system of girdles has been developed, constituting a
second (outer) cortical shell of lentelliptical form, concentric with the
first. Also in this second system the transverse girdle is first developed,
secondly the lateral girdle, thirdly the sagittal girdle. The three genera
_Amphipyle_, _Tetrapyle_ (with _Octopyle_), and _Pylonium_ represent these
three different stages of growth.

Commonly the growth of the Pylonida stops with the completion of the second
system but sometimes the same process is once repeated and a third system
of girdles is formed, constituting a third lentelliptical shell; in this
case also the succession of the three latticed girdles is the same; firstly
the (third) transverse girdle is formed, secondly the (third) lateral
girdle, and thirdly the (third) sagittal girdle. Each of these three
girdles of the third system encloses concentrically the corresponding
girdles of the second and first system. The three corresponding genera of
this third subfamily (Triplozonaria) are _Amphipylonium_, _Tetrapylonium_,
and _Pylozonium_. But in general this highest number of girdles (nine) is
very seldom reached; commonly the growth of the Pylonida stops with five
girdles (_Tetrapyle_ and _Octopyle_). More than nine girdles I have never
observed, though there remains the possibility of the apposition of a
fourth system owing to the peculiar imperfect character of the growth
itself.

The central or primordial chamber of the shell, with which in all Pylonida
the shell-building commences, is a quite simple, very small fenestrated
shell. Commonly one sees on the surface only five to ten small pores (three
to four on the diameter). Its form seems to be sometimes spherical,
sometimes elongated, ellipsoidal or probably lentelliptical. It may be
originally a small _Cenolarcus_. This simple central chamber, the true
"medullary shell" of the small Haplozonaria, is quite different from the
medullary shell of the larger Diplozonaria, and particularly of the
well-known _Tetrapyle_. The former observers, J. Müller as well as R.
Hertwig, have described in these forms also the medullary shell as a simple
spherical or oblong body. But a careful {630}comparison of many hundred
specimens of them and of their dimensions, has convinced me that this was
an error, and that the small spherical or elliptical medullary shell of
_Tetrapyle_ and the other Diplozonaria possesses already the same complex
structure, composed of a system of three girdles, as _Trizonium_ and
_Larnacilla_. Whilst in the Haplozonaria probably the simple central
chamber only represents the medullary shell (enclosed in the central
capsule), and the first system of girdles (complete in _Trizonium_) the
external cortical shell, with the progressive growth this latter becomes
enclosed in the central capsule and so constitutes the "trizonal medullary
shell" of the Diplozonaria and Triplozonaria.

A very difficult matter is the mode of connection between the cortical and
medullary shell. In most of the Pylonida it seems that the first or
transverse girdle (in each system) is produced by the formation of two
lateral wings or chambers (one on each side of the medullary shell), so
that each wing (or half girdle) represents a short and wide, nearly
cylindrical tube, the axis of which (with free openings on both poles) is
parallel to the principal axis of the medullary shell. In this case
(probably the ordinary one) both principal faces of the medullary shell
itself (dorsal and ventral face) constitute the middle part of the first
girdle whilst its lateral parts are formed by the wings (comparable to the
lateral chambers of _Amphitholus_).

In the second case (probably a much rarer one) there is a free ring-shaped
space between the medullary shell and the first (transverse) girdle, and
both are connected by a small number of very short and small radial beams
(R. Hertwig, _loc. cit._, p. 52, line 19 to 21 from above). This mode of
connection would be the same as is common between the concentric shells of
the #Sphæroidea# and #Prunoidea#. The distinction between these two
different modes of connection is often very difficult.

The second or lateral girdle is commonly not in direct connection with the
medullary shell, or only by some scattered radial beams (mainly in the
principal axis). This lateral guide arises by prolongation of both wings of
the transverse girdle in the lateral plane, so that from both sides (right
and left) they become united on the poles of the principal axis. The minor
axis of the elliptical lateral ring (thus formed) is therefore the major
axis of the foregoing (transverse) elliptical ring; the major axes of both
are perpendicular one to another. The major axis of the lateral ring is the
principal (or longitudinal) axis of the whole body.

The third or sagittal girdle becomes developed from the second almost in
the same manner as the second from the first. On both poles of the
principal axis two latticed wings arise from the lateral girdle, growing
further in the direction of an elliptical ring, which represents the
perimeter of the sagittal plane or median plane. These wings are already
mentioned by J. Müller as "prominent roofs, protecting the gates of the
_Tetrapyle_-shell." If these roofs grow towards the equatorial plane of the
shell and become united in pairs on the poles of the sagittal axis, the
third girdle becomes {631}complete. R. Hertwig supposes that the minor axis
of this sagittal girdle is constantly at the same time the major axis of
the lateral girdle, but this is not always the case. Very often the size of
both these girdles is nearly the same, or one is not much larger than the
other. In this case the principal axis of the body is the major axis of the
second as well as of the third girdle.

The characteristic "gates" of the Pylonida, or the large wide openings in
their cortical shell, remaining between the crossed latticed girdles, are
in general roundish, sometimes nearly circular, commonly more elliptical,
kidney-shaped or semilunar, their special form varying much according to
the different form of the girdles. The narrowest part of each girdle, or
its "isthmus," in the case of the halves of the transverse girdle is
commonly at their origin from the medullary shell, in the case of the
halves of the lateral girdle at the poles of the principal axis, and in the
case of the halves of the sagittal girdle at the poles of the sagittal
axis. The number of the gates is quite constant in the different genera. If
only one girdle (the transverse) be developed, we find only two large
gates, between the two wings on the poles of the principal axis (in
_Monozonium_, _Amphipyle_, _Amphipylonium_). In all other cases there are
four gates (determining the original name "_Tetrapyle_"), as well if only
two or if all three girdles be completed. If two girdles be complete (in
_Dizonium_, _Tetrapyle_, _Tetrapylonium_) the four gates lie opposite in
pairs on the sagittal faces (two anterior and two posterior gates), and are
limited by the transverse and lateral girdles. If all three girdles be
complete (in _Trizonium_, _Pylonium_, _Pylozonium_) the four gates lie
opposite in pairs on the lateral faces (two dorsal and two ventral gates),
and are limited by the sagittal and the transverse girdles. If we turn the
shell through an angle of 90°, we have the same aspect as in the former
group. A sagittal septum sometimes becomes developed, beginning with two
polar beams, rising from the poles of the principal axis. If these polar
beams become branched and connected with the middle part of the lateral
girdle, we get a latticed vertical septum, which divides the four gates of
_Tetrapyle_ into eight gates, _Octopyle_.

The lattice-work of the Pylonida is commonly very variable and irregular,
with roundish meshes of very unequal size, therefore without value in the
determination of the species. Commonly the outside of the shell is thorny,
and often distinguished by larger radial spines, symmetrically disposed. We
can separate these into two groups; "dimensive" spines, lying in one of the
three dimensive axes (principal, transverse, or sagittal), and "diagonal"
spines, lying crossed in pairs in diagonal axes. Among these latter eight
diagonal wing-spines, which arise from the lateral edges of the four gates,
are particularly remarkable; they are not only characteristic of _Tetrapyle
octacantha_, but also of a large number of other Pylonida, and form the
starting-point for many specific forms.

The shell of the Pylonida is characterised by extraordinary variability and
great {632}inclination to individual abnormalities, formation of varieties
and transitions into other families, hence derived, as Larnacida,
Tholonida, Lithelida, &c.

The central capsule in all Pylonida, in which I could observe it, was a
true lentellipsis (or a "triaxial ellipsoid" in the geometrical sense, with
the three unequal isopolar axes of the "rhombic octahedron"). In the living
Pylonida it is commonly coloured pink or scarlet. During growth its
dimensions are probably more or less changed, and perhaps the axes
alternate. Regarding the relation of the central capsule to the skeleton,
we can distinguish two different groups, quite as in the nearly allied
Larnacida. In the Haplozonaria (as also in the Larnacillida) the central
capsule encloses only the central chamber and is enveloped by the first
system of girdles, whereas in the Diplozonaria and Triplozonaria (as in the
Larnacalpida) that "trizonal shell" becomes enclosed (as the "medullary
shell") in the central capsule, which is now enveloped by the second system
of girdles as the "cortical shell."

_Synopsis of the Genera of Pylonida._

-------------------------------------------------------------------------
I. Subfamily Haplozonaria.
One system of girdles. Medullary shell simple, spherical or
lentelliptical; cortical shell simple, with one, two, or three
girdles.
-------------------------------------------------------------------------
Cortical shell only with one latticed (transverse)
girdle, 279. _Monozonium_.

Cortical shell with two perfect girdles (transverse
and lateral), 280. _Dizonium_.

Cortical shell with three perfect girdles
(transverse, lateral, and sagittal), 281. _Trizonium_.
-------------------------------------------------------------------------
II. Subfamily Diplozonaria.
Two systems of concentric girdles. Medullary shell trizonal, with
three perfect girdles; cortical shell simple, with one, two or three
girdles.
-------------------------------------------------------------------------
Cortical shell only with one perfect (transverse)
girdle, 282. _Amphipyle_.

Cortical shell with two { Four gates simple, 283. _Tetrapyle_.
perfect girdles (the {
transverse and lateral).{ Four gates bisected by
{ a sagittal septum, 284. _Octopyle_.

Cortical shell with three perfect girdles
(transverse, lateral, and sagittal), 285. _Pylonium_.
-------------------------------------------------------------------------
III. Subfamily Triplozonaria.
Three systems of concentric girdles. Medullary shell trizonal, with
three perfect girdles, quite as the inner cortical shell; outer
cortical shell with one, two, or three girdles.
-------------------------------------------------------------------------
Outer cortical shell (third system) only with one
perfect (transverse) girdle, 286. _Amphipylonium_.

Outer cortical shell with two perfect girdles
(transverse and lateral), 287. _Tetrapylonium_.

Outer cortical shell with three perfect girdles
(transverse, lateral, and sagittal), 288. _Pylozonium_.

Subfamily 1. HAPLOZONARIA, Haeckel.

_Definition._--#Pylonida# with one single system of fenestrated girdles
(with one, two, or three girdles, lying in one lentelliptical face).

{633}Genus 279. _Monozonium_,[318] n. gen.

_Definition._--#Pylonida# with simple, spherical or subspherical, central
chamber, surrounded by one single (transverse) latticed girdle.

The genus _Monozonium_ is the most simple and primitive of all Pylonida,
and may be regarded as their common ancestral form; it may probably be
derived phylogenetically either directly from _Cenosphæra_ or from
_Lentellipsis_, either by apposition of two imperfect lateral chambers, or
by surrounding it with an equatorial latticed girdle. This transverse
girdle is composed of three parts, the central chamber and two lateral
wings, which represent two short hollow latticed tubes, the axes of which
are parallel to the principal axis. On both principal sides (on the
anterior and posterior faces) there are two large open gates as in
_Amphipyle_. If we imagine the openings of the tube-shaped lateral wings
closed by lattice-work, _Monozonium_ becomes transformed into _Tholartus_,
the most simple form of Tholonida. Probably in all Pylonida the ontogeny of
the shell begins with the formation of a _Monozonium_.

Subgenus 1. _Monozonaris_, Haeckel.

_Definition._--Shell smooth or rough, without radial spines or thorns.

1. _Monozonium primordiale_, n. sp.

Central chamber of the shell spherical, smooth, with three to four pores on
the half equator. Both wings of the girdle of the same breadth, but of
twice the length of the central chamber, with three to four longitudinal
rows of pores. No radial spines.

_Dimensions._--Diameter of the central chamber 0.02; breadth of the wings
0.02, length 0.04; pores and bars 0.003.

_Habitat._--Pacific, central area, Station 271, surface.

2. _Monozonium alatum_, n. sp. (Pl. 9, fig. 1).

Central chamber of the shell lentelliptical, smooth, one and a half times
as long as broad, with three to four pores on the half equator. Both wings
of the girdle twice as broad, and somewhat longer than the central chamber,
with five to six longitudinal rows of pores. No radial spines.

_Dimensions._--Length of the central chamber 0.03, breadth 0.02; breadth of
the wings 0.04, length 0.05; pores and bars 0.004.

_Habitat._--North Pacific, Station 241, surface.

_Monozonium hartingii_ = _Haliomma amphiaspis_, Harting, 1863 (L. N. 18, p.
15, pl. 2, fig. 43).

{634}Subgenus 2. _Monozonitis_, Haeckel.

_Definition._--Shell with radial spines or thorns, symmetrically disposed.

3. _Monozonium pleurostylum_, n. sp.

Central chamber spherical, smooth, with four to five pores on the half
equator. Both wings of the girdle of the same breadth, but somewhat longer
than the central chamber, with four to five longitudinal rows of pores. On
the poles of the lateral axis two opposite strong conical spines (one in
the middle of each wing).

_Dimensions._--Diameter of the central chamber 0.025; breadth of the wings
0.025, length 0.03; pores and bars 0.005.

_Habitat._--Pacific, central area, Station 265, surface.

4. _Monozonium amphistylum_, n. sp.

Central chamber lentelliptical, one and a half times as long as broad, with
three to four pores on the half equator. Breadth and length of each wing
somewhat greater than that of the central chamber. On the latter two strong
conical spines, opposite on the poles of the principal or longitudinal
axis.

_Dimensions._--Length of the central chamber 0.03, breadth 0.02; breadth of
the wings 0.025, length 0.04; pores and bars 0.004.

_Habitat._--South Atlantic, Station 332, surface.

5. _Monozonium staurostylum_, n. sp.

Central chamber lentelliptical, one and a third times as long as broad,
with four to five pores on the half equator. Both wings of the girdle have
the same breadth, but one and a half times the length of the central
chamber. Four conical radial spines, two opposite on the poles of the
lateral, two on the poles of the principal axis.

_Dimensions._--Length of the central chamber 0.028, breadth 0.021; breadth
of the wings 0.02, length 0.04.

_Habitat._--South Pacific, Station 295, surface.

Genus 280. _Dizonium_,[319] n. gen.

_Definition._--#Pylonida# with simple, spherical or subspherical, central
chamber, surrounded by two crossed latticed girdles, one smaller (primary)
transverse and one larger (secondary) lateral girdle.

The genus _Dizonium_ differs from the preceding _Monozonium_ in the
possession of two crossed elliptical lattice-girdles. The smaller girdle,
immediately surrounding the {635}central chamber, is the transverse girdle
(the single girdle of _Monozonium_). On the poles of its major axis (the
transverse axis) it is connected with the larger girdle, the elliptical
perimeter of which circumscribes the lateral plane. The minor axis of this
latter is the major axis of the former. Between the two crossed girdles
remain four larger openings or gates, quite as in _Tetrapyle_. But whilst
in _Tetrapyle_ the enclosed medullary shell is a trizonal
_Larnacilla_-shell, here in _Dizonium_ it is a simple spherical or
lentelliptical chamber.

Subgenus 1. _Dizonaris_, Haeckel.

_Definition._--Shell smooth or rough, without radial spines or thorns.

1. _Dizonium circulare_, n. sp.

Central chamber of the shell spherical, smooth, with four to five pores on
the half equator. Lateral girdle circular, three times as broad as the
former. Four gates semilunar, twice as broad as high. No radial spines.

_Dimensions._--Diameter of the central chamber 0.02, of the lateral girdle
0.06; height of the gates 0.02, breadth 0.04.

_Habitat._--Pacific, central area, Station 274, surface.

2. _Dizonium ellipticum_, n. sp.

Central chamber of the shell elliptical, smooth, with three to four pores
on the half equator. Lateral girdle elliptical, three times as long and
broad as the central chamber. Four gates kidney-shaped, twice as broad as
high. No radial spines.

_Dimensions._--Length of the central chamber 0.03, breadth 0.02; length of
the lateral girdle 0.09, breadth 0.06; height of the gates 0.02, breadth
0.038.

_Habitat._--South Pacific, Station 288, surface.

3. _Dizonium transversum_, n. sp.

Central chamber of the shell spherical, rough, with five to six pores on
the half equator. Lateral girdle transverse-elliptical, so that its longer
axis corresponds to the lateral axis of the transverse girdle, and equals
four times the diameter of the central chamber. Four gates kidney-shaped,
four times as broad as high. No radial spines.

_Dimensions._--Diameter of the central chamber 0.02; length of the lateral
girdle 0.05, breadth 0.08; height of the gates 0.01, breadth 0.04.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

{636}Subgenus 2. _Dizonitis_, Haeckel.

_Definition._--Shell with radial spines or thorns, symmetrically disposed.

4. _Dizonium pleuracanthum_, n. sp. (Pl. 9, fig. 2).

Central chamber of the shell nearly spherical, smooth. Lateral girdle
elliptical, one and a third times as long as broad, three times as long as
the central chamber. Four gates transverse-elliptical, one and a half times
as broad as high. Two conical spines, opposite on the poles of the lateral
axis.

_Dimensions._--Diameter of the central chamber 0.03; length of the lateral
girdle 0.1, breadth 0.075; height of the gates 0.026, breadth 0.04.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

5. _Dizonium amphacanthum_, n. sp.

Central chamber elliptical. Lateral girdle elliptical, twice as long as
broad. Four gates heart-shaped, about as high as broad. Two conical
spines, opposite on the poles of the principal axis.

_Dimensions._--Length of the central chamber 0.03, breadth 0.02; length of
the lateral girdle 0.1, breadth 0.05; height and breadth of the gates 0.03.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

6. _Dizonium stauracanthum_, n. sp. (Pl. 9, fig. 3).

Central chamber spherical. Lateral girdle elliptical, one and a third times
as long as broad. Four gates elliptical, one and a half times as broad as
high, halved by an axial beam (as in _Octopyle_). Four conical radial
spines opposite in pairs, two on the poles of the principal, two on the
poles of the transverse axis.

_Dimensions._--Diameter of the central chamber 0.025; length of the lateral
girdle 0.09, breadth 0.06; height of the gates 0.024, breadth 0.036.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

7. _Dizonium octacanthum_, n. sp.

Central chamber spherical. Lateral girdle elliptical, one and a half times
as long as broad. Four gates kidney-shaped, twice as broad as high. Eight
radial spines, opposite in pairs in two crossed diagonal planes.

_Dimensions._--Diameter of the central chamber 0.02; length of the lateral
girdle 0.075, breadth 0.05; height of the gates 0.02, breadth 0.04.

_Habitat._--Pacific, central area, Station 272, surface.

{637}Genus 281. _Trizonium_,[320] n. gen.

_Definition._--#Pylonida# with simple, spherical or subspherical, central
chamber, surrounded by three latticed girdles, one smaller (primary)
transverse, one larger (secondary) lateral, and one (tertiary) sagittal
girdle.

The genus _Trizonium_ represents the most highly developed form of the
Haplozonaria, with three complete elliptical latticed girdles, crossed at
right angles and lying in the perimeter of the three dimensive planes.
Commonly the first (transverse) girdle is the smallest, immediately
connected with the spherical or lentelliptical central chamber. The minor
axis of the second (lateral) girdle is identical with the major axis of the
first girdle. The third (sagittal) girdle is either larger than both
others, or intermediate between them. If the four gates between the girdles
become afterwards closed by lattice-work, the "trizonal shell" of
_Trizonium_ passes over into the typical _Larnacilla_, the ancestral form
of a great many #Larcoidea#.

Subgenus 1. _Trizonaris_, Haeckel.

_Definition._--Shell smooth or rough, without radial spines or thorns.

1. _Trizonium tricinctum_, n. sp. (Pl. 9, fig. 4).

Central chamber of the shell elliptical. Lateral girdle elliptical, one and
a half times as long as broad. Four gates kidney-shaped, one and a half
times as broad as high. Ten to eleven pores on the half equator, fourteen
to sixteen on the half meridian of the lentelliptical shell. No radial
spines on the surface.

_Dimensions._--Length of the central chamber 0.024, breadth 0.016; length
of the lateral girdle 0.075, breadth 0.05; height of the four gates 0.026,
breadth 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Trizonium constrictum_, n. sp.

Central chamber elliptical. Lateral girdle violin-shaped, on both sides in
the middle constricted, twice as long as broad. Four gates
transverse-elliptical, twice as broad as high. Eight to nine pores on the
half equator, twelve to thirteen on the half meridian. No radial spines.

_Dimensions._--Length of the central chamber 0.03, breadth 0.02; length of
the lateral girdle 0.09, breadth (in the middle) 0.045; height of the gates
0.022, breadth 0.044.

_Habitat._--Pacific, central area, Station 263, surface.

{638}3. _Trizonium hexagonium_, n. sp.

Central chamber spherical. Lateral girdle hexagonal, with parallel sides,
twice as long as broad. Four gates triangular, twice as broad as high.
Eleven to twelve pores on the half equator, sixteen to eighteen on the half
meridian. No radial spines.

_Dimensions._--Diameter of the central chamber 0.025; length of the lateral
girdle 0.08, breadth 0.04; height of the gates 0.018, breadth 0.036.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

4. _Trizonium octogonium_, n. sp.

Central chamber spherical. Lateral girdle octagonal; two lateral sides of
the octagon twice as long as the two polar sides and the four diagonal
sides. Four gates hexagonal, one and a half times as broad as high. Nine to
ten pores on the half equator, twelve to fourteen on the half meridian. No
radial spines.

_Dimensions._--Diameter of the central chamber 0.02; length of the lateral
girdle 0.01, breadth 0.07; height of the gates 0.04, breadth 0.06.

_Habitat._--Indian Ocean, surface; Ceylon, Haeckel.

Subgenus 2. _Trizonitis_, Haeckel.

_Definition._--Shell with radial spines or thorns, symmetrically disposed.

5. _Trizonium pleurobelonium_, n. sp.

Central chamber spherical. Lateral girdle elliptical, one and a third times
as long as broad. Four gates nearly circular. Ten to eleven pores on the
half equator, fifteen to sixteen on the half meridian. Two opposite conical
spines on the poles of the lateral axis.

_Dimensions._--Diameter of the central chamber 0.02; length of the lateral
girdle 0.08, breadth 0.06; height and breadth of the gates 0.025.

_Habitat._--South Atlantic, off Buenos Ayres, Station 323, depth 1900
fathoms.

6. _Trizonium amphibelonium_, n. sp.

? _Echinosphæra datura_, R. Hertwig (partim), 1879, Organismus d.
Radiol., p. 54, Taf. iv. figs. 8, 8_a_.

Central chamber lentelliptical. Lateral girdle lanceolate, nearly one and a
half times as long as broad. Four gates subtriangular. Eight to nine pores
on the half equator, ten to twelve on the half meridian. Two opposite thin
and long spines on the poles of the principal axis, numerous shorter radial
spines on the whole surface. Possibly this species is identical with one of
the three different #Larcoidea# which R. Hertwig has described as
_Echinosphæra datura_. His fig. 8, Taf. iv., would be the aspect from the
sagittal girdle.

{639}_Dimensions._--Length of the central chamber 0.04, breadth 0.025;
length of the lateral girdle 0.11, breadth 0.08; height of the gates 0.03,
breadth 0.04.

_Habitat._--Mediterranean (Genoa), Haeckel, (Messina), R. Hertwig, surface;
Tropical Atlantic, Station 348, surface.

7. _Trizonium staurobelonium_, n. sp.

Central chamber spherical, with two opposite polar beams. Lateral girdle
elliptical, one and a half times as long as broad. Four gates
transverse-elliptical, half as high as broad. Ten to eleven pores on the
half equator, fourteen to fifteen on the half meridian. Four conical radial
spines, two on the poles of the lateral, two on the poles of the principal
axis.

_Dimensions._--Diameter of the central chamber 0.02; length of the lateral
girdle 0.075, breadth 0.05; height of the gates 0.014, breadth 0.028.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

8. _Trizonium hexabelonium_, n. sp.

Central chamber elliptical. Lateral girdle hexagonal, one and a half times
as long as broad. Four gates rhombic, two-thirds as high as broad. Eight to
nine pores on the half equator, twelve to thirteen on the half meridian.
Six radial spines in the lateral plane, opposite in pairs on the six
corners of the lateral girdle (two principal and four diagonal).

_Dimensions._--Length of the central chamber 0.024, breadth 0.018; length
of the lateral girdle 0.06, breadth 0.04; height of the gates 0.02, breadth
0.03.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

9. _Trizonium octobelonium_, n. sp.

Central chamber elliptical. Lateral girdle hexagonal, one and one-third
times as long as broad. Four gates kidney-shaped or nearly rhombic, twice
as broad as high. Eleven to twelve pores on the half equator, sixteen to
seventeen on the half meridian. Eight radial spines, opposite in pairs in
two crossed diagonal planes. (An intermediate stage between _Dizonium
octacanthum_ and _Amphipyle octostyle_.)

_Dimensions._--Length of the central chamber 0.03, breadth 0.024; length of
the lateral girdle 0.09, breadth 0.07; height of the gates 0.025, breadth
0.05.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

10. _Trizonium decabelonium_, n. sp.

Central chamber elliptical. Lateral girdle hexagonal, one and a half times
as long as broad. Four gates rhombic, twice as broad as high. Twelve to
thirteen pores on the half equator, seventeen to eighteen on the half
meridian. Ten radial spines, two opposite on the two poles of the principal
axis, eight others opposite in pairs in two crossed diagonal planes.

{640}_Dimensions._--Length of the central chamber 0.04, breadth 0.03;
length of the lateral girdle 0.12, breadth 0.08; height of the gates 0.03,
breadth 0.06.

_Habitat._--North Atlantic, Station 353, surface.

11. _Trizonium dodecabelos_, n. sp.

Central chamber spherical. Lateral girdle octagonal, nearly twice an long
as broad. Four gates hexagonal, one and a half times as broad as high. Nine
to ten pores on the half equator, fifteen to sixteen on the half meridian.
Twelve radial spines, four in the lateral plane, in pairs on both sides of
the poles of the principal axis, eight others opposite in pairs in two
crossed diagonal axes.

_Dimensions._--Diameter of the central chamber 0.025; length of the lateral
girdle 0.1, breadth 0.06; height of the gates 0.033, breadth 0.05.

_Habitat._--Pacific, central area, Station 272, surface.

Subfamily 2. DIPLOZONARIA, Haeckel.

_Definition._--#Pylonida# with two concentric systems of fenestrated
girdles, lying in two concentric lentelliptical faces (every one system
with one to three girdles, lying in one lentelliptical face).

Genus 282. _Amphipyle_,[321] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell,
surrounded by one single (transverse) latticed cortical girdle.

The genus _Amphipyle_ opens the large series of Diplozonaria, comprising
all Pylonida, the shell of which is composed of two concentric systems of
latticed girdles; the first system constituting the characteristic
"trizonal medullary shell" or "_Larnacilla_-shell;" the second system
composed of one to three girdles of the second order. The first system lies
inside, the second outside the central capsule. In _Amphipyle_ only the
first (transverse) girdle of the second system becomes developed, and
therefore on both poles of the principal axis are two large open gates.
_Amphipyle_ repeats the two-winged form of _Monozonium_; but whilst the
medullary shell in this latter is a simple central chamber, it is here a
trizonal _Larnacilla_-shell.

Subgenus 1. _Amphipylissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without large,
symmetrically disposed spines.

{641}1. _Amphipyle aceros_, n. sp.

Cortical shell quite smooth, without any spines or thorns. Both lateral
wings (or opposite half girdles) semilunar, with convex lateral crest,
three times as long as broad; twice as broad as the trizonal lentelliptical
medullary shell.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.03; length of
each lateral wing (or principal dimension of the latticed girdle) 0.18,
breadth of it (or transverse dimension of each girdle-tube) 0.06.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

2. _Amphipyle stenoptera_, n. sp.

Cortical shell smooth, without spines. Both lateral wings of the same
breadth as the lentelliptical medullary shell, but five times as long as
broad, prolonged on both ends into cylindrical latticed tubes, with
rectilinear lateral crest.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.04; length of
each cylindrical lateral wing 0.2, breadth 0.04.

_Habitat._--South Atlantic, off Ascension Island, Station 343, surface.

3. _Amphipyle platyptera_, n. sp.

Cortical shell thorny, but without larger spines. Both lateral wings nearly
triangular, very broad and short, four times as broad as the medullary
shell between them, and only twice as long as broad, with rectilinear
lateral crest.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.25; length of
each lateral wing 0.18, breadth 0.09.

_Habitat._--Indian Ocean, Ceylon, Haeckel, surface.

Subgenus 2. _Amphipylura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed
spines.

4. _Amphipyle amphiceros_, n. sp.

Cortical shell with two conical spines on the poles of the longitudinal
axis (of the lentelliptical medullary shell), without other large spines,
but with small thorns on the surface; two to three longitudinal rows of
irregular, roundish pores on each half wing of the cortical girdle.

_Dimensions._--Length of the medullary shell (or principal axis) 0.06,
breadth (or transverse axis) 0.04; length of each lateral wing 0.2, breadth
of it 0.05.

_Habitat._--Western Tropical Pacific, Station 222, surface.

{642}5. _Amphipyle tetraceros_, n. sp. (Pl. 9, fig. 5).

Cortical shell with smooth surface, and with four strong conical spines in
the lateral plane, opposite in pairs on the ends of the concave lateral
crests (one pair on the end-points of the lateral line of each wing); four
to five longitudinal rows of irregular, roundish pores on each half wing of
the cortical girdle. Surface of the medullary shell spiny.

_Dimensions._--Length of the medullary shell 0.08, breadth of the same
0.05; length of each lateral wing 0.16, breadth 0.04.

_Habitat._--Pacific, central area, Station 271, surface.

6. _Amphipyle stauroceros_, n. sp.

Cortical shell with four strong conical spines, two on the poles of the
longitudinal axis, two opposite on the poles of the transverse axis
(perpendicular to the former). Three to four longitudinal rows of
irregular, roundish pores on each half wing of the cortical girdle, about
six pores in the longest row. Lateral crest convex.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.03; length of
each lateral wing 0.15, breadth 0.08.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

7. _Amphipyle amphiptera_, n. sp. (Pl. 9, fig. 7).

Cortical shell with six strong conical spines, lying in the lateral plane
in three parallel longitudinal lines; two on the poles of the longitudinal
axis, two others on the poles of the wing axes, which are parallel to the
former. Two to three longitudinal rows of irregular, roundish pores on each
half wing of the cortical girdle, about eight pores in the longest row.
Lateral crest convex, thorny.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.045; length of
each lateral wing 0.09, breadth 0.035.

_Habitat._--Western Tropical Pacific, Station 224, surface.

8. _Amphipyle hexaceros_, n. sp.

Cortical shell thorny, with six strong conical spines lying in the lateral
plane; two on the poles of the lateral axis (in the central point of each
wing), two on the end-points of the triangular crest of each wing (on both
poles of the longitudinal wing-axis).

_Dimensions._--Length of the medullary shell 0.06, breadth 0.04; length of
each lateral wing 0.13, breadth 0.07.

_Habitat._--South Pacific, Station 291, surface.

{643}9. _Amphipyle octoceros_, n. sp.

Cortical shell thorny, with eight long radial spines, lying in two crossed
diagonal planes, opposite in pairs. These eight horns are the
prolongations of the anterior and posterior edges of each wing, on both
sides of its lateral plane; they are of great importance, as appearing (by
heredity) in many other Pylonida (_e.g._, in _Tetrapyle octacantha_).
Lateral crest of each wing without spines, slightly convex.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.05; length of
each lateral wing 0.15, breadth 0.06.

_Habitat._--Pacific, central area, Stations 270 to 274, surface.

10. _Amphipyle cladoceros_, n. sp.

Cortical shell spiny, with eight strong ramified radial spines, lying in
the same two crossed diagonal planes as in the preceding species. It
differs from this in the ramification of the eight horns, which bear six to
nine irregular lateral branches, partly simple, partly bifurcate. Lateral
crest of each wing spiny, convex.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.045; length of
each lateral wing (without spines) 0.2, breadth 0.09.

_Habitat._--Indian Ocean, surface, Madagascar, Rabbe.

11. _Amphipyle decaceros_, n. sp.

Cortical shell spiny, with ten strong conical spines; eight lying in two
crossed diagonal planes, in the same disposition (opposite in pairs) as in
both preceding species; two others opposite on the poles of the principal
axis, arising from both poles of the medullary shell (columella-beams).
Spiny crest of the wings convex, semilunar.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of
each lateral wing 0.12, breadth 0.04.

_Habitat._--North Atlantic, Station 354, surface.

12. _Amphipyle dodecaceros_, n. sp.

Cortical shell thorny, with twelve large cylindrical spines; eight lying in
two crossed diagonal planes, in the same disposition (opposite in pairs) as
in the three preceding species; four others crossed at right angles in the
lateral plane, two opposite on the poles of the principal, two on the poles
of the lateral axis (the latter shorter). Both lateral wings nearly
rhombic.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.04; length of
the lateral wings 0.15, breadth 0.05.

_Habitat._--South Atlantic, coast of Brazil, Rabbe, surface.

{644}13. _Amphipyle callizona_, n. sp. (Pl. 9, fig. 6).

Cortical shell smooth, with sixteen strong and short conical spines, lying
opposite in pairs in two crossed diagonal planes; each lateral wing
four-sided prismatic, its lateral face concave, both ends truncated, and
each end provided with four divergent spines. Length of the wings somewhat
greater, but breadth smaller, than that of the medullary shell.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.05; length of
the lateral wings 0.09, breadth 0.03.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 283. _Tetrapyle_,[322] J. Müller, 1858, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 154.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell,
surrounded by two crossed latticed cortical girdles, one smaller (primary)
transverse, and one larger (secondary) lateral girdle. Four gates between
the two cortical girdles simple, without a sagittal septum.

The genus _Tetrapyle_, till 1881 the only known genus of the whole family,
was founded by Johannes Müller in 1858, and clearly illustrated by the
Mediterranean (and common cosmopolitan) _Tetrapyle octacantha_, hitherto
the best known type of this family. Afterwards (1860, Monatsber. d. k.
preuss. Akad. d. Wiss. Berlin, p. 832) Ehrenberg founded the genus
_Schizomma_ for a nearly allied form, which exhibits only slight specific
differences (compare my Monograph, 1862, p. 434). Some good remarks on the
structure of this typical genus and its relations to other Pylonida are to
be found in Richard Hertwig's Organismus, &c., 1879, p. 52, but the true
trizonal structure of the medullary shell in this genus was not recognised
by him, so that his description agrees more with _Dizonium_. We confine
here the genus _Tetrapyle_ to those Pylonida for which _Tetrapyle
octacantha_ of J. Müller remains the determining type; the cortical shell
is composed only of two perfect lattice-girdles (the transverse and
lateral), between which four wide gates remain open. This structure is
similar to that of _Dizonium_; but whilst here the medullary shell is a
simple central chamber, in _Tetrapyle_ it is a complete trizonal or
_Larnacilla_-shell.

Subgenus 1. _Tetrapylissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without large,
symmetrically disposed spines.

{645}1. _Tetrapyle circularis_, n. sp. (Pl. 9, fig. 8).

Cortical shell rough, without radial spines. Lateral girdle circular,
therefore the longitudinal axis is equal to the transverse. Four gates
kidney-shaped, twice as broad as high. In each half wing of the transverse
girdle six to seven longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.15
long and broad; gates 0.04 high, 0.08 broad.

_Habitat._--Pacific, central area, Station 263, surface.

2. _Tetrapyle circopyle_, n. sp.

Cortical shell smooth, without radial spines. Lateral girdle elliptical,
one and a half times as long as broad. Four gates nearly circular. On each
half wing of the transverse girdle four to five rows of subregular,
circular pores. (Similar to _Tetrapyle pleuracantha_, Pl. 9, fig. 9, but
without lateral spines.)

_Dimensions._--Medullary shell 0.04 long, 0.03 broad; cortical shell 0.16
long, 0.11 broad; gates 0.05 high and broad.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Tetrapyle quadriloba_, Haeckel.

_Tetrapyle quadriloba_, Haeckel, 1862, Monogr. d. Radiol., p. 436.

_Schizomma quadrilobum_, Ehrenberg, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 815; Abhandl. d. k. Akad. d. Wiss. Berlin, 1872, Taf.
x. figs. 12-14.

Cortical shell thorny, without regular, radial spines. Lateral girdle
elliptical, one and a half times as long as broad. Four gates nearly
circular. On each half wing of the transverse girdle three to four rows of
irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.16
long, 0.1 broad; diameter of the gates 0.05.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian Ocean, Pacific,
surface.

4. _Tetrapyle nephropyle_, n. sp.

Cortical shell thorny, without radial spines. Lateral girdle elliptical,
one and a third times as long as broad. Four gates kidney-shaped, nearly
twice as broad as high, with two prominent polar spines on the poles of the
longitudinal axis of the medullary shell. On each half wing of the
transverse girdle three to four longitudinal rows of large roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.24
long, 0.18 broad; gates 0.05 high, 0.08 broad.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{646}5. _Tetrapyle cardiopyle_, n. sp.

Cortical shell rough, without radial spines. Lateral girdle with hexagonal
contour, nearly cylindrical in the middle part, conical at both ends, twice
as long as broad. Four gates heart-shaped, about as high as broad. On each
half wing of the transverse girdle three to four longitudinal rows of
subregular, polygonal pores.

_Dimensions._--Medullary shell 0.04 long, 0.02 broad; cortical shell 0.14
long, 0.07 broad; gates 0.05 high and broad.

_Habitat._--Eastern Tropical Atlantic, Station 348, depth (2450) fathoms.

Subgenus 2. _Tetrapylura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed
spines.

6. _Tetrapyle fusiformis_, n. sp.

Cortical shell rough, with two strong conical spines on the poles of the
longitudinal axis. Lateral girdle spindle-shaped, one and a half times as
long as broad. Four gates kidney-shaped, twice as broad as high. On each
half wing of the transverse girdle four to five longitudinal rows of
irregular, roundish pores.

_Dimensions._--Medullary shell 0.07 long, 0.05 broad; cortical shell 0.18
long, 0.12 broad; gates 0.04 high, 0.08 broad.

_Habitat._--Western Tropical Atlantic, Mexican Gulf Stream, depth 1500
fathoms, Schaffner.

7. _Tetrapyle pleuracantha_, n. sp. (Pl. 9, fig. 9).

Cortical shell rough, with two strong conical spines on the poles of the
transverse axis. Lateral girdle elliptical, one and half times as long as
broad. Four gates subcircular or nearly hexagonal, about as high as broad.
On each half wing of the transverse girdle three to four longitudinal rows
of very irregular pores.

_Dimensions._--Medullary shell 0.045 long, 0.035 broad; cortical shell 0.16
long, 0.11 broad; gates 0.045 high and broad.

_Habitat._--Western Tropical Pacific, off New Guinea, Stations 222 to 224,
depths 1850 to 2450 fathoms.

8. _Tetrapyle transversa_, n. sp.

Cortical shell thorny, with two strong conical lateral spines on the poles
of the transverse axis. Lateral girdle elliptical, only four-fifths as long
as broad. Four gates elliptical, twice as broad as high, with two prominent
spines on the poles of the longitudinal axis of the medullary shell. On
each half wing of the transverse girdle five to six longitudinal rows of
subregular, roundish pores. (Similar to _Octopyle transversa_, but without
sagittal septum and with stronger lateral spines.)

{647}_Dimensions._--Medullary shell 0.05 long, 0.035 broad; cortical shell
0.12 long, 0.15 broad; gates 0.03 high, 0.06 broad.

_Habitat._--Southern Pacific, Station 290, surface.

9. _Tetrapyle cruciata_, n. sp.

Cortical shell smooth, with four strong radial spines, two on the poles of
the longitudinal axis, two on the poles of the transverse axis. Lateral
girdle nearly circular, quite as long as broad. Four gates kidney-shaped,
twice as broad as high. On each half wing of the transverse girdle five to
six rows of subregular, circular pores.

_Dimensions._--Medullary shell 0.06 long, 0.05 broad; cortical shell 0.16
long and broad; gates 0.05 high, 0.09 broad.

_Habitat._--South Atlantic, Station 330, surface.

10. _Tetrapyle staurophora_, n. sp.

Cortical shell thorny, with four strong radial spines, two on the poles of
the longitudinal axis, two on the poles of the transverse axis. Lateral
girdle elliptical, one and a half times as long as broad. Four gates nearly
hexagonal, quite as high as broad. On each half wing of the transverse
girdle three to four longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.18
long, 0.12 broad; gates 0.08 high and broad.

_Habitat._--Western Tropical Atlantic, coast of Brazil, Rabbe, surface.

11. _Tetrapyle quadricornis_, n. sp.

Cortical shell thorny, with four strong horn-like curved spines, lying in
the lateral plane and converging in pairs towards the poles of the
transverse axis. Lateral girdle elliptical, one and a third times as long
as broad. Four gates kidney-shaped, twice as broad as high. On each half
wing of the transverse girdle four to five rows of irregular, roundish
pores. (May be only the young form of _Pylonium quadricorne_, Pl. 9, fig.
14.)

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.16
long, 0.12 broad; gates 0.03 high, 0.07 broad.

_Habitat._--Pacific, central area, Stations 270 to 274, surface.

12. _Tetrapyle tetracantha_, n. sp.

Cortical shell thorny, with four strong radial spines lying in the lateral
plane and in pairs in its two crossed diagonal axes. Lateral girdle
elliptical, constricted in the middle, nearly twice as long as broad. Four
gates kidney-shaped, one and a half times as broad as high. On each half
wing of the transverse girdle three to four longitudinal rows of large,
irregular, roundish pores. {648}(Similar to _Tetrapylonium quadrangulare_,
Pl. 49, fig. 15, but without a sagittal girdle and with stronger lateral
spines.)

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.18
long, 0.1 broad; gates 0.007 high, 0.01 broad.

_Habitat._--North Atlantic, Canary Islands, Station 353, surface.

13. _Tetrapyle quadrigata_, n. sp.

Cortical shell thorny, with four brush-like groups of radial spines on the
four diagonal corners of the lateral plane, opposite in pairs in diagonal
axes. Lateral girdle nearly square, somewhat broader than long. Four gates
kidney-shaped, twice as broad as high. On each half wing of the transverse
girdle five to six longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.14
long, 0.15 broad; gates 0.03 high, 0.07 broad.

_Habitat._--Pacific, central area, Station 270, surface.

14. _Tetrapyle octacantha_, J. Müller.

_Tetrapyle octacantha_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 33, Taf. ii. figs. 1-6.

_Tetrapyle octacantha_, Haeckel, 1862, Monogr. d. Radiol., p. 435.

_Tetrapyle octacantha_, R. Hertwig, 1879, Organismus d. Radiol., p. 52,
Taf. iv. fig. 7, Taf. vi. figs. 2, 5, 5_a._

Cortical shell thorny, with eight long and thin, cylindrical radial spines,
lying in two crossed diagonal planes, opposite in pairs. These eight
characteristic diagonal spines (or "angular spines") are the same as in
_Amphipyle octoceros_ and many other Pylonida, and arise as prolongations
of the proximal edges of the four gates (or of the lateral wings on the
eight points, where they are intersected by the edges of the lateral ring).
This cosmopolitan, widely distributed and very variable species was a long
time the only known species of all the Pylonida, and very accurately first
described (1858) by Johannes Müller, afterwards (1879) by R. Hertwig. But
in the descriptions of these authors also some different species (such as
the following) may be confounded with the true typical _Tetrapyle
octacantha_. The four gates of this species are transverse-elliptical or
almost triangular, nearly twice as broad as high.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of
the cortical shell 0.18, breadth 0.13; height of the gates 0.05, breadth
0.08.

_Habitat._--Cosmopolitan; very common in all warmer seas, Mediterranean,
Atlantic, Indian, Pacific, surface.

15. _Tetrapyle cladacantha_, n. sp.

Cortical shell thorny, with eight strong ramified radial spines, lying
opposite in pairs in two crossed diagonal planes, as in the preceding
species. It differs from this by the peculiar ramification of the eight
spines, which bear two to eight simple or furcate lateral branches arising
under {649}right angles (already commencing in a transitional variety of
_Tetrapyle octacantha_, J. Müller, _loc. cit._, Taf. ii. figs. 5, 6). Four
gates transverse-elliptical, one and a half times as broad as high.

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.16
long, 0.13 broad; gates 0.05 high, 0.08 broad.

_Habitat._--Mediterranean (Corfu), Haeckel, surface.

16. _Tetrapyle pluteus_, Haeckel.

_Tetrapyle octacantha_, var., J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, Taf. iii. figs. 7-12.

Cortical shell thorny, with eight strong radial spines, lying opposite in
pairs, as in the two foregoing species. On the distal edge each of the four
triangular gates is protected by a prominent roof of lattice-work (or
"pluteus"), which arises from the distal part of the lateral girdle and
connects the two angular diagonal spines of each gate. Therefore the shell
assumes the characteristic form very well represented by J. Müller in his
fig. 11 (seen from the lateral side) and fig. 7 (seen from the pole of the
principal axis).

_Dimensions._--Medullary shell 0.04 long, 0.03 broad; cortical shell 0.15
long, 0.11 broad; gates 0.045 high, 0.07 broad.

_Habitat._--Mediterranean, Atlantic, Stations 348 to 353, &c., surface.

17. _Tetrapyle turrita_, n. sp. (Pl. 9, fig. 10).

Cortical shell smooth, but with ten strong conical spines, two on the poles
of the principal axis, eight others symmetrically distributed on both sides
of the distal edge of the four roundish gates, and directed towards the
transverse axis. Lateral girdle lanceolate, twice as long as broad.

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.2
long, 0.1 broad; gates 0.05 high, 0.07 broad.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

18. _Tetrapyle dodecaceros_, n. sp.

Cortical shell thorny, with twelve stronger conical spines, eight diagonal
spines in two crossed planes (as in _Tetrapyle octacantha_), and four
others in the lateral plane, in pairs on both sides of a deep sagittal
constriction of the quadrangular lateral girdle. Four gates kidney-shaped,
twice as broad as high. On each half wing of the transverse girdle five to
six longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.18
long, 0.14 broad; gates 0.04 high, 0.08 broad.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{650}Genus 284. _Octopyle_,[323] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell,
surrounded by two crossed, latticed, cortical girdles; one smaller
(primary) transverse, and one larger (secondary) lateral girdle. Four gates
between the two cortical girdles divided by a sagittal septum into eight
gates.

The genus _Octopyle_ comprises those Pylonida which are distinguished from
the nearly allied _Tetrapyle_ by the development of a sagittal septum,
dividing the four gates of the latter into eight separate gates. The septum
begins with the formation of two axial rods or columellæ, which afterwards
become branched; the branches communicating one with another and with the
middle parts of the lateral girdle (on the poles of the principal axis),
there is formed a latticed septum in the sagittal plane, which separates
more or less incompletely the right and left halves of the shell. The four
gates of _Tetrapyle_ become halved by this septum, and their number
doubled.

Subgenus 1. _Octopylissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without larger
symmetrically disposed spines.

1. _Octopyle ovulina_, n. sp.

Cortical shell lentelliptical, smooth, without thorns. Lateral girdle
elliptical, one and a third times as long as broad. Transverse girdle
narrow, with two pores on the isthmus (or on the narrowest part of each
quadrant). Sagittal septum as long as the elliptical medullary shell. Eight
gates egg-shaped.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.035; length of
the cortical shell 0.16, breadth 0.12.

_Habitat._--Pacific, central area, Station 274, surface.

2. _Octopyle quadrata_, n. sp.

Cortical shell thorny, nearly square, of equal length and breadth. Lateral
girdle quadrangular, with rounded edges, of equal length and breadth.
Transverse girdle narrow, with three pores on the isthmus. Sagittal septum
of the same length as the subspherical medullary shell. Eight gates nearly
circular.

_Dimensions._--Diameter of the medullary shell 0.04, of the cortical shell
0.14.

_Habitat._--Northern Pacific, Station 244, depth 2900 fathoms.

{651}3. _Octopyle subglobosa_, n. sp.

Cortical shell nearly spherical, thorny, of equal length and breadth.
Lateral girdle broad, nearly circular. Transverse girdle broad, with four
pores on the isthmus. Sagittal septum shorter than the subspherical
medullary shell. Eight gates egg-shaped.

_Dimensions._--Diameter of the medullary shell 0.05, of the cortical shell
0.13.

_Habitat._--Southern Pacific, Station 300, depth 1375 fathoms.

4. _Octopyle transversaria_, n. sp.

Cortical shell thorny, transverse-elliptical. Lateral girdle very broad,
one and a third times as broad as long. Transverse girdle also very broad,
with five pores on the isthmus. Sagittal septum shorter than the elliptical
medullary shell. Eight gates small, roundish.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.04; length of
the cortical shell 0.11, breadth 0.15.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

Subgenus 2. _Octopylura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed
spines.

5. _Octopyle amphistyle_, n. sp.

Cortical shell smooth, nearly spindle-shaped, about twice as long as broad.
Transverse girdle broad, with four large pores on the isthmus. Lateral
girdle lanceolate. Sagittal septa about as long as the medullary shell,
prolonged at both poles of the principal axis into two opposite, strong,
angular spines. Eight gates obliquely quadrangular.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.05; length of
the cortical shell 0.2, breadth 0.12.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

6. _Octopyle staurostyle_, n. sp.

Cortical shell nearly rhombic, smooth, one and a half times as long as
broad, with four strong, conical, radial spines, two on the poles of the
principal axis (as prolongations of the sagittal septa), two others on the
poles of the transverse axis. Transverse girdle narrow, with two pores on
the isthmus. Sagittal septa longer than the medullary shell. Eight gates
triangular.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of
the cortical shell 0.18, breadth 0.12.

_Habitat._--Tropical Atlantic, Station 347, depth 2250 fathoms.

{652}7. _Octopyle tetrastyle_, n. sp.

Cortical shell lentelliptical, with rough surface and four angular radial
spines in the lateral plane, opposite in pairs in two crossed diagonals.
Lateral girdle nearly rectangular (each angle with one spine), one and a
third times as long as broad. Sagittal septum longer than the hexagonal
medullary shell. Eight gates egg-shaped.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.03; length of
the cortical shell 0.16, breadth 0.12.

_Habitat._--North Atlantic, Station 354, surface.

8. _Octopyle tetraptera_, n. sp.

Cortical shell quadrangular, with thorny surface and four latticed
wing-like prolongations on the corners of the quadrangle, opposite in pairs
in two crossed diagonals, each wing supported by a strong radial spine.
Lateral girdle quadrangular, one and a half times as long as broad. On the
isthmus of the transverse girdle four pores. Sagittal septum longer than
the hexagonal medullary shell. Eight gates triangular.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.04; length of
the cortical shell 0.24, breadth 0.16.

_Habitat._--Pacific, central area, Station 271, surface.

9. _Octopyle stenozona_, n. sp. (Pl. 9, fig. 11).

Cortical shell quadrangular, with thorny surface, and four conical spines
in the lateral plane, on the points, where both cortical girdles are
crossed. Lateral girdle very narrow, nearly square, with rounded corners,
of equal length and breadth. On the isthmus of the narrow transverse girdle
only one pore. Sagittal septum of the same length as the subspherical
medullary shell. Eight gates irregular roundish or nearly circular.

_Dimensions._--Diameter of the medullary shell 0.04, of the cortical shell
0.14.

_Habitat._--Tropical Atlantic, Station 338, depth 1990 fathoms.

10. _Octopyle euryzona_, n. sp.

Cortical shell nearly quadrangular, very thorny, with four bunches of
stronger radial spines on the four corners. Lateral girdle broad, with
rounded edges of the quadrangle, little longer than broad. On the isthmus
of the broad transverse girdle four pores. Sagittal septum of the same
length as the elliptical medullary shell, with many radial beams. Eight
gates nearly square.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.035; length of
the cortical shell 0.16, breadth 0.14.

_Habitat._--Southern Atlantic, Station 332, depth 2200 fathoms.

{653}11. _Octopyle hexastyle_, n. sp.

Cortical shell nearly rectangular, thorny, with six strong radial spines in
the lateral plane, two opposite on the poles of the principal axis
(prolongations of the principal beam at the sagittal septum), four others
on the truncate corners of the quadrangular lateral girdle, which is one
and a half times as long as broad. On the isthmus of the broad transverse
girdle three pores. Sagittal septum longer than the hexagonal medullary
shell. Eight egg-shaped gates.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.04; length of
the cortical shell 0.18, breadth 0.12.

_Habitat._--Northern Atlantic, surface; Canary Islands, Station 9, depth
3150 fathoms.

12. _Octopyle sexangulata_, n. sp. (Pl. 9, fig. 12).

Cortical shell thorny, hexagonal, with six strong conical radial spines on
the six corners of the lateral girdle, lying in the lateral plane, two
opposite on the poles of the transverse axis, four others in pairs on both
sides of the annular sagittal constriction, which is only twice as long as
the elliptical medullary shell. Transverse girdle broad, with five pores on
the isthmus. Eight gates roundish, smaller than the medullary shell.

_Dimensions._--Length of the medullary shell 0.045, breadth 0.035; length
of the cortical shell 0.11, breadth 0.13.

_Habitat._--Pacific, central area, Station 270, surface.

13. _Octopyle octostyle_, n. sp.

Cortical shell elliptical, thorny, with eight long cylindrical radial
spines, lying in two crossed diagonal planes, opposite in pairs, quite as
in the nearly related _Tetrapyle octacantha_. It differs from this species
by the perfect, latticed, vertical sagittal septum, which divides every
gate into two triangular halves. Lateral girdle elliptical, one and a third
times as long as broad. Transverse girdle on the isthmus with three pores.
Septum longer than the elliptical medullary shell.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.045; length of
the cortical shell 0.16, breadth 0.12.

_Habitat._--South Pacific, Station 288, surface.

14. _Octopyle obtecta_, n. sp.

Cortical shell quadrangular, thorny, with eight strong diagonal spines,
lying in two crossed planes, and arising from the proximal edges of the
gates in the same way as in the foregoing species. It differs from this by
four prominent latticed roofs, which arise from the distal edge of the
gates and connect the diagonal spines of one gate. _Octopyle obtecta_ bears
therefore the same relation to _Octopyle octostyle_ that _Tetrapyle
pluteus_ does to _Tetrapyle octacantha_.

_Dimensions._--Length of the medullary shell 0.055, breadth 0.04; length of
the cortical shell 0.16, breadth 0.12.

_Habitat._--Indian Ocean, surface, Madagascar, Rabbe.

{654}15. _Octopyle decastyle_, n. sp. (Pl. 9, fig. 13).

Cortical shell thorny, nearly quadrangular, with ten stronger radial
spines, two opposite on the poles of the principal axis (as prolongations
of the axial beams), eight others opposite in pairs in two diagonal planes
(as in the foregoing species). Lateral girdle nearly square, with slightly
convex lateral faces, slightly concave principal faces. Transverse girdle
broad, with four pores on the isthmus. Sagittal septum of the same length
as the hexagonal medullary shell. Eight gates elliptical or irregular
roundish.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.032; diameter
of the cortical shell 0.15.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

Genus 285. _Pylonium_,[324] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell
surrounded by three crossed latticed cortical girdles, one smaller
(primary) transverse, one larger (secondary) lateral, and one (tertiary)
sagittal girdle.

The genus _Pylonium_ is the most highly developed among the Diplozonaria,
with six complete latticed girdles, three on the medullary shell, three on
the cortical shell, the latter corresponding to the former. In each of the
two systems the transverse girdle is smaller than the lateral; the sagittal
girdle may be smaller or larger than the lateral girdle. _Pylonium_ repeats
the typical form of _Trizonium_; but whilst in this latter the medullary
centre of the shell is a simple spherical chamber, here in _Pylonium_ it is
a true trizonal or _Larnacilla_-shaped medullary shell.

Subgenus 1. _Pylonissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without large,
symmetrically disposed spines.

1. _Pylonium circozonium_, n. sp.

Cortical shell thorny, without radial spines; all its three girdles nearly
of the same size, subcircular, very narrow, only with one to two rows of
pores. Four gates subcircular.

_Dimensions._--Principal axis 0.12, transverse axis 0.11; diameter of the
subspherical medullary shell 0.05.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{655}2. _Pylonium hexazonium_, n. sp.

Cortical shell smooth, without symmetrical radial spines. Transverse girdle
elliptical, one and a half times as long as broad. Lateral and sagittal
girdles nearly circular. Four gates egg-shaped.

_Dimensions._--Principal axis 0.16, transverse axis 0.15; diameter of the
subspherical medullary shell 0.04.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

3. _Pylonium nephropylium_, n. sp.

Cortical shell thorny, without radial spines. All its three girdles
elliptical, one and a third times as long as broad. Four gates
kidney-shaped, nearly twice as broad as high.

_Dimensions._--Principal axis 0.17, transverse axis 0.13; length of the
lentelliptical medullary shell 0.04, breadth 0.03.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

Subgenus 2. _Pylonura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed
spines.

4. _Pylonium quadricorne_, n. sp. (Pl. 9, fig. 14).

Cortical shell thorny, with four strong horn-like curved spines in the
lateral plane. All its three girdles elliptical, one and a third times as
long as broad. Four gates kidney-shaped, twice as broad as high. (This
species is a further developmental stage of _Tetrapyle quadricornis_, with
perfect sagittal girdle.)

_Dimensions._--Principal axis 0.16, transverse axis 0.12; length of the
medullary shell 0.05, breadth 0.04.

_Habitat._--Pacific, central area, Stations 270 to 274, surface; Indian
Ocean, Madagascar, Rabbe, surface.

5. _Pylonium octacanthum_, n. sp.

Cortical shell thorny, with eight long and thin cylindrical radial spines,
lying in two crossed diagonal planes, opposite in pairs, and arising from
the eight points where the transverse girdle is crossed by the lateral
girdle. All three girdles elliptical, one and a half times as long as
broad; four gates triangular. (This species is a further developmental
stage of _Tetrapyle octacantha_, with perfect sagittal girdle.)

_Dimensions._--Principal axis 0.18, transverse axis 0.13; length of the
medullary shell 0.03 to 0.04.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

{656}6. _Pylonium stenozonium_, n. sp.

Cortical shell thorny, with eight short conical radial spines, lying in two
crossed diagonal planes (as in the foregoing species). All three girdles
nearly of the same size, subcircular, very narrow, only with one to two
rows of pores. Four gates kidney-shaped.

_Dimensions._--Principal axis 0.15, transverse axis 0.14; diameter of the
quadrangular medullary shell 0.04.

_Habitat._--Equatorial Atlantic, Station 347, surface.

Subfamily 3. TRIPLOZONARIA, Haeckel.

_Definition._--#Pylonida# with three concentric systems of fenestrated
girdles, lying in three concentric lentelliptical faces (every system with
one to three girdles lying in one lentelliptical face).

Genus 286. _Amphipylonium_,[325] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell,
surrounded by a double latticed cortical shell; inner cortical shell
_Pylonium_-shaped, with three perfect crossed girdles; outer cortical shell
only represented by a single (transverse) girdle.

The genus _Amphipylonium_ opens the series of Triplozonaria, or of those
Pylonida in which the shell is composed of three concentric systems of
latticed girdles. The first (and innermost) system represents the complete
trizonal medullary shell, which is probably a lentelliptical
_Larnacilla_-shell. The first and intermediate system is formed of a
complete trizonal cortical shell of the same form, but much larger (like
_Pylonium_). The third (and outermost) system is represented by one to
three latticed girdles, corresponding to the former and forming an outer or
second cortical shell. In _Amphipylonium_ (as the most simple form of the
Triplozonaria) there is only developed the first (transverse) girdle of the
third system. It repeats therefore the form of _Amphipyle_, the cortical
shell of which is here double (Prodromus, 1881, p. 463).

1. _Amphipylonium semilunare_, n. sp.

Inner cortical shell lentelliptical, one and a half times as long as broad,
with smooth surface and four semilunar gates. Transverse girdle of the
outer cortical shell with two smooth semilunar wings, with convex, smooth
lateral crests; each wing as broad as the transverse girdle of the inner
cortical shell and twice as long as its lateral girdle.

_Dimensions._--Length of the lentelliptical medullary shell 0.04, breadth
0.03; length of the inner cortical shell 0.15, breadth 0.1; length of each
lateral wing of the outer cortical shell 0.3, breadth 0.1.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{657}2. _Amphipylonium spinosissimum_, n. sp.

Inner cortical shell lentelliptical, one and a third times as long as
broad, with thorny surface and four kidney-shaped gates. Transverse girdle
of the outer cortical shell very spiny, with two semilunar wings, with
convex thorny lateral crests; each wing one and a half times as broad as
the transverse girdle of the inner cortical shell, and one and a half times
as long as its lateral girdle.

_Dimensions._--Length of the lentelliptical medullary shell 0.05, breadth
0.03; length of the inner cortical shell 0.16, breadth 0.12; length of each
lateral wing of the outer cortical shell 0.24, breadth 0.18.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

3. _Amphipylonium tetraceros_, n. sp.

Inner cortical shell hexagonal, one and a half times as long as broad, with
smooth surface and four transverse elliptical gates. Transverse girdle of
the outer cortical shell smooth, with rectilinear smooth lateral crests,
which are prolonged at both ends into strong conical spines (therefore four
spines in the lateral plane); each wing of the same breadth as the
transverse girdle of the inner cortical shell, and twice as long as its
lateral girdle.

_Dimensions._--Length of the hexagonal medullary shell 0.03, breadth 0.02;
length of the inner cortical shell 0.12, breadth 0.08; length of each
lateral wing of the outer cortical shell 0.24, breadth 0.08.

_Habitat._--Northern Pacific, Station 253, depth 3125 fathoms.

4. _Amphipylonium octoceros_, n. sp.

Inner cortical shell quadrangular, one and a third times as long as broad,
with thorny surface and four kidney-shaped gates. Transverse girdle of the
outer cortical shell thorny, constricted at the equator, with concave
thorny lateral crests, and with eight strong radial spines, opposite in
pairs and lying in two crossed diagonal planes; each wing narrower than the
transverse girdle of the inner cortical shell and twice as long as its
lateral girdle.

_Dimensions._--Length of the subspherical medullary shell 0.04, breadth
0.035; length of the inner cortical shell 0.14, breadth 0.11; length of
each lateral wing of the outer cortical shell 0.26, breadth 0.09.

_Habitat._--Southern Pacific, Station 295, depth 1500 fathoms.

Genus 267. _Tetrapylonium_,[326] Haeckel, 1881, Prodromus, p. 464.

{658}The genus _Tetrapylonium_ represents a further developmental stage of
the foregoing _Amphipylonium_; whilst in this latter the outer cortical
shell (or the third system of girdles) is formed only by a transverse
girdle, here this is crossed by a lateral girdle. _Tetrapylonium_ repeats
therefore the typical form of _Tetrapyle_, but with doubled cortical shell
(Prodromus, 1881, p. 464).

1. _Tetrapylonium pantellipticum_, n. sp.

Outer cortical shell elliptical, one and a third times as long as broad,
with smooth surface and four elliptical gates. Inner cortical shell
elliptical, with smooth surface and four elliptical gates which repeat the
form of the four outer gates, but are of half the size. Transverse girdle
broad, with five pores on the isthmus.

_Dimensions._--Length (or principal axis) of the first shell (medullary
shell) 0.04, breadth (or transverse axis) 0.03; length of the second (or
inner cortical) shell 0.12, breadth 0.09; length of the third (or outer
cortical) shell 0.18, breadth 0.14.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

2. _Tetrapylonium reniforme_, n. sp.

Outer cortical shell elliptical, one and a half times as long as broad,
with thorny surface and four large kidney-shaped gates. Inner cortical
shell of the same form, but one-third smaller, also with four kidney-shaped
gates. Transverse girdle small, but with two pores on the isthmus.

_Dimensions._--Length of the first (innermost) shell 0.03, breadth 0.02;
length of the second (middle) shell 0.14, breadth 0.08; length of the third
(outermost) shell 0.2, breadth 0.11.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

3. _Tetrapylonium quadrangulare_, n. sp. (Pl. 9, fig. 15).

Outer cortical shell nearly quadrangular, one and a third times as long as
broad, with four rounded corners, from which arise in the lateral plane
four three-sided pyramidal radial spines, opposite in pairs in two crossed
diagonal axes. Surface thorny; four gates elliptical or nearly
quadrangular. Inner cortical shell more elliptical, by one half smaller,
with broader transverse girdle and four kidney-shaped gates. Medullary
shell nearly spherical, of half the size.

_Dimensions._--Length of the first shell 0.045, breadth 0.035; length of
the second shell 0.1, breadth 0.07; length of the third shell 0.16, breadth
0.11.

_Habitat._--South Atlantic, off Tristan da Cunha, Station 332, depth 2200
fathoms.

4. _Tetrapylonium octacanthum_, n. sp.

Outer cortical shell elliptical, one and a third times as long as broad,
with thorny surface, and eight long and thin radial spines arising from the
corners of the four triangular gates, and lying {659}opposite in pairs in
two crossed diagonal planes. Inner cortical shell of the same shape, but
two-thirds smaller, also thorny. Transverse girdle broad, with four pores
on the isthmus. (This species appears to be the common _Tetrapyle
octacantha_, with doubled cortical shell.)

_Dimensions._--Length of the first shell 0.04, breadth 0.03; length of the
second shell 0.16, breadth 0.12; length of the third shell 0.24, breadth
0.18.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

5. _Tetrapylonium armatum_, n. sp.

Outer cortical shell nearly quadrangular, one and a fourth times as long as
broad, with very spiny surface. Between numerous smaller (simple or
branched) thorns arise twelve larger radial spines, symmetrically
distributed, eight wing-spines crossed in two diagonal planes (as in
_Tetrapylonium octacanthum_) and four corner spines on the four rounded
corners of the lateral plane (as in _Tetrapylonium quadrangulare_).
Therefore this species combines the armature of both foregoing species.
Four gates kidney-shaped. Inner cortical shell elliptical, smooth,
three-fourths smaller.

_Dimensions._--Length of the first shell 0.045, breadth 0.035; length of
the second shell 0.18, breadth 0.15; length of the third shell 0.25,
breadth 0.2.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Genus 288. _Pylozonium_,[327] n. gen.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell,
surrounded by double latticed cortical shell; inner cortical shell as well
as the outer _Pylonium_-shaped, each with three perfect crossed girdles
(transverse, lateral, and sagittal).

The genus _Pylozonium_ is the most highly developed form of the Pylonida,
as all three systems of concentric girdles here become fully developed,
each system with three perfect girdles. In this genus only among all the
Pylonida we find nine complete latticed girdles, and in each of the three
dimensive planes three concentric elliptical girdles. The first system
forms the medullary shell, the second system the inner cortical shell, and
the third system the outer cortical shell.

1. _Pylozonium novemcinctum_, n. sp.

Outer cortical shell lentelliptical, thorny, one and a third times as long
as broad, without radial spines, with four kidney-shaped gates. Inner
cortical shell of the same shape, but one-third smaller, four times as
large as the lentelliptical medullary shell.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of
the inner cortical shell 0.16, breadth 0.12; length of the outer cortical
shell 0.24, breadth 0.18.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

{660}2. _Pylozonium octacanthum_, n. sp. (Pl. 9, fig. 16).

Outer cortical shell lentelliptical, one and a third times as long as
broad, with thorny surface and eight long and thin radial spines arising
from the corners of the four elliptical gates, and lying opposite in pairs
in two crossed diagonal planes. Inner cortical shell of the same shape but
one-fourth smaller, about twice as large as the lentelliptical medullary
shell.

_Dimensions._--Length of the medullary shell 0.065, breadth 0.045; length
of the inner cortical shell 0.18, breadth 0.12; length of the outer
cortical shell 0.24, breadth 0.18.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

Family XXVII. #THOLONIDA#, n. fam. (Pl. 10).

_Definition._--#Larcoidea# with regular, completely latticed cortical
shell, which is composed of two to six or more hemispherical or cap-shaped
domes (vaulted chambers or cupolas). The domes lie opposite in pairs on the
poles of the three dimensive axes, are separated by annular constrictions,
and surround a simple or _Larnacilla_-shaped central chamber.

The family #Tholonida# represents a peculiar and very remarkable group of
the #Larcoidea#, distinguished from the other groups of this suborder by
the characteristic form of the shell, composed of a variable number of
hemispherical domes or cupolas. The middle and original part of the shell
is constantly formed of an elliptical or subspherical central chamber,
which often, but not constantly, contains a small medullary shell. An even
number (two, four, six, or more) of domes is attached to the poles of the
three dimensive axes of the central chamber; according as only one, or two,
or all three axes develop cupolas, we distinguish in this family three
different subfamilies (the Amphitholida, Staurotholida, and Cubotholida).
The Amphitholida (or Tholonida monaxonia) form cupolas only on the two
poles of one single axis, and this axis corresponds to the minor (or
transverse) axis of the central chamber, we find here therefore constantly
at least two lateral cupolas (Pl. 10, figs. 1-7). The Staurotholida (or
Tholonida diaxonia) form cupolas on the poles of two axes perpendicular one
to another; these two axes are the major (principal) and the minor
(transverse) axis of the central chamber, we find here therefore constantly
at least four cupolas crossed in pairs (Pl. 10, figs. 8-11). The
Cubotholida (or Tholonida triaxonia) form cupolas on the poles of all three
dimensive axes (perpendicular one to another); corresponding to the
principal, transverse, and sagittal axes of the lentelliptical central
chamber; therefore we find here constantly at least six cupolas, attached
in pairs on the six sides of the central chamber (Pl. 10, figs. 12-17).

The number of genera and species in the family #Tholonida# is rather large,
but the number of individuals is much smaller than in most of the other
SPUMELLARIA, and {661}particularly than in the nearly allied Pylonida; the
greater number of the species are very rare. Besides this they seem to be
very variable and inclined to produce numerous abnormalities. Very often
intermediate forms of transition are to be found between the Tholonida and
other #Larcoidea#, particularly the Pylonida and Lithelida. In many species
the thick-walled shell is very opaque, and offers great difficulties to the
clear study; in the greater number the structure of the shell cannot be
understood completely without rolling the shell to the different sides; and
the distinction between the Staurotholida and Cubotholida is often very
difficult.

The primordial chamber of the Tholonida, or the central chamber in which
its growth begins, is either a simple lentelliptical shell (without
enclosed medullary shell), like _Cenolarcus_, or it is a trizonal shell,
like _Larnacilla_, and contains a small concentric medullary shell. But
this important inner shell of the central chamber offers peculiar
difficulties for study. In many cases (probably in the greater number)
there is no doubt the same characteristic trizonal medullary shell, which
we found in the Larnacida and Pylonida, and this is our principal argument,
if we regard the Tholonida as #Larcoidea#, which are most nearly allied to
both these families, and in which the fenestrated open cortical girdles of
the Pylonida are replaced by fenestrated closed cupolas or domes; the
characteristic "gates" or large fissures in the cortical shell of the
former are therefore here perfectly closed by network. The Tholonida agree
in this point with the Larnacida, but are distinguished from them by the
prominent vaultings of the hemispherical cupolas or domes, which give them
a peculiar appearance. Each pair of domes (opposite on both poles of one
dimensive axis) corresponds to one single girdle of the Pylonida.

Regarding the absence or presence of a medullary shell in the central
chamber, we may divide the Tholonida into two groups, Cenotholida (without
medullary shell) and Coccotholida (with medullary shell). The absence of
the medullary shell in many Tholonida may be either primary (original) or
secondary (by reduction and loss of it). Probably in the larger proportion
(if not always?) its absence is the consequence of reduction and loss, and
in this case the Cenotholida must be derived phylogenetically from the
Coccotholida, but possibly often (or always?) the contrary may also be the
case. At present we cannot find certain arguments for one opinion or the
other.

The connection between the medullary (internal) and the cortical (external)
shell of the central chamber in the Coccotholida is effected by two lateral
wings of a latticed transverse girdle; on both sides of the latter remain
the four internal open "gates" of the Pylonida and Larnacida. Very often
the limiting edges of these four gates are prolonged into eight external
radial spines which lie opposite in pairs in two crossed diagonal planes,
and correspond to the eight portal-spines of _Tetrapyle octacantha_.
Besides these, we often find four other beams opposite in pairs in the two
crossed axes (principal and lateral). Very often also these radial beams
(between medullary and {662}cortical shells) are prolonged on the outside
into prominent free radial spines. But other spines may also arise from the
surface. Very remarkable is the presence of twenty symmetrically disposed
radial spines in some forms of Amphitholida (_Amphitholus acanthometra_ and
_Amphitholonium acanthonium_, &c.). It recalls the twenty radial spines of
the #Acanthonida#, though the laws of distribution in the two similar cases
are quite different.

The cortical shell of the Tholonida remains either simple, or it becomes
doubled by formation of an outer veil or envelope. This outer cortical
shell or "veil" may either repeat completely the typical form of the inner
with its cupolas, or it may form an ellipsoidal envelope without
dome-shaped partitions. Its network is either like that of the inner, or it
is very delicate and irregular, spider-web like. The connection between the
two cortical shells is effected by a variable number of radial beams, often
prolonged on the outside into radial spines.

The growth of the Tholonida by apposition of new chambers is very
characteristic, constantly pairs of chambers originating at both opposite
poles of the three dimensive axes, firstly on the transverse axis, secondly
on the principal axis, thirdly on the sagittal axis. It is possible that by
repeated apposition of new chambers the Tholonida are transformed into
#Discoidea#, but commonly the number of domes is restricted, two in the
Amphitholida, four in the Staurotholida, six in the Coccotholida. If new
chambers in an irregular manner or in a spiral order be opposed to those
first formed pairs of chambers, the Tholonida may be transformed into
Soreumida, Streblemida, or Lithelida. The phylogenetic connection between
these families is probably very complex and very worthy of further
research.

The network of the shell in the Tholonida is sometimes regular, with
circular (often hexagonally framed) pores of equal size, sometimes
irregular, with roundish pores of unequal size. Commonly the shell is thick
and compact; in those genera in which the cortical shell is doubled, the
inner is commonly compact, the outer a very delicate veil.

The central capsule of the Tholonida is originally always enclosed by the
central chamber, and like this of lentelliptical form, a triaxial
ellipsoid. With the apposition of cupolas the growing central capsule may
form dome-shaped protuberances which enter into the former. So we may find
in the Amphitholida a central capsule with three joints (separated by two
annular parallel constrictions), in the Staurotholida a cross-shaped
capsule (with four caps around the central lentellipsis), in the
Cubotholida a clustered capsule with six caps, surrounding six sides of the
cubical central mass. In the Coccotholida (with medullary shell in the
central chamber) the latter encloses in the centre the nucleus of the cell.
The calymma, or the jelly-veil between the central capsule and the
enveloping shell, probably always exhibits annular constrictions,
corresponding to those which separate the cupolas of the shell. All these
anatomical peculiarities in the Tholonida require a further accurate study.

{663}_Synopsis of the Genera of Tholonida._

-------------------------------------------------------------------------
I. Subfamily Amphitholida.
Cupolas developed only in the direction of one single axis (commonly
two cupolas).
(Tholonida unidimensiva.)
-------------------------------------------------------------------------
{Two simple cupolas
{ (without veil), 289. _Tholartus_.
Central chamber simple, {
without medullary shell. {Two double cupolas
{ (with veil), 290. _Tholodes_.

{Two simple cupolas
Central chamber { (without veil), 291. _Amphitholus_.
_Larnacilla_-shaped, with {
enclosed medullary shell. {Two double cupolas
{ (with veil), 292. _Amphitholonium_.
-------------------------------------------------------------------------
II. Subfamily Staurotholida.
Cupolas developed in the direction of two axes perpendicular one to
another (commonly four cupolas).
(Tholonida bidimensiva.)
-------------------------------------------------------------------------
{Four simple cupolas
{ (without veil), 293. _Tholostaurus_.
Central chamber simple, {
without medullary shell. {Four double cupolas
{ (with veil), 294. _Tholoma_.

{Four simple cupolas
Central chamber { (without veil), 295. _Staurotholus_.
_Larnacilla_-shaped, with {
enclosed medullary shell. {Four double cupolas
{ (with veil), 296. _Staurotholonium_.
-------------------------------------------------------------------------
III. Subfamily Cubotholida.
Cupolas developed in the direction of three axes perpendicular one
to another (commonly six cupolas).
(Tholonida tridimensiva.)
-------------------------------------------------------------------------
{Six simple cupolas
{ (without veil), 297. _Tholocubus_.
Central chamber simple, {
without medullary shell. {Six double cupolas
{ (with veil), 298. _Tholonium_.

{Six simple cupolas
Central chamber { (without veil), 299. _Cubotholus_.
_Larnacilla_-shaped, with {
enclosed medullary shell. {Six double cupolas
{ (with veil), 300. _Cubotholonium_.

Subfamily 1. AMPHITHOLIDA, Haeckel.

_Definition._--#Tholonida# with monaxial growth; cupolas opposite on the
poles of one axis. (Shell commonly three-chambered, with two domes on both
sides of the central chamber.)

Genus 289. _Tholartus_,[328] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external
veil); with two hemispherical cupolas, opposite on the poles of one axis;
central chamber between them simple (without medullary shell).

{664}The genus _Tholartus_ (Pl. 10, fig. 1) represents the most simple form
of all Tholonida, a three-jointed simple cortical shell without medullary
shell; two hemispherical cupolas or domes are attached on both lateral
sides of a simple triaxial ellipsoidal central chamber, corresponding to
the poles of its lateral or transverse axis. _Tholartus_ may be regarded as
the common ancestral form of all Tholonida, and may be derived either from
the Pylonida _Monozonium_ (by complete lattice-locking of the lateral
wings) or from the Ellipsida _Cenellipsis_ (by development of two lateral
protuberances on the poles of the transverse axis). But it is also possible
that the absence of the medullary shell is produced by reduction, and in
this case _Tholartus_ may be descended from _Amphitholus_.

Subgenus 1. _Tholartella_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Tholartus tricolus_, n. sp. (Pl. 10, fig. 1).

Central chamber one and a half times as high and as broad as both cupolas.
Surface of the shell rough. Pores regular, circular, without hexagonal
frames, three times as broad as the bars; about sixteen on the half
meridian of the central chamber (or its vertical diameter).

_Dimensions._--Length of the shell (major axis of the central chamber,
vertical) 0.1, breadth of the shell (major axis of the whole
three-chambered shell, horizontal) 0.13; pores 0.01, bars 0.003.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

2. _Tholartus paniscus_, n. sp.

Central chamber of the same breadth, but of the double height of both
cupolas. Surface of the shell rough. Pores regular, circular, with
hexagonal frames, of the same breadth as the bars; eight to ten on the half
meridian of the central chamber.

_Dimensions._--Length of the shell (major axis of the central chamber,
vertical) 0.12, breadth of the shell (major axis of the whole
three-chambered shell, horizontal) 0.15; pores and bars 0.008.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

3. _Tholartus isocolus_, n. sp.

Central chamber of the same breadth and height as both cupolas. Surface of
the shell smooth. Pores irregular, roundish, once to three times as broad
as the bars; ten to fifteen on the half meridian of the central chamber.

_Dimensions._--Length of the shell 0.11, breadth 0.14; pores 0.004 to 0.01,
bars 0.003.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

{665}Subgenus 2. _Tholartissa_, Haeckel.

_Definition._--Surface of the shell with radial spines.

4. _Tholartus tripanis_, n. sp.

Central chamber of the same size as both cupolas. Surface of the shell
thorny, everywhere covered with short conical radial spines, about as long
as the cupolas. Pores regular or subregular, circular, twice as broad as
the bars; eight to ten on the half meridian.

_Dimensions._--Length of the shell 0.1, breadth 0.12; pores 0.01, bars
0.005.

_Habitat._--South Pacific, Station 295, surface.

5. _Tholartus sagitta_, n. sp.

Central chamber twice as large as both cupolas. Surface of the shell spiny,
with eight to twelve regularly (?) distributed radial spines, radiating
from the two constrictions between the three chambers. Spines
needle-shaped, very thin and long. Pores subregular or irregular, roundish
or circular, about three times as broad as the bars; twelve to sixteen on
the half meridian.

_Dimensions._--Length of the shell 0.11, breadth 0.15; pores 0.01, bars
0.003.

_Habitat._--North Pacific, Station 244, surface.

Genus 290. _Tholodes_,[329] n. gen.

_Definition._--#Tholonida# with double cortical shell (with external veil),
with two hemispherical cupolas, opposite on the poles of one axis; central
chamber between them simple (without medullary shell).

The genus _Tholodes_ (Pl. 10, fig. 2) differs from _Tholartus_, its
probable ancestral form, only in the duplication of the shell; the outer
shell has exactly the same three-jointed form as the inner; both are
connected by eight radial beams, lying in two diagonal planes and
corresponding to the eight frontal spines of _Tetrapyle octacantha_.
Possibly _Tholodes_ may also be descended from _Amphitholonium_ by loss of
the medullary shell. I have observed only one single specimen of this
genus.

1. _Tholodes cupula_, n. sp. (Pl. 10, fig. 2).

Outer shell of the same form and structure as the inner shell. Central
chamber larger than both cupolas. Surface a little rough, without radial
spines. Distance of both shells equals the height of the outer cupolas.
Pores of both shells regular, circular, three times as broad as the
{666}bars; six to seven in the semicircle of one inner cupola, ten to
twelve in the semicircle of one outer cupola. Both shells are connected by
eight radial beams, crossed in two diagonal planes.

_Dimensions._--Major axis of the outer shell 0.1, minor 0.065; major axis
of the inner shell 0.05, minor 0.04; pores 0.006, bars 0.002.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 291. _Amphitholus_,[330] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external
veil), with two hemispherical cupolas, opposite on the poles of one axis;
central chamber (between them) _Larnacilla_-shaped, with medullary shell.

The genus _Amphitholus_ is the most simple form of all Coccotholida (or all
Tholonida provided with medullary shell). Its simple cortical shell
exhibits, like _Tholartus_, two lateral hemispheric cupolas, attached on
both sides of a lentelliptical central chamber (on the poles of its
transverse axis). The central chamber has the form of a triaxial ellipsoid,
and contains, like _Larnacilla_, a small medullary shell, connected with it
by the wings of a latticed transverse girdle. _Amphitholus_ may be derived
either from _Tholartus_ by secondary production of a cortical shell, or
(more probably) from _Larnacilla_ by apposition of two lateral dome-shaped
protuberances. In the latter case it may be regarded as an _Amphipyle_, the
lateral open wings of which are closed by lattice-work.

Subgenus 1. _Amphitholissa_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Amphitholus artiscus_, n. sp. (Pl. 10, fig. 3).

Central chamber of the same height as both cupolas. Surface of the cortical
shell rough. Pores regular, circular, with prominent hexagonal frames,
twice as broad as the bars; eight to ten in the basal semicircle of one
cupola.

_Dimensions._--Major axis of the shell 0.16, minor axis 0.1; pores 0.008,
bars 0.004.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Amphitholus artidium_, n. sp.

Central chamber one-fourth higher than both cupolas. Surface of the
cortical shell smooth. Pores irregular, roundish, once to three times as
broad as the bars. (Very similar to _Tholartus_ tricolus, Pl. 10, fig. 1,
but different by the medullary shell.)

_Dimensions._--Major axis of the shell 0.15, minor axis 0.12; pores 0.004
to 0.01, bars 0.003.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

{667}Subgenus 2. _Amphitholura_, Haeckel.

_Definition._--Surface of the shell with radial spines or thorns.

3. _Amphitholus octacanthus_, n. sp.

Central chamber about twice as high and broad as both cupolas. From the
surface arise, symmetrically distributed, eight stout conical radial
spines, lying in two diagonal planes, as prolongations of the eight
wing-beams, which connect the _Tetrapyle_-shaped medullary shell with the
two ring-like constrictions of the cortical shell. Pores of the latter
regular, circular, twice as broad as the bars; ten to twelve in the basal
semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.15, minor 0.12; pores 0.01, bars
0.005.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

4. _Amphitholus dodecanthus_, n. sp.

Central chamber a little larger than both cupolas. From the surface arise
twelve strong conical radial spines, four on the poles of the major and the
minor axis of the shell, lying in the lateral plane; eight others lying in
two diagonal planes, as prolongations of the eight wing-beams, which
connect the _Tetrapyle_-shaped medullary shell with the two ring-like
constrictions of the cortical shell. Pores of the latter irregular,
roundish, twice to four times as broad as the bars; sixteen to eighteen in
the basal semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.15, minor 0.11; pores 0.004 to
0.008, bars 0.002.

_Habitat._--North Pacific, Station 256, surface.

5. _Amphitholus acanthometra_, n. sp. (Pl. 10, figs. 5, 6).

Central chamber about twice as broad and as high as both cupolas. On the
surface quite symmetrically distributed twenty long and strong cylindrical
radial spines; eight in the sagittal plane or in the meridian plane of the
central chamber (four opposite in pairs in the principal and the sagittal
axes, four others in the middle between the principal and the sagittal
spines); four in the horizontal transverse plane, on both sides of the
poles of the major or lateral axis; and eight in diagonal planes
(corresponding to the eight wing-spines of _Tetrapyle octacantha_). In the
centre of the central chamber is a distinct trizonal medullary shell like
that of _Tetrapyle_, with two vertical columella beams. Pores of the
cortical shell subregular, circular, with elevated hexagonal frames, about
four times as broad as the bars; five to seven in the semicircle of one
cupola. (This remarkable species differs from _Tholartus vicenus_ in the
possession of a medullary shell, from _Amphitholonium acanthometra_ by the
simple cortical shell. Compare these species).

_Dimensions._--Major axis of the shell 0.14 to 0.16, minor axis 0.10 to
0.12; pores 0.015, bars 0.004; major axis of the medullary shell 0.04 to
0.05, minor (horizontal) axis 0.02 to 0.03; length of the twenty radial
spines 0.18 to 0.24, basal breadth 0.01.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

{668}6. _Amphitholus armatus_, n. sp.

Central chamber twice as broad, but of the same height, as the flat vaulted
cupolas. Surface armed with thirty to forty very large three-sided
prismatic spines, longer than the major axis of the shell. Pores irregular,
roundish, twice to five times as broad as the bars; eight to nine on the
basal semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.15, minor axis 0.1; pores 0.006 to
0.015, bars 0.003; length of the spines 0.2 to 0.25, breadth 0.01.

_Habitat._--North Atlantic, Station 353, surface.

7. _Amphitholus polyacanthus_, n. sp.

Central chamber of the same breadth and height as both cupolas. Surface
armed with very numerous (sixty to eighty) strong conical spines, about as
long as the minor axis of the shell. Pores subregular, circular, twice as
broad as the bars; sixteen to eighteen in the basal semicircle of one
cupola.

_Dimensions._--Major axis of the shell 0.13, minor 0.08; pores 0.006, bars
0.003; length of the radial spines 0.09, breadth 0.006.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

8. _Amphitholus panicium_, n. sp. (Pl. 10, fig. 4).

Central chamber ellipsoidal, nearly twice as broad, but of the same height,
as both cupolas. Whole surface thorny; the largest radial thorns about as
long as the medullary shell. Pores irregular, roundish, twice to four times
as broad as the bars. On the base of each cupola a circle of ten to twelve
larger square pores, separated by radial beams, which are prolonged into
stouter free spines. This remarkable formation of the cupolas, very rare in
this family, recalls the characteristic formation of the distal chambers of
the Panartida and Zygartida (_e.g._, Pl. 40, figs. 4, 8, &c.), where it is
very common.

_Dimensions._--Major axis of the shell 0.15, minor 0.1; pores 0.005 to
0.012, bars 0.003; large square pores 0.02; axes of the medullary shell
0.03 and 0.02.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

Genus 292. _Amphitholonium_,[331] n. gen.

_Definition._--#Tholonida# with double cortical shell (with external veil),
with two hemispherical cupolas, opposite on the poles of one axis; central
chamber (between them) _Larnacilla_-shaped, with medullary shell.

The genus _Amphitholonium_ differs from the nearly allied _Amphitholus_
(probably its ancestral form) only in the duplication of the cortical
shell; the outer has the {669}same three-jointed form as the inner, and is
connected with it by a variable number of radial beams. From the similar
_Tholodes_ it differs in the possession of a medullary shell.

1. _Amphitholonium tricolonium_, n. sp. (Pl. 10, fig. 7).

Outer cortical shell smooth, of the same three-jointed form as the inner,
at an equal distance from it throughout the whole circumference; central
chamber higher-vaulted than both cupolas. Network of the outer shell
delicate, with subregular, circular pores. Pores of the thick-walled inner
shell subregular, circular, with hexagonal frames, three times as broad as
the bars; eight to ten in the basal semicircle of one cupola.

_Dimensions._--Major axis of the outer cortical shell 0.2, minor axis 0.15;
major axis of the inner cortical shell 0.16, minor axis 0.11; pores 0.01,
bars 0.0035.

_Habitat._--South Pacific, Station 302, depth 1450 fathoms.

2. _Amphitholonium octostylium_, n. sp.

Outer cortical shell lentelliptical, smooth, not articulated, at a varying
distance from the three-jointed inner shell; central chamber of the latter
twice as high as both hemispherical cupolas. Network of the outer shell
very thin, cob-web like. Pores of the inner shell subregular, roundish,
twice as broad as the bars; six to seven in the semicircle of one cupola.
Eight thin and long, bristle-shaped, radial spines, opposite in pairs in
two crossed diagonal planes.

_Dimensions._--Major axis of the outer cortical shell 0.16, minor 0.12;
major axis of the inner cortical shell 0.12, minor 0.09; pores 0.01, bars
0.005.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

3. _Amphitholonium acanthonium_, n. sp.

Outer cortical shell of the same three-jointed form as the inner, at a
uniform distance from it; central chamber higher vaulted than both cupolas.
Network of the outer shell irregular, delicate, of the inner regular,
strong, with circular, hexagonally framed pores, three times as broad as
the bars; eight to nine in the semicircle of one cupola. On the surface,
quite symmetrically disposed, twenty long and strong, cylindrical, radial
spines; eight in the sagittal plane, four in the transverse plane; eight in
two diagonal planes between the former and the latter. The remarkable
geometric disposition of the twenty spines is in this species quite the
same as in _Amphitholus acanthometra_. It differs from this nearly allied
species in the double cortical shell.

_Dimensions._--Major axis of the outer cortical shell 0.18, minor 0.14;
major axis of the inner cortical shell 0.14, minor 0.1; pores 0.012, bars
0.004.

_Habitat._--South Pacific, Station 295, surface.

{670}Subfamily 2. STAUROTHOLIDA, Haeckel.

_Definition._--#Tholonida# with diaxial growth; cupolas crossed in pairs,
opposite on the poles of two axes, perpendicular one to another. (Shell
commonly five-chambered, with four domes, cross-wise surrounding the
central chamber.)

Genus 293. _Tholostaurus_,[332] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external
veil), composed of four hemispherical cupolas in cross-form, opposite in
pairs on the poles of two axes perpendicular one to another; central
chamber simple (without medullary shell).

The genus _Tholostaurus_ is the most simple form of the Staurotholida, or
the Tholonida with four crossed hemispherical cupolas, lying on the poles
of two axes perpendicular one to another. The central chamber communicates
by four wide openings with the four domes, and contains no medullary shell.
_Tholostaurus_ may originate either from _Tholartus_ by apposition of two
opposite cupolas between the first pair, or from _Staurotholus_ by loss of
the medullary shell.

Subgenus 1. _Tholostaurantha_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Tholostaurus quadrigatus_, n. sp.

All four cupolas nearly of the same size and form, subregular. Surface
smooth, without radial spines. Pores subregular, circular, four times as
broad as the bars; eight to ten pores in the basal semicircle of one
cupola.

_Dimensions._--Diameter of the shell 0.12; pores 0.008, bars 0.002.

_Habitat._--Pacific, central area, Station 272, surface.

2. _Tholostaurus cruciformis_, n. sp.

Two opposite cupolas larger than the two others. Surface rough, without
radial spines. Pores regular, circular, hexagonally framed, twice as broad
as the bars; twelve to fourteen in the basal semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.16, minor axis 0.12; pores 0.006,
bars 0.003.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

{671}Subgenus 2. _Tholostauroma_, Haeckel.

_Definition._--Surface of the shell with radial spines or thorns.

3. _Tholostaurus tetrabelonis_, n. sp.

All four cupolas nearly of the same size and form, subregular. Pores
regular, circular, hexagonally framed, twice as broad as the bars; twelve
to fourteen in the semicircle of one cupola. Four long and stout conical
radial spines, two vertical on the poles of the principal axis, two
horizontal on the poles of the transverse axis.

_Dimensions._--Major axis of the shell 0.15, minor 0.13; pores 0.006, bars
0.003.

_Habitat._--South Pacific, Station 288, surface.

4. _Tholostaurus octobelonis_, n. sp.

Two opposite cupolas larger than the two others. Pores subregular,
circular, three times as broad as the bars; ten to twelve in the semicircle
of one cupola. Eight long and thin, needle-shaped, radial spines, in pairs
crossed in two diagonal planes.

_Dimensions._--Major axis of the shell 0.16, minor 0.12; pores 0.01, bars
0.003.

_Habitat._--South Pacific, Station 291, surface.

5. _Tholostaurus dodecabelos_, n. sp.

Two opposite cupolas larger than the two others. Pores irregular, roundish,
twice to three times as broad as the bars; six to eight in the semicircle
of one cupola. Twelve thin and long radial spines, four on the poles of the
two crossed axes (principal and lateral), eight crossed in two diagonal
planes.

_Dimensions._--Major axis of the shell 0.14, minor 0.11; pores 0.006 to
0.01, bars 0.003.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

6. _Tholostaurus polybelonis_, n. sp.

Two opposite cupolas larger than the two others. Pores circular,
hexagonally framed, of the same breadth as the bars; ten to twelve in the
semicircle of one cupola. Numerous (twenty to thirty or more) thin,
bristle-like, radial spines, about as long as the radius of the shell.

_Dimensions._--Major axis of the shell 0.15, minor axis 0.12; pores and
bars 0.007.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

Genus 294. _Tholoma_,[333] n. gen.

_Definition._--#Tholonida# with double cortical shell (with external veil),
composed of four hemispherical cupolas in cross-form, opposite in pairs on
the poles of two axes perpendicular one to another; central chamber simple
(without medullary shell).

{672}The genus _Tholoma_ (Pl. 10, figs. 10, 13) differs from the preceding
_Tholostaurus_ (its probable ancestral form) only in the duplication of the
shell. The outer (secondary) shell has the same cross-form as the inner
(primary) shell. In the two observed species the growth seems to be
different, in the first species all four cupolas of each cross being of the
same size, form, and age; in the second two opposite cupolas, larger and
apparently older than the other two. _Tholoma_ is possibly the offspring of
_Staurotholonium_, from which it may have been produced by loss of the
medullary shell.

Subgenus 1. _Tholomantha_, Haeckel.

_Definition._--Surface of the shell smooth, without radial spines.

1. _Tholoma quadrigeminum_, n. sp. (Pl. 10, fig. 10).

All four chambers in each cortical shell nearly equal, subregular. Surface
smooth, without radial spines. Structure of the network in both shells
similar. Pores regular, circular, twice as broad as the bars; six to eight
in the basal semicircle of one inner, ten to twelve in the semicircle of
one outer cupola.

_Dimensions._--Diameter of the outer shell 0.14, of the inner 0.09; pores
0.006, bars 0.003.

_Habitat._--South Pacific, Station 302, surface.

Subgenus 2. _Tholomura_, Haeckel.

_Definition._--Surface of the shell with radial spines.

2. _Tholoma metallasson_,[334] n. sp. (Pl. 10, fig. 13).

Two opposite chambers in each cortical shell larger than the two others;
the larger chambers of the inner shell corresponding to the smaller
chambers of the outer shell. Network in both shells of the same structure,
regular, with circular, hexagonally framed pores of the same breadth as the
bars; eight to ten in the semicircle of an inner, fourteen to sixteen in
the semicircle of an outer cupola. Numerous (twenty to thirty or more)
thin, cylindrical, bristle-shaped, long radial spines, symmetrically
disposed.

_Dimensions._--Major axis of the outer shell 0.26, minor axis 0.2; major
axis of the inner shell 0.16, minor axis 0.13; pores and bars 0.006.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Genus 295. _Staurotholus_,[335] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external
veil), composed of four hemispherical cupolas in cross-form, opposite in
pairs on the poles of {673}two axes perpendicular one to another; central
chamber _Larnacilla_-shaped (with enclosed medullary shell).

The genus _Staurotholus_ differs from _Tholostaurus_ in the possession of a
medullary shell in the central chamber, and may be derived from this genus
by its production. But it may also be derived from _Amphitholus_ by
apposition of two secondary opposite cupolas between the two primary
cupolas. The symmetrical position of an increasing number of radial spines
in the different species is remarkable (resembling _Tholostaurus_ as well
as _Amphitholus_).

Subgenus 1. _Staurotholissa_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Staurotholus quadratus_, n. sp.

Surface of the cortical shell smooth. All four cupolas nearly of the same
size and form; therefore principal and lateral axes equal. Pores regular,
circular, with hexagonal frames, twice as broad as the bars; eight to ten
on the basal semicircle of one cupola. Medullary shell square.

_Dimensions._--Diameter of the cortical shell 0.15; pores 0.01, bars 0.005.

_Habitat._--South Atlantic, Station 323, depth 1900 fathoms.

2. _Staurotholus cruciatus_, n. sp.

Surface of the cortical shell rough. The two principal cupolas somewhat
larger than the two lateral cupolas; therefore the longitudinal axis longer
than the transverse. Pores irregular, roundish, twice to three times as
broad as the bars; twelve to sixteen in the semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.16, minor 0.13; pores 0.005 to
0.009, bars 0.003.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Subgenus 2. _Staurotholura_, Haeckel.

_Definition._--Surface of the shell with radial spines.

3. _Staurotholus tetrastylus_, n. sp. (Pl. 10, fig. 8).

The two principal cupolas smaller than the two lateral cupolas. Pores
subregular, circular, three times as broad as the bars; six to eight in the
semicircle of one cupola. Medullary shell elliptical. On the surface four
long cylindrical radial spines; two in the principal and two in the lateral
axis.

_Dimensions._--Major axis of the cortical shell 0.15, minor 0.12; pores
0.012, bars 0.004.

_Habitat._--South Pacific, Station 166, surface.

{674}4. _Staurotholus octostylus_, n. sp.

The two lateral cupolas larger than the two principal cupolas. Pores
subregular, circular, twice as broad as the bars; ten to twelve in the
semicircle of one cupola. Medullary shell hexagonal, connected with each
ring-like constriction by eight strong cylindrical radial spines (longer
than the whole shell), lying in two crossed meridian planes, and
corresponding to the eight diagonal wing-spines of _Tetrapyle octacantha_.

_Dimensions._--Major axis of the cortical shell 0.12, minor 0.1; pores
0.008, bars 0.004.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

5. _Staurotholus decastylus_, n. sp.

The two principal cupolas twice as broad and high as the two lateral
cupolas. Pores subregular, circular, four times as broad as the bars; six
to eight in the semicircle of one cupola. Ten long and thin radial spines
with thickened conical bases, about as long as the major axis of the shell,
by inner prolongations connected with the lentelliptical medullary shell;
two polar spines in the principal axis, eight wing-spines in two crossed
diagonal planes.

_Dimensions._--Major axis 0.12, minor 0.1; pores 0.012, bars 0.003.

_Habitat._--Western Tropical Pacific; Station 224, depth 1850 fathoms.

6. _Staurotholus dodecastylus_, n. sp. (Pl. 10, fig. 9).

The two lateral cupolas larger than the two principal cupolas. Pores
subregular, circular, three times as broad as the bars; five to seven in
the semicircle of one cupola. Twelve strong cylindrical radial spines, four
on the poles of the two larger axes (two principal and two lateral), eight
wing-spines in two crossed diagonal planes. Medullary shell hexagonal.

_Dimensions._--Major (lateral) axis 0.11, minor (principal) 0.09; pores
0.009, bars 0.003.

_Habitat._--Tropical Atlantic, Station 348, surface.

7. _Staurotholus polystylus_, n. sp.

The two principal cupolas larger than the two lateral cupolas. Pores
irregular, roundish, twice to three times as broad as the bars; ten to
twelve in the semicircle of one cupola. On the surface numerous thin
bristle-shaped spines (twenty to thirty or more).

_Dimensions._--Major axis 0.14, minor 0.11; pores 0.006 to 0.01, bars
0.003.

_Habitat._--North Pacific, Station 231, surface.

Genus 296. _Staurotholonium_,[336] n. gen.

{675}The genus _Staurotholonium_ differs from the nearly allied
_Staurotholus_ (its probable ancestral form) only in the duplication of the
cortical shell. The outer shell commonly repeats the cross-form of the
inner, with four corresponding cupolas; but sometimes the four cupolas of
the outer shell alternate in size and form with those of the inner, or the
outer shell forms a simple lenticular envelope around the inner. From the
similar _Tholoma_, _Staurotholonium_ differs in the possession of a
medullary shell in the central chamber.

Subgenus 1. _Staurotholodes_, Haeckel.

_Definition._--Surface of the outer cortical shell smooth or rough, without
radial spines.

1. _Staurotholonium biquadratum_, n. sp.

Outer cortical shell smooth, of the same regular crucial form as the inner,
equidistant from it everywhere. Form and structure of both cortical shells
nearly the same, but the outer about twice as large as the inner; both
connected only by eight diagonal beams. All four cupolas of each cortical
shell regular, of the same size. Pores regular, circular, twice as broad
as the bars; six to eight in the basal semicircle of each cupola. Medullary
shell square. (Similar to _Staurotholus quadratus_, but differs by the
double cortical shell.)

_Dimensions._--Diameter of the outer cortical shell 0.16, of the inner
0.08, of the medullary shell 0.04; pores of the inner cortical shell 0.006,
bars 0.003.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

2. _Staurotholonium bicruciatum_, n. sp.

Outer cortical shell rough, twice as large as the inner, of the same form
and structure, at an equal distance from it. In both shells the principal
cupolas are larger than the lateral. Pores irregular, roundish, in the
outer shell four times, in the inner twice as large as the bars; eight to
ten in the basal semicircle of one cupola. (Similar to _Staurotholus
cruciatus_, but differs mainly in the double cortical shell.) Medullary
shell lentelliptical.

_Dimensions._--Major (longitudinal) axis of the outer cortical shell 0.016,
of the inner 0.08; minor (transverse) axis of the former 0.14, of the
latter 0.07; medullary shell 0.03 to 0.04.

_Habitat._--Pacific, central area, Station 267, depth 2700 fathoms.

3. _Staurotholonium alternatum_, n. sp.

Outer cortical shell smooth, very different from the inner; in the outer
the two principal cupolas are larger than the two lateral; in the inner
inversely smaller. Pores subregular, circular, in the outer shell four
times, in the inner shell twice as broad as the bars; ten to twelve pores
in the basal semicircle of one cupola.

{676}_Dimensions._--Major axis (length) of the outer cortical shell 0.15,
minor (breadth) 0.13; pores 0.012, bars 0.003; major axis (length) of the
inner cortical shell 0.11, minor axis (breadth) 0.09; pores 0.006, bars
0.003.

_Habitat._--Indian Ocean, surface, Madagascar, Rabbe.

4. _Staurotholonium lenticulare_, n. sp.

Outer cortical shell smooth, lenticular, with circular circumference, with
very delicate irregular network and small roundish pores. Its distance from
the inner much greater in the four diagonal points than in the four polar
points. Inner cortical shell regular, cross-like, with four cupolas of
similar size and form. Pores subregular, circular, twice as broad as the
bars; eight to ten in the basal semicircle of one cupola. Medullary shell
lenticular.

_Dimensions._--Diameter of the outer cortical shell 0.16, of the inner
0.12; pores of the inner 0.008, bars 0.004.

_Habitat._--North Atlantic, Station 353, surface.

Subgenus 2. _Staurotholoma_, Haeckel.

_Definition._--Surface of the outer cortical shell with radial spines or
thorns.

5. _Staurotholonium octodoratium_, n. sp.

Outer cortical shell of the same form and structure as the inner, but twice
as large, both principal domes somewhat larger than both lateral. Pores
subregular, circular, of the same breadth as the bars; five to six in the
semicircle of one inner, eight to ten of one outer cupola. Eight thin and
long bristle-like spines, opposite in pairs in two crossed diagonal planes.

_Dimensions._--Major axis of the outer cortical shell 0.15, minor 0.12;
major axis of the inner cortical shell 0.07, minor 0.06; pores and bars
0.005.

_Habitat._--North Pacific, Station 231, surface.

6. _Staurotholonium octodoronium_, n. sp. (Pl. 10, fig. 11).

Outer cortical shell of the same form as the inner, both little distant;
network of the outer irregular and delicate. Pores of the inner regular,
circular, twice as broad as the bars; seven to nine in the basal semicircle
of one cupola. Both lateral domes larger than the principal. Eight long and
thin, cylindrical radial spines opposite in pairs in two crossed diagonal
planes.

_Dimensions._--Major axis of the outer cortical shell 0.2, minor 0.18;
major axis of the inner cortical shell 0.16, minor 0.14; pores 0.008, bars
0.004.

_Habitat._--South Pacific, Station 288, surface.

{677}Subfamily 3. CUBOTHOLIDA, Haeckel.

_Definition._--#Tholonida# with triaxial growth; cupolas lying in pairs on
the six sides of a cubical central chamber, opposite at the poles of three
axes perpendicular one to another. (Shell commonly seven-chambered, with
six domes surrounding the central chamber.)

Genus 297. _Tholocubus_,[337] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external
veil), composed of six hemispherical cupolas, opposite in pairs on the
poles of three axes perpendicular one to another, covering six sides of the
simple cuboidal central chamber (without medullary shell).

The genus _Tholocubus_ is the most simple form of the Cubotholida, or of
the Tholonida with domes situated in three axes perpendicular one to
another; six hemispherical cupolas lying on the six sides of a cuboidal
central chamber; this latter contains no medullary shell. _Tholocubus_ may
be derived phylogenetically either from _Tholostaurus_ by apposition of two
opposite domes on the flat sides of the cross-shell, or from _Cubotholus_
by loss of the medullary shell.

Subgenus 1. _Tholocubulus_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Tholocubus tessellatus_, n. sp. (Pl. 10, fig. 12).

Surface of the shell smooth, without radial spines. Pores regular,
circular, hexagonally framed, twice as broad as the bars; eight to twelve
pores on the semicircle of one cupola. Principal cupolas smaller than the
lateral, larger than the sagittal cupolas.

_Dimensions._--Diameter of the shell 0.15; pores 0.01, bars 0.005.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

2. _Tholocubus tesserarius_, n. sp.

Surface of the shell rough, without radial spines. Pores irregular,
roundish, once to three times as broad as the bars; twelve to sixteen pores
in the semicircle of one cupola. All six cupolas nearly of the same size.

_Dimensions._--Diameter of the shell 0.16; pores 0.004 to 0.008, bars
0.003.

_Habitat._--North Atlantic, Station 353, depth 2965 fathoms.

{678}Subgenus 2. _Tholocubitus_, Haeckel.

_Definition._--Surface of the shell with radial spines or thorns.

3. _Tholocubus tesseralis_, n. sp. (Pl. 10, fig. 16).

Surface of the shell with numerous (eight to sixteen) thin and long,
bristle-shaped radial spines (the greater part broken off in the figured
specimen); pores subregular, circular, three to four times as broad as the
bars; ten to twelve in the semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.16, minor axis 0.14; pores 0.01,
bars 0.0027.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 298. _Tholonium_,[338] n. gen.

_Definition._--#Tholonida# with double cortical shell (with external veil),
composed of six hemispherical cupolas, opposite in pairs on the poles of
three axes perpendicular one to another, covering six sides of the simple
cuboidal central chamber (without medullary shell).

The genus _Tholonium_ (Pl. 10, fig. 17) differs from the preceding
_Tholocubus_ only in the duplication of the cortical shell. The outer
(secondary) shell forms either a simple (spheroidal or ellipsoidal) thin
veil around the inner (primary) shell, or both shells are of the same form,
with six corresponding cupolas. Possibly _Tholonium_ is the offspring of
_Cubotholonium_, having originated by loss of the medullary shell.

Subgenus 1. _Tholonetta_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines
or thorns.

1. _Tholonium bicubicum_, n. sp.

Outer shell with six hemispherical dome-shaped protuberances, corresponding
in those of the inner shell. Both shells connected by numerous radial
beams. Surface of the outer shell smooth; its network nearly of the same
shape as that of the inner, with subregular, circular pores, twice as broad
as the bars; ten to fifteen pores on the semicircle of one cupola.

_Dimensions._--Diameter of the outer shell 0.14, of the inner 0.12; pores
of the latter 0.006, bars 0.003.

_Habitat._--South Atlantic, Station 325, depth 2650 fathoms.

{679}2. _Tholonium ellipticum_, n. sp.

Outer shell ellipsoidal, without dome-shaped protuberances, with smooth
surface, without radial spines; network delicate, with subregular, circular
pores. Inner shell with regular, circular, hexagonally framed pores, twice
as broad as the bars; ten to twelve pores on the semicircle of one cupola.

_Dimensions._--Major axis of the outer shell 0.16, minor axis 0.14; major
axis of the inner shell 0.14, minor axis 0.12; pores 0.008, bars 0.004.

_Habitat._--Pacific central area, Station 267, depth 2700 fathoms.

3. _Tholonium sphæricum_, n. sp.

Outer shell spherical, without dome-shaped protuberances, with smooth
surface, without radial spines; network very delicate, with very small
subregular, circular pores. Inner shell with regular, circular pores of the
same breadth as the bars; fourteen to sixteen on the basal semicircle of
one cupola.

_Dimensions._--Diameter of the spherical outer shell 0.15, inner shell
0.12; pores and bars of the inner shell 0.005.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

Subgenus 2. _Tholonilla_, Haeckel.

_Definition._--Surface of the outer shell covered with radial spines or
thorns.

4. _Tholonium hexonium_, n. sp. (Pl. 10, fig. 17).

Outer shell ellipsoidal, without dome-shaped protuberances, covered with
numerous (thirty to fifty) thin and long, bristle-shaped radial spines;
network very delicate, irregular, with roundish pores. Inner shell with six
marked hemispherical domes of somewhat different sizes; pores regular,
circular, with prominent hexagonal frames, twice as broad as the bars; ten
to twelve on the basal semicircle of one cupola.

_Dimensions._--Major axis of the outer shell 0.15, minor axis 0.14; major
axis of the inner shell 0.13, minor axis 0.12; pores 0.008, bars 0.004.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

5. _Tholonium sphæronium_, n. sp.

Outer shell very thin walled, spherical, without dome-shaped protuberances,
with smooth surface, excepting eight large radial spines, rising from the
eight corners of the inner cubical central chamber. Pores of the outer
shell very small, subcircular. Inner shell very thick walled, with six
marked hemispherical domes; pores subregular, circular, with prominent
hexagonal frames, twice as broad as the bars; six to eight on the
semicircle of one cupola.

{680}_Dimensions._--Diameter of the spherical outer shell 0.15, inner shell
0.11; pores of the inner 0.01, bars 0.005.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

Genus 299. _Cubotholus_,[339] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external
veil), composed of six hemispherical cupolas, opposite in pairs on the
poles of three axes perpendicular one to another, covering six sides of the
cuboidal _Larnacilla_-shaped central chamber (with medullary shell).

The genus _Cubotholus_ differs from _Tholocubus_ in the possession of a
medullary shell in the central chamber, and may be derived from this genus
by its production; but it may also be derived from _Staurotholus_ by
apposition of two opposite domes on the flat sides of the cross-shell.
Sometimes all six domes are of the same size and form, but commonly
different in pairs.

Subgenus 1. _Cubotholissa_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Cubotholus regularis_, n. sp. (Pl. 10, fig. 14).

On the six sides of the cubical central chamber six hemispherical cupolas
of the same size and form. Surface smooth. Pores regular, circular, twice
as broad as the bars; eight in the basal semicircle of each cupola.
Medullary shell apparently spherical (?), connected with the eight corners
of the central chamber by eight radial beams, regularly disposed. (This
species is remarkable for the perfect symmetry of the shell, the six sides
of which appear to be quite similar. It differs from the similar
_Tholocubus regularis_ in the possession of a medullary shell.)

_Dimensions._--Diameter of the cortical shell (equal in all three dimensive
axes) 0.15; pores 0.01, bars 0.005; diameter of the medullary shell 0.04.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

2. _Cubotholus quadraticus_, n. sp.

Surface of the shell smooth. Both principal cupolas (on the poles of the
longitudinal axis) larger than the four other cupolas, which have the same
size and lie cross-wise in the equatorial plane. (Therefore two of the
three fundamental axes equal, the third larger.) Pores subregular,
circular, three times as broad as the bars; ten to twelve in the semicircle
of each cupola-basis.

_Dimensions._--Major axis 0.16, minor axis 0.12; pores 0.012, bars 0.004.

_Habitat._--Indian Ocean, Zanzibar, Pullen; depth 2200 fathoms.

{681}3. _Cubotholus rhombicus_, n. sp.

Surface of the shell smooth. Both principal cupolas (on the poles of the
longitudinal axis) larger than the two lateral (on the poles of the
transverse axis), and these larger than the two sagittal cupolas (on the
poles of the sagittal axis). Therefore all three fundamental axes unequal.
Pores irregular, roundish, twice to four times as broad as the bars; eight
to twelve in the basal semicircle of each cupola. Medullary shell
lentelliptical.

_Dimensions._--Major (principal) axis 0.16, middle (lateral) axis 0.14,
minor (sagittal) axis 0.12; pores 0.006 to 0.012, bars 0.003.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

Subgenus 2. _Cubotholura_, Haeckel.

_Definition._--Surface of the shell with radial spines or thorns.

4. _Cubotholus octoceras_, n. sp.

All six cupolas in pairs of different sizes. Both principal domes larger
than the lateral domes, and these larger than the sagittal domes. Pores
subregular, circular, twice as broad as the bars; eight to ten in the
semicircle of each cupola. From the surface arise (at the intersecting
points of every three cupolas) eight strong conical radial spines, about as
long as the shell-axis; they lie in two diagonal planes, and are the
external free prolongations of eight inner beams (homologous with the eight
wing-spines of _Tetrapyle octacantha_), which connect the lentelliptical
medullary shell with the eight corners of the cuboidal central chamber.

_Dimensions._--Major axis of the cortical shell 0.15, middle 0.13, minor
0.11; pores 0.008, bars 0.004; diameters of the medullary shell
corresponding to 0.05, 0.04, 0.03.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

Genus 300. _Cubotholonium_,[340] n. gen.

_Definition._--#Tholonida# with double (or sometimes triple) cortical shell
(with external veil), composed of six hemispherical cupolas, opposite in
pairs on the poles of three axes perpendicular one to another, covering six
sides of the cuboidal _Larnacilla_-shaped central chamber (with medullary
shell).

The genus _Cubotholonium_ differs from the nearly allied _Cubotholus_ (its
probable ancestral form) only in duplication of the cortical shell. I have
observed only two species of this rare form, both rather different. In the
first species the outer cortical shell forms a simple spherical thin veil
around the inner, the six cupolas of which are nearly of the same form and
size. In the second species each of the domes of the {682}inner cortical
shell is protected by an outer larger cupola, and besides this the whole
shell is enveloped by a thin ellipsoidal veil (Pl. 10, fig. 15). Therefore
this species may be the representative of a peculiar genus, the most highly
developed of all Tholonida--_Tholothauma_.

1. _Cubotholonium sphæroides_, n. sp.

Outer cortical shell (or veil) spherical, with smooth surface; network very
delicate, with very thin bars and very small irregular, roundish pores.
Inner cortical shell simple, composed of six hemispherical cupolas,
surrounding the six sides of the cubical central chamber, which encloses a
spherical medullary shell (one-third as large as itself). Network of the
inner cortical shell regular, with circular pores of the same breadth as
the bars, twelve to fourteen in the basal semicircle of one cupola.

_Dimensions._--Diameter of the spherical outer shell 0.2, of the inner
cortical shell 0.15; pores and bars of the latter 0.006.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

2. _Cubotholonium ellipsoides_, n. sp. (Pl. 10, fig. 15).

_Tholothauma ellipsoides_, Haeckel, 1883, MS.

Outer cortical shell (or veil) ellipsoidal, with very thin irregular
network and thorny surface. Inner cortical shell double, with six double,
flatly vaulted cupolas, surrounding the six sides of the
_Larnacilla_-shaped central chamber; the double domes of each shell are in
opposite pairs somewhat larger than the alternating pairs. Pores
subregular, circular, about the same breadth as the bars; eight to twelve
in the basal semicircle of one cupola. Central chamber with ellipsoidal
medullary shell. Radial spines short, very numerous.

_Dimensions._--Major axis of the outer cortical shell 0.28, minor 0.24;
major axis of the inner cortical shell 0.16, minor axis 0.14; pores and
bars 0.006; medullary shell 0.03.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Family XXVIII. #ZONARIDA#, n. fam. (Pl. 50, figs. 9-12).

_Definition._--#Larcoidea# with regular, completely latticed cortical
shell, distinguished by two to four or more annular constrictions, which
lie (all or partly) in the dimensive planes (sagittal, transverse, or
lateral), and by which four to eight or more vaulted cupolas or dome-like
chambers become separated. In the centre of this chambered cortical shell
lies constantly a trizonal or _Larnacilla_-shaped medullary shell.

The family #Zonarida# comprises a small number of peculiar #Larcoidea#,
resembling the Tholonida in the composition of the polythalamous cortical
shell by a number of cupolas or dome-shaped protuberances. But the
disposition and origin of these latter {683}are quite different. Whilst in
the Tholonida the axes of the domes are dimensive axes, and these are
separated by annular constrictions lying in diagonal planes, in the
Zonarida we find the contrary; the axes of the domes are here diagonal
axes, and these are separated by annular constrictions lying in dimensive
planes. However, this definition agrees absolutely only in the
four-chambered _Zonarium_ and in the eight-chambered _Zonidium_, whilst in
the six chambered _Zoniscus_ only four domes are disposed according to this
law, two others, however, in the same manner as in the Tholonida. Therefore
this genus is intermediate between both families.

_The Cortical Shell_ of the Zonarida is in all cases completely latticed
and of regular lentelliptical fundamental form, as in the nearly allied
Larnacida and Tholonida. The three dimensive axes are constantly of
different sizes, each with two equal poles; commonly (as in the human body)
the principal or longitudinal axis is the longest, the sagittal (or
dorso-ventral) axis the shortest; the transverse (or lateral) axis being
intermediate between them. Of the three dimensive planes the lateral plane
is the largest (determined by the principal and transverse axes); the
smallest is the equatorial plane (crossed by the transverse and sagittal
axes); the sagittal plane (determined by the sagittal and principal axes)
being intermediate between them.

The annular constrictions of the cortical shell which produce the
dome-shaped protuberances are different in number in the three known
genera--two, three, or four. To each constriction often (but not always)
corresponds an internal latticed septum, which connects the cortical with
the medullary shell. The number of the cupolas is always double the number
of the annular constrictions by which they are separated, therefore four,
six, or eight.

In all known Zonarida the sagittal septum is quite constant, but derived
from the original axial rod, which lies in the principal axis. By
ramification of this axial beam and reticular connection with the sagittal
girdle arises the sagittal septum, which we found first in _Octopyle_,
halving the four gates of _Tetrapyle_. Whilst this sagittal septum (between
right and left halves of the body) is common to all three known genera of
this family, the number and shape of the other annular constrictions are
different. In _Zonarium_ (Pl. 50, fig. 9) we find only one transverse
constriction (in the equatorial plane), in _Zoniscus_ (figs. 10, 11) two
parallel transverse constrictions (parallel to the equatorial plane, on
both sides of it). _Zonidium_ (fig. 12) is a combination of both foregoing
genera; it has three parallel transverse constrictions (one in the
equatorial plane, and one on each side of it).

_The Latticed Domes_ (cupolas or chambers) of the cortical shell exhibit
correspondingly a different number and disposition in the three known
genera. In _Zonarium_ are found only four crossed chambers, separated by
the sagittal and transverse septa; the axes of the four crossed domes are
diagonal axes, whilst in the similar _Staurotholus_ they are dimensive axes
(principal and transverse axes). In _Zonidium_ we find eight domes, each
cupola of _Zonarium_ being halved by a diagonal septum. _Zoniscus_ is
intermediate {684}between the two foregoing genera, and has six cupolas,
three on each side of the sagittal septum.

The inner communication of the cupolas or chambers is more or less free,
the lattice-work of the separating septa between them commonly remaining
more or less imperfect, or represented only by some isolated beams or
meshes. The outer network of the cupolas is commonly irregular (as in the
majority of #Larcoidea#), but sometimes distinguished by a small number of
regularly disposed larger apertures (similar to the "gates" of the
Pylonida). From the surface radial spines often arise in characteristic
number and symmetrical disposition, commonly as prolongations of the septal
axes or of the constricted edges.

The _Medullary Shell_ in all Zonarida is a true trizonal or
_Larnacilla_-shaped lattice-shell (compare above, p. 600); its perimeter
(or the first lateral girdle) is sometimes more elliptical, at other times
more hexagonal; the hexagon is amphithect; both its lateral sides are often
concave and commonly longer than the four other sides.

The _Central Capsule_ in all Zonarida is in a strict geometrical sense a
true lentellipsis (compare above, p. 599); its principal axis is commonly
one and a third to one and a half times as great as the transverse axis,
and twice to three times as great as the sagittal axis. The lentelliptical
central capsule encloses the trizonal medullary shell, whilst it is
externally enveloped by the chambered cortical shell.

_Synopsis of the Genera of Zonarida._

Two annular constrictions and four cupolas, 301. _Zonarium_.

Three annular constrictions and six cupolas, 302. _Zoniscus_.

Four annular constrictions and eight cupolas, 303. _Zonidium_.

Genus 301. _Zonarium_,[341] n. gen.

_Definition._--#Zonarida# with four dome-shaped chambers of the cortical
shell, separated by two annular constrictions (one sagittal and one
transverse).

The genus _Zonarium_ is the most simple form of the Zonarida, and differs
from the nearly allied _Larnacalpis_ by two ring-like constrictions, which
are crossed at right angles, one in the sagittal (or median) plane, and one
in the transverse (or equatorial) plane. By these two annular constrictions
four egg-shaped or kidney-shaped chambers become imperfectly separated,
which correspond to the quadrants of the lateral plane. The first cause of
the marked constrictions may be the formation of the latticed sagittal
septum, which is found first in _Octopyle_, as halving the four gates of
{685}_Tetrapyle_. Between this septum and the constricted narrow transverse
girdle the cortical shell grows out in the form of four vaulted cupolas;
every two opposite domes are congruent, two neighbouring are symmetrically
equal.

1. _Zonarium quadrigatum_, n. sp.

Cortical shell quadrangular, one and a half times as long as broad, with
four rounded corners. Surface thorny, with numerous short radial spines.
Sagittal constriction twice as long as the hexagonal medullary shell. Four
cupolas kidney-shaped.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.11; length of
the medullary shell 0.06, breadth 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Zonarium quadrispinum_, n. sp.

Cortical shell quadrangular, nearly rectangular, about twice as long as
broad, with four corners, from which arise four strong, three-sided
pyramidal, radial spines (crossed in two diagonals of the lateral plane).
Surface thorny, only smooth in the transverse constriction. Sagittal
constriction three times as long as the lentelliptical medullary shell.
Four cupolas elliptical or nearly quadrangular.

_Dimensions._--Length of the cortical shell 0.22, breadth 0.1; length of
the medullary shell 0.07, breadth 0.05.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

3. _Zonarium octangulum_, n. sp. (Pl. 50, fig. 9).

Cortical shell octangular, one and a fourth times as long as broad, with
eight strong conical spines on the eight corners; these are separated by
four deep constrictions on the poles of the principal and transverse axes,
and by four truncated planes on the poles of the crossed diagonal axes
between the former. Sagittal constriction twice as long as the hexagonal
medullary shell. Four cupolas kidney-shaped or nearly pentagonal.

_Dimensions._--Length of the cortical shell 0.15, breadth 0.12; length of
the medullary shell 0.07, breadth 0.04.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

4. _Zonarium tetratholium_, n. sp.

Cortical shell quadrangular, with four rounded corners. Surface thorny,
with sixteen stronger radial spines; eight of these lie in the lateral
plane, in the same symmetrical disposition as in the foregoing species;
eight others lie on both sides of the lateral plane, opposite in pairs in
two crossed diagonal planes, in the same symmetrical disposition as in
_Tetrapyle octacantha_. Sagittal {686}constriction three times as long as
the lentelliptical medullary shell. Four cupolas obliquely elliptical.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.12; length of
the medullary shell 0.04, breadth 0.03.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

Genus 302. _Zoniscus_,[342] n. gen.

_Definition._--#Zonarida# with six dome-shaped chambers of the cortical
shell, separated by three annular constrictions (one sagittal and two
transverse, parallel to the equatorial plane).

The genus _Zoniscus_ differs from _Zonarium_ as well as from _Zonidium_ by
the development of the transverse girdle, which is not constricted, but on
the contrary prominently vaulted in the equatorial plane. Therefore both
wings of the transverse girdle form here two opposite lateral or
"equatorial cupolas," as in _Amphitholus_. These are separated from four
other domes (the "corner cupolas") by two transverse annular constrictions,
which correspond to the free edges of the original transverse girdle. The
corner domes of each pair are separated from each other by the sagittal
septum.

1. _Zoniscus rectangulus_, n. sp.

Cortical shell nearly rectangular, with rounded corners, nearly one and a
half times as long as broad. Surface thorny, without larger radial spines.
Sagittal constriction scarcely half as long as the hexagonal medullary
shell. Both equatorial cupolas (or wings of the transverse girdle) scarcely
half as large as the four corner cupolas.

_Dimensions._--Length of the cortical shell 0.15, breadth 0.11; length of
the medullary shell 0.07, breadth 0.045.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

2. _Zoniscus hexathalamus_, n. sp.

Cortical shell nearly elliptical, one and a third times as long as broad.
Surface nearly smooth, without radial spines. Sagittal constriction three
times as long as the lentelliptical medullary shell. Both equatorial
cupolas about as large as the four corner cupolas.

_Dimensions._--Length of the cortical shell 0.12, breadth 0.09; length of
the medullary shell 0.035, breadth 0.025.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{687}3. _Zoniscus tetracanthus_, n. sp. (Pl. 50, fig. 10).

Cortical shell nearly quadrangular, one and a half times as long as broad,
with four prominent corners, from which arise in the lateral plane four
strong, three-sided pyramidal radial spines (opposite in pairs in two
crossed diagonal axes). Surface thorny, with exception of the concave
lateral sides. Sagittal constriction two and a half times as long as the
hexagonal medullary shell. Both equatorial cupolas scarcely half as large
as the four corner cupolas.

_Dimensions._--Length of the cortical shell 0.18, breadth 0.12; length of
the medullary shell 0.07, breadth 0.04.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

4. _Zoniscus octacanthus_, n. sp.

Cortical shell nearly four-sided, prismatic, one and a third times as long
as broad, with spiny surface. Eight longer thin radial spines opposite in
pairs in two crossed diagonal planes (as in _Tetrapyle octacantha_).
Sagittal constriction two and a half times as long as the hexagonal
medullary shell. Both equatorial cupolas somewhat larger than the four
corner cupolas.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.12; length of
the medullary shell 0.055, breadth 0.04.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

5. _Zoniscus hexatholius_, n. sp. (Pl. 50, fig. 11).

Cortical shell nearly rectangular, with four rounded corners and deep
sagittal constriction, one and a fourth times as long as broad with thorny
surface. Twelve longer edged radial spines; eight opposite in pairs in two
crossed diagonal planes (as in the foregoing species), four others in the
lateral plane, opposite in pairs on both sides of the sagittal
constriction, which is scarcely twice as long as the hexagonal, in the
equatorial plane constricted medullary shell. Both equatorial cupolas
nearly of the same size as the four corner cupolas.

_Dimensions._--Length of the cortical shell 0.16, breadth 0.13; length of
the medullary shell 0.07, breadth 0.04.

_Habitat._--South Pacific, Station 300, depth 1375 fathoms.

Genus 303. _Zonidium_,[343] n. gen.

_Definition._--#Zonarida# with eight dome-shaped chambers of the cortical
shell, separated by four annular constrictions (one sagittal, one
transverse, and two others parallel to the latter).

The genus _Zonidium_ must be regarded as a _Zonarium_, in which the four
cupolas (in the quadrants of the lateral plane) are halved by diagonal
beams or latticed septa, {688}lying opposite in pairs in two crossed
diagonal planes. Therefore the number of the septa (four) and the domes
(eight) is here doubled. Among the eight cupolas we distinguish four median
(on both sides of the sagittal plane) and four lateral (on both sides of
the equatorial plane); both groups are of different size and form. Probably
_Zonidium_ is derived from _Zonarium_ by lattice-connection between the
eight diagonal wing-spines, which in both species of this genus are
present, the same as in _Tetrapyle octacantha_.

1. _Zonidium octostylium_, n. sp.

Cortical shell nearly quadrangular, with rounded corners and thorny
surface. Eight long and thin radial spines on both sides of the lateral
plane opposite in pairs and lying in two crossed diagonal planes (as in
_Tetrapyle octacantha_). Sagittal constriction three times as long as the
lentelliptical medullary shell. Four medial cupolas (on both sides of the
sagittal plane) somewhat larger than the four lateral cupolas (on both
sides of the equatorial plane).

_Dimensions._--Length of the cortical shell 0.15, breadth 0.12; length of
the medullary shell 0.05, breadth 0.03.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

2. _Zonidium octotholium_, n. sp. (Pl. 50, fig. 12).

Cortical shell nearly octangular, with spiny surface; twenty long and stout
radial spines between numerous smaller spines; eight wing-spines opposite
in pairs in two crossed diagonal planes (as in the preceding species);
twelve other strong spines in the lateral plane (four longer opposite in
pairs on the poles of the principal and transverse axes, eight others
smaller, alternating between these and the diagonal spines). Sagittal
constriction twice as long as the hexagonal medullary shell. Four median
cupolas somewhat smaller than the four lateral cupolas.

_Dimensions._--Length of the cortical shell 0.18, breadth 0.15; length of
the medullary shell 0.07, breadth 0.05.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

Family XXIX. #LITHELIDA#, Haeckel (Pl. 49, figs. 1-7).

_Lithelida_, Haeckel, 1862, Monogr. d. Radiol., p. 515.

_Definition._--#Larcoidea# with symmetrical spiral shell, divided by the
spiral plane into two symmetrical halves; all windings of the spiral lie in
this plane. Primordial chamber either simple or _Larnacilla_-shaped.

The family #Lithelida# comprises all those #Larcoidea# in which the growth
of the latticed shell is spirally winding in one plane, _Nautilus_-like.
They agree in the spiral growth with the following family. But in the
Streblonida the spiral is screw-shaped, {689}ascending (like _Helix_).
Therefore in these latter the geometrical fundamental form of the shell is
asymmetrical or "dysdipleural," whereas in the Lithelida
bilateral-symmetrical or "eudipleural." The lentelliptical or nearly
spherical shell may be divided by a median section into two symmetrical
halves; the right half is the mirror image of the left half.

When in 1862 I founded the family Lithelida in my Monograph (p. 515), I
knew only one genus, _Lithelius_, with two species. The rich material of
the Challenger collection contains a great number of similar spirally
constructed #Larcoidea#, so that at the present time we may distinguish at
least six genera. These belong to two different subfamilies, which may
possibly be afterwards better separated as families. The first subfamily,
Spiremida, possess a simple, spherical or subspherical, medullary shell;
the second subfamily, Larcospirida, possess a trizonal or
_Larnacilla_-shaped medullary shell. No doubt these latter must be derived
from Pylonida, as we observe all stages of development starting from a
simple _Trizonium_; but perhaps also the Spiremida have the same origin,
their simple, spherical or subspherical, medullary shell being derived from
a trizonal or _Larnacilla_-shaped medullary shell by reduction.

The general appearance in both subfamilies of the Lithelida is quite the
same, and it requires a careful study of the medullary shell to distinguish
certainly the Spiremida from the Larcospirida. This distinction is often
not easy, particularly in the larger forms; the shell is often very opaque
and difficult to understand. Only in one position, if the spiral axis be
parallel to the axis of the eye of the observer, and the spiral plane be
therefore fully seen in the optical plane of the microscope, the spiral
line (or the axial section of the latticed spiral lamella) is distinctly
observed; in all other positions the figure of the spiral is more or less
indistinct, and the whole microscopical image often quite intricate and
confused. The sufficient study of this family requires therefore the
contemplation of the shell from different sides, and is the more difficult,
as the variability of the Lithelida--as of the Pylonida--is extraordinarily
great.

The description which I gave of _Lithelius_ (1862) in my Monograph is in
some points erroneous, and was afterwards (1879) corrected by R. Hertwig,
who explained particularly the near relation of it to _Tetrapyle_. Indeed
the intermediate forms between the Lithelida and the Pylonida are so
numerous and so evident in all stages of development, that the derivation
of the former (at least of the Larcospirida) from the latter is quite
clear. The analogy between the structure of the Lithelida and the
calcareous (foraminiferous) Alveolinida is not so complete as I supposed it
to be in my Monograph (1862); particularly the formation of the small
chambers between the turnings of the spiral lamella is much more complete
in the Alveolinida than in the Lithelida.

The cortical shell of all Lithelida has the same geometrical fundamental
form as _Nautilus_ or as the nautiloid Polythalamia (_Polystomella_,
_Nummulites_, &c.); therefore the shell is dipleural, being divided by the
median plane into two symmetrical lateral halves. {690}Since the spiral
line lies in the median plane, we will call it the spiral plane; it
separates the right half from the left. The axis of the body, around which
the spiral turns (without touching it), is the spiral axis. The latticed
part of the cortical shell, which turns around them, is the spiral lamella.
Only in one genus of our family, viz., _Tholospira_, are the spiral axis,
the lateral axis, the spiral plane, and the sagittal plane quite as in
_Nautilus_. In all other genera this disposition is different or is
uncertain. This depends on the different part of the cortical shell, from
which the spiral growth begins. In this respect we can distinguish four
different modes.

In the Larcospirida (or the Lithelida with _Larnacilla_-shaped medullary
shell) the spiral growth exhibits four quite different forms. It begins
here with _Larcospira_, in which already the first cortical girdle of the
Diplozonaria determines the spiral growth; one wing of this girdle, the
transverse girdle of _Amphipyle_, grows more swiftly than the other,
overgrows it, and thus turns around the principal axis. In _Pylospira_ the
first or transverse girdle is already perfectly formed (as in _Amphipyle_),
and the spiral growth is introduced by the second or lateral girdle of
_Tetrapyle_; one wing of it (the right or the left) grows more swiftly than
the other, overgrows it, and thus turns around the sagittal axis. In
_Tholospira_ also the second girdle is complete, and the spiral growth
begins from the third or sagittal girdle. One of its wings grows more
swiftly than the other, overgrows it, and thus turns around the transverse
axis. Consequently we see that each of the three dimensive planes of the
lentelliptical Larcoid-body may be the spiral plane: in _Larcospira_ the
transverse plane, in _Pylospira_ the lateral plane, in _Tholospira_ the
sagittal plane. Correspondingly the spiral axis in the first genus is the
principal, in the second the sagittal, in the third the transverse axis of
the central _Larnacilla_-shell. Therefore in these three genera the spiral
plane is the plane of the latticed girdle, which determines the spiral
growth, one of both its wings overgrowing the other.

In each of the three above mentioned genera the spiral may be simple or
double; it remains simple if only one of both wings of the turning girdle
overgrow the other, and this latter remain a simple half girdle (or
tube-like wing). Whereas the spiral becomes double if the second wing of
the girdle afterwards follow the example of the first wing and now turn
around it in the same direction. As this happens in all three genera, we
can subdivide them into six subgenera.

A quite peculiar form of spiral growth is produced in _Spironium_, in which
the direction of growth in both lateral wings of the transverse girdle is
inverse from the beginning. The left wing grows against the posterior, the
right wing against the anterior pole of the principal axis, turning around
it in crossed, eight-like spirals. The whole shell afterwards assumes a
lentelliptical form.

Commonly between the embracing spiral turnings or convolutions a great
number of radial beams is developed, irregularly disposed and often
branching; they support the {691}thin spiral lamellæ and give to the whole
shell more solidity. Often these beams form imperfect radial septa, by
which the spiral cavity of the turnings is divided into a variable number
of chambers. But these chambers never become so regular and perfect as in
the analogous nautiloid Polythalamia.

In many Lithelida the growth of the shell reaches a certain limit,
concluding with the formation of a superficial latticed lamella of
lentelliptical or nearly spherical form. In many other forms of the family
this seems not to be the case; but these may possibly be younger forms,
afterwards reaching the same limit.

The network of the shell in the Lithelida is commonly quite irregular, and
so variable that its special conformation has usually no value in the
determination of the species. The surface of the shell is often covered
with radial spines, which are sometimes arborescent.

The central capsule seems always to preserve the same lentelliptical form
(or triaxial ellipsoid) as in all other #Larcoidea#. With the increase of
growth it encloses successively a larger part of the spiral cortical shell,
but on the outside is constantly protected by the last turnings of the
spiral, or by the lattice-lamella of the surface.

_Synopsis of the Genera of Lithelida._

-------------------------------------------------------------------------
I. Subfamily Spiremida.
Central medullary shell simple, spherical or lentelliptical.
-------------------------------------------------------------------------
Surface of the cortical shell smooth or thorny,
without radial spines, 304. _Spirema_.

Surface of the cortical shell covered with numerous
simple or branched radial spines, 305. _Lithelius_.
-------------------------------------------------------------------------
II. Subfamily Larcospirida.
Central medullary shell double, trizonal or _Larnacilla_-shaped.
-------------------------------------------------------------------------
The transverse girdle turns around the principal axis, 306. _Larcospira_.

The lateral girdle turns around the sagittal axis, 307. _Pylospira_.

The sagittal girdle turns around the transverse axis, 308. _Tholospira_.

Both wings of the transverse girdle turn around the
principal axis in an opposite diagonal direction, 309. _Spironium_.

Subfamily 1. SPIREMIDA, Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Lithelida# with simple, spherical or subspherical,
medullary shell.

Genus 304. _Spirema_,[344] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Lithelida# with simple, spherical or subspherical,
medullary shell, and lentelliptical or subspherical, spirally constructed
cortical shell; surface smooth or thorny, without radial spines.

{692}The genus _Spirema_ begins the series of the Spiremida, or of those
Lithelida in which the medullary shell presents a simple latticed sphere or
ellipsoid, never composed of a double, trizonal or _Larnacilla_-shaped
shell. In the present state of our knowledge we cannot say whether this
simple medullary shell be a primary formation, or effected by secondary
means, by reduction of a double _Larnacilla_-shaped medullary shell, which
is constantly found in the Larcospirida. The species of this genus (as of
all Lithelida) are difficult to distinguish, are transformistic, and
incline very much to variations and abnormalities. The spiral may be simple
or double.

Subgenus 1. _Spiremarium_, Haeckel.

_Definition._--Spiral convolutions of the cortical shell simple.

1. _Spirema lentellipsis_, n. sp.

Cortical shell lentelliptical, with smooth surface; proportion of its three
dimensive axes = 4 : 5 : 6. In the median plane are visible four perfect
turnings of the simple spiral, the breadth of which gradually increases
towards the third convolution, finally decreasing; the broadest (third)
turning three times as broad as the simple spherical medullary shell.

_Dimensions._--Length of the lentelliptical cortical shell 0.18, breadth
0.15, height 0.12; diameter of the spherical medullary shell 0.02.

_Habitat._--North Atlantic, Station 353, surface.

2. _Spirema melonia_, n. sp. (Pl. 49, fig. 1).

Cortical shell nearly spherical, with smooth surface; proportion of its
three dimensive axes = 1.4 : 1.5 : 1.6. In the median plane are visible
three perfect turnings of the simple spiral, all of the same breadth as the
simple spherical medullary shell; the breadth of each convolution somewhat
greater at the poles of the principal than at the poles of the sagittal
axis.

_Dimensions._--Length of the shell 0.16, breadth 0.15, height 0.14;
medullary shell 0.018.

_Habitat._--Pacific, central area, Station 271, surface.

3. _Spirema flustrella_, Haeckel.

_Flustrella haliomma_, Ehrenberg (1861), Abhandl. d. k. Akad. d. Wiss.
Berlin, 1872, p. 293, Taf. ii. fig. 6.

Cortical shell egg-shaped, with thorny surface; proportion of its three
axes = 4 : 5 : 6. In the median plane are visible three perfect turnings of
the simple spiral, the first and second of about the same breadth as the
simple spherical medullary shell, the third suddenly increasing, and
finally three to four times as broad. Network of the surface irregular,
with roundish pores.

_Dimensions._--Length of the shell 0.2, breadth 0.17, height 0.14;
medullary shell 0.013.

_Habitat._--North Atlantic, Greenland, 1000 fathoms, Ehrenberg; Færöe
Channel, John Murray.

{693}Subgenus 2. _Spiremidium_, Haeckel.

_Definition._--Spiral convolutions of the cortical shell double.

4. _Spirema diplospira_, n. sp.

Cortical shell lentelliptical, with smooth surface; proportion of its three
dimensive axes = 6 : 7 : 8. In the median plane are visible three perfect
turnings of a double spiral, the breadth of which gradually increases; the
broadest (third) convolution three times as broad as the simple
lentelliptical medullary shell.

_Dimensions._--Length of the shell 0.24, breadth 0.21, height 0.18;
medullary shell 0.02.

_Habitat._--Tropical Pacific, Philippines, Station 200, depth 250 fathoms.

5. _Spirema subglobosum_, n. sp.

Cortical shell nearly spherical, with thorny surface; proportion of its
three axes = 2 : 2.1 : 2.2. In the median plane are visible two perfect
turnings of a double spiral, the breadth of which in the second convolution
is four times as great as that of the first convolution and the simple
spherical medullary shell.

_Dimensions._--Length of the shell 0.22, breadth 0.21, height 0.2;
medullary shell 0.02.

_Habitat._--Tropical Atlantic, off Sierra Leone, Station 348, depth (2450)
fathoms.

Genus 305. _Lithelius_,[345] Haeckel, 1862, Monogr. d. Radiol., p. 519.

_Definition._--#Lithelida# with simple, spherical or subspherical,
medullary shell, and lentelliptical or subspherical, spirally constructed
cortical shell; surface covered with numerous, simple or branched, radial
spines.

The genus _Lithelius_, founded by me in 1862, and represented by two
Mediterranean species, was at that time the only known form of this family,
which now contains six genera and twenty-seven species. It differs from the
foregoing _Spirema_ in the possession of numerous radial spines on the
surface. These may be either simple or branched. The spiral may be simple
or double, and according to this latter modification we distinguish two
different subgenera.

Subgenus 1. _Lithospira_, Haeckel.

_Definition._--Spiral convolutions of the cortical shell simple.

1. _Lithelius spiralis_, Haeckel.

_Lithelius spiralis, Haeckel_, 1862, Monogr. d. Radiol., p. 519, Taf.
xxvii. figs. 6, 7.

Cortical shell lentelliptical, one and a third times as long as broad,
covered with very numerous (one hundred to one hundred and fifty or more)
simple, bristle-shaped radial spines, about as long {694}as the shell.
Spiral turnings simple, all nearly of the same breadth and scarcely broader
than the simple spherical medullary shell.

_Dimensions._--Length of the cortical shell (with six spiral convolutions)
0.15, breadth 0.13; diameter of the medullary shell 0.012.

_Habitat._--Mediterranean, Messina, Haeckel, surface; Atlantic, Stations
348 to 353, surface.

2. _Lithelius primordialis_, R. Hertwig.

_Lithelius primordialis_, R. Hertwig, 1879, Organism. d. Radiol., p. 54,
Taf. vi. figs. 4, 4_a_.

Cortical shell subspherical, covered with numerous simple, bristle-shaped
radial spines, longer than the shell. Spiral turnings simple, with
gradually increasing breadth, so that the beginning of the third spiral is
twice as broad as the first and as the simple spherical medullary shell.

_Dimensions._--Diameter of the cortical shell (with two spiral
convolutions) 0.12; diameter of the medullary shell 0.02.

_Habitat._--Mediterranean, Messina, R. Hertwig, surface.

3. _Lithelius alveolina_, Haeckel, 1862.

_Lithelius alveolina_, Haeckel, 1862, Monogr. d. Radiol., p. 520, Taf.
xxvii. figs. 8, 9.

Cortical shell spherical, covered with simple, very numerous (two hundred
to three hundred or more), short, bristle-shaped radial spines, scarcely
half as long as the radius of the shell. Spiral turnings simple, with
gradually increasing breadth, so that the beginning of the third spiral is
three times as broad as the first and as the simple spherical medullary
shell.

_Dimensions._--Diameter of the cortical shell (with four spiral
convolutions) 0.2; diameter of the medullary shell 0.01.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, on many
Stations, surface.

4. _Lithelius capreolus_, n. sp.

Cortical shell lentelliptical, one and a fourth times as long as broad,
covered with numerous (eighty to one hundred and twenty or more) short,
branched radial spines, about half as long as the radius of the shell; each
spine once or twice dichotomous, with curved divergent branches. Spiral
turnings simple, with gradually increasing breadth, so that the beginning
of the third spiral is twice as broad as the first, and as the simple
spherical medullary shell.

_Dimensions._--Length of the cortical shell (with four spiral convolutions)
0.18, breadth 0.14; diameter of the medullary shell 0.015.

_Habitat._--Pacific, central area, Station 274, surface.

Subgenus 2. _Drymospira_, Haeckel, 1881, Prodromus, p. 464.

_Definition._--Spiral convolutions of the cortical shell double.

{695}5. _Lithelius solaris_, n. sp. (Pl. 49, fig. 2).

Cortical shell spherical, covered with simple, very numerous (two hundred
to three hundred) bristle-shaped radial spines, longer than the diameter of
the shell. Spiral turnings double, both of the same breadth, gradually
increasing with the growth of the shell and several times surpassing the
diameter of the simple spherical medullary shell. (Pl. 49, fig. 2, exhibits
only the first convolutions in the centre of the shell.)

_Dimensions._--Diameter of the cortical shell (with four spiral
convolutions) 0.18; diameter of the medullary shell 0.02.

_Habitat._--Pacific, central area, Stations 266 to 272, surface and in
various depths.

6. _Lithelius arborescens_, n. sp.

Cortical shell lentelliptical, one and a half times as long as broad,
covered with numerous (fifty to eighty or more) branched radial spines,
about as long as the greatest diameter of the shell; each spine with two to
four lateral branches, which are again branched or dichotomous. Spiral
turnings double, both of little different breadth, which increases
considerably with the growth of the shell, so that the third turn is four
times as broad as the simple spherical medullary shell.

_Dimensions._--Length of the cortical shell (with three spiral
convolutions) 0.18, breadth 0.12; diameter of the medullary shell 0.01.

_Habitat._--North Atlantic, Færöe Channel, surface, John Murray.

Subfamily 2. LARCOSPIRIDA, Haeckel.

_Definition._--#Lithelida# with double, trizonal, or _Larnacilla_-shaped
medullary shell.

Genus 306. _Larcospira_,[346] n. gen.

_Definition._--#Lithelida# with double, trizonal, or _Larnacilla_-shaped
medullary shell; cortical shell subspherical or lentelliptical, constructed
of a simple or double spiral of the transverse girdle (or primary cortical
girdle); the spiral lamella revolving around the principal axis.

The genus _Larcospira_ begins the interesting series of the Larcospirida,
or of those Lithelida in which the medullary shell is formed by a trizonal
or _Larnacilla_-shaped lattice-shell, and the cortical shell by spiral
turnings of one of the three girdles, which compose the cortical shell of
the Pylonida. In _Larcospira_, as the oldest and most simple form of
Larcospirida, the spiral is formed by the transverse girdle, or the first
girdle of the Diplozonaria, the only cortical girdle of _Amphipyle_. If in
this genus one of both wings of the transverse girdle grow stronger than
the other and overgrow the latter, turning {696}around the principal axis,
we reach the characteristic form of _Larcospirema_, the first subgenus of
_Larcospira_; but if afterwards the second wing follow the example of the
first, and overgrow it from the other side, we reach the typical form of
the second subgenus, _Larcospironium_. In this latter subgenus the spiral
becomes double, whilst in the former it remains simple.

Subgenus 1. _Larcospirema_, Haeckel.

_Definition._--Spiral convolutions or turns of the cortical shell simple;
only one single wing of the transverse girdle turning around the principal
axis.

1. _Larcospira lentelliptica_, n. sp.

Cortical shell with smooth surface, four times as long as the
lentelliptical medullary shell. Perimeter of the lateral plane elliptical,
one and a half times as long as broad, without constrictions. One lateral
wing of the transverse girdle is more strongly developed and turns around
the other in two to three simple spiral turns.

_Dimensions._--Length of the cortical shell 0.2, breadth 0.16; length of
the medullary shell 0.05, breadth 0.04.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

2. _Larcospira quadrangula_, n. sp. (Pl. 49, fig. 3).

Cortical shell with thorny surface, three times as long as the
lentelliptical medullary shell. Perimeter of the lateral plane nearly
quadrangular, with four rounded corners (on the poles of two crossed
diagonal axes), one and a third times as long as broad, with one sagittal
constriction at the poles of the principal axis. One lateral wing of the
transverse girdle turns around the other in one and a half to two simple
spiral turns.

_Dimensions._--Length of the cortical shell 0.18, breadth 0.14; length of
the medullary shell 0.06, breadth 0.04.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

3. _Larcospira sexangula_, n. sp.

Cortical shell with spiny surface, four times as long as the hexagonal
medullary shell. Perimeter of the lateral plane nearly hexagonal, one and a
half times as long as broad, with six rounded corners (two on the poles of
the principal axis, four on the poles of two crossed diagonal axes), with
three slight ring-like constrictions. One lateral wing of the transverse
girdle turns around the other in two to two and a half simple spiral turns.

_Dimensions._--Length of the cortical shell 0.24, breadth 0.16; length of
the medullary shell 0.06, breadth 0.04.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{697}Subgenus 2. _Larcospironium_, Haeckel.

_Definition._--Spiral convolutions or turns of the cortical shell double;
both wings of the transverse girdle turning around the principal axis.

4. _Larcospira oliva_, n. sp.

Cortical shell with smooth surface, six times as long as the lentelliptical
medullary shell. Perimeter of the lateral plane elliptical, one and a
fourth times as long as broad, without constrictions. Both lateral wings of
the transverse girdle turn round one another and form one and a half to two
double spiral turns.

_Dimensions._--Length of the cortical shell 0.25, breadth 0.2; length of
the medullary shell 0.04, breadth 0.035.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Genus 307. _Pylospira_,[347] n. gen.

_Definition._--#Lithelida# with double, trizonal or _Larnacilla_-shaped
medullary shell; cortical shell subspherical or lentelliptical, constructed
of a single or double spiral of the lateral girdle (or second cortical
girdle); the spiral lamella revolving round the sagittal axis.

The genus _Pylospira_ follows after _Larcospira_ as the second genus of
Larcospirida; but in this latter the spiral of the cortical shell is formed
by the transverse girdle (or the first lattice-girdle of the Diplozonaria),
whilst in _Pylospira_ it is produced by the lateral girdle, or the second
lattice-girdle of that group. Therefore _Pylospira_ may be derived
phylogenetically from _Tetrapyle_ in the same manner as _Larcospira_ from
_Amphipyle_. Whilst in this latter the first cause of the spiral turning,
the unequal growth of both girdle-wings, proceeds from the transverse
girdle, in _Pylospira_ it proceeds from the lateral girdle. One of its
wings overgrows the other, turning around the sagittal axis. If the second
wing do not become developed, the spiral remains simple and represents the
subgenus _Pylospirema_; but if afterwards the second wing follow the
example of the first and overgrow it from the other side, we reach the
typical form of the second subgenus, _Pylospironium_, with a double spiral.

Subgenus 1. _Pylospirema_, Haeckel.

_Definition._--Spiral convolutions or turns of the cortical shell simple,
only one single wing of the lateral girdle turning around the sagittal
axis.

{698}1. _Pylospira tetrapyle_, n. sp.

Cortical shell with thorny surface, five times as long as the
lentelliptical medullary shell. Perimeter of the lateral plane elliptical,
one and a half times as long as broad. Four internal gates (as in
_Tetrapyle_, between the complete lateral wings of the transverse girdle)
kidney-shaped. One principal wing of the lateral girdle turns around the
other in one and a half to two simple spiral turns.

_Dimensions._--Length of the cortical shell 0.21, breadth 0.15; length of
the medullary shell 0.04, breadth 0.03.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

2. _Pylospira octopyle_, n. sp. (Pl. 49, fig. 4).

Cortical shell with thorny surface, three times as long as the
lentelliptical medullary shell. Perimeter of the lateral plane elliptical
or nearly hexagonal, with four rounded corners (on the poles of two crossed
diagonal axes), one and a third times as long as broad. Eight internal
egg-shaped gates (as in _Octopyle_), between the complete lateral wings of
the transverse girdle, and two axial beams in the principal axis. One
single wing of the lateral girdle turns around the other in two to three
simple spiral turns.

_Dimensions._--Length of the cortical shell 0.18, breadth 0.14; length of
the medullary shell 0.06, breadth 0.035.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

Subgenus 2. _Pylospironium_, Haeckel.

_Definition._--Spiral convolutions or turns of the cortical shell double;
both wings of the lateral girdle turning around the sagittal axis.

3. _Pylospira cymbium_, n. sp.

Cortical shell with smooth surface, seven times as long as the
lentelliptical medullary shell. Perimeter of the lateral plane elliptical,
one and a third times as long as broad. Four internal kidney-shaped gates
between the complete lateral wings of the transverse girdle (as in
_Tetrapyle_). Both principal wings of the lateral girdle turn round one
another in two to two and a half double spiral turns.

_Dimensions._--Length of the cortical shell 0.22, breadth 0.17; length of
the medullary shell 0.03, breadth 0.025.

_Habitat._--Indian Ocean, Madagascar, Rabbe, surface.

{699}Genus 308. _Tholospira_,[348] n. gen.

_Definition._--#Lithelida# with double, trizonal or _Larnacilla_-shaped
medullary shell; cortical shell subspherical or lentelliptical constructed
of a simple or double spiral of the sagittal girdle (or third cortical
girdle); the spiral lamella revolving around the transverse axis.

The genus _Tholospira_ represents the third genus of Larcospirida. Whilst
the spiral growth of the cortical shell is produced in _Larcospira_ by the
first (transverse) girdle of the Diplozonaria, in _Pylospira_ by the second
(lateral) girdle, in _Tholospira_ it is effected by the third or sagittal
girdle, which we found complete in _Pylonium_. If in this genus one wing of
the sagittal girdle overgrow the other remaining one and turn around the
transverse axis, we get _Tholospirema_, the first subgenus of our genus,
with simple spiral; but if afterwards the second wing follow the example of
the first, and overgrow it from the other side, we get _Tholospironium_,
with double spiral.

Subgenus 1. _Tholospirema_, Haeckel.

_Definition._--Spiral convolutions or turns of the cortical shell simple,
only one single wing of the sagittal girdle turning around the transverse
axis.

1. _Tholospira nautiloides_, n. sp.

Cortical shell with smooth surface, five times as long as the
lentelliptical medullary shell. Perimeter of the lateral plane elliptical,
one and a half times as long as broad. Four internal kidney-shaped gates
(between the lateral wings of the transverse girdle, as in _Tetrapyle_).
One wing of the sagittal girdle turns around the other in two to two and a
half simple spiral turns.

_Dimensions._--Length of the cortical shell 0.24, breadth 0.18; length of
the medullary shell 0.05, breadth 0.04.

_Habitat._--Indian Ocean, between Ceylon and Socotra, Haeckel, surface.

2. _Tholospira spinosa_, n. sp.

Cortical shell covered with numerous (sixty to eighty or more) simple,
bristle-like radial spines, about the length of the shell. Perimeter of the
lateral plane nearly quadrangular, with four rounded corners (on the poles
of two crossed diagonal axes), one and a half times as long as broad. Eight
internal egg-shaped gates (between the lateral wings of the transverse
girdle and two axial {700}beams in the principal axes, as in _Octopyle_).
One wing of the sagittal girdle turns around the other in two to three
simple spiral turns.

_Dimensions._--Length of the cortical shell 0.27, breadth 0.18; length of
the medullary shell 0.04, breadth 0.03.

_Habitat._--South Pacific, Station 288, surface.

3. _Tholospira dendrophora_, n. sp. (Pl. 49, fig. 6).

Cortical shell covered with numerous (forty to fifty or more) branched
radial spines, about half as long as the shell; each spine with two to six
dichotomous branches. Perimeter of the lateral plane elliptical, one and a
third times as long as broad. Eight internal egg-shaped gates, as in the
foregoing species. One single wing of the sagittal girdle turns around the
other in three to four spiral turns.

_Dimensions._--Length of the cortical shell 0.22, breadth 0.17; length of
the medullary shell 0.05, breadth 0.04.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Subgenus 2. _Tholospironium_, Haeckel.

_Definition._--Spiral convolutions of the cortical shell double; both wings
of the sagittal girdle turning around the transverse axis.

4. _Tholospira hystrix_, n. sp.

Cortical shell covered with numerous (sixty to eighty or more) simple
conical spines, about half as long as the shell. Perimeter of the lateral
plane hexagonal, one and a half times as long as broad. Four internal
kidney-shaped gates, as in _Tetrapyle_. Both wings of the sagittal girdle
turn round one another in two to two and a half double spiral turns.

_Dimensions._--Length of the cortical shell 0.21, breadth 0.14; length of
the medullary shell 0.04, breadth 0.03.

_Habitat._--North Atlantic, Station 354, surface.

5. _Tholospira cervicornis_, n. sp. (Pl. 49, fig. 5).

Cortical shell covered with numerous (forty to fifty or more) branched
radial spines; each spine antler-shaped, about as long as the medullary
_Larnacilla_-shell, with eight to twelve dichotomous branches (similar to
_Cromyodrymus abietinus_, Pl. 30, fig. 6). Perimeter of the lateral plane
elliptical, one and a third times as long as broad. Four internal
kidney-shaped gates, as in _Tetrapyle_. Both wings of the sagittal girdle
turn round one another in one and a half to two double spiral turns.

_Dimensions._--Length of the cortical shell 0.2, breadth 0.15; length of
the medullary shell 0.03, breadth 0.025.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{701}Genus 309. _Spironium_,[349] n. gen.

_Definition._--#Lithelida# with double, trizonal or _Larnacilla_-shaped
medullary shell; cortical shell subspherical or lentelliptical, constructed
of two crossed spirals, which arise from both lateral wings of one girdle
(commonly the lateral girdle) and revolve in an opposite diagonal direction
around the principal axis.

The genus _Spironium_ differs in a very remarkable manner from all
foregoing Lithelida, and is distinguished by a quite peculiar mode of
growth. It is most nearly allied to _Larcopyle_, and may, like this, be
derived from _Amphipyle_ (or rather from _Larnacilla_, beginning to
transform into _Amphipyle_). But whilst in _Larcopyle_ one of the two wings
of the lateral girdle overgrows the other in the direction of the
transverse axis (turning around the principal axis), here in _Spironium_
both lateral wings begin at the same time to grow out from the lateral
sides of the _Larnacilla_-shaped medullary shell; the most remarkable thing
is, however, that the direction of growth in the wings is diverse from the
beginning: the left wing grows downwards and turns around the lower
(aboral) pole of the principal axis, the right wing grows upwards and turns
around the upper (oral) pole of the same axis. Thus both wings of the
lateral girdle are crossed in diagonal axes, and with increasing growth one
overgrows the other in the direction of these diagonals, so as to resemble
the figure 8 in shape. The open gates remaining between the turnings of the
girdle become afterwards closed on the surface by irregular lattice-work,
and so the whole cortical shell assumes finally a spherical, ellipsoidal,
or lentelliptical form. Its surface sometimes becomes covered with simple
or branched radial spines. In the interior the eight characteristic
egg-shaped gates of _Octopyle_ are commonly (or constantly?) visible, two
strong radial beams in the principal axis arising from the poles of the
lentelliptical medullary shell.

Subgenus 1. _Spironetta_, Haeckel.

_Definition._--Surface of the shell smooth or rough, but not with radial
spines.

1. _Spironium octonium_, n. sp. (Pl. 49, fig. 7).

Cortical shell lentelliptical, its breadth (or transverse axis) surpassing
considerably the length (or the principal axis). Surface thorny and
somewhat hump-backed. The spiral wings of the transverse girdle have about
the same breadth as the eight internal gates between them and the axial
beams.

_Dimensions._--Length of the cortical shell (or principal axis) 0.15,
breadth (or transverse axis) 0.2; length of the medullary shell 0.05,
breadth 0.04.

_Habitat._--South Atlantic, west of Tristan da Cunha, Station 332, depth
2200 fathoms.

{702}2. _Spironium diagonale_, n. sp.

Cortical shell nearly spherical, four times as great as the subspherical
medullary shell. Surface rough. The spiral wings of the transverse girdle
about half as broad as the eight internal gates between them and the axial
beams.

_Dimensions._--Diameter of the cortical shell 0.16, of the medullary shell
0.04.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

Subgenus 2. _Spironilla_, Haeckel.

_Definition._--Surface of the shell covered with simple or branched radial
spines.

3. _Spironium spinosum_, n. sp.

Cortical shell subspherical, five times as great as the subspherical
medullary shell. Surface covered with numerous (sixty to eighty or more)
simple, bristle-like radial spines, longer than the shell. The spiral wings
of the transverse girdle of about the same breadth as the eight internal
gates between them and the axial beams.

_Dimensions._--Diameter of the cortical shell 0.2, of the medullary shell
0.04.

_Habitat._--Pacific, central area, Station 274, surface.

4. _Spironium arbustum_, n. sp.

Cortical shell lentelliptical, its breadth surpassing its length
considerably. Surface covered with numerous (forty to sixty or more) thin
radial spines, about as long as the greatest diameter of the shell; each
spine with two to six lateral branches, which are either simple or again
branched (similar to _Cromyodrymus abietinus_, Pl. 30, fig. 6). The spiral
wings of the transverse girdle only half as broad as the eight internal
gates between them and the strong beams of the principal axis.

_Dimensions._--Length of the cortical shell 0.12, breadth 0.15; length of
the hexagonal medullary shell 0.05, breadth 0.04.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Family XXX. #STREBLONIDA#, n. fam. (Pl. 49, figs. 8, 9).

_Definition._--#Larcoidea# with asymmetrical, spiral, polythalamous shell,
composed of a variable number of roundish chambers, which form together an
ascending spiral; both halves of the shell unequal. Primordial chamber
either simple or _Larnacilla_-shaped.

The family #Streblonida# comprises those #Larcoidea# in which a number of
chambers is arranged in an ascending spiral, round a simple or trizonal
primordial chamber, like winding stairs. They show the same spiral
structure as in the foregoing {703}family, but whilst in the Lithelida the
spiral line lies in one plane (as in _Nautilus_), in the Streblonida it
ascends like a screw (as in _Helix_). Therefore the former have the same
relation, regarding the spiral structure, to the nautiloid Polythalamia as
the latter to the turbinoid Foraminifera. Indeed the single forms of
Streblemida repeat in their special structure the characteristic genera of
Turbinoida, such as _Globigerina_, _Rosalina_, _Pulvinulina_,
_Hastigerina_, &c. As in these calcareous turbinoid Foraminifera, so also
in the analogous siliceous Streblonida the distinction of species is very
difficult and open to many objections.

The number of species in this family is very small; all are rare and for
the most part very opaque and difficult to understand, so that the
following distinction of a dozen species can have only a provisional value.
To get a complete idea of their peculiar structure, the shell must be
turned and observed from different sides, and thus their full study
requires yet much time and work. There are to be found evident transitional
forms between them and the Lithelida on the one hand and the Soreumida on
the other. Besides this, most species of Streblonida seem to have more
inclination to individual varieties and abnormalities than the majority of
the other Radiolaria.

The general form of the whole shell is in the Streblonida sometimes more
egg-shaped or even subspherical, at other times more top-like or conical,
sometimes nearly discoidal. The height of the shell (or the vertical axis
of the ascending spiral) is occasionally larger, at other times smaller
than the breadth (or the greatest horizontal diameter, perpendicular to the
height). Some very flat forms seem to approach the Lithelida. With regard
to the internal screw-formation, the shell of all Streblonida is
asymmetrical.

The number of the aggregated incomplete chambers is commonly between ten
and twenty, but ascends sometimes to thirty, forty, or more. Sometimes the
size of all the chambers is nearly the same, sometimes they increase
gradually, occasionally also very rapidly. The primordial chamber (or the
first and oldest) seems to be commonly the smallest, and inversely, the
last and youngest chamber, the largest. But sometimes (in _Streblopyle_)
also the contrary may be the case. The form of the single chambers is very
variable, from the spherical or hemispherical through all transitions
leading to irregular roundish or longish forms. The network is commonly
irregular, with small roundish pores of different sizes, but sometimes also
regular, circular. The surface of the shell is commonly smooth or rough,
rarely covered with radial spines. In most species the reticulation and
particularly the separation of the chambers is more or less incomplete.

As in the Lithelida, so also in the Streblonida we can distinguish two
subfamilies. In the Streblacanthida (_Streblonia_, _Streblacantha_) the
primordial chamber is a simple, spherical, subspherical, or lentelliptical
latticed shell. In the Streblopylida (_Streblopyle_) the primordial chamber
is trizonal or _Larnacilla_-shaped, as in the greater number of
#Larcoidea#, composed of three elliptical latticed girdles of unequal size,
perpendicular one to another, and surrounding a simple central chamber. As
in the Lithelida, so also {704}here we cannot certainly say whether the
former have originated from the latter by reduction of the
_Larnacilla_-shell, or whether both groups be of different origin. The
latter is perhaps more probable. This family as well as the foregoing
requires a much more careful study than I could give to it.

_Synopsis of the Genera of Streblonida._

-------------------------------------------------------------------------
I. Subfamily Streblacanthida.
Primordial chamber simple, spherical or lentelliptical.
-------------------------------------------------------------------------
Shell without radial spines, 310. _Streblonia_.

Shell with radial spines, 311. _Streblacantha_.
-------------------------------------------------------------------------
II. Subfamily Streblopylida.
Primordial chamber trizonal or _Larnacilla_-shaped.
-------------------------------------------------------------------------
Shell without radial spines, 312. _Streblopyle_.

Genus 310. _Streblonia_,[350] n. gen.

_Definition._--#Streblonida# with simple, spherical, subspherical, or
lentelliptical primordial chamber, beginning the screw-like series of
spirally ascending chambers. Surface smooth or thorny, without radial
spines.

The genus _Streblonia_ contains those Streblonida in which a variable
number of roundish, subspherical, or longish chambers form a screw-like
aggregate, beginning with a quite simple primordial chamber. The special
order of the complex spiral offers interesting resemblances to different
genera of the calcareous Foraminifera, from which I have taken the
corresponding name of the species. The whole form of the shell is sometimes
more egg-shaped or subspherical, at other times more top-like or conical,
occasionally very flat. Its surface is smooth or rough, but not covered
with radial spines.

1. _Streblonia globigerina_, n. sp.

Shell subspherical, thick walled, clustered, with eight to ten nearly
spherical chambers, of rapidly increasing size, the tenth chamber about
twelve times as broad as the first. Breadth of the shell nearly equal to
the height. Pores subregular, circular, hexagonally formed, of about the
same breadth as the bars; about sixteen on the breadth of the tenth
chamber. (Resembles very much the common _Globigerina_.)

_Dimensions._--Breadth of the shell 0.18, height 0.16.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

2. _Streblonia uvigerina_, n. sp.

Shell nearly egg-shaped, clustered, with eight to eleven subspherical
chambers of gradually increasing size, the tenth chamber about six times as
broad as the first. Breadth of the shell about {705}half its height. Pores
of the shell subregular, circular, about twice as broad as the bars; about
twenty on the breadth of the tenth chamber. (Resembles some species of
_Uvigerina_.)

_Dimensions._--Breadth of the shell 0.11, height 0.23.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

3. _Streblonia polymorphina_, n. sp.

Shell egg-shaped, thin walled, with twelve to fourteen roundish chambers of
rapidly increasing size, the tenth chamber about seven times as broad as
the first. Breadth of the shell about two-thirds of the height. Pores
irregular roundish, twice as broad as the bars; about twenty on the tenth
chamber. (Resembles certain forms of _Polymorphina_.)

_Dimensions._--Breadth of shell 0.22, height 0.14.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

4. _Streblonia bulimina_, n. sp.

Shell nearly egg-shaped, thick walled, clustered, with fourteen to eighteen
egg-shaped chambers of rapidly increasing size, the tenth chamber about
eight times as broad as the first. Breadth of the shell about two-thirds of
the height. Pores irregular, roundish, half as broad as the bars; about
twelve on the breadth of the tenth chamber. (Resembles closely _Bulimina_.)

_Dimensions._--Breadth of the shell 0.17, height 0.24.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

5. _Streblonia rosalina_, n. sp.

Shell top-shaped or flatly conical, with twelve to sixteen chambers of
gradually increasing size, the tenth chamber about four times as broad as
the first. Breadth of the shell twice as large as the height. Pores
subregular, circular, very small, of the same breadth as the bars; about
twelve on the breadth of the tenth chamber. (Resembles some forms of
_Rosalina_.)

_Dimensions._--Breadth of the shell 0.27, height 0.13.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

6. _Streblonia planorbulina_, n. sp.

Shell very flatly conical, nearly lenticular or discoidal, with twenty to
twenty-five chambers of nearly equal size, the tenth chamber a little
broader than the first. Breadth of the shell exceeds five to six times the
height. Pores subregular, circular, very small, half as broad as the bars;
about eight on the breadth of the tenth chamber. (Resembles closely
_Planorbulina_.)

_Dimensions._--Breadth of the shell 0.3 to 0.4, height 0.06 to 0.07.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

{706}7. _Streblonia pulvinulina_, n. sp.

Shell top-shaped or flatly conical, with thirty to forty chambers of slowly
increasing size, the tenth chamber about three times as broad as the first.
Breadth of the shell exceeds three times the height. Pores circular,
subregular, very small, about one-third as broad as the bars; about thirty
in the breadth of the tenth chamber. (Resembles closely _Pulvinulina_.)

_Dimensions._--Breadth of the shell 0.25, height 0.08.

_Habitat._--Pacific, central area, Station 263, depth 2650 fathoms.

Genus 311. _Streblacantha_,[351] n. gen.

The genus _Streblacantha_ differs from the nearly allied _Streblonia_ only
in the covering of radial spines, and bears therefore the same relation to
it as _Hastigerina_ has to _Globigerina_ amongst the similar calcareous
Polythalamia.

1. _Streblacantha siderolina_, n. sp. (Pl. 49, figs. 8, 8_a_).

_Streblonia siderolina_, Haeckel, 1883, MS.

Shell flatly conical, with fourteen to sixteen nearly hemispherical
chambers of gradually increasing size, the tenth chamber about six times as
broad as the first. Breadth of the shell nearly equal to the height. Pores
subregular, circular, hexagonally framed, twice as broad as the bars; about
nine pores on the breadth of the tenth chamber. Surface covered with
numerous short conical radial spines, one-fourth to one-sixth as long as
the diameter of the shell.

_Dimensions._--Breadth of the shell 0.15, height 0.17.

_Habitat._--South Pacific, Station 298, depth 2225 fathoms.

2. _Streblacantha calcarina_, n. sp.

Shell conical, with sixteen to eighteen roundish chambers of gradually
increasing size, the tenth chamber about three times as broad as the first.
Breadth of the shell about one and a half times the height. Pores
irregular, roundish. Surface covered with numerous strong conical radial
spines, about half as long as the diameter of the shell.

_Dimensions._--Breadth of the shell 0.24, height 0.17.

_Habitat._--South Pacific, Station 285, depth 2375 fathoms.

{707}3. _Streblacantha hastigerina_, n. sp.

Shell nearly spherical, clustered, with nine to eleven nearly spherical
chambers of rapidly increasing size, the tenth chamber about six times as
broad as the first. Breadth of the shell nearly equal to the height. Pores
subregular, circular, of about the same breadth as the bars. Surface
bristly, covered with numerous very thin and long, needle-shaped radial
spines, longer than the diameter of the shell. (Resembles closely
_Hastigerina_.)

_Dimensions._--Breadth of the shell 0.18, height 0.15.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 312. _Streblopyle_,[352] n. gen.

_Definition_.--#Streblonida# with trizonal lentelliptical medullary shell
(composed like _Larnacilla_ of three elliptical dimensive girdles
surrounding one simple central primordial chamber). From this begins a
screw-like series of spirally ascending chambers. Surface smooth or thorny,
without radial spines.

The genus _Streblopyle_ presents externally the same appearance and contour
as _Streblonia_, and is composed like this of a variable number of
chambers, ascending screw-like around the axis of the spiral shell. The
first or primordial chamber, however, in which the growth begins, is in
_Streblonia_ a simple spherical shell, but in _Streblopyle_ a trizonal
shell or _Larnacilla_-shell (compare above, p. 600). The chambers are very
incompletely separated, and comparatively much larger, their number much
smaller than in _Streblonia_. The structure in the species of this genus is
difficult to understand.

1. _Streblopyle helicina_, n. sp. (Pl. 49, fig. 9).

Shell helicoid, one and a third times as high as broad, with eight to
twelve incomplete semizonal chambers, ascending spirally from the lateral
half girdle of the lentelliptical medullary shell, octopyle-shaped, and
enveloping it in three to four spiral turnings. The height of the whole
cortical shell equals nearly five times the height of the trizonal
medullary shell. Pores irregular, roundish. Surface of the shell rough or
nearly smooth. (This species seems to be nearly allied to _Spironium
octonium_.)

_Dimensions._--Breadth of the spiral cortical shell 0.18, height 0.24;
breadth of the medullary shell 0.04, height 0.05.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Streblopyle spirulina_, n. sp.

Shell egg-shaped or nearly spherical, about as high as broad, with eight to
nine semizonal chambers, ascending spirally from the subspherical trizonal
medullary shell, and enveloping it in {708}four to five turnings. The sixth
chamber twice as broad as the trizonal medullary shell. Pores irregular,
roundish. Surface of the shell thorny.

_Dimensions._--Breadth of the spiral cortical shell 0.27, height 0.25;
breadth of the medullary shell 0.05, height 0.06.

_Habitat._--Pacific, central area, Station 265, depth 2900 fathoms.

Family XXXI. #PHORTICIDA#, Haeckel (Pl. 49, figs. 10, 11).

_Phorticida_, Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Larcoidea# with quite irregular monothalamous shell,
representing irregular modifications of an original lentelliptical latticed
shell; the irregular cortical shell encloses a regular or subregular,
lentelliptical or trizonal medullary shell.

The family Phorticida comprises a small number of #Larcoidea# in which a
subregular, trizonal, lentelliptical medullary shell is enclosed by an
irregular simple or spongy cortical shell. The lattice-work of the latter
is sometimes simple and complete, at other times incomplete, with open
gates (as in the Pylonida), sometimes also spongy. Its form is always more
or less irregular, roundish, often dimply or tuberous; different from most
other #Larcoidea#.

The medullary shell is constantly a regular or subregular
_Larnacilla_-shell, composed of three elliptical latticed girdles of
different sizes, perpendicular one to another. This leaves no doubt that
the Phorticida are true #Larcoidea#. The connection of it with the cortical
shell is rarely effected by radial beams, commonly by two opposite latticed
wings, which are identical with the lateral halves of the transverse girdle
in the Pylonida diplozonaria (_Amphipyle_, _Tetrapyle_). Often also between
this transverse and a second (lateral) girdle there remain large open
gates, so that the affinity between these Phorticida and the Pylonida
cannot be doubted. In other cases these gates become closed, so that they
more nearly approach the Larnacida. From both families they differ by the
irregularity of the papillate or tuberous cortical shell. The network is
more or less irregular, its surface often thorny, but never covered with
symmetrically disposed radial spines.

The central capsule is lentelliptical, encloses the medullary shell, and is
enveloped by the cortical shell, as in the nearly allied Pylonida and
Larnacida, of which the Phorticida may be regarded as irregular aberrant
forms.

_Synopsis of the Genera of Phorticida._

Cortical shell simply latticed, 313. _Phorticium_.

Cortical shell spongy, 314. _Spongophortis_.

{709}Genus 313. _Phorticium_,[353] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Phorticida# with irregular cortical shell of simple
lattice-work, enclosing a lentelliptical _Larnacilla_-shaped medullary
shell.

The genus _Phorticium_ comprises all Phorticida in which the irregular
cortical shell is formed by simple lattice-work, not by spongy framework.
We can divide this genus into two subgenera: in _Phortopyle_ (as in the
_Pylonida_) the lattice-work of the cortical shell exhibits large openings
or gates; in _Phortolarcus_ these gates are perfectly closed by network;
the former may be regarded as abnormal or irregular Pylonida, the latter as
modifications of Larnacida.

Subgenus 1. _Phortopyle_, Haeckel.

_Definition._--Lattice-work of the irregular cortical shell incomplete,
with large openings or gates.

1. _Phorticium pylonium_, n. sp. (Pl. 49, fig. 10).

Cortical shell irregular, roundish, about three times as large as the
enclosed lentelliptical, regular, _Larnacilla_-shell, connected with it by
some radial beams and irregularly latticed girdles; between these remain
four to eight large open gates of irregular roundish form and size; and
these gates are the same as in _Tetrapyle_ and _Octopyle_. This very
variable species may be regarded as a monstrosity of those genera of
Pylonida; it is very common, but all individuals are more or less unequal;
some specimens approach to some common species of _Tetrapyle_. The surface
of the shell is more or less spiny.

_Dimensions._--Diameter of the irregular cortical shell 0.12 to 0.18;
length of the lentelliptical medullary shell 0.05 to 0.06, breadth 0.035 to
0.045.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, &c., common,
surface and in various depths.

2. _Phorticium spironium_, n. sp.

Cortical shell irregular, roundish, tuberous, about four times as large as
the enclosed subregular _Larnacilla_-shell, connected with it by some
irregular radial beams, and by opposite, spirally begining, irregularly
latticed girdles, comparable to those of _Spironium_; between them remain
six to twelve large open gates of irregular size and form. Surface rough.
The resemblance to some forms of _Spironium_ makes it probable that this
species is a deformity or monstrosity of that genus.

_Dimensions._--Diameter of the irregular cortical shell 0.12 to 0.2, of the
lentelliptical medullary shell 0.03 to 0.06.

_Habitat._--Pacific, central area, Stations 270 to 274, surface, and in
various depths.

{710}Subgenus 2. _Phortolarcus_, Haeckel.

_Definition._--Lattice-work of the irregular cortical shell complete,
without large openings or gates.

3. _Phorticium deforme_, n. sp.

Cortical shell irregular, roundish or longish, three times as large as the
enclosed subregular, lentelliptical _Larnacilla_-shell, connected with it
by two opposite latticed wings (the halves of the transverse girdle of
_Tetrapyle_). Network of the cortical shell irregular, dense, perfectly
closed, without larger openings or gates. Surface thorny. (May be regarded
as a monstrous form of _Larnacalpis_.)

_Dimensions._--Diameter of the irregular cortical shell 0.15 to 0.18, of
the medullary shell 0.04 to 0.06.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

4. _Phorticium abnorme_, n. sp.

Cortical shell irregular, roundish, tuberous, with five to ten quite
irregular or nearly hemispherical protuberances, which resemble the cupolas
of Zonarida. The regular lentelliptical _Larnacilla_-shell is one-third to
one-fourth as large as the enclosing cortical shell, and is connected with
it by some irregular radial beams. Lattice-work completely closed, without
gates. Surface spiny. (May be regarded as an anomalous form of _Zonidium_;
as in the other species of this variable genus, the individuals are very
unequal.)

_Dimensions._--Diameter of the irregular cortical shell 0.12 to 0.2, of the
medullary shell 0.03 to 0.05.

_Habitat._--Atlantic and Pacific, tropical zone, surface, and at various
depths.

Genus 314. _Spongophortis_,[354] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Phorticida# with irregular cortical shell of spongy
framework, enclosing a lentelliptical _Larnacilla_-shaped medullary shell.

The genus _Spongophortis_ differs from _Phorticium_ in the spongy framework
of the cortical shell. This encloses the inner _Larnacilla_-shaped
medullary shell either directly, or both shells are separated by a hollow
interval, and connected either by radial beams or by latticed lamellæ.
Perhaps both these subgenera might be better separated as genera.

{711}Subgenus 1. _Stypophorticium_, Haeckel.

_Definition._--Spongy cortical shell immediately enclosing the
lentelliptical medullary shell, without hollow interval.

1. _Spongophortis spongiosa_, n. sp.

Cortical shell irregular, roundish, rough or tuberous, composed of loose
spongy framework, which immediately envelops the lentelliptical central
_Larnacilla_-shell; the diameter of the former becomes about five to six
times as large as that of the latter.

_Dimensions._--Diameter of the spongy cortical shell 0.15 to 0.25, of the
trizonal medullary shell 0.03 to 0.04.

_Habitat._--Pacific, central area, Station 274, surface.

Subgenus 2. _Spongophorticium_, Haeckel.

_Definition._--Spongy cortical shell separated by a hollow interval from
the lentelliptical medullary shell.

2. _Spongophortis radiosa_, n. sp.

Cortical shell irregular, roundish, four to five times as large as the
enclosed lentelliptical _Larnacilla_-shell, with which it is connected by
ten to twenty irregularly disposed radial beams. Spongy framework compact,
about as thick as the medullary shell. Surface covered with numerous short,
bristle-shaped, radial spines.

_Dimensions._--Diameter of the spongy cortical shell 0.15 to 0.2, of the
trizonal medullary shell 0.035 to 0.045.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

3. _Spongophortis larnacilla_, n. sp. (Pl. 49, figs. 11_a_-11_d_).

Cortical shell irregular, roundish, tuberous, three to four times as large
as the enclosed lentelliptical _Larnacilla_-shell, connected with it by two
opposite latticed wings (the halves of the transverse girdle of
_Tetrapyle_). Spongy framework compact, about half as thick as the
medullary shell. Surface rough. (May be regarded as an abnormal _Tetrapyle_
or _Larnacalpis_, with an irregular spongy cortical shell.)

_Dimensions._--Diameter of the spongy cortical shell 0.16 to 0.2, of the
trizonal medullary shell 0.04 to 0.06.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{712}Family XXXII. #SOREUMIDA#, Haeckel (Pl. 49, figs. 12, 13).

_Soreumida_, Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Larcoidea# with quite irregular polythalamous shell,
composed of a variable number of chambers, aggregated without any definite
order. Primordial chamber either simple or _Larnacilla_-shaped.

The family #Soreumida# contains a small number of #Larcoidea#, different
from most other #Sphærellaria# in the complete irregularity of the
polythalamous shell, which is composed of a variable number of roundish
chambers or subspherical latticed shells, aggregated in the form of an
irregular heap. We can distinguish in this family only two genera, with
very different structure of the central medullary shell or the first
chamber beginning the growth; and these correspond to the two subfamilies
of the nearly allied Streblonida (p. 704). In _Soreuma_ (as in
_Streblonia_) the first or primordial chamber, from which the growth
begins, is like the others, a simple spherical or irregular roundish
lattice-shell. In _Sorolarcus_, however (as in _Streblopyle_), the first or
primordial chamber is a trizonal or _Larnacilla_-shell. It is not
improbable that the former originated phylogenetically from _Streblonia_,
the latter from _Streblopyle_, by loss of the original spiral order of
growth. But it is also possible that these groups have no nearer relation.
Among the calcareous Foraminifera a very similar form is represented by
_Acervulina_ and its allies.

The general form of the whole shell in the Soreumida is sometimes more
egg-shaped or lentelliptical, at other times even subspherical,
occasionally quite irregular, tuberous, or clustered. The number of the
aggregated chambers is very variable, in _Sorolarcus_ between ten and
thirty, in _Soreuma_ ascending to fifty to eighty, sometimes from one
hundred to one hundred and fifty and more. Their size is sometimes nearly
equal, at other times very different, their form commonly very irregular,
roundish, but sometimes also subspherical or egg-shaped. The network of the
shell is also commonly irregular, with roundish pores of different sizes.
The surface is usually smooth or rough, rarely covered with radial spines.

The central capsule is not known, as I observed only a few skeletons of
this family.

_Synopsis of the Genera of Soreumida._

Primordial chamber of the shell simple, subspherical
or roundish, 315. _Soreuma_.

Primordial chamber of the shell trizonal or
_Larnacilla_-shaped, 316. _Sorolarcus_.

Genus 315. _Soreuma_,[355] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Soreumida# with numerous chambers, aggregated without any
regularity around one simple, spherical or subspherical, central chamber.

{713}The genus _Soreuma_ contains those Soreumida in which no trace of any
regular structure is found, but all the chambers of the irregular shell are
without any order, aggregated around a simple spherical or subspherical
central chamber or medullary shell. _Soreuma_ may have originated either
from _Sorolarcus_ by loss of the central _Larnacilla_-shell or from
_Cenolarcus_ by irregular apposition of new chambers around the
lentelliptical central chamber or simple Larcoid-shell. Some species seem
to exhibit a transition to _Sorolarcus_. Owing to the absolute irregularity
of the polythalamous shell _Soreuma_ resembles _Acervulina_ among the
Foraminifera.

Subgenus 1. _Soreumium_, Haeckel.

_Definition._--Shell without radial spines.

1. _Soreuma irregulare_, n. sp. (Pl. 49, fig. 12).

Shell irregular, clustered, or tuberous, composed of a large number (one
hundred and twenty to one hundred and fifty or more) of irregular, roundish
chambers of very different sizes, the largest four to five times as broad
as the smallest. Network very delicate, with very small roundish pores, to
five times as broad as the bars. Surface thorny.

_Dimensions._--Diameter of the shell 0.3, of the single chambers 0.02 to
0.08.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

2. _Soreuma acinosum_, n. sp.

Shell irregularly lentelliptical, with different growth in the three
dimensions, composed of a large number (forty to sixty or more) of
irregular, roundish chambers of very different sizes, the largest six to
eight times as broad as the smallest. Pores subregular, circular, twice as
broad as the bars. Surface smooth.

_Dimensions._--Length of the shell 0.21, breadth 0.17, height 0.13;
diameter of the largest chambers 0.03, of the smallest 0.004.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

3. _Soreuma subglobosum_, n. sp.

Shell nearly spherical, composed of a variable number (twelve to fifteen or
more) of irregularly aggregated subspherical chambers of nearly equal size.
Pores subregular, circular, twice as broad as the bars. Surface thorny.

_Dimensions._--Diameter of the shell 0.25, of the largest chambers 0.04, of
the smallest 0.005.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

{714}4. _Soreuma acervulina_, n. sp.

Shell quite irregular, cloddy, or tuberous, composed of twenty to thirty
(or more) roundish chambers of almost uniform size, the largest twice to
three times as broad as the smallest. Pores irregular, roundish. Surface
smooth.

_Dimensions._--Diameter of the shell 0.18 to 0.24, of the largest chambers
0.06, of the smallest 0.02.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

Subgenus 2. _Soreumidium_, Haeckel.

_Definition._--Shell with radial spines.

5. _Soreuma spinosum_, n. sp.

Shell quite irregular, cloddy, or tuberous, composed of thirty to forty
subspherical chambers of nearly the same size. Pores subregular, circular,
twice as broad as the bars; on the equator of each chamber six to eight
pores. Surface thorny, covered with irregularly scattered conical radial
spines, about as long as the diameter of the chambers.

_Dimensions._--Diameter of the shell 0.17 to 0.25, of the chambers 0.04.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

6. _Soreuma setosum_, n. sp.

Shell nearly spherical, composed of sixty to seventy (or more) irregular,
roundish chambers of very different sizes, the largest five to six times as
broad as the smallest. Pores irregular, roundish. Surface bristly, covered
with very numerous, long and thin, bristle-shaped radial spines, about as
long as the diameter of the shell.

_Dimensions._--Diameter of the shell 0.28, of the chambers 0.005 to 0.03.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

Genus 316. _Sorolarcus_,[356] n. gen.

_Definition._--#Soreumida# with numerous chambers, aggregated irregularly
around a trizonal medullary shell or _Larnacilla_-shell.

The genus _Sorolarcus_ comprises those Soreumida in which the heap of
irregularly aggregated chambers encloses a central trizonal medullary
shell, by which they demonstrate clearly their descent from Larnacida or
Pylonida. The lentelliptical medullary {715}shell exhibits quite the same
characteristic structure as that of _Larnacilla_, being composed of three
elliptical latticed girdles, perpendicular one to another. In some species
also the beginning of a second system of girdles is clearly indicated, so
that there can be no doubt as to their derivation from _Amphipyle_ or
_Tetrapyle_.

Subgenus 1. _Sorolarcium_, Haeckel.

_Definition._--Shell without radial spines.

1. _Sorolarcus larnacillifer_, n. sp. (Pl. 49, fig. 13).

Shell irregular, clustered, or tuberous, composed of twenty to thirty
irregular, roundish chambers of very different size, the largest four to
eight times as broad as the smallest, aggregated without order around a
central, lentelliptical, _Larnacilla_-shaped medullary shell. Pores
irregular, roundish, twice to four times as broad as the bars. Surface
smooth or a little spiny.

_Dimensions._--Diameter of the whole shell 0.18, of the central
_Larnacilla_-shell 0.05.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

2. _Sorolarcus tetrapylifer_, n. sp.

Shell irregularly roundish, clustered, composed of ten to twelve irregular
rather long chambers of almost uniform size, the largest twice as broad as
the smallest, aggregated without order around a central shell of the
structure of _Tetrapyle_, which encloses an inner trizonal
_Larnacilla_-shell of half the size. Pores irregular, roundish, twice to
four times as broad as the bars. Surface spiny.

_Dimensions._--Diameter of the whole shell 0.25, of the outer
(_Tetrapyle_-like) medullary shell 0.12, of the inner (_Larnacilla_-like)
shell 0.06.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

Subgenus 2. _Sorolarcidium_, Haeckel.

_Definition._--Shell with radial spines.

3. _Sorolarcus terminalis_, n. sp.

Shell nearly spherical, composed of fifteen to eighteen irregularly
aggregated roundish chambers of nearly equal size; in the centre a
lentelliptical _Larnacilla_-shell. Surface covered with numerous thin,
bristle-like radial spines, somewhat longer than the shell.

_Dimensions._--Diameter of the whole shell 0.21, of the central
_Larnacilla_-shell 0.05.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{716}----

LEGION II. #ACANTHARIA#,

VEL ACTIPYLEA, VEL ACANTHOMETREA (PLS. 129-140).

_Acantharia_, Haeckel, 1881.
_Actipylea_, Haeckel, 1882.
_Acanthometrea_, Hertwig, 1879.
_Panacantha_, Haeckel, 1878.

_Definition._--Radiolaria with simple membrane bounding the central
capsule, which is everywhere perforated by innumerable fine pores (disposed
either equally or symmetrically). Extracapsulum without phæodium. Skeleton
centrogenous (its growth proceeding from the centre), acanthinic (organic,
not siliceous). Fundamental form originally spherical.

The legion ACANTHARIA vel ACTIPYLEA, to the extent here defined, was
constituted by me, 1878, in my Protistenreich (p. 102) under the name
"PANACANTHA." A more accurate definition of this group was given in 1879 by
Hertwig under the name ACANTHOMETREA. Both names were replaced by me, 1881,
in my Prodromus (pp. 421, 465) by the more convenient name ACANTHARIA. This
legion comprises all those Radiolaria which were first described by
Johannes Müller, 1858, as _Acanthometrae_, and also an important part of
his _Haliomma_. In my Monograph (1862, pp. 371-424) I disposed them in
three families, Acanthometrida, Diploconida, and Dorataspida.

Although the number of genera and species in this legion is much increased
by the rich collection of the Challenger, we can divide all ACANTHARIA into
two different orders: #Acanthometra# (without complete lattice-shell) and
#Acanthophracta# (provided with a complete lattice-shell).

The ACANTHARIA agree with the SPUMELLARIA in the structure of the simple
capsule-membrane, which is perforated by numerous small pores (but
constantly devoid of the large main opening, which the NASSELLARIA and
PHÆODARIA possess, being hence united as "Merotrypasta"). We can therefore
unite both former legions as "Holotrypasta" (compare above, pp. 5, 6); but
in many ACANTHARIA (if not in all?) the numerous small pores of the
capsule-membrane exhibit a certain peculiar arrangement not observed in the
SPUMELLARIA; therefore the latter can be regarded as true "Peripylea" in
opposition to the former as "Actipylea."

The peculiar main character of all ACTIPYLEA or ACANTHARIA is determined by
the chemical constitution of their skeleton, which is not silex, but a
peculiar organic substance, called by me in 1862 "acanthin" (Monogr. d.
Radiol., pp. 30, 32). In all other Radiolaria the skeleton is composed of
silex or of a silicate. But besides this {717}chemical difference, an
important morphological character of the skeleton also separates the
ACANTHARIA from all other Radiolaria: in the latter the skeleton is never
centrogenous or arising from the centre of the capsule; in strict
opposition to this general fact the skeleton of all ACANTHARIA is
centrogenous, composed of radial spines, which arise from the central point
of the capsule and pierce its membrane. These characteristic "radial spines
of acanthin," arising from the centre, are never hollow (as formerly was
supposed), but constantly solid. Their form is extremely variable, and most
important for the distinction of genera and species; but more interesting
from a general point of view is their peculiar arrangement or disposition.

The regular disposition of twenty radial spines has general value almost
for all ACANTHARIA, with the exception only of the small group of
#Actinelida#. In this latter group the number of radial spines is either
more or less than twenty, and their disposition is either quite irregular
or follows a peculiar rule. The number of individuals of these
#Actinelida#, compared with that of the other ACANTHARIA, may be scarcely 1
per cent., whilst the latter have more than 99 per cent.; the number of
observed species is in the former about 5 per cent., in the latter about 95
per cent. Nevertheless the small group of #Actinelida# is very important,
being probably the ancestral group from which all other ACANTHARIA have
been phylogenetically derived. These other ACANTHARIA, with twenty
regularly disposed radial spines, represent the two large groups of
#Acanthonida# and #Acanthophracta#. For short and clear distinction of
these two groups of ACANTHARIA, we will call the #Actinelida# (with
irregular number and disposition of radial spines) Adelacantha, in
opposition to the Icosacantha (#Acanthonida# and #Acanthophracta#), which
all possess twenty regularly disposed radial spines.

Johannes Müller, the great zoologist, to whom we are indebted for the first
detection and accurate knowledge of the #Acanthometra#, already recognised
the regularity in the peculiar disposition of their twenty radial spines
(Abhandl. d. k. Akad. d. Wiss. Berlin, 1858, pp. 12, 37). In honour of my
great master I have called this regular disposition the "Müllerian law of
spine disposition," and have given a full explanation of it in my Monograph
(1862, pp. 40-45, 371, 372). With regard to its general value for all
Icosacantha (#Acanthonida# and #Acanthophracta#), we might also call this
promorphological Müllerian law "the Icosacanthan law."

In 1862 I had already given the following precise definition of this
"Icosacanthan law" (_loc. cit._, p. 40):--"Between two poles of a spineless
axis are regularly disposed five parallel zones, each with four radial
spines; the four spines of each zone are equidistant one from another, and
also equidistant from each pole; and the four spines of each zone are so
alternating with those of each neighbouring zone, that all twenty spines
together lie in four meridian planes, which intersect one another at an
angle of 45°." For the clear conception of this remarkable Müllerian law,
and for the complete understanding of its high value for the complicated
{718}morphology of all Icosacantha, it is the most profitable way to retain
constantly in mind for comparison the figure of a terrestrial globe with
its axis and zones. The axis of the globe is the spineless axis of all
Icosacantha, around which all twenty spines are symmetrically disposed; it
is perpendicular to the bisecting equatorial plane, in which lies the
middle of the five parallel zones; therefore the four spines, crossed
perpendicularly in this equatorial plane, are called the equatorial spines
(_c_1 to _c_4 in the figures of Pls. 131-140); often, and mainly in the
family Quadrilonchida (Pl. 131), these four equatorial spines are much
larger or of a peculiar form, different from that of the sixteen other
spines. Each pair of the four equatorial spines lies in one equatorial
axis, and this latter is perpendicular to the crossing axis, in which lies
the other pair of opposite spines. We may regard these two equatorial
diameters, perpendicular one to another and to the spineless axis, as the
two perradial axes or primary axes. Correspondingly the two meridian
planes, which are determined by one perradial axis and the spineless axis,
may be called the two primary or perradial meridian planes.

The globe is divided by the equatorial plane into two equal halves, the
northern and the southern hemisphere. In each hemisphere there are disposed
quite symmetrically eight radial spines, the distal ends of which fall in
two parallel circles, a larger tropical circle (nearer to the equator) and
a smaller polar circle (nearer to the pole of the spineless axis).
Therefore we call the four spines of the former the "tropical spines" and
the four spines of the latter the "polar spines." The angle between the
former and the equatorial plane is about 30°, the angle between the latter
and that plane about 60°.

The eight polar spines (four northern and four southern) lie in the same
two meridian planes as the four equatorial spines. Therefore in each of
these two perradial planes lie six radial spines, opposite in pairs; two
equatorial and four polar spines. Commonly all eight polar spines are of
the same size and form; and often they are also equal to the eight tropical
spines; but in some cases (e.g., in some species of Quadrilonchida) they
are much smaller than the twelve other spines, and sometimes even
rudimentary. In all figures of the Pls. 131-140 (and also in my Monograph,
1862, Taf. xv.-xxii.) the polar spines of the northern circle are marked by
the characters _a_1 to _a_4, the polar spines of the southern circle by the
characters _e_1 to _e_4. In the first perradial meridian plane lie _a_1 and
_a_3, _e_1 and _e_3, in the second _a_2 and _a_4, _e_2 and _e_4.

The eight tropical spines lie between the eight polar and the four
equatorial spines, four in each hemisphere; their distal points fall in two
parallel circles, which correspond exactly to the two tropics of the globe.
Therefore the four northern tropical spines may be called "canceral spines"
(as their ends fall in the Tropic of Cancer) and the four southern
correspondingly "capricornal spines" (as their points lie {719}in the
Tropic of the Capricorn). In the figures of the Pls. 131-140 (as well as in
my Monograph, 1862, Taf. xv.-xxii.) the four northern or canceral spines
are marked by the characters _b_1 to _b_4, and the four southern or
capricornal spines by the characters _d_1 to _d_4. Also the eight tropical
spines lie (crossed in pairs) in two meridian planes; they do not lie,
however, in those perradial planes, in which are placed the twelve other
spines; but in two different meridian planes, crossing the former at angles
of 45°; we call these the "secondary" or "interradial" meridian planes.
Each of these planes is determined by the spineless axis and by two crossed
interradial or secondary axes; in each of the latter lie two opposite
tropical spines. In the first interradial meridian plane lie _b_1 and _b_3,
_d_1 and _d_3, in the second _b_2 and _b_4, _d_2 and _d_4.

It is a most interesting and important fact, that in all Icosacantha
(#Acanthonida# and #Acanthophracta#) this regular disposition of the twenty
spines (in five parallel zones and four meridian planes) becomes constantly
preserved by heredity, whilst the form and size of the different spines are
extremely varied by adaptation.

Only in a minority of the Icosacantha are all twenty spines perfectly
equal or nearly equal in size and form; and then it is often very difficult
to distinguish the different zones in their disposition. But in far the
greater part the size or the form of the twenty spines becomes different in
different zones; and then we can commonly distinguish easily the five
different zones. Firstly, in all Quadrilonchida and Dorataspida, the four
equatorial are distinguished from the sixteen other spines either by form
or by size, and often in a very remarkable degree. As soon as these four
principal spines are recognised, it is easy to determine also the sixteen
others; for the eight polar spines lie in the same two (perradial) meridian
planes as the former, whilst the eight tropical spines lie in two different
(interradial) meridian planes, intersecting the two former at angles of
45°. Commonly, therefore, this distinction is rather easy.

In the majority of the Icosacantha all four equatorial spines are exactly
of the same form and size. But in four families the two opposite spines of
one equatorial axis are much larger, or of another form, than those of the
crossing axis. This is the case in the Amphilonchida, Belonaspida,
Hexalaspida, and Diploconida. Therefore we here call the major equatorial
axis (with larger spines) the "hydrotomical axis," and the minor axis (with
smaller spines) the "geotomical axis." Correspondingly, the meridian plane,
in which the two larger equatorial spines are placed (_c_1, _c_3) and the
appertaining four polar spines (_a_1, _a_3, _e_1, _e_3) may be called the
"hydrotomical plane"; in the remarkable family of Hexalaspida (Pl. 139) all
six spines of this hydrotomical plane are much larger than the other
fourteen. Perpendicular to this plane is the second perradial meridian
plane, which we call the "geotomical plane"; in it lie the two smaller
equatorial spines (c2, c4) and the corresponding four polar spines (_a_2,
_a_4, _e_2, _e_4). In some Hexalaspida (_Hexonaspis_ and _Hexacolpus_) the
six spines of the hydrotomical plane become so preponderant that {720}the
other fourteen spines appear rudimentary; and in some of them the two
equatorial spines of the hydrotomical plane are much larger than the four
polar spines of the same plane. This curious relation reaches its maximum
in the Diploconida (Pl. 140).

The different development of the two equatorial axes (of the larger
hydrotomical and the smaller geotomical axis) is the first and most
important cause of the peculiar forms, which are produced in the four cited
families. We derive these terms also from the metaphor of the terrestrial
globe. The hydrotomical plane is that meridian plane of the globe which
intersects almost only the water-hemisphere (the island of Ferro in the
Atlantic, the island of Pandora in the Pacific). Perpendicular to this is
the geotomical plane, the meridian of which intersects great land-masses in
both hemispheres (Bombay in India, Athabasca in Canada). Both poles of the
smaller geotomical axis are everywhere equal (the East Indian and the
Western American). However, both poles of the larger hydrotomical axis (the
eastern Atlantic and the western Pacific) are in some genera very
different, _e.g._, in _Amphibelone_ among the Amphilonchida, and in
_Zygostaurus_ among the Quadrilonchida. In this case we call the anterior
(commonly more developed) pole of the hydrotomical axis the frontal pole,
the opposite posterior (commonly smaller) the caudal pole (Pl. 131, figs.
7, 8; Pl. 132, figs. 9, 10). On both sides of these (right and left) lie
symmetrically the two equal poles of the geotomical lateral axis.

The promorphology of the ACANTHARIA demonstrates that the geometrical
fundamental form in those groups is different. In the majority of the
ACANTHARIA, where the two equatorial axes are equal, that form is a double
square-pyramid or a "quadrate octahedron"; the four equal equatorial spines
indicate the two diagonals of the square, which is the common base of the
united regular four-sided pyramids; their common axis is the spineless axis
of the body; the ends of the polar spines fall on the edges of the
pyramids, while the ends of the tropical spines fall on the halving lines
of their faces. However, in those ACANTHARIA in which the two equatorial
axes become different, the square double pyramid becomes changed into a
rhombic double pyramid; the common base of the united pyramids is thus a
rhombus; the hydrotomical axis is the larger, the geotomical axis the
smaller diagonal of the rhombus.

Opposed to the Icosacantha, under the name "Adelacantha," is the small
group of Actinelida, in which the number and disposition of the radial
spines is variable, not determined by the Müllerian law. Probably this
group is the common ancestral stock, from which the Icosacantha have been
derived by gradual development of their peculiar disposition. Probably the
oldest and most primitive form of all ACANTHARIA is _Actinelius_, in which
a variable and undetermined (often very large) number of radial spines is
united in one common central point, and therefore forms a needle-sphere.
Whilst here all spines (often more than a hundred) are of equal size and
form, in the nearly allied _Astrolophus_ large and small spines are
intermingled. Both genera together form the small ancestral family of
Astrolophida. In the strange family of Litholophida the radial spines do
not {721}radiate within a spherical space (equally disposed in all
directions), but within a quadrant or even an octant, forming a conical
brush or pencil.

One very remarkable form of #Actinelida# is _Actinastrum_, forming the
transition from these Adelacantha to the common regular Icosacantha. In the
two observed species of _Actinastrum_ we find thirty-two radial spines,
twenty of which are disposed after the Müllerian law, as in the
Icosacantha. The other twelve are four interradial equatorial spines (lying
in the two secondary meridian planes) and eight perradial tropical spines
(lying in the two primary meridian planes). Therefore here in each primary
meridian plane are placed ten spines (two equatorial, four tropical, and
four polar spines), whereas in each secondary meridian plane are placed six
spines (two equatorial and four tropical). But here also all thirty-two
spines are so regularly placed that their free distal ends fall into five
parallel zones, four in each polar zone, eight in each tropical zone, and
eight in the equatorial zone.

_The Central Junction_ of the radial spines in the ACANTHARIA becomes
effected in four different ways:--(1) by simple apposition of the pyramidal
central ends or bases; (2) by a basal leaf-cross, or by broad wings, four
on each spine, supported one upon the other; (3) by a central concrescence
of the meeting bases of all the twenty spines, growing perfectly together;
and (4) by a concrescence in pairs of every two opposite spines. The most
common and probably the original mode of junction is the first--by
pyramidal apposition; the spines at the central base are pointed in the
form of a pyramid, and the triangular faces of the neighbouring pyramids
are simply placed upon one another. Often the small basal pyramids are
imperfectly separated from the spines by an annular constriction. Commonly
the basal pyramids of the four equatorial spines are six-sided, those of
the sixteen other spines five-sided.

The second mode of junction, by a basal leaf-cross, is developed from the
first and appears as a strengthening or a mechanical elaboration of it.
Immediately above the basal pyramid arise from its radial edges four thin
and broad triangular leaves or wings, and the meeting edges of the
neighbouring wings are in apposition one with the other, so that between
the bases of every three or four neighbouring spines a hollow pyramidal
space remains open. The apex of such a pyramidal space is directed towards
the centre of the body, but separated from it by the small basal pyramid;
its open base is directed outwards. The twenty-two hollow pyramidal spaces
are disposed regularly in four different groups:--(A) Four equatorial
spaces, four-sided, each limited by two equatorial and two tropical spines
(one canceral and one capricornal); (B) eight perizonal spaces (four
northern and four southern), four-sided, each limited by one equatorial,
two tropical, and one polar spine; (C) eight peripolar spaces (four
northern and four southern), three-sided, each limited by one tropical and
two polar spines; (D) two polar spaces (one northern and one southern),
four-sided, each limited by four neighbouring polar spines.

{722}The third mode of junction, by central concrescence of all twenty
spines, was formerly regarded by me as an important peculiarity, sufficient
for the separation of subfamilies and genera (Monogr. d. Radiol., 1862, pp.
399, 401; Prodromus, 1881, p. 466). But I found afterwards that in many
species where the twenty spines commonly remain separated, accidentally
they grow perfectly together and form one single piece of acanthin--a
starrulet with twenty rays. Therefore I now think it is more natural to
divide those species only into different subgenera.

A fourth and a very different mode of junction, quite sufficient for the
distinction of different families, is the concrescence in pairs of every
two opposite spines, lying in one diameter (in _Acanthochiasma_ and
_Chiastolus_). Here we obtain a number of "diametral spines" (each composed
of two originally opposed radial spines) and all these diametral spines are
crossed loosely near the central point of the body without any solid and
permanent apposition (Chiastolida). However, in some species of this
peculiar family the central part of the diametral spines is twisted like a
screw or spirally convoluted (Pl. 129, figs. 2, 3).

_The Form of the Radial Spines_ in the ACANTHARIA is extremely varied, and
constitutes the main characters for the distinction of nearly four hundred
species. But all these different forms may be reduced phylogenetically to
three different fundamental forms:--(_a_) the cylindrical (with circular
transverse section), (_b_) the two-edged (with elliptical or lanceolate
transverse section), and (_c_) the four-edged (with square transverse
section). No doubt the first (_a_) is the original primitive form, from
which the two others are secondarily derived. Triangular spines never occur
in the ACANTHARIA, whilst, however, they are common in the #Sphærellaria#.
The first and original form, the cylindrical spine, is either a true
cylinder of equal thickness in its whole length, or it is more or less
conical. Rarely the spine is in the distal half spindle-shaped, and thicker
than in the basal half. The second form, the two-edged spine, is more or
less compressed from two opposite sides; its two edges are either more
blunt, rounded, or more acute, sharp; its transverse section in the former
case is elliptical, in the latter case lanceolate or rhomboidal. Sometimes
the two edges are broader and in the form of two thin opposite wings. The
two-edged spines may be occasionally shorter, triangular or lanceolate, at
other times longer sword-shaped or linear. The third form, the four-edged
spine, has constantly a square transverse section; the sides of this square
are either even or concave; in the latter case the four edges are broadened
and wing-like, but in the former case not. The quadrangular spines are
either prismatic (of equal breadth throughout their whole length) or
pyramidal (becoming gradually thinner towards the distal apex).

_The Apex of the Radial Spines_, or their free distal end, is in the
majority of ACANTHARIA simple, conical. In the minority it is either
truncated or bifid, or four-sided pyramidal, often with two, rarely with
four prominent parallel teeth. In some forms {723}the bifid spines are so
deeply cleft that they become forked. Much more interesting and more varied
than these different forms of the distal end are those of the apophyses of
the radial spines.

_The Apophyses of the Radial Spines_, or their "lateral transverse
processes," are of the greatest importance for the morphological
development of the whole subclass. Only in sixteen among the sixty-five
genera of ACANTHARIA are the apophyses perfectly wanting; in the other
genera they determine in the first place their general character. In the
#Acanthometra# the apophyses remain perfectly free, whilst in the
#Acanthophracta# their meeting ends or branches compose the latticed shell.
All differences in form and shape of the apophyses can be reduced to only
two primary modes; either the spine bears two opposite or four crossed
apophyses; correspondingly all Acantharia apophysaria may be divided into
two different main groups, the Zygapophysica (with two opposite lateral
processes) and the Staurapophysica (with four crossed lateral processes
opposite in pairs). Both groups have probably no direct phylogenetic
connection, but seem to be derived independently from different stocks, and
produce different families. The Zygapophysica are probably derived from
Astrolonchida with two-edged spines (_Zygacantha_), and from this group
arise the Diporaspida, the ancestral group of the majority of
#Acanthophracta#. On the other hand the Staurapophysica are probably
derived from Astrolonchida with four-edged spines (_Acanthonia_), and from
this group arise the Tessaraspida. The apophyses of the #Acanthonida# are
partly simple, partly branched or even latticed; the apophyses of the
#Acanthophracta# are never simple, constantly branched and commonly
latticed.

_The Malacoma_ (or the whole soft body of the ACANTHARIA as opposed to the
skeleton) exhibits some peculiarities which distinguish them from the other
Radiolaria, as well in the structure of the central capsule and its nucleus
as in that of the enveloping extracapsular body and the pseudopodia.

The _Central Capsule_ is constantly spherical in the far greater number of
the ACANTHARIA, viz., in the following six families:--Astrolophida,
Chiastolida, Astrolonchida, Dorataspida, Sphærocapsida, and Phractopeltida.
Among these six families the Astrolonchida and Dorataspida are far greater
and far richer in different forms than all the other families. The central
capsule becomes ellipsoidal or cylindrical, prolonged in one axis, in the
three families, Amphilonchida, Belonapsida, and Diploconida; it becomes
discoidal or lenticular, by the shortening of one axis, in two families,
viz., in the Quadrilonchida and Hexalaspida. Finally, the peculiar family
Litholophida is distinguished by the conical form of its central capsule.

_The Membrane_ of the central capsule in all ACANTHARIA is simple, commonly
thin, sometimes very delicate; in some species it seems to be developed
late, just immediately before the formation of the spores; but in no
species is it completely missing. The membrane is constantly pierced by
innumerable fine pores, for the emission of the {724}pseudopodia; but in
many species (and probably more or less in all ACANTHARIA) there is
recognisable a certain regularity in the disposition of the numerous
pseudopodia and of the pores by which they radiate from the capsule.
Sometimes these pores are disposed in a regular network of ramified lines,
whilst the meshes of this network are devoid of pores; in other cases they
form regular tufts or bushes between the radial spines. Probably in no
ACANTHARIA are the pores of the capsule membrane so numerous and so equally
distributed throughout as in the SPUMELLARIA; we may therefore call the
former ACTIPYLEA (in opposition to the latter, as PERIPYLEA).

_The Nucleus_ of the ACANTHARIA is constantly excentric, whilst it is
originally constantly central in the SPUMELLARIA. This excentric position
is a necessary consequence of the centrogenous development of the radial
spines. Probably connected with this peculiarity is the other, that the
nucleus assumes a peculiar, complicated structure, and that in the greater
number of ACANTHARIA it becomes cleft very early, and that this cleavage is
effected by a peculiar kind of gemmation, first detected and very
accurately described by R. Hertwig (compare his Organismus d. Radiol.,
1879, pp. 10-24). However, in the young ACANTHARIA the nucleus is
constantly simple, and in a certain number of species its cleavage takes
place late (as in the greater number of SPUMELLARIA).

_The Endoplasm_, or the intracapsular sarcode, exhibits in the greater
number of ACANTHARIA a more or less distinct radial arrangement; but this
is often concealed by the different enclosed products of the
endoplasm--oil-globules, vacuoles, red or different coloured
pigment-granules, crystals, &c. Often it encloses a variable number of
"yellow cells" (becoming green by mineral acids) to be considered as
symbiotic xanthellæ.

_The Calymma_ or the jelly-veil, including the central capsule, in the
ACANTHARIA is more or less voluminous, and commonly envelops the skeleton
perfectly. In its surface is sometimes developed a peculiar network of
"supporting fibres." A very peculiar product are the remarkable
"Myophrisca" of the #Acanthometra#, which are wanting in the
#Acanthophracta#; they were first detected by Johannes Müller, and figured
as "Cilien-Kränze," afterwards explained by Hertwig as "contractile Fäden,"
similar to muscular fibrillæ (compare below).

_The Matrix_, placed between the calymma and central capsule, in the
majority of the ACANTHARIA is a rather thin layer of granular exoplasm.

_The Pseudopodia_ arising from it are not so numerous as in the
SPUMELLARIA, and not so equally disposed over the whole surface. Also their
tendency to ramify, anastomose, and form networks seems to be much less
developed. Commonly they are simple or little ramified. In many cases (and
perhaps everywhere) there may be distinguished two different kinds of
pseudopodia:--(1) Axopodia, or permanent pseudopodia (with axial
filaments?), piercing the wall of the central capsule, and arising from the
central mass of endoplasm; and (2) Collopodia, or variable pseudopodia
(without axial filaments), arising outside the capsule from the matrix of
extracapsular sarcode or from the {725}exoplasm on the surface of the
calymma. These and other differentiations seem to indicate that the
pseudopodia in the ACANTHARIA are more highly developed than in the
SPUMELLARIA, and justify the denomination of the former as "Actipylea."

_Synopsis of the Orders and Suborders of_ ACANTHARIA.

-------------------------------------------------------------------------
I. ACANTHOMETRA.
Skeleton composed only of acanthinic radial spines not forming a
complete lattice-shell.
-------------------------------------------------------------------------
Radial spines in variable and indefinite number,
disposed irregularly, 1. #Actinelida#.

Radial spines constantly twenty, disposed regularly
after the Müllerian law of Icosacantha, 2. #Acanthonida#.
-------------------------------------------------------------------------
II. ACANTHOPHRACTA.
Skeleton composed of twenty acanthinic radial spines (disposed after
the Müllerian law) and of a spherical or variously shaped complete
lattice-shell.
-------------------------------------------------------------------------
Radial spines all twenty of equal size; shell and
central capsule spherical, 3. #Sphærophracta#.

Radial spines of different sizes; shell and central
capsule ellipsoidal, discoidal, or heteromorphous, 4. #Prunophracta#.

----

Order III. ACANTHOMETRA, Johannes Müller, 1855.

_Acanthometra_, J. Müller, 1855, Monatsber. d. k. preuss. Akad. d. Wiss.
Berlin.
_Acanthometrida_, Haeckel, 1862, Monogr. d. Radiol., p. 371.
_Acanthometrea_, R. Hertwig, 1879, Organismus d. Radiol., p. 133.
_Acanthonida et Litholophida_, Haeckel, 1881, Prodromus, pp. 465, 469.

_Definition._--ACANTHARIA without complete latticed shell.

The order #Acanthometra#, the third order of Radiolaria, comprises all
those ACANTHARIA in which the acanthinic skeleton is only composed of
radial spines arising from one common central point, but never forms a
complete latticed shell. By the absence of such a latticed or fenestrated
shell the #Acanthometra# differ principally from the nearly allied
#Acanthophracta#, the second order of ACANTHARIA, which constantly possess
such a complete shell.

Johannes Müller, who first detected and described the #Acanthometra# (in
1855, _loc. cit._), defined them as follows:--"Radiolaria without shell,
with siliceous radial spines" (1858, Abhandl. d. k. Akad. d. Wiss. Berlin,
p. 46). He described and figured eighteen species of them, disposed in four
genera (_Acanthometra_ with fifteen species, and _Zygacantha_,
_Lithophyllium_, _Lithoptera_, each with a single species). Among those
eighteen species, however, were two "Acanthometræ cataphractæ,"
appertaining to the following order, the #Acanthophracta#.

In my Monograph (1862, p. 371) all true #Acanthometra# were united into a
single family, Acanthometrida, with the following definition:--"Skeleton
composed of a number of radial spines, piercing the central capsule and
united in its centre, without {726}latticed shell." In the majority of them
I observed that the skeleton did not consist of silex, but of a very
peculiar organic substance, which I called "acanthin." At that time I
divided the family Acanthometrida into four subfamilies:--(1)
Acanthostaurida, (2) Astrolithida, (3) Litholophida, (4) Acanthochiasmida.
The two former now represent the suborder #Acanthonida#, the two latter the
suborder #Actinelida#. The number of genera which I distinguished in my
Monograph amounted to nine, the number of species to fifty. By the rich
collections of the Challenger this number is so much increased that we can
here describe twenty-seven genera and one hundred and sixty species.

Richard Hertwig in his work on the Organismus der Radiolarien (1879, pp.
6-25) adopted my family Acanthometrida, and gave a very accurate
description of its anatomical structure. He confirmed my observations that
the radial spines of this family are never hollow, but solid, and that
their chemical substance is not silex, but the organic matter "acanthin."
He found that the simple nucleus of the Acanthometrida is commonly very
early cleft, and that the peculiar brushes of filaments on the calymma,
described by Johannes Müller and by me as "Gallert-cilien," are peculiar
"contractile filaments," comparable to the "muscle-fibrillæ" of some
Infusoria, or the "Myophan-filaments" (Myophrisca).

The order #Acanthometra# is here divided into two different suborders of
very unequal extent and value, the #Actinelida# and #Acanthonida#. The
first may be regarded as the common ancestral stock, not only of the
second, but of all ACANTHARIA. In the small group of #Actinelida# the
number of radial spines is variable and commonly indefinite, often very
large (more than a hundred); they are therefore Adelacantha. The second
suborder, the #Acanthonida#, comprise by far the greatest part of the
order, and possess constantly twenty radial spines, regularly disposed
after the Müllerian law; they are therefore (like all #Acanthophracta#)
Icosacantha (compare above, p. 717).

The #Actinelida# possess constantly simple radial spines, without any
apophyses; their form is commonly very simple and primitive. This suborder
comprises three small but very different families, the Astrolophida,
Litholophida, and Chiastolida. The first family, the Astrolophida, is the
original ancestral group. A large and variable, commonly indefinite number
of radial spines is here united in the centre of the spherical central
capsule and radiating within a spherical space. In the second family, the
Litholophida, a small and variable number of radial spines (between ten and
twenty) is united in the apex of a conical central capsule and radiating
within the quadrant or octant of a spherical space. In the third family,
the Chiastolida, a variable number of radial spines is grown together by
pairs, in such a manner that every two opposite spines (placed originally
in one axis of the spherical central capsule) forms a single "diametral
spine"; all these diametral spines are not united in the centre of the
central capsule but only crossed loosely near the centre.

{727}The #Acanthonida#, the second suborder of #Acanthometra#, embraces by
far the greatest number in this order, viz., all those forms in which
twenty radial spines are regularly disposed after the Müllerian
law--Icosacantha (compare above, p. 717). The radial spines of this
suborder are either simple or provided with transverse processes (either
two opposite or four crossed apophyses). They are commonly united in the
middle of the central capsule by their opposed basal ends, forming small
pyramids; the meeting triangular faces of the neighbouring pyramids being
propped one upon another. Above these small basal pyramids often arises a
basal leaf-cross formed by four broad triangular leaves or wings with
straight edges; the meeting thin edges of the neighbouring spines serve for
strengthening the basal junction and form hollow pyramidal spaces or
compartments, filled with the contents of the central capsule (compare p.
721). The suborder #Acanthonida# comprises three different families, the
Astrolonchida, Quadrilonchida, and Amphilonchida. The first family, the
Astrolonchida, comprises by far the greater number of the #Acanthonida#;
those genera in which all twenty spines are perfectly equal or nearly equal
in size and form. In the second family, the Quadrilonchida, the four
equatorial spines are much larger (and often also of another form) than the
sixteen other spines (often also the eight tropical larger than the eight
polar spines). The third family, the Amphilonchida, is distinguished by the
preponderating development of only two opposite equatorial spines, which
are much larger (and often also of another form) than the eighteen other
spines.

_Synopsis of the Suborders and Families of_ #Acanthometra#.

-------------------------------------------------------------------------
Suborder I. ACTINELIDA.
Number of the radial spines variable, either more or less than twenty,
commonly disposed irregularly and not according to the Müllerian law.
-------------------------------------------------------------------------
Radial spines very numerous (thirty to a hundred
or more), radiating from a common centre
within a spherical space, 1. ASTROLOPHIDA.

Radial spines between ten and twenty, radiating
from one common point within a sphere-quadrant, 2. LITHOLOPHIDA.

Radial spines of variable number; every two
opposite spines grown together in the centre;
therefore numerous diametral spines are
crossed freely in the centre, 3. CHIASTOLIDA.
-------------------------------------------------------------------------
Suborder II. ACANTHONIDA.
Number of the radial spines constantly twenty, disposed regularly
according to the Müllerian law.
-------------------------------------------------------------------------
All twenty radial spines nearly equal, and of
the same size and form, 4. ASTROLONCHIDA.

Four equatorial spines much larger than (and
often also of different form from) the sixteen
other spines, 5. QUADRILONCHIDA.

Two opposite equatorial spines (or principal
spines) much larger than (and often also of
different form from) the eighteen other
spines, 6. AMPHILONCHIDA.

{728}----

Suborder I. ACTINELIDA, Haeckel, 1882.

_Definition._--#Acanthometra# with a variable number of radial spines,
which are commonly irregularly disposed, not according to the Icosacantha.

Family XXXIII. #ASTROLOPHIDA#, Haeckel.

_Astrolophida_, Haeckel, 1881, Prodromus, p. 469.

_Definition._--#Acantharia# with a variable number of simple radial spines,
radiating within a spherical space from one common central point which is
the centre of the spherical central capsule. No lattice shell.

The family #Astrolophida# comprises the simplest and the most primitive
forms among all ACANTHARIA, and may therefore be regarded as the common
ancestral stock of this whole legion or subclass of Radiolaria. The
acanthinic skeleton is composed of a variable number of quite simple radial
spines, which are united in the centre of the spherical central capsule and
radiate, piercing its walls and the surrounding jelly-veil, within a
spherical space.

The first observed form of this family is the ancestral genus _Actinelius_,
two different species of which I detected in 1864 in the northern
Mediterranean, at Villafranca, near Nice (compare Zeitschr. f. wiss. Zool.,
1865, Bd. xv. p. 364, Taf. xxvi. fig. 4). Three other species of the same
genus were afterwards found by me in the Challenger collections. Whilst in
this _Actinelius_ all radial spines are of the same size, a new nearly
allied genus, _Astrolophus_ (with two species), differs from it by the
different size of the radial spines, a small number of very large spines
being intermingled with a very large number of small spines. In these two
genera, _Actinelius_ and _Astrolophus_ (the true "Astrolophida" _sensu
strictiori_), the number of the radial spines is quite indeterminable and
their arrangement quite irregular and variable.

A third remarkable genus, _Actinastrum_, differs from these two genera in
the definite number and regular order of thirty-two radial spines, and may
therefore perhaps better represent a peculiar family, Actinastrida. In this
genus (of which two species were observed) the thirty-two radial spines are
disposed in such a regular manner that they lie in four meridian planes,
and that their distal ends fall into five parallel zones. These five zones
and these four planes are the same as we find in all Icosacantha (compare
above, p. 717). Also the constant twenty spines of these latter are present
in _Actinastrum_; but their number is here enlarged by twelve other spines
missing in the Icosacantha; four of these are secondary or interradial
equatorial spines, lying opposite in pairs between the four primary or
perradial equatorial spines; and eight are perradial tropical spines, lying
between the eight interradial tropical spines. Therefore the {729}distal
ends of the thirty-two radial spines are disposed regularly in five
parallel zones, and while two zones (the two polar) contain only the points
of every four spines, three zones (the single equatorial and the two
tropical) contain the points of every eight spines. The four meridian
planes are in _Actinastrum_ the same as in the Icosacantha, crossed in the
spineless axis at angles of 45°. But in the Icosacantha each of the two
perradial meridian planes contains six radial spines (two equatorial and
four polar), each of the two interradial meridian planes only four tropical
spines. Whereas in _Actinastrum_ each of the two primary or perradial
meridian planes contains ten spines (two equatorial, four tropical, and
four polar), each of the two secondary or interradial meridian planes six
spines (two equatorial and four tropical). We find therefore altogether
thirty-two radial spines in three orders; eight equatorial, sixteen
tropical, and eight polar spines.

Only one other genus of Radiolaria exhibits the same characteristic
disposition of thirty-two radial spines as _Actinastrum_, and this is
_Chiastolus_; but here the two opposite spines of each pair are grown
together and form one diametral spine; and the sixteen diametral spines are
crossed in the centre of the capsule. In _Actinastrum_, as in _Astrolophus_
and _Actinelius_, the central ends or bases of all the spines are
pyramidal, and the triangular faces of the neighbouring spines rest one
upon another (as in the greater number of #Acanthonida#). The form of the
radial spines in all Astrolophida is quite simple, without lateral
processes or apophyses; chiefly cylindrical, more rarely compressed,
two-edged or quadrangular.

The central capsule in all Astrolophida is spherical, and in the younger
specimens contains a single large concentric and lobed nucleus, but in the
older specimens a large number of small nuclei. The surrounding jelly-veil
or calymma seems commonly to envelop the spines perfectly. The piercing
pseudopodia radiate everywhere between the spines, and are very numerous
and thin. The circulating granules in them are sometimes red (_Actinelius
purpureus_).

_Synopsis of the Genera of Astrolophida._

Radial spines of indefinite {Spines of equal size, 317. _Actinelius_.
number and of irregular {
disposition. {Spines of unequal size, 318. _Astrolophus_.

Radial spines thirty-two, disposed regularly
in five parallel zones, 319. _Actinastrum_.

Genus 317. _Actinelius_,[357] Haeckel, 1865, Zeitschr. f. wiss. Zool., Bd.
xv. p. 364.

_Definition._--#Astrolophida# with a variable and undetermined number of
simple radial spines, all of equal size, united in the centre of the
spherical central capsule.

{730}The genus _Actinelius_ comprises the most simple and primitive forms
among all ACANTHARIA, and may be regarded as the common ancestral stock of
this whole legion. The spherical central capsule is pierced by numerous
simple radial spines of equal size, the pyramidal bases of which are
supported one upon another with their triangular faces in the centre of the
capsule. The number and position of the spines are quite indefinite and
variable. We may derive _Actinelius_ either from _Actissa_ (#Colloidea#) by
development of acanthinic radial spines, or directly from _Actinosphærium_
(Heliozoa) by formation of a central capsule.

Subgenus 1. _Actinelarium_, Haeckel.

_Definition._--Radial spines cylindrical, conical, or spindle-shaped, their
transverse section circular.

1. _Actinelius primordialis_, n. sp. (Pl. 129, fig. 1).

Spines sixty to eighty or more, cylindrical, at the distal end thickened,
spindle-shaped. Apex simple. Base a small slender pyramid. Central capsule
yellow. Granules of the sarcode colourless.

_Dimensions._--Length of the spines 0.3 to 0.4, breadth in the distal part
0.02, in the basal part 0.008.

_Habitat._--Central Pacific, Stations 265 to 274, surface.

2. _Actinelius purpureus_, Haeckel.

_Actinelius purpureus_, Haeckel, 1865, Zeitschr. f. wiss. Zool., Bd. xv.
p. 364, Taf. xxvi. fig. 4.

Spines thirty to forty or more, cylindrical, very thin, a little thinner
towards both ends. Apex simple. Base a small sulcate pyramid. Central
capsule opaque, purple. Granules of the sarcode also purple.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth 0.002.

_Habitat._--Mediterranean (Nice), Haeckel.

Subgenus 2. _Actinelidium_, Haeckel.

_Definition._--Radial spines compressed, two-edged; their transverse
section elliptical or lanceolate.

3. _Actinelius protogenes_, n. sp.

Spines fifty to sixty, compressed, two-edged, gradually broadened towards
the truncated distal end. Basal or proximal end thin, pyramidal. The spines
of this species are similar to those of {731}_Actinastrum pentazonium_ (p.
733) and of _Chiastolus amphicopium_ (Pl. 129, fig. 3), but much more
numerous, smaller, and not regularly disposed. These latter two Actinelida
must be separated on account of the regular disposition of the thirty-two
spines.

_Dimensions._--Length of the spines 0.2, basal breadth 0.008, distal
breadth 0.02.

_Habitat._--South Pacific, Station 165, surface.

Subgenus 3. _Actinelonium_, Haeckel.

_Definition._--Radial spines quadrangular, prismatic, or pyramidal, their
transverse section square.

4. _Actinelius pallidus_, Haeckel.

_Actinelius pallidus_, Haeckel, 1865, Zeitschr. f. wiss. Zool., Bd. xv.
p. 364.

Spines eighty to one hundred and twenty or more, quadrangular, prismatic,
of equal breadth throughout their whole length. Apex simple, truncate or
pyramidal. Base a four-sided slender pyramid. Central capsule pale
yellowish. Granules of the sarcode colourless.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth 0.005.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific.

5. _Actinelius polyacanthus_, n. sp.

Spines two hundred to three hundred or more, quadrangular, pyramidal,
gradually thinned towards the simple apex. Base a small three-sided
pyramid. Central capsule opaque.

_Dimensions._--Length of the spines 0.12 to 0.18, basal breadth 0.012.

_Habitat._--South Pacific, Station 291, surface.

Genus 318. _Astrolophus_,[358] Haeckel, 1881, Prodromus, p. 469.

_Definition._--#Astrolophida# with a variable and undetermined number of
simple radial spines of different sizes (large and small spines
intermingled), which are united in the centre of the spherical central
capsule.

The genus _Astrolophus_ differs from the nearly allied ancestral genus
_Actinelius_ only in the unequal size of the numerous radial spines. In
both observed species very numerous small spines are intermingled with a
small number of large spines, and between them numerous spines of medium
size. The small spines fill up the hollow spaces between the basal parts of
the large spines.

{732}1. _Astrolophus stellaris_, n. sp.

Radial spines from one hundred to two hundred, of very different sizes, but
of similar form; about sixteen to twenty very large spines, forty to fifty
of medium size, and one hundred to one hundred and twenty much smaller. All
spines cylindrical in the greater part of their length, with simple apex,
gradually thickened towards the central part, conical, without edges. The
base itself is a slender pyramid with four to eight edges.

_Dimensions._--Length of the largest spines 0.3 to 0.4, of the majority 0.1
to 0.2, of the smallest 0.05 to 0.1.

_Habitat._--South Pacific, Station 288, surface.

2. _Astrolophus solaris_, n. sp. (Pl. 132, figs. 12_a_, 12_b_).

Radial spines from two hundred to three hundred, of very different sizes,
but of similar form; about twenty to thirty very large spines, sixty to
eighty of medium size, and one hundred and twenty to one hundred and fifty
much smaller. All spines cylindrical in the greater part of their length,
with simple apex, gradually thickened and four-edged towards the central
base. The base itself is a slender pyramid with four to eight edges; partly
the faces, partly the edges of these basal pyramids rest one upon another,
the points of the larger spines meeting in the centre.

_Dimensions._--Length of the largest spines 0.4 to 0.5, of the majority 0.2
to 0.3, of the smallest 0.1 to 0.16; basal thickness of the largest spines
0.015.

_Habitat._--South-east Pacific (off Juan Fernandez), Station 296, surface.

Genus 319. _Actinastrum_,[359] n. gen.

_Definition._--#Astrolophida# with thirty-two simple radial spines,
regularly disposed within four meridian planes in such an order that their
distal ends fall into five parallel zones. Central ends of the thirty-two
spines supported one upon another in the centre of the spherical central
capsule.

The genus _Actinastrum_ differs from the two preceding genera in the
definite number and order of the thirty-two radial spines, which are
disposed in a very remarkable manner. Twenty radial spines are disposed
after the Müllerian law of Icosacantha (compare above, p. 717). The
remaining twelve spines are four equatorial spines lying in the two
secondary meridian planes, and eight tropical spines lying in the two
primary meridian planes. We have therefore together eight equatorial,
sixteen tropical, and eight polar spines (compare above, p. 729).

{733}1. _Actinastrum legitimum_, n. sp.

All thirty-two radial spines of equal size and similar form, cylindrical,
conical at the distal end, at the central base pyramidal. Central capsule
pellucid, colourless.

_Dimensions._--Length of the radial spines 0.3, breadth 0.004.

_Habitat._--South Pacific, Station 288, surface.

2. _Actinastrum pentazonium_, n. sp.

All thirty-two radial spines of equal size and similar form, compressed,
two-edged, gradually becoming broader and thinner from the pyramidal
central base towards the truncated distal end. Central capsule dark,
opaque. (Compare the similar _Chiastolus amphicopium_, Pl. 129, fig. 3.)

_Dimensions._--Length of the radial spines 0.2, breadth at the base 0.005,
at the distal end 0.02.

_Habitat._--South Pacific (west coast of Patagonia), Station 302, surface.

Family XXXIV. LITHOLOPHIDA, Haeckel.

_Litholophida_, Haeckel, 1862, Monogr. d. Radiol., p. 401.

_Definition._--ACANTHARIA with a variable number of simple radial spines
radiating within a conical space (or within the quadrant of a sphere) from
one common central point, which is the apex of the conical central capsule.
No lattice-shell.

The family #Litholophida#, represented only by a single genus,
_Litholophus_, differs from all other ACANTHARIA in the remarkable fact
that the common point, from which the radial spines arise, is not the
geometrical central point of the whole body, but is quite excentric in
position, the apex of the conical or pyramidal central capsule. Therefore
the spines form together a kind of brush or broom.

When I founded the family Litholophida in my Monograph (1862, p. 401) I
knew only a single species, _Litholophus rhipidium_, observed very
frequently in Messina. Another species, _Litholophus ligurinus_, was
afterwards (1864) found by me at Nice. Six other species were detected in
the preparations of the Challenger, some of them very frequent. All these
eight species of _Litholophus_ are very nearly allied, and exhibit only
slight differences in the form and number of the radial spines; their mode
of excentric connection and the structure of the peculiar soft body is
everywhere the same.

The radial spines in all observed Litholophida possess the form of the
genus _Acanthonia_, _i.e._, they are quite simple, four-sided prismatic or
quadrangular, with square transverse section; their four edges are
sometimes smooth, at other times elegantly denticulate, commonly more or
less prominent or wing-shaped. In the greater number of species they are
very long and of nearly equal breadth, prismatic; in some species they are
more pyramidal, thinned towards the distal end; the latter {734}is commonly
truncated or broken off, sometimes pyramidal. The central end is everywhere
thinned, more or less pyramidal, and the neighbouring spines are propped
one upon another by the triangular faces of their small basal pyramids. A
slight pressure is sufficient to destroy their connection.

The number and disposition of the radial spines seem to be variable and
irregular, but require further researches. In four of the observed eight
species I found constantly ten spines, in two other species from ten to
twenty (commonly twelve or sixteen), and in two species twenty or more. A
certain order or disposition of the spines within the conical space in
which they radiate could nowhere be ascertained.

When I first observed _Litholophus_, I supposed that it might only be a
mutilated or altered form of an _Acanthonia_. Afterwards, observing many
specimens with ten spines, I was led to the suggestion that they were
produced by self-division of an _Acanthonia_, and that the number of the
spines in each half of the body might be afterwards doubled. But this
suggestion seems to be refuted by the fact that in no other genus of the
numerous ACANTHARIA is self-division observed, and that many hundreds of
_Litholophus_ which I observed exhibit quite constantly only a single form
of radial spines, that of _Acanthonia_--simple quadrangular spines without
any apophyses.

Genus 320. _Litholophus_,[360] Haeckel, 1862, Monogr. d. Radiol., p. 401.

_Definition._--#Litholophida# with a variable number of quadrangular
diverging radial spines, united with pyramidal bases in the apex of the
conical central capsule.

The genus _Litholophus_, the only one of this family, exhibits the
peculiarities just described, but might more nearly be defined as a typical
"genus" by the quadrangular form of the radial spines, identical with those
of _Acanthonia_.

The central capsule of _Litholophus_ is constantly conical or pyramidal,
commonly opaque, of a dark brownish or reddish colour; it contains many
small nuclei. It envelops the basal half of all radial spines in such a
manner that their basal parts are united in its apex, and their distal
parts pierce the rounded base of the conical capsule (Pl. 129, fig. 2).

The calymma or the jelly envelope of the central capsule is only developed
at its base, where the spines radiate; at the conical mantle of the capsule
it is very thin. The spines seem to be perfectly enclosed in the calymma
and connected with it by the same contractile retinacula or "myophrisca"
which we observe in the #Acanthonida#. The pseudopodia arise only from the
rounded base of the conical capsule, and radiate between the spines,
piercing the calymma, diverging within the conical space occupied by the
fascicle of spines.

{735}Subgenus 1. _Litholopharium_, Haeckel.

_Definition._--Ten radial spines.

1. _Litholophus decimalis_, n. sp.

Ten radial spines, four-sided prismatic, with prominent smooth edges, of
equal breadth throughout their whole length.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth 0.006.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

2. _Litholophus pyramidalis_, n. sp.

Ten radial spines, four-sided pyramidal, with prominent smooth edges,
gradually thickened from the small pyramidal base towards the truncated
distal end.

_Dimensions._--Length of the spines 0.3 to 0.4, breadth in the basal part
0.002, in the middle part 0.006, in the distal part 0.012 to 0.02.

_Habitat._--Central Pacific, Station 266, surface.

3. _Litholophus decapristis_, n. sp. (Pl. 129, fig. 2).

Ten radial spines, four-sided prismatic, with prominent, elegantly
denticulated edges, of equal breadth in their whole length.

_Dimensions._--Length of the spines 0.2 to 0.4, breadth 0.008.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

4. _Litholophus decastylus_, n. sp.

Ten radial spines, four-winged pyramidal, with broad and thin, elegantly
denticulated edges, gradually thickened from the small pyramidal base
towards the truncated distal end.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth in the basal part
0.002, in the middle part 0.005, in the distal part 0.015.

_Habitat._--South Atlantic, Station 332, surface.

Subgenus 2. _Litholophidium_, Haeckel.

_Definition._--Number of the radial spines variable, between ten and
twenty, commonly twelve to sixteen.

{736}5. _Litholophus ligurinus_, Haeckel.

_Litholophus ligurinus_, Haeckel, 1865, Zeitschr. f. wiss. Zool., Bd. xv.
p. 366.

Spines of variable number, from eleven to twenty, commonly twelve to
sixteen, four-sided prismatic, with smooth thin edges, of equal breadth in
their whole length or a little thinner towards the proximal end.

_Dimensions._--Length of the spines 0.3, breadth 0.005.

_Habitat._--Mediterranean (Nice); Central Pacific, Station 274, surface.

6. _Litholophus rhipidium_, Haeckel.

_Litholophus rhipidium_, Haeckel, 1862, Monogr. d. Radiol., p. 402, Taf.
xix. fig. 6.

Spines of variable number, from eleven to twenty, commonly twelve to
sixteen, four-sided prismatic, with distantly denticulated edges, of equal
breadth in their whole length or a little thinner towards both ends.

_Dimensions._--Length of the spines 0.3, breadth 0.006.

_Habitat._--Mediterranean (Messina); North Atlantic, Station 352, surface.

Subgenus 3. _Litholophonium_, Haeckel.

_Definition._--Number of the radial spines twenty (or more?).

7. _Litholophus fasciculus_, n. sp.

Spines constantly (?) twenty, four-sided prismatic, with smooth prominent
edges, nearly of equal breadth throughout their whole length.

_Dimensions._--Length of the spines 0.3 to 0.5, breadth 0.008.

_Habitat._--South Pacific, Station 291, surface.

8. _Litholophus penicillus_, n. sp.

Spines constantly twenty (or more?), four-sided prismatic, with distantly
denticulated edges, gradually thickened from the small pyramidal base to
the middle part, of equal breadth in the distal half.

_Dimensions._--Length of the spines 0.3 to 0.4, breadth 0.005 to 0.007.

_Habitat._--North Pacific, Station 252, surface.

Family XXXV. #CHIASTOLIDA#, Haeckel.

_Acanthochiasmida_, Haeckel, 1862, Monogr. d. Radiol., p. 402.

_Definition._--ACANTHARIA with a variable number of simple radial spines,
which are grown together in pairs (two opposite spines of each pair
representing together a single diametral spine). Diametral spines crossed
loosely in the centre of the spherical or irregular roundish central
capsule. No lattice-shell.

{737}The family #Chiastolida# (or Acanthochiasmida) differs from all other
ACANTHARIA in the peculiar mode of the radial spines; these grown together
in pairs in the centre of the body, so that every two spines opposite in
one axis of the body form together one single diametral spine. All
diametral spines are loosely crossed in the middle of the central capsule,
or connected by a peculiar screw-like winding, but not united firmly.

I established the family Acanthochiasmida in my Monograph (1862, p. 402)
upon the single genus _Acanthochiasma_ (with three species), in which only
ten diametral spines are constantly found; I derived these from the twenty
radial spines of the common #Acanthometra#, supposing that every two
opposite spines of the latter (lying in one axis) were grown together in
the centre, whilst the intimate connection of the twenty radial spines in
the common centre was dissolved. This opinion was afterwards confirmed by
Richard Hertwig, who observed _Acanthochiasma_ intact in the living state.
Although two species of this genus are cosmopolitan and very common, the
number of species is very small; I could add to those three older known
forms only a single new species.

Another genus of this family, _Chiastolus_, was observed by me only in a
single specimen but it is extremely interesting. It has sixteen diametral
spines, disposed quite regularly after the same law of the thirty-two
spines of _Actinastrum_ which we described above (compare above, p. 729).
Therefore we cannot doubt that the former is derived from the latter in the
same way, every two opposite radial spines (of one axis) being grown
together to form a single diametral spine. As we place _Acanthometron_
(with twenty spines) and _Actinastrum_ (with thirty-two spines) in two
different families, it would perhaps be more convenient to separate also
_Acanthochiasma_ and _Chiastolus_ as representatives of two different
families--Acanthochiasmida (with ten diametral spines) and Chiastolida
(with sixteen diametral spines).

As we derive _Acanthochiasma_ from _Acanthometron_ by concrescence in pairs
of the twenty radial spines, the Müllerian law of Icosacantha must be
employed also to the ten diametral spines of the former, therefore two of
them are equatorial, four tropical, and four polar spines. In the same way
we may employ the new law of disposition found in the thirty-two radial
spines of _Actinastrum_ equally to the sixteen diametral spines of
_Chiastolus_, which we derive from the former, four of them are equatorial,
eight tropical, and four polar spines (compare above, p. 732).

_The Central Capsule_ of the Chiastolida is spherical, and exhibits in
general the same shape as in the #Acanthonida#, and specially in the
Astrolonchida. Of course every diametral spine pierces the capsule twice,
at two points diametrically opposed. In some species of _Acanthochiasma_
the central capsule is formed very late, so that it seems often to be
absent. A very accurate description of the capsule and its nucleus, as well
as of the calymma and the pseudopodia, is given by Richard Hertwig in his
Organismus der Radiolarien (1879, pp. 10-18). The pseudopodia are very
numerous, and sometimes bear reddish granules.

{738}_Synopsis of the Genera of Chiastolida._

I. Subfamily Chiastolidina. Sixteen diametral
spines, derived by concrescence of thirty-two
radial spines, 321. _Chiastolus_.

II. Subfamily Acanthochiasmida. Ten diametral
spines, derived by concrescence of twenty
radial spines, 322. _Acanthochiasma_.

Genus 321. _Chiastolus_,[361] n. gen.

_Definition._--#Chiastolida# with sixteen diametral spines, derived from
thirty-two radial spines opposite and grown together in pairs.

The genus _Chiastolus_, hitherto known only by a single observed specimen,
comprises the Chiastolida with sixteen diametral spines, which are loosely
crossed in the centre of the body. Four of these spines are equatorial,
four polar, and eight tropical. We derive _Chiastolus_ from _Actinastrum_
by concrescence of every two opposite spines in one axis of the body
(compare above, pp. 729, 732).

1. _Chiastolus amphicopium_, n. sp. (Pl. 129, figs. 3, 3_a_, 3_b_).

Spines in the central part cylindrical, spirally convoluted in a very
peculiar manner, broadened towards both ends, strongly compressed,
two-edged; the broadest parts are the two truncated distal ends, five to
seven times as broad as the thinnest central part, which is placed between
two spindle-shaped intumescences. These cochleary central parts of the
sixteen spines seem to be resting one upon another. Each spine (composed of
two opposite equal radial spines) has nearly the form of a double oar. The
single observed specimen (preserved in glycerine) exhibited a most regular
disposition of the thirty-two spines (grown together in pairs in the
centre). The diameter of the dark non-transparent spherical central capsule
equalled one-fifth to one-fourth of the total length of the double spines.
When the soft parts of the body were destroyed by sulphuric acid, the
sixteen single spines were suddenly dispersed.

_Dimensions._--Length of the sixteen double spines 0.5, distal breadth (of
the truncated ends) 0.05, central breadth 0.01; diameter of the central
capsule 0.12.

_Habitat._--South-east Pacific (near Juan Fernandez), Station 297, surface.

Genus 322. _Acanthochiasma_,[362] Krohn, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 810.

_Definition._--#Chiastolida# with ten diametral spines, derived from twenty
radial spines opposite and grown together in pairs.

{739}The genus _Acanthochiasma_ with a small number of common species,
comprises the Chiastolida with ten diametral spines, which are loosely
crossed in the centre of the body. I could distinguish only four species,
two of which are cosmopolitan and very widely distributed. In all four
species the diametral spines are quite simple, cylindrical; only in one
species distinguished by a spiral winding or torsion in the middle part,
where they are crossed one to another. We derive _Acanthochiasma_ from
_Acanthometron_ by concrescence of every two spines opposite in one axis of
the body.

1. _Acanthochiasma krohnii_, Haeckel.

_Acanthochiasma krohnii_, Haeckel, 1862, Monogr. d. Radiol., p.403, Taf.
xix. fig. 7.

_Acanthochiasma krohnii_, R. Hertwig, 1879, Organismus d. Radiol., Taf.
ii. fig. 6.

Spines needle-shaped, cylindrical, very thin and long, of equal breadth in
their whole length, distinguished by a high degree of elasticity. Central
capsule colourless or yellowish-white, transparent. Granules of the sarcode
colourless.

_Dimensions._--Length of the spines 0.5 to 1.0, breadth 0.001 to 0.002.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific, very
common.

2. _Acanthochiasma rubescens_, Krohn.

_Acanthochiasma rubescens_, Haeckel, 1862, Monogr. d. Radiol., p. 403.

Spines cylindrical, of equal breadth in their whole length, not very
elastic, pointed at the two ends. Central capsule intransparent, reddish,
with violin-shaped concretions. Granules of the sarcode red coloured.

_Dimensions._--Length of the spines 0.2 to 0.6, breadth 0.004 to 0.006.

_Habitat._--North Atlantic, Madeira, Krohn; Lanzerote, Haeckel.

3. _Acanthochiasma fusiforme_, Haeckel.

_Acanthochiasma fusiforme_, Haeckel, 1862, Monogr. d. Radiol., p. 404,
Taf. xix. fig. 8.

Spines spindle-shaped, from the thicker central part thinned towards the
two thin conical ends, perfectly straight and smooth, rigid, inelastic.
Central capsule non-transparent, brown.

_Dimensions._--Length of the spines 0.3 to 0.5, breadth in the central part
0.006 to 0.009.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific.

4. _Acanthochiasma spirale_, n. sp.

Spines spindle-shaped, tapering from the thicker central part towards the
two thin conical ends, rigid, inelastic; their central part is spirally
convoluted in a very peculiar cochlea-like manner, as in _Chiastolus
amphicopium_ (Pl. 129, figs. 3_a_, 3_b_). The ten spines are propped one
upon another by the central screw. Central capsule dark, opaque.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth of the central
spiral part 0.01.

_Habitat._--Central Pacific, Station 266, surface.

{740}----

Suborder II. ACANTHONIDA, Haeckel (Pls. 130-132).

_Acanthonida_, Haeckel, 1881, Prodromus, p. 465.

_Definition._--#Acanthometra# with twenty radial spines, disposed according
to the Müllerian or Icosacanthan law in five zones each of four spines.

Family XXXVI. #ASTROLONCHIDA#, Haeckel (Pl. 130).

_Astrolonchida_, Haeckel, 1881, Prodromus, p. 465.

_Definition._--#Acanthometra# with twenty radial spines of nearly equal
size and similar form, disposed according to the law of the Icosacantha. No
lattice-shell.

The family #Astrolonchida#, the first and oldest of the #Acanthonida#, is
no doubt the ancestral stock not only of this suborder but also all
#Acanthophracta#, _i.e._, of all Icosacantha or all ACANTHARIA in which
twenty radial spines are regularly disposed according to the Müllerian law,
forming five zones each of four alternating spines (compare above, p. 717).
The Astrolonchida differ from the #Acanthophracta# in the absence of a
complete lattice-shell, from the other two families of #Acanthonida# (the
Quadrilonchida and Amphilonchida) in the equal size and similar form of all
the spines. Probably this equality is nowhere quite perfect, since in all
Icosacantha the central bases of the twenty spines exhibit originally
certain slight differences of form and junction, effected by the regular
disposition itself. But setting aside this slight difference, only
recognisable by means of a very accurate investigation of the central
junction (and in thinner spines often not at all recognisable), the twenty
spines of the Astrolonchida appear perfectly equal. Therefore the four
equatorial spines are not distinguished from the sixteen other spines, as
is constantly the case in the two following families.

The number of genera (eleven) and of species (seventy-six) in the
Astrolonchida is far larger than in the five other families of
#Acanthometra#, and requires a distinction into three different
subfamilies. (A) In the Zygacanthida the form of the radial spines is quite
simple, without apophyses or transverse processes; (B) in the
Phractacanthida each spine bears two opposite apophyses (rarely two
longitudinal rows of these opposite apophyses); (C) in the Stauracanthida
each spine bears a cross of four apophyses, opposite in pairs (rarely four
longitudinal crossed rows of apophyses, opposite in pairs). The
Phractacanthida and Stauracanthida appear as two divergent branches of the
pedigree, derived independently from the common ancestral stock of
Zygacanthida.

In the Zygacanthida, constantly devoid of apophyses, we can distinguish
only three genera, characterised by the different fundamental form of the
radial spines; these are:--(1) _Acanthometron_, with cylindrical or conical
spines (without edges); (2) _Zygacantha_, with compressed and two-edged
spines; (3) _Acanthonia_, with four-edged, prismatic or pyramidal spines.
The transverse section of the spines is in the first case {741}circular, in
the second elliptical or lanceolate, in the third square. All the different
forms of spines, which we find in the numerous ACANTHARIA, may be reduced
to these three forms, and among these the second and third are derived from
the first.

The development of apophyses or of lateral transverse processes (wanting in
the Zygacanthida) is of the greatest value for the further differentiation
of the ACANTHARIA. For from the Phractacanthida (with two opposite
apophyses on each spine) we must derive the Phrastaspida, the common
ancestral stock of the Diporaspida (and therefore also the Belonaspida,
Hexalaspida, Diploconida, and Phractopeltida). On the other hand the
Stauracanthida (with four crossed apophyses on each spine) have produced
the Stauraspida, or the ancestral group of the Tessaraspida and
Sphærocapsida. From all these #Acanthophracta#, possessing a complete
lattice-shell, the Astrolonchida differ in the absence of such a complete
shell. Also in the few cases in which the apophyses become latticed
(_Doracantha_ among the Phractacanthida, and _Phatnacantha_ among the
Stauracanthida), the lattice-plates of the neighbouring spines never meet
with their edges, as is the case in all #Acanthophracta#. But in a
phylogenetic as well as in an ontogenetic sense the former are the
ancestral stock of the latter.

_The Central Capsule_ in the Astrolonchida is commonly spherical, sometimes
with twenty roundish elevations or conical papillæ, extending radially to
the basal half of the radial spines. The calymma is voluminous, and forms
around the radial spines conical or cylindrical "jelly-sheaths," which are
connected with the spines by coronas of Myophrisca (or of the bodies
formerly called "Gallert-cilien," afterwards recognised as "contractile
Filamente").

_Synopsis of the Genera of Astrolonchida._

-------------------------------------------------------------------------
I. Subfamily Zygacanthida.
Twenty radial spines simple, without apophyses or lateral transverse
processes.
-------------------------------------------------------------------------
Spines cylindrical, with circular transverse
section, 323. _Acanthometron_.

Spines compressed, two-edged or lamellar, with
elliptical or rhomboidal transverse section, 324. _Zygacantha_.

Spines quadrangular (prismatic or pyramidal), with
four edges, with square transverse section, 325. _Acanthonia_.

-------------------------------------------------------------------------
II. Subfamily Phractacanthida.
Twenty radial spines provided each with two opposite apophyses (or
two longitudinal rows of apophyses).
-------------------------------------------------------------------------
{ Apophyses simple, 326. _Lithophyllium_.
{
Two apophyses opposite on { Apophyses branched, 327. _Phractacantha_.
each spine. {
{ Apophyses latticed, 328. _Doracantha_.

Two opposite longitudinal rows of apophyses (four
to eight or more apophyses on each spine,
opposite in pairs), 329. _Astrolonche_.
-------------------------------------------------------------------------
III. Subfamily Stauracanthida.
Twenty radial spines provided each with four crossed apophyses (or
four crossed longitudinal rows of apophyses).
-------------------------------------------------------------------------
{ Apophyses simple, 330. _Xiphacantha_.
Four apophyses (in cross {
form) opposite in pairs { Apophyses branched, 331. _Stauracantha_.
on each spine. {
{ Apophyses latticed, 332. _Phatnacantha_.

Four longitudinal rows of apophyses, opposite in
pairs in cross form on each spine, 333. _Pristacantha_.

{742}Subfamily 1. ZYGACANTHIDA, Haeckel.

_Definition._--#Astrolonchida# with twenty simple radial spines, without
apophyses or lateral transverse processes.

Genus 323. _Acanthometron_,[363] J. Müller, 1855, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 229.

_Definition._--#Astrolonchida# with simple cylindrical or needle-shaped
radial spines, without edges and without apophyses; their transverse
section is circular.

The genus _Acanthometron_, with the restricted definition here given, is
the most simple form of all #Acanthonida#, and may be regarded as the
common ancestral form not only of this suborder but also of all
#Acanthophracta#, in general of all Icosacantha, or all ACANTHARIA in which
twenty radial spines are regularly disposed after the Müllerian law (p.
717). In the wider sense, given originally to #Acanthometra# by Johannes
Müller, its discoverer, this genus comprised all ACANTHARIA constituting
here our order "#Acanthometra#" (Radiolaria without lattice-shell, with
radial spines united in the centre). In my Monograph (1862, p. 375) I
restricted this genus to those "Acanthometrida" in which twenty simple
spines of equal size (and without apophyses) are supported one upon another
in the centre, and I separated as _Astrolithium_ those forms in which they
are grown together in the centre. But this difference now appears not so
important, and I restrict here the genus _Acanthometron_ (not
#Acanthometra#) to those most simple forms in which the simple radial
spines are cylindrical or conical, without edges.

Subgenus 1. _Acanthometrella_, Haeckel.

_Definition._--Spines at the central base without leaf-cross, united by the
opposed triangular faces of their pyramidal bases, resting one upon
another.

1. _Acanthometron elasticum_, Haeckel.

_Acanthometra elastica_, Haeckel, 1862, Monogr. d. Radiol., p. 376, Taf.
xv. fig. 1, Taf. xviii. fig. 1.

_Acanthometra elastica_, R. Hertwig, 1879, Organismus d. Radiol., Taf. i.
figs. 2, 2_a_, 2_b_.

Spines cylindrical, very thin and long, needle-shaped, at the central base
four-sided pyramidal, without leaf-cross. Distal apex conical. The spines
are very elastic, of nearly equal thickness in their whole length. Central
capsule quite pellucid, colourless, with a variable number of yellow
pigment-bodies (xanthellæ?).

_Dimensions._--Length of the spines 0.3 to 0.6, breadth 0.001 to 0.002.

_Habitat._--Cosmopolitan, very common in all warmer seas; Mediterranean,
Atlantic, Indian, Pacific, surface.

{743}2. _Acanthometron cylindricum_, n. sp. (Pl. 130, fig. 2).

Spines cylindrical, thick and long, at the central base thickened with a
pear-shaped knob, and with very small central fulcral pyramid, without
leaf-cross. Distal apex rounded or truncated. Central capsule opaque,
filled with red pigment-bodies.

_Dimensions._--Length of the spines 0.4 to 0.8, breadth 0.01 to 0.015.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

3. _Acanthometron fuscum_, J. Müller.

_Acanthometra fusca_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 47, Taf. xi. fig. 4.

_Acanthometra fusca_, Haeckel, 1862, Monogr. d. Radiol, p. 377.

Spines very thin and long, in the proximal half cylindrical, in the distal
half conical, gradually thinned towards the simple conical apex. Central
base a small four-sided pyramid, without leaf-cross. Central capsule
opaque, filled with brown pigment-bodies.

_Dimensions._--Length of the spines 0.2 to 0.4, breadth 0.002 to 0.004.

_Habitat._--Mediterranean (Cette, Portofino, Messina).

4. _Acanthometron bulbosum_, Haeckel.

_Acanthometra bulbosa_, Haeckel, 1862, Monogr. d. Radiol., p. 377, Taf.
xv. fig. 2, Taf. xviii fig. 2.

Spines very thin and long, cylindrical, with simple needle-shaped apex; in
the basal part conical, towards the centre much thickened, with a short
four-sided fulcral pyramid, without leaf-cross. Central capsule opaque,
filled with brown pigment-bodies.

_Dimensions._--Length of the spines 0.3 to 0.5, breadth in the middle and
outer part 0.001, in the basal part 0.008 to 0.012.

_Habitat._--Mediterranean; Atlantic, Station 354, surface.

5. _Acanthometron dolichoscion_, Haeckel (Pl. 129, figs. 6-8).

_Acanthometra dolichoscia_, Haeckel, 1862, Monogr. d. Radiol., p. 377,
Taf. xviii. figs. 3_a_, 3_b_.

Spines very long and thin, cylindrical, in the thickened outer half about
twice as thick as in the thin inner half, at the simple apex compressed,
two-edged. Central base little thickened, with a small four-sided pyramid,
without leaf-cross. Central capsule transparent, whitish or yellowish.

_Dimensions._--Length of the spines 0.6 to 0.8, breadth in the proximal
part 0.004, in the distal part 0.008.

_Habitat._--Mediterranean (Messina, Portofino), surface.

6. _Acanthometron conicum_, n. sp.

Spines short and thick, conical, gradually thinner from the thick conical
base towards the simple apex. Central part of the base with a large fulcral
pyramid, but without leaf-cross. Central capsule transparent, colourless
(?).

_Dimensions._--Length of the spines 0.08 to 0.12, basal breadth 0.02.

_Habitat._--North Pacific, Station 244, surface.

{744}7. _Acanthometron pellucidum_, J. Müller.

_Acanthometra pellucida_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 47, Taf. xi. figs. 1-3.

_Acanthometra pellucida_, Haeckel, 1862, Monogr. d. Radiol., p. 378.

Spines cylindrical, very thin and long, needle-shaped, nearly of equal
thickness in their whole length. Central base with a small fulcral pyramid,
without leaf-cross. Distal apex bifid or bifurcate, with two thin parallel
teeth. Central capsule pellucid, colourless, with yellow pigment-bodies.
(Differs from _Acanthometron elasticum_ mainly in the bifid apex.)

_Dimensions._--Length of the spine 0.1 to 0.3, breadth 0.002 to 0.004.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

8. _Acanthometron wageneri_, Haeckel.

_Acanthometra wageneri_, Haeckel, 1862, Monogr. d. Radiol., p. 378.

Spines cylindrical, in the thickened outer half about twice as broad as in
the thin inner half. Central base a little thickened, with large fulcral
pyramid, but without leaf-cross. Distal apex bifid, with two divergent,
often denticulated teeth. Central capsule pellucid, with yellow
pigment-bodies. (Differs from _Acanthometron dolichoscion_ mainly in the
bifid apex.)

_Dimensions._--Length of the spines 0.2 to 0.5, breadth in the inner part
0.003, in the outer 0.006.

_Habitat._--Adriatic Sea; Triest, Wagner; Corfu, Haeckel, surface.

Subgenus 2. _Phyllostaurus_, Haeckel, 1862, Monogr. d. Radiol., p. 381.

_Definition._--Spines at the central base with a broad leaf-cross, composed
of four prominent triangular lamellæ; the meeting edges of the neighbouring
lamellæ are propped one upon another in such a manner that there are formed
twenty-two hollow pyramidal spaces or compartments (compare p. 721).

9. _Acanthometron siculum_, Haeckel.

_Acanthometra sicula_, Haeckel, 1862, Monogr. d. Radiol., p. 382, Taf.
xvii. figs. 1, 2; Taf. xviii. fig. 8.

Spines elongate, conical, tapering gradually from the thick base towards
the simple distal apex. Conical circular base supported by a basal
leaf-cross of double the breadth. Central capsule yellowish-brown, opaque.

_Dimensions._--Length of the spines 0.3 to 0.4, basal breadth 0.008 to
0.012, leaf-cross 0.024.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

{745}10. _Acanthometron catervatum_, Haeckel.

_Acanthometra brevispina_, Haeckel, 1862, Monogr. d. Radiol., p. 382,
Taf. xv. fig. 5, Taf. xviii. fig. 9.

Spines cylindrical, nearly of equal breadth throughout their whole length.
Apex either simple, conical, or bifid. Base with a large leaf-cross, four
to six times as broad as the spine itself. Central capsule transparent,
yellow. The Atlantic specimens have much longer spines than those figured
from the Mediterranean, but are otherwise not different. Therefore I have
changed the inconvenient name _brevispinum_ into _catervatum_.

_Dimensions._--Length of the spines 0.1 to 0.3, breadth 0.005 to 0.008;
basal leaf-cross 0.02 to 0.032.

_Habitat._--Mediterranean (Messina); North Atlantic, Station 354, Gulf
Stream (Færöe Channel) in great abundance, John Murray, surface.

Subgenus 3. _Astrolithium_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 810.

_Definition._--Spines in the basal part grown perfectly together, so that
the whole skeleton forms a single piece of acanthin; a star with twenty
equal rays.

11. _Acanthometron bulbiferum_, n. sp. (Pl. 130, fig. 1).

_Astrolithium bulbiferum_, Haeckel, 1881, Prodromus, p. 466.

Spines needle-shaped, cylindrical, very thin and long, with simple apex;
suddenly thickened and forming a broad regular cone at the central base;
all twenty conical bulbs have their broad bases grown together and forming
a central icosahedron of acanthin; from its twenty faces arise the circular
bases of the cones in regular disposition.

_Dimensions._--Length of the spines 0.2 to 0.5, breadth 0.001 to 0.003;
height of the basal cones 0.06, basal breadth of them 0.02.

_Habitat._--Central Pacific, Stations 265 to 274, surface.

12. _Acanthometron bifidum_, Haeckel.

_Astrolithium bifidum_, Haeckel, 1862, Monogr. d. Radiol., p. 400, Taf.
xx. fig. 5.

Spines needle-shaped, cylindrical, of equal breadth in their whole length.
Distal apex bifid, with two parallel straight teeth. Central bases of all
twenty spines grown perfectly together and forming a central sphere of
acanthin. Central capsule brown opaque.

_Dimensions._--Length of the spines 0.2, breadth 0.003.

_Habitat._--Mediterranean (Messina), Haeckel, surface.

{746}Genus 324. _Zygacantha_,[364] J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 51.

_Definition._--#Astrolonchida# with simple, compressed, and two-edged
radial spines, without apophyses; their transverse section is elliptical or
rhomboidal.

The genus _Zygacantha_ comprised in the original definition of J. Müller
only a single species, _Zygacantha furcata_, distinguished from the other
#Acanthometra# by forked spines with two long parallel teeth. It seems now
advisable to unite in this genus all those Astrolonchida in which the
simple spines are two-edged, compressed, or leaf-shaped. The term
_Zygacantha_ may be conceived as the general expression of the important
fact, that in all Icosacantha the twenty spines are opposite in pairs.

Subgenus 1. _Zygacantharium_, Haeckel.

_Definition._--Spines at the central base without leaf-cross and without
hollow pyramidal compartments, united by the opposed triangular faces of
their pyramidal bases, resting one upon another.

1. _Zygacantha lanceolata_, Haeckel.

_Acanthometra lanceolata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 48, Taf. xi. fig. 12.

_Acanthometra lanceolata_, Haeckel, 1862, Monogr. d. Radiol., p. 380.

Spines lanceolate, from the broader middle part equally thinned towards the
two ends. Apex simple. Base pyramidal, without leaf-cross. Each flat
lamellar spine exhibits an elevated middle rib (like a lanceolate leaf),
and is therefore compressed quadrangular.

_Dimensions._--Length of the spines 0.1 to 0.15, greatest breadth (in the
width) 0.03 to 0.04.

_Habitat._--Mediterranean (Saint Tropez, French shore), J. Müller; North
Atlantic (Canary Islands), Haeckel, surface.

2. _Zygacantha costata_, n. sp.

Spines compressed, two-edged, linear, of nearly equal breadth in their
whole length. Apex truncate. Base pyramidal, without leaf-cross. Each flat
lamellar spine exhibits an elevated middle rib, which in the distal half is
cleft into two divergent rods ending in the corners of the truncated apex.
(Similar to _Zygacantha dicopa_, but with broader free spines, which are
not grown together in the centre.)

_Dimensions._--Length of the spines 0.1 to 0.15, breadth 0.02.

_Habitat._--Central Pacific, Stations 265 to 274, surface.

{747}3. _Zygacantha compressa_, Haeckel.

_Acanthometra compressa_, Haeckel, 1862, Monogr. d. Radiol., p. 378, Taf.
xviii. figs. 4_a_, 4_b_.

Spines in the proximal half compressed, broad lanceolate, four to six times
as broad as in the thin cylindrical distal half. Apex simple or short
bifid. Base pyramidal, thickened, without leaf-cross. No middle rib.
Central capsule opaque, yellowish or reddish-brown.

_Dimensions._--Length of the spines 0.3 to 0.5, breadth of the inner half
0.015 to 0.02, of the outer half 0.002 to 0.003.

_Habitat._--Mediterranean (Messina); Atlantic (Canary Islands), Station
352, surface.

4. _Zygacantha furcata_, J. Müller.

_Zygacantha furcata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 51, Taf. ix. fig. 6.

_Acanthometra furcata_, J. Müller, 1856, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 499.

_Acanthometra furcata_, Haeckel, 1862, Monogr. d. Radiol., p. 380.

Spines compressed, fork-shaped, divided by two thickened knots into three
sections of nearly equal length; middle section broader than the proximal,
but smaller than the distal section, which is cleft by a deep fissure into
two long parallel straight branches. Base pyramidal, without leaf-cross.
Central capsule purple, with yellow bodies.

_Dimensions._--Length of the spines 0.1 to 0.15, basal breadth 0.003 to
0.005, distal breadth 0.02 to 0.03.

_Habitat._--Mediterranean (Cette). J. Müller, surface.

Subgenus 2. _Zygacanthidium_, Haeckel.

_Definition._--Spines at the central base with a cross of four prominent
leaves; the meeting edges of the neighbouring lamellæ so rest one upon
another that twenty-two hollow pyramidal compartments are formed (compare
p. 721).

5. _Zygacantha dichotoma_, Haeckel.

_Zygacantha dichotoma_, Haeckel, 1862, Monogr. d. Radiol., p. 381.

_Acanthometra dichotoma_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 50, Taf. ix. fig. 5.

Spines compressed, pincer-shaped, cleft nearly throughout their whole
length into two thin parallel straight rods or fork branches, which are
united only in their middle by a narrow bridge, and at their central base
by the pyramidal small central apex; above this pyramid each rod is divided
into two broad triangular leaves, forming a large basal leaf-cross. Central
capsule purple, with yellow bodies.

_Dimensions._--Length of the spines 0.1 to 0.15, breadth 0.01 to 0.02.

_Habitat._--Mediterranean (Nice), J. Müller, surface.

{748}6. _Zygacantha complanata_, n. sp.

Spines compressed, two-edged, linear, of equal breadth throughout their
whole length. Apex bifid. Base thickened, with a large leaf-cross. (Similar
to _Amphilonche complanata_, but different in the equal size of all twenty
spines, which are somewhat broader.)

_Dimensions._--Length of the spines 0.12 to 0.18, breadth 0.002.

_Habitat._--North Pacific, Station 244, surface.

7. _Zygacantha semicompressa_, Haeckel.

_Acanthometra hemicompressa_, Car, 1884, Zool. Anzeiger, p. 94, with
woodcut.

Spines in the proximal half compressed, two-edged, linear, three to four
times as broad as in the needle-shaped cylindrical distal half. Apex
simple. Base pyramidal, with a small leaf-cross. Central capsule
transparent.

_Dimensions._--Length of the spines 0.1 to 0.2, breadth in the proximal
half 0.02, in the distal half 0.006.

_Habitat._--Mediterranean (Genoa), Haeckel; Adriatic Sea (Trieste), Car,
surface.

8. _Zygacantha foliacea_, n. sp.

Spines lanceolate, tapering from the broader middle part towards the two
ends, with two dentated or serrated edges and a prominent middle rib. Apex
simple. Base with a small leaf-cross. The spines like the leaves of _Agave
americana_.

_Dimensions._--Length of the spines 0.15 to 0.2, middle breadth 0.02 to
0.03.

_Habitat._--Indian Ocean, near the Island of Socotra, Haeckel.

Subgenus 3. _Zygacanthonium_, Haeckel.

_Definition._--Spines in the centre grown perfectly together and forming a
single star-shaped piece of acanthin.

9. _Zygacantha dicopa_, Haeckel.

_Astrolithium dicopum_, Haeckel, 1862, Monogr. d. Radiol., p. 400, Taf.
xx. figs. 3, 4.

Spines compressed, two-edged, linear, of nearly equal breadth throughout
their whole length, with a prominent middle rib, which in the distal third
is cleft into two divergent teeth, ending in the two corners of the broad,
obliquely truncated apex. All twenty spines with their central bases grown
together and forming one single piece of acanthin--a star with twenty rays.

_Dimensions._--Length of the spines 0.12 to 0.2, breadth 0.01 to 0.02.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, many Stations,
surface.

{749}Genus 325. _Acanthonia_,[365] Haeckel, 1881, Prodromus, p. 465.

_Definition._--#Astrolonchida# with simple, four-edged, prismatic or
pyramidal radial spines, without apophyses; their transverse section is
square.

The genus _Acanthonia_ comprises all those Astrolonchida (formerly united
with _Acanthometron_) in which the simple spines either in their whole
length or in the greatest part of it are four-edged, with square transverse
section. They are sometimes more prismatic (with equal breadth), at other
times more pyramidal (with decreasing breadth towards the distal apex). If
_Acanthometron_ be the common simple ancestral form of the Acanthonida,
then the two-edged _Zygacantha_, and the four-edged _Acanthonia_ may be
regarded as two divergent main lines arising from it; the former leading to
the Phractacanthida and Diporaspida, the latter leading to the
Stauracanthida and Tessaraspida.

Subgenus 1. _Acanthonarium_, Haeckel.

_Definition._--Spines at the central base, without leaf-cross and without
hollow pyramidal compartments, united by the triangular faces of their
pyramidal bases, resting one upon another.

1. _Acanthonia tetracopa_, Haeckel (Pl. 129, figs. 9-11).

_Acanthometra tetracopa_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 47, Taf. vii. figs. 3 to 5, Taf. xi. fig. 5.

_Acanthometra tetracopa_, Haeckel, 1862, Monogr. d. Radiol., p. 379, Taf.
xviii. fig. 5.

Spines quadrangular prismatic, with prominent lamellar edges, of equal
breadth in their whole length. Base pyramidal, without leaf-cross. Apex
truncated, or with four short teeth (sometimes only two teeth). Central
capsule opaque, brown or yellow.

_Dimensions._--Length of the spines 0.15 to 0.2, breadth 0.01 to 0.012.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific, many
Stations, surface.

2. _Acanthonia prismatica_, n. sp.

Spines quadrangular prismatic, without prominent edges, with four plane
lateral faces, of equal breadth throughout their whole length. Base
pyramidal, without leaf-cross. Apex truncate, with square apical face.

_Dimensions._--Length of the spines 0.1 to 0.16, breadth 0.005 to 0.008.

_Habitat._--Antarctic Ocean, Station 154, surface.

{750}3. _Acanthonia denticulata_, n. sp.

Spines quadrangular prismatic, with prominent elegantly denticulated edges,
of equal breadth throughout their whole length. Both ends pyramidal,
without leaf-cross. (Similar to _Amphilonche denticulata_, but with the
twenty spines all equal, and with spherical central capsule.)

_Dimensions._--Length of the spines 0.12 to 0.16, breadth 0.008.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

4. _Acanthonia mülleri_, Haeckel.

_Acanthometra mülleri_, Haeckel, 1862, Monogr. d. Radiol., p. 379, Taf.
xv. fig. 3, Taf. xviii. fig. 6.

Spines quadrangular, nearly prismatic, but tapering gradually from the
pyramidal base towards the distal bifid apex, which bears two thin,
parallel, or little divergent teeth; four edges elegantly denticulated;
base without leaf-cross. Central capsule yellowish or reddish.

_Dimensions._--Length of the spines 0.12 to 0.18, basal breadth 0.005.

_Habitat._--Mediterranean (Messina); Atlantic, Stations 348 to 352,
surface.

5. _Acanthonia fragilis_, Haeckel.

_Acanthometra fragilis_, Haeckel, 1862, Monogr. d. Radiol., p. 380, Taf.
xv. fig. 4, Taf. xviii. fig. 7.

Spines quadrangular, nearly prismatic, but gradually thickened from the
pyramidal base towards the distal truncated end; four edges regularly
denticulated; base without leaf-cross. Central capsule opaque.

_Dimensions._--Length of the spines 0.5 and more, basal breadth 0.002 to
0.003, distal breadth 0.008 to 0.016 or more.

_Habitat._--Mediterranean (Messina); North Atlantic, Station 354, surface.

6. _Acanthonia convexa_, n. sp.

Spines quadrangular, with four broad lamellar, convex, prominent edges,
which from the broader middle part are thinned towards the two emarginated
ends; each end with a small quadrangular pyramid, without leaf-cross. All
the twenty spines of this species exhibit nearly the same form, which is
seen in the caudal or posterior (geotomical) spine of _Amphilonche anomala_
(Monogr. d. Radiol., Taf. xviii. fig. 23, _b_).

_Dimensions._--Length of the spines 0.12 to 0.16, breadth in the middle
part 0.02 to 0.025.

_Habitat._--North Pacific, Station 244, surface.

7. _Acanthonia concava_, n. sp.

Spines quadrangular, with four broad lamellar, concave, prominent edges,
which from the smaller middle part are broadened towards the two
emarginated ends; each end with a small {751}quadrangular pyramid, without
leaf-cross. All the twenty spines of this species exhibit nearly the same
form, which is seen in the frontal or anterior (hydrotomical) spine of
_Amphilonche anomala_ (Monogr. d. Radiol., Taf. xviii. fig. 23, _a_).

_Dimensions._--Length of the spines 0.15, breadth in the middle part 0.012,
on both ends 0.025.

_Habitat._--South Pacific, Station 288, surface.

8. _Acanthonia quadrangula_, n. sp.

Spines quadrangular prismatic, with smooth prominent straight edges, but of
very different breadth in both halves; the inner or proximal half (inside
the central capsule) twice to four times as broad as the outer or distal
half (outside the central capsule); both ends suddenly separated by a
constriction, in which is inserted the membrane of the capsule. The latter
is pellucid, with a number of yellow bodies. Apex of the spines simple,
base pyramidal, without leaf-cross.

_Dimensions._--Length of the spines 0.2 to 0.4, breadth of the basal half
0.01, of the distal half 0.002 to 0.004.

_Habitat._--Atlantic, Canary Islands, Azores, Station 354, surface.

Subgenus 2. _Acanthonidium_, Haeckel.

_Definition._--Spines at the central base with a broad leaf-cross, composed
of four prominent triangular lamellæ; the meeting edges of these crossed
lamellæ between every three or four neighbouring spines so rest one upon
another that twenty-two hollow pyramidal compartments are formed (compare
p. 721).

9. _Acanthonia echinoides_, Haeckel.

_Acanthometra echinoides_, Claparède et Lachmann, 1858, Études sur les
Infusoires et les Rhizopodes, &c., p. 459, pl. xxiii. figs. 1-5.

_Acanthometra echinoides_, Haeckel, 1862, Monogr. d. Radiol., p. 383.

Spines quadrangular prismatic without prominent edges, with four plane
lateral faces, of equal breadth throughout their whole length. Central
base three to four times as broad, with large leaf-cross. Apex truncated or
bifid (sometimes with four short teeth).

_Dimensions._--Length of the spines 0.3 to 0.5, breadth 0.004 to 0.008;
leaf-cross 0.02 to 0.03.

_Habitat._--North Atlantic, west coast of Norway (Claparède et Lachmann);
Færöe Channel (Gulf Stream), John Murray, surface.

10. _Acanthonia claparedei_, Haeckel.

_Acanthometra claparedei_, Haeckel, 1862, Monogr. d. Radiol., p. 383,
Taf. xviii. fig. 12.

Spines quadrangular prismatic, with four prominent lamellar edges, of equal
breadth throughout their whole length. Apex bifid. Central base twice as
broad, with large leaf-cross. Central capsule opaque, reddish-brown.

_Dimensions._--Length of the spines 0.6, breadth 0.016; leaf-cross 0.032.

_Habitat._--Mediterranean (Messina); Atlantic, Station 347, surface.

{752}11. _Acanthonia cuspidata_, Haeckel.

_Acanthometra cuspidata_, Haeckel, 1862, Monogr. d. Radiol., p. 383, Taf.
xviii. fig. 11.

Spines quadrangular pyramidal, with four prominent lamellar edges, tapering
gradually from the broad base towards the simple distal apex. Basal
leaf-cross as broad, with a small central pyramid.

_Dimensions._--Length of the spines 0.5, basal breadth 0.012; leaf-cross
0.025.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

12. _Acanthonia quadrifolia_, Haeckel.

_Acanthometra quadrifolia_, Haeckel, 1862, Monogr. d. Radiol., p. 382,
Taf. xviii. fig. 10.

Spines in the basal part quadrangular pyramidal, with four prominent
lamellar edges and a large basal leaf-cross; in the distal part three to
six times as long, cylindrical, of equal breadth. Apex simple, conical, or
bifid. Central capsule yellow or whitish. This common species differs from
the nearly allied _Acanthometron catervatum_ mainly by the strong
development of the large basal leaves or wings.

_Dimensions._--Length of the spines 0.3 to 0.5, breadth in the distal half
0.002, in the basal part 0.02.

_Habitat._--Mediterranean; North Atlantic, Stations 252 to 254; Færöe
Channel, Gulf Stream, in enormous numbers, John Murray, surface and at
various depths.

13. _Acanthonia diplopyramis_, n. sp.

Spines formed like a quadrangular double pyramid or an irregular
octahedron; the basal leaf-cross being extremely developed, with four very
large and thin lamellar leaves; the basal halves of the twenty double
pyramids are united by the meeting edges of those leaves, while their
distal halves are free, with simple apices. Therefore the four triangular
leaves of each spine are equally thinned from the middle towards the two
ends.

_Dimensions._--Length of the spines 0.1 to 0.2, greatest breadth 0.04 to
0.08.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

14. _Acanthonia multispina_, Haeckel.

_Acanthometra multispina_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 47, Taf. vii. figs. 6-9.

_Acanthometra multispina_, Haeckel, 1862, Monogr. d. Radiol., p. 384.

Spines quadrangular, with four lamellar prominent edges, in the proximal
half nearly prismatic, in the distal half pyramidal; both halves separated
by a prominent short tooth on each edge. Here in the middle part the
breadth (including the four teeth) is equal to the basal leaf-cross.
Central capsule opaque, brown.

_Dimensions._--Length of the spines 0.2 to 0.3, greatest breadth 0.01 to
0.02.

_Habitat._--Mediterranean (Messina); Tropical Atlantic, Station 348.

{753}15. _Acanthonia serrulata_, n. sp.

Spines quadrangular pyramidal, with simple apex; the large basal leaf-cross
nearly half as long as the prolonged distal part. The four prominent
triangular edges of the latter are very thin and broad lamellæ, finely
dentate or serrate.

_Dimensions._--Length of the spines 0.2 to 0.3, greatest breadth 0.05 to
0.06.

_Habitat._--Central Pacific, Station 272, surface.

Subgenus 3. _Acantholithium_, Haeckel.

_Definition._--Spines in the basal part grown together, so that the whole
skeleton is not composed of twenty separated pieces, but represents a
single piece of acanthin--a star with twenty equal rays.

16. _Acanthonia stellata_, n. sp.

Spines quadrangular pyramidal, with simple distal apex, in the basal part
grown perfectly together, so that the whole skeleton forms a single piece
of acanthin--a starlet with twenty equal rays. The free pyramidal part of
each spine is twice to four times as long as the basal part.

_Dimensions._--Length of the spines 0.1 to 0.15, greatest breadth (on the
surface of the central solid sphere) 0.02 to 0.04.

_Habitat._--Central Pacific, Station 271, surface.

Subfamily 2. PHRACTACANTHIDA, Haeckel.

_Definition._--#Astrolonchida# with twenty radial spines, each of which
bears two opposite apophyses or lateral transverse processes; sometimes two
longitudinal rows of opposite apophyses.

Genus 326. _Lithophyllium_,[366] J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 52.

_Definition._--#Astrolonchida# with two simple, not branched, opposite
apophyses on each radial spine.

The genus _Lithophyllium_ was founded by J. Müller for a single species
(_Lithophyllium foliosum_), which we also here retain as the type of the
genus. It is the first observed Astrolonchid, which bears two opposite
lateral apophyses on each spine, and may therefore be regarded as the
ancestral form of the subfamily Phractacanthida. The two opposite apophyses
are here simple, whilst in the other genera of the subfamily they are
branched or multiplied.

{754}1. _Lithophyllium cruciatum_, Haeckel.

_Acanthometra cruciata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 49, Taf xi. fig. 11.

_Xiphacantha cruciata_, Haeckel, 1862, Monogr. d. Radiol., p. 385, Taf.
xviii. fig. 13.

Spines cylindrical, very thin, crossed perpendicularly in the distal third
by a thin transverse beam; both lateral rods of the cross have the same
length as the distal end. Apex simple. Base pyramidal, without leaf-cross.

_Dimensions._--Length of the spines 0.05 to 0.15, breadth 0.001 to 0.002.

_Habitat._--Mediterranean, Atlantic, Stations 352 to 354, &c., surface.

2. _Lithophyllium gladiatum_, n. sp. (Pl. 130, fig. 3).

Spines compressed, sword-shaped, two-edged, crossed perpendicularly in the
middle part by a broad, somewhat curved transverse beam; both lateral rods
of the cross are triangular compressed, and have the same length as the
proximal end. Apex simple. Base pyramidal, without leaf-cross.

_Dimensions_.--Length of the spines 0.2 to 0.25, breadth 0.01.

_Habitat_.--North Pacific, Station 241, surface.

3. _Lithophyllium condylatum_, n. sp.

Spines compressed quadrangular, tapering from the broader middle towards
the two ends; from the middle part arise two opposite perpendicular
apophyses, which bear on the rounded end a thickened condyle. Apex simple.
Base pyramidal, with a small leaf-cross.

_Dimensions._--Length of the spines 0.1 to 0.12, breadth in the middle part
0.02.

_Habitat._--Central Pacific, Station 266 to 272, surface.

4. _Lithophyllium foliosum_, J. Müller.

_Lithophyllium foliosum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 52, Taf. xi. figs. 6-10.

_Xiphacantha foliosa_, Haeckel, 1862, Monogr. d. Radiol., p. 385.

Spines lanceolate, tapering from the broader middle towards the two ends.
From the middle part or from the outer third arise two opposite triangular
apophyses, which are not perpendicular to the axis of the spine, but form
an acute angle with its distal part. Therefore each spine represents a
broad leaf with a middle rib and with three lobes or truncated teeth. The
distal apex is distinguished by a violet colour. Base pyramidal, without
leaf-cross. Central capsule yellow.

_Dimensions._--Length of the spines 0.05 to 0.1, breadth 0.02 to 0.03.

_Habitat._--Mediterranean, French shore, Saint Tropez, J. Müller.

{755}Genus 327. _Phractacantha_,[367] Haeckel, 1881, Prodromus, p. 465.

_Definition_.--#Astrolonchida# with two branched, but not latticed,
opposite apophyses on each radial spine.

The genus _Phractacantha_ differs from its ancestral form, the preceding
_Lithophyllium_ by the ramification of the apophyses, which are either
forked or bear lateral branches. If the prolonged fork-branches of the
neighbouring spines meet and form a lattice-shell, this genus passes over
into _Phractaspis_, the common ancestral form of the Diporaspida.

1. _Phractacantha_ bifurca, n. sp.

Spines cylindrical, thin, of equal breadth throughout their whole length.
Apex simple. Base pyramidal, without leaf-cross. From the outer third arise
two opposite, thin, forked apophyses; ends of the fork branches acute.

_Dimensions._--Length of the spines 0.1, breadth 0.006.

_Habitat._--Central Pacific, Station 274, surface.

2. _Phractacantha bipennis_, n. sp.

Spines compressed, two-edged, tapering from the broader middle towards the
two ends. Apex simple. Base pyramidal, without leaf-cross. From the middle
arise two opposite broad, forked apophyses; ends of the fork-branches broad
and obtuse.

_Dimensions._--Length of the spines 0.15, breadth in the middle 0.012.

_Habitat._--Central Pacific, Station 266, surface.

Genus 328. _Doracantha_,[368] Haeckel, 1881, Prodromus, p. 465.

_Definition_.--#Astrolonchida# with two latticed or fenestrated apophyses
on each radial spine.

The genus _Doracantha_ arises from the foregoing _Phractacantha_ by union
of the fork-branches of the apophyses on each spine. By this concrescence
is formed a polygonal or roundish plate with two pores, pierced by the
radial spine between them. _Doracantha_ may be regarded as a _Dorataspis_,
in which the twenty small fenestrated plates have not become united.

{756}1. _Doracantha dorataspis_, n. sp.

Spines compressed, two-edged, tapering from the broader middle towards the
two ends. Apex simple. Base pyramidal, without leaf-cross. From the middle
part of each spine arise two opposite forked apophyses; the neighbouring
fork-branches are recurved and united in the tangential plane; so that each
spine bears a plate or shield with two elliptical pores; the margin of the
roundish plate bears a variable number of short teeth.

_Dimensions._--Length of the spines 0.2, breadth 0.02.

_Habitat._--Central Pacific, Station 271, surface.

Genus 329. _Astrolonche_,[369] Haeckel, 1881, Prodromus, p. 465.

_Definition._--#Astrolonchida# with numerous simple apophyses (four to
eight or more on each radial spine), which are arranged in two opposite
longitudinal rows (rarely in six such rows, opposite in three parallel
planes).

The genus _Astrolonche_ differs from the preceding Phractacanthida in the
multiplication of the simple apophyses. Commonly these are opposite in
pairs in two longitudinal rows (on the two edges of the compressed spine,
at least two on each side). But sometimes (in the subgenus
_Astrolonchidium_) there are six instead of two longitudinal rows of teeth,
and these are placed in three parallel planes.

Subgenus 1. _Astroloncharium_, Haeckel.

_Definition._--Each spine with two longitudinal rows of apophyses, opposite
in one meridian plane.

1. _Astrolonche bicruciata_, n. sp.

Spines compressed, two-edged, almost of equal breadth throughout their
whole length, with simple apex and small leaf-cross at the base. From the
two edges of the middle part of each spine arise two pairs of opposite
apophyses, which are compressed and a little curved (convex on the distal,
concave on the proximal margin). All four apophyses are of the same size
and form; the radial distance of each pair much smaller than their common
distance from the centre.

_Dimensions._--Length of the spines 0.25, breadth 0.02.

_Habitat._--Central Pacific, Station 270, surface.

2. _Astrolonche mucronata_, Haeckel.

_Acanthometra mucronata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 49, Taf. x. fig. 9.

_Aspidomma mucronatum_, Haeckel, 1862, Monogr. d. Radiol., p. 424.

Spines conical, little compressed, with simple apex and small leaf-cross at
the base. From the basal half of each spine arise two pairs of opposite
apophyses, which exhibit a very different form. {757}The upper or distal
apophyses (nearly in the middle of the spine) are flat, leaf-shaped,
broadened in the periphery, often lobed, and sometimes branched or even
fenestrated. (Transition to _Phractaspis_ and _Dorataspis_, or to
_Phractopelta_?) The lower or proximal apophyses are thick, simple, all
slightly curved; they are equidistant from the former and from the centre.

_Dimensions._--Length of the spines 0.2 to 0.3, greatest breadth 0.01.

_Habitat._--Mediterranean (Cette on the French shore), surface, J. Müller.

3. _Astrolonche pectinata_, Haeckel.

_Acanthometra pectinata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 48, Taf. x. figs. 1, 2.

_Xiphacantha pectinata_, Haeckel, 1862, Monogr. d. Radiol., p. 386.

Spines compressed quadrangular, with short, simple or bifid apex, pyramidal
on the base, without leaf-cross. From the two broader edges of the proximal
half arise two longitudinal rows of opposite apophyses; three to four
slender teeth in each row.

_Dimensions._--Length of the spines 0.2, breadth 0.08.

_Habitat._--Mediterranean, Cette (French shore), surface, J. Müller.

4. _Astrolonche pinnata_, n. sp.

Spines in the distal half compressed, linear, two-edged, with bifid apex;
in the proximal half three times as broad lanceolate, at the base
pyramidal, without leaf-cross. From the two broader edges of the proximal
half arise two longitudinal rows of opposite apophyses; four to six broad
triangular teeth in each row.

_Dimensions._--Length of the spines 0.3 to 0.4, breadth in the outer half
0.012, in the inner half 0.03 to 0.04.

_Habitat._--South Atlantic, Station 332, surface.

Subgenus 2. _Astrolonchidium_, Haeckel.

_Definition._--Each spine with three parallel double rows of opposite
apophyses.

5. _Astrolonche serrata_, Haeckel.

_Xiphacantha serrata_, Haeckel, 1862, Monogr. d. Radiol., p. 386, Taf.
xvii. fig. 3, Taf. xviii. figs. 14_a_, 14_b_.

_Xiphacantha serrata_, R. Hertwig, 1879, Organismus d. Radiol., p. 11,
Taf. ii. fig. 4.

_Acanthometra serrata_, Haeckel, 1860, Monogr. d. Radiol., p. 807.

Spines in the distal half thin, nearly cylindrical or a little compressed,
thinned towards the short simple or bifid apex; in the proximal half three
to six times as broad, four-winged, with six longitudinal rows of opposite
apophyses. From the edges of the two broader (lateral) wings arise three
{758}to four pairs of strong conical teeth; from the edges of the two
smaller (sagittal) wings arise also three to four pairs of teeth, but very
short and broad, triangular; each of these teeth bears on both its
(lateral) sides two opposite slender conical teeth, which are parallel to
the large conical teeth of the lateral wings. Therefore all teeth (eighteen
to twenty-four) are placed opposite in pairs in three parallel planes. Base
of the spines pyramidal, with a small leaf-cross. The central capsule of
this large and very remarkable species commonly entirely includes the
apophyses of the spines, and is opaque, whitish.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth of the distal half
0.002 to 0.004, of the proximal half 0.01 to 0.02.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

Subfamily 3. STAURACANTHIDA, Haeckel.

_Definition._--#Astrolonchida# with a cross of four free apophyses (or four
crossed longitudinal rows of apophyses) on each radial spine.

Genus 330. _Xiphacantha_,[370] Haeckel, 1862, Monogr. d. Radiol., p. 384.

_Definition._--#Astrolonchida# with four simple apophyses on each radial
spine, opposite in pairs in the form of a cross.

The genus _Xiphacantha_ was founded by me in 1862 for all those
_Acanthometrida_ which bear simple or branched apophyses on their twenty
equal spines. I restrict here the genus to those Astrolonchida which bear
on each spine a cross of four simple, not branched, apophyses. These are
either conical teeth or broad wings, sometimes extremely thin leaves.
_Xiphacantha_ may be regarded as the ancestral form not only of the
subfamily Stauracanthida, but also of the Tessaraspida, derived from the
latter.

Subgenus 1. _Xiphacanthonia_, Haeckel.

_Definition._--Apophyses of the radial spines small, formed like a tooth or
a hook, not wing-shaped. Edges of the spines commonly narrow, little
prominent.

1. _Xiphacantha quadridentata_, Haeckel.

_Xiphacantha quadridentata_, Haeckel, 1862, Monogr. d. Radiol., p. 387,
Taf. xviii. figs. 15_a_, 15_b_.

_Acanthometra quadridentata_, J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 48, Taf. x. fig. 3.

Spines slender, four-sided prismatic, gradually thinner towards the simple
pyramidal apex. Base with large wing-cross. Four apophyses about in the
middle of each spine, conical, straight, smooth, about as long as the basal
breadth of the spine. Central capsule opaque, reddish-brown.

{759}_Dimensions._--Length of the spines 0.2 to 0.3, breadth in the middle
part 0.012, on the base 0.02; length of the apophyses 0.02 to 0.03.

_Habitat._--Mediterranean, Atlantic, Stations 348, 354, surface.

2. _Xiphacantha crucifera_, n. sp.

Spines slender, four-sided prismatic, thin, of nearly equal breadth
throughout the whole length. Base with small wing-cross. Four apophyses
much nearer the proximal than the distal end, thin, straight smooth, three
to six times as long as the basal breadth of the spine. Central capsule
pellucid, with yellow bodies.

_Dimensions._--Length of the spines 0.4 to 0.5, breadth 0.002 to 0.004;
length of the apophyses 0.01 to 0.02.

_Habitat._--Central Pacific, Station 266, surface.

3. _Xiphacantha spinulosa_, Haeckel.

_Xiphacantha spinulosa_, Haeckel, 1862, Monogr. d. Radiol., p. 388, Taf.
xvii. fig. 4.

_Acanthometra spinulosa_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 807.

Spines stout, four-sided prismatic, of nearly equal breadth throughout the
whole length, on the distal apex truncated, two-edged. Base with large
wing-cross. Four apophyses about in the middle of each spine, conical,
straight, spinulated, twice to four times as long as the basal breadth of
the spine. Central capsule opaque, yellow.

_Dimensions._--Length of the spines 0.27, breadth 0.014; length of the
apophyses 0.05.

_Habitat._--Mediterranean (Messina), surface.

4. _Xiphacantha emarginata_, n. sp.

Spines stout, four-sided, in the proximal half prismatic, in the distal
half pyramidal, with emarginated apex. Base with very large wing-cross.
Four apophyses about in the middle of each spine, compressed, straight,
two-edged, with emarginated ends, about twice as long as the basal breadth
of the spine. Central capsule small, pellucid.

_Dimensions._--Length of the spines 0.2 to 0.3, breadth 0.02 to 0.03;
length of the apophyses 0.04 to 0.06.

_Habitat._--South Pacific, Station 291, surface.

5. _Xiphacantha falcata_, n. sp.

Spines slender, four-sided prismatic, gradually thickened from both ends
towards the middle part, with simple apex. Base with very small wing-cross.
Four apophyses about in the middle part of each spine, falcated,
compressed, recurved, twice to three times as long as the greatest breadth
of the spine.

{760}_Dimensions._--Length of the spines 0.3 to 0.4, greatest breadth
0.015; length of the apophyses 0.03 to 0.05.

_Habitat._--Central Pacific, Station 270, surface.

6. _Xiphacantha ancorata_, n. sp.

Spines slender, four-sided prismatic, gradually thickened from the narrow
base towards the short, simple, pyramidal apex. Base with large wing-cross.
Four apophyses on the distal end, immediately below the pyramidal apex,
falcated, strongly recurved, like an anchor with four strong teeth.

_Dimensions._--Length of the spines 0.4 to 0.5, basal breadth 0.005, distal
breadth 0.015; length of the apophyses 0.04.

_Habitat._--Indian Ocean, Madagascar, Rabbe.

Subgenus 2. _Xiphacanthidium_, Haeckel.

_Definition._--Apophyses of the radial spines broad, compressed,
wing-shaped, formed like a large thin plate. Edges of the spines commonly
broad, strongly prominent.

7. _Xiphacantha stauroptera_, n. sp.

Spines four-winged, from the middle part little thinner towards both ends.
Apex simple pyramidal. Base with small wing-cross. Four apophyses
wing-shaped, isosceles triangular or truncated quadrangular; their base is
about half as broad as their length, and occupies in the basal half of each
spine about one-sixth to one-eighth of its length.

_Dimensions._--Length of the spines 0.1 to 0.15; greatest breadth (diagonal
of the cross of the apophyses) 0.006 to 0.009.

_Habitat._--South Atlantic, Station 332, surface.

8. _Xiphacantha trigonoptera_, n. sp.

Spines four-winged, tapering rapidly from the broad middle part towards
both ends. Apex simple pyramidal. Base with a small wing-cross. Four
apophyses equilateral triangular; their base occupies the middle of each
spine, extending to about one-fifth of its length.

_Dimensions._--Length of the spines 0.15 to 0.2, greatest breadth (diagonal
of the cross of the apophyses) 0.06 to 0.08.

_Habitat._--Central Pacific, Station 272, surface.

9. _Xiphacantha macroptera_, n. sp.

Spines four-winged, from the broad middle part gradually thinner towards
both ends. Apex simple pyramidal. Base with small wing-cross. Four
apophyses wing-shaped, very long and broad, {761}triangular; their base
occupies about the middle third of each spine; the proximal side of each
wing is the shortest, truncated or concave.

_Dimensions._--Length of the spines 0.4 to 0.5, greatest breadth (diagonal
of the middle leaf-cross) 0.08 to 0.12.

_Habitat._--North Pacific, Stations 253 to 256, surface.

10. _Xiphacantha platyptera_, n. sp.

Spines four-winged, nearly prismatic in the basal and distal parts. Apex
truncated. Base with small wing-cross. Four apophyses wing-shaped, very
broad and thin, extremely delicate, of irregular quadrangular or nearly
rhomboidal form; their base occupies the middle half of the length of the
spines (second and third quarter). Very variable in form. Often the edges
of the wing-apophyses of the neighbouring spines seem to meet.

_Dimensions._--Length of the spines 0.2 to 0.3, greatest breadth (diagonal
of the leaf-cross) 0.06 to 0.08.

_Habitat._--Pacific, widely distributed, Stations 239, 253, 272, 288, &c.,
surface.

11. _Xiphacantha ciliata_, n. sp. (Pl. 129, figs. 4, 5).

Spines four-winged, prismatic in the basal and the distal third. Apex
pyramidal, short. Base with large leaf-cross. Four apophyses wing-shaped,
nearly semicircular, with dentated edges and rough spinulate or porous
faces; their base occupies the middle third of the length of each spine.

_Dimensions._--Length of the spines 0.2 to 0.3, greatest breadth (diagonal
of the apophysial cross) 0.08 to 0.12.

_Habitat._--Tropical Atlantic, Stations 342 to 352, surface.

12. _Xiphacantha alata_, Haeckel.

_Xiphacantha alata_, Haeckel, 1862, Monogr. d. Radiol., p. 388.

_Acanthometra alata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 48, Taf. ix. figs. 1-3.

Spines four-winged, prismatic in the basal quarter, more slender in the
distal half. Apex simple pyramidal. Base with small wing-cross. Four
apophyses wing-shaped, nearly semicircular, with denticulated edges (and
often also with spinulate faces); their base occupies the second quarter of
the length of each spine.

_Dimensions._--Length of the spines 0.3 to 0.4, greatest breadth (diagonal
of the apophysial cross) 0.05 to 0.06.

_Habitat._--Mediterranean (Nice), Müller, (Portofino), Haeckel.

Genus 331. _Stauracantha_,[371] Haeckel, 1881, Prodromus, p. 465.

_Definition._--#Astrolonchida# with four branched (but not latticed)
apophyses on each radial spine, opposite in pairs in the form of a cross.

{762}The genus _Stauracantha_ differs from its ancestral form,
_Xiphacantha_, in the ramification of the four crossed apophyses on each
spine. These bear either lateral branches, which are parallel to the cross
axes of the radial spine itself (subgenus _Stauracanthonium_), or they are
forked, with divergent branches not parallel to those cross axes (subgenus
_Stauracanthidium_). Both subgenera may perhaps be better separated as
genera. They form the transition to the _Stauraspida_.

Subgenus 1. _Stauracanthonium_, Haeckel.

_Definition._--Apophyses of the radial spines not forked, but crossed by
perpendicular branches, which are placed in tangential planes and parallel
to the cross axes of the quadrangular spine itself.

1. _Stauracantha orthostaura_, n. sp. (Pl. 130, fig. 5).

Spines four-sided prismatic, with simple apex and small leaf-cross at the
base; with four thin slender conical apophyses in the proximal third. Each
apophysis represents a regular rectangular cross, being intersected in its
middle by one perpendicular rod of its own length.

_Dimensions._--Length of the spines 0.3, breadth 0.02; distance of the
apophyses from the apex 0.15 to 0.2.

_Habitat._--South Pacific, Station 169, surface.

2. _Stauracantha tetrastaura_, n. sp.

Spine four-winged prismatic, with four broad prominent edges, pyramidal
apex, and large basal leaf-cross, with four broad compressed apophyses
about in the middle. Each apophysis represents a vertical lamella (placed
in a meridian) with convex distal and concave proximal edge, and is crossed
in its apical part by one perpendicular short rod.

_Dimensions._--Length of the spines 0.5, breadth 0.03; distance of the
apophyses from the centre 0.2.

_Habitat._--Central Pacific, Station 272, surface.

3. _Stauracantha diplostaura_, n. sp.

Spines four-sided prismatic, with truncated apex and broad basal
leaf-cross; in the proximal half with four slender conical apophyses. Each
apophysis is crossed in its apical half at right angles by two short
parallel transverse rods.

_Dimensions._--Length of the spines 0.32, breadth 0.012; distance of the
apophyses from the centre 0.11.

_Habitat._--South Pacific, Station 291, surface.

{763}4. _Stauracantha scalaris_, n. sp.

Spines four-winged prismatic, with four broad prominent edges, truncated
apex, and small basal leaf-cross; in the proximal third with four
compressed lamellar apophyses. Each apophysis is pinnate with opposite
pinnulæ, or crossed at right angles by three to six parallel transverse
rods.

_Dimensions._--Length of the spines 0.24, breadth 0.016; central distance
of the apophyses 0.12.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

5. _Stauracantha johannis_, n. sp. (Pl. 132, fig. 11).

Spines four-sided pyramidal, with simple apex and broad basal leaf-cross,
with four large conical apophyses about in their middle. Each apophysis
bears two irregular rows of alternating perpendicular lateral branches, the
length of which decreases from the base towards the apex of the apophysis.

_Dimensions._--Length of the spines 0.2, middle breadth 0.02; central
distance of the apophyses 0.1.

_Habitat._--South-east Pacific (off Juan Fernandez), Station 300, surface.

6. _Stauracantha murrayana_, Haeckel.

_Xiphacantha species_, Wyville Thomson, 1877, The Atlantic, vol. i. p.
235, fig. 53.

_Xiphacantha murrayana_, Haeckel, 1878, Protistenreich, p. 45, fig. 33.

Spines quadrangular prismatic, with pyramidal apex and small basal
leaf-cross, with four regular conical apophyses in the proximal part. Each
apophysis is crossed at right angles by two parallel transverse rods; the
smaller apical rod is simple; the larger basal rod is crossed again on each
side by a secondary perpendicular branch, which therefore is parallel to
the apophysis itself.

_Dimensions._--Length of the spines 0.3, breadth 0.01, distance of the
apophyses 0.08.

_Habitat._--South Atlantic, Station 332, surface.

7. _Stauracantha pinnulata_, n. sp.

Spines four-winged pyramidal, with prominent edges, simple apex and large
basal leaf-cross, in the middle with doubly pinnate triangular apophyses.
Each apophysis is crossed at right angles by three to four transverse
parallel rods, which bear again perpendicular secondary branches; the
outline of the doubly pinnate apophysis is an isosceles triangle.

_Dimensions._--Length of the spines 0.3, middle breadth 0.02; distance of
the apophyses 0.12.

_Habitat._--North Pacific, Station 244, surface.

{764}Subgenus 2. _Stauracanthidium_, Haeckel.

_Definition._--Apophyses of the radial spines forked, each with two
divergent terminal branches, which are not parallel to the cross axes of
the quadrangular spine.

8. _Stauracantha stauraspis_, Haeckel.

_Dorataspis polyancistra, juvenis_, Haeckel, 1862, Monogr. d. Radiol., p.
418, Taf. xxi. fig. 7.

Spines slender, quadrangular prismatic, with simple apex and pyramidal base
(without basal leaf-cross); with four slender apophyses, which are simply
forked in the outer third. Each apophysis with two recurved fork branches.

_Dimensions._--Length of the spines 0.12, breadth 0.004; distance of the
apophyses 0.06.

_Habitat._--Mediterranean (Messina), surface.

9. _Stauracantha bifurca_, n. sp.

Spines stout, four-winged prismatic, with pyramidal apex and pyramidal base
(without leaf-cross); in their middle with four compressed, little curved
apophyses (convex on the outer, concave on the inner edge), which are
simply forked at the end. Each apophysis with two short straight fork
branches.

_Dimensions._--Length of the spines 0.2, breadth 0.006; distance of the
apophyses 0.08.

_Habitat._--Central Pacific, Station 266, surface.

10. _Stauracantha quadrifurca_, n. sp. (Pl. 130, fig. 4).

Spines slender, four-sided prismatic, with simple apex and pyramidal base
(without leaf-cross); with four slender apophyses, which are doubly forked
about in their middle. Each apophysis with four short and thin terminal
branches.

_Dimensions._--Length of the spines 0.2, breadth 0.002; distance of the
apophyses from the centre 0.08.

_Habitat._--South Atlantic, Stations 325 to 330, surface.

Genus 332. _Phatnacantha_,[372] Haeckel, 1881, Prodromus, p. 465.

_Definition._--#Astrolonchida# with four apophyses on each radial spine,
opposite in pairs in cross form, and forming a lattice-plate by
communicating branches.

The genus _Phatnacantha_ has been derived from the preceding _Stauracantha_
by concrescence of the branches of the apophyses. Therefore each spine
bears a lattice-plate or a fenestrated shield. If the growing plates of the
neighbouring spines {765}meet together, then the characteristic
lattice-shell of the Tessaraspida is perfect. Either each plate bears four
crossed pores (like _Tessaraspis_), or a larger number of pores (four
aspinal and four to eight or more coronal) like _Icosaspis_.

1. _Phatnacantha tessaraspis_, n. sp.

Spines quadrangular prismatic, with simple pyramidal apex and small basal
leaf-cross. Each spine bears in the basal half a square plate, which is
perforated by four square pores; margin of the plate with twelve short
straight teeth.

_Dimensions._--Length of the spines 0.12, breadth 0.005.

_Habitat._--Central Pacific, Station 265, surface.

2. _Phatnacantha icosaspis_, n. sp. (Pl. 130, fig. 6).

Spines quadrangular with prominent edges; prismatic in the basal half, with
a broad basal leaf-cross, pyramidal in the distal half, with a simple or
truncate apex. Each spine bears in the middle part a square plate, which is
perforated by eight to sixteen or more square pores (four aspinal and four
to eight or more coronal); margin of the plate with twenty-four to
forty-eight or more short straight irregular compressed teeth.

_Dimensions._--Length of the spines 0.18, breadth 0.008.

_Habitat._--Central Pacific, Station 273, surface.

Genus 333. _Pristacantha_,[373] n. gen.

_Definition._--#Astrolonchida# with numerous simple apophyses (eight to
sixteen or more on each radial spine) which are arranged in four
longitudinal rows opposite in pairs in the form of a cross.

The genus _Pristacantha_ differs from all other Stauracanthida in the
multiplication of the crossed apophyses (at least eight on each spine), and
exhibits therefore to them the same relation as _Astrolonche_ exhibits to
the other Phractacanthida. The remarkable _Astrolonchidium serratum_
appears intermediate between both groups.

1. _Pristacantha octodon_, n. sp. (Pl. 130, fig. 9).

Spines four-sided prismatic, a little broader in the middle part than at
either end, with thin prominent edges. Apex truncate or pyramidal. Base
pyramidal, with a small leaf-cross. From the four edges arise in the basal
half (between first and second third of the length) eight slender, conical,
or triangular apophyses (two on each edge).

_Dimensions._--Length of the spines 0.4 to 0.6, breadth 0.02 to 0.04.

_Habitat._--Central Pacific, Station 271, surface.

{766}2. _Pristacantha dodecodon_, n. sp. (Pl. 130, fig. 8).

Spines four-winged, of lanceolate outline, tapering from the broader middle
towards the two ends. Apex pyramidal. Base with a large leaf-cross. From
the four wings arise in the basal part (between first and second third of
the length) twelve triangular apophyses (three from each wing).

_Dimensions._--Length of the spines 0.3, breadth in the middle 0.03.

_Habitat._--North Pacific, Station 244, surface.

3. _Pristacantha polyodon_, n. sp. (Pl. 130, fig. 7).

Spines in the distal half four-sided prismatic, thin, with pyramidal apex;
in the basal half much broader, four-winged, with a large leaf-cross on the
base. From the four wings of the basal half arises a variable number of
slender teeth or triangular apophyses, commonly sixteen to twenty-four,
often irregular (four to six from each wing).

_Dimensions._--Length of the spines 0.3 to 0.4, breadth in the distal part
0.007 to 0.01, in the basal part 0.02 to 0.03.

_Habitat._--South Pacific (off New Zealand), Station 169, surface.

Family XXXVII. #QUADRILONCHIDA#, Haeckel (Pl. 131).

_Acanthostaurida_, Haeckel, 1881, Prodromus, p. 466.

_Definition._--ACANTHARIA with twenty radial spines of very unequal size,
disposed according to the law of the Icosacantha; four equatorial spines
much larger than the sixteen others. No lattice-shell.

The family #Quadrilonchida# differs from the foregoing ancestral family,
the Astrolonchida, in the unequal development of the twenty radial spines.
The four equatorial spines are constantly much larger, and often also of
another form and shape, than the sixteen other spines; often also among
these the eight tropical spines are larger and of another form than the
eight polar spines. Therefore the five parallel girdles or zones of every
four spines, which in the Astrolonchida are equal, are here distinctly
unequal. The whole body is flattened and compressed in the direction of the
spineless axis, so that the equatorial plane is larger than any other
plane, laid through the centre. In consequence of this flattening the
central capsule is also commonly compressed and flattened, lenticular or
discoidal, rarely spherical. In the Astrolonchida the "promorph" or the
"geometrical fundamental form" is constantly a square double pyramid, the
axes of which are of equal length. In the Quadrilonchida it becomes a
square double pyramid, the two equatorial axes of which (or the diagonals
of the square) are constantly longer than all other axes.

In the simpler forms of Quadrilonchida are found only two different kinds
of spines, the four larger equatorial spines being of the same size and
form, and the sixteen smaller {767}spines also not differing from one
another (_Acanthostaurus_, &c.). But in the majority of this family there
are three different kinds: four larger equatorial spines, eight tropical
spines of middle size, and eight smaller polar spines (_Belonostaurus_,
&c.); the latter become sometimes rudimentary, so that only twelve spines
are developed (four larger equatorial and eight smaller tropical spines).
In this case the development of the flat discoidal body is much stronger in
the equatorial plane than in all other planes. These discoidal or
lenticular Quadrilonchida exhibit a relation to the spherical Astrolonchida
similar to that which the #Discoidea# exhibit to the #Sphæroidea# among the
#Sphærellaria#.

A further morphological differentation takes place in the remarkable genera
_Lonchostaurus_ and _Zygostaurus_. In the former (Pl. 131, figs. 4-6) the
four larger equatorial spines becomes differentiated in pairs, so that the
opposite equal spines of one pair, _c_1, _c_3 (in the longitudinal or
hydrotomical axis), are larger, and often also of another form, than the
opposite equal spines of the other pair, _c_2, _c_4 (in the transverse or
geotomical axis). The most peculiar form is the rather common _Zygostaurus_
(Pl. 131, figs. 7, 8). Here the two opposite spines of one equatorial axis
(of the longitudinal axis) become very different, so that the anterior or
frontal spine (_c_1) is very unequal to the posterior or caudal spine
(_c_3), whereas the two opposite spines, of the other equatorial axis (of
the transverse axis) remain equal (_c_2, _c_4). Therefore the fundamental
forms become here "amphithect," as in the Ctenophora.

The numerous Quadrilonchida may be disposed in two different subfamilies:
in the Acanthostaurida all twenty radial spines are simple, without
apophyses; in the Lithopterida all twenty spines (or only one part of
them) bear two opposite apophyses (or lateral transverse processes). The
former correspond to the Zygacanthida, the latter to the Phractacanthida
among the Astrolonchida. The two opposite apophyses are simple in
_Quadrilonche_ (Pl. 133, fig. 1). In _Xiphoptera_ they are provided on
their distal side with lateral branches which are parallel to the spine
itself. In _Lithoptera_ the spines bear two to four parallel pairs of
transverse apophyses, and these are crossed by perpendicular branches,
parallel to the spine itself, so that there arise fenestrated wings or
latticed plates, comparable to the sails of a wind-mill. The lattice-work
of these plates lies in the same meridian plane with the radial spine
itself, and is therefore not comparable to the fenestrated apophyses of
_Doracantha_, of _Phatnacantha_, and of the Dorataspida; in these the
lattice-plates lie in tangential planes, perpendicular to the radial spine.

The apophyses of the Lithopterida may be developed either on all twenty
spines equally, or only on twelve spines (four equatorial and eight
tropical, whilst the eight polar spines are simple, Pl. 131, fig. 10), or
only on the four equatorial spines (whilst the sixteen others are simple,
Pl. 131, fig. 9).

_The Central Capsule_ of the Quadrilonchida is rarely spherical, commonly
more or {768}less compressed from both poles of the spineless axis,
lenticular or discoidal, sometimes square. It is enveloped by a voluminous
calymma constantly bearing coronals of "Myophrisca" (compare p. 724).

_Synopsis of the Genera of Quadrilonchida._

-------------------------------------------------------------------------
I. Subfamily Acanthostaurida.
All twenty spines simple, without lateral apophyses (sometimes forked,
but neither branched nor latticed).
-------------------------------------------------------------------------
{ Eight tropical and
{ eight polar spines
{ nearly equal, 334. _Acanthostaurus_.
Four equatorial spines of {
equal size and form. { Eight tropical and
{ eight polar spines
{ very different, 335. _Belonostaurus_.

{ Two principal spines
Four equatorial spines of { of equal size and
very different size or { form, 336. _Lonchostaurus_.
form (the two lateral {
constantly equal). { Two principal spines
{ (frontal and caudal)
{ very different, 337. _Zyostaurus_.
-------------------------------------------------------------------------
II. Subfamily Lithopterida.
Either all twenty spines or a part of them provided with two opposite
lateral branches or apophyses.
-------------------------------------------------------------------------
Apophyses simple, neither branched nor latticed, 338. _Quadrilonche_.

Apophyses branched or pinnate, but not latticed, 339. _Xiphoptera_.

Apophyses latticed, with fenestrated network, 340. _Lithoptera_.

Subfamily 1. ACASTHOSTAURIDA, Haeckel, 1881, Prodromus, p. 466.

_Definition._--#Quadrilonchida# with simple radial spines, without
apophyses.

Genus 334. _Acanthostaurus_,[374] Haeckel, 1862, Monogr. d. Radiol., p.
395.

_Definition._--#Quadrilonchida# with four equatorial spines of equal size
and form, which are much larger than the sixteen other spines. Eight
tropical and eight polar spines nearly equal. No apophyses.

The genus _Acanthostaurus_ is the most simple and primitive form of the
Quadrilonchida, and the common ancestral genus of this family; it is at the
same time its most common and widely distributed form. Some species appear
in astonishing numbers in different seas. It has been derived from
_Acanthometron_ by stronger development of the four equatorial spines,
which are all of equal size and much larger than the sixteen others.

{769}Subgenus 1. _Acostaurus_, Haeckel.

_Definition._--All twenty spines separated, in the centre united only by
the triangular faces or the meeting leaf-shaped edges of their pyramidal
bases.

1. _Acanthostaurus aequatorialis_, n. sp.

Spines cylindrical, of nearly equal breadth throughout their whole length.
Apex simple conical. Base with a large leaf-cross. Four equatorial spines
of the same form as the sixteen others, but much longer and about three
times as broad.

_Dimensions._--Length of the four major spines 0.2 to 0.25, breadth 0.008;
length of the sixteen minor spines 0.01 to 0.15, breadth 0.003.

_Habitat._--Equatorial Pacific, Station 271, surface.

2. _Acanthostaurus bipennis_, n. sp.

Spines linear, sword-shaped, strongly compressed, two edged; of nearly
equal breadth in their whole length. Apex bifid. Base with a small
leaf-cross. Four equatorial spines of the same form as the sixteen others,
but twice as long and as broad.

_Dimensions._--Length of the four major spines 0.4 to 0.5, of the sixteen
minor 0.2 to 0.3; breadth of the former 0.02, of the latter 0.01.

_Habitat._--North Pacific, Station 244, surface.

3. _Acanthostaurus conacanthus_, n. sp.

Spines conical, short, gradually tapering from the thick base towards the
simple apex. Base with large leaf-cross. Four equatorial spines more
cylindrical, twice to three times as long and as broad as the sixteen
others.

_Dimensions._--Length of the four major spines 0.15 to 0.2, of the sixteen
minor 0.05 to 0.08; basal breadth of the former 0.03, of the latter 0.012.

_Habitat._--South Atlantic, Station 332, surface.

4. _Acanthostaurus purpurascens_, Haeckel.

_Acanthostaurus purpurascens_, Haeckel, 1862, Monogr. d. Radiol., p. 395,
Taf. xix. figs. 1, 2.

_Acanthostaurus purpurascens_, R. Hertwig, 1879, Organism. d. Radiol.,
Taf. i. figs. 8, 9, Taf. iii. figs. 13, 15.

_Acanthometra purpurascens_, Haeckel, 1860, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 809.

Spines nearly cylindrical, with four blunt (often scarcely visible) edges,
tapering slightly from the central to the distal end. Apex bifid, with two
short parallel teeth. Base with a broad leaf-cross. {770}Four equatorial
spines one and a half times to twice as long and broad as the sixteen
others. Central capsule cruciate, with four arms (enveloping the basal part
of the four large spines); filled up with yellow bodies and purple
granules. Calymma large, with a network of purple granules. The four main
spines are constantly much larger than the sixteen others, but in variable
proportion. The eight polar spines are sometimes rudimentary.

_Dimensions._--Length of the four major spines 0.3 to 0.4, of the sixteen
minor 0.15 to 0.3; breadth of the former 0.012, of the later 0.008 or less.

_Habitat._--Mediterranean (Messina); Atlantic, Stations 348 to 354,
surface.

5. _Acanthostaurus pallidus_, Haeckel.

_Acanthostaurus pallidus_, Haeckel, 1862, Monogr. d. Radiol., p. 396.

_Acanthometra pallida_, Claparède et Lachmann, 1858, Études sur les
Infusoires, &c., p. 461, Taf. xxiv. fig. 6.

Spines four-sided prismatic, with four blunt edges, of nearly equal breadth
throughout their whole length. Apex bifid. Base with a small leaf-cross.
Four equatorial spines much larger than the sixteen others. Central capsule
spherical, transparent, containing some yellow bodies.

_Dimensions._--Length of the four main spines 0.12 to 0.2, of the sixteen
others 0.04 to 0.08; breadth of the former 0.01, of the latter 0.004.

_Habitat._--North Atlantic, west coast of Norway, Claparède; Færöe Channel,
John Murray.

6. _Acanthostaurus forceps_, Haeckel.

_Acanthostaurus forceps_, Haeckel, 1862, Monogr. d. Radiol., p. 396, Taf.
xix. figs. 3, 4.

_Acanthometra forceps_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 809.

Spines pincer-shaped, linear, compressed, cleft nearly throughout their
whole length into two parallel thin, straight branches, which are united
only at the pyramidal base and by means of a small bridge in the middle
part. Four main spines twice as long and as broad as the sixteen others.
Central capsule square, filled up with red pigment, and containing yellow
bodies. The diagonals of the square are the two equatorial axes.

_Dimensions._--Length of the four main spines 0.1, of the sixteen others
0.05; breadth of the former 0.01, of the latter 0.005.

_Habitat._--Mediterranean (Messina); North Atlantic, Canary Islands,
Station 354, surface.

7. _Acanthostaurus hastatus_, Haeckel.

_Acanthostaurus hastatus_, Haeckel, 1862, Monogr. d. Radiol., p. 397,
Taf. xix. fig. 5.

_Acanthometra hastata_, Haeckel, 1860, Monatsber. d. k. preuss. Akad. d.
Wiss. Berlin, p. 809.

Spines of very different size and form; four equatorial spines cylindrical
in the basal half, spear-shaped or lanceolate in the distal half, with thin
edges, little longer, but five to eight times as {771}broad as the sixteen
smaller spines, which are cylindrical, of equal breadth throughout their
whole length, with bifid apex. Base of all twenty spines pyramidal, without
leaf-cross. Central capsule yellow, spherical, or lenticular.

_Dimensions._--Length of the four major spines 0.066, breadth in the middle
0.005, on the apex 0.008; length of the sixteen smaller spines 0.054,
breadth 0.001.

_Habitat._--Mediterranean (Messina), surface.

Subgenus 2. _Staurolithium_, Haeckel, 1862, Monogr. d. Radiol., p. 401.

_Definition._--All twenty spines in the centre melted and grown together,
forming one single, star-like piece of acanthin.

8. _Acanthostaurus cruciatus_, Haeckel.

_Staurolithium cruciatum_, Haeckel, 1862, Monogr. d. Radiol., p. 401,
Taf. xx. fig. 6.

_Astrolithium cruciatum_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 811.

Spines cylindrical, of equal breadth throughout their whole length. Apex
simple conical. Four equatorial spines twice as long and broad as the
sixteen other spines. All twenty spines in the centre melted and grown
together, forming a single piece of acanthin. Central capsule spherical,
red-brown, opaque.

_Dimensions._--Length of the four equatorial spines 0.12, breadth 0.006;
length of the sixteen smaller spines 0.06, breadth 0.003.

_Habitat._--Mediterranean (Messina), surface.

Genus 335. _Belonostaurus_,[375] n. gen.

The genus _Belonostaurus_ differs from the preceding _Acanthostaurus_ in
the unequal size and form of the eight tropical and the eight polar spines;
the latter are much smaller than the former and often quite rudimentary, so
that the skeleton appears composed only of twelve spines, four larger
equatorial and eight smaller tropical spines. The central bases of the
polar spines are constantly preserved. The central capsule is a flattened
square disk.

1. _Belonostaurus quadratus_, n. sp.

Four equatorial spines little compressed, almost cylindrical in the basal
half, lanceolate, broader, with simple apex in the distal half. Eight
tropical spines a little shorter, but only half as broad,
{772}sword-shaped, two-edged, of equal breadth, with simple apex. Eight
polar spines very short, conical, also with simple apex. Base without
leaf-cross, a simple pyramid.

_Dimensions._--Length of the four equatorial spines 0.18, of the eight
tropical spines 0.14, of the eight polar spines 0.04.

_Habitat._--Central Pacific, Station 271, surface.

2. _Belonostaurus bicuspis_, n. sp.

Four equatorial spines compressed, almost lanceolate in the basal half,
bifid, with two large, bent, little divergent horns in the distal half.
Eight tropical spines nearly of the same form, but only half as large, with
much smaller horns. Eight polar spines rudimentary, very short, conical,
with simple apex. Base with a large leaf-cross.

_Dimensions._--Length of the four equatorial spines 0.2, of the eight
tropical spines 0.1, of the eight polar spines 0.02.

_Habitat._--Central Pacific, Station 263, surface.

Genus 336. _Lonchostaurus_,[376] Haeckel, 1862, Monogr. d. Radiol., p. 397
(_sensu emendato_).

_Definition._--#Quadrilonchida# with four equatorial spines of unequal size
and form; the principal spines (in the longitudinal axis) larger than the
lateral spines (in the transverse axis); both opposite spines of each pair
equal. Sixteen other spines much smaller (often the eight tropical larger
than the eight rudimentary polar spines). No apophyses.

The genus _Lonchostaurus_ differs from the two preceding genera in the
unequal size and shape of both equatorial pairs of spines, whilst the two
opposite spines of each pair are equal. Therefore the equatorial plane is
here a rhombus, not a square, as in the preceding genera. We encounter here
for the first time the differentiation of the two equatorial axes, the
longitudinal (or hydrotomical) and the transverse (or geotomical) axis. The
sixteen other spines are constantly smaller (often also the eight polar
more or less rudimentary).

1. _Lonchostaurus rhomboides_, n. sp. (Pl. 131, fig. 6).

Four equatorial spines cylindrical, twice to three times as broad as the
sixteen smaller spines, conical at the distal apex; the two opposite spines
of the longitudinal axis twice as long as those of the transverse axis.
Sixteen smaller spines conical at the base, with simple bristle-shaped
apex. The surface of the rhombical calymma, including the spines, was
covered in the figured specimen with small plates like those of
_Sphærocapsa_.

_Dimensions._--Length of the cross (longitudinal axis) 0.24, breadth
(transverse axis) 0.12.

_Habitat._--South Pacific, Station 302, depth 1450 fathoms.

{773}2. _Lonchostaurus lanceolatus_, n. sp.

Four equatorial spines compressed, lanceolate, tapering equally from the
middle towards the two ends; the two opposite spines of the longitudinal
axis one and a half times as long as those of the transverse axis. Sixteen
smaller spines compressed, linear, two-edged, half as long and only
one-fourth as broad as the two longitudinal spines.

_Dimensions._--Length of the equatorial cross 0.3, breadth 0.2.

_Habitat._--Central Pacific, Station 272, surface.

3. _Lonchostaurus bifidus_, n. sp.

Four equatorial spines compressed, nearly rectangular, forked, with two
long and thin, bent, little divergent horns; the two opposite spines of the
longitudinal axis about one-third longer and broader than those of the
transverse axis. Sixteen smaller spines of the same form, also forked, but
the eight tropical only half as large, and the eight polar spines scarcely
one-fourth as large as the two longitudinal spines.

_Dimensions._--Length of the equatorial cross 0.36, breadth 0.26.

_Habitat._--South-west Pacific, Station 166, surface.

4. _Lonchostaurus bifurcus_, n. sp. (Pl. 131, fig. 4).

Four equatorial spines nearly rhomboidal, about one and a half times as
large as the eight tropical and three times as large as the eight polar
spines; the two opposite spines of the longitudinal axis only with two
large divergent straight horns; the two spines of the transverse axis
somewhat shorter, with four short horns (the two outer horns longer than
the two inner). Eight tropical spines doubly forked, each with four thin
bristle-shaped horns. Eight polar spines very short, each with two thin
bristle-shaped, much divergent horns.

_Dimensions._--Length of the equatorial cross 0.34, breadth 0.28.

_Habitat._--North-west Pacific, Station 236, surface.

5. _Lonchostaurus rhombicus_, n. sp.

Four equatorial spines nearly rhomboidal, tapering from the broader middle
towards the two ends. Leaf-cross of their base very large. The two opposite
spines of the longitudinal axis about one-third larger than the two spines
of the transverse axis. Eight tropical spines of the same form, but only
half as large. Eight polar spines rudimentary, very short, conical.

_Dimensions._--Length of the equatorial cross 0.4, breadth 0.3.

_Habitat._--North Pacific, Station 253, surface.

6. _Lonchostaurus crystallinus_, n. sp. (Pl. 131, fig. 5).

Four equatorial spines lanceolate or rhomboidal, with broad leaf-cross at
the base, of a peculiar crystalline structure and a lamellated surface,
both opposite spines of the longitudinal axis twice as {774}large as the
two spines of the transverse axis. Eight tropical spines of similar form,
but only half as large, and with a very large, extremely prominent,
lamellated leaf-cross. Eight polar spines quite rudimentary, scarcely
prominent.

_Dimensions._--Length 0.36, breadth 0.2.

_Habitat._--Central Pacific, Station 271, surface.

Genus 337. _Zygostaurus_,[377] n. gen.

_Definition._--#Quadrilonchida# with four equatorial spines of unequal size
and form; both lateral spines (in the transverse axis) equal; but both
principal spines (in the longitudinal axis) very different, the frontal
larger than the caudal. Sixteen other spines much smaller (often the eight
tropical larger than the eight rudimentary polar spines). No apophyses.

The genus _Zygostaurus_ differs from all other Staurolonchida in the
peculiar differentiation of the four equatorial spines; the two opposite
spines of the hydrotomical or longitudinal axis being very different in
size and form (the frontal spine forked, the caudal spine simple); whilst
the two opposite spines of the geotomical or lateral axis (perpendicular to
the former) are equal, forked, but different in shape from the former.
Therefore the geometrical fundamental form of the body in this remarkable
genus becomes "amphithect" or "bilateral" in the widest signification of
this term (comp. my General Morphology, vol. i. pp. 480, 482). Of the three
different dimensive axes one (the longitudinal) exhibits unequal poles,
whilst the two other (the sagittal and lateral) axes exhibit equal poles.

1. _Zygostaurus amphithectus_, n. sp. (Pl. 131, fig. 7).

Frontal spine (_c_1) little different from the two lateral spines; each
with two divergent curved horns, which are equal and of about the same
length as the simple broad basal part. Caudal spine (_c_3) simple,
spindle-shaped, about as long as the frontal spine. Tropical spines little
smaller than the three former, and of the same symmetrical forked form.
Eight polar spines much smaller, but also forked, with thin bristle-shaped
branches.

_Dimensions._--Length of the cross (longitudinal axis) 0.5, breadth
(lateral axis) 0.4.

_Habitat._--Central Pacific, Station 272, surface.

2. _Zygostaurus longicornis_, n. sp.

Frontal spine (_c_1) little different from the two lateral spines, each
with two divergent curved horns, which are equal and of about the same
length as the simple broad basal part. Caudal {775}spine (_c_3) simple,
lanceolate, shorter than the frontal spine. Eight tropical spines in the
basal part smaller than the three former, but with much longer
fork-branches, which are prolonged into very thin and long bristles. Eight
polar spines rudimentary, very short.

_Dimensions._--Length of the equatorial cross 0.4, breadth 0.3.

_Habitat._--South Pacific, Station 295, surface.

3. _Zygostaurus cornutus_, n. sp.

Frontal spine (_c_1) little different from the two lateral spines, each
with two divergent straight horns, which are shorter than the basal part.
Caudal spine (_c_3) simple, triangular, shorter than the three former.
Eight tropical spines of equal size and form, doubly forked, each with four
thin, bristle-shaped, little divergent teeth. Eight polar spines short,
simply forked.

_Dimensions._--Length of the equatorial cross 0.5, breadth 0.4.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

4. _Zygostaurus caudatus_, n. sp.

Frontal spine (_c_1) very different from the others, pincer-shaped, with
two long, nearly parallel slightly bent horns. Lateral spines (_c_2 and
_c_4) only half as long, with two short, nearly parallel horns. Caudal
spine (_c_3) very long and stout, spindle-shaped, about twice as long as
the frontal spine. Eight tropical spines of equal size and form,
symmetrical, with two long and thin, divergent horns. Eight polar spines
short and stout, with two divergent horns.

_Dimensions._--Length of the equatorial cross 0.8, breadth 0.3.

_Habitat._--Central Pacific, Station 265, surface.

5. _Zygostaurus frontalis_, n. sp.

Frontal spine (_c_1) very different from the others, with two very long,
divergent, stout branches. Lateral spines (_c_2 and _c_4) only half as
long, each with two stout, nearly parallel horns of equal length. Caudal
spine (_c_3) simple, sword-like, shorter than the three former. Eight
tropical spines of equal size and form, symmetrical, of the same shape as
the two lateral, but only half as large. Eight polar spines very small,
rudimentary, each with two short teeth.

_Dimensions._--Length of the equatorial cross 0.7, breadth 0.5.

_Habitat._--Central Pacific, Station 274, surface.

6. _Zygostaurus sagittalis_, n. sp. (Pl. 131, fig. 8).

Frontal spine (_c_1) very different from the others, with two equal,
strongly divergent, bent horns. Lateral spines (_c_2 and _c_4) little
smaller, but with two very unequal horns (the anterior shorter than the
posterior). Caudal spine (_c_3) simple, sword-like or triangular,
two-edged, longer {776}than the frontal spine. Eight tropical spines
different, the four anterior (_b_1, _b_4, _d_1, _d_4) smaller, with longer
horns; the four posterior (_b_2, _b_3, _d_2, _d_3) broader, with shorter
horns; the anterior horn of each tropical spine is longer than the
posterior. Eight polar spines rudimentary, simple, very short.

_Dimensions._--Length of the equatorial spine-cross 0.6, breadth 0.4.

_Habitat._--Central Pacific, Station 271, surface.

Subfamily 2. LITHOPTERIDA, Haeckel.

_Definition._--#Quadrilonchida# with two opposite transverse apophyses
either on all twenty spines or only on a part of them.

Genus 338. _Quadrilonche_,[378] n. gen.

_Definition._--#Quadrilonchida# with two simple opposite apophyses either
on each radial spine or only on a part of the twenty spines.

The genus _Quadrilonche_ is the ancestral form of the Lithopterida, or of
those Quadrilonchida which bear two opposite apophyses or transverse
processes. In _Quadrilonche_ these apophyses are simple, whilst they are
branched in _Xiphoptera_ and fenestrated in _Lithoptera_. Each of these
three genera may be divided into three subgenera; in the first only the
four equatorial spines are provided with apophyses, in the second twelve
spines (four equatorial and eight tropical), in the third subgenus all
twenty spines.

Subgenus 1. _Quadriloncharium_, Haeckel.

_Definition._--Four large equatorial spines provided with transverse
apophyses; sixteen other smaller spines simple, without apophyses.

1. _Quadrilonche tetrastaura_, n. sp.

Four equatorial spines very large, compressed, two-edged, each crossed in
the distal third by two opposite simple apophyses. Sixteen other spines
also compressed, linear, somewhat shorter than the former and only half as
broad, without apophyses.

_Dimensions._--Length of the four major spines 0.12, breadth 0.02; length
of the sixteen minor spines 0.08, breadth 0.01.

_Habitat._--Central Pacific, Station 266, surface.

{777}2. _Quadrilonche platystaura_, n. sp. (Pl. 131, fig. 2).

Four equatorial spines lanceolate, compressed, with two opposite triangular
simple apophyses in the broadest middle part. Sixteen other spines also
lanceolate, much smaller, of about half the length, but only of one-fourth
the breadth, without apophyses.

_Dimensions._--Length of the four major spines 0.2, breadth 0.018; length
of the sixteen minor spines 0.1, breadth 0.004.

_Habitat._--North Pacific, Station 235, surface.

Subgenus 2. _Quadrilonchidium_, Haeckel.

_Definition._--All twenty spines provided with transverse apophyses.

3. _Quadrilonche mesostaura_, n. sp. (Pl. 131, fig. 1).

Four equatorial spines almost of the some form and length as the sixteen
others, but of double or triple the breadth; each spine in its middle third
with two opposite broad triangular apophyses, in the basal third
rectangular, compressed, in the distal third isosceles triangular, with
simple apex.

_Dimensions._--Length of the four major spines 0.16, breadth 0.01 to 0.015;
length of the sixteen minor spines 0.12, breadth 0.005.

_Habitat._--Tropical Atlantic, Station 345, surface.

4. _Quadrilonche telostaura_, n. sp.

Four equatorial spines one and a half times as long and three times as
broad as the sixteen others; all twenty spines cylindrical, of equal
breadth throughout their whole length, with simple conical apex; each spine
crossed in the distal third by two opposite, simple, conical apophyses.

_Dimensions._--Length of the four major spines 0.3, breadth 0.012; length
of the sixteen minor spines 0.2, breadth 0.004.

_Habitat._--South Atlantic, Station 325, surface.

Genus 339. _Xiphoptera_,[379] Haeckel, 1881, Prodromus, p. 466.

_Definition._--#Quadrilonchida# with two opposite branched (but not
latticed) apophyses, either on each radial spine or only on a part of the
twenty spines.

The genus _Xiphoptera_ differs from the preceding ancestral genus
_Quadrilonche_ in the ramification of the apophyses, which bear
perpendicular branches on their distal side. These are therefore parallel
to the spine itself. If the branches become united by transverse beams then
we get _Lithoptera_.

{778}1. _Xiphoptera tessaractena_, n. sp.

Four equatorial spines in the outer third crossed by two opposite
transverse apophyses, each of which bears on its distal side two to three
branches, perpendicular to the apophysis and parallel to the spine itself.
Sixteen other spines much smaller, in the outer third crossed by two simple
opposite transverse apophyses.

_Dimensions._--Length of the four major spines 0.18, of the sixteen minor
0.05 to 0.1.

_Habitat._--North Pacific, Station 238, surface.

2. _Xiphoptera dodecactena_, n. sp. (Pl. 131, fig. 3).

Four equatorial spines one and a half times to twice as long and broad as
the eight tropical spines. These twelve spines have the same form and are
crossed in their distal third by two large opposite transverse apophyses,
each of which bears on its distal side two to four branches, perpendicular
to the apophysis and parallel to the spine. Eight polar spines much smaller
than the twelve others, simple, without apophyses. The central capsule of
this species exhibited a conical protuberance around the base of each
individual spine.

_Dimensions._--Length of the twelve larger spines 0.1 to 0.2, of the eight
smaller 0.02 to 0.06.

_Habitat._--Central Pacific, Station 272, surface.

3. _Xiphoptera icosactena_, n. sp.

Four equatorial spines about twice as long and four times as broad as the
sixteen other spines. All twenty spines crossed in the outer third by two
large opposite transverse apophyses, each of which bears on its distal side
two to four branches perpendicular to the apophysis and parallel to the
spine itself.

_Dimensions._--Length of the four equatorial spines 0.26, of the sixteen
smaller 0.11 to 0.14.

_Habitat._--South Pacific, Station 288, surface.

Genus 340. _Lithoptera_,[380] J. Müller, 1858, Monatsber. d. k. preuss.
Akad. d. Wiss. Berlin, p. 155.

_Definition._--#Quadrilonchida# with two opposite branched and latticed
apophyses, either on each radial spine or only on a part of the twenty
spines.

The genus _Lithoptera_, founded by Johannes Müller in 1858 for the first
observed Mediterranean species, _Lithoptera fenestrata_, differs from all
other Quadrilonchida in the fenestrated form of the apophyses, which he
compared to the sails of a windmill. This peculiar fenestration is effected
by two to four parallel pairs of opposite apophyses, which are crossed by
perpendicular branches, parallel to the spine itself. {779}Therefore the
wings or lattice-plates have quadrangular meshes and lie in one meridian
plane of the spine, not in a tangential plane (as in the _Acanthophracta_).
Commonly the lateral ends of the four broad equatorial wings are so crossed
that one lateral corner of each wing lies on the upper, the other corner on
the under side of both its neighbours; but sometimes the meeting corners
have grown together.

Subgenus 1. _Lithopteranna_, Haeckel.

_Definition._--Four equatorial spines with transverse apophyses; sixteen
others (eight tropical and eight polar spines) simple, without apophyses.

1. _Lithoptera tetraptera_, n. sp. (Pl. 131, fig. 9).

Four equatorial spines spindle-shaped, with latticed apophyses, each
crossed by two transverse beams which are connected at equal distances by
four rods parallel to the spine (therefore each wing with four square
meshes in a single row). Lateral corners of the neighbouring wings not
meeting. Sixteen smaller spines (eight tropical and eight polar) simple,
conical, without apophyses.

_Dimensions._--Diagonal of the square body 0.24; length of the inner square
0.16.

_Habitat._--Tropical Atlantic, Station 348, surface.

2. _Lithoptera mülleri_, Haeckel.

_Lithoptera mülleri_, Haeckel, 1862, Monogr. d. Radiol., p. 398, Taf. xx.
figs. 1, 2.

Four equatorial spines four-edged, with large latticed apophyses, each
crossed by three transverse beams, which are connected at irregular
distances by eight to twelve rods parallel to the spine (therefore each
wing with two rows of rectangular meshes). The wings are placed not
perfectly in the equatorial plane, but a little obliquely, so that each
wing lies with one lateral corner on the upper, with the other corner on
the under side of its neighbours. Sixteen smaller spines simple, thin,
cylindrical, without apophyses.

_Dimensions._--Diagonal of the square body 0.3; length of the inner square
0.17.

_Habitat._--Mediterranean (Messina), surface.

3. _Lithoptera lamarckii_, n. sp.

Four equatorial spines four-edged, with large latticed apophyses; each
crossed by three transverse beams, which are connected at short distances
by twelve to sixteen rods parallel to the spine (therefore each wing with
two rows of hexagonal or nearly elliptical meshes). Wings placed as in
_Lithoptera mülleri_. Sixteen smaller spines simple, thin, prismatic,
without apophyses.

_Dimensions._--Diagonal of the square body 0.35; length of the inner square
0.2.

_Habitat._--Central Pacific, Station 271, surface.

{780}4. _Lithoptera tetragona_, n. sp.

Four equatorial spines compressed, four-edged, with large latticed
apophyses, each crossed by four transverse beams, which are connected at
regular distances by eight to twelve rods parallel to the spine. (Therefore
each wing with three rows of square meshes.) Wings placed in the equatorial
plane and grown together by their meeting lateral ends, forming a square
equatorial girdle of lattice-work. Sixteen smaller spines simple, thin,
prismatic, without apophyses.

_Dimensions._--Diameter of the square body 0.4; length of the inner square
0.22.

_Habitat._--North Pacific, Station 244, surface.

Subgenus 2. _Lithopterella_, Haeckel.

_Definition._--Twelve spines (four equatorial and eight tropical) with
transverse apophyses; the eight polar spines simple, without apophyses.

5. _Lithoptera quadrata_, n. sp. (Pl. 131, fig. 10).

Twelve spines with transverse apophyses; eight (polar) spines simple,
small, without apophyses. Four equatorial spines very large and stout,
compressed, each crossed by four transverse beams, which are connected by
eight to ten rods parallel to the spine (therefore each wing with three
rows of irregular rectangular meshes). Wings placed in the equatorial plane
and grown together by their meeting lateral ends, forming a square
equatorial girdle of lattice-work. Eight tropical spines thin, crossed by a
long and thin transverse beam, which bears on its distal side eight to ten
rods parallel to the spine.

_Dimensions._--Diagonal of the square body 0.35; length of the inner square
0.18.

_Habitat._--South Atlantic, Station 326, surface.

6. _Lithoptera dodecaptera_, n. sp.

Twelve spines with transverse apophyses; eight (polar) spines simple,
small, without apophyses. Four equatorial spines very large, quadrangular;
each crossed by three transverse beams, which are connected by ten to
twelve rods parallel to the spine (therefore each wing with two rows of
square meshes). Wings placed as in _Lithoptera mülleri_. Eight tropical
spines much thinner, crossed each by two transverse beams, which are
connected by six to eight rods parallel to the spine (therefore each wing
with one row of rectangular meshes).

_Dimensions._--Diagonal of the square body 0.4; length of the inner square
0.25.

_Habitat._--Central Pacific, Station 271, surface.

Subgenus 3. _Lithopteromma_, Haeckel.

_Definition._--All twenty spines with transverse apophyses.

{781}7. _Lithoptera darwinii_, Haeckel.

_Lithoptera darwinii_, Haeckel, 1879, Natürl. Schöpfungsgesch., Aufl.
vii. p. 706, Taf. xvi. fig. 12.

All twenty spines with transverse apophyses; these are quite simple in the
eight small cross-shaped polar spines; also simple, but bearing some
perpendicular rods, in the eight thin tropical spines. Four equatorial
spines very large and stout, each crossed by four transverse beams, which
are connected by eight to ten rods parallel to the spine (therefore each
wing with three rows of irregular rectangular meshes). Wings placed in the
equatorial plane and grown partly together by their meeting lateral ends.

_Dimensions._--Diagonal of the square body 0.4; length of the inner square
0.2.

_Habitat._--South-east Pacific, west coast of Patagonia, Station 302,
surface.

8. _Lithoptera icosaptera_, n. sp.

All twenty spines with transverse apophyses, bearing one to three rows of
rectangular meshes; eight polar spines small, each with a single row; eight
tropical spines long and thin, each with two parallel rows. Four equatorial
spines much larger and thicker, each with three parallel transverse rows of
meshes. All twenty wings free, not grown together by their lateral ends.

_Dimensions._--Diagonal of the square body 0.48; length of the inner square
0.24.

_Habitat._--Indian Ocean, Belligemma, Ceylon (Haeckel), surface.

9. _Lithoptera fenestrata_, J. Müller.

_Lithoptera fenestrata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 154, Taf. xi. fig. 13.

All twenty spines with transverse apophyses, bearing one to three rows of
irregular, rectangular meshes; eight polar and eight tropical spines small,
each with a single row of meshes (between two parallel transverse beams).
Four equatorial spines little larger, each with two rows (between three
parallel transverse beams). Central capsule filled up with green pigment.

_Dimensions._--Diagonal of the square body 0.3; length of the inner square
0.2.

_Habitat._--Mediterranean (Saint Tropez, Nice).

Family XXXVIII. #AMPHILONCHIDA#, Haeckel (Pl. 132).

_Acantholonchida_ (Amphilonchida et Amphilithida), Haeckel, 1881,
Prodromus, p. 466.

_Definition._--ACANTHARIA with twenty radial spines of very unequal size,
disposed according to the law of the Icosacantha; two opposite equatorial
spines (in the longitudinal axis) much larger than the eighteen others. No
lattice-shell.

The family #Amphilonchida# is distinguished from all other #Acanthometra#
by the excessive development of two opposite equatorial spines, which are
much larger {782}than the eighteen other spines. In my Monograph (1862, p.
389) I had united all these Acanthonida in a single genus _Amphilonche_
(with ten species). Some species of it are very common and widely
distributed; but in general the number of different forms in this family is
much smaller than in the two foregoing families.

The two principal spines, which in all Amphilonchida are much larger than
the eighteen other spines, characterise the "hydrotomical axis" or the
larger equatorial axis. The two other equatorial spines or the "geotomical
spines" are much smaller, and commonly of the same size as the eight
tropical and the eight polar spines. In the genera _Amphilonche_ and
_Amphibelone_ these eighteen smaller spines are rather equally developed;
sometimes they are very small or quite rudimentary, so that the skeleton
seems to be represented only by the two very long principal spines (Pl.
132, fig. 4). The genus _Acantholonche_ is distinguished by the unequal
size of the eight tropical and the eight polar spines, the latter being
more or less rudimentary. However, the central bases of all twenty spines,
by which they are united in the centre, are constantly present. The genus
_Amphibelone_ is distinguished by the unequal size and form of the two
principal spines, one of them the "caudal spine," being larger (and often
of another form) than the opposite "frontal spine."

The form of the radial spines is in the Amphilonchida far less varied and
complicated than in the other Acanthonida (the Astrolonchida and
Quadrilonchida). Apophyses or lateral transverse processes are never
developed. The three main forms of spines are the same as in the other
Acanthonida; they are (1) either cylindrical or conical (like
_Acanthometron_), or (2) compressed or two-edged (like _Zygacantha_), or
(3) quadrangular or four-edged (like _Acanthonia_). Often the spines are
angular in the inner or proximal, roundish in the outer or distal part. The
distal apex is commonly simple, conical or pyramidal. The central base is
commonly also pyramidal, as in the majority of the Acanthonida; and the
triangular faces of the neighbouring bases are simply propped one upon
another. More rarely a basal leaf-cross is developed above the basal
pyramid. Very rarely the central bases of the united spines grow together
in the centre, so that the whole skeleton forms a single piece of acanthin.

_The Central Capsule_ is rarely spherical, commonly prolonged in the
direction of the hydrotomical axis; ellipsoidal or cylindrical, sometimes
also four-sided prismatic; it commonly envelops the greater part of the two
principal spines; its structure and contents are the same as in the other
Acanthonida.

_Synopsis of the Genera of Amphilonchida._

{ Two opposite large
{ principal spines
Eighteen smaller spines { (frontal and caudal)
of nearly equal size { equal, 341. _Amphilonche_.
and similar form. {
{ Caudal spine larger than
{ the frontal spine, 342. _Amphibelone_.

Eight tropical and eight polar spines of different
sizes (the latter often rudimentary), 343. _Acantholonche_.

{783}Genus 341. _Amphilonche_,[381] Haeckel, 1862, Monogr. d. Radiol., p.
389.

_Definition._--#Amphilonchida# with two equal principal spines (frontal and
caudal spines not different); the eighteen smaller spines nearly equal.

The genus _Amphilonche_ represents the original and at the same time the
most common form of Amphilonchida; the two opposite principal spines are of
equal size and form, much larger (and often also of another form) than the
eighteen smaller spines; these latter exhibit no marked differences in size
and form.

Subgenus 1. _Amphiloncharium_, Haeckel.

_Definition._--Spines in the centre united by triangular faces of their
pyramidal bases, propped one upon another. No basal leaf-cross.

1. _Amphilonche belonoides_, Haeckel.

_Amphilonche belonoides_, Haeckel, 1862, Monogr. d. Radiol., p. 392, Taf.
xvi. fig. 6, Taf. xviii. fig. 21.

_Acanthometra belonoides_, Haeckel, 1860, Monatsber. d. k. Akad. d. Wiss.
Berlin, p. 808.

Two principal spines cylindrical, of equal breadth throughout their whole
length, with simple conical apex; base a small pyramid, without leaf-cross.
Eighteen smaller spines conical, much shorter, with simple bristle-shaped
apex. Central capsule spindle-shaped or cylindrical, yellow.

_Dimensions._--Length of the two major spines 0.4 to 0.8, of the eighteen
minor 0.05 to 0.2.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

2. _Amphilonche lanceolata_, n. sp. (Pl. 132, fig. 1).

Two principal spines compressed, lanceolate, two-edged, gradually tapering
from the broader middle towards the two ends; apex simple; base a large
pyramid, with broad leaf-cross. Eighteen smaller spines about half as long,
pyramidal, with simple conical or bristle-shaped apex. Central capsule
lentelliptical, pink, opaque.

_Dimensions._--Length of the two major spines 0.2, of the eighteen minor
0.1.

_Habitat._--Tropical Atlantic, Station 343, surface.

3. _Amphilonche diodon_, n. sp. (Pl. 132, fig. 3).

Two principal spines compressed, lanceolate, two-edged, tapering from the
broader middle towards the two ends; apex simple; base a small pyramid,
without leaf-cross. Eighteen smaller spines {784}sword-shaped, only
two-thirds or one-half as long, and one-third or one-fourth as broad, each
with two opposite teeth in the distal part. Central capsule ellipsoidal.

_Dimensions._--Length of the two major spines 0.13, breadth 0.02; length of
the eighteen minor spines 0.1.

_Habitat._--North Atlantic, Station 352, surface.

4. _Amphilonche tenuis_, Haeckel.

_Amphilonche tenuis_, Haeckel, 1862, Monogr. d. Radiol., p. 389, Taf.
xvi. fig. 1, Taf. xviii. fig. 16.

_Acanthometra tenuis_, Haeckel, 1860, Monatsber. d. k. Akad. d. Wiss.
Berlin, p. 807.

Two principal spines quadrangular prismatic, with smooth edges, slightly or
not at all prominent, of equal breadth throughout their whole length; apex
simple or bifid; base a small pyramid, without leaf-cross. Eighteen smaller
spines of the same form, but only one-fourth or one-third as long. Central
capsule spherical or ellipsoidal, colourless.

_Dimensions._--Length of the two major spines 0.2 to 0.3, of the eighteen
minor spines 0.05 to 0.1.

_Habitat._--Mediterranean (Messina); Atlantic, Station 354, Canary Islands,
surface.

5. _Amphilonche denticulata_, Haeckel.

_Amphilonche denticulata_, Haeckel, 1862, Monogr. d. Radiol., p. 390,
Taf. xvi. fig. 2, Taf. xviii. fig. 17.

Two principal spines quadrangular prismatic, with prominent, elegantly
denticulated edges, of equal breadth throughout their whole length; apex
simple or truncated; base a small pyramid, without leaf-cross. Eighteen
smaller spines of the same form, but only one-fourth or one-third as long.
Central capsule ellipsoidal, opaque, yellowish-brown.

_Dimensions._--Length of the two major spines 0.3 to 0.4, of the eighteen
minor 0.1; breadth of the former 0.009, of the latter 0.006.

_Habitat._--Mediterranean (Messina), Haeckel, surface.

6. _Amphilonche heteracantha_, Haeckel.

_Amphilonche heteracantha_, Haeckel, 1862, Monogr. d. Radiol., p. 293,
Taf. xvi. fig. 7.

_Acanthometra heteracantha_, Haeckel, 1860, Monatsber. d. k. Akad. d.
Wiss. Berlin, p. 808.

Two principal spines quadrangular prismatic, very large, with four broad,
prominent, lamellar edges, of equal breadth throughout their whole length.
Apex as well as the base a four-sided pyramid. Eighteen smaller spines very
thin, conical, with bristle-shaped apex; base a small pyramid, without
leaf-cross. Central capsule cylindrical or violin-shaped, opaque yellow.

_Dimensions._--Length of the two major spines 0.2 to 0.3, breadth 0.015 to
0.02; length of the eighteen minor spines 0.1 to 0.15, basal breadth 0.004
to 0.008.

_Habitat._--Mediterranean (Messina); Central Pacific, Station 271, surface.

{785}7. _Amphilonche elongata_, Haeckel.

_Amphilonche elongata_, Haeckel, 1862, Monogr. d. Radiol., p. 392, Taf.
xviii. figs. 22_a_, 22_b_.

_Acanthometra elongata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 48, Taf. vii. fig. 13.

Two principal spines quadrangular prismatic in the proximal half,
cylindrical or spindle-shaped in the distal half, with simple apex; base a
small pyramid, without leaf-cross. Eighteen smaller spines very thin,
bristle-shaped, conical on the base, with small pyramid. Central capsule
spindle-shaped, transparent, yellow.

_Dimensions._--Length of the two major spines 0.3 to 0.5, breadth 0.01 to
0.03; length of the eighteen smaller spines 0.05 to 0.3.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian, Pacific,
surface.

Subgenus 2. _Amphilonchidium_, Haeckel.

_Definition._--Spines at the central base with a broad leaf-cross, composed
of four prominent triangular lamellæ; between the meeting edges of the
latter twenty-two pyramidal compartments or basal funnels.

8. _Amphilonche ovata_, Haeckel.

_Amphilonche ovata_, Haeckel, 1862, Monogr. d. Radiol., p. 390.

_Acanthometra ovata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 47, Taf. vii. fig. 10, Taf. ix. fig. 4.

Two principal spines cylindrical, of equal breadth throughout their whole
length, with simple conical or bifid apex; base with a large leaf-cross of
double the breadth. Eighteen smaller spines of the same form, but thinner
and only half as long. Central capsule ellipsoidal, brown, opaque.

_Dimensions._--Length of the two major spines 0.3 to 0.4, breadth 0.008 to
0.012; length of the eighteen minor spines 0.1 to 0.2.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

9. _Amphilonche conica_, n. sp. (Pl. 132, fig. 6).

Two principal spines conical, elongate, with simple apex; base thickened,
with large leaf-cross. Eighteen smaller spines also conical, of about the
same basal breadth, but only one-fourth to one-half as long. Central
capsule ellipsoidal, pellucid.

_Dimensions._--Length of the two major spines 0.15 to 0.2, basal breadth
0.03 to 0.04; length of the eighteen minor spines 0.05 to 0.1.

_Habitat._--Central Pacific, Stations 265 to 274, surface.

{786}10. _Amphilonche lancetta_, n. sp.

Two principal spines compressed, lanceolate, two edged, gradually tapering
from the broader middle towards the two ends; apex simple; base of double
the breadth, with a large leaf-cross. Eighteen smaller spines of the same
form, but shorter and only half as broad. Central capsule lentelliptical,
pellucid.

_Dimensions._--Length of the two major spines 0.2 to 0.3, breadth in the
middle part 0.02 to 0.03; length of the eighteen smaller spines 0.1 to 0.2.

_Habitat._--South Pacific, Station 288, surface.

11. _Amphilonche complanata_, Haeckel.

_Amphilonche complanata_, Haeckel, 1862, Monogr. d. Radiol., p. 390, Taf.
xvi. fig. 3, Taf. xviii. fig. 18, _a_, _b_.

_Acanthometra complanata_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 808.

Two principal spines compressed, two-edged, linear, of equal breadth
throughout their whole length; apex emarginate or bifid; base with a large
leaf-cross of double the breadth. Eighteen smaller spines of the same form,
but only one-fourth to one-half as large. Central capsule ellipsoidal,
yellow, pellucid.

_Dimensions._--Length of the two major spines 0.15 to 0.25, breadth 0.004
to 0.006; length of the eighteen minor spines 0.05 to 0.1.

_Habitat._--Mediterranean (Messina), surface.

12. _Amphilonche messanensis_, Haeckel.

_Amphilonche messanensis_, Haeckel, 1862, Monogr. d. Radiol., p. 391,
Taf. xvi. fig. 4, Taf. xviii. fig. 19.

_Acanthometra messanensis_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 808.

Two principal spines quadrangular prismatic, often a little compressed from
two sides, of nearly equal breadth throughout their whole length; apex
either truncate or emarginate, with two opposite teeth; base with a large
leaf-cross of double the breadth. Eighteen smaller spines of similar form
or more compressed, much shorter. Central capsule spherical or ellipsoidal,
yellow, transparent.

_Dimensions._--Length of the two major spines 0.12 to 0.18, breadth 0.016;
length of the minor spines 0.05 to 0.09.

_Habitat._--Mediterranean (Messina, Corfu), Haeckel, surface.

13. _Amphilonche hydrotomica_, n. sp. (Pl. 132, fig. 2).

Two principal spines quadrangular prismatic, with four broad prominent
lamellar edges or wings, of equal breadth throughout their whole length;
apex pyramidal; base with a large leaf-cross. Eighteen smaller spines
cylindrical or bristle-shaped, only half as long and very thin. Central
capsule cylindrical or spindle-shaped, very long, opaque.

{787}_Dimensions._--Length of the two major spines 0.1 to 0.2, breadth 0.02
to 0.03; length of the eighteen minor spines 0.05 to 0.1.

_Habitat._--Central Pacific, Stations 270 to 274, surface.

14. _Amphilonche violina_, n. sp. (Pl. 132, fig. 5).

Two principal spines quadrangular prismatic, nearly violin-shaped, with
four very broad, prominent, lamellar wings, which are constricted in the
middle part, and broadened towards the two ends; apex truncate pyramidal;
base with a large leaf-cross. Eighteen smaller spines much shorter, of
equal breadth at the base, assuming the form of a quadrangular pyramid,
thin prismatic in the distal half. Central capsule spindle-shaped, opaque.

_Dimensions._--Length of the two major spines 0.15 to 0.18, breadth 0.02 to
0.025; length of the eighteen minor spines 0.04 to 0.08.

_Habitat._--Central Pacific, Station 271, surface.

15. _Amphilonche tetraptera_, Haeckel.

_Amphilonche tetraptera_, Haeckel, 1862, Monogr. d. Radiol., p. 391, Taf.
xvi. fig. 5, Taf. xviii. fig. 20.

_Acanthometra tetraptera_, Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 808.

Two principal spines four-sided pyramidal, with four broad lamellar
prominent edges; apex simple or bifid; base with a large leaf-cross.
Eighteen smaller spines of similar form, but only half as large. Central
capsule spherical, opaque.

_Dimensions._--Length of the two major spines 0.2, breadth on the base
0.025; length of the eighteen minor spines 0.1.

_Habitat._--Mediterranean (Messina); North Atlantic, Canary Islands,
Stations 352 to 354, surface.

Subgenus 3. _Amphilithium_, Haeckel, 1881, Prodromus, p. 466.

_Definition._--Spines in the basal part grown together, so that the whole
skeleton represents a single piece of acanthin; a star with two larger and
eighteen smaller rays.

16. _Amphilonche concreta_, n. sp. (Pl. 132, figs. 4, 4_a_).

Two principal spines cylindrical, very long, of equal breadth throughout
their whole length, with simple conical apex. Eighteen smaller spines
short, conical or bristle-shaped, scarcely one-fourth or one-tenth as long,
often quite rudimentary. All twenty spines perfectly grown together in the
centre, forming a single piece of acanthin (derived from _Amphilonche
belonoides_ by central concrescence; often the sutures of the concreted
bases are visible, fig. 4_a_).

_Dimensions._--Length of the two major spines 0.1 to 0.4, breadth 0.005 to
0.015; length of the eighteen minor spines 0.005 to 0.15.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

{788}17. _Amphilonche acufera_, n. sp.

Two principal spines thick, four-sided prismatic in the basal half,
cylindrical or spindle-shaped in the distal half, with simple conical apex.
Eighteen smaller spines shorter, thin, bristle-shaped or conical on the
base. All twenty spines in the centre perfectly grown together, forming a
single piece of acanthin. (Derived from _Amphilonche elongata_ by central
concrescence.)

_Dimensions._--Length of the two major spines 0.3 to 0.5, breadth 0.01 to
0.03; length of the eighteen minor spines 0.08 to 0.2.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

Genus 342. _Amphibelone_,[382] Haeckel, 1862, Monogr. d. Radiol., p. 392.

_Definition._--#Amphilonchida# with two unequal principal spines (the
frontal spine very different from the caudal spine); the eighteen smaller
spines nearly equal.

The genus _Amphibelone_ exhibits among the Amphilonchida the same
remarkable differentiation of the two principal or longitudinal spines, as
_Zygostaurus_ among the Quadrilonchida; the frontal spine differs commonly
from the caudal spine not only in its size, but also in its peculiar form;
commonly one pole of the longitudinal axis is much more strongly developed
than the other. The eighteen smaller spines are nearly equal.

Subgenus 1. _Amphibelonium_, Haeckel.

_Definition._--All twenty spines separate, but in contact in the centre and
resting one against another by the triangular sides of their pyramidal
bases, without a prominent basal leaf-cross.

1. _Amphibelone aciculata_, n. sp.

Two principal spines thick, without edges and wings, the frontal spine
short, spindle-shaped, the caudal three to six times as long, cylindrical,
both of equal breadth, with simple conical apex and simple pyramidal base,
without leaf-cross. Eighteen smaller spines very thin, bristle-shaped, of
the same length as the frontal spine. Central capsule cylindrical or
spindle-shaped, enveloping the two principal spines nearly throughout their
whole length.

_Dimensions._--Length of the frontal spine 0.2 to 0.4, of the caudal spine
1.0 to 2.0; breadth of both 0.01; length of the eighteen smaller spines 0.1
to 0.3.

_Habitat._--South Atlantic, Station 325, surface.

{789}2. _Amphibelone cultellata_, n. sp. (Pl. 132, fig. 10).

Two principal spines broad, two-edged, knife-shaped; two opposite in the
equatorial plane, thin and broad wings or lamellæ are developed, and these
enclose the proximal part of both spines, whilst their distal part is free,
cylindrical, with conical apex: the two wings of the longer caudal spine
are lanceolate, twice to four times as broad as the two wings of the
shorter frontal spine; their base is a simple pyramid without leaf-cross.
Eighteen smaller spines conical, with bristle-shaped prolongation,
one-fourth to one-half as long as the frontal spine. Central capsule green,
semitransparent.

_Dimensions._--Length of the frontal spine 0.1 to 0.2, breadth 0.02 to
0.03; length of the caudal spine 0.2 to 0.4, breadth 0.04 to 0.08; length
of the eighteen smaller spines 0.05 to 0.15.

_Habitat._--Indian Ocean, Maldive Islands (Haeckel), surface.

3. _Amphibelone pyramidata_, n. sp. (Pl. 132, fig. 9).

Two principal spines four-winged, with four crossed, very thin and broad
wings, which are somewhat broader in the convex middle part than at either
end; both ends of each spine four-sided pyramidal, with four concave edges
and a very small terminal pyramid; base without leaf-cross. The longer
caudal spine is twice as broad at the distal end as at its proximal end,
and three times as broad as the distal end of the shorter frontal spine.
Eighteen smaller spines bristle-shaped, with four-sided pyramidal bases.
Central capsule pyramidal, enveloping the two principal spines throughout
their whole length; the base of the slender quadrangular pyramid is on the
caudal, the apex on the frontal pole of the longitudinal axis.

_Dimensions._--Length of the frontal spine 0.2 to 0.25, distal breadth
0.01; length of the caudal spine 0.3 to 0.4, distal breadth 0.03; length of
the eighteen smaller spines 0.04 to 0.08.

_Habitat._--Cape of Good Hope, Station 143, depth 1900 fathoms.

4. _Amphibelone anomala_, Haeckel.

_Amphilonche anomala_, Haeckel, 1862, Monogr. d. Radiol., p. 394, Taf.
xvi. fig. 8, Taf. xviii. figs. 23_a_, 23_b_.

_Acanthometra anomala_, Haeckel, 1860, Monatsber. d. k. Akad. d. Wiss.
Berlin, p. 808.

Two principal spines four-winged, nearly prismatic, with four crossed, very
broad and thick wings; both ends of each spine four-sided pyramidal, with
four concave edges and a very small terminal pyramid; base without
leaf-cross. The edges of the four wings are concave on the shorter frontal,
convex on the longer caudal spine. Eighteen smaller spines only one-fourth
to one-half as long, linear, two-edged, with two parallel teeth on the
distal end. Central capsule four-sided prismatic, olive-green, enveloping
almost completely the two principal spines.

_Dimensions._--Length of the frontal spine 0.12, middle breadth 0.016;
length of the caudal spine 0.14, middle breadth 0.024; length of the
eighteen smaller spines 0.04 to 0.06.

_Habitat._--Mediterranean (Messina), surface.

{790}Subgenus 2. _Amphibelithium_, Haeckel.

_Definition._--All twenty spines grown together in the centre, forming a
single star-shaped piece of acanthin.

5. _Amphibelone clavaria_, n. sp.

Two principal spines roundish, without edges and wings; the smaller frontal
spine elongate conical, the larger caudal spine two to four times as long,
cylindrical, in the distal part spindle-shaped or club-shaped, with conical
apex. Central base simple pyramidal, without leaf-cross. Eighteen smaller
spines much shorter, bristle-shaped. Central capsule very long,
club-shaped. All twenty spines perfectly grown together in the centre.

_Dimensions._--Length of the frontal spine 0.12, of the caudal spine 0.4 to
0.8; frontal breadth 0.01, caudal breadth 0.03; length of the eighteen
smaller spines 0.05.

_Habitat._--Central Pacific, Station 272, surface.

Genus 343. _Acantholonche_,[383] Haeckel, 1881, Prodromus, p. 466.

_Definition._--#Amphilonchida# with two equal principal spines (frontal and
caudal spines not different). The eighteen smaller spines are very unequal,
ten of them (eight tropical and two transverse equatorial spines) much
larger than the rudimentary eight polar spines.

The genus _Acantholonche_ differs from its ancestral genus _Amphilonche_ in
the different shape of the eight tropical and the eight polar spines; these
latter are much smaller than the former, which are almost equal to the two
transverse equatorial spines. The two principal spines are equal, but in
size and shape very different from the others.

1. _Acantholonche amphipolaris_, n. sp. (Pl. 132, fig. 7).

Two principal spines stout, quadrangular prismatic in the basal half,
cylindrical or spindle-shaped in the distal half, with simple conical apex;
base a small pyramid without leaf-cross. Two transverse and eight tropical
spines, about half as long as the former, very thin, bristle-shaped,
conical at the basal part. Eight polar spines very small, scarcely
one-eighth or one-fourth as long as the latter, short pyramidal or conical,
often quite rudimentary. Central capsule cylindrical, enveloping the basal
half of the two principal spines.

_Dimensions._--Length of the two principal spines 0.4 to 0.5, of the ten
smaller spines 0.2 to 0.3, of the eight rudimentary polar spines 0.01 to
0.06; breadth of the two large spines 0.02 to 0.03.

_Habitat._--Central Pacific, Stations 266 to 274, surface.

{791}2. _Acantholonche peripolaris_, n. sp. (Pl. 132, fig. 8).

Two principal spines quadrangular prismatic, with four broad prominent
lamellar wings, of increasing breadth towards the pyramidal distal apex.
Both ends of each spine four-sided pyramidal, base without leaf-cross. Two
transverse and eight tropical spines about two-thirds as long as the
former, four-sided pyramidal in the basal half, conical in the distal half,
often curved. Eight polar spines very small, about one-fourth as long as
the latter, short conical or pyramidal. Central capsule four-sided
prismatic, enveloping both principal spines.

_Dimensions._--Length of the two principal spines 0.2, of the ten smaller
spines 0.12, of the eight rudimentary polar spines 0.04.

_Habitat._--Central Pacific, Station 274, surface.

----

Order IV. ACANTHOPHRACTA, Richard Hertwig, 1879.

_Acanthometræ cataphractæ_, Johannes Müller, 1858, Abhandl. d. k.
Akad. d. Wiss. Berlin, pp. 12, 22, 49.
_Dorataspida et Diploconida_, Haeckel, 1862, Monogr. d. Radiol.,
pp. 404, 412.
_Acanthophractida_, Richard Hertwig, 1879, Organismus d. Radiol.,
pp. 25, 137.
_Dorataspida, Diploconida, et Sphærocapsida_, Haeckel, 1881,
Prodromus, p. 467.

_Definition._--ACANTHARIA with complete latticed shell.

The order #Acanthophracta#, the fourth order of Radiolaria, comprises all
those ACANTHARIA in which the acanthinic skeleton is a complete latticed or
fenestrated shell, supported by radial spines arising from one common
central point. By the possession of such a complete shell the
#Acanthophracta# differ from their ancestral group, the nearly allied
#Acanthometra#, which represent the older and simpler, first order of
ACANTHARIA. All #Acanthophracta# are Icosacantha (like the #Acanthonida#,
their ancestral group), and possess twenty radial spines disposed according
to the Müllerian law (compare above, p. 717).

Johannes Müller, who first observed five representatives of this order,
called a part of them "_Acanthometræ cataphractæ_," and united these with
the true #Acanthometra# (_Acanthometra costata_ and _Acanthometra
cataphracta_; Abhandl. d. k. Akad. d. Wiss. Berlin, 1858, pp. 12, 49).
Another part was united by him with the true _Haliomma_ (_Haliomma
echinoides_, _Haliomma hystrix_, _Haliomma tabulatum_; Abhandl. d. k. Akad.
d. Wiss. Berlin, 1858, pp. 36, 37). He supposed that these latter formed
the immediate transition from the true #Acanthometra# to the true
_Haliomma_, and that their skeleton was siliceous.

{792}In my Monograph (1862, p. 412) I founded a separate subfamily,
Dorataspida, for the "_Acanthometræ cataphractæ,_" which I considered as
the first subfamily of the "Ommatida." That subfamily contained at that
time only two genera, _Dorataspis_ (with seven species) and
_Haliommatidium_ (with five species). A third genus, _Aspidomma_ (with two
species), was united by me with the Haliommatida (because of its double
shell). For a fourth genus (_Diploconus_) with a single species I founded
the peculiar family of Diploconida. Therefore the whole number of
#Acanthophracta# described in my Monograph amounted only to four genera and
fifteen species. Now the rich collections of the Challenger have added such
a great number of new forms, that we may distinguish here thirty-eight
genera and two hundred and twelve species.

Richard Hertwig in his excellent work (Der Organismus der Radiolarien,
1879, p. 25) separated his "Acanthophractida" perfectly from the "Ommatida"
(or the siliceous #Sphæroidea#), and united them with the "Acanthometrida"
in his order "Acanthometrea." But he separated them also from the nearly
allied Diploconida, following my former arrangement. He distinctly noted
that the skeleton in all these Acanthophractida (as well as in the
Acanthometrida) consists not of silex but of the organic substance
"acanthin."

The astonishing number of new and interesting forms of #Acanthophracta#
which I have found in the rich collection of the Challenger enables me to
distinguish now in this suborder six different families, two of which are
perfectly new (the Sphærocapsida and the Hexalaspida). But the four other
families also are so much enlarged that their interesting morphology
appears in quite a new and clear light. Far the largest and most important
of these six families is that of the true Dorataspida, which embraces
seventeen genera and one hundred and eight species (more than the other
five families together). From this largest and oldest ancestral family four
other families have afterwards arisen, whilst a single family, the
Sphærocapsida, seems to possess no direct phylogenetic connection with the
five other families.

The peculiar and quite new family of Sphærocapsida (Pl. 133, figs. 7-11;
Pl. 135, figs. 6-10) differs from all other #Acanthophracta# in the
singular structure of the spherical acanthinic shell, composed of
innumerable small plates or aglets, each of which is pierced by a very
small porule. This peculiar pavemented shell (enclosing the central capsule
and separated from it by the jelly-like calymma) seems to be produced on
the surface of the spherical calymma, immediately by secretion of the
pseudopodia, and independently from the twenty radial spines, united in the
centre of the sphere. On the twenty points, where the spines perforate the
shell, there are originally eighty larger pores (four around each piercing
spine); but there is no certain indication that the shell is produced by
the meeting apophyses of the twenty spines, as is the case in the five
other families of #Acanthophracta#. Therefore perhaps it is {793}more
natural to unite these latter into another suborder as #Cladophracta#, and
to separate them from the Sphærocapsida, which may be called Capsophractæ.

The Dorataspida (Pls. 134-138), the common ancestral stock of the
#Cladophracta#, in the definition here restricted embraces all those
#Acanthophracta# in which the spherical lattice-shell is simple and
composed of the meeting branches of twenty radial spines united in its
centre. As already pointed out above, this family is probably diphyletic,
and embraces two subfamilies which have been derived originally from two
different forms of #Acanthonida#--the Diporaspida (with two opposite
apophyses on each spine) derived from the Phractacanthida, and the
Tessaraspida (with four crossed apophyses on each spine) derived from the
Stauracanthida; in the former we find originally forty apophyses, in the
latter eighty apophyses, by the meeting branches of which the spherical
lattice-shell originates. The four following families of #Acanthophracta#
have probably been derived from the Diporaspida.

The Phractopeltida (Pl. 133, figs. 1-6) differ from all other
#Acanthophracta# in the possession of a double lattice-shell, composed of
two concentric spheres which are united by the twenty radial spines meeting
in the centre. As all Phractopeltida possess originally only two apophyses
on each radial spine, they must be derived from the Diporaspida
(_Orophaspis_), and bear to them the same relation as the Dyosphærida do to
the Monosphærida. As the spherical central capsule of the Phractopeltida is
enclosed between both shells, smaller than the outer, larger than the inner
shell, the latter may be called "medullary shell," the former "cortical
shell." This family represents among the #Acanthophracta# only the
"Diplophracta," whilst all others are "Haplophracta."

The three families here characterised may be called together
"#Sphærophracta#," as their central capsule and the enveloping shell are
constantly spherical (or the shell sometimes an "endospherical
polyhedron"). On the contrary the following three families of
#Acanthophracta# may be united as "#Prunophracta#," as their central
capsule and shell are never spherical, but either ellipsoidal or lenticular
or of another form. The common ancestral stock of this suborder are the
Belonaspida, in which the form of the central capsule and the enclosing
lattice-shell is ellipsoidal; they are derived from the Dorataspida (and
probably all from the subfamily Diporaspida) by the prolongation of two
opposite radial spines which are larger than the eighteen others; they are
the two equatorial spines of the "hydrotomical axis" (compare above, p.
719, and Pl. 136, figs. 6-9).

The Hexalaspida (Pl. 139) represent a new and very remarkable family,
distinguished from all other #Acanthophracta# by the preponderating
development of six stout radial spines, which are much larger than the
fourteen others. These six principal spines lie in one meridian plane of
the shell (in the "hydrotomical plane," p. 720), and are the two opposite
equatorial spines and the four appertaining polar spines of the same plane.
{794}As the fourteen smaller spines develop their apophyses at smaller
distances from the centre, the shell assumes a peculiar lenticular or
discoidal form, and the margin of this disk bears the six larger spines.
Moreover the enclosed small central capsule is lenticular. The Hexalaspida
may be derived immediately from the Belonaspida.

The Diploconida (Pl. 140) form the last and the most modified family of all
#Acanthophracta#. The remarkable shell exhibits the strange form of a
double cone, bearing in its axis two very large opposite spines; these are
the two equatorial spines of the "hydrotomical axis" (p. 719). The
double-conical or nearly cylindrical shell is composed of three different
parts or segments; the small middle part is the true lattice-shell of the
Hexalaspida and Belonaspida, and bears the eighteen smaller (often quite
rudimentary) radial spines. The two other parts (opposite on both poles of
its hydrotomical axis) are the conical or cylindrical, solid, basal sheaths
of the two large equatorial spines, enveloping their major part. In
consequence of this peculiar metamorphosis of the shell the Diploconida
represent the last and the most aberrant group of all ACANTHARIA.

_Synopsis of the Suborders and Families of_ #Acanthophracta#.

-------------------------------------------------------------------------
Suborder I. SPHÆROPHRACTA.
Twenty radial spines of equal size. Shell spherical (or an
endospherical polyhedron).
-------------------------------------------------------------------------
Shell spherical, simple, pierced by twenty or
eighty aspinal pores and composed of a
pavement of innumerable very small plates
or aglets, each pierced by one porule, 1. SPHÆROCAPSIDA.

Shell spherical, simple, composed of the
meeting branches of two or four apophyses
of the twenty radial spines, 2. DORATASPIDA.

Shell spherical, double, composed of two
concentric lattice-spheres, which are connected
by the twenty radial spines and composed
of the meeting branches of their apophyses, 3. PHRACTOPELTIDA.
-------------------------------------------------------------------------
Suborder II. PRUNOPHRACTA.
Twenty radial spines of unequal size; two or six hydrotomical spines
much larger than the eighteen or fourteen others. Shell not spherical.
-------------------------------------------------------------------------
Shell ellipsoidal, with prolonged hydrotomical
axis, the two spines of which are larger
than the eighteen others, 4. BELONASPIDA.

Shell lenticular or discoidal, with six larger
spines placed in the hydrotomical plane
(fourteen other spines much smaller), 5. HEXALASPIDA.

Shell diploconical or nearly cylindrical, with
two opposite large funnels, the sheaths of
the enlarged two spines of the hydrotomical
axis (eighteen other spines much
smaller or rudimentary), 6. DIPLOCONIDA.

{795}----

Suborder I. SPHÆROPHRACTA, Haeckel.

_Definition._--Shell spherical, with twenty radial beams of equal size.

Family XXXIX. #SPHÆROCAPSIDA#, Haeckel (Pl. 133, figs. 7-11; Pl. 135, figs.
6-10).

_Sphærocapsida_, Haeckel, 1881, Prodromus, p. 469.

_Definition._--ACANTHARIA with simple spherical porous shell, composed of
innumerable very small plates, each of which is pierced by one radial
porule. Twenty radial spines of equal size meeting in the centre of the
shell and disposed according to the Müllerian law of the Icosacantha,
sometimes short and enclosed in the shell, at other times long and piercing
it (rarely rudimentary or quite absent). Shell pierced therefore either by
twenty larger perspinal pores or by eighty smaller aspinal pores. Central
capsule spherical, enclosed in the porous shell.

The family #Sphærocapsida#, founded by me in 1881 for the single genus
_Sphærocapsa_, represents a very peculiar and remarkable group of the
#Acanthophracta#, very different from the five other families of this
suborder, and probably derived, independently of them, directly from the
#Acanthonida#. Whilst the lattice-shell of the five other families is
composed of the meeting branches of lateral apophyses of the twenty spines,
and its meshes are all or partly the intervals between these apophyses, in
the Sphærocapsida the spherical shell has quite another structure, and is
composed of innumerable small plates (each with one pore) which are
secreted on the surface of the spherical calymma, independently of the
twenty radial spines, which do not possess true apophyses.

In all Sphærocapsida the structure of the spherical shell is quite peculiar
and different from that of all other Radiolaria. It is composed everywhere
of innumerable very small plates or aglets, which are connected irregularly
like paving-stones, and form a single continuous layer or pavement on the
surface of the spherical calymma (Pl. 133, fig. 11, _a_; Pl. 135, figs. 8,
10). The small plates or paving-stones, which we will call "aglets," are
connected at their meeting edges by a kind of cement, and form together
with it a continuous thick capsule of acanthin. The form of the aglets is
commonly more or less irregular, roundish or polygonal, sometimes longish
(Pl. 133, fig. 11, _a_), more rarely it becomes rather regular, hexagonal,
square, or roundish (Pl. 135, fig. 8). Usually all aglets of one and the
same individual are of nearly equal size, between 0.01 and 0.02 in
diameter, rarely less or more. The outer face of the aglets is more or less
concave, so that the elevated meeting edges of the neighbouring aglets
commonly form together a prominent network of crests (Pl. 135, figs. 8,
10); rarely the meeting edges partly cover one another like squamules (Pl.
133, fig. 11, _a_). {796}Each small plate or aglet is pierced in its centre
by a single radial canalicule or porule. The dimpled surface, so produced,
resembles somewhat the dimpled plates of _Ceriaspis_, &c. Different from
these innumerable very small dimples of the surface are the twenty larger
"spinal dimples," or the concave larger plates, which are originally
pierced by the twenty radial spines. Before we describe these, we must
examine the spines themselves.

The twenty radial spines of all observed Sphærocapsida (sixteen species)
agree perfectly with those of the genus _Acanthonia_ (p. 749), and
especially with _Acanthonia tetracopa_, _Acanthonia denticulata_, &c. All
twenty spines, regularly disposed according to the Müllerian law of the
Icosacantha, are of equal size, constantly four-edged prismatic, of equal
breadth throughout their whole length. The prominent four edges are
parallel, sometimes smooth, at other times elegantly denticulated. The
central bases of the twenty spines are pyramidal, without leaf-cross, and
propped one upon another with their triangular faces, as in the majority of
the #Acanthonida#.

The relation of the twenty radial spines to the spherical shell exhibits in
the five genera described very peculiar and important differences. In the
first described genus, in _Sphærocapsa_, the spines are exactly as long as
the shell-radius, and therefore are not prominent over the surface of the
shell, with which they are firmly connected; the truncated distal end of
the spine lies therefore here in the surface of the shell itself, and is
connected with it by its four edges, between which four open aspinal pores
remain, as in _Tessaraspis_, &c. (Pl. 135, figs. 6-10). In the next allied
genus, _Astrocapsa_ (Pl. 133, figs. 9, 10), the spines are longer than the
shell-radius, and therefore more or less prominent over its surface; the
piercing part of each spine is also surrounded by four aspinal pores. In
the two following genera, _Porocapsa_ and _Cannocapsa_ (Pl. 133, figs. 7,
8), the radial spines are shorter than the shell-radius and therefore quite
hidden and withdrawn inside the shell, which they do not reach. But in the
ideal prolongation of each spine the shell is pierced by a single large
opening, the "perspinal pore" or "perspinal hole," composed of the four
united aspinal pores. Whilst in _Porocapsa_ the perspinal pores are simple,
they are prolonged in _Cannocapsa_ into cylindrical tubes, open at both
ends. The twenty perspinal holes of these Porocapsida are therefore derived
by confluence of the eighty original aspinal pores of the Astrocapsida and
preserve the same regular disposition, according to the Müllerian law of
the Icosacantha. Finally, the same law as is valid also in the last genus
is found in _Cenocapsa_; here the radial spines have completely
disappeared, and the whole skeleton is a simple sphere, but of the same
structure, and with the same twenty perspinal pores as in _Porocapsa_. It
is very interesting that this spineless _Cenocapsa_ among the ACANTHARIA
exhibits the same shell (a simple hollow sphere) as a last reduced form,
which _Cenosphæra_ among the #Sphærellaria# produces as a primitive
ancestral form of numerous genera.

{797}_The Central Capsule_ of the Sphærocapsida is spherical, constantly
smaller than the enclosing concentric shell, and separated from it by the
calymma. Its structure seems to be the same as in the #Acanthonida#, and
specially in the Astrolonchida. The pseudopodia (not yet observed) are
probably protruded only through the twenty perspinal holes or the eighty
aspinal pores.

_Synopsis of the Genera of Sphærocapsida._

-------------------------------------------------------------------------
I. Subfamily Astrocapsida.
Radial spines connected with the porous shell, as long or longer than
its radius. Eighty aspinal pores.
-------------------------------------------------------------------------
Spines as long as the radius,
without external prolongation, 344. _Sphærocapsa_.

Spines longer than the radius,
with external prolongation, 345. _Astrocapsa_.
-------------------------------------------------------------------------
II. Subfamily Porocapsida.
Radial spines not connected with the porous shell, shorter than its
radius. Twenty perspinal pores.
-------------------------------------------------------------------------
Perspinal holes of the shell
simple, without external
prolongation, 346. _Porocapsa_.

Perspinal holes of the shell
prolonged into radial
centrifugal tubuli, 347. _Cannocapsa_.
-------------------------------------------------------------------------
III. Subfamily Cenocapsida.
Radial spines disappeared.
-------------------------------------------------------------------------
Twenty perspinal holes of the
shell simple, without tubular
prolongation, 348. _Cenocapsa_.

Subfamily 1. ASTROCAPSIDA, Haeckel.

_Definition._--Radial spines connected with the porous shell, as long as or
longer than its radius. Therefore the shell pierced by eighty aspinal pores
(four around each spine).

Genus 344. _Sphærocapsa_,[384] Haeckel, 1881, Prodromus, p. 469.

_Definition._--#Sphærocapsida# with twenty radial spines as long as the
radius of the shell, without external prolongation; therefore their distal
ends inserted in the perspinal holes, each of which is composed of four
aspinal pores.

The genus _Sphærocapsa_ is the most common form of the Sphærocapsida, and
comprises those species in which the radial spines are as long as the
radius of the shell, and therefore are connected with the margin of its
aspinal holes, but not prolonged beyond its surface.

{798}1. _Sphærocapsa cruciata_, n. sp. (Pl. 135, figs. 6, 7).

Aspinal holes nearly circular, with flat radially striated margin. Four
aspinal pores of each hole rounded equilateral triangular. Porules of the
shell simple, without ring and dimple. Four edges of the spines smooth.

_Dimensions._--Diameter of the shell 0.4 to 0.5, of the central capsule 0.3
to 0.4.

_Habitat._--North Atlantic, Færöe Channel (Gulf Stream), 1880, John Murray,
surface.

2. _Sphærocapsa dentata_, n. sp. (Pl. 135, fig. 9).

Aspinal holes four-lobed, with broad concave margin denticulated on the
periphery. Four aspinal pores of each hole pear-shaped, oblong, elevated in
the centre. Porules of the shell simple, without ring and dimple. Four
edges of the spines denticulate.

_Dimensions._--Diameter of the shell 0.3 to 0.4, of the central capsule 0.2
to 0.3.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

3. _Sphærocapsa quadrata_, n. sp. (Pl. 135, fig. 8).

Aspinal holes square, with concave umbilicus in the centre, surrounded by a
convex denticulated margin. Four aspinal pores of each hole nearly square.
Shell-porules with a polygonal elevated smooth ring, in the bottom of a
shallow dimple. Four edges of the spines smooth.

_Dimensions._--Diameter of the shell 0.22, of the central capsule 0.17.

_Habitat._--Indian Ocean, south of Australia, Station 159, surface.

4. _Sphærocapsa pavimentata_, n. sp. (Pl. 135, fig. 10).

Aspinal holes four-lobed, with broad concave, irregularly crenated and
figured margin. Four aspinal pores of each hole violin-shaped. Porules of
the shell surrounded by an irregularly oblong ring with thick elevated,
elegantly crenated margin. Four edges of the spines smooth.

_Dimensions._--Diameter of the shell 0.36, of the central capsule 0.3.

_Habitat._--South-east Pacific (off Valparaiso), Station 298, surface.

Genus 345. _Astrocapsa_,[385] n. gen.

_Definition._--#Sphærocapsida# with twenty radial spines longer than the
radius of the shell, piercing its perspinal holes, with free external
prolongation; therefore with four aspinal pores around each spine.

The genus _Astrocapsa_ differs from the preceding _Sphærocapsa_ in the
external prolongation of the radial spines piercing the perspinal holes; it
assumes therefore the common shape of the Dorataspida more than the other
Sphærocapsida do.

{799}1. _Astrocapsa tritonis_, n. sp.

Aspinal holes circular, with smooth convex margin and four circular aspinal
pores. Porules of the shell simple, without ring and dimple. Four edges of
the spines smooth, their outer free part about as long as the diameter of
the shell.

_Dimensions._--Diameter of the shell 0.25; outer length of the spines 0.3.

_Habitat._--North Atlantic, Færöe Channel, Gulf Stream (expedition of H.M.S
"Triton," August, 1882), John Murray, surface.

2. _Astrocapsa stellata_, n. sp. (Pl. 133, fig. 10).

Aspinal holes cruciform, with high crenated margin. Four aspinal pores of
each hole egg-shaped. Porules of the shell simple, without ring and dimple.
Four edges of the spines denticulate; their outer free part twice to three
times as long as the diameter of the shell.

_Dimensions._--Diameter of the shell 0.36; outer length of the spines 0.5
to 0.8.

_Habitat._--Antarctic Ocean (near Kerguelen), Station 152, surface.

3. _Astrocapsa quadrifida_, n. sp.

Aspinal holes four-lobed, with four prominent teeth between the four
roundish aspinal pores. Porules of the shell surrounded by a high polygonal
smooth ring. Four edges of the spines denticulate; their outer free part
about as long as the radius of the shell.

_Dimensions._--Diameter of the shell 0.28; outer length of the spines 0.15.

_Habitat._--North Pacific, Station 241, surface.

4. _Astrocapsa coronata_, n. sp. (Pl. 133, fig. 9).

Aspinal holes circular, with a coronet of numerous thin parallel teeth.
Porules of the shell in dimples on irregular polygonal small plates, with
coronated ring. Four edges of the spines denticulate; their outer free part
about twice as long as the diameter of the shell.

_Dimensions._--Diameter of the shell 0.4 to 0.5 ; outer length of the
spines 1.0 to 1.2.

_Habitat._--North Atlantic, Færöe Channel, Gulf Stream (expedition of the
"Knight Errant," 1880), John Murray, surface and at depths varying from 10
to 200 fathoms.

Subfamily 2. POROCAPSIDA, Haeckel.

_Definition._--Radial spines not connected with the porous shell, shorter
than its radius; therefore the shell pierced by twenty perspinal pores
(each one in the ideal radial prolongation of one spine).

{800}Genus 346. _Porocapsa_,[386] n. gen.

_Definition._--#Sphærocapsida# with twenty radial spines shorter than the
radius of the shell; therefore their distal ends not connected with the
twenty perspinal holes, which are simple, not prolonged into radial tubes.

The genus _Porocapsa_ and the following _Cannocapsa_ form together the
small sub-family of Porocapsida, distinguished by the peculiar reduction or
retrograde development of the twenty radial spines; all these twenty are
present and disposed according to the Müllerian law of the Icosacantha, but
they are shorter than the radius of the shell and therefore do not reach
it. In the ideal prolongation of the spines the shell is pierced by twenty
simple quadrangular or circular perspinal holes.

1. _Porocapsa murrayana_, n. sp. (Pl. 133, fig. 7).

Perspinal holes cruciform, with smooth thickened margin or with four short
teeth between the four lobes. Porules of the shell simple, without ring and
dimple. Four edges of the spines smooth. (Differs from _Sphærocapsa
cruciata_, Pl. 135, figs. 6, 7, living in the same locality, mainly in the
reduction of the radial spines, which do not reach the shell.)

_Dimensions._--Diameter of the shell 0.28; length of the spines 0.2.

_Habitat._--North Atlantic, Færöe Channel, Gulf Stream (expedition of
H.M.S. "Triton," August 1882), John Murray, surface and in depths from 40
to 640 fathoms.

2. _Porocapsa tetrodon_, n. sp.

Perspinal holes cruciform, with four triangular prominent teeth between the
four lobes of the cross. Porules of the shell with an elevated polygonal
ring, in the bottom of a dimple. Four edges of the spines elegantly
denticulate.

_Dimensions._--Diameter of the shell 0.36; length of the spines 0.12.

_Habitat._--North Pacific, Station 253, surface.

3. _Porocapsa octodon_, n. sp.

Perspinal holes square, with four larger prominent teeth on the sides of
the square, and four smaller teeth on its corners. Porules of the shell
surrounded by an elevated ring with elegantly crenated irregular margin.
Four edges of the spines smooth.

_Dimensions._--Diameter of the shell 0.48; length of the spines 0.18.

_Habitat._--Arctic Ocean (Greenland), in the contents of the stomach of the
Peromedusa, _Periphylla hyacinthina_.

{801}4. _Porocapsa coronodon_, n. sp.

Perspinal holes circular, with ciliated margin, which forms a crown of
sixteen to twenty-four thin parallel teeth. Poruli of the shell surrounded
by an elevated ring with high crenated margin. Four edges of the spines
denticulate.

_Dimensions._--Diameter of the shell 0.55; length of the spines 0.15.

_Habitat._--Antarctic Ocean, Station 154, surface.

Genus 347. _Cannocapsa_,[387] n. gen.

The genus _Cannocapsa_, exhibits the same peculiar reduction of the radial
spines as the foregoing _Porocapsa_; the spines are also here shorter than
the shell-radius and do not therefore reach the perspinal holes of the
shell. But whilst these latter are simple in _Porocapsa_, they are
prolonged into radial tubules in _Cannocapsa_; the outer surface bears
therefore twenty such cylindrical tubules, separated by a short distance
from the inner enclosed spines, but disposed quite regularly according to
the law of Icosacantha.

1. _Cannocapsa osculata_, n. sp.

Perspinal holes prolonged into short cylindrical tubuli, the length of
which about equals their diameter. Both ends of the tubuli with smooth
thickened margin. Poruli of the shell simple, without ring and dimple. Four
edges of the spines smooth; their length equals about four-fifths of the
shell-radius.

_Dimensions._--Diameter of the shell 0.24; length of the spines 0.09, of
the tubuli 0.01.

_Habitat._--North Atlantic, Færöe Channel (Gulf Stream), surface, John
Murray.

2. _Cannocapsa stethoscopium_, n. sp. (Pl. 133, fig. 8).

Perspinal holes prolonged into cylindrical tubuli, half as long as the
shell radius. Both ends of the tubuli with smooth thin trumpet-shaped
margins. Poruli of the shell simple, without ring and dimple. Four edges of
the spines smooth; their length about equals three-fourths of the
shell-radius.

_Dimensions._--Diameter of the shell 0.2; length of the spines 0.08, of the
tubuli 0.05.

_Habitat._--South Atlantic (west of Tristan da Cunha), Station 333,
surface.

3. _Cannocapsa tubulosa_, n. sp.

Perspinal holes prolonged into cylindrical tubuli, which are as long as or
longer then the shell-radius. Both ends of the tubuli with smooth thickened
margin. Poruli of the shell simple, {802}without ring and dimple. Four
edges of the spines smooth; their length scarcely equals half the
shell-radius.

_Dimensions._--Diameter of the shell 0.15; length of the spines 0.03, of
the tubuli 0.08.

_Habitat._--South Atlantic (east coast of Patagonia), Station 318, surface.

Subfamily 3. CENOCAPSIDA, Haeckel.

_Definition._--Radial spines completely reduced and absent; shell cavity
therefore simple; shell pierced by twenty perspinal pores (each placed in
the direction of one radial spine which has disappeared).

Genus 348. _Cenocapsa_,[388] n. gen.

_Definition._--#Sphærocapsida# without radial spines, with simple cavity of
the spherical shell, which is pierced by twenty perspinal holes (each one
placed in the radial direction of one spine which has disappeared).

The genus _Cenocapsa_ comprises only a single species, but is very
remarkable in that it is the most reduced form among all Sphærocapsida. The
twenty radial spines of the Icosacantha have perfectly disappeared by
complete retrograde metamorphosis, and the only evidence of their former
existence (in the ancestral genus _Porocapsa_) are the twenty perspinal
holes remaining in the shell. _Cenocapsa_ is the only form of ACANTHARIA
which possesses no radial spines.

1. _Cenocapsa nirvana_, n. sp. (133, fig. 11, 11_a_, 11_c_).

Perspinal holes four-lobed, cruciform, with four short triangular teeth
between the four lobes of the cross. Poruli of the shell in the bottom of
an elliptical dimple surrounded by an elevated ring.

_Dimensions._--Diameter of the shell 0.2 to 0.3, of the aspinal holes 0.02.

_Habitat._--North Pacific, Station 248, surface.

Family XL. #DORATASPIDA#, Haeckel (Pls. 134-138).

_Dorataspida_, Haeckel, 1862, Monogr. d. Radiol., p. 412.

_Definition._--ACANTHARIA with simple spherical lattice-shell, composed of
the branched apophyses of twenty equal radial spines meeting in its centre
and disposed according to the Müllerian law of Icosacantha. Central capsule
spherical, enclosed in the fenestrated shell.

The family #Dorataspida# is the most important family of the
#Acanthophracta#, or of those ACANTHARIA in which the radial spines are
connected by a complete extracapsular lattice-shell. The Dorataspida
represent probably the ancestral {803}stock of this whole order, with the
exception of the Sphærocapsida. The four following families of the order
may be easily derived from the Dorataspida. The number of genera
(seventeen) and of species (one hundred and eight) is in this family
greater than in the other five families together. When I constituted that
family in my Monograph 1862, it comprised only one genus, _Dorataspis_,
with seven species. The nearly allied genus _Haliommatidium_ (_Phatnaspis_)
belongs to the Belonaspida.

The Dorataspida differ from the other #Acanthophracta# in the simple
spherical lattice-shell, which is composed of the meeting apophyses of the
twenty radial spines. In three other families of the suborder the shell is
not spherical, but ellipsoidal (Belonaspida), discoidal (Hexalaspida), or
diploconical (Diploconida). In the Phractopeltida the spherical shell is
double, composed of two concentric lattice-spheres. In the Sphærocapsida
the simple spherical shell is not composed of the apophyses of the spines,
but of innumerable small plates.

The family Dorataspida may be divided into two very different subfamilies,
which are probably derived, independently of one another, from two
different subfamilies of the Astrolonchida. The first subfamily,
Diporaspida, exhibits on each radial spine two opposite apophyses, like its
ancestral group, the Phractacanthida (p. 753); whereas the second
subfamily, Tessaraspida, possesses on each radial spine four crossed
apophyses (opposite in pairs), like its ancestral group, the Stauracanthida
(p. 758). Therefore the composition of the spherical shell, produced by the
meeting branches of the tangential apophyses, is essentially different in
the two subfamilies: in the Diporaspida each radial spine is surrounded by
two opposite primary aspinal meshes, in the Tessaraspida by four crossed
primary aspinal meshes.

Another principle of division may be established for the whole family by
the different mode of composition of the shell, and regarding this
important difference we may distinguish also two different subfamilies as
#Cladophracta# and #Peltophracta#. In the first and simpler subfamily, the
#Cladophracta#, the shell is composed totally (or sometimes partially) of
the meeting branches of the apophyses of the neighbouring spines; but in
each single spine (or in the most part of them) the branches of the
apophyses are not united, and form no lattice-plate (Pl. 137, figs. 1 to
8). Whereas in the #Peltophracta# the shell is composed constantly of
twenty perforated plates, as in each single spine the branches of its
apophyses are united and form a fenestrated shield with two or four (and
sometimes numerous) pores (Pls. 135, 136, 138).

In the Diporaspida as well as in the Tessaraspida we find numerous
representatives of the two groups of the #Cladophracta# and of the
#Peltophracta#; therefore the whole family of Dorataspida may be divided
into four different tribes. The Diporaspida (with two opposite apophyses on
each spine) are partly #Cladophracta# (the Phractaspida, Pl. 137, figs.
1-4), partly #Peltophracta# (the Ceriaspida, Pl. 138). On the other hand
the Tessaraspida (with four crossed apophyses on each {804}spine) are also
partly #Cladophracta# (the Stauraspida, Pl. 137, figs. 5-8), partly
#Peltophracta# (the Lychnaspida, Pls. 135-136). The differences and
relations of these tribes are placed synoptically in the following table:--

All Dorataspida are true Icosacantha, and the twenty spines, composing the
spherical shell, are equally developed, regularly disposed according to the
Müllerian law, and of equal size; also the distance of their plates from
the common centre is equal. Nevertheless they are never of perfectly the
same form; in consequence of their peculiar disposition in five zones (each
with four spines) certain slight differences are effected, so that with
accurate knowledge of the peculiar shell-composition it is generally not
difficult to distinguish the spines of the equatorial, the two tropical,
and the two polar zones.

Already the central bases, by which the twenty spines are united in the
centre of the sphere, exhibit certain differences in the five zones.
Commonly these bases are small pyramids, all meeting with their apex in the
centre, and the triangular faces of the neighbouring pyramids are supported
one upon another. The four equatorial pyramids are commonly six-sided, the
other sixteen five-sided; but sometimes there are eight six-sided and
twelve five-sided basal pyramids; two opposite polar spines on each pole
having a six-sided base (like the four equatorial), the other two polar
spines on each pole having a five-sided base (like the eight tropical).
Rarely the central bases are perfectly grown together, forming a single
spherical central piece of acanthin.

The three different fundamental forms of radial spines, which are found in
all ACANTHARIA, the cylindrical, the two-edged, and the four-edged (spines
with circular, with elliptical, and with square transverse section
respectively) occur also in the different groups of Dorataspida; but
commonly the two-edged or compressed form is prevalent in the Diporaspida,
the four-edged or quadrangular form in the Tessaraspida. In the majority of
species the spines are thickened in the shell-face, where the apophyses
arise, and thinner towards the two ends. Usually the outer or distal part
of the spine (outside the shell) is longer than the inner or proximal part
(inside {805}the shell). The distal apex is commonly simple, conical or
pyramidal, rarely bifid or truncate. The edges of the spines are commonly
smooth, rarely denticulate or serrate.

The apophyses, or the lateral transverse processes of the radial spines, in
the Dorataspida assume the greatest variety and complexity in form, size,
mode of ramification, and in composition of the shell. An expert and
practised observer may determine easily the range of each spine, whether it
be an equatorial (_c_), or a tropical (_b_, _d_), or a polar spine (_a_,
_e_, Pls. 133-138). The two opposite apophyses of the Diporaspida, as well
as the four crossed apophyses of the Tessaraspida, lie constantly in
certain meridian planes of the spine, which have a legitimate signification
for each of the five zones. The comparative morphology of this regular
disposition of the apophyses and the regular meeting of their branches is
of the greatest interest, and necessary for the complete understanding of
the complicated structure of these wonderful shells.

The pores or meshes of the spherical shell, offering the most varied forms,
may generally be divided into two different groups, into sutural and parmal
meshes. The sutural pores are bordered by the meeting branches of the
apophyses of two, three, or four neighbouring spines, and therefore also by
the sutures in which they meet. The parmal pores on the other hand are
bordered only by the united branches of the apophyses of a single spine and
pierce the shield or lattice-plate formed by them. Therefore the
shell-meshes of the #Cladophracta# are all sutural pores (Pl. 137, figs.
1-8; rarely and only in a part of the spines parmal pores also: _Zonaspis_,
_Dodecaspis_); whereas the shell-meshes of the #Peltophracta#, piercing the
shields or lattice-plates of all twenty spines, are always partly sutural,
partly parmal pores (Pls. 135, 136, 138). The parmal pores again may be
divided into two different groups--aspinal and coronal pores. Aspinal pores
("ad spinam") are those which lie immediately on the sides of the radial
spine and are bordered by the primary branches of its apophyses; therefore
constantly only two in the Diporaspida, four in the Tessaraspida. Coronal
pores on the contrary are those which lie in the periphery of the
lattice-plates, surrounding in a circle or crown the aspinal pores and not
touching the spine itself. In _Dorataspis_, _Ceriaspis_, _Tessaraspis_,
_Lychnaspis_, &c., all parmal meshes are only aspinal pores (Pl. 135, figs.
2-5; Pl. 136); whilst in _Coscinaspis_, _Acontaspis_, _Icosaspis_,
_Hylaspis_, &c., one part of the parmal pores is aspinal, one part coronal
(Pl. 136). The number, form, and size of the coronal pores is very variable
and often very large (sometimes more than a hundred in one plate).

The #Cladophracta# exhibit a comparatively simple shell-formation; either
all twenty spines or at least a part of them not forming lattice-plates.
The most primitive form among these is _Phractaspis_ (Pl. 137, figs. 1, 2).
The forty apophyses of its twenty spines are simply forked, and their
eighty fork-branches united by forty sutures, enclosing twenty-two sutural
meshes: two square polar meshes (between the four polar spines on the poles
of the spineless axis, _a a a a_ and _e e e e_); eight triangular
{806}circumpolar meshes (each between two polar and one tropical spine, _a
b a_ and _e d e_); eight quadrangular tropical meshes (each between one
polar, one equatorial, and two tropical spines, _a b c b_ and _e d c d_);
and four rhomboidal equatorial meshes (between two tropical and two
equatorial spines, _c b c d_). If the fork-branches be again forked
(_Phractaspidium_, Pl. 137, fig. 3), the number of the sutures and sutural
meshes is doubled, and the same is the case in _Stauraspis_, the most
simple form of the Tessaraspida.

A peculiar small group, and an interesting transition for the
#Cladophracta# to the #Peltophracta#, is presented by the Zonaspida among
the Tessaraspida (_Zonaspis_ and _Dodecaspis_). Here only one part of the
radial spines bears lattice-plates, the other part not. In _Zonaspis_ the
four equatorial spines bear lattice-plates, the sixteen other only free
branches of the apophyses. In _Dodecaspis_ twelve spines are provided with
lattice-plates (four equatorial and eight polar spines), whilst the eight
other (tropical) spines are devoid of them.

The #Peltophracta# exhibit a great variety in the form and composition of
their twenty lattice-plates or fenestrated shields. In the most simple case
(a part of _Dorataspis_ and _Diporaspis_) the shell is composed of four
(equatorial) hexagonal plates, and sixteen pentagonal plates (four tropical
and four polar); in this case the four polar plates meet on each pole in
one common point. More commonly, however, the shell seems to be composed of
eight hexagonal plates (four equatorial and the four polar spines of the
hydrotomical plane) and twelve pentagonal plates (eight tropical and the
four polar spines of the geotomical plane); in this case only two
(hexagonal) polar plates meet on each pole in a suture which separates the
two other (pentagonal) polar plates (Pl. 138, fig. 4). In the majority of
the Dorataspida the composition of the shell is much more complicated and
often very difficult to understand. Often the surface of the plates is
covered with a network of elevated crests, by which concave blind dimples
are separated (_Ceriaspis_, _Hystrichaspis_, Pl. 138); and sometimes these
dimples become pierced by coronal pores (Pl. 138, fig. 11, &c.).

Peculiar by-spines or "accessory spines" cover the outer surface of the
shell in a great part of Dorataspida, and commonly these most
characteristic by-spines are not placed radially, but parallel to the
radial spine, from the lattice-plate of which they arise (Pl. 135, figs. 1,
5; Pl. 137, figs. 4-8; Monogr. d. Radiol., 1862, Taf. xxi. figs. 8, 9).
They are commonly placed perpendicular to the sutural condyles, or the
branch-ends of the apophyses; so that close to each suture arises a pair of
divergent by-spines, belonging to the meeting apophyses of the two
neighbouring spines, which meet in the suture (Pl. 137, fig. 4). Rarely
these thin, bristle-shaped by-spines are quite simple and straight,
commonly they undulate or are zigzag and often armed with recurved hooks.
Sometimes they are also forked or arborescent (Pl. 138, fig. 7).

_The Central Capsule_ of the Dorataspida is constantly spherical and about
one-third smaller than the enclosing shell, from which it is separated by
the spherical calymma. {807}The membrane of the central capsule is commonly
rather thin, and pierced by the twenty radial spines, meeting in the centre
of the capsule. Between these lie innumerable small pores for the radiating
pseudopodia; however, in many cases (and perhaps constantly) these pores
exhibit a certain regular disposition. In many species the central capsule
encloses Xanthellæ or symbiotic yellow unicellular Algæ. The nucleus
becomes cleft very early in the majority of Dorataspida.

_Synopsis of the Genera of Dorataspida._

-------------------------------------------------------------------------
I. Subfamily Diporaspida.
Each radial spine with two opposite primary apophyses; therefore the
whole shell with forty primary apophyses.
--------------------
I. Tribe Phractaspida.
Twenty radial spines without lattice-plates; no normal pores.
-------------------------------------------------------------------------
No by-spines, 349. _Phractaspis_.

With by-spines, 350. _Pleuraspis_.
-------------------------------------------------------------------------
II. Tribe Ceriaspida.
Twenty radial spines all with lattice-plates produced by union of the
branched apophyses of each plate.
-------------------------------------------------------------------------
{ { No by-spines, 351. _Dorataspis_.
{ Plates not {
Forty parmal { dimpled, { With by-spines, 352. _Diporaspis_.
pores (two { without {
in each { network { No by-spines, but
plate)--no { of crests. { free apophyses, 353. _Orophaspis_.
coronal {
pores in { Plates dimpled, } No by-spines, 354. _Ceriaspis_.
the plates. { with a network }
{ of crests. } With by-spines, 355. _Hystrichaspis_.

Eighty to two hundred or more } No by-spines, 356. _Coscinaspis_.
(in each plate two aspinal }
and two to ten or more } With by-spines, 357. _Acontaspis_.
coronal pores). }
-------------------------------------------------------------------------
II. Subfamily Tessaraspida.
Each radial spine with four crossed primary apophyses; therefore the
whole shell with eighty primary apophyses.
--------------------
III. Tribe Stauraspida.
Twenty radial spines all or partly without lattice-plates.
-------------------------------------------------------------------------
{ No by-spines, 358. _Stauraspis_.
All twenty spines without {
lattice-plates. { With by-spines, 359. _Echinaspis_.

Four plates with, sixteen }
without, lattice-plates. } With by-spines, 360. _Zonaspis_.

Twelve plates with, eight }
without, lattice-plates. } With by-spines, 361. _Dodecaspis_.
-------------------------------------------------------------------------
IV. Tribe Lychnaspida.
Twenty radial spines all with lattice-plates (produced by union of
the branched apophyses of each plate).
-------------------------------------------------------------------------
Eighty parmal pores (four } No by-spines, 362. _Tessaraspis_.
on each plate)--no coronal }
pores in the plates. } With by-spines, 363. _Lychnaspis_.

One hundred and sixty to three }
hundred or more parmal pores } No by-spines, 364. _Icosaspis_.
(in each plate four aspinal }
and four to twelve or more } With by-spines, 365. _Hylaspis_.
coronal pores). }

{808}Subfamily 1. DIPORASPIDA, Haeckel.

_Definition._--#Dorataspida# with twenty radial spines, each of which bears
two opposite apophyses. The spherical shell is composed either of the
meeting branches of these apophyses (Phractaspida), or of twenty perforated
plates, produced by concrescence of their branches (Ceriaspida).

A. Tribe I. #Phractaspida#, Haeckel.

_Definition._--#Dorataspida# without perforated plates; the spherical shell
is composed only of the meeting branches of the two opposite apophyses,
which arise from each radial spine. Therefore the meshes of the shell are
all sutural.

Genus 349. _Phractaspis_,[389] Haeckel, 1881, Prodromus, p. 467.

_Definition._--#Dorataspida# without perforated plates; shell composed only
of the meeting branches of the two opposite apophyses, which arise from
each radial spine. Condyles of the branch-ends without by-spines.

The genus _Phractaspis_ is the most simple and primitive form among all
Dorataspida, and may be regarded as the common ancestral form either of
this whole family, or at least of its first subfamily, the Diporaspida. In
all members of this subfamily the spherical shell is composed of twenty
radial spines, each of which bears two opposite apophyses; but the mode of
composition is different in the two tribes of the subfamily, in the
Phractaspida and Ceriaspida. In the simpler tribe, the Phractaspida, the
shell is composed only of the meeting branches of the apophyses of
neighbouring spines; there are no peculiar perforated plates or shields. In
the Ceriaspida, however, both apophyses of each single spine form a
perforated plate or shield by union of their branches, and the shell is
formed of the meeting edges of these shields. Of course the Ceriaspida must
be derived from the simpler Phractaspida. _Phractaspis_, as the common
ancestral form of both, exhibits a very simple structure of the shell (Pl.
137, figs. 1, 2). Commonly, if the fork-branches of each apophysis be not
again branched, the shell possesses only twenty-two large meshes and forty
sutures. More rarely their number increases, the fork-branches of the
apophyses being again branched (_Phractaspidium_, Pl. 137, fig. 3).

Subgenus 1. _Phractasparium_, Haeckel.

_Definition._--Shell with twenty-two meshes, and forty sutures, each spine
with only four branches, its two apophyses being simply forked.

{809}1. _Phractaspis prototypus_, n. sp. (Pl. 137, fig. 2).

Radial spines cylindrical, thin, pointed; the outer and inner halves of
nearly equal size. Each spine bears in its middle part two opposite
apophyses, which are simply forked; the four condyles of each spine (or the
thickened ends of the fork-branches) are united with the meeting condyles
of the neighbouring spines by sutures. Therefore the network of the
spherical shell is composed of twenty-two large meshes: two square polar
meshes on each pole (_a a a a_ and _e e e e_); eight triangular circumpolar
meshes (each between two polar and one tropical spine, _a b a_ and _e d
e_); eight tropical rhomboidal meshes (each between one polar, one
equatorial, and two tropical spines: _a b c b_ and _e d c d_); and four
rhomboidal equatorial meshes (each between two tropical and two equatorial
spines _c b c d_).

_Dimensions._--Diameter of the shell 0.1, of the meshes 0.03 to 0.04;
breadth of the spines and bars 0.005.

_Habitat._--Cosmopolitan; Mediterranean (Corfu), Atlantic (Stations 348,
354), Indian Ocean (Ceylon), Pacific (Stations 253, 265, 274), &c.,
surface.

2. _Phractaspis complanata_, n. sp. (Pl. 137, fig. 1).

Radial spines leaf-shaped, strongly compressed, two-edged, pointed; their
outer part longer than the inner. Each spine with two opposite apophyses
which are simply forked, therefore with four condyles. The network with
forty sutures and twenty-two large meshes, as in the foregoing species. The
broad faces of the eight polar spines lie in two meridian planes, of the
four equatorial spines in the equatorial plane, of the eight tropical
spines in two planes parallel to the latter. The planes of the leaf-shaped
fork-branches lie in the spherical face.

_Dimensions._--Diameter of the shell 0.1, of the meshes 0.02 to 0.03;
breadth of the fork-branches 0.012.

_Habitat._--South Atlantic, Station 343, surface.

3. _Phractaspis condylophora_, n. sp.

Radial spines quadrangular, thin; their outer pyramidal part shorter than
the inner. Each spine with four branches, its two opposite apophyses being
simply forked. The eighty condyles (or sutural ends of the branches) much
thickened, twice to four times as broad as the branches themselves. Network
with forty sutures and twenty-two large meshes, as in both foregoing
species.

_Dimensions._--Diameter of the shell 0.12, of the meshes 0.03 to 0.04;
breadth of the condyles 0.01.

_Habitat._--South Pacific, Station 288, surface.

4. _Phractaspis bipennis_, Haeckel.

_Dorataspis bipennis_, Haeckel, 1862, Monogr. d. Radiol., p. 413, Taf.
xxi. figs. 1, 2.

_Phractasplenium bipenne_, Haeckel, 1882, Manuscript.

Radial spines very thin, quadrangular; their outer pyramidal part shorter
than the inner. Each spine with four bent branches, its two opposite
apophyses being simply forked. Eighty {810}condyles, very thin, pointed. In
the specimens of this remarkable species, which I first observed in
Messina, two opposite equatorial spines had quite free apophyses, not
connected with the neighbouring spines; therefore the thin lattice-work of
the shell exhibited only thirty-six sutures and twenty meshes (two meshes
with six sutures, six meshes with four sutures, and twelve meshes with
three sutures). In similar specimens, which I afterwards observed in the
Canary Islands, all four equatorial spines were connected in the same
manner with the neighbouring spines; therefore they possessed forty sutures
and twenty-two meshes, like _Phractaspis prototypus_ (Pl. 137, fig. 2).
Perhaps the Mediterranean species represents a peculiar genus,
_Phractasplenium bipenne_.

_Dimensions._--Diameter of the shell 0.1, of the meshes 0.03 to 0.05;
breadth of the spines 0.002.

_Habitat._--Mediterranean (Messina), Canary Islands (Lanzerote), surface.

Subgenus 2. _Phractaspidium_, Haeckel.

_Definition._--Shell with forty to eighty or more meshes, and eighty to one
hundred or more sutures. Each spine with six to eight or more branches, its
two apophyses being doubly forked or more ramified.

5. _Phractaspis constricta_, n. sp. (Pl. 137, fig. 3).

Radial spines strongly compressed, two-edged, pointed; their outer half
twice constricted and somewhat longer than the inner half. Each spine with
two opposite forked apophyses, the branches of which are again forked;
therefore eight condyles on each spine. The network of the spherical shell
with eighty sutures and sixty-two meshes (twenty-two large primary meshes
and forty smaller secondary meshes, the latter between the distal
fork-branches).

_Dimensions._--Diameter of the shell 0.11, of the large meshes 0.04 to
0.05, of the small meshes 0.01; breadth of the spines 0.01.

_Habitat._--South Atlantic, Station 348, depth 2450 fathoms.

6. _Phractaspis cataphracta_, Haeckel.

_Acanthometra cataphracta_, J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 49, Taf. x. figs. 7, 8.

_Dorataspis cataphracta_, Haeckel, 1862, Monogr. d. Radiol., p. 415.

Radial spines thin, quadrangular; their outer pyramidal part shorter than
the inner. Each spine with six to eight condyles, the fork-branches of
their two opposite apophyses being (all or partly) again forked. The
network with sixty to eighty sutures and meshes: sometimes as regular as in
the preceding species, at other times more or less irregular.

_Dimensions._--Diameter of the shell 0.1, of the meshes 0.02 to 0.04;
breadth of the bars 0.004 to 0.008.

_Habitat._--Mediterranean (Cette), Müller (Messina), Haeckel; North
Atlantic, Station 353, surface.

{811}Genus 350. _Pleuraspis_,[390] Haeckel, 1881, Prodromus, p. 467.

The genus _Pleuraspis_ has quite the same structure of the shell as the
foregoing _Phractaspis_, and differs from it only in the development of
external by-spines; commonly each condyle of the branch end of the
apophyses bears one zigzag by-spine, which is directed parallel to the
radial main-spine from which the apophyses arise. Therefore each suture of
the shell is armed with two divergent by-spines (Pl. 137, fig. 4).

Subgenus 1. _Pleurasparium_, Haeckel.

_Definition._--Shell regularly developed with twenty-two meshes and forty
sutures (sometimes twenty to twenty-four meshes and thirty-six to
forty-eight sutures); each spine commonly with four branches, its two
apophyses being simply forked.

1. _Pleuraspis horrida_, n. sp. (Pl. 137, fig. 4).

Radial spines roundish, somewhat compressed, very thick, conical, pointed
at both ends, outer part nearly twice as long as the inner. Two apophyses
of each spine simply forked, with short and broad branches and thickened
condyles. Forty sutures, very broad. Twenty-two large meshes three to
four times as broad as the bars. By-spines much developed, as long as the
radius, zigzag, with alternating recurved hooks.

_Dimensions._--Diameter of the shell 0.15, of the meshes 0.04 to 0.06;
breadth of the bars 0.01 to 0.015.

_Habitat._--Tropical Pacific (east of Philippines), Station 215, surface.

2. _Pleuraspis amphithecta_, n. sp.

Radial spines two-edged, leaf-shaped, strongly compressed, pointed at both
ends; outer half longer than the inner. Two apophyses of each spine simply
forked, with short and very broad branches; condyles not thickened. Forty
sutures, broad. Twenty-two large meshes twice to three times as broad as
the bars. By-spines zigzag, half as long as the radius. (Resembles
_Phractaspis complanata_, Pl. 137, fig. 1.)

_Dimensions._--Diameter of the shell 0.12, of the meshes 0.02 to 0.03; bars
0.012.

_Habitat._--Central Pacific, Station 274, surface.

{812}3. _Pleuraspis costata_, Haeckel.

_Acanthometra costata_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 49, Taf. ii. fig. 1, Taf. x. figs. 4-6.

_Dorataspis costata_, Haeckel, 1862, Monogr. d. Radiol., p. 414, Taf.
xxiii. fig. 1.

Radial spines roundish, conical, pointed at both ends; outer and inner
halves of nearly equal length. Two apophyses of each spine simply forked,
with broad branches and thickened condyles. Forty sutures, broad.
Twenty-two large meshes, four to five times as broad as the bars. By-spines
straight, denticulate, half as long as the radius. (This common species is
rather variable in size and details.)

_Dimensions._--Diameter of the shell 0.1 to 0.13, meshes 0.03 to 0.06, bars
0.005 to 0.01.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

4. _Pleuraspis pyramidalis_, n. sp.

Radial spines quadrangular, their outer part pyramidal, very thick, about
half as long as the prismatic inner part. Two apophyses of each spine
simply forked, with very short and broad branches. Condyles much thickened.
Forty sutures, very broad. Twenty-two meshes small, the largest twice as
broad as the bars; the smaller only half as broad. By-spines short,
straight, denticulate, half as long as the radius.

_Dimensions._--Diameter of the shell 0.14, meshes 0.005 to 0.03.

_Habitat._--Western Tropical Pacific, Station 222, surface.

Subgenus 2. _Pleuraspidium_, Haeckel.

_Definition._--Shell with forty to eighty or more meshes and eighty to one
hundred or more sutures; each spine with six to eight or more branches, its
two apophyses being forked twice or oftener.

5. _Pleuraspis ramosa_, n. sp.

Radial spines roundish, stout, somewhat compressed; their inner and outer
half nearly of equal length. Two apophyses of each spine doubly forked or
more irregularly branched; therefore commonly eight (sometimes six, seven,
nine, or ten to twelve) thickened condyles on each spine. Sutures eighty to
one hundred, broad. Irregular meshes sixty to eighty; three to five times
as broad as the bars. By-spines zigzag, ramified, half as long as the
radius.

_Dimensions._--Diameter of the shell 0.15, of the meshes 0.007 to 0.014,
breadth of the bars 0.003.

_Habitat._--Central Pacific, Station 272, surface.

{813}B. Tribe II. #Ceriaspida#, Haeckel.

_Definition._--#Dorataspida#, the spherical shell of which is composed of
twenty perforated plates, produced by union of the branches of the two
opposite apophyses, which arise from each radial spine. Therefore the
meshes of the shell are partly sutural, partly parmal.

Genus 351. _Dorataspis_,[391] Haeckel, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 811.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
forty aspinal pores (two pores in each plate). Surface of the shell without
combs, dimples, and by-spines.

The genus _Dorataspis_ opens the long series of the Ceriaspida, or of those
Dorataspida in which the shell is composed of twenty plates, each of which
is perforated by two primary aspinal pores. This tribe has arisen from the
simpler Phractaspida by reunion of the branches of the apophyses in each
single spine. If in _Phractaspis_, their common ancestral form (Pl. 137,
figs. 1, 2), the neighbouring fork-branches of both opposite apophyses in
each radial spine become recurved and united, they will produce a shield,
which is perforated by two pores and between them by the spine itself.
These two "primary aspinal pores" are characteristic of all Ceriaspida;
among these _Dorataspis_ itself is the most simple form. Its shell exhibits
therefore constantly forty aspinal pores, and besides these a variable
number of "sutural pores" (in the sutures between the twenty plates). If in
each suture there be only a single pore, we get on the whole fifty-two or
fifty-four pores; if in each suture there be two or three pores, that
number becomes doubled or tripled.

Subgenus 1. _Doratasparium_, Haeckel.

_Definition._--Shell with fifty-two sutures; four polar plates on each pole
of the main axis meeting in one point (in the pole itself); therefore all
eight polar plates pentagonal and of equal size. Spherical shell therefore
composed of four (equatorial) hexagonal plates and of sixteen pentagonal
(eight tropical and eight polar plates).

1. _Dorataspis fusigera_, n. sp. (Pl. 138, fig. 2).

Shell with fifty-two sutures and fifty-two sutural meshes, with four
hexagonal and sixteen pentagonal plates. Both aspinal meshes of each plate
elliptical, twice to four times as broad as the {814}sutural meshes. Radial
spines in the outer half fusiform, somewhat longer than the inner
cylindrical half.

_Dimensions._--Diameter of the shell 0.08 to 0.09, of the parmal meshes 0.1
to 0.2, of the sutural meshes 0.004 to 0.01.

_Habitat._--Central Pacific, Station 271, surface.

2. _Dorataspis macracantha_, n. sp.

Shell with fifty-two sutures and fifty-two sutural meshes; with four
hexagonal and sixteen pentagonal plates. Both aspinal meshes of each plate
circular, small, twice as broad as the small sutural meshes. Radial spines
quadrangular, prismatic, their outer half twice to three times as long as
the inner half.

_Dimensions._--Diameter of the shell 0.12, of the parmal pores 0.008, of
the sutural meshes 0.004.

_Habitat._--South Pacific, Station 288, surface.

3. _Dorataspis macropora_, n. sp.

Shell with fifty-two sutures and one hundred to one hundred and fifty
sutural meshes, with four hexagonal and sixteen pentagonal plates. Both
aspinal meshes nearly circular, very large, five to ten times as broad as
the small circular sutural meshes; the number of the latter is in this
species multiplied, in each suture being two to three (commonly three)
small pores. Radial spines in the outer half leaf-shaped, compressed,
nearly lanceolate, longer than in the inner cylindrical half.

_Dimensions._--Diameter of the shell 0.16, of the parmal pores 0.03 to
0.04, of the sutural pores 0.004 to 0.006.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

Subgenus 2. _Dorataspidium_, Haeckel.

_Definition._--Shell with fifty-four sutures; four polar plates on each
pole of the main axis different in pairs; two major hexagonal meeting in a
polar suture (the "hydrotomical suture"); two minor pentagonal, not meeting
together (separated by that hydrotomical suture). Spherical shell therefore
composed of eight hexagonal plates (four equatorial and four polar) and by
twelve pentagonal plates (eight tropical and four polar plates).

4. _Dorataspis loricata_, Haeckel.

_Dorataspis loricata_, Haeckel, 1862, Monogr. d. Radiol., p. 415, Taf.
xxi. figs. 3-6.

Shell with fifty-four sutures and fifty-four sutural meshes, with eight
hexagonal and twelve pentagonal plates. Both aspinal meshes of each plate
kidney-shaped or roundish, nearly of the {815}same size as the sutural
meshes. Radial spines cylindrical; their outer half on the pointed distal
end two-edged and longer than the inner half.

_Dimensions._--Diameter of the shell 0.14, of its meshes 0.008 to 0.016.

_Habitat._--Mediterranean (Messina), North Atlantic (Canary Islands),
Azores, surface.

5. _Dorataspis typica_, n. sp. (Pl. 138, figs. 4, 4_a_).

Shell with fifty-four crest-like sutures and fifty-four circular sutural
meshes, with eight hexagonal and twelve pentagonal plates. Both aspinal
meshes of each plate elliptical, twice to three times as broad as the
sutural meshes. Aspinal meshes of the four equatorial plates twice as broad
as those of the sixteen other plates. The meshes are surrounded by elevated
and denticulated crests (incipient spine-sheaths). Radial spines
compressed, more or less two-edged; outer pointed half somewhat larger than
the inner half. (In this typical species the composition of the shell
prevailing in the majority of Ceriaspida is very clear.)

_Dimensions._--Diameter of the shell 0.15 to 0.18, of the aspinal pores
0.02 to 0.03, of the sutural pores 0.01.

_Habitat._--Central Pacific, Stations 270 to 274, surface and in various
depths.

6. _Dorataspis micropora_, n. sp. (Pl. 138, fig. 3).

Shell with fifty-four sutures and fifty-four very small sutural meshes,
with eight hexagonal and twelve pentagonal plates. Both aspinal meshes of
each plate elliptical, four to six times as broad as the sutural meshes.
Radial spines in the outer half conical, somewhat shorter than in the inner
cylindrical half.

_Dimensions._--Diameter of the shell 0.11, of the aspinal pores 0.01 to
0.02, of the sutural pores 0.003.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

7. _Dorataspis gladiata_, n. sp.

Shell with fifty-four sutures and one hundred to one hundred and twenty
sutural meshes, with eight hexagonal and twelve pentagonal plates. Both
aspinal meshes elliptical, three times as broad as the small sutural
meshes; commonly two meshes in each suture. Radial spines sword-shaped,
two-edged, tapering from the shell towards the two ends. (Resembles
_Belonaspis datura_, Pl. 139, fig. 9, but differs in the spherical shell,
the equal size of the four equatorial spines, and the absence of
by-spines.)

_Dimensions._--Diameter of the shell 0.12, of the aspinal pores 0.012, of
the sutural pores 0.004.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

8. _Dorataspis polypora_, n. sp.

Shell with fifty-four sutures and one hundred and sixty to two hundred
sutural meshes, with eight hexagonal and twelve pentagonal plates. Commonly
three meshes (sometimes four) on {816}each suture. Both aspinal meshes
kidney-shaped, four times as broad as the small sutural meshes. Radial
spines thick, in the inner longer part cylindrical, in the outer shorter
part conical. (Differs from all other species in the multiplication of the
sutural pores.)

_Dimensions._--Diameter of the shell 0.2, of the aspinal pores 0.016, of
the sutural pores 0.004.

_Habitat._--Equatorial Atlantic, Station 347, surface.

Genus 352. _Diporaspis_,[392] n. gen.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
forty aspinal pores (two pores in each plate). Surface of the shell without
combs and dimples, but armed with numerous by-spines.

The genus _Diporaspis_ has the same characteristic structure of the shell
as the typical _Dorataspis_, and differs from it only in the development of
numerous by-spines on the surface. The number of the sutures between the
twenty plates is sometimes fifty-two, at other times fifty-four, and in
each suture we find occasionally a single pore, at other times two or three
such pores.

Subgenus 1. _Diporasparium_.

_Definition._--Shell with fifty-two sutures, four polar plates on each pole
of the main axis meeting in one common point; therefore all eight polar
plates pentagonal and of equal size. Shell therefore composed of four
(equatorial) hexagonal plates, and of sixteen pentagonal (eight tropical
and eight polar) plates.

1. _Diporaspis nephropora_, n. sp. (Pl. 134, fig. 15).

Shell with fifty-two sutures and fifty-two sutural pores, with four
hexagonal and sixteen pentagonal plates. Both aspinal pores of each plate
kidney-shaped, about twice as broad as the circular sutural pores. Radial
spines thin, cylindrical, longer than the radius. By-spines forked, only
one-third as long as the radius.

_Dimensions._--Diameter of the shell 0.13, parmal pores 0.03, sutural pores
0.07.

_Habitat._--North Pacific, Station 244, surface.

2. _Diporaspis circopora_, n. sp.

Shell with fifty-two sutures and one hundred to one hundred and fifty
sutural pores, with four hexagonal and sixteen pentagonal plates. Both
aspinal pores of each plate circular, very large, six to eight times as
broad as the small circular sutural pores (in each suture two to three
pores). {817}Radial spines strongly compressed, two-edged; outer half
shorter than the inner. By-spines undulate, half as long as the radius.

_Dimensions._--Diameter of the shell 0.12, of the parmal pores 0.02 to
0.03, of the sutural pores 0.003 to 0.004.

_Habitat._--South Pacific, Station 287, surface.

Subgenus 2. _Diporaspidium_, Haeckel.

_Definition._--Shell with fifty-four sutures, four polar plates on each
pole of the main axis different in pairs: two major hexagonal meeting in a
polar ("geotomical") suture, two minor pentagonal, not meeting together
(separated by that suture). Shell therefore composed of eight hexagonal
plates (four equatorial and four polar) and of twelve hexagonal plates
(eight tropical and four polar).

3. _Diporaspis zygopora_, n. sp.

Shell with fifty-four sutures and fifty-four circular sutural pores: with
eight hexagonal and twelve pentagonal plates. Both aspinal pores of each
plate elliptical, three times as broad as the sutural pores. Radial spines
compressed, two-edged; outer half shorter than the inner. By-spines very
numerous, simple, one-third as long as the radius, forming coronels or
elegant circles around the pores (a small coronel around each sutural pore,
a large one around each couple of aspinal pores).

This typical species is nearly allied to _Dorataspis typica_ (Pl. 138, fig.
4), and may be derived from it by development of the coronels of by-spines.

_Dimensions._--Diameter of the shell 0.16, aspinal pores 0.03, sutural
pores 0.01.

_Habitat._--Central Pacific, Station 271, surface

Genus 353. _Orophaspis_,[393] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
forty aspinal pores (two pores in each plate). Surface of the shell without
combs, dimples, and by-spines. Each radial spine bears outside of the shell
two opposite free apophyses, which are either simple or branched.

The genus _Orophaspis_ differs not only from its ancestral form,
_Dorataspis_, but from all other Dorataspida in the development of peculiar
free apophyses on the radial spines, outside the shell. These apophyses,
two being opposite on each spine, appear as a repetition of the primary
apophyses of _Phractaspis_; they are either simple or branched, and
sometimes the branches are united together, forming an outer free shield
with two or four pores. These outer plates represent the beginning of a
second outer shell and form the transition to _Phractopelta_, the ancestral
form of the Phractopeltida.

{818}Subgenus 1. _Orophasparium_, Haeckel.

_Definition._--Free apophyses of the radial spines simple, not branched.

1. _Orophaspis astrolonche_, n. sp.

Parmal pores of the shell circular, twice as large as the sutural pores and
as the breadth of the bars. Radial spines very long, three to six times as
long as the diameter of the shell, compressed, two-edged; each with two
simple, opposite, triangular apophyses; their distance from the shell equal
to its diameter.

_Dimensions._--Diameter of the shell 0.05, parmal pores 0.004, sutural
pores 0.002.

_Habitat._--North Pacific, Station 239, surface.

2. _Orophaspis gladiata_, n. sp.

Parmal pores of the shell circular, of the same size as the sutural pores
and the bars. Radial spines thick, spindle-shaped, scarcely longer than the
diameter of the shell, each with two simple opposite conical apophyses;
their distance from the shell about equal to its radius.

_Dimensions._--Diameter of the shell 0.06, pores 0.003.

_Habitat._--Central Pacific, Station 265, surface.

Subgenus 2. _Orophaspidium_, Haeckel.

_Definition._--Free apophyses of the radial spines branched, their branches
free (not anastomosing).

3. _Orophaspis furcata_, n. sp. (Pl. 133, fig. 6).

Parmal pores of the shell roundish or elliptical, three times as broad as
the sutural pores and the bars. Radial spines very long, compressed, each
with two opposite apophyses, which are simply forked; their distance from
the shell somewhat greater than its diameter.

_Dimensions._--Diameter of the shell 0.06 to 0.08, parmal pores 0.006,
sutural pores 0.002.

_Habitat._--Central Pacific, Station 274, surface.

4. _Orophaspis ramosa_, n. sp.

Parmal pores of the shell circular, of the same size as the sutural pores
and the bars. Radial spines compressed, very long, each with two opposite
apophyses, which are more or less irregularly branched (commonly between
eight and twelve thin branches on each spine); their distance from the
shell smaller than its diameter.

_Dimensions._--Diameter of the shell 0.08, pores 0.006.

_Habitat._--North-West Pacific, Station 235, surface.

{819}Subgenus 3. _Stegaspis_, Haeckel, 1881, Prodromus, p. 468.

_Definition._--Free apophyses of the radial spines branched, and by reunion
of their anastomosing branches forming perforated shields.

5. _Orophaspis diporaspis_, n. sp.

Parmal pores of the shell circular, of the same size as the sutural pores
and the bars. Radial spines sword-shaped, two-edged, very broad; each spine
bears an elliptical free shield with two longish pores (opposite on the two
flat sides of the sword). Distance of the twenty free two-pored shields
from the shell about equal to its diameter.

_Dimensions._--Diameter of the shell 0.04, of the pores 0.004.

_Habitat._--South Pacific, Station 295, surface.

6. _Orophaspis tessaraspis_, n. sp.

Parmal pores of the shell roundish, somewhat larger than the sutural pores
and the bars. Radial spines very long, compressed; each spine bears a
roundish free shield with four irregular pores disposed in a cruciform
manner, the two larger pores being opposite on the flat sides of the spine,
the two smaller being opposite but on its two edges. Distance of the twenty
free four-pored shields from the shell about equal to its radius.

_Dimensions._--Diameter of the shell 0.05, of the pores 0.003.

_Habitat._--North Pacific, Station 239, surface.

Genus 354. _Ceriaspis_,[394] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
forty aspinal pores (two pores in each plate). Surface of the shell with
numerous dimples, separated by a network of elevated combs. No by-spines.

The genus _Ceriaspis_ has the same structure of the shell, as its ancestral
form _Dorataspis_, differing from it only in the development of high combs
or crests, which form on the surface of the shell a peculiar elevated
network. The dimples or funnel-shaped pits between these combs are either
all or partly perforated by the pores of the shell. Both the aspinal pores
of each plate are usually placed in one common dimple, whilst each sutural
pore is placed in its peculiar smaller dimple.

Subgenus 1. _Ceriasparium_, Haeckel.

_Definition._--Shell-surface with seventy to one hundred or more small
funnel-shaped dimples, each of which opens on the bottom by one pore or by
a couple of {820}apertures; twenty larger dimples in the centre of the
plates (each with a couple of aspinal pores) and fifty to one hundred or
more smaller dimples, each of which contains one sutural pore. No blind
dimples between the perforated dimples.

1. _Ceriaspis lacunosa_, n. sp.

Shell spherical with seventy-two funnel-shaped dimples, each of which is
perforated on the bottom by one or two apertures; twenty larger dimples in
the centre of the plates, each with two elliptical aspinal pores, and
fifty-two smaller sutural dimples between them, each with one circular pore
of half the size. No blind dimples. Radial spines quadrangular, stout;
their outer part shorter than the inner.

_Dimensions._--Diameter of the shell 0.1, of the forty parmal pores 0.01,
of the fifty-two sutural pores 0.005.

_Habitat._--South Atlantic, Station 330, surface.

2. _Ceriaspis scrobiculata_, n. sp.

Shell spherical, with seventy-four funnel-shaped dimples, each of which is
perforated on the bottom by one or two apertures; twenty larger dimples in
the centre of the plates, each with two kidney-shaped large pores, and
fifty-four smaller dimples on the sutures, each with one circular pore of
one-fourth of the size of the reniform pores. No blind dimples. Radial
spines cylindrical, the outer part longer than the inner.

_Dimensions._--Diameter of the shell 0.12, of the forty parmal pores 0.016,
of the fifty-four sutural pores 0.004.

_Habitat._--Central Pacific, Station 270, surface.

Subgenus 2. _Ceriaspidium_, Haeckel.

_Definition._--Shell-surface with funnel-shaped dimples (commonly one
hundred and seventy-six or one hundred and eighty-two), which on the bottom
are partly closed, partly perforated by one aperture (or by a pair of
pores). The blind dimples are placed on the corners of the twenty plates,
and are therefore either one hundred and four or one hundred and eight; if
there be no polar suture, the blind dimples are one hundred and four
(twenty-four on the four hexagonal equatorial plates, forty on the eight
pentagonal tropical plates, and forty on the eight pentagonal polar
plates); if, however, there be a polar suture on both main poles, the
number of blind dimples is one hundred and eight (twenty-four on the four
hexagonal equatorial and twenty-four on the four hexagonal polar plates,
two opposite on each pole; forty on the eight pentagonal tropical plates
and twenty on the four pentagonal polar plates, two opposite on each pole).
Between the blind dimples there are usually seventy-two to seventy-four
perforated {821}dimples, twenty larger parmal dimples (enclosing a radial
spine and a couple of aspinal pores) and fifty-two to fifty-four sutural
dimples (sometimes one hundred or more), each of which encloses one
sutural pore.

3. _Ceriaspis inermis_, n. sp. (Pl. 138, fig. 5).

Shell spherical, with one hundred and seventy-six funnel-shaped dimples,
one hundred and four of which are blind and seventy-two perforated; of the
latter, each of the fifty-two smaller contains a single sutural pore, each
of the twenty larger a couple of aspinal pores. The elliptical or
kidney-shaped aspinal pores are about twice as broad as the circular
sutural pores. Radial spines thin, compressed, two-edged; their outer
prolongation very short, rudimentary, scarcely higher than the axis of the
surrounding funnel, and projecting but slightly from its aperture.

_Dimensions._--Diameter of the shell 0.15, of the forty parmal pores 0.016,
of the fifty-two sutural pores 0.008.

_Habitat._--South Pacific, Station 289, surface.

4. _Ceriaspis favosa_, n. sp. (Pl. 138, fig. 6).

Shell polyhedral (icosahedral), with one hundred and eighty-two
funnel-shaped dimples, one hundred and eight of which are blind and
seventy-four perforated; of the latter, each of the fifty-four smaller
contains a single sutural pore, each of the twenty larger a couple of
aspinal pores. All pores roundish, nearly of the same size. Radial spines
thin, compressed, two-edged; their outer part somewhat longer than the
inner.

_Dimensions._--Diameter of the shell 0.12 to 0.13, of the pores 0.01 to
0.015.

_Habitat._--Central Pacific, Station 274, surface.

5. _Ceriaspis icosahedra_, n. sp.

Shell polyhedral (icosahedral), with one hundred and eighty-two
funnel-shaped dimples, one hundred and eight of which are blind and
seventy-four perforated; of the latter, each of the fifty-four smaller
contains one small sutural pore, each of the twenty larger a couple of
aspinal pores. The kidney-shaped aspinal pores are very large, four to six
times as broad as the small circular sutural pores. Radial spines strong,
quadrangular, their outer part from two to three times as long as the inner
part.

_Dimensions._--Diameter of the shell 0.16, of the parmal pores 0.015 to
0.02, of the sutural pores 0.003 to 0.004.

_Habitat._--Tropical Atlantic, Station 349, surface.

6. _Ceriaspis cicatricosa_, n. sp.

Shell spherical, with two hundred and fifty to three hundred (or more)
small funnel-shaped dimples, the majority of which are blind, the minority
perforated by pores; of the latter forty {822}are aspinal pores (two at the
base of each spine), the others sutural pores. All pores nearly of the same
size. As the shell of this species is very dark and thick-walled, it was
impossible to obtain more information with regard to the pores. Radial
spines very stout; the outer conical part shorter than the inner
cylindrical part.

_Dimensions._--Diameter of the shell 0.2, of the pores 0.006 to 0.008.

_Habitat._--North Pacific, Station 236, surface.

Genus 355. _Hystrichaspis_,[395] n. gen.

The genus _Hystrichaspis_ has the same structure with regard to the shell,
as its ancestral form _Ceriaspis_, and differs from the latter only in
possessing numerous by-spines. Moreover, in the former the dimples of the
shell-surface are either all perforated by pores, or some of them are
blind.

Subgenus 1. _Hystrichasparium_, Haeckel.

_Definition._--Shell with seventy to one hundred or more funnel-shaped
dimples, each of which opens on the bottom by one aperture or by a couple
of pores; twenty larger dimples in the centre of the plates (each with a
couple of pores, sometimes also three couples) and fifty to one hundred or
more smaller dimples, each of which contains one sutural pore. No blind
dimples between the perforated dimples.

1. _Hystrichaspis pectinata_, n. sp. (Pl. 138, fig. 8).

Shell with numerous (one hundred to two hundred?) funnel-shaped dimples,
each of which is pierced at the bottom by one or two pores; twenty larger
dimples in the centre of the plates (each with two aspinal pores), and
fifty to one hundred and fifty or more smaller dimples (each with one
sutural pore). No blind dimples. Crests between the dimples armed with a
continuous series of simple by-spines. Radial main spines stout, in the
inner half cylindrical, in the outer conical, more or less compressed.

_Dimensions._--Diameter of the shell 0.13, of the pores 0.004 to 0.006.

_Habitat._--Tropical Atlantic, Station 352, surface.

2. _Hystrichaspis furcata_, n. sp. (Pl. 138, fig. 9).

Shell with numerous (one hundred to two hundred?) funnel-shaped dimples,
each of which is pierced at the bottom by one or two pores; twenty larger
dimples in the centre of the plates (each {823}with two large aspinal
pores), and fifty to one hundred and fifty or more smaller dimples (each
with one small sutural pore). No blind dimples. Crests between the
dimples armed with forked by-spines. Radial main spines stout, leaf-shaped,
tapering towards both ends.

_Dimensions._--Diameter of the shell 0.12, parmal pores 0.01, sutural pores
0.005.

_Habitat._--Central Pacific, Station 276, surface.

3. _Hystrichaspis cristata_, n. sp. (Pl. 138, fig. 11).

_Siphonasphis cristata_, Haeckel, 1882, Manuscript.

Shell with numerous (one hundred to two hundred?) funnel-shaped dimples,
each of which is pierced at the bottom by one or more pores. Twenty larger
dimples in the centre of the plates are pierced by the radial main-spines;
among these fourteen contain each a couple of aspinal pores; six others are
much larger, and contain each six larger pores; these six plates are two
opposite equatorial plates and four polar plates, placed in the same
meridian plane (the "hydrotomical plane"); in each of these six
"hydrotomical dimples" two pores are placed opposite to one another on the
two edges of the leaf-shaped spine, four others being opposite in pairs on
both flat sides of it. By this peculiar structure this species connects the
true _Hystrichaspis_ with _Hexalaspis_ and _Diploconus_; however the twenty
spines are of equal length, and the shell continues to be spherical. The
twenty radial main-spines are leaf-like and compressed. The crests between
the dimples are dentated by a series of small by-spines.

_Dimensions._--Diameter of the shell 0.15, of the aspinal pores 0.01, of
the sutural pores 0.005.

_Habitat._--North Pacific, Station 240, surface.

Subgenus 2. _Hystrichaspidium_, Haeckel.

_Definition._--Shell-surface with numerous funnel-shaped dimples (commonly
one hundred and seventy-six to one hundred and eighty-two), which on the
bottom are partly closed, partly perforated by one aperture (or by a pair
of pores). The blind dimples are situated on the corners of the twenty
plates; their number is commonly one hundred and four or one hundred and
eight, sometimes more. The perforated dimples, alternating with the former,
are usually seventy-two to seventy-four, sometimes more; twenty larger
parmal dimples (each with a couple of aspinal pores, sometimes also with
three such couples) and fifty-two to fifty-four sutural dimples, sometimes
one hundred or more (each with one sutural pore). (Compare the definition
of _Ceriaspidium_, p. 820.)

4. _Hystrichaspis dorsata_, n. sp. (Pl. 138, fig. 10).

Shell with one hundred and seventy-six funnel-shaped dimples, one hundred
and four of which are blind and seventy-two perforated; of the latter, each
of the fifty-two smaller contains a single {824}sutural pore, each of the
twenty larger a couple of aspinal pores; the elliptical aspinal pores are
twice as broad as the circular sutural pores. The crests between the
dimples are armed with simple scattered short by-spines; usually in the
nodal points, where three combs meet, there exist three or two divergent
by-spines. The twenty radial main-spines are thin and long, compressed.

_Dimensions._--Diameter of the shell 0.22, of the aspinal pores 0.02, of
the sutural pores 0.01.

_Habitat._--Central Pacific, Station 271, surface.

5. _Hystrichaspis armata_, n. sp.

Shell with one hundred and seventy-six funnel-shaped dimples, one hundred
and four of which are blind and seventy-two perforated; of the latter, each
of the fifty-two smaller contains a single sutural pore, each of the twenty
larger a couple of aspinal pores. Crests between the dimples comb-like,
armed with a series of simple by-spines. Twenty main-spines very stout, in
the inner part cylindrical, in the outer shorter part conical.

_Dimensions._--Diameter of the shell 0.25, of the aspinal pores 0.02, of
the sutural pores 0.01.

_Habitat._--South-East Pacific, Station 300, surface.

6. _Hystrichaspis sulcata_, n. sp.

Shell with one hundred and eighty-two funnel-shaped dimples, one hundred
and eight of which are blind and seventy-four perforated; of the latter,
each of the fifty-four smaller contains a single sutural pore, each of the
twenty larger a couple of aspinal pores. All pores nearly of the same size,
very small. Crests between the dimples very high, sulcated, on the free
edge serrate with short very numerous, simple by-spines. Radial main-spines
very stout and short, conical.

_Dimensions._--Diameter of the shell 0.2, of the pores 0.003.

_Habitat._--South Atlantic, Station 325, surface.

7. _Hystrichaspis foveolata_, n. sp.

Shell with very numerous (three hundred to four hundred?) small and deep
funnel-shaped dimples, the majority of which are blind, the minority
perforated; among the latter there are forty larger aspinal pores; the
other smaller (scarcely half as large) pores are sutural. Crests between
the dimples densely armed with simple short by-spines. Radial main-spines
stout, leaf-shaped.

_Dimensions._--Diameter of the shell 0.12, of the aspinal pores 0.01, of
the sutural pores 0.004.

_Habitat._--Tropical Atlantic, Station 338, surface.

8. _Hystrichaspis divaricata_, n. sp.

Shell with numerous (two hundred to three hundred?) funnel-shaped dimples,
the majority of which are blind, the minority perforated; among the latter
there are forty larger aspinal pores. {825}Crests between the dimples armed
with long forked by-spines with divergent fork-branches. Radial spines
cylindrical, very long and thick, longer than the diameter of the shell.

_Dimensions._--Diameter of the shell 0.14, aspinal pores 0.08, sutural
pores 0.03.

_Habitat._--Central Pacific, Station 266, surface.

9. _Hystrichaspis fruticata_, n. sp. (Pl. 138, fig, 7).

Shell with numerous (one hundred and fifty to two hundred?) funnel-shaped
dimples, the majority of which are blind, the minority perforated; forty
aspinal pores elliptical, of the same size as the circular sutural pores.
Crests between the dimples bearing elegant arborescent by-spines. Twenty
radial main-spines long and thin, cylindrical or a little compressed.

_Dimensions._--Diameter of the shell 0.17, pores 0.01.

_Habitat._--Tropical Pacific, Station 275, surface.

10. _Hystrichaspis serrata_, n. sp.

Shell with numerous (two hundred to three hundred?) funnel-shaped dimples,
the majority of which are blind, the minority perforated; forty aspinal
pores of the same size as the sutural pores. Crests between the dimples
covered with denticulated by-spines. Twenty radial main-spines compressed,
two-edged, with serrated edges, about as long as the diameter of the shell.

_Dimensions._--Diameter of the shell 0.21, pores 0.012.

_Habitat._--North Pacific, Station 238, surface.

Genus 356. _Coscinaspis_,[396] n. gen.

_Definition._--#Doratispida# with twenty plates, which are perforated by
eighty to two hundred or more parmal pores (two aspinal and two to ten or
more coronal pores in each plate). Surface without by-spines.

The genus _Coscinaspis_, together with the following nearly allied genus
_Acontaspis_, may be separated from the other Ceriaspida as a peculiar
tribe, _Coscinaspida_. This tribe is characterised by the larger number of
the parmal pores. Whilst in all other Ceriaspida this number is constantly
forty (only two pores in each plate), here it may be from eighty to two
hundred or more; in each shield the two primary "aspinal pores" are
surrounded by a circle of two to ten or more (commonly eight to twelve)
"coronal pores." The number of sutural pores in this group is also usually
larger.

{826}Subgenus 1. _Coscinasparium_, Haeckel.

_Definition._--Plates of the shell smooth, without crests, not dimply.

1. _Coscinaspis peripora_, n. sp. (Pl. 138, fig. 1).

Shell thin walled, with smooth surface, without crests and dimples,
perforated by pores of very different sizes. Forty aspinal pores, roundish
or elliptical, of about the same size as the fifty or sixty violin-shaped
sutural pores, and three to four times as broad as the small roundish
coronal pores, which are irregularly formed and distributed, four to eight
occurring on each plate (altogether one hundred to one hundred and twenty).
Radial spines roundish, somewhat compressed; inner and outer half nearly of
the same length.

_Dimensions._--Diameter of the shell 0.12, aspinal pores 0.01 to 0.012,
coronal pores 0.003.

_Habitat._--North-West Pacific, Station 235, surface.

2. _Coscinaspis stigmopora_, n. sp.

Shell very thin walled, with smooth surface, without crests and dimples,
perforated by pores of very different sizes. Forty aspinal pores, roundish,
of about the same size as the fifty or sixty violin-shaped sutural pores,
their diameter being about one-fifth of that of the plates. Coronal pores
very numerous (two hundred to four hundred) and very small, irregularly
scattered, ten to twenty occurring in each plate. Sutures strongly
denticulated. Radial spines very thin and short, cylindrical.

_Dimensions._--Diameter of the shell 0.12 to 0.15, aspinal and sutural
pores 0.01, coronal pores 0.001 to 0.003.

_Habitat._--North Pacific, Station 257, surface.

3. _Coscinaspis rhacopora_, n. sp.

Shell very thin walled, with smooth surface, without crests and dimples,
perforated by very irregular pores of very different sizes and forms;
commonly more or less lobed or sinuate. Forty aspinal pores and fifty to
eighty sutural pores, larger than the numerous (one hundred to two hundred)
irregularly scattered coronal pores. Sutures very sinuate. Radial spines
cylindrical, thin and long.

_Dimensions._--Diameter of the shell 0.15 to 0.18, aspinal and sutural
pores 0.01 to 0.02, coronal pores 0.001 to 0.005.

_Habitat._--Central Pacific, Station 274.

4. _Coscinaspis coscinopora_, n. sp.

Shell thin walled, with smooth surface, without crests and dimples,
perforated by very numerous pores of circular form, but of different sizes.
Forty aspinal pores and one hundred to two hundred {827}sutural pores much
larger than the coronal pores, which are very small and very numerous
(fifty to sixty on each plate). Sutures sinuate. Radial spines compressed,
outer and inner half nearly of equal length.

_Dimensions._--Diameter of the shell 0.3, aspinal and sutural pores 0.01 to
0.015, coronal pores 0.001 to 0.002.

_Habitat._--Tropical Atlantic, Station 338, surface.

5. _Coscinaspsis polypora_, n. sp. (Pl. 136, fig. 8).

Shell very thin walled, with smooth surface, without crests and dimples,
the two aspinal pores of each plate narrow, lanceolate, ten to twelve times
as long as broad and half as long as the radius of each plate. Coronal
pores irregular, polygonal, very numerous (two hundred to three hundred on
each plate), commonly arranged more or less regularly in ten to twelve
series parallel to the longitudinal diameter of the aspinal spines (ten to
twenty pores in each series). Sutural pores irregular, polygonal, very
numerous. Radial spines very thin and long, cylindrical.

_Dimensions._--Diameter of the shell 0.24; length of the aspinal pores
0.04, breadth 0.004; coronal and sutural pores 0.002 to 0.01.

_Habitat._--Tropical Atlantic, Station 349, surface.

6. _Coscinaspis orthopora_, n. sp.

Shell very thin walled, with smooth surface, without crest and dimples. All
pores of nearly equal size and form; quadrangular, mostly rectangular; one
hundred to two hundred, separated by two systems of parallel bars,
perpendicular one to another, occur in each plate. Sutural pores mostly
triangular. Radial spines very thin and long, cylindrical, somewhat
compressed. (Similar to those of _Phatnaspis lacunaria_, Pl. 136, fig. 9,
but spherical, not ellipsoidal.)

_Dimensions._--Diameter of the shell 0.2, pores 0.008 to 0.012.

_Habitat._--Central Pacific, Station 272, surface.

7. _Coscinaspis parmipora_, n. sp. (Pl. 137, fig. 9).

_Craniaspis parmipora_, Haeckel, 1866, Manuscript.

_Dorataspis parmipora_, Haeckel, 1881, Prodromus, Atlas.

Shell thin walled, with smooth surface, without crests and dimples. There
are no sutural pores, since all twenty plates are connected by perfect
sinuate sutures (therefore this excellent species may perhaps better
represent a peculiar genus, called by me in 1866 _Craniaspis_). All pores
are parmal pores; each plate with two elliptical aspinal pores, which are
twice to five times broader than the numerous, roundish irregularly
scattered coronal pores (thirty to fifty occurring on each plate). The
radial spines are quite internal, that is, not prolonged on the outside of
the shell; in this respect they resemble those of _Sphærocapsa_.

_Dimensions._--Diameter of the shell 0.16, aspinal pores 0.01, coronal
pores 0.002 to 0.005.

_Habitat._--North Atlantic, Canary Islands (Lanzerote), surface.

{828}8. _Coscinaspis isopora_, n. sp. (Pl. 134, figs. 13, 14).

Shell thick walled, with smooth surface, without crests and dimples. All
pores of the shell nearly of the same size and of similar form, about as
broad as the bars between them, and about two hundred in number, viz.,
fifty to sixty (regularly fifty-two or fifty-four) sutural pores and one
hundred and forty to one hundred and fifty parmal pores (forty aspinal
kidney-shaped pores, and one hundred to one hundred and ten coronal
circular pores: six in the angles of each equatorial plate, and five in the
angles of each tropical and polar plate; if the disposition be quite
regular, one hundred and four or one hundred and eight). Radial spines
short, rudimentary, conical.

_Dimensions._--Diameter of the shell 0.2, of the pores and bars 0.012.

_Habitat._--Tropical Pacific, Station 218 (off New Guinea), surface.

Subgenus 2. _Coscinaspidium_, Haeckel.

_Definition._--Surface of the shell dimply, with a network of prominent
crests.

9. _Coscinaspis ceriopora_, n. sp.

Shell thick-walled, with numerous (one hundred and sixty to two hundred?)
funnel-shaped dimples, which are separated by prominent crests; on the
bottom of each dimple there is a simple or double circular pore. If this
species be quite regularly developed, it closely resembles the preceding,
differing from it mainly in the prominent combs of the surface. It
resembles also _Ceriaspis favosa_, Pl. 138, fig. 6; but whilst in this
latter the majority of the dimples are blind, here they are all perforated.
The twenty aspinal dimples (in the centre of each plate) present at the
bottom a couple of pores, all other dimples a single pore. Among the latter
there are fifty to sixty sutural pores and one hundred to one hundred and
ten coronal pores, viz., six in each equatorial plate, and five in each of
the other plates; but the number is not quite constant. Radial spines
strong, in the inner part cylindrical, in the outer conical.

_Dimensions._--Diameter of the shell 0.15, of the pores and bars 0.01.

_Habitat._--Tropical Pacific, Station 215, surface.

Genus 357. _Acontaspis_,[397] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
eighty to two hundred or more parmal pores (in each plate two aspinal and
two to ten or more coronal pores). Surface covered with by-spines.

The genus _Acontaspis_ has the same characteristic structure of the shell
as _Ceriaspis_, differing from it only in the presence of numerous
by-spines. Each plate is perforated by four to sixteen or more (commonly
ten to twelve) parmal pores, the two central of which are primary "aspinal
pores," all the others being secondary "coronal pores."

{829}Subgenus 1. _Acontasparium_, Haeckel.

_Definition._--Plates of the shell not dimply, without prominent crests.

1. _Acontaspis lanceolata_, n. sp.

Shell thin walled, even, without crests and dimples between them,
perforated by about three hundred pores of different sizes: forty aspinal
pores elliptical, about as large as the irregular (fifty to sixty) sutural
pores, and two to four times as broad as the small circular coronal pores
(eight to twelve being on each plate, altogether about two hundred).
Between the pores numerous short conical by-spines. Radial main spines
lanceolate, about as long as the radius. (Similar to _Coscinaspis
peripora_, Pl. 138, fig. 1, but with broad lanceolate spines and numerous
short by-spines.)

_Dimensions._--Diameter of the shell 0.15, aspinal and sutural pores 0.012
to 0.015, coronal pores 0.004 to 0.008.

_Habitat._--South Atlantic, Station 325, surface.

Subgenus 2. _Acontaspidium_, Haeckel.

_Definition._--Surface of the shell dimply, with a network of prominent
crests.

2. _Acontaspis hastata_, n. sp. (Pl. 134, fig. 16).

Shell thick walled, with numerous (one hundred and seventy to one hundred
and ninety) dimples, which are separated by an elevated network of
prominent crests. In the centre of each plate a larger dimple with a couple
of aspinal pores, surrounded by a corona of ten or twelve smaller dimples,
each of which contains a single pore (a coronal and sutural alternately).
All pores circular or roundish, nearly of the same size. If the shell be
quite regularly developed, there are fifty to sixty sutural pores and one
hundred to one hundred and ten coronal pores (six in each equatorial plate,
five in each of the other plates). The knobs of the meeting crests are
conical, and bear each a simple short by-spine. Radial main spines
compressed, at the distal end spear-shaped, with a rhomboidal plate below
the apex, about as long as the radius.

_Dimensions._--Diameter of the shell 0.18, of the pores and bars 0.01.

_Habitat._--Western Tropical Pacific (off the Philippines), Station 215.

3. _Acontaspis furcata_, n. sp.

Shell thick walled, dimply, with a network of crests, of the same
composition as in the foregoing species, with one hundred and seventy to
one hundred and ninety dimples (twenty aspinal, one hundred to one hundred
and ten coronal and fifty to sixty sutural dimples). All dimples and pores
nearly of the same size. Each node of the crested network bears a stout
by-spine, which is forked {830}on the base, with two divergent straight
branches (similar to those of _Hystrichaspis furcata_, Pl. 138, fig. 9).
Radial main spines two-edged, sword-like, nearly as long as the diameter of
the shell.

_Dimensions._--Diameter of the shell 0.13, of the pores 0.008.

_Habitat._--Central Pacific, Station 272, surface.

4. _Acontaspis capillata_, n. sp.

Shell very dark and thick walled, non-transparent, with very numerous
(three hundred to four hundred or more?) deep funnel-shaped dimples, each
of which is perforated by a small circular pore (forty aspinal, two hundred
to three hundred coronal, and fifty to one hundred sutural pores?). The
high crests between the dimples bear very numerous simple by-spines, nearly
half as long as the radius, so that the shell appears covered with hairs.
Radial main spines very long and thin, cylindrical, longer than the
diameter of the shell.

_Dimensions._--Diameter of the shell 0.2, of the pores 0.012.

_Habitat._--North Pacific, Station 244, surface.

Subfamily 2. TESSARASPIDA, Haeckel.

_Definition._--#Dorataspida# with twenty radial spines, each of which bears
four crossed apophyses (opposite in pairs). The spherical shell is composed
either of the meeting branches of these apophyses (Stauraspida), or of
twenty perforated plates, produced by concrescence of their branches
(Lychnaspida).

A. Tribe II. #Stauraspida#, Haeckel, 1881, Prodromus, p. 467.

_Definition._--#Dorataspida# with spherical shell, which is composed either
of the meeting branches of the four crossed apophyses only, or exhibits
four to twelve perforated plates which are produced by the crossed
apophyses of four to twelve radial spines (but never of all twenty spines).
Each plate bears four crossed pores.

Genus 358. _Stauraspis_,[398] Haeckel, 1881, Prodromus, p. 467.

_Definition._--#Dorataspida# without perforated plates; shell composed only
of the meeting branches of the four crossed apophyses, which arise
(opposite in pairs) from each radial spine. Condyles of the branch-ends
without by-spines.

The genus _Stauraspis_ is the most simple and primitive form among all
Tessaraspida, or that subfamily of Dorataspida, in which the shell is
composed of twenty {831}radial spines, each of which bears four crossed
apophyses. The subfamily may be divided into two different tribes, the
Stauraspida and Lychnaspida. In the Stauraspida either all twenty spines,
or a part of them, bear no perforated plates, and the shell is composed
wholly or partially of the meeting branches of their apophyses. In the
Lychnaspida, however, the four apophyses of each single spine form, by
reunion of their recurved branches, a plate or shield with four crossed
aspinal pores. The Lychnaspida represent therefore a more developed stage
in the shell-formation than the simpler Stauraspida. _Stauraspis_, as the
common ancestral form of both, may be derived phylogenetically from
_Xiphacantha_ or _Stauracantha_, which differ only by the apophyses or
branches of the apophyses not meeting. These branches (originally eight on
each spine) are either simple or again branched.

Subgenus 1. _Staurasparium_, Haeckel.

_Definition._--Apophyses of the spines simple, not branched; therefore each
spine with four sutural condyles.

1. _Stauraspis cruciata_, n. sp. (Pl. 134, fig. 5).

Radial spines thin, quadrangular, prismatic; outer and inner half nearly of
equal length. Central bases pyramidal, with wing-like edges. Four apophyses
of each spine simple, not branched, with thin condyles. Large meshes of the
shell ten to twenty times as broad as the bars. This and the following
species greatly resemble the simplest forms of _Phractaspis_ (Pl. 137,
figs. 1, 2); they differ from these, however, by the equal size and
distance of the four branches of each spine, which thus form a rectangular
cross.

_Dimensions._--Diameter of the shell 0.1; breadth of the spines and bars
0.002.

_Habitat._--Central Pacific, Station 268, surface.

2. _Stauraspis xiphacantha_, n. sp.

Radial spines stout, cylindrical in the inner half, conical in the shorter
outer half. Four apophyses of each spine simple, not branched, broad, with
thick condyles. Meshes of the shell six to eight times as broad as the
bars.

_Dimensions._--Diameter of the shell 0.12; breadth of the spines and bars
0.008 to 0.01.

_Habitat._--South Pacific, Station 290, surface.

Subgenus 2. _Stauraspidium_, Haeckel.

_Definition._--Apophyses of the spines branched; therefore each spine with
eight to twenty or more sutural condyles.

{832}3. _Stauraspis furcata_, n. sp.

Radial spines thin, quadrangular, prismatic; outer and inner halves nearly
of equal length. The four apophyses of each spine simply forked (or partly
with bifid fork-branches); each spine with eight to twelve sutural
condyles. Meshes of the shell ten to twenty times as broad as the bars.

_Dimensions._--Diameter of the shell 0.15; breadth of the spines and bars
0.003.

_Habitat._--Central Pacific, Station 266, surface.

4. _Stauraspis stauracantha_, n. sp. (Pl. 137, figs. 5, 6).

Radial spines thin, in the inner longer half cylindrical, in the outer half
conical with thickened base. Four apophyses of each spine doubly forked or
dichotomously (more or less irregularly) branched; each spine with sixteen
to twenty-four sutural condyles. Meshes of the shell of very different
sizes and forms; the largest ten to fifteen, the smallest two to three
times as broad as the bars.

_Dimensions._--Diameter of the shell 0.14; breadth of of the spines 0.002
to 0.01, of the bars 0.002.

_Habitat._--Tropical Atlantic, Station 343, surface.

Genus 359. _Echinaspis_,[399] Haeckel, 1881, Prodromus, p. 467.

_Definition_--#Dorataspida# without perforated plates; shell composed only
of the meeting branches of the four crossed apophyses, which arise
(opposite in pairs) from each radial spine. Condyles of the branch-ends
bearing by-spines.

The genus _Echinaspis_ exhibits the same structure of the shell as its
ancestral form _Stauraspis_, and differs from it only in the development of
by-spines on the sutural condyli.

1. _Echinaspis dichotoma_, n. sp.

Radial spines cylindrical, thin, outer half longer than the inner. Four
apophyses of each spine simply forked (or partly with bifid fork-branches);
therefore each spine usually possesses eight (sometimes ten to twelve)
sutural condyles. Meshes of the shell ten to twelve times as broad as the
bars. Each condyle bears a zigzag by-spine, half as long as the radius.

_Dimensions._--Diameter of the shell 0.12, of the spines 0.004 to 0.006.

_Habitat._--North Pacific, Station 253, surface.

{833}2. _Echinaspis diadema_, n. sp.

Radial spines thin and long, quadrangular, prismatic; outer half two to
three times as long as the inner. Four apophyses of each spine simply
forked (or partly with bifid fork-branches); each spine with eight to
twelve sutural condyles. Meshes of the shell fifteen to twenty-five times
as broad as the bars. Each condyle bears a simple bristle-shaped by-spine,
nearly as long as the radius.

_Dimensions._--Diameter of the shell 0.11; breadth of the spines and bars
0.002.

_Habitat._--South Atlantic, Station 333, surface.

3. _Echinaspis echinoides_, n. sp. (Pl. 137, figs. 7,8).

Radial spines roundish, in the outer half conical, and two to three times
as broad as in the inner half. Four apophyses of each spine doubly forked
or dichotomously (more or less irregularly) branched; each spine with
sixteen to twenty-four sutural condyles. Meshes of the shell four to eight
times as broad as the bars. Each condyle bears a zigzag by-spine, one-third
as long as the radius.

_Dimensions._--Diameter of the shell 0.15; breadth of the spines 0.004 to
0.01, of the bars 0.002.

_Habitat._--Central Pacific, Station 266, surface.

Genus 360. _Zonaspis_,[400] n. gen.

_Definition._--#Dorataspida# with four plates, each of which is perforated
by four crossed aspinal pores; these four plates are formed by the united
branches of the other spines (eight polar and eight tropical) form no
lattice-plates by union. Each condyle bears a by-spine.

The genus _Zonaspis_ differs from all other Dorataspida in the peculiar
formation of the four equatorial spines, which form by union of the
recurved branches of their apophyses four perforated plates (each with four
crossed pores), whilst the branches of the sixteen other plates do not
unite to form plates, but simply meet the branches of the neighboring
spines.

1. _Zonaspis fragilis_, n. sp.

Radial spines very thin and long, cylindrical. Four meshes of each
equatorial plate pentagonal, ten to twelve times as broad as the bars.
By-spines zig-zag, nearly as long as the radius.

_Dimensions._--Diameter of the shell 0.16, of the parmal pores 0.018.

_Habitat._--South Atlantic, Station 332, surface.

{834}2. _Zonaspis cingulata_, n. sp. (Pl. 134, figs. 3, 4).

Radial spines cylindrical in the inner half, with thickened pyramidal bases
(fig. 3), conical in the outer half; both halves of equal length. The four
meshes of each equatorial plate egg-shaped, four to six times as broad as
the bars. By-spines zigzag, half as long as the radius.

_Dimensions._--Diameter of the shell 0.15, of the parmal meshes 0.015.

_Habitat._--Tropical Pacific (east coast off Philippines), Station 275,
surface.

3. _Zonaspis æquatorialis_, n. sp. (Pl. 135, fig. 5).

Radial spines stout, cylindrical in the shorter inner half, conical in the
longer outer half. The four meshes of each equatorial plate circular, only
twice as broad as the bars. By-spines zigzag, nearly as long as the radius.

_Dimensions._--Diameter of the shell 0.11, of the parmal meshes 0.008.

_Habitat._--Equatorial Atlantic, Station 347, surface.

Genus 361. _Dodecaspis_,[401] n. gen.

_Definition._--#Dorataspida# with twelve plates, each of which is
perforated by four crossed aspinal pores; these twelve plates are formed by
the united branches of the apophyses of four equatorial and eight polar
spines. The branches of the apophyses of the eight tropical spines form no
lattice-plates by union. Each condyle bears a by-spine.

The genus _Dodecaspis_ differs from all other Dorataspida in the peculiar
composition of its shell; twelve spines (four equatorial and eight polar
spines) form by union of the branches of their apophyses twelve plates,
each of which exhibits four crossed pores, whilst the eight other
(tropical) spines form no perforated plates, but simply unite with the
neighbouring spines by meeting branches. Some irregularity is often to be
found in this genus; instead of four pores in each plate there may be two
closed, the other two which are opposite being open. Several times I
observed a peculiar "hemihedral" variety: only six spines of one meridian
plane (the two equatorial and the four polar spines of the "hydrotomical"
plane) exhibited complete shields, whilst the branches of the other
fourteen plates remained open. This "hemihedral" form may perhaps represent
a peculiar genus, _Hemiaspis_.

1. _Dodecaspis tricinata_, n. sp. (Pl. 134, fig. 1).

Radial spines thin and long, cylindrical. The four meshes of each
equatorial and each polar plate of equal size, pentagonal, about eight
times as broad as the bars. By-spines zigzag, nearly as long as the radius.
In this species the hemihedral variety, _Hemiaspis_, often occurs.

_Dimensions._--Diameter of the shell 0.15, of the parmal pores 0.016.

_Habitat._--South-east Pacific (west coast of Patagonia), Station 302,
surface.

{835}2. _Dodecaspis trizonia_, n. sp.

Radial spines thin, cylindrical in the longer inner half, conical in the
shorter outer half. The four meshes of each equatorial and each polar plate
of equal size, roundish or nearly circular, five to six times as broad as
the bars. By-spines straight, denticulated, scarcely half as long as the
radius.

_Dimensions._--Diameter of the shell 0.12, of the parmal pores 0.012.

_Habitat._--South-west Pacific (east coast of New Zealand), Station 169,
surface.

B. Tribe II. #Lychnaspida#, Haeckel, 1881, Prodromus, p. 467.

_Definition._--#Dorataspida# with twenty perforated plates or fenestrated
shields (each plate at least with four pores), produced by union of the
branches of the four crossed apophyses, which arise, opposite in pairs,
from each radial spine. The spherical shell is composed of the twenty
plates united by sutures (rarely by concrescence).

Genus 362. _Tessaraspis_,[402] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
eighty aspinal pores (four crossed pores in each plate). Surface smooth,
without by-spines.

The genus _Tessaraspis_ introduces the series of Lychnaspida, which
comprise all those Dorataspida in which the shell is composed of twenty
plates, each of which is perforated by four primary aspinal pores. In
_Tessaraspis_ and _Lychnaspis_ each plate exhibits only these four primary
pores, whilst in _Icosaspis_ and _Hylaspis_ they become surrounded by a
circle of secondary or coronal pores. If in _Stauraspis_, the common
ancestral form of the Tessaraspida, the four crossed apophyses of each
single radial spine became recurved and united together, we should have the
typical plate of _Tessaraspis_, in which the piercing radial spine is
surrounded by four crossed pores of equal size. The number of sutural
pores, between the neighbouring plates, is variable; usually each plate is
surrounded by a circle of eight to twelve sutural pores. The sutures
between the meeting condyles of the apophyses usually remain open; but in
some species they become obliterated (subgenus _Tessaraspidium_).

Subgenus 1. _Tessarasparium_, Haeckel.

_Definition._--Condyles of the neighbouring plates connected by permanent
open sutures; therefore the whole shell is composed of twenty separated
pieces of acanthin.

{836}1. _Tessaraspis arachnoides_, n. sp. (Pl. 136, fig. 1).

Parmal pores pentagonal, ten to twenty times as broad as the thin
thread-like bars, on an average of about the same size as the irregular
sutural meshes; the majority of the latter are either triangular or
hexagonal. Radial spines very thin and long, cylindrical, their outer part
two to four times as long as the inner. As the insertion of the spines is
on the highest point of the plates, the shell becomes polyhedral
(dodecahedral?).

_Dimensions._--Diameter of the shell 0.15 to 0.17, of the parmal meshes
0.02 to 0.025, sutural meshes 0.01 to 0.03, bars 0.002.

_Habitat._--Equatorial Atlantic, Station 347, surface.

2. _Tessaraspis pentagonalis_, n. sp.

Parmal meshes pentagonal, three to four times as broad as the thick bars,
and on an average smaller than the irregular polygonal sutural meshes.
Radial spines stout, quadrangular; their outer pyramidal part shorter than
the inner prismatic part.

_Dimensions._--Diameter of the shell 0.12, of the parmal pores 0.01 to
0.012, bars 0.003.

_Habitat._--Central Pacific, Station 269, surface.

3. _Tessaraspis tetragonalis_, n. sp.

Parmal meshes tetragonal, or nearly square, six to eight times as broad as
the thin bars, and on an average larger than the irregular polygonal
sutural meshes. Radial spines thin, quadrangular, prismatic, their outer
part longer than their inner.

_Dimensions._--Diameter of the shell 0.16, of the parmal pores 0.012 to
0.016, bars 0.002.

_Habitat._--South Pacific, Station 288, surface.

4. _Tessaraspis hexagonalis_, n. sp.

Parmal meshes hexagonal, ten to twelve times as broad as the thin bars, and
on an average larger than the polygonal sutural meshes (the two proximal
sides of each hexagonal parmal mesh two to three times as long as the four
distal sides). Radial spines cylindrical, thin; their outer part longer
than the inner.

_Dimensions._--Diameter of the shell 0.15, of the parmal pores 0.01 to
0.015, bars 0.001.

_Habitat._--North Pacific, Station 254, surface.

5. _Tessaraspis trigonalis_, n. sp.

Parmal meshes triangular, three to four times as broad as the thick bars,
and on an average smaller than the irregular sutural meshes (in each plate
all four parmal meshes of the same size, formed like an isosceles triangle,
the distal base of which is somewhat curved, and convex towards {837}the
periphery, whilst the two proximal sides are straight). Radial spines
thick, rounded; their outer conical part shorter than the inner cylindrical
part.

_Dimensions._--Diameter of the shell 0.1, of the parmal pores 0.01, of the
sutural pores 0.01 to 0.02.

_Habitat._--Tropical Atlantic (near Ascension), Station 345, depth 2010
fathoms.

6. _Tessaraspis circularis_, n. sp.

Parmal meshes circular, all eighty of equal size, quite regular, five to
six times as broad as the thick bars, on an average larger than the
roundish irregular sutural meshes. Radial spines quadrangular, prismatic,
the outer part longer than the inner.

_Dimensions._--Diameter of the shell 0.18, parmal pores 0.015 to 0.018,
bars 0.003.

_Habitat._--South-east Pacific, Station 300, surface.

7. _Tessaraspis micropora_, n. sp.

Parmal meshes circular, very small, all eighty of the same size, regular,
scarcely as broad as the separating thick bars, and much smaller than the
large irregular sutural meshes. Radial spines rounded, their outer conical
part about as long as the inner cylindrical part.

_Dimensions._--Diameter of the shell 0.08, of the parmal pores 0.003, of
the sutural pores 0.005 to 0.015, bars 0.004.

_Habitat._--Western Tropical Pacific, Station 225, surface.

8. _Tessaraspis diodon_, Haeckel.

_Dorataspis diodon_, Haeckel, 1862, Monogr. d. Radiol., p. 417, Taf.
xxii. figs. 1-5.

Parmal meshes roundish, for the most part nearly circular, three to four
times as broad as the bars, and on an average smaller than the roundish
sutural meshes. Radial spines in the inner part cylindrical; the outer part
very short (only one-third or one-fourth of the radius), divided completely
into two parallel conical pointed teeth by a deep incision.

_Dimensions._--Diameter of the shell 0.12, of the parmal pores 0.01 to
0.012, of the sutural pores 0.01 to 0.03, bars 0.004.

_Habitat._--Mediterranean (Messina), surface.

9. _Tessaraspis quadriforis_, n. sp.

Parmal meshes irregular, roundish, of nearly equal size, four to six times
as broad as the bars, and on an average of the same size as the roundish
sutural meshes. Radial spines quadrangular, prismatic in the inner part,
which is somewhat longer than the conical outer part.

_Dimensions._--Diameter of the shell 0.16, of the pores 0.012 to 0.018,
bars 0.003.

_Habitat._--North Atlantic, Canary and Cape Verde Islands, Station 353,
surface.

{838}10. _Tessaraspis irregularis_, n. sp.

Parmal meshes irregular, roundish, of unequal size, three to six times as
broad as the bars, and on an average larger than the irregular sutural
meshes. Radial spines cylindrical, the outer part longer than the inner
part.

_Dimensions._--Diameter of the shell 0.13, parmal pores 0.006 to 0.012,
sutural pores 0.005 to 0.01; bars 0.002.

_Habitat._--Central Pacific, Station 270, surface.

Subgenus 2. _Tessaraspidium_, Haeckel.

_Definition._--Condyles of the neighbouring plates grown together and
sutures obliterated; therefore the whole shell forms a single piece of
acanthin.

11. _Tessaraspis quadrata_, n. sp.

_Tessaraspidium quadratum_, Haeckel, 1882, Manuscript.

Parmal meshes square, four times as broad as the thin bars, on an average
of the same size as the polygonal sutural meshes. Radial spines stout,
tetrapterous, prismatic, in the outer part longer than in the inner.
Sutures of the shell completely obliterated, therefore the whole shell
forms one piece.

_Dimensions._--Diameter of the shell 0.16, of the parmal pores 0.012,
sutural pores 0.01 to 0.015.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

12. _Tessaraspis rotunda_, n. sp.

Parmal meshes circular, of equal size, twice as broad as the thick bars, on
an average smaller than the roundish sutural meshes. Radial spines
cylindrical in the inner part, which is somewhat longer than the outer
conical part. Sutures of the shell completely obliterated, therefore the
whole shell forms one piece.

_Dimensions._--Diameter of the shell 0.12, parmal pores 0.008, sutural
pores 0.005 to 0.015.

_Habitat._--North Atlantic, Station 354, surface.

13. _Tessaraspis concreta_, n. sp. (Pl. 136 fig. 5).

Parmal meshes irregular, polygonal or roundish, six to eight times as broad
as the narrow and high bars, much larger than the irregular sutural meshes.
Radial spines leaf-shaped, compressed, two-edged, pointed at both ends;
inner and outer part nearly of equal size. Shell very thick walled; meshes
therefore funnel-shaped; sutures completely obliterated, therefore the
whole shell forms a single piece.

_Dimensions._--Diameter of the shell 0.09, parmal pores 0.02, sutural pores
0.01.

_Habitat._--Central Pacific, Station 274, depth 2750 fathoms.

{839}Genus 363. _Lychnaspis_,[403] Haeckel, 1862, Prodromus, p. 468.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
eighty aspinal pores (four crossed pores in each plate). Surface covered
with numerous by-spines.

The genus _Lychnaspis_, the largest and most common of all Dorataspida,
exhibits the same structure of the shell as its ancestral form
_Tessaraspis_, and differs from it only in the development of by-spines on
the sutural condyles. Many species of this genus are very widely
distributed, and appear in large numbers, and some of them are amongst the
most graceful and elegant of the Radiolaria.

Subgenus 1. _Lychnasparium_, Haeckel.

_Definition._--Condyles of the neighbouring plates connected by permanent
open sutures; therefore the whole shell is composed of twenty separate
pieces of acanthin.

1. _Lychnaspis giltschii_, n. sp. (Pl. 135, fig. 3).

Parmal meshes pentagonal, about ten or twelve times as broad as the thin
bars, on an average of the same size as the irregular polygonal sutural
meshes. By-spines (two hundred to three hundred) very delicate, half as
long as the radius, barbed, and zigzag. Radial main-spines very thin and
long, straight, cylindrical; their outer part longer than the inner part.

_Dimensions._--Diameter of the shell 0.2, of the parmal pores 0.02, sutural
pores 0.01 to 0.03, bars 0.002.

_Habitat._--Tropical Atlantic, Stations 338 to 348, surface.

2. _Lychnaspis capillaris_, n. sp.

Parmal meshes pentagonal, twenty to thirty times as broad as the very thin
thread-like bars, on an average of the same size as the irregular polygonal
sutural meshes. By-spines (two hundred to two hundred and fifty) very
delicate, zigzag, with very small denticles, one-third as long as the
radius. Radial main-spines very thin and long, cylindrical, more or less
undulated.

_Dimensions._--Diameter of the shell 0.25, parmal pores 0.025, sutural
pores 0.02 to 0.04, bars 0.001.

_Habitat._--North Pacific, Station 250, surface.

3. _Lychnaspis maxima_, n. sp.

Parmal meshes pentagonal, six to eight times as broad as the thick bars,
for the most part larger than the irregular sutural meshes. By-spines
(four hundred to five hundred) nearly as {840}long as the radius,
zigzag, with short denticles. Radial spines very long and stout,
quadrangularly-prismatic, with four smooth edges.

_Dimensions._--Diameter of the shell 0.3, parmal pores 0.03, sutural pores
0.01 to 0.025, bars 0.004.

_Habitat._--Central Pacific, Stations 271 to 274, surface.

4. _Lychnaspis serrata_, n. sp.

Parmal meshes pentagonal, three to four times as broad as the thick bars,
smaller than the irregular sutural meshes. By-spines (two hundred to three
hundred) scarcely one-third as long as the radius, zigzag, with strong
denticles. Radial spines very long, stout, quadrangularly-prismatic, with
four serrated edges.

_Dimensions._--Diameter of the shell 0.22, parmal pores 0.015, sutural
pores 0.012 to 0.03, bars 0.004.

_Habitat._--South Atlantic, Station 330, surface.

5. _Lychnaspis wagenschieberi_, n. sp.

Parmal meshes tetragonal, nearly rhombic, three to four times as broad as
the thick bars, smaller than the large irregular sutural meshes. By-spines
(about two hundred) large, as long as the radius, with long recurved
denticles, very zigzag. Radial main-spines very long, four-sided prismatic.
This species differs by the form and size of the spines, and by the square
form of the parmal pores from the similar _Lychnaspis polyancistra_, of
which the late excellent engraver Wagenschieber, of Berlin, has given such
a beautiful figure in my monograph (Taf. xxi. fig. 8).

_Dimensions._--Diameter of the shell 0.16, parmal pores 0.014, sutural
pores 0.015 to 0.02, bars 0.004.

_Habitat._--Central Pacific, Station 266, surface.

6. _Lychnaspis polyancistra_, Haeckel.

_Dorataspis polyancistra_, Haeckel, 1862, Monogr. d. Radiol., p. 418,
Taf. xxi. figs. 7-9.

Parmal meshes roundish, tetragonal, or nearly square, two to three times as
broad as the thick bars, smaller than the irregular sutural meshes.
By-spines (about two hundred) thin, half as long as the radius, zigzag,
with blunt denticles. Radial main-spines stout, in the inner half
cylindrical, in the outer half (of the same length) conical, pointed.

_Dimensions._--Diameter of the shell 0.12, parmal pores 0.012, sutural
pores 0.01 to 0.04, bars 0.004.

_Habitat._--Mediterranean (Messina), surface.

{841}7. _Lychnaspis rottenburgii_, n. sp. (Pl. 135, fig. 4).

Parmal meshes roundish, about twice as broad as the thick bars, and for the
most part smaller than the irregular sutural meshes. By-spines (about two
hundred to two hundred and fifty) thin, zigzag, half as long as the radius.
Radial main-spines very strong, cylindrical in the inner half, in the outer
half much thicker and conical, pointed. I call this species in honour of my
learned friend, the great patron of zoological studies, Mr. Paul
Rottenburg, of Glasgow.

_Dimensions._--Diameter of the shell 0.15, parmal pores 0.012, sutural
pores 0.01 to 0.03, bars 0.006.

_Habitat._--Central Pacific, Station 270, depth 2925 fathoms.

8. _Lychnaspis undulata_, n. sp. (Pl. 135, fig. 2).

Parmal meshes circular, twice as broad as the thick bars, on an average of
the same size as the irregular sutural meshes. By-spines (one hundred and
fifty to two hundred) very thin, barbed and zigzag, as long as the radius.
Radial main-spines cylindrical; their outer pointed part longer than the
inner part.

_Dimensions._--Diameter of the shell 0.1, parmal pores 0.01, sutural pores
0.01, bars 0.005.

_Habitat._--Tropical Atlantic, Station 343, surface.

9. _Lychnaspis longissima_, n. sp. (Pl. 134, fig. 6).

Parmal meshes circular, very small, of the same breadth as the thick bars,
much smaller than the irregular sutural meshes. By-spines (one hundred to
one hundred and ten) very long and thin, zigzag, twice as long as the
diameter of the shell. Radial main-spines very long and strong, cylindrical
(at the base quadrangular, pyramidal), four to six times as long as the
diameter of the shell, and one fourth as thick as its radius.

_Dimensions._--Diameter of the shell 0.08, parmal pores 0.002, sutural
pores 0.01 to 0.015, bars 0.003; length of the radial spines 0.3 to 0.5.

_Habitat._--Tropical Pacific (Philippines), Stations 200 to 215, surface.

10. _Lychnaspis minima_, n. sp. (Pl. 134, figs. 2, 7, 8).

Parmal meshes circular, very small, only half as broad as the thick bars,
much smaller than the irregular sutural meshes. By-spines (one hundred to
one hundred and ten) half as long as the radius, zigzag. Radial main-spines
thick, in the inner part cylindrical, in the outer shorter part conical, of
very variable size.

_Dimensions._--Diameter of the shell 0.05, parmal pores 0.002, sutural
pores 0.006 to 0.012, bars 0.004.

_Habitat._--Antarctic Ocean, Station 154, surface.

{842}Subgenus 2. _Lychnaspidium_, Haeckel.

_Definition._--Condyles of the neighbouring plates grown together and
sutures obliterated, therefore the whole shell forms a single piece of
acanthin.

11. _Lychnaspis echinoides_, Haeckel.

_Haliomma echinoides_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p.36, Taf. v. figs. 3, 4.

_Haliommatidium echinoides_, J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, p. 22.

_Haliommatidium echinoides_, Haeckel, 1862, Monogr. d. Radiol., p. 422.

Parmal meshes pentagonal or somewhat roundish, four times as broad as the
bars, and of about the same size as the polygonal meshes. By-spines (about
two hundred) short, zigzag. Radial spines thin; their outer conical part
shorter than the inner cylindrical part. Sutures perfectly obliterated, but
recognisable by the characteristic pair of divergent by-spines. (Some
recent observations on this species, made during 1880 in Portofino, have
convinced me that the interpretation of it given in my Monograph, 1862,
_loc. cit._, was quite correct.)

_Haliomma ligurinum_, J. Müller (= _Haliommatidium ligurinum_, Haeckel, L.
N. 16, p. 423) seems to be closely allied to the preceding.

_Dimensions._--Diameter of the shell 0.12 to 0.14, parmal pores 0.015,
sutural pores 0.01 to 0.02, bars 0.004.

_Habitat._--Mediterranean, Nice, Saint Tropez (J. Müller); Portofino near
Genoa (Haeckel).

12. _Lychnaspis haliommidium_, n. sp.

_Lychnaspidium haliommidium_, Haeckel, 1882, Manuscript.

Parmal meshes circular, twice as broad as the bars, smaller than the
irregular sutural meshes. By-spines (about two hundred) barbed and zigzag,
as long as the radius. Radial main-spines four-sided; their outer pyramidal
part shorter than the inner prismatic part. Sutures perfectly obliterated.

_Dimensions._--Diameter of the shell 0.1, parmal pores 0.01, sutural meshes
0.015 to 0.02.

_Habitat._--South Atlantic, Station 335, depth 1425 fathoms.

13. _Lychnaspis rabbeana_, n. sp.

Parmal meshes circular, very small, of the same breadth as the bars, and
much smaller than the irregular sutural meshes. By-spines (about one
hundred) very long and thin, zigzag, about as long as the diameter of the
shell. Radial main-spines cylindrical, thick, twice to three times as long
as the diameter of the shell. Sutures perfectly obliterated, with thickened
condyles. Named in honour of Captain Henrik Rabbe (of Bremen), to whom I am
indebted for many new Indian and Atlantic Radiolaria.

_Dimensions._--Diameter of the shell 0.086, parmal pores 0.002, sutural
pores 0.012, bars 0.002.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

{843}14. _Lychnaspis cataplasta_, n. sp.

Parmal pores very small, circular, half as broad as the bars, and much
smaller than the irregular sutural pores. By-spines zigzag, as long as the
diameter of the shell. Radial main-spines very thin and long,
needle-shaped, cylindrical, five to six times as long as the diameter of
the shell. Sutures perfectly obliterated. (This stunted species is one of
the smallest of the Dorataspida.)

_Dimensions._--Diameter of the shell 0.05, parmal pores 0.0015, sutural
pores 0.01, bars 0.002.

_Habitat._--Antarctic Ocean (off Kerguelen Island), Station 149, surface.

Genus 364. _Icosaspis_,[404] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Dorataspida# with twenty plates, which are perforated by
one hundred and sixty to three hundred or more parmal pores (in each plate
four crossed aspinal pores, and around them four to twelve or more coronal
pores). Surface without by-spines.

The genus _Icosaspis_ and the closely allied _Hylaspis_ differ from all
other Tessaraspida in the increased number of the parmal pores. Whilst this
number in all other genera is eighty (only four crossed pores in each
plate), here it amounts to one hundred and sixty to three hundred or more
(sometimes more than a thousand); in each shield four primary, crossed
"aspinal pores" being surrounded by a circle of four to twelve or more
"coronal pores." The number of sutural pores in these two genera is also
increased.

Subgenus 1. _Icosasparium_, Haeckel.

_Definition._--Condyles of the neighboring plates connected by permanent
open sutures; therefore the whole shell composed of twenty separated pieces
of acanthin.

1. _Icosaspis tabulata_, n. sp. (Pl. 136, fig. 2).

Parmal meshes all of nearly equal size and form, square, four times as
broad as the bars, little larger than the triangular or polygonal sutural
meshes. In each plate fifty to seventy (regularly sixty-four) quadrangular
pores, viz., four primary square aspinal meshes, forming together a regular
square surrounded by two to three coronas of rectangular (not quite
regular) coronal meshes (six to eight in each transverse row). Radial
spines tetrapterous, prismatic, with four thin and broad wings, from which
arise the crossed bars between the four primary pores. Outer part of the
spines longer than the inner. Commonly the condyles of the plates are only
contiguous; sometimes they grow together, and this form approaches
_Icosaspis tetragonopa_.

_Dimensions._--Diameter of the shell 0.25 to 0.3, of the pores 0.02, bars
0.005.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{844}2. _Icosaspis elegans_, n. sp. (Pl. 136, fig. 4; Pl. 134, fig. 9).

_Tessaraspis elegans_, Haeckel, 1882, Manuscript et Atlas.

Parmal meshes of very different size and form; in the centre of each plate
a cross of four primary, pear-shaped "aspinal pores" (the largest of all);
between them four secondary, little smaller, crossed, egg-shaped "angular
pores"; around this rosette of eight larger meshes an inner complete circle
of sixteen to twenty polygonal coronal pores, and an outer incomplete
circle of thirty to forty very small marginal pores. The latter are smaller
than the irregular sutural meshes, which are constricted in the middle,
about forty to sixty around each plate. Radial spines thin, cylindrical, or
a little compressed; their outer part longer than the inner. In this
elegant and very common species the condyles usually remain separated by
sutures; but sometimes the latter become obliterated, and the whole shell
then forms a single piece, _Icosaspidium elegans_.

_Dimensions._--Diameter of the shell 0.2 to 0.3, commonly 0.25; larger
parmal pores 0.02, smaller 0.001 to 0.003; sutural pores 0.005 to 0.015;
bars 0.003.

_Habitat._--Tropical and Subtropical Atlantic, Canary Islands to Ascension
Island, Stations 340 to 354, surface.

3. _Icosaspis cruciata_, n. sp. (Pl. 134, fig. 10).

Parmal meshes very different; in the centre of each plate a cross of four
primary, nearly oblong, rectangular "aspinal pores"; between these four
secondary, triangular, egg-shaped "angular pores" (the largest of all), and
around this rosette a single circle of twelve to twenty-four small "coronal
pores." The latter are of about the same size as the irregular sutural
meshes, of which there are twenty to thirty around each plate. Radial
spines thin, cylindrical, or a little compressed; their outer part longer
than the inner.

_Dimensions._--Diameter of the shell 0.2 to 0.3, commonly 0.25; larger
parmal pores 0.025, smaller 0.005 to 0.01; bars 0.005.

_Habitat._--Tropical and Subtropical Pacific, Sandwich to Marquesas
Islands, Stations 256 to 274, surface.

4. _Icosaspis ornata_, n. sp.

Parmal meshes very different; in the centre of each plate a cross of four
primary octagonal aspinal pores (the largest of all); between them four
secondary, rhombic angular pores, and around this rosette a circle of
twelve to sixteen smaller, polyhedral coronal pores, which are however
larger than the irregular sutural pores (surrounding each plate to the
number of twenty to thirty). Radial spines thin, quadrangular, prismatic;
the outer part longer than the inner.

_Dimensions._--Diameter of the shell 0.25; larger parmal pores 0.03,
smaller 0.01; sutural pores 0.004 to 0.008; bars 0.006.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

{845}5. _Icosaspis spectabilis_, n. sp.

Parmal meshes very different; in the centre of each plate four very large,
pentagonal aspinal pores, and around these two to three circles of smaller
polygonal coronal pores, which are very numerous, and not larger than the
small sutural pores. Radial spines quadrangular, prismatic, stout, very
long; the outer part two to three times as long as the inner.

_Dimensions._--Diameter of the shell 0.4, of the larger aspinal meshes
0.03, of the outer meshes 0.002 to 0.02, bars 0.005.

_Habitat._--South Atlantic, Station 333, surface.

6. _Icosaspis multiforis_, n. sp.

Parmal meshes very numerous, more than one hundred in each plate; in the
centre four larger pear-shaped pores, and around these four to five circles
of smaller pores, gradually smaller towards the margin of the plate; the
sutural meshes also very small and numerous (more than fifty around each
plate), so that the number of all the pores together amounts to two
thousand or even more. Radial spines thin, cylindrical, very long.

_Dimensions._--Diameter of the shell 0.32; larger aspinal pores 0.02,
smaller 0.002 to 0.01; bars 0.003.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

Subgenus 2. _Icosaspidium_, Haeckel.

_Definition._--Condyles of the neighbouring plates grown together, and
sutures obliterated; therefore the whole shell forms a single piece of
acanthin.

7. _Icosaspis tetragonopa_, Haeckel.

_Haliommatidium tetragonopum_, Haeckel, 1862, Monogr. d. Radiol., p.421,
Taf. xxii. fig. 13.

Parmal meshes all of nearly equal size and form, square, three times as
broad as the bars, little larger than the sutural meshes. In each plate
commonly sixteen equal square meshes, viz., four primary aspinal and twelve
secondary, surrounding the former as a square corona. Radial spines
tetrapterous, stout; the outer pyramidal half somewhat longer than the
inner. This species differs from the similar _Icosaspis tabulata_ (Pl. 136,
fig. 2) in the concrescence of the sutures, the smaller number of pores,
and the form of the stouter spines. The figure in my Monograph, drawn from
a broken fragment, is not quite correct.

_Dimensions._--Diameter of the shell 0.18, pores 0.009, bars 0.003.

_Habitat._--Mediterranean (Messina, Corfu), surface.

8. _Icosaspis icosahedra_, n. sp.

Parmal meshes of different size and form; in the centre of each plate a
cross of four pentagonal, primary aspinal pores, surrounded by a complete
corona of twelve to sixteen polygonal coronal {846}pores and an incomplete
corona of thirty to forty very small marginal pores; the latter are not to
be distinguished from the pores of the obliterated sutures. Radial spines
quadrangular, the outer pyramidal part shorter than the inner prismatic
part. Condyles grown together. As the plates are quite even, the shell
becomes icosahedral.

_Dimensions._--Diameter of the shell 0.16, pores 0.002 to 0.02, bars 0.005.

_Habitat._--North Pacific, Station 241, surface.

9. _Icosaspis icosastaura_, n. sp. (Pl. 136, fig. 3).

_Tessaraspis icosastaura_, Haeckel, 1882, Manuscript et Atlas.

Parmal plates of different size and form; in the centre of each plate a
cross of four larger primary, pyriform aspinal pores; between these four
smaller roundish angular pores, and around this rosette a circle of ten to
twenty (commonly sixteen) coronal pores, little larger than the very small
sutural pores. Radial spines very thin and long, cylindrical or
bristle-shaped. Condyles grown together; no suture visible.

_Dimensions._--Diameter of the shell 0.14; larger pores of the cross 0.01,
smaller pores 0.002 to 0.008; bars 0.002 to 0.004.

_Habitat._--Central Pacific, Station 268, surface.

Genus 365. _Hylaspis_,[405] n. gen.

The genus _Hylaspis_ exhibits the same structure of the shell as the nearly
allied ancestral genus _Icosaspis_, and differs from it only in the
development of by-spines. Some species of these two genera exhibit the
highest degree of complication in the structure of the shell seen among the
Dorataspida.

1. _Hylaspis serrulata_, n. sp. (Pl. 135, fig. 1).

Parmal meshes four hundred to five hundred; in the centre of each plate a
cross of four very large pentagonal or roundish aspinal pores, and around
this a circle of sixteen to twenty much smaller irregular, polygonal,
coronal pores; the latter of about the same size as the sutural pores. On
each condyle one thin zigzag-shaped by-spine, nearly as long as the radius.
Twenty radial spines very long, quadrangular, prismatic; on the inside
thinner and smooth, on the outside thickened, and armed with four rows of
recurved teeth, serrated.

_Dimensions._--Diameter of the shell 0.18, aspinal spines 0.02, other pores
0.002 to 0.01, bars 0.003.

_Habitat._--South Atlantic, Station 326, surface.

{847}2. _Hylaspis coronata_, n. sp.

Parmal meshes five hundred to six hundred; in the centre of each plate a
cross of four long rectangular aspinal pores, between these four larger
egg-shaped angular pores; around this rosette a circle of sixteen to twenty
much smaller, irregular, roundish, coronal pores; the latter about of the
same size as the sutural pores. On each condyle one bearded by-spine about
one-third or one-fourth as long as the radius. Twenty radial spines, very
long, smooth, quadrangular, prismatic.

_Dimensions._--Diameter of the shell 0.25, aspinal pores 0.02, other pores
0.005 to 0.01, bars 0.004.

_Habitat._--Central Pacific, Station 271, surface.

3. _Hylaspis barbata_, n. sp.

Parmal meshes twelve hundred to sixteen hundred; in the centre of each
plate a cross of four large, somewhat oblong, octahedral aspinal pores,
between these four rhombic smaller angular pores; around this rosette an
inner circle of twelve to sixteen larger and an outer circle of fifty to
sixty very small coronal pores; the latter smaller than the sutural pores.
On each condyle one bristle-shaped zigzag by-spine, with recurved thin
hooks, half as long as the radius. Twenty radial spines, very long,
cylindrical, smooth.

_Dimensions._--Diameter of the shell 0.25 to 0.3, aspinal pores 0.022,
other pores 0.002 to 0.015, bars 0.002.

_Habitat._--South Pacific, Station 295, surface.

Family XLI. #PHRACTOPELTIDA#, Haeckel (Pl. 133, figs. 1-6).

_Phractopeltida_, Haeckel, 1881, Prodromus, p. 468.

_Definition._--ACANTHARIA with double spherical lattice-shell, composed of
the branched apophyses of twenty radial spines meeting in its centre, and
disposed according to the Müllerian law of Icosacantha. Central capsule
spherical, enclosing the inner and surrounded by the outer concentric
shell.

The family #Phractopeltida# differs from all other ACANTHARIA in the
development of a double spherical shell, composed of two concentric
lattice-spheres, which are united by twenty radial spines meeting in the
common centre. We could therefore oppose the Phractopeltida as
_Diplophracta_ to all other #Acanthophracta# as _Haplophracta_ (with simple
shell). The former exhibit a relation to the latter, similar to that
exhibited by the Dyosphærida to the simple Monosphærida among the
#Sphæroidea#.

In my Monograph (1862, p. 423) I described only one genus appertaining to
this family, _Aspidomma_. I founded it upon the singular _Phractopelta_,
described by {848}J. Müller as _Haliomma hystrix_. A second species of
_Aspidomma_, the _Acanthometra mucronata_ of J. Müller, was probably an
_Astrolonche_. At that time I placed _Aspidomma_ among the Haliommatida,
led by the erroneous opinion that it might represent a transition-form
between _Dorataspis_ and _Haliomma_. But I afterwards gave up this view, as
I was convinced that there is no true phylogenetic connection between the
acanthinic Dorataspida (_Actipylea_) and the siliceous Haliommatida
(_Peripylea_). Therefore in my Prodromus (1881, p. 468) I placed
_Aspidomma_ among the Dorataspida and changed its name to _Phractopelta_,
to avoid further confusion with the unrelated Ommatida (#Sphæroidea#). It
formed there, with three nearly related genera, the "subfamily
Phractopeltida," which we now advance to the higher rank of a separate
family. (By a typographical mistake the words are printed in the Prodromus
_Phractopelma_ and Phractopelmida, &c., instead of _Phractopelta_ and
Phractopeltida, &c.). The detection of other new species appertaining to
this family, and a closer anatomical investigation of them, has now led to
the distinction of five different genera, characterised by other
differences than were employed in 1881 in the provisional system of the
"Prodromus."

The two concentric spherical lattice-shells of the Phractopeltida,
connected by radial beams, correspond perfectly to those of the
double-shelled Dyosphærida (_Haliomma_, _Diplosphæra_, &c.), and in both
cases we may call the smaller inner the "medullary shell," and the larger
outer the "cortical shell." There is no doubt that the double-shelled
Phractopeltida must be derived phylogenetically from the simple-shelled
Dorataspida (just as we derive the double Dyosphærida from the simple
Monosphærida). But it is not yet possible to decide positively which of the
two shells is the first formed. Probably the small inner or medullary shell
of the Phractopeltida is the first formed, and corresponds to the simple
spherical lattice-shell of the Dorataspida; and the larger outer or
cortical shell of the former is a later new formation, absent in the latter
family. This opinion seems to be confirmed by the genus _Orophaspis_, the
only form among the Dorataspida, in which the radial spines outside the
shell bear free latticed apophyses. If these twenty apophyses grow further
and meet one another, the second or outer shell of _Phractopelta_ may be
formed. But some objections may be raised to this opinion from the peculiar
structure and the very small size of the inner shell; and there is some
possibility that this latter is a secondary later product inside of the
primary cortical shell. The probably phylogenetic series which reveals the
origin of the Phractopeltida is the following:--_Acanthometron_,
_Zygacantha_, _Lithophyllium_, _Phractacantha_, _Doracantha_, _Dorataspis_,
_Orophaspis_, _Phractopelta_.

The twenty radial spines exhibit in all Phractopeltida the same
characteristic position and relation as in all other Icosacantha, and are
constantly arranged according to the Müllerian law in four meridian planes,
their distal ends falling into five parallel zones. Their distinction in
the majority of the Phractopeltida is not difficult, {849}since the spines
of the different zones bear apophyses of different shapes. Sometimes the
four equatorial spines are stouter than the sixteen other spines, and often
the eight tropical spines are somewhat different in form from the eight
polar and from the four equatorial spines. The length of all twenty spines
is commonly equal. Their form is usually more or less compressed, two-edged
(as in _Zygacantha_), more rarely cylindrical (as in _Acanthometron_), or
somewhat quadrangular (but not truly prismatic); therefore the transverse
section of the spines is commonly elliptical or lanceolate, rarely circular
or rhombic, never square; this seems to indicate their origin from
_Zygacantha_. As in all ACANTHARIA, the spines consist of acanthin, not of
silex. Their central ends are either perfectly grown together, and form a
single star of acanthin, or the triangular faces of their small pyramidal
bases are supported one upon another, without true concrescence.

The apophyses of the radial spines, by which the two concentric spherical
shells are formed, seem to be constantly four on each spine, two being
opposite in each shell. The proximal pair of opposite apophyses, forming
the inner or medullary shell, is constantly much smaller than the distal
pair composing the outer or cortical shell (Pl. 133, fig. 5). The proximal
pair corresponds probably to the two primary apophyses of the Diporaspida
(_Phractaspis_, _Dorataspis_, &c.), whilst the distal pair corresponds to
the free apophyses of _Orophaspis_ (Pl. 133, fig. 6). Therefore the
Phractopeltida may be derived phylogenetically from the Diporaspida (not
from the Tessaraspida). In the common ancestral genus of this family,
_Phractopelta_, the free part of the radial spines (outside the outer
shell) is quite simple, without free apophyses; in all other genera of the
family that free part of the spines (either in all twenty spines or only in
some of them) bears a third pair of lateral apophyses; these may be either
simple or branched or even latticed; but the outer apophyses (of the third
rank) remain constantly free, and a third lattice-shell is never formed by
union of their edges (Pl. 133, figs. 2-4).

The inner lattice-shell of the Phractopeltida, or their "medullary shell,"
is constantly very small (commonly 0.03 to 0.05 mm. in diameter, rarely
more). Its structure is difficult to make out; in the unbroken shell it is
concealed by the dense network of the outer shell; in the broken shell it
is commonly destroyed. Usually the pores of the inner shell are very small,
circular or subcircular, scarcely as broad as the small separating bars. In
the majority of Phractopeltida their number seems to be about forty, being
probably the forty primary aspinal pores of the Diporaspida; in some
species this number seems to be exceeded, so that perhaps some sutural
pores may exist between the aspinal pores; but commonly the twenty plates
composing the inner shell (each with two aspinal pores) seems to grow
together perfectly by their meeting edges, so that there are no sutural
pores between them. Evident sutures were not recognisable in the inner
shell of any Phractopeltida.

{850}The outer lattice-shell of the Phractopeltida, or their "cortical
shell," is at least twice as broad, commonly about three times as broad, as
the enclosed inner shell; it is much more varied in composition than the
latter. Like the greater part of the Dorataspida we may distinguish here in
the lattice-work two kinds of pores--parmal pores and sutural pores. The
parmal pores are produced by the union of the meeting branches of the
apophyses of each single spine, and are therefore visible on each isolated
spine; whilst the sutural pores are formed by the meeting branches of the
apophyses of neighbouring spines. The distinction of the parmal and the
sutural pores, easy in most Dorataspida, is difficult in most
Phractopeltida, because the sutures between the meeting branches are
usually very early obliterated. However, the place of the obliterated
suture is often indicated by the thickened condyles of the apophyses on
both sides of the suture. Commonly also the form of the sutural pores is
much more irregular than that of the parmal pores; the former are more or
less constricted in the middle by the intumescence of the sutural condyles,
whilst the latter are more roundish, elliptical, kidney-shaped, or square.
The number of the pores in the outer shell in the typical normal form of
Phractopeltida seems to be the same as in the most species of _Dorataspis_,
_Diporaspis_, &c., between ninety and one hundred, viz., forty parmal pores
and from fifty to sixty sutural pores. However, in many species this number
is increased. Since in all Phractopeltida, each of the twenty plates is
composed only of the meeting branches of two opposite apophyses, we find
originally in each plate only two primary parmal pores or "aspinal pores."
But in some species there occur four, six, or more pores in each plate; in
this case two of them only are aspinal pores, all the others being "coronal
pores." Moreover, in those species which exhibit on the base of each spine
in the outer shell four crossed pores (Pl. 133, fig. 2), there are not four
equivalent aspinal pores (as in the Tessaraspida), but the two opposite are
primary or aspinal pores and the other two (different from them in size and
form) coronal pores. However, the number of coronal pores in the
Phractopeltida is never so large as in many Dorataspida, and the same holds
good also for the increasing number of the irregular sutural pores. In none
of the species observed does the total number of the pores in the outer
shell reach two hundred.

The original mode of development of the apophyses composing the outer shell
seems to be imitated by the free apophyses of the third order, which are
developed from the radial spines outside the outer shell in all
Phractopeltida, with the single exception of the simple ancestral genus
_Phractopelta_. These apophyses of the third rank are also originally
constantly two, opposite to one another (after the type of _Lithophyllium_,
_Dorataspis_, &c.). Commonly they do not remain simple, but become
branched, and by communication of the neighbouring branches small
lattice-plates arise. Originally each of these free lattice-plates has only
two parmal pores, but the number of the parmal pores increases afterwards,
so that we may distinguish two (primary) {851}aspinal pores, and two,
four, or more (secondary) coronal pores. In the majority of species the two
opposite apophyses are first crossed at right angles by a transverse beam,
and the two parallel transverse beams are again crossed by perpendicular
tertiary branches (again parallel to the apophyses). In this case the
network of the free lattice-plates becomes more or less rectangular. But in
other species the ramification of the apophyses assumes more the form of
bifurcation or of irregular branching. As already said, the neighbouring
free lattice-plates of this third order never meet, and therefore a
complete third shell is never formed.

The different genera of Phractopeltida exhibit very remarkable differences
in the development of free apophyses (or lattice-plates of the third
order). Whilst in the numerous species of the ancestral genus
_Phractopelta_ all twenty spines remain simple, without such apophyses,
only in a single observed species (representing the genus _Pantopelta_)
were all twenty spines protected by them. In the three other genera only
one part of the spines bears free apophyses, but not the other part. The
most frequent form is _Dorypelta_ (Pl. 133, fig. 2); here eight spines are
simple (four equatorial and four polar spines of the hydrotomical plane),
whilst twelve spines bear apophyses (eight tropical and four polar spines
of the geotomical plane). In _Octopelta_ the eight tropical spines only
bear apophyses, whilst the twelve other (four equatorial and eight polar)
are simple. In _Stauropelta_ finally the four equatorial spines only are
simple, whilst the sixteen other bear free apophyses (eight tropical and
eight polar spines).

_The Central Capsule_ of the Phractopeltida is constantly spherical, and
enclosed between the two concentric spherical shells; it is therefore
larger than the inner and smaller than the outer shell. Its wall is pierced
by the twenty radial beams connecting the two shells. The shape of the
central capsule and of the enveloping calymma is the same as in the other
#Acanthophracta# and specially in the Dorataspida.

_Synopsis of the Genera of Phractopeltida._

{All twenty spines without
{ apophyses in the free
All twenty spines of the { external part, 366. _Phractopelta_.
same form, {
{All twenty spines with
{ apophyses in the free
{ part, 367. _Pantopelta_.

{Eight tropical spines with
{ apophyses, twelve others
{ (eight polar and four
{ equatorial) simple, 368. _Octopelta_.
{
{Twelve radial spines
Twenty radial spines, { (eight tropical and four
partly without, partly { polar) with apophyses,
with apophyses in the { eight others (four polar
free external part, { and four equatorial)
{ simple, 369. _Dorypelta_.
{
{Sixteen radial spines
{ (eight tropical and
{ eight polar) with
{ apophyses, four
{ equatorial, simple, 370. _Stauropelta_.

{852}Genus 366. _Phractopelta_,[406] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Phractopeltida# with twenty simple radial spines, bearing
no free apophyses outside the outer shell.

The genus _Phractopelta_ is the most simple form among the Phractopeltida,
and may be regarded as the common ancestral form of this family. All twenty
spines are of nearly equal form and size, and bear no free apophyses on
their free part, outside the two concentric shells. _Phractopelta_ may be
derived from _Orophaspis_ by further development of the free apophyses,
which by union of their branches form a second outer shell around the
primary shell of that Dorataspid.

Subgenus 1. _Phractopeltaris_, Haeckel.

_Definition._--Outer shell composed of twenty separated plates, the sutures
of their meeting condyles not being grown together.

1. _Phractopelta dorataspis_, n. sp. (Pl. 133, fig. 1).

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with two elliptical
aspinal pores, which are two to three times as broad as the irregular
sutural pores. Radial spines (in the outer free part) compressed, linear,
twice as long as the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.11, of the inner 0.05.

_Habitat._--North Pacific (off Japan), Station 239, surface.

2. _Phractopelta dyadopora_, n. sp.

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with two kidney-shaped
aspinal pores, which are three to four times as broad as the irregular
sutural pores. Radial spines conical, about as long as the radius of the
shell.

_Dimensions._--Diameter of the outer shell 0.08, of the inner 0.04.

_Habitat._--North Pacific, Station 256, surface.

3. _Phractopelta diporaspis_, n. sp.

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with two quadrangular
aspinal pores, which are four to five times {853}as broad as the irregular
sutural pores. Radial spines compressed, sword-shaped, about as long as the
diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.09, of the inner 0.04.

_Habitat._--South Atlantic, Station 332, surface.

4. _Phractopelta tessaraspis_, n. sp.

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with four crossed,
quadrangular, aspinal pores, the two opposite of which are much larger than
the two others. Sutural pores small, roundish. Radial spines compressed,
linear, about twice as long as the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.084, of the inner 0.032.

_Habitat._--North Pacific (off Japan), Station 238, surface.

5. _Phractopelta tetradopora_, n. sp.

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with four crossed
quadrangular (or nearly circular) aspinal pores, all of nearly the same
size. Sutural pores polygonal or roundish. Radial spines cylindrical, two
to three times as long as the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.11, of the inner 0.045.

_Habitat._--North Pacific, Station 253, surface.

6. _Phractopelta hexadopora_, n. sp.

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with six aspinal pores,
the two opposite of which are much larger than the four others. Sutural
pores small, roundish. Radial spines compressed, two-edged, larger than the
diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.04.

_Habitat._--Central Pacific, Station 272, surface.

7. _Phractopelta octadopora_, n. sp.

Outer shell composed of twenty plates, the meeting condyles of which are
separated by permanent sutures. Each plate commonly with eight aspinal
pores, the four crossed being larger than the four others alternating with
them. Sutural pores irregular. Radial spines conical, shorter than the
diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.04.

_Habitat._--Indian Ocean (Cocos Islands, surface), Rabbe.

{854}Subgenus 2. _Phractopeltidium_, Haeckel.

_Definition._--Outer shell composed of twenty united plates, the sutures of
their meeting condyles being grown together.

8. _Phractopelta aspidomma_, n. sp.

Outer shell composed of twenty united plates, the meeting condyles of which
are grown together. On the base of each radial spine (where its outer free
part arises from the surface of the outer shell), two large elliptical
aspinal pores, two to four times as large as the other roundish pores.
Radial spines compressed, sword-shaped, about as long as the radius of the
shell.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.05.

_Habitat._--Equatorial Atlantic, Station 347, surface.

9. _Phractopelta haliomma_, n. sp.

Outer shell composed of twenty united plates, the meeting condyles of which
are grown together. On the base of each radial spine two large
kidney-shaped aspinal pores, little larger than the other irregular pores.
Radial spines compressed, linear, two-edged, much longer than the diameter
of the shell.

_Dimensions._--Diameter of the outer shell 0.09, of the inner 0.03.

_Habitat._--South Pacific, Station 295, surface.

10. _Phractopelta tessaromma_, Haeckel.

Outer shell composed of twenty united plates, the meeting condyles of which
are grown together. On the base of each radial spine four crossed
egg-shaped aspinal pores, two opposite of which are much larger than the
two others. Sutural pores irregular. Radial spines cylindrical or little
compressed, thin, longer than the diameter of the shell.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.04.

_Habitat._--Central Pacific, Station 266, surface.

11. _Phractopelta hystrix_, Haeckel.

_Haliomma hystrix_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 37, Taf. v. figs. 1, 2.

_Aspidomma hystrix_, Haeckel, 1862, Monogr. d. Radiol., p. 424.

Outer shell composed of twenty united plates, the meeting condyles of which
are grown together. On the base of each radial spine four crossed circular
aspinal spines of equal size. Sutural pores roundish, of about the same
size. Radial spines conical, about as long as the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.07, of the inner 0.024.

_Habitat._--Mediterranean (Nice), J. Müller, surface.

{855}Genus 367. _Pantopelta_,[407] n. gen.

_Definition._--#Phractopeltida# with twenty radial spines, each of which is
protected by two free external apophyses.

The genus _Pantopelta_ differs from all other Phractopeltida in the
development of free protecting apophyses on all twenty radial spines. Only
a single specimen of this rare form was seen, and in this all twenty spines
exhibited no marked differences in form and size, but were more or less
irregularly developed.

1. _Pantopelta icosaspis_, n. sp. (Pl. 133, fig. 4).

Apophyses of all twenty radial spines with forked anastomosing branches,
forming a more or less irregular cup-like fenestrated shield, pierced by a
variable number of pores. Distance of the apophyses from the outer shell
equal to half its radius. Pores of the outer shell irregular, roundish,
four to six times as broad as those of the inner shell. Radial spines
compressed, sword-shaped, longer than the radius of the shell.

_Dimensions._--Diameter of the outer shell 0.08, of the inner 0.03.

_Habitat._--Antarctic Ocean, Station 157, depth 1950 fathoms.

Genus 368. _Octopelta_,[408] n. gen.

_Definition._--#Phractopeltida# with twelve simple spines (four equatorial
and eight polar), and with eight tropical spines protected by external free
apophyses.

The genus _Octopelta_ differs from the other Phractopeltida in the
possession of eight shields or pairs of free apophyses outside the outer
lattice-shell; the spines bearing these apophyses are the eight tropical
spines. The twelve other spines (eight polar and four equatorial) are quite
simple, without apophyses.

1. _Octopelta cultella_, n. sp.

Apophyses of the eight tropical spines simple, compressed, knife-shaped,
about as long as their distance from the outer shell. On the base of each
spine (in the network of the outer shell) two orthogonal aspinal pores, two
to three times as long as the other polygonal pores.

_Dimensions._--Diameter of the outer shell 0.08, of the inner 0.03.

_Habitat._--Tropical Atlantic, Station 338, surface.

{856}2. _Octopelta furcella_, n. sp.

Apophyses of the eight tropical spines forked, each with two parallel
simple fork-branches, about as long as their distance from the outer shell.
On the base of each spine (in the outer shell) two kidney-shaped aspinal
pores, about twice as broad as the other irregular pores.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.04.

_Habitat._--South Atlantic (off Tristan d'Acunha), Station 332, surface.

3. _Octopelta scutella_, n. sp. (Pl. 133, fig. 5).

Apophyses of the eight tropical spines crossed by a transverse beam, which
is again crossed by two perpendicular branches; by union of these branches
each tropical spine forms a square shield with four crossed pores and
twelve to sixteen marginal spikes. On the base of each spine (in the
surface of the outer shell) four crossed aspinal pores which correspond to
those of the free shields; these are somewhat larger than the other pores.

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.06.

_Habitat._--Tropical Atlantic, Station 348, surface.

Genus 369. _Dorypelta_,[409] Haeckel, Prodromus, p. 369.

_Definition._--#Phractopeltida# with eight simple spines (four equatorial
and four hydrotomical polar spines) and with twelve spines protected by
external free apophyses (eight tropical and four geotomical polar spines).

The genus _Dorypelta_, the most common of all Phractopeltida which are
protected by free apophyses, exhibits a very peculiar differentiation of
its twenty radial spines. There are constantly eight simple spines and
twelve spines with apophyses. The eight simple spines are the four
equatorial spines and four polar spines placed in the hydrotomical median
plane. The four other polar spines (placed in the geotomical meridian
plane) and the eight tropical spines are protected by two opposite
apophyses, which are now simple, now branched or shield-shaped. A similar
differentiation occurs in no other Acanthophractida.

Subgenus 1. _Dorypeltarium_, Haeckel.

_Definition._--Free apophyses of the radial spines simple, not branched.

{857}1. _Dorypelta stauroptera_, n. sp.

Apophyses of the long compressed radial spines simple, conical, about as
long as their distance from the shell. At the base of each spine (in the
outer shell) two elliptical aspinal pores, two to three times as large as
the other pores.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.044.

_Habitat._--North Pacific, Station 238, surface.

2. _Dorypelta gladiata_, n. sp.

Apophyses of the long cylindrical radial spines simple, triangular, about
half as long as their distance from the shell. At the base of each spine
(in the outer shell) four crossed aspinal pores, the two opposite of which
are much larger than the other two pores.

_Dimensions._--Diameter of the outer shell 0.09, of the inner 0.04.

_Habitat._--Central Pacific, Station 266, surface.

Subgenus 2. _Dorypeltidium_, Haeckel.

_Definition._--Free apophyses of the radial spines branched, with free (not
anastomosing) branches.

3. _Dorypelta furcata_, n. sp.

Apophyses of the long compressed spines simply forked, each with two simple
parallel fork-branches. At the base of each spine two elliptical aspinal
pores, much larger than the other roundish pores.

_Dimensions._--Diameter of the outer shell 0.08, of the inner 0.04.

_Habitat._--Central Pacific, Station 274, surface.

4. _Dorypelta tetrodon_, n. sp.

Apophyses of the long compressed spines crossed by a transverse beam, which
bears four simple parallel fork-branches or spikes (perpendicular to the
beam), two larger medial and two smaller lateral. At the base of each spine
four crossed aspinal pores of nearly equal size, little different from the
other pores.

_Dimensions._--Diameter of the outer shell 0.09, of the inner 0.035.

_Habitat._--South Pacific, Station 284, surface.

5. _Dorypelta ramosa_, n. sp.

Apophyses of the long cylindrical spines crossed by a transverse beam,
which bears four to six irregularly branched spikes or fork-branches, the
medial of which are larger than the lateral. At {858}the base of each spine
four crossed aspinal pores, two opposite of which are much larger than the
two other pores.

_Dimensions._--Diameter of the outer shell 0.09, of the inner 0.04.

_Habitat._--South Pacific, Station 302, surface.

Subgenus 3. _Dorypeltonium_, Haeckel.

_Definition._--Free apophyses of the radial spines branched, and forming
perforated shields by union of their anastomosing branches.

6. _Dorypelta lithoptera_, n. sp.

Apophyses of the long compressed radial spines with anastomosing branches;
each spine bearing two free separated shields (each with two roundish
aspinal pores). At the base of each spine (in the network of the outer
shell) two elliptical aspinal pores, larger than the other pores.

_Dimensions._--Diameter of the outer shell 0.09, of the inner 0.03.

_Habitat._--North Pacific, Station 239, surface.

7. _Dorypelta tessaraspis_, n. sp. (Pl. 133, fig. 2).

Apophyses of the long cylindrical spines with anastomosing branches; each
spine bearing a single free shield with four crossed aspinal pores of equal
size (on the margin of the shield twelve to sixteen spikes). At the base of
each spine (in the outer shell) four crossed aspinal pores, about as large
as the other roundish pores.

_Dimensions._--Diameter of the outer shell 0.11, of the inner 0.05.

_Habitat._--Central Pacific, Station 263, surface.

8. _Dorypelta dodecaspis_, n. sp.

Apophyses of the long two-edged spines with anastomosing branches; each
spine bearing a single free shield with six to eight parmal pores (four
crossed aspinal pores alternating with four outer coronal pores), on the
margin of each shield twelve to sixteen spikes. At the base of each spine
(in the outer shell) four crossed aspinal pores, about as large as the
other irregular pores.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.04.

_Habitat._--North Pacific (off Japan), Station 235, surface.

Genus 370. _Stauropelta_,[410] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Phractopeltida# with four simple equatorial spines and
sixteen spines protected by external free apophyses (eight tropical and
eight polar spines).

{859}The genus _Stauropelta_ is distinguished from the other Phractopeltida
by the possession of sixteen pairs of free apophyses (on the eight tropical
and the eight polar spines); only the other four equatorial spines remain
simple, without apophyses, and form a simple cross in the equatorial plane.

1. _Stauropelta cruciata_, n. sp. (Pl. 133, fig. 3).

Apophyses of the long polar and tropical spines crossed by a transverse
beam, which is again crossed by two perpendicular branches parallel to the
apophyses; branches with free ends, not united by concrescence. At the base
of each spine (in the outer shell) two large elliptical aspinal pores,
larger than the other irregular pores. Four equatorial spines simple,
without apophyses, of the same size as the other sixteen spines.

_Dimensions._--Diameter of the outer shell 0.12, of the inner 0.05.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

2. _Stauropelta stauropora_, n. sp.

Apophyses of the long polar and tropical spines crossed by a transverse
beam, which is again crossed by two perpendicular branches parallel to the
apophyses; these branches are united by concrescence, and form a square
shield with four crossed pores, the centre of which is pierced by the
spine. At the base of each spine (in the outer shell) four crossed aspinal
pores, corresponding to those of the shield, of about the same size as the
other roundish pores. Four equatorial spines, simple, without apophyses,
somewhat larger than the other sixteen spines.

_Dimensions._--Diameter of the outer shell 0.1, of the inner 0.04.

_Habitat._--Indian Ocean, Sunda-Archipelago, Singapore, Trebing, surface.

----

Suborder II. PRUNOPHRACTA.

_Definition._--Shell ellipsoidal, lenticular or diploconical, with radial
beams of different size.

Family XLII. #BELONASPIDA#, n. fam. (Pl. 136, figs. 6-9; Pl. 139, figs. 8,
9).

_Definition._--ACANTHARIA with a simple ellipsoidal lattice-shell, composed
of the branched apophyses of twenty radial spines meeting in its centre and
disposed according to the Müllerian law of Icosacantha. Two opposite
equatorial spines larger than the two others. Central capsule ellipsoidal,
enclosed in the fenestrated shell.

The family #Belonaspida#, formerly united by me with the Dorataspida (in
the Prodromus, 1881, p. 468), differs from it in the ellipsoidal form of
the lattice-shell and {860}the enclosed central capsule. In the spherical
Dorataspida, their ancestral family, all twenty radial spines are of equal
size, whilst here in the Belonaspida two opposite spines are larger than
the eighteen others. These two larger or principal spines are both
equatorial spines, placed in the longitudinal or major axis of the
ellipsoid, or the "hydrotomical axis" (compare above, p. 719). The two
other equatorial spines are constantly smaller, and lie in the transverse
or minor axis of the ellipsoid, or the "geotomical axis." The geometrical
proportion of these two determining axes of the ellipsoidal shell is very
variable (even in one and the same species), commonly 4 : 3 or 3 : 2,
rarely 2 : 1, often only 5 : 4 or 6 : 5. All meridian planes, passing
through the principal spines (or the longitudinal axis of the shell) are
elliptical, and of equal size. All transverse planes, perpendicular to that
axis, are circular; the largest of these circular parallel planes is the
geotomical plane, which passes through the smaller equatorial spines and
the spineless axis.

In the spherical Dorataspida the internal length of the radial spines (or
the distance between the shell and the centre) is equal in all twenty
spines. In the ellipsoidal Belonaspida this internal length is different,
and commonly exhibits four different degrees; in the two principal spines
it is of first rank, in the eight tropical spines of second rank, in the
four (hydrotomical) polar spines of third rank, and in the six spines (four
geotomical polar spines and two transverse equatorial spines) of fourth
rank. These differences of the internal length become more important the
more the hydrotomical axis is prolonged. Regarding all other qualities (in
form, disposition, and mode of junction at the centre) the ellipsoidal
Belonaspida do not differ from their ancestral group, the spherical
Dorataspida (compare above, p. 802). In both families the pyramidal central
bases of the twenty spines are commonly supported one upon another with
their triangular neighbouring faces; but sometimes also here (particularly
in _Phatnaspis_) the central bases are perfectly grown together. In this
case also the sutures of the meeting branches of the apophyses are
obliterated, whilst usually they remain open. Such forms, with spines and
plates perfectly grown together, form a single piece of acanthin, and were
formerly separated by me as a peculiar genus _Haliommatidium_ (Monogr. d.
Radiol., 1862, p. 419). But as I now find this concrescence to be an
accidental and inconstant peculiarity of some species, it no longer seems
to me to be of generic importance.

The apophyses of the twenty radial spines, the branches of which compose
the ellipsoidal lattice-shell, exhibit in all Belonaspida the same
appearance as in the Diporaspida (and especially the Ceriaspida) among the
spherical Dorataspida. Everywhere each radial spine gives off only two
opposite primary apophyses, the fork-branches of which unite together and
form a plate or shield with two aspinal pores (as in _Dorataspis_).
Commonly the shell exhibits only these forty parmal pores, the other meshes
between them being sutural pores. Only in one genus, _Phatnaspis_ (Pl. 136,
{861}figs. 6-9), is this number multiplied; the two opposite primary
apophyses are here crossed by numerous perpendicular lateral branches, and
these are again united by secondary perpendicular ramules which are
parallel to the apophyses themselves; therefore each plate here forms a
rhombic shield pierced by very numerous (often more than one hundred) small
quadrangular or circular pores. Only two of these numerous parmal pores are
the primary aspinal pores (sometimes, as in Pl. 136, fig. 8, distinguished
by their larger size); all others are secondary or coronal pores. Therefore
_Phatnaspis_ repeats the characteristic formation of _Coscinaspis_, from
which it differs in its ellipsoidal shell.

Moreover, the four other genera of the ellipsoidal Belonaspida correspond
perfectly to certain genera of the spherical Dorataspida, and may be
derived from these by hypertrophy or stronger development of two opposite
equatorial spines. In _Thoracaspis_ and _Belonaspis_ (Pl. 139, figs. 8, 9)
the surface of the shell-plates is without crests and dimples (in the
former without, in the latter with accessory by-spines); they thus
correspond to _Dorataspis_ and _Diporaspis_ among the Dorataspida. In two
other genera the surface of the shell-plates bears a network of elevated
crests, separating funnel-shaped dimples, in _Dictyaspis_ without, in
_Coleaspis_ with by-spines; these correspond to _Ceriaspis_ and
_Hystrichaspis_ among the Dorataspida. The small by-spines, covering the
surface, exhibit the same forms as in the Dorataspida; but they are usually
smaller, and less developed in the Belonaspida.

The twenty radial spines are commonly more or less compressed or two-edged,
sometimes very flat, broad, and triangular (Pl. 139, figs. 8, 9). In some
species (mainly of _Phatnaspis_) they are very long, thin, and
needle-shaped (Monogr. d. Radiol., 1862, Taf. xxii. figs. 10-12). Rarely
they are cylindrical, four-edged, or prismatic. The Belonaspida testify by
this and other peculiarities to their near relation to the Diporaspida and
Phractacanthida and their older origin from _Zygacantha_.

_The Central Capsule_ is in the Belonaspida ellipsoidal, and more or less
prolonged in the hydrotomical or longitudinal axis of the shell. It is
constantly smaller than the enveloping ellipsoidal shell, and separated
from it by the calymma. Its structure and the shape of its nucleus are the
same as in the nearly allied Dorataspida.

_Synopsis of the Genera of Belonaspida._

-------------------------------------------------------------------------
I. Subfamily Coleaspida.
Forty parmal pores (two aspinal pores on each plate, no coronal
pores).
-------------------------------------------------------------------------
Plates not dimpled, without { No by-spines, 371. _Thoracaspis_.
prominent crests. {
{ With by-spines, 372. _Belonaspis_.

Plates dimpled, with a { No by-spines, 373. _Dictyaspis_.
network of prominent {
crests, { With by-spines, 374. _Coleaspis_.
-------------------------------------------------------------------------
II. Subfamily Phatnaspida.
Eighty to two thousand or more parmal pores (two aspinal pores on
each plate, surrounded by two to one hundred or more coronal pores).
-------------------------------------------------------------------------
No by-spines (plates not dimpled), 375. _Phatnaspis_.

{862}Genus 371. _Thoracaspis_,[411] Haeckel, 1862, Prodromus, p. 468.

_Definition._--#Belonaspida# with forty parmal pores (two on each plate),
without dimples and crests, also without by-spines, therefore with smooth
surface.

The genus _Thoracaspis_ is the simplest form among the Belonaspida, and
repeats in this family the nearly allied genus _Dorataspis_ among the
Dorataspida. It differs from this genus only in the characteristic
prolongation of one equatorial axis, which effects an ellipsoidal
transformation of the spherical central capsule and the enclosing
lattice-shell.

1. _Thoracaspis ellipsoides_, n. sp.

Parmal pores large, elliptical, five or six times as large as the small
circular sutural pores. Each plate surrounded by five or six sutural pores
(a single one on each side). Spines compressed triangular, sword-like,
two-edged; inner and outer half of nearly equal length.

_Dimensions._--Length of the ellipsoidal shell (or major axis) 0.15,
breadth (or minor axis) 0.12; length of the spines 0.07, basal breadth
0.016.

_Habitat._--North Pacific, Station 253, surface.

2. _Thoracaspis nephropora_, n. sp.

Parmal pores large, kidney-shaped, three to four times as large as the
roundish small sutural pores. Each plate surrounded by five or six sutural
pores (a single one on each side). Spines in the inner longer part
cylindrical, in the outer shorter part conical, not compressed.

_Dimensions._--Length of the shell 0.2, breadth 0.15; length of the spines
0.06, basal breadth 0.018.

_Habitat._--Central Pacific, Station 268, surface.

3. _Thoracaspis circopora_, n. sp.

Parmal pores circular, of the same size as the circular sutural pores. Each
plate surrounded by five or six sutural pores (a single one on each side).
Spines compressed, thin, linear, two-edged; outer part one and a half times
as long as the inner.

_Dimensions._--Length of the shell 0.17, breadth 0.13; length of the spines
0.12, basal breadth 0.003.

_Habitat._--South Pacific, Station 288, surface.

4. _Thoracaspis bipennis_, n. sp. (Pl. 139, fig. 8).

Parmal pores elliptical, about twice as large as the circular sutural
pores, which are of rather variable size. Each plate surrounded by eight to
twelve sutural pores (commonly two on each side). {863}Spines compressed,
triangular, gradually tapering towards both ends; outer half a little
longer, and much broader than the inner.

_Dimensions._--Length of the shell 0.14, breadth 0.12; length of the spines
0.08, basal breadth 0.025.

_Habitat._--Central Pacific, Station 274, surface.

Genus 372. _Belonaspis_,[412] Haeckel, 1862, Prodromus, p. 468.

_Definition._--#Belonaspida# with forty parmal pores (two on each plate),
without dimples and crests, but with numerous by-spines on the surface.

The genus _Belonaspis_ differs from its ancestral genus _Thoracaspis_ only
in the possession of numerous superficial by-spines, and bears therefore to
it the same relation as _Diporaspis_ does to _Dorataspis_. The two latter
differ from the two former in the spherical form of the central capsule and
the enclosing shell, which here becomes ellipsoidal.

1. _Belonaspis pandanus_, n. sp.

Parmal pores elliptical, three or four times as large as the circular
sutural pores. Each plate surrounded by five or six sutural pores (a single
one on each side). Spines compressed, triangular; outer part half as long
as the inner, and twice as long as the numerous, simple, bristle-shaped
by-spines.

_Dimensions._--Length of the ellipsoidal shell (or major axis) 0.12,
breadth (or minor axis) 0.1; length of the spines 0.03, basal breadth 0.02.

_Habitat._--South Pacific, Station 295, surface.

2. _Belonaspis furcata_, n. sp.

Parmal pores elliptical, twice as broad as the circular sutural pores. Each
plate surrounded by five or six sutural pores (a single one on each side).
Spines compressed, linear, very thin; outer part longer than the inner.
By-spines very numerous, half as long as the radius, furcate, with
divergent fork-branches.

_Dimensions._--Length of the shell 0.1, breadth 0.08; length of the spines
0.12, breadth 0.003.

_Habitat._--Indian Ocean (Cocos Islands), Rabbe, surface.

3. _Belonaspis datura_, n. sp. (Pl. 139, fig. 9).

Parmal pores elliptical, three or four times as large as the circular
sutural pores. Each plate surrounded by ten or twelve sutural pores (two on
each side). Spines triangular, two-edged, about {864}as long as the radius
of the shell; gradually tapering from its surface towards both ends.
By-spines very short and numerous, simple.

_Dimensions._--Length of the shell 0.13, breadth 0.11; length of the spines
0.08, basal breadth 0.025.

_Habitat._--Central Pacific, Station 271, surface.

4. _Belonaspis lanceolata_, n. sp.

Parmal pores elliptical, six or eight times as large as the small circular
sutural pores. Each plate surrounded by ten to twelve sutural pores (two on
each side). Spines lanceolate, flat, in the distal part needle-shaped,
about as long as the diameter of the shell. By-spines very numerous,
zigzag, half as long as the shell-radius.

_Dimensions._--Length of the shell 0.15 breadth 0.13; length of the spines
0.14, basal breadth 0.03; length of the by-spines 0.04.

_Habitat._--North Pacific, Station 235, surface.

5. _Belonaspis conifera_, n. sp.

Parmal pores kidney-shaped, three or four times as large as the small
circular pores. Each plate surrounded by ten to twelve sutural pores (two
on each side). Spines very thick, half as long in the outer conical part as
in the inner cylindrical part. By-spines very short, conical.

_Dimensions._--Length of the shell 0.2, breadth 0.16; length of the spines
0.05, basal breadth 0.03.

_Habitat._--Central Pacific, Station 267, surface.

6. _Belonaspis multiforis_, n. sp.

Parmal pores circular, of the same size as the circular sutural pores. Each
plate surrounded by fifteen to eighteen sutural pores (three on each side).
Spines compressed, about as long as the radius. By-spines very numerous,
zigzag, half as long as the radius.

_Dimensions._--Length of the shell 0.22, breadth 0.18; length of the spines
0.12, basal breadth 0.012; length of the by-spines 0.06.

_Habitat._--Equatorial Atlantic, Station 347, surface.

Genus 373. _Dictyaspis_,[413] n. gen.

_Definition._--#Belonaspida# with forty parmal pores (two on each plate),
without by-spines, but with a network of prominent crests on the dimply
surface.

The genus _Dictyaspis_ repeats among the Belonaspida the characteristic
structure of _Ceriaspis_ (among the Dorataspida), by the development of
prominent crests forming a {865}network with dimples on the outer surface
of the shields. Either all the dimples or only a part of them are pierced
by a pore. The forty aspinal pores are united in pairs in twenty larger
dimples. The shell is usually very thick-walled and non-transparent.

1. _Dictyaspis solidissima_, Haeckel.

_Dorataspis solidissima_, Haeckel, 1862, Monogr. d. Radiol., p. 416, Taf.
xxii. figs. 6-9.

Surface of the shell with seventy to eighty funnel-shaped dimples, each of
which is perforated at the base by one or two small meshes; twenty larger
dimples in the centre of the plates (each with two elliptical aspinal
pores), and fifty to sixty smaller sutural dimples between them (each with
a single circular pore). All pores of nearly the same size. No blind
dimples. Outer conical part of the thick radial spines of the same length
as the inner cylindrical part. Radius of the shell four times as large as
the thickness of its wall.

_Dimensions._--Length of the shell 0.12, breadth 0.1; length of the spines
0.06, basal breadth 0.012.

_Habitat._--Mediterranean (Messina), surface.

2. _Dictyaspis favosa_, n. sp.

Surface of the shell with one hundred and seventy to one hundred and ninety
deep funnel-shaped dimples, seventy to eighty of which are perforated, the
others blind; among the former each of the fifty to sixty smaller contains
a single sutural pore, each of the twenty larger a couple of aspinal pores.
All pores of nearly the same size. Outer part of the compressed two-edged
spines triangular, half as long as the inner part. Radius of the shell six
times as large as the thickness of its wall. (Differs from _Ceriaspis
favosa_, Pl. 138, fig. 6, mainly in the ellipsoidal form of the shell and
the different size and form of the spines.)

_Dimensions._--Length of the shell 0.16, breadth 0.13; length of the spines
0.04, basal breadth 0.025.

_Habitat._--Central Pacific, Station 265, surface.

3. _Dictyaspis compacta_, n. sp.

Surface of the shell with one hundred and seventy to one hundred and ninety
deep funnel-shaped dimples, seventy to eighty of which are perforated, the
others blind; among the former each of fifty to sixty each contain a single
sutural pore, twenty each a couple of aspinal pores. All dimples nearly of
the same size, very deep. Outer part of the compressed sword-like spines
somewhat longer than the inner part. Radius of the dark shell only twice as
large as the thickness of its compact wall.

_Dimensions._--Length of the shell 0.18, breadth 0.15; length of the spines
0.12, basal breadth 0.02.

_Habitat._--Central Pacific, Station 274, surface.

{866}Genus 374. _Coleaspis_,[414] Haeckel, 1881, Prodromus, p. 468.

_Definition._--#Belonaspida# with forty parmal pores (two on each plate),
with numerous by-spines and with a network of prominent crests on the
dimpled surface.

The genus _Coleaspis_ differs from its ancestral genus _Dictyaspis_ in the
development of numerous by-spines, and bears therefore among the
ellipsoidal Belonaspida the same relation to it as _Hystrichaspis_ does to
_Coscinaspis_ among the spherical Dorataspida. If the four equatorial
spines in _Coleaspis_ become different in pairs, it passes over into
_Hexalaspis_. The shell is usually very dark and thick-walled.

1. _Coleaspis coronata_, n. sp.

All twenty spines of nearly equal size and form, about as long as the
radius of the shell, compressed, sword-like, two-edged; both equatorial
spines of the hydrotomical axis little longer than the eighteen others.
Crests between the dimples of the shell-surface dentated, forming around
the basal half of each spine a cylindrical sheath with a dentated,
crown-like mouth, the teeth of which are prolonged into simple by-spines.
(Similar to _Hexaconus coronatus_, Pl. 139, fig. 5, but differing in the
nearly equal size of the spines and their sheaths, and in the longer
by-spines.)

_Dimensions._--Major axis of the shell 0.2, minor 0.17; length of the
spines 0.12, basal breadth 0.02.

_Habitat._--Central Pacific, Station 266, surface.

2. _Coleaspis vaginata_, n. sp. (Pl. 140, fig. 13).

All twenty spines of nearly equal size and form, nearly as long as the
diameter of the shell, conical; both equatorial spines of the hydrotomical
axis about one-third longer than the eighteen others. Crests between the
dimples of the shell-surface serrated, forming around the basal half of
each spine a conical truncate sheath with crested surface and serrated
mouth, the teeth of which are prolonged into short by-spines. (Similar to
_Hexaconus vaginatus_, Pl. 139, fig. 7, but differing in the nearly equal
size of all the spines and in the double thickness of the shell-wall.)

_Dimensions._--Major axis of the shell 0.12, minor 0.1, length of the
spines 0.08, basal breadth 0.01.

_Habitat._--North Pacific, Station 253, surface.

3. _Coleaspis obscura_, n. sp.

All twenty spines of nearly equal size and form, little longer than their
large cylindrical sheaths, which are nearly as long as the radius of the
shell, and armed on the truncated mouth with strong triangular by-spines,
both equatorial spines of the hydrotomical axis one and a half times as
long as the eighteen others. Shell very dark and thick walled, quite
opaque.

{867}_Dimensions._--Major axis of the shell 0.1, minor 0.08; length of the
spines 0.06, of their sheaths 0.04.

_Habitat._--Arctic Ocean, Greenland, Koch, surface.

4. _Coleaspis occulta_, n. sp.

All twenty spines of nearly equal size and form, quite concealed in their
long cylindrical sheaths, which are somewhat longer than the radius of the
shell, and armed on the truncated mouth with acute simple teeth; each
sheath deeply sulcated, apparently resulting from the concrescence of
numerous parallel leaf-shaped by-spines. Small shell very dark and
thick-walled, quite opaque.

_Dimensions._--Major axis of the shell 0.08, minor 0.06; length of the
spines and their sheaths 0.05 to 0.06.

_Habitat._--Antarctic Ocean (off Kerguelen), Station 159, surface.

Subgenus 2. _Coleaspidium_, Haeckel.

_Definition._--Both equatorial spines of the hydrotomical axis much larger
and of peculiar form, different from the eighteen other spines.

5. _Coleaspis amphilonche_, n. sp.

Both equatorial spines of the hydrotomical axis much longer than the
eighteen others, and of very different form; one and a half times as long
as the diameter of the shell, prismatic, with six prominent edges,
pyramidal at the distal point. The eighteen other spines triangular,
compressed, two-edged, scarcely as long as the radius of the thick-walled
shell. Pores of the shell irregular polygonal, separated by ciliated
crests, which bear simple by-spines (one-third as long as the radius).

_Dimensions._--Major axis of the shell 0.15, minor 0.12; length of the two
larger spines 0.22, of the eighteen smaller 0.06 to 0.08; basal breadth of
the former 0.03, of the latter 0.008.

_Habitat._--South Atlantic, Station 341, surface.

6. _Coleaspis hydrotomica_, n. sp. (Pl. 140, fig. 14).

Both equatorial spines of the hydrotomical axis much larger than the
eighteen others, and of very different form; about as long as the diameter
of the shell, in the basal half four-sided pyramidal, with four thick
prominent edges, in the middle part constricted, in the distal half
lanceolate, two-edged. Both equatorial spines of the geotomical axis of
similar form, but much smaller, scarcely one-third as long. The sixteen
other spines sword-like, thin, two-edged, about as long as the radius of
the thick-walled shell. Pores of the shell irregular roundish, separated by
high dentated crests, which bears zigzag by-spines (half as long as the
radius).

_Dimensions._--Major axis of the shell 0.12, minor axis 0.1; length of the
two larger spines 0.11, of the eighteen others 0.04 to 0.06; basal breadth
of the former 0.04, of the latter 0.01.

_Habitat._--Cape of Good Hope, Station 143, surface.

{868}Genus 375. _Phatnaspis_,[415] Haeckel, 1881, Prodromus, p. 468

_Definition._--#Belonaspida# with eighty to two thousand or more parmal
pores (four to one hundred or more on each plate), without by-spines on the
surface.

The genus _Phatnaspis_ corresponds to the spherical _Coscinaspis_ among the
Dorataspida, and differs from the other Belonaspida in the great number of
the parmal pores; whilst the four other preceding genera exhibit only two
opposite aspinal pores in the centre of each plate, in this there are
constantly numerous coronal pores in addition to these, and the plates
always possess the characteristic form of a wainscotted or panelled work,
with quadrangular meshes. The number of these parmal pores amounts in each
plate to from ten to twenty, often one hundred and twenty or more;
therefore the number of parmal pores in the whole shell amounts to two
thousand or more. Sometimes the pores are circular, but in this case too
they are surrounded by quadrangular frames. The quadrangles are sometimes
quite regular squares, sometimes more or less irregular. The thin and
fragile bars between the quadrangular pores form in each plate two peculiar
systems of parallel crests, which cross at right angles. Commonly the
parallel crests of one system (parallel to the major diameter of the
compressed radial spines) are equidistant, and pierce from one edge of the
plate to the opposite, whilst the parallel crests of the other system
(parallel to the minor diameter of the spines) are interrupted and at
different distances (Pl. 136, fig. 9); but in other species both crossed
systems are quite regular. In each plate there is one primary diagonal rib
(often stronger than the parallel crests) which connects the two opposite
corners of the rhomboidal plate. We can distinguish in this genus three
different subgenera: A. in _Phatnasparium_ the primary diagonal rib arises
from both flat sides of the compressed sword-like, radial spines; B. in
_Phatnasplenium_ from both sharp edges of them; C. in _Phatnaspidium_ two
crossed diagonal ribs arise from four edges of the spines (combination of A
and B). Therefore in the first subgenus (A) two primary aspinal pores are
placed opposite on the sharp edges of the spines, but in the second (B)
inversely on their flat sides; in the third (C) there are apparently four
primary aspinal pores, which are probably derived from B or A by division
of the two pores.

Subgenus 1. _Phatnasparium_, Haeckel.

_Definition._--In the centre of each rhomboidal plate two primary aspinal
pores, opposite on both edges of the compressed spines, from both flat
sides of which the primary diagonal crest arises.

{869}1. _Phatnaspis lacunaria_, n. sp. (Pl. 136, fig. 9).

Parmal pores irregular quadrangular, of unequal size and form, ten to
twelve on each side of the primary diagonal rib, which arises from both
flat sides of the two-edged spines; two primary aspinal pores opposite on
both edges of the latter. Spines much compressed, sword-like, their outer
half shorter than the inner.

_Dimensions._--Length of the shell 0.21 to 0.23, breadth 0.18 to 0.2; basal
breadth of the spines 0.01.

_Habitat._--Central Pacific, Station 268, surface.

2. _Phatnaspis ensiformis_, n. sp.

Parmal pores irregular quadrangular, of unequal size and form, six to eight
on each side of the primary diagonal rib, which arises from both flat sides
of the two-edged spines; two primary aspinal pores opposite on the two
edges of the latter. Spines very broad, strongly compressed, sword-like,
their outer half larger than the inner.

_Dimensions._--Length of the shell 0.16, breadth 0.11; basal breadth of the
spines 0.02.

_Habitat._--Tropical Atlantic, Station 348, surface.

3. _Phatnaspis loculata_, n. sp.

Parmal pores regular quadrangular, of nearly equal size and form, twelve to
sixteen on each side of the primary diagonal rib, which arises from both
flat sides of the two-edged spines; two primary aspinal pores on the two
edges of the latter. Spines linear, compressed, very long and thin, their
outer half three to four times as long as the inner.

_Dimensions._--Length of the shell 0.24, breadth 0.18; breadth of the
spines 0.004.

_Habitat._--North Atlantic, Station 354, surface.

4. _Phatnaspis fenestrata_, Haeckel.

_Haliommatidium fenestratum_, Haeckel, 1862, Monogr. d. Radiol., p. 421.

Parmal pores regular quadrangular, of nearly equal size and form, six to
eight on each side of the primary diagonal rib, which arises from both flat
sides of the two-edged spines; two primary aspinal pores on the two edges
of the latter. Spines linear, little compressed, or nearly needle-shaped,
very long and thin, their outer half five to ten times as long as the
inner.

_Dimensions._--Length of the shell 0.11 to 0.12, breadth 0.07 to 0.08;
breadth of the spines 0.002.

_Habitat._--Mediterranean (Messina), surface.

5. _Phatnaspis cristata_, n. sp. (Pl. 136, fig. 6).

Parmal pores circular, of very different size, separated by high square
crests, ten to twelve on each side of the high, comb-like, primary diagonal
rib, which arises from both flat sides of the {870}twoedged spines; two
primary aspinal pores on the two edges of the latter. Each circular pore is
surrounded by a square frame. Spines sword-like, strongly compressed, their
outer part longer than the inner.

_Dimensions._--Length of the shell 0.2 to 0.22, breadth 0.16 to 0.18; basal
breadth of the spines 0.01.

_Habitat._--North Pacific, Station 254, surface.

6. _Phatnaspis coscinoides_, n. sp.

Parmal pores circular, regular, all of nearly equal size, twelve to sixteen
on each side of the primary diagonal rib, which arises from both flat sides
of the two-edged spines; two primary pores on the two edges of the latter.
Spines linear, little compressed, their outer part much longer than the
inner.

_Dimensions._--Length of the shell 0.18, breadth 0.15; breadth of the
spines 0.006.

_Habitat._--South Atlantic, Station 325, surface.

Subgenus 2. _Phatnasplenium_, Haeckel.

_Definition._--In the centre of each rhomboidal plate two primary aspinal
pores, opposite on the two flat sides of the compressed spines, from the
two edges of which the primary diagonal crest arises.

7. _Phatnaspis orthopora_, n. sp.

Parmal pores irregular quadrangular, of unequal size and form, four to six
on each side of the primary diagonal rib, which arises from both edges of
the compressed spines; two larger primary aspinal pores opposite on the two
flat sides of the latter. Spines very thin and long, linear, their outer
half three to four times as long as the inner.

_Dimensions._--Length of the shell 0.08, breadth 0.06; breadth of the
spines 0.003.

_Habitat._--North Atlantic, Station 353, surface.

8. _Phatnaspis polypora_, n. sp.

Parmal pores irregular polygonal, of very unequal size and form, sixteen to
twenty on each side of the primary diagonal rib, which arises from both
edges of the compressed spines; two very large, lanceolate, primary aspinal
pores opposite on the two flat sides of the latter. Spines very thin,
linear, on the outside of the shell rudimentary. (This remarkable species
somewhat resembles _Coscinaspis polypora_, Pl. 136, fig. 8.)

_Dimensions._--Length of the shell 0.32, breadth 0.24; breadth of the
spines 0.002.

_Habitat._--South Atlantic, Station 332, surface.

{871}9. _Phatnaspis quadratura_, n. sp.

Parmal pores regular, square, all of nearly equal size and form, ten to
twelve on each side of the primary diagonal rib, which arises from both
edges of the compressed spines; two primary aspinal pores opposite on the
two flat sides of the latter, not different from the other pores. Spines
sword-like, their outer part about as long as the inner.

_Dimensions._--Length of the shell 0.2, breadth 0.16; basal breadth of the
spines 0.004.

_Habitat._--Central Pacific, Station 274, surface.

10. _Phatnaspis tabulata_, Haeckel.

_Haliomma tabulatum_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 37, Taf. v. figs. 5-8.

_Haliomma tabulatum_, Haeckel, 1862, Monogr. d. Radiol., p. 429.

Parmal pores regular, circular, with square frames, all of nearly equal
size, eight to ten on each side of the primary diagonal rib, which arises
from both edges of the compressed spines; two primary aspinal pores
opposite on the flat sides of the latter. Spines sword-like, short,
scarcely half as long as the radius of the shell, their central ends are
thickened and cause by their union the deceptive appearance of an enclosed
"medullary shell."

_Dimensions._--Length of the shell 0.12 to 0.15, breadth 0.1 to 0.12;
breadth of the spines 0.006.

_Habitat._--Mediterranean; Nice, Müller; Portofino, near Genoa, Haeckel,
surface.

Subgenus 3. _Phatnaspidium_, Haeckel.

_Definition._--In the centre of each rhomboidal plate four primary aspinal
spines, forming a regular cross, the centre of which receives the
cylindrical or four-sided prismatic spine.

11. _Phatnaspis haliommidium_, n. sp. (Pl. 136, fig. 7).

Parmal pores irregular quadrangular, of unequal size and form, eight to ten
on each side of the crossed diagonal ribs, which arise at right angles from
the four edges of the prismatic spines; four primary aspinal pores not
different from the others. Outer part of the strong spines scarcely longer
than the inner. (Resembling _Icosaspis tabulata_, Pl. 136, fig. 2, which,
however, is distinguished by the spherical shell and the larger pores.)

_Dimensions._--Length of the shell 0.22, breadth 0.17; breadth of the
spines 0.016.

_Habitat._--North Pacific, Station 244, surface.

12. _Phatnaspis mülleri_, n. sp.

_Haliommatidium mülleri_, Haeckel, 1862, Monogr. A. Radiol, p. 419, pl.
xiii. figs. 10-12.

Parmal pores regular, square, all of nearly equal size and form, twelve to
sixteen on each side of the crossed diagonal ribs, which arise at right
angles from the needle-shaped spines; the four {872}primary aspinal pores
not different from the others. Spines very thin and long, cylindrical; four
to six times longer in the outer than in the inner part.

_Dimensions._--Length of the shell 0.24, breadth 0.16; breadth of the
spines 0.002.

_Habitat._--Mediterranean (Messina), surface.

Family XLIII. #HEXALASPIDA#, n. fam. (Pl. 139).

_Definition._--ACANTHARIA with a simple discoidal or lenticular
lattice-shell, composed of the branched apophyses of twenty radial spines
meeting in the centre and disposed according to the Müllerian law of
Icosacantha. Six larger spines in the hydrotomical plane, prominent on the
margin of the circular or elliptical biconvex lens. Fourteen other spines
much smaller or rudimentary. Central capsule biconvex lenticular, enclosed
in the fenestrated shell.

The family #Hexalaspida# represents a new small, but very interesting group
of Acanthophracta, which differs from all others in the lentelliptical or
triaxial form of the lenticular lattice-shell, the margin of which bears
six larger spines placed in the hydrotomical plane (compare above, p. 719).
They may therefore be characterised shortly as "_Acanthophracta
lentelliptica_," with three different dimensive axes and six larger
marginal spines. A closer comparison with the other ACANTHARIA leaves no
doubt that the Hexalaspida must be derived from the Belonaspida by stronger
development of six radial spines placed in the hydrotomical plane, namely,
two equatorial and four associated polar spines; whilst the six spines of
the geotomical plane (perpendicular to the former) are much smaller; the
eight tropical spines are intermediate in size between the former and the
latter.

The geometrical fundamental form of the Hexalaspida (of the central capsule
as well as of the enclosing shell) is therefore lentelliptical, with three
different dimensive axes, and they exhibit among the ACANTHARIA a relation
to the spherical Dorataspida and the ellipsoidal Belonaspida similar to
that which the lentelliptical #Larcoidea# exhibit to the spherical
#Sphæroidea# and the ellipsoidal #Prunoidea# among the #Sphærellaria#
(compare above, p. 599). The largest of the three dimensive axes (which are
perpendicular to one another) is here the hydrotomical axis, the shortest,
on the contrary, the geotomical axis; the intermediate in size being the
spineless axis. The development of the whole body is strongest in the
hydrotomical meridian plane, in which the six principal spines are placed;
it is weakest in the geotomical plane, in which the six smallest spines are
placed; the eight tropical spines are intermediate in size between the
others. This peculiar development is illustrated by the figures of Pl. 139,
where the four equatorial spines are everywhere marked by _c_, the eight
tropical spines by _b_ and _d_, the eight polar spines by _a_ and _e_.

{873}Rarely the six hydrotomical or principal spines are of equal size, and
thus the margin of the shell may be quite circular or regularly hexagonal
(Pl. 139, figs. 1-3); the two equatorial spines of the hydrotomical plane
are usually larger than its four polar spines, and thus the margin of the
shell becomes more or less elliptical (Pl. 139, figs. 4-7). Usually (almost
constantly) these six larger spines are more or less compressed,
triangular, often very broad and flat; their two edges lie in the
hydrotomical plane. Their two flat surfaces are often furrowed, with
longitudinal ribs or crests converging towards the simple apex of the
spines.

The fourteen smaller spines are regularly disposed according to the
Müllerian law of Icosacantha on both convex sides of the lenticular shell;
they are not only smaller than the six principal spines, but often also of
different form, much thinner and shorter, sometimes needle-shaped. In the
genera _Hexonaspis_ and _Hexacolpus_ (Pl. 139, figs. 1, 2) only their inner
part (inside of the shell) is developed, whilst their outer part is quite
rudimentary and not prominent on the surface. Therefore these genera appear
to possess only six marginal spines externally.

_The Lenticular Shell_ itself offers in the Hexalaspida great difficulties
in the way of accurate study, as its wall is constantly very thick and
dark, often quite opaque and non-transparent. However, prolonged accurate
researches have convinced me that its structure is essentially the same as
in the Belonaspida and especially in the genera _Dictyaspis_ and
_Coleaspis_. As in these latter the twenty plates of the shell bear high
crests or combs on the outer surface, and by these funnel-shaped dimples
are separated. The network of these crests is more or less regular (Pl.
139, figs. 1-7). Around the base of each radial spine the shell is usually
elevated in the form of a conical or cylindrical sheath; the crests are
prolonged into the sheaths as longitudinal ribs, parallel to the spine or
convergent towards its apex. Whilst in _Hexalaspis_ and _Hexonaspis_ these
basal sheaths are not at all or but little prominent (Pl. 139, fig. 2; Pl.
140, fig. 16), in _Hexaconus_ and _Hexacolpus_ they envelop the basal half
(or even more) of the spines, and very often the circular or elliptical
free distal edge of the sheath is elegantly denticulated or serrated (Pl.
139, figs. 1, 3-7; pl. 140, figs. 9-16).

_The Pores_ of the shell exhibit in the Hexalaspida the same shape as in
the majority of the Belonaspida. Each spine bears only two broad opposite
apophyses, the fork-branches of which unite to form a polygonal shield with
two pores. The number of parmal pores is constantly (?) forty, as each
plate possesses only two primary aspinal pores; there are no secondary or
coronal pores. The numerous (between fifty and one hundred, rarely more)
smaller pores between the forty parmal pores are probably always sutural
pores; however, their number and position is very difficult to determine,
on account of the high protecting crests; the majority of the funnel-shaped
dimples between the latter seem to be blind, not perforated. Sometimes all
the dimples, except the twenty spinal ones, seem to be blind and the
sutural pores appear {874}to have disappeared completely, so that there
remain only twenty aspinal pores (Pl. 139, fig. 4).

The internal cavity of the shell in all Hexalaspida is very small, on
account of the thickness of the massive wall; the latter is often greater
than the diameter of the cavity. Setting aside this disproportion, the
space of the cavity is further much reduced by the internal parts of the
thick radial spines, which are united in the centre by their pyramidal
bases (Pl. 140, fig. 15). Usually the twenty bases seem to remain free
(supported one upon another by means of their triangular faces); but
sometimes they seem to grow together perfectly and to form a single central
star of acanthin. In some species, too, the sutures of the meeting branches
of the apophyses of neighbouring spines seem to grow together, so that the
whole shell exhibits a single piece of acanthin.

_The Central Capsule_ of the Hexalaspida is therefore very small and seems
to fill up the greatest part of the shell-cavity. Its form is constantly
more or less lenticular, sometimes lentelliptical. On account of the
opacity of the shell I could not make out its shape more closely.

_Synopsis of the Genera of Hexalaspida._

All twenty spines externally } Sheaths of the spines
developed, prominent on } not prominent, 376. _Hexalaspis_.
the surface of the }
lenticular shell. } Sheaths of the spines
} prominent, 377. _Hexaconus_.

Only six spines (the }
hydrotomical spines) } Sheaths of the spines
externally developed and } not prominent, 378. _Hexonaspis_.
prominent on the margin }
of the disk; the other } Sheaths of the spines
fourteen spines not } prominent, 379. _Hexacolpus_.
prominent. }

Genus 376. _Hexalaspis_,[416] n. gen.

_Definition._--#Hexalaspida# with twenty prominent radial spines, which are
not surrounded by prominent sheaths; the six hydrotomical spines much
larger than the fourteen others.

The genus _Hexalaspis_ is the simplest form among the Hexalaspida, and may
be derived directly from _Dictyaspis_ among the Belonaspida, by stronger
development of the six hydrotomical spines. As in the following genus
_Hexaconus_ all twenty spines are prominent externally.

Subgenus 1. _Hexalasparium_, Haeckel.

_Definition._--Six hydrotomical spines of equal size.

{875}1. _Hexalaspis heliodiscus_, n. sp. (Pl. 139, fig. 2).

All six hydrotomical spines of nearly equal size (or sometimes the two
equatorial a little larger than the four polar spines), isosceles
triangular, compressed, smooth; about as long as the radius of the shell,
and half as broad at the base. The fourteen other spines very small, also
triangular and compressed, but little prominent on the two convex sides of
the lenticular shell.

_Dimensions._--Diameter of the shell 0.11; length of the six hydrotomical
spines 0.05, basal breadth 0.03.

_Habitat._--Central Pacific, Station 271 to 274, surface.

2. _Hexalaspis stellata_, n. sp.

All six hydrotomical spines of nearly equal size, lanceolate compressed,
with two longitudinal furrows on each flat side, about as long as the
diameter of the shell, and one-fourth as broad at the base. The fourteen
other spines very thin, also compressed, half as long and only one-fourth
as broad as the six large spines.

_Dimensions._--Diameter of the shell 0.14; length of the six hydrotomical
spines 0.13, basal breadth 0.035.

_Habitat._--South Pacific, Station 284, surface.

Subgenus 2. _Hexalaspidium_, Haeckel.

_Definition._--Six hydrotomical spines of unequal size, two opposite
(equatorial) much larger than the four other (polar) spines.

3. _Hexalaspis sexalata_, n. sp.

Six hydrotomical spines of unequal size; the two equatorial spines about as
long as the shell-radius and twice as long as the four polar spines, which
are isosceles triangular. The fourteen other spines are only half as long
and one-fourth as broad as the latter, little prominent. (Resembles
_Hexonaspis hastata_, Pl. 140, fig. 16, which is distinguishable by the
furrows on the six spines and by the total absence of the fourteen external
rudimentary spines.)

_Dimensions._--Diameter of the shell 0.15; length of the two equatorial
spines 0.08, of the four polar spines 0.04, of the fourteen other spines
0.02.

_Habitat._--North Pacific, Station 240, surface.

4. _Hexalaspis hexalastrum_, n. sp.

Six hydrotomical spines of unequal size; the two equatorial spines somewhat
longer than the diameter of the shell and three times as long as the four
polar spines, all six triangular, smooth, of the same basal breadth (equal
to half the radius). The fourteen other spines very thin, conical at the
base, nearly as long as the radius.

{876}_Dimensions._--Diameter of the shell 0.21; length of the two
equatorial spines 0.24, of the eighteen others 0.08 to 0.1; basal breadth
of the six larger spines 0.05, of the fourteen others 0.01.

_Habitat._--Western Tropical Pacific, Station 224, surface.

5. _Hexalaspis hexaglypha_, n. sp.

Six hydrotomical spines of unequal size; the two equatorial very large,
six-sided prismatic, twice as long as the diameter of the shell and four
times as long as the four pyramidal polar spines; each of these six spines
with six deep furrows between the six prominent edges. The fourteen other
spines very thin, compressed, two-edged, about as long as the radius of the
shell.

_Dimensions._--Diameter of the shell 0.11; length of the two equatorial
spines 0.2, of the eighteen others 0.04 to 0.06; basal breadth of the six
larger spines 0.03, of the fourteen others 0.01.

_Habitat._--Tropical Atlantic, Station 352, surface.

Genus 377. _Hexaconus_,[417] n. gen.

_Definition._--#Hexalaspida# with twenty prominent radial spines, which are
surrounded at the base by prominent sheaths; the six hydrotomical spines
much larger than the fourteen others.

The genus _Hexaconus_ differs from the preceding _Hexalaspis_ in the
development of conical or cylindrical sheaths surrounding the basal parts
of the radial spines; these sheaths are developed sometimes around all
twenty spines, sometimes only around the six larger hydrotomical spines.

Subgenus 1. _Hexaconarium_, Haeckel.

_Definition._--Six hydrotomical spines of equal size.

1. _Hexaconus ciliatus_, n. sp. (Pl. 139, fig. 3).

All six hydrotomical spines of nearly equal size, pyramidal, with six
prominent edges, somewhat longer than the radius of the shell. Sheaths
sulcated, finely ciliated at the mouth, twice as broad as long and only
one-fourth as long as the spines. The fourteen smaller spines very thin,
bristle-shaped, shorter than the radius of the shell.

_Dimensions._--Diameter of the shell 0.16; length of the six hydrotomical
spines 0.11; basal breadth 0.03.

_Habitat._--Central Pacific, Station 270, surface.

{877}2. _Hexaconus coronatus_, n. sp. (Pl. 139, fig. 5).

All six hydrotomical spines of equal size, conical, somewhat compressed,
with two prominent edges, scarcely as long as the radius of the shell.
Sheaths crested, strongly dentated on the mouth, three times as broad as
long and only one-sixth as long as the spines. Fourteen smaller spines
about half as large as the six principal spines, of the same form, but
without coronated sheaths.

_Dimensions._--Diameter of the shell 0.2; length of the six hydrotomical
spines 0.1; basal breadth 0.02.

_Habitat._--North Pacific, Station 253, surface.

3. _Hexaconus velatus_, n. sp. (Pl. 139, fig. 6).

All six hydrotomical spines of equal size, conical, somewhat compressed,
shorter than the radius of the shell. Sheaths very large, truncated
conical, enveloping the spines almost entirely, with crested wall, only
half as broad at the constricted mouth as at the base. The fourteen smaller
spines about half as large as the six principal spines, of the same form,
but without large sheaths.

_Dimensions._--Diameter of the shell 0.12; length of the six hydrotomical
spines 0.05; basal breadth 0.01.

_Habitat._--North Atlantic, Station 354, surface.

Subgenus 2. _Hexaconidium_, Haeckel.

_Definition._--Six hydrotomical spines of unequal size, two opposite
(equatorial) much larger than the four other (polar) spines.

4. _Hexaconus serratus_, n. sp. (Pl. 139, fig. 4).

Six hydrotomical spines of unequal size, the two equatorial being as long
as the radius of the shell and one and a half times as large as the four
polar; all of the same form, triangular, compressed, with six prominent
edges. Sheaths cylindrical, half as long as the spines, with prominent
crests, and with strong serrated teeth at the distal mouth. The fourteen
smaller spines scarcely one-fourth or one-sixth as large as the six
principal spines, without prominent sheaths.

_Dimensions._--Diameter of the shell 0.2; length of the equatorial spines
0.1, basal breadth 0.05; length and breadth of the sheaths 0.05.

_Habitat._--South Pacific, Station 288, surface.

5. _Hexaconus vaginatus_, n. sp. (Pl. 139, fig. 7).

Six hydrotomical spines of unequal size; the two equatorial very stout,
pyramidal, six-edged, nearly as long as the diameter of the shell, and on
the base three to four times as broad as the eighteen other spines, which
are much thinner, all nearly of equal length, and two-edged. All
{878}twenty spines are provided with prominent basal sheaths, which are
truncate conical, sulcate, and dentate on the narrowed distal mouth. The
sheaths of the six hydrotomical spines are twice to three times as large as
those of the fourteen smaller spines.

_Dimensions._--Diameter of the shell 0.15; length of the spines 0.12; basal
breadth of the equatorial spines 0.04, of the other spines 0.02; length of
the hydrotomical sheaths 0.05, of the other sheaths 0.02.

_Habitat._--Central Pacific, Station 274, surface.

6. _Hexaconus echinatus_, n. sp. (Pl. 140, fig. 12).

Six hydrotomical spines of unequal size; the two equatorial, and their
sheaths twice as large as the four polar spines. These six spines are
six-edged, pyramidal, and their basal half enveloped by very large conical
sheaths which are sulcate, and twice as broad on the dentate distal mouth
as on the narrower base. The other fourteen spines are very thin,
two-edged, half as long, with low sheaths. Approaches some forms of
_Diploconus_.

_Dimensions._--Diameter of the shell 0.15; length of the equatorial spines
0.13, of the polar spines 0.08.

_Habitat._--Indian Ocean, Belligemma, Ceylon, Haeckel, surface.

Genus 378. _Hexonaspis_,[418] n. gen.

_Definition._--#Hexalaspida# with six prominent radial spines (in the
hydrotomical plane) which are not surrounded by prominent sheaths; the
fourteen other spines quite rudimentary, not prominent.

The genus _Hexonaspis_ and the following _Hexacolpus_ differ from the two
preceding genera in the rudimentary shape of the fourteen reduced and
stunted smaller spines; these are only developed inside the shell, and are
not prominent outside over its surface.

Subgenus 1. _Hexonasparium_, Haeckel.

_Definition._--Six hydrotomical spines of equal size.

1. _Hexonaspis heliosestrum_, n. sp.

All six hydrotomical spines of nearly equal size, isosceles triangular,
compressed, smooth, about as long as the diameter of the shell and three
times as long as broad at the base. This species is very similar to
_Hexalaspis heliodiscus_ (Pl. 139, fig. 2), but differs in the larger size
of the six marginal spines and in the complete external absence of the
fourteen smaller spines.

{879}_Dimensions._--Diameter of the shell 0.13; length of the six marginal
spines 0.12, basal breadth 0.04.

_Habitat._--Tropical Atlantic, Station 348, surface.

2. _Hexonaspis hexapleura_, n. sp. (Pl. 140, fig. 15).

All six hydrotomical spines of nearly equal size, triangular, compressed,
six-edged, about as long as the radius of the shell and twice as long as
broad on the base. Two strong prominent ribs on each flat side of the
spines. Shell very thick-walled, with a very small cavity.

_Dimensions._--Diameter of the shell 0.11; length of the six marginal
spines 0.06, basal breadth 0.03.

_Habitat._--South Atlantic, Station 332, surface.

3. _Hexonaspis hexagona_, n. sp.

All six hydrotomical spines of equal size, very short, triangular, only
half as long as broad at the base. The whole shell accordingly forms a
regular hexagon, the six corners of which are formed by the distal points
of the rudimentary spines, the six sides by their straight lateral edges.
Shell very dark.

_Dimensions._--Diameter of the shell 0.2; length of the six marginal spines
0.03, basal breadth 0.06.

_Habitat._--South Atlantic (east of Patagonia), Station 318, surface.

Subgenus 2. _Hexonaspidium_, Haeckel.

_Definition._--Six hydrotomical spines of unequal size, two opposite
(equatorial) much larger than the four other (polar) spines.

4. _Hexonaspis hastata_, n. sp. (Pl. 140, fig. 16).

Six hydrotomical spines of unequal size; two larger equatorial spines with
six prominent wings, about as long as the radius of the shell and half as
broad at the base; the four polar spines quite as broad, but only half as
long, nearly equilateral triangular, with two shallow furrows on each flat
side at the broader base. The fourteen smaller spines not visible on the
surface, quite rudimentary. Crests of the surface elegantly denticulated.

_Dimensions._--Diameter of the shell 0.14; length of the two equatorial
spines 0.08, of the four polar spines 0.04; basal breadth 0.04.

_Habitat._--Central Pacific, Station 271, surface.

{880}Genus 379. _Hexacolpus_,[419] n. gen.

_Definition._--#Hexalaspida# with six prominent radial spines (in the
hydrotomical plane), which are surrounded by prominent sheaths at the base;
the fourteen other spines quite rudimentary, not prominent.

The genus _Hexacolpus_ differs from the preceding _Hexonaspis_ in the
development of conical or cylindrical sheaths around the basal parts of the
radial spines. It bears to the latter the same relation as _Hexaconus_
exhibits to _Hexalaspis_.

Subgenus 1. _Hexacolparium_, Haeckel.

_Definition._--Six hydrotomical spines all nearly of equal size.

1. _Hexacolpus nivalis_, n. sp. (Pl. 139, fig. 1).

_Hexalaspis nivalis_, Haeckel, 1882, Manuscript et Atlas.

All six hydrotomical spines of nearly equal size and equidistant, thin,
lanceolate, compressed, somewhat shorter than the diameter of the hexagonal
shell. Sheaths nearly prismatic, conical in the distal third, each with
twelve prominent parallel crests, which are separated by twelve deep
furrows; their contracted distal opening or mouth denticulate. The sheaths
are as long as the radius of the shell, and envelop two-thirds of the
spines. (Resembles certain forms of snow-crystals.)

_Dimensions._--Diameter of the shell 0.15, thickness of its wall 0.08;
length of the spines 0.1, of the sheath 0.007; breadth of the latter 0.04.

_Habitat._--West Tropical Pacific, Station 225, surface.

2. _Hexacolpus conifer_, n. sp.

All six hydrotomical spines of equal size and equidistant, triangular,
compressed, about as long as the radius of the circular shell. Sheaths
conical, sulcate, half as broad on the serrate distal end as on the base.
The sheaths envelop the basal half of the spines.

_Dimensions._--Diameter of the shell 0.18; length of the spines 0.1, of the
sheaths 0.06; basal breadth of the latter 0.07, distal breadth 0.04.

_Habitat._--North Pacific, Station 236, surface.

Subgenus 2. _Hexacolpidium_, Haeckel.

_Definition._--Six hydrotomical spines of unequal size, two opposite
(equatorial) much larger than the four other (polar) spines.

{881}3. _Hexacolpus trypanon_, n. sp. (Pl. 140, fig. 11).

Six hydrotomical spines of unequal size, two equatorial about as long as
the diameter of the shell and nearly twice as large as the four polar; all
of the same form, quadrangular prismatic, with prominent edges and pointed
distal ends. Sheaths six-sided prismatic, with strong prominent edges and
thinner parallel ribs between them; their distal mouth with six strong
denticulated teeth. The sheaths envelop two-thirds of the spines, and are
as broad as the radius of the shell.

_Dimensions._--Diameter of the shell 0.13; length of the equatorial spines
0.12, of their sheaths 0.08; breadth of the latter 0.06.

_Habitat._--South Pacific, Station 288, surface.

4. _Hexacolpus dodecodus_, n. sp.

Six hydrotomical spines of unequal size, two equatorial about as long as
the radius of the shell and twice as large as the four polar; all of the
same form, triangular, compressed, sulcated. Sheaths prismatic, with twelve
prominent, parallel edges, and twelve strong serrated triangular teeth on
the mouth. The sheaths envelop the basal half of the spines. This species
greatly resembles _Hexaconus serratus_ (Pl. 139, fig. 4), but differs in
the larger size of the equatorial spines and the complete reduction of the
fourteen smaller spines.

_Dimensions._--Diameter of the shell 0.2; length of the equatorial spines
0.1, of their sheaths 0.06; breadth of the latter 0.08.

_Habitat._--North Pacific, Station 248, surface.

5. _Hexacolpus infundibulum_, n. sp. (Pl. 140, fig. 10).

Six hydrotomical spines of unequal size, two equatorial about twice as long
as the diameter of the shell and as the four polar spines; all six spines
of the same form, quadrangular prismatic, pointed at the distal pyramidal
end. Sheaths conical, two to three times as broad at the denticulate distal
mouth as at the narrow base, sulcate; the mantle of the cone concavely
vaulted. The large sheaths envelop two-thirds or three-fourths of the
spines.

_Dimensions._--Diameter of the shell 0.1 to 0.13; length of the equatorial
spines 0.15 to 0.2, of their sheaths 0.1 to 0.15; basal breadth of the
latter 0.04, distal breadth 0.12.

_Habitat._--Central Pacific, Station 272, surface.

Family XLIV. #DIPLOCONIDA#, Haeckel (Pl. 140).

_Diploconida_, Haeckel, 1862, Monogr. d. Radiol., p.404.

_Definition._--ACANTHARIA with simple diploconical shell, composed of two
very large equatorial spines which are opposite in the hydrotomical axis,
are surrounded by conical or cylindrical, often compressed sheaths, and
arise from a small central {882}lattice-shell. Eighteen other spines
(disposed according to the Müllerian law of Icosacantha) much smaller,
often rudimentary. Central capsule ellipsoidal or diploconical.

The family #Diploconida#, founded by me in 1862 for a single Mediterranean
species (_Diploconus fasces_), appears to be the most aberrant and strange
form among the ACANTHARIA. As I had met with only a single specimen, very
dark and intransparent in its central part, my observations on its
structure were imperfect and the explanation of it partly erroneous
(compare my Monograph, pp. 46, 404, Taf. xx. figs. 7, 8). However, I
regarded _Diploconus_ as the representative of quite a peculiar family,
derived from the Acanthometrida, and I correctly compared the large
opposite radial spines of one equatorial axis with the corresponding parts
in _Amphilonche_.

Afterwards Richard Hertwig observed some specimens of _Diploconus fasces_
in the same locality (Messina), and gave an accurate description of its
central capsule, including numerous small nuclei (1879, Organismus d.
Radiol., p. 28, Taf. ii. fig. 3). He found also that the peculiar
diploconical skeleton is not composed of silex, but of acanthin. In the
explanation of the shell-structure he adopted my opinion.

In the rich collections of the Challenger I detected ten different forms of
Diploconida, all very rare, and for the most part represented only by
single specimens. A twelfth species was found by me in the collection of
Captain Rabbe from the Indian Ocean. By the study of these new forms, and
particularly by their comparison with the most nearly allied Hexalaspida
and Belonaspida, it was possible for me to correct some errors in my former
description and to give a much more correct description and natural
explanation of this very peculiar and strange family of Radiolaria (compare
Pl. 140).

The most characteristic and the most voluminous part of the acanthinic
skeleton in all Diploconida appears as the diploconical or nearly
cylindrical solid "mantle" giving them their name and odd appearance (Pl.
140). Usually this mantle is broader on its two opposite terminal openings
than in its more or less constricted middle part. This latter is now more
spherical or ellipsoidal, now more lenticular, and usually separated from
the two cones by two slight transverse strictures. On the surface of this
middle part twelve to eighteen radial spines, which in _Diplocolpus_ are
rudimentary or absent, are visible in _Diploconus_. The longitudinal axis
of this shell is constantly occupied by a very large pair of opposite stout
prismatic or cylindrical principal spines, which are united in the centre
and usually more or less prominent with their distal apex over the two
openings of the double cone.

In my first communication on the _Diploconus_ (1862, _loc. cit._) I
correctly compared these two large spines in the prolonged main axis of the
shell to the principal equatorial spines of _Amphilonche_ (or to the
"hydrotomical spines," _c_1, _c_3); but my explanation of the two peculiar
cones enveloping them was erroneous. I supposed at that time that they were
formed by the eight flattened and leaf-shaped curved {883}tropical spines,
so that around each principal spine the four neighbouring tropical spines
(two of the northern and two of the southern hemisphere) had grow together
by their edges and formed the peculiar conical sheath. I can now say that
this opinion (afterwards adopted also by Hertwig) was quite erroneous, the
two conical or funnel-shaped sheaths being the enlarged basal sheaths of
the two hydrotomical spines, which we have already seen in the Hexalaspida
(Pl. 139). But whilst in these latter all six principal spines of the
hydrotomical meridian plane are hypertrophied (two equatorial and four
polar spines), in the nearly allied Diploconida only the two opposite
equatorial spines are developed, whilst all other eighteen spines are more
or less atrophied or quite rudimentary. In _Diploconus_ the latter are more
or less evident, whilst in _Diplocolpus_ they disappear externally.

The true lattice-shell of the #Acanthophracta# (constantly composed of the
meeting apophyses of twenty radial spines) is therefore represented in the
Diploconida by the small roundish middle part of the whole shell, which is
usually much smaller than the two opposite cones, and separated from them
by the two slight transverse strictures. Usually this small but most
important middle part of the shell is very dark and opaque, on account of
its very thick wall and small pores; but in some species it is clear enough
to ascertain that the structure of this lattice-shell is the same as in the
lenticular Hexalaspida, there being a network of thick crests on the outer
surface and small pores in the dimples between them. Indeed, in many (and
probably in all) Diploconida the forty aspinal pores are present which we
found in all Hexalaspida, Belonaspida, and Diporaspida, so that these four
families of #Acanthophracta# represent one continuous phylogenetical
series; _Phractaspis_ among the Diporaspida is at the beginning, and
_Diplocolpus_ among the Diploconida at the end of this remarkably
transformed morphological series.

The twenty radial spines in all Diploconida are probably united very firmly
(or even perfectly grown together) in the centre of the small thick-walled
lattice-shell, the inner space of which is extremely reduced. Probably,
too, the sutures between the meeting apophyses of the thick radial spines
are often (or even constantly) obliterated by concrescence, so that the
whole shell forms a single piece of acanthin. But I regret that I cannot
ascertain these and other points in the structure of the shell, as the
small number of specimens observed did not permit an anatomical examination
to be made. I have no doubt, however, that the structure of the whole of
the middle main part of the shell is quite the same as in the lenticular
shell of the thick-walled Hexalaspida, and that in both families each of
the twenty radial spines bears originally only two opposite apophyses.

The characteristic mantle of the double cone of the Diploconida, or the
basal sheath of their two large, perfectly developed principal spines is
usually much larger than the shell itself, and more or less compressed from
both poles of the shortened geotomical axis. Therefore the transverse
section of the two cones is usually elliptical, more {884}rarely circular.
Their widest part is generally the distal opening; more rarely this is a
little constricted. The thin transparent lamella of acanthin, representing
the mantle of the double cone, is commonly ribbed or furrowed by
longitudinal, parallel or divergent crests, and elegantly denticulated on
the edge of the distal opening.

The two conical or cylindrical halves of the mantle are connected with the
two enclosed principal spines not only at the base, where they arise from
the small central lattice-shell, but also throughout a certain part of
their length, by means of two, four, or six wings or leaves, which lie
opposite and in pairs in the meridian planes of those spines. These
meridian wings are more or less triangular (with broader concave outer
bases), and connected by their axial edge with the spine and by their
peripheral edge with the mantle. They separate two, four, or six conical
spaces or pyramidal compartments in each cone. But these aspinal
compartments and the separating septa are not new productions of the
Diploconida, but are inherited from their ancestral family, the Hexalaspida
(compare above, p. 873).

The eighteen smaller spines in _Diploconus_ are either of nearly equal size
or more or less differentiated. The eight tropical spines are often much
larger than the eight polar spines. The two geotomical spines, (or the two
opposite equatorial spines of the shortened geotomical axis) are often
quite rudimentary. In _Diplocolpus_ the external part (outside the shell)
is in all eighteen smaller spines rudimentary or atrophied.

_The Central Capsule_, as shown by Hertwig, contains numerous small nuclei,
and is divided into three parts by the above named two transverse
strictures; the smaller central part (in the original lenticular
lattice-shell) and the two opposite larger parts, filling up the greater
part of the two conical or cylindrical sheaths, and more or less adopting
their form. Corresponding to the shell itself the enclosed capsule is often
more or less flattened, being compressed at both poles of the geotomical
axis. The pseudopodia seem to proceed only from the two large polar
apertures of the sheaths, and form therefore two opposite conical tufts or
bunches.

_Synopsis of the Genera of Diploconida._

All twenty spines more or less developed (sometimes
eight of them rudimentary), 380. _Diploconus_.

Only the two hydrotomical spines developed (all the
eighteen others rudimentary), 381. _Diplocolpus_.

Genus 380. _Diploconus_,[420] Haeckel, 1862, Monogr. d. Radiol., p. 404.

_Definition._--#Diploconida# with two very large spines (opposite in the
hydrotomical axis) and ten to eighteen other much smaller spines externally
visible.

{885}The genus _Diploconus_ must be derived from _Hexacolpus_ (among the
_Hexalaspida_) by the stronger development of the two hydrotomical spines
and their large sheaths; all the other eighteen spines are much smaller,
and usually devoid of prominent sheaths. Sometimes the eight tropical
spines are rudimentary.

Subgenus 1. _Diploconulus_, Haeckel.

_Definition._--Mantle of the double cone not compressed; its transverse
section therefore circular.

1. _Diploconus amalla_, n. sp. (Pl. 140, fig. 1).

Mantle of the double cone thick walled, not compressed, its transverse
section circular; its contour little convex; its surface nearly smooth; its
distal margin regularly denticulated. Diameter of its mouth twice as large
as the equatorial diameter of the shell and one-third as long as its total
length. The two large spines prismatic, one-fourth longer than their
conical sheath. The eighteen smaller spines very thin, about half as long
as the former, a little curved.

_Dimensions._--Length of the shell 0.3; equatorial breadth 0.05, polar
breadth 0.1.

_Habitat._--Central Pacific, station 271, surface.

2. _Diploconus cyathiscus_, n. sp. (Pl. 140, fig. 3).

Mantle of the double cone thin walled, not compressed; its transverse
section circular; its contour strongly convex; its surface with six
stronger and many smaller ribs; its margin with numerous straight and long,
parallel denticles. Diameter of its mouth half as long as the whole shell
and one and a half times as long as its equatorial diameter. The two
principal spines one-fourth longer than their conical sheath. The eighteen
smaller spines about half as long, thin, straight. Both geotomical spines
thick and short.

_Dimensions._--Length of the shell 0.2; equatorial breadth 0.07, polar
breadth 0.1.

_Habitat._--West Tropical Pacific, Station 225, surface.

3. _Diploconus cylindrus_, n. sp.

Mantle of the double cone thick walled, dark, cylindrical, not compressed;
its transverse section circular; its contours straight, parallel; its
surface with strong parallel straight longitudinal ribs; its margin
irregularly dentated. Diameter of its mouth one-fifth as long as the whole
shell and two-thirds as long as the diameter of the equatorial
intumescence. The two principal spines prismatic, nearly twice as long as
their cylindrical sheath. The smaller spines about half as long, thin,
conical, straight. (Resembles the medial part of _Hexacolpus trypanon_, Pl.
140, fig. 11.)

_Dimensions._--Length of the shell 0.25; equatorial breadth 0.08, polar
breadth 0.05.

_Habitat._--North Pacific, Station 241, surface.

{886}Subgenus 2. _Diploconium_, Haeckel.

_Definition._--Mantle of the double cone compressed from both poles of the
shortened geotomical axis; its transverse section therefore elliptical.

4. _Diploconus fasces_, Haeckel.

_Diploconus fasces_, Haeckel, 1862, Monogr. d. Radiol., p. 405, Taf. xx.
figs. 7, 8.

_Diploconus fasces_, R. Hertwig, 1879, Organismus d. Radiol., p. 28, Taf.
ii. fig. 3.

Mantle of the double cone compressed, with elliptical transverse section;
its contour straight; its surface with numerous longitudinal furrows; its
margin finely denticulated. Diameter of its mouth two-fifths as long as the
whole shell and three times as broad as its equatorial diameter. The two
main spines one-third longer than their conical sheath, four-edged. The
eighteen smaller spines thick and short, cylindrical or a little
compressed.

_Dimensions._--Length of the shell 0.18; equatorial breadth 0.025, polar
breadth 0.07.

_Habitat._--Mediterranean (Messina), surface.

5. _Diploconus cotyliscus_, n. sp. (Pl. 140, fig. 4).

Mantle of the double cone compressed, thin walled, with elliptical
transverse section; its contour convex; surface and margin nearly smooth.
Diameter of its mouth half as long as the whole shell and twice as long as
its equatorial diameter. The two main spines scarcely longer than their
hemispherical sheath. The smaller spines conical, straight, nearly of the
same length.

_Dimensions._--Length of the shell 0.3; equatorial breadth 0.08, polar
breadth 0.15.

_Habitat._--Tropical Atlantic, Station 347, surface,

6. _Diploconus saturnus_, Haeckel.

_Diploconus saturnus_, Haeckel, 1879, Natürl. Schöpfungsgesch., Aufl.
vii. p. 706, Taf. xvi. fig. 11.

Mantle of the double cone compressed, thick walled, with elliptical
transverse section; its contour convex; its surface with numerous deep and
irregular longitudinal furrows and marginal incisions. Diameter of its
mouth one-third as long as the whole shell and twice as broad as its
equatorial diameter. The two hydrotomical spines short, scarcely longer
than their sheath. The eighteen other spines thin, cylindrical, nearly of
the same length, curved. (Resembles a sheaf.)

_Dimensions._--Length of the shell 0.24; equatorial breadth 0.04, polar
breadth 0.07.

_Habitat._--South Pacific, Station 300, surface.

7. _Diploconus hexaphyllus_, n. sp. (Pl. 140, fig. 2).

Mantle of the double cone compressed, thick walled, with elliptical
transverse section; its contour straight; surface with six stronger and
numerous smaller straight ribs and six marginal {887}denticulate incisions.
Diameter of its mouth one-third as long as the whole shell and somewhat
smaller than the equatorial diameter. Hydrotomical spines about twice as
long as their sheath, pyramidal, with emarginate point and six wings.
Geotomical spines short and strong, pyramidal. Sixteen other spines thin,
straight, conical.

_Dimensions._--Length of the shell 0.3; equatorial breadth 0.11, polar
breadth 0.1.

_Habitat._--Central Pacific, Station 266, surface.

Genus 381. _Diplocolpus_,[421] n. gen.

_Definition._--#Diploconida# with only two developed, very large spines
(opposite in the hydrotomical axis); all the other eighteen spines quite
rudimentary or externally atrophied.

The genus _Diplocolpus_ is the last and the most modified genus among the
ACANTHARIA. The eighteen smaller spines of _Diploconus_ are here quite
rudimentary or have even perfectly disappeared by atrophy, so that the
shell seems to consist only of the large hydrotomical spines and their
enveloping sheaths.

Subgenus 1. _Diplocolpulus_, Haeckel.

_Definition._--Mantle of the double cone not compressed; its transverse
section therefore circular.

1. _Diplocolpus costatus_, n. sp. (Pl. 140, fig. 7).

Mantle of the double cone thin walled, not compressed; its transverse
section circular or nearly hexagonal; its contour little convex; surface
with six thick longitudinal equidistant ribs and many thinner ribs between
them; margin denticulate. Transverse diameter of the shell one-third as
long as the longitudinal. The two hydrotomical spines thick and short,
little prominent over the polar mouth. No rudiments of other spines visible
externally.

_Dimensions._--Length of the shell 0.15; equatorial and polar breadth 0.05.

_Habitat._--Indian Ocean (Madagascar), Rabbe, surface.

2. _Diplocolpus cristatus_, n. sp. (Pl. 140, fig. 6).

Mantle of the double cone thick walled, not compressed; its transverse
section circular or hexagonal; its contour nearly straight; surface with
six thick prominent longitudinal equidistant ribs; margin irregularly
dentated. Transverse diameter of the shell half as long as the
longitudinal. {888}The two hydrotomical spines very thick and short,
pyramidal, little prominent. Short rudiments of the eighteen other spines
present, bristle-shaped.

_Dimensions._--Length of the shell 0.1, breadth 0.05.

_Habitat._--South Atlantic, Station 335, surface.

Subgenus 2. _Diplocolpium_, Haeckel.

_Definition._--Mantle of the double cone compressed from both poles of the
shortened geotomical axis; its transverse section therefore elliptical.

3. _Diplocolpus serratus_, n. sp. (Pl. 140, fig. 5).

Mantle of the double cone thick walled, compressed, with elliptical
transverse section; its surface with numerous delicate ribs; contour little
convex; margin regularly serrate. Hydrotomical spines short, pyramidal,
with six serrate wings. Transverse diameter of the shell nearly twice as
great at the polar mouth as at the equator and half as long as the whole
shell.

_Dimensions._--Length of the shell 0.15; breadth on the equator 0.035, on
the poles 0.07.

_Habitat._--Central Pacific, Station 274, surface.

4. _Diplocolpus dentatus_, n. sp. (Pl. 140, fig. 9).

Mantle of the double cone thin walled, compressed, with elliptical
transverse section; its contour straight; surface with six deeper and
numerous shallower furrows; margin denticulated, with six triangular
prominent larger teeth. Hydrotomical spines very large prismatic, pyramidal
at the ends. Transverse diameter of the shell a little larger at the polar
mouth than at the equator and half as long as the whole shell.

_Dimensions._--Length of the shell 0.2; breadth on the equator 0.08, on the
poles 0.1.

_Habitat._--North Pacific, Station 244, surface.

5. _Diplocolpus sulcatus_, n. sp. (Pl. 140, fig. 8).

Mantle of the double cone thin walled, compressed, with elliptical
transverse section; its contour straight; surface with twelve to sixteen
deep longitudinal furrows, which are separated by thin, undulating, double
edged ribs; margin little dentated. Hydrotomical spines short, pyramidal,
with six dentate wings. Transverse diameter of the shell a little larger at
the polar mouth than at the equator, about one-third of the whole length of
the shell.

_Dimensions._--Length of the shell 0.17; breadth on the equator 0.05, on
the poles 0.06.

_Habitat._--South Pacific, Station 291, surface.

Notes.

[1] The numbers preceded by L. N. refer to the list of names of authors
in the Bibliography on p. clxxvi.

[2] _Sitzungsb. med.-nat. Gesellsch. Jena_, February 16, 1883.

[3] _Loc. cit._

[4] _Abhandl. d. k. Akad. d. Wiss. Berlin_, 1858, p. 28.

[5] Monogr. d. Radiol., 1862, Taf. ii. p. 253.

[6] Histologie der Radiolarien, pp. 43-73, Taf. iii.-v.

[7] _Sitzungsb. med.-nat. Gesellsch. Jena_, February 16, 1883.

[8] _Actissa_ = Radiant, [Greek: aktis].

[9] _Thalassolampe_ = Sea-scum; [Greek: thalassa] and [Greek: lampê].

[10] _Thalassopila_ = Sea-ball; [Greek: thalassa], [Greek: pila].

[11] _Thalassicolla_ = Sea-jelly, [Greek: thalassa], [Greek: kolla].

[12] _Thalassophysa_ = Sea-bladder; [Greek: thalassa], [Greek: physa.]

[13] _Collozoum_ = Jelly-animal; [Greek: kolla], [Greek: zôon].

[14] _Thalassosphæra_ = Sea-sphere; [Greek: thalassa], [Greek: sphaira].

[15] Atlantic, vol. i. p. 233, fig. 51, 1877.

[16] _Thalassoxanthium_ = Sea-burdock; [Greek: thalassa], [Greek:
xanthion.]

[17] _Physematium_ = Small vesicle; [Greek: Physêmation].

[18] _Thalassoplancta_ = Floating on the sea; [Greek: thalassoplankta].

[19] Compare _Thalassoplancta cavispicula_, Monogr. d. Radiol., 1862, p.
261, Taf. iii. figs. 10-13.

[20] _Lampoxanthium_ = Scum-burdock; [Greek: lampe], [Greek: xanthion].

[21] _Loc. cit._, Taf. xxxii., xxxiii.

[22] _Belonozoum_ = Needle-animal; [Greek: belonê], [Greek: zôon].

[23] _Sphærozoum_ = Spherical animal; [Greek: sphaira], [Greek: zôon].

[24] _Rhaphidozoum_ = Needle-animal; [Greek: rhaphis], [Greek: zôon].

[25] Ethmosphærida = Liosphærida simplicia = Monosphærida anacantha.

[26] _Cenosphæra_ = Hollow sphere; [Greek: kenos], [Greek: sphaira].

[27] _Stigmosphæra_ = Sphere with central point; [Greek: stigma], [Greek:
sphaira].

[28] _Ethmosphæra_ = Sieve-sphere; [Greek: êthmos], [Greek: sphaira].

[29] _Sethosphæra_ = Sieve-sphere; [Greek: sêthos], [Greek: sphaira].

[30] _Carposphærida_ = Liosphærida duplicia = Dyosphærida anacantha.

[31] _Carposphæra_ = Fruit-shaped sphere, [Greek: karpos], [Greek:
sphaira].

[32] _Liosphæra_ = Smooth sphere; [Greek: leios], [Greek: sphaira].

[33] Thecosphærida = Liosphærida triplicia = Triosphærida anacantha.

[34] _Thecosphæra_ = Capsule-sphere; [Greek: thêkê], [Greek: sphaira].

[35] _Rhodosphæra_ = Rose-sphere; [Greek: rhodon], [Greek: sphaira].

[36] Cromyosphærida = Liosphærida tetraplicia = Tetrasphærida anacantha.

[37] _Cromyosphæra_ = Onion-sphere; [Greek: chromyon], [Greek: sphaira].

[38] Caryosphærida = Liosphærida multiplicia = Polysphærida anacantha.

[39] _Caryosphæra_ = Nut-sphere; [Greek: karyon], [Greek: sphaira].

[40] Plegmosphærida = Liosphærida spongiosa = Spongosphærida anacantha.

[41] _Styptosphæra_ = Hemp-sphere; [Greek: styptos], [Greek: sphaira].

[42] _Plegmosphæra_ = Sphere of wicker-work; [Greek: plegma], [Greek:
sphaira].

[43] _Spongoplegma_ = Spongy wickerwork; [Greek: spongos], [Greek:
plegma].

[44] _Spongodictyon_ = Spongy network; [Greek: spongos], [Greek: diktyon].

[45] _Loc. cit._, pp. 17, 55, 1858.

[46] _Abhandl. d. k. Akad. d. Wiss. Berlin_, p. 55, 1858.

[47] _Loc. cit._, p. 530, 1862.

[48] _Loc. cit._, pp. 30, 133, 1879.

[49] _Loc. cit._, p. 471, 1881.

[50] _Loc. cit._, p. 535, Taf. xxxiv. fig. 1.

[51] _Collosphæra_ = Jelly-sphere; [Greek: kolla], [Greek: sphaira].

[52] _Tribonosphæra_ = Cloak-sphere; [Greek: tribôn], [Greek: sphaira].

[53] _Pharyngosphæra_ = Throat-sphere; [Greek: pharynx], [Greek: sphaira].

[54] _Buccinosphæra_ = Trumpet-sphere; [Greek: bykanê], [Greek: sphaira].

[55] _Acrosphæra_ = Pointed-sphere; [Greek: akros], [Greek: sphaira].

[56] _Odontosphæra_ = Teeth-sphere; [Greek: odous], [Greek: sphaira].

[57] _Choenicosphæra_ = Shell with coronel trepans; [Greek: choinikê],
[Greek: sphaira].

[58] _Siphonosphæra_ = Sphere with tubes; [Greek: siphôn], [Greek:
sphaira].

[59] _Mazosphæra_ = Teat-sphere; [Greek: mazos], [Greek: sphaira].

[60] _Trypanosphæra_ = Auger-sphere; [Greek: trypanon], [Greek: sphaira].

[61] _Caminosphæra_ = Chimney-sphere; [Greek: kaminos], [Greek: sphaira].

[62] _Solenosphæra_ = Sphere with tubules; [Greek: sôlên], [Greek:
sphaira].

[63] _Otosphæra_ = Shell with ears; [Greek: otosphaira].

[64] _Coronosphæra_ = Coronal-sphere; [Greek: koronê], [Greek: sphaira].

[65] _Clathrosphæra_ = Lattice-sphere.

[66] _Xanthiosphæra_ = Burdock-sphere; [Greek: xanthion], [Greek:
sphaira].

[67] Stylosphærida = Sphæroidea dissacantha, Prodromus, p. 449.

[68] _Xiphosphæra_ = Sword-sphere; [Greek: xiphos], [Greek: sphaira].

[69] _Xiphostylus_ = Sword style; [Greek: xiphos], [Greek: stylos].

[70] _Saturnalis_ = Similar to _Saturnus_, with a ring.

[71] _Stylosphæra_ = Sphere with styles; [Greek: stylos], [Greek:
sphaira].

[72] _Sphærostylus_ = Sphere with styles; [Greek: sphaira], [Greek:
stylos].

[73] _Saturnulus_ = Small _Saturnus_, with a ring.

[74] _Amphisphæra_ = Sphere with spines on both poles; [Greek: amphi],
[Greek: sphaira].

[75] _Amphistylus_ = Shell with styles on both poles; [Greek: amphi],
[Greek: stylos].

[76] _Saturninus_ = Similar to Saturnus, with a ring.

[77] _Stylocromyum_ = Onion with styles; [Greek: stylos], [Greek:
kromyon].

[78] _Cromyostylus_ = Onion with styles; [Greek: kromyon], [Greek:
stylos].

[79] _Spongolonchis_ = Spongy shell with spears; [Greek: spongos], [Greek:
lonchis].

[80] _Spongostylus_ = Spongy shell with styles; [Greek: spongos], [Greek:
stylos].

[81] _Spongostylidium_, diminutive of _Spongostylus_.

[82] Staurostylida = Staurosphærida simplicia = Monosphærida tetracantha.

[83] _Staurosphæra_ = Cross-sphere; [Greek: stauros], [Greek: sphaira].

[84] _Staurostylus_ = Cross-style; [Greek: stauros], [Greek: stylos].

[85] _Stylostaurus_, Style-cross; [Greek: stylos], [Greek: stauros].

[86] Staurolonchida = Staurosphærida duplicia = Dyosphærida tetracantha.

[87] _Staurolonche_ = Crossed spear; [Greek: stauros], [Greek: lonchê].

[88] _Staurancistra_ = Crossed fish-hook; [Greek: stauros], [Greek:
ankistra].

[89] _Staurolonchidium_ = Diminutive from _Staurolonche_.

[90] _Stauroxiphos_ = Sword with cross; [Greek: stauros], [Greek: xiphos].

[91] Stauracontida = Staurosphærida triplicia = Triosphærida tetracantha.

[92] _Stauracontium_ = Crossed dart; [Greek: stauros], [Greek: akontion].

[93] Staurocromyida = Staurosphærida quadruplicia = Tetrasphærida
tetracantha.

[94] _Staurocromyum_ = Cross-onion; [Greek: stauros], [Greek: kromyon].

[95] _Cromyostaurus_ = Onion-cross; [Greek: kromyon], [Greek: stauros].

[96] Staurocaryida = Staurosphærida multiplicia = Polysphærida
tetracantha.

[97] _Staurocaryum_ = Cross-nut; [Greek: stauros], [Greek: karyon].

[98] Staurodorida = Staurosphærida spongiosa = Spongosphærida tetracantha.

[99] _Staurodoras_ = Crossed spear; [Greek: stauros], [Greek: doras].

[100] Hexastylida = Cubosphærida simplicia = Monosphærida hexacantha.

[101] _Hexastylus_ = Shell with six styles; [Greek: hexa], [Greek: stylos].

[102] _Hexastylarium_ = Shell with six styles; derivation from
_Hexastylus_.

[103] _Hexastylidium_ = Shell with six styles; derivation from
_Hexastylus_.

[104] Hexalonchida = Cubosphærida duplicia = Dyosphærida hexacantha.

[105] _Hexalonche_ = Shell with six spears; [Greek: hexalonchê].

[106] _Hexancistra_ = Shell with six fish-hooks; [Greek: hexankistra].

[107] _Hexaloncharium_ = Shell with six spears; derivation from
_Hexalonche_.

[108] _Hexalonchidium_ = Shell with six spears; derivation from
_Hexalonche_.

[109] Hexacontida = Cubosphærida triplicia = Triosphærida hexacantha.

[110] _Hexacontion_ = Shell with six darts; [Greek: hexakontion].

[111] _Hexadendron_ = Shell with six trees; [Greek: hexadendron].

[112] _Hexacontarium_ = Shell with six darts; derivation, from
_Hexacontion_.

[113] Hexacromyida = Cubosphærida quadruplicia = Tetrasphærida hexacantha.

[114] _Hexacromyum_ = Onion with six rays; [Greek: hexa], [Greek: kromyon].

[115] Hexacaryida = Cubosphærida multiplicia = Polysphærida hexacantha.

[116] _Cubosphæra_ = Sphere with three axes of a cubus; [Greek: kubos],
[Greek: sphaira].

[117] _Hexacaryum_ = Nut with six spines; [Greek: hexa], [Greek: karyon].

[118] Hexadorida = Cubosphærida spongiosa = Spongosphærida hexacantha.

[119] _Cubaxonium_ = Shell with three axes like those of a cube; [Greek:
kubos], [Greek: axônion].

[120] _Hexadoras_ = Shell with six spears; [Greek: hexadoras].

[121] _Hexadoridium_ = Shell with six small spears; derivation from
_Hexadoras_.

[122] _Acanthosphæra_ = Spiny sphere; [Greek: akantha], [Greek: sphaira].

[123] _Heliosphæra_ = Sun sphere; [Greek: hêlios], [Greek: sphaira].

[124] _Conosphæra_ = Sphere with cones; [Greek: kônos], [Greek: sphaira].

[125] _Coscinomma_ = Sieve eye; [Greek: koskinon], [Greek: omma].

[126] _Cladococcus_ = Nucleus with branches; [Greek: klados], [Greek:
kokkos].

[127] _Elaphococcus_ = Shell with harts' horn-shaped spines; [Greek:
elaphos], [Greek: kokkos].

[128] Haliommida = Astrosphærida duplicia = Dyosphærida polycantha.

[129] _Haliomma_ = Sea-eye; [Greek: hals], [Greek: omma].

[130] _Heliosoma_ = Sun-body; [Greek: hêlios], [Greek: sôma].

[131] _Elatomma_ = Pine-eye; [Greek: elatê], [Greek: omma].

[132] _Leptosphæra_ = Delicate sphere; [Greek: leptos], [Greek: sphaira].

[133] _Diplosphæra_ = Double sphere; [Greek: diploos], [Greek: sphaira].

[134] _Drymosphæra_ = Woody sphere; [Greek: drymos], [Greek: sphaira].

[135] _Astrosphæra_ = Star-sphere; [Greek: astron], [Greek: sphaira].

[136] Actinommida = Astrosphærida triplicia = Triosphærida polyacantha.

[137] _Actinomma_ = Radiant eye; [Greek: aktis], [Greek: omma].

[138] _Echinomma_ = Urchin-eye; [Greek: echinos], [Greek: omma].

[139] _Pityomma_ = Pine-eye; [Greek: pitys], [Greek: omma].

[140] Cromyommida = Astrosphærida quadruplicia = Tetrasphæria polyacantha.

[141] _Cromyomma_ = Onion-eye; [Greek: kromyon], [Greek: omma].

[142] _Cromyechinus_ = Onion-shaped Urchin; [Greek: kromyon], [Greek:
echinos].

[143] _Cromyodrymus_ = Onion with trees; [Greek: kromyon], [Greek: drymos].

[144] Caryommida = Astrosphærida multiplica = Polysphæria polyacantha.

[145] _Caryomma_ = Nut-eye; [Greek: karyon], [Greek: omma].

[146] _Arachnopila_ = Cobweb ball; [Greek: arachnê], [Greek: pilos].

[147] _Arachnopegma_ = Cobweb-building; [Greek: arachnê], [Greek: pêgma].

[148] _Arachnosphæra_ = Cobweb-sphere; [Greek: arachnê], [Greek: sphaira].

[149] Spongiommida = Astrosphærida spongiosa = Spongosphærida polyacantha.

[150] _Spongiomma_ = Spongy-eye; [Greek: spongia], [Greek: omma].

[151] _Spongodrymus_ = Spongy wood; [Greek: spongos], [Greek: drymos].

[152] _Spongechinus_ = Spongy Urchin; [Greek: spongos], [Greek: echinos].

[153] _Spongothamnus_ = Spongy shrub; [Greek: spongos], [Greek: thamnos].

[154] _Spongopila_ = Spongy ball; [Greek: spongos], [Greek: pilos].

[155] _Rhizoplegma_ = Root-shaped wicker-work; [Greek: rhiza], [Greek:
plegma].

[156] _Lychnosphæra_ = Lantern-sphere; [Greek: lychnos], [Greek: sphaira].

[157] _Centrocubus_ = Shell with a central cube; [Greek: kentron], [Greek:
kubos].

[158] _Octodendron_ = Shell with eight trees; [Greek: oktô], [Greek:
dendron].

[159] _Spongosphæra_ = Spongy sphere; [Greek: spongos], [Greek: sphaira].

[160] _Rhizosphæra_ = Root-sphere; [Greek: rhiza], [Greek: sphaira].

[161] _Cenellipsis_ = Hollow ellipsoid; [Greek: kenos], [Greek: elleipsis].

[162] _Axellipsis_ = Ellipsoid with an axial beam; [Greek: axis], [Greek:
elleipsis].

[163] _Ellipsidium_ = Small ellipsoid; [Greek: elleipsidion].

[164] _Ellipsoxiphus_ = Ellipsoid with swords; [Greek: elleipsis], [Greek:
xiphos].

[165] _Axoprunum_ = Plum with axis; [Greek: axis], [Greek: prounon].

[166] _Ellipsostylus_ = Ellipsoid with styles; [Greek: elleipsis], [Greek:
stylos].

[167] _Lithomespilus_ = Siliceous medlar; [Greek: lithos], [Greek:
mespilos].

[168] _Lithapium_ = Siliceous pear; [Greek: lithos], [Greek: apion].

[169] _Pipettella_ = Small pipette.

[170] _Druppula_ = Small drupe or [Greek: druppa], ripe Olive.

[171] _Druppocarpus_ = Olive-fruit; [Greek: druppa], [Greek: karpos].

[172] _Prunulum_ = Little-plum.

[173] _Prunocarpus_ = Plum fruit; [Greek: prounon], [Greek: karpos].

[174] _Cromyodruppa_ = Onion-olive; [Greek: kromyon], [Greek: druppa].

[175] _Cromyocarpus_ = Onion fruit; [Greek: kromyon], [Greek: karpos].

[176] _Lithatractus_ = Spindle of silex; [Greek: lithos], [Greek:
atraktos].

[177] _Druppatractus_ = Spindle-like olive; [Greek: druppa], [Greek:
atraktos].

[178] _Stylatractus_ = Spindle with styles; [Greek: stylos], [Greek:
atraktos].

[179] _Xiphatractus_ = Sword-spindle; [Greek: xiphos], [Greek: atraktos].

[180] _Cromyatractus_ = Onion-spindle; [Greek: kromyon], [Greek: atraktos].

[181] _Pipetta_ = Small pipe.

[182] _Pipettaria_ = _Pipetta_-like.

[183] _Spongellipsis_ = Spongy ellipsoid; [Greek: spongos], [Greek:
elleipsis].

[184] _Spongurus_ = Spongy tail; [Greek: spongos], [Greek: oura].

[185] _Spongocore_ = Spongy puppet; [Greek: spongos], [Greek: korê].

[186] _Spongoprunum_ = Spongy plum; [Greek: spongos], [Greek: prounon].

[187] _Spongodruppa_ = Spongy olive-fruit; [Greek: spongos], [Greek:
druppa].

[188] _Spongatractus_ = Spongy-spindle; [Greek: spongos], [Greek:
atraktos].

[189] _Spongoliva_ = Spongy olive; [Greek: spongos], [Greek: eleia].

[190] _Spongoxiphus_ = Spongy sword; [Greek: spongos], [Greek: xiphos].

[191] _Artiscus_ = Small loaf; [Greek: artiskos].

[192] _Stylartus_ = Bread with styles; [Greek: stylos], [Greek: artos].

[193] _Cannartus_ = Leaf with tubes; [Greek: kanna], [Greek: artos].

[194] _Cyphanta_ = [Greek: Kyphanta], Mediterranean port in Laconia.

[195] _Cyphonium_ = Roundish vessel; [Greek: kyphônion].

[196] _Cypassis_ = [Greek: kypassis]; A girl's girdle.

[197] _Cyphocolpus_ = Vaulted-bosom; [Greek: kyphos], [Greek: kolpos].

[198] _Cyphinus_ = [Greek: kyphinos]; derivation from [Greek: kyphos] =
roundish vessel, a kind of fruit.

[199] _Cyphinidium_ = Small Cyphinus; [Greek: kyphinidion].

[200] _Cannartiscus_ = Small loaf with tubes; [Greek: kanna], [Greek:
artiskos].

[201] _Cannartidium_ = Small loaf with tubules; [Greek: kanna], [Greek:
artidion].

[202] _Panartus_ = quite bread-like; [Greek: panartos].

[203] _Peripanartus_ = _Panartus_ with mantle; [Greek: peripanartos].

[204] _Panicium_ = A kind of bread.

[205] _Peripanicium_ = _Panicium_ with mantle.

[206] _Panarium_ = Bread-basket.

[207] _Peripanarium_ = _Panarium_ (or bread-basket) with mantle.

[208] _Ommatocampe_ = Caterpillar with eyes; [Greek: omma], [Greek: kampê].

[209] _Ommatartus_ = Loaf with eyes; [Greek: omma], [Greek: artos].

[210] _Desmocampe_ = Chain-like caterpillar; [Greek: desmos], [Greek:
kampê].

[211] _Desmartus_ = Chain-like loaf; [Greek: desmos], [Greek: artos].

[212] _Zygocampe_ = Caterpillar with paired joints; [Greek: zygon], [Greek:
kampê].

[213] _Zygartus_ = Loaf with paired joints; [Greek: zygon], [Greek: artos].

[214] _Cenodiscus_ = Hollow disk; [Greek: kenos], [Greek: diskos].

[215] _Zonodiscus_ = Disk with girdle; [Greek: zônê], [Greek: diskos].

[216] _Stylodiscus_ = Disk with styles; [Greek: stylos], [Greek: diskos].

[217] _Theodiscus_ = Divine disk; [Greek: theos], [Greek: diskos].

[218] _Crucidiscus_ = Disk with cross.

[219] _Trochodiscus_ = Wheel-disk; [Greek: trochos], [Greek: diskos].

[220] _Sethodiscus_ = Sieve-disk; [Greek: sêthos], [Greek: diskos].

[221] _Phacodiscus_ = Lenticular disk; [Greek: phakos], [Greek: diskos].

[222] _Periphæna_ = Shell with transparent girdle; [Greek: periphaina].

[223] _Perizona_ = Shell with surrounding girdle; [Greek: peri], [Greek:
zônê].

[224] _Sethostylus_ = Sieve with styles; [Greek: sêthos], [Greek: stylos].

[225] _Phacostylus_ = Lens with styles; [Greek: phakos], [Greek: stylos].

[226] _Triactiscus_ = Shell with three rays; [Greek: triaktiskos].

[227] _Sethostaurus_ = Sieve-cross; [Greek: sêthos], [Greek: stauros].

[228] _Phacostaurus_ = Lens with cross; [Greek: phakos], [Greek: stauros].

[229] _Distriactis_ = Shell with twice three rays; [Greek: distriaktis].

[230] _Heliosestrum_ = Sun-sieve; [Greek: hêlios], [Greek: sêstron].

[231] _Astrosestrum_ = Stellated sieve; [Greek: astron], [Greek: sêstron].

[232] _Heliodiscus_ = Sun-disk; [Greek: hêlios], [Greek: diskos].

[233] _Heliodrymus_ = Sun-forest; [Greek: hêlios], [Greek: drymos].

[234] _Astrophacus_ = Star-lens; [Greek: astron], [Greek: phakos].

[235] _Lithocyclia_ = Circular stone; [Greek: lithos], [Greek: kyklion].

[236] _Coccodiscus_ = Disk with nucleus; [Greek: kokkos], [Greek: diskos].

[237] _Stylocyclia_ = Circular shell with styles; [Greek: stylos], [Greek:
kyklion].

[238] _Amphicyclia_ = Circular shell with spines on both sides; [Greek:
amphi], [Greek: kyklion].

[239] _Trigonocyclia_ = Triangular shell with circular rings; [Greek:
trigonon], [Greek: kyklion].

[240] _Staurocyclia_ = Circular shell with four crossed spines; [Greek:
stauros], [Greek: kyklia].

[241] _Astrocyclia_ = Stellated circular shell; [Greek: astron], [Greek:
kyklion].

[242] _Coccocyclia_ = Circular shell with nucleus; [Greek: kokkos], [Greek:
kyklios].

[243] _Diplactura_ = Stellated shell with double tail; [Greek: diplous],
[Greek: aktis], [Greek: oura].

[244] _Amphiactura_ = Stellated shell with tail on both sides; [Greek:
amphi], [Greek: aktis], [Greek: oura].

[245] _Trigonactura_ = Triangular shell with three rays; [Greek: trigonon],
[Greek: aktis], [Greek: oura].

[246] _Hymenactura_ = Star-shaped shell with membrane between the rays;
[Greek: hymên], [Greek: aktis], [Greek: oura].

[247] _Astractura_ = Star-shaped shell with many rays; [Greek: astron],
[Greek: aktis], [Greek: oura].

[248] _Stauractura_ = Cruciform shell with four rays; [Greek: stauros],
[Greek: aktis], [Greek: oura].

[249] _Pentactura_ = Stellated shell with five rays; [Greek: pente],
[Greek: aktis], [Greek: oura].

[250] _Echinactura_ = _Echinus_-like shell with five rays; [Greek:
echinos], [Greek: aktis], [Greek: oura].

[251] _Archidiscus_ = Primordial disk; [Greek: archidiskos].

[252] _Axodiscus_ = Disk with certain axes; [Greek: axis], [Greek: diskos].

[253] _Porodiscus_ = Porous disk; [Greek: poros], [Greek: diskos].

[254] _Perichlamydium_ = Shell surrounded by a mantle; [Greek: peri],
[Greek: chlamydion].

[255] _Ommatodiscus_ = Disk with eyes; [Greek: omma], [Greek: diskos].

[256] _Stomatodiscus_ = Disk with openings; [Greek: stoma], [Greek:
diskos].

[257] _Xiphodictya_ = Net with swords; [Greek: xiphos], [Greek: diktyon].

[258] _Tripodictya_ = Net with tripod; [Greek: tripous], [Greek: diktyon].

[259] _Staurodictya_ = Cross-net; [Greek: stauros], [Greek: diktyon].

[260] _Stylodictya_ = Net with styles; [Greek: stylos], [Greek: diktyon].

[261] _Stylochlamydium_ = Shell with styles and mantle; [Greek: stylos],
[Greek: chlamydion].

[262] _Amphibrachium_ = Shell with two arms; [Greek: amphi], [Greek:
brachiôn].

[263] _Amphymenium_ = Shell with veil on both sides; [Greek: amphi],
[Greek: hymênion].

[264] _Amphirrhopalum_ = Shell with clubs on both sides; [Greek: amphi],
[Greek: rhopalon].

[265] _Amphicraspedum_ = Shell with borders on both sides; [Greek: amphi],
[Greek: kraspedon].

[266] _Dictyastrum_ = Reticulated star; [Greek: diktyon], [Greek: astron].

[267] _Rhopalastrum_ = Club-star; [Greek: rhopalon], [Greek: astron].

[268] _Hymeniastrum_ = Membranous star; [Greek: hymên], [Greek: astron].

[269] _Euchitonia_ = Nice shell; [Greek: eu], [Greek: chitônia].

[270] _Chitonastrum_ = Star-shell; [Greek: chitônia], [Greek: astron].

[271] _Trigonastrum_ = Triangular star; [Greek: trigônon], [Greek: astron].

[272] _Stauralastrum_ = Crossed sea-star; [Greek: stauros], [Greek: hals],
[Greek: astron].

[273] _Hagiastrum_ = Holy starrulet; [Greek: hagion], [Greek: astron].

[274] _Histiastrum_ = Star with enveloping tissue; [Greek: histion],
[Greek: astron].

[275] _Tessarastrum_ = Starrulet with four rays; [Greek: tessara], [Greek:
astron].

[276] _Stephanastrum_ = Garland-starrulet; [Greek: stephanos], [Greek:
astron].

[277] _Dicranastrum_ = Fork-starrulet; [Greek: dikranon], [Greek: astron].

[278] _Myelastrum_ = Medullary starrulet; [Greek: myelos], [Greek: astron].

[279] _Pentalastrum_ = Little sea-star with five rays; [Greek: pente],
[Greek: hals], [Greek: astron].

[280] _Pentinastrum_ = Starrulet with five rays; [Greek: pente], [Greek:
inos], [Greek: astron].

[281] _Pentophiastrum_ = Starrulet with five snakes; [Greek: pente],
[Greek: ophis], [Greek: astron].

[282] _Hexalastrum_ = Little sea-star with six rays; [Greek: hexa], [Greek:
hals], [Greek: astron].

[283] _Hexinastrum_ = Starrulet with six rays; [Greek: hexa], [Greek:
inos], [Greek: astron].

[284] _Triolena_ = Shell with three arms; [Greek: tria], [Greek: ôlenê].

[285] _Triopyle_ = With three gate-openings; [Greek: tria], [Greek: pylê].

[286] _Triodiscus_ = Disk with three openings; [Greek: triodiskos].

[287] _Pylolena_ = Disk with alternating gates and arms; [Greek: pylê],
[Greek: ôlenê].

[288] _Hexapyle_ = With six gate-openings; [Greek: hexa], [Greek: pylê].

[289] _Pylodiscus_ = Disk with gates; [Greek: pylê], [Greek: diskos].

[290] _Discozonium_ = Disk with girdle; [Greek: diskos], [Greek: zônion].

[291] _Discopyle_ = Disk with gate; [Greek: diskos], [Greek: pylê].

[292] _Spongodiscus_ = Spongy disk; [Greek: spongos], [Greek: diskos].

[293] _Spongophacus_ = Spongy lens; [Greek: spongos], [Greek: phakos].

[294] _Spongolonche_ = Spongy disk with spontoons; [Greek: spongos],
[Greek: lonchê].

[295] _Spongotripus_ = Spongy disk with tripod; [Greek: spongos], [Greek:
tripous].

[296] _Spongostaurus_ = Spongy disk with crossed spines; [Greek: spongos],
[Greek: stauros].

[297] _Stylotrochus_ = Wheel with styles; [Greek: stylos], [Greek:
trochos].

[298] _Spongotrochus_ = Spongy wheel; [Greek: spongos], [Greek: trochos].

[299] _Spongolena_ = Spongy disk with two arms; [Greek: spongos], [Greek:
ôlenê].

[300] _Spongobrachium_ = Spongy shell with two arms; [Greek: spongos],
[Greek: brachiôn].

[301] _Rhopalodictyum_ = Network with clubs; [Greek: rhopalon], [Greek:
diktyon].

[302] _Dictyocoryne_ = Net with clubs; [Greek: diktyon], [Greek: korynê].

[303] _Spongasteriscus_ = Spongy star; [Greek: spongos], [Greek:
asteriskos].

[304] _Spongaster_ = Spongy star; [Greek: spongos], [Greek: astêr].

[305] _Cenolarcus_ = Hollow basket; [Greek: kenos], [Greek: larkos].

[306] _Larcarium_ = A kind of basket; [Greek: larkarion].

[307] _Coccolarcus_ = Basket with kernel; [Greek: kokkos], [Greek: larkos].

[308] _Larcidium_ = Little basket, diminutive of _Larcus_; [Greek:
larkidion].

[309] _Spongolarcus_ = Spongy basket; [Greek: spongos], [Greek: larkos].

[310] _Stypolarcus_ = Hemp basket; [Greek: stypê], [Greek: larkos].

[311] _Larnacilla_ = Little chest, diminutive of _Larnax_; [Greek: larnax].

[312] _Larnacidium_ = Little chest, diminutive of Larnax; [Greek: larnax].

[313] _Larnacalpis_ = Tankard-shaped chest; [Greek: larnax], [Greek:
kalpis].

[314] _Larnacantha_ = Chest with spines; [Greek: larnax], [Greek: akantha].

[315] _Larnacoma_ = Shell of chest form; [Greek: larnax].

[316] _Larnacospongus_ = Spongy chest; [Greek: larnax], [Greek: spongos].

[317] _Larnacostupa_ = Chest with hemp envelop; [Greek: larnax], [Greek:
stupê].

[318] _Monozonium_ = With one girdle; [Greek: monozônion].

[319] _Dizonium_ = With two girdles; [Greek: dizônion].

[320] _Trizonium_ = With three girdles; [Greek: trizônion].

[321] _Amphipyle_ = With one gate on both sides; [Greek: amphi], [Greek:
pylê].

[322] _Tetrapyle_ = With four gate-openings; [Greek: tetrapylê].

[323] _Octopyle_ = With eight gate-openings; [Greek: oktô], [Greek: pylê].

[324] _Pylonium_ = Building with gates; [Greek: pylônion].

[325] _Amphipylonium_ = With one large gate on either side; [Greek: amphi],
[Greek: pylônion].

[326] _Tetrapylonium_ = Building with four gates; [Greek: tetra], [Greek:
pylônion].

[327] _Pylozonium_ = Shell with gates and girdles; [Greek: pylê], [Greek:
zônion].

[328] _Tholartus_ = Cupola-bread; [Greek: tholos], [Greek: artos].

[329] _Tholodes_ = Cupola-shaped; [Greek: tholôdês] _vel_ [Greek:
tholoeidês].

[330] _Amphitholus_ = Shell with two opposite cupolas; [Greek: amphi],
[Greek: tholos].

[331] _Amphitholonium_ = Small shell with two opposite cupolas; [Greek:
amphi], [Greek: tholônion].

[332] _Tholostaurus_ = Cross of four cupolas; [Greek: tholos], [Greek:
stauros].

[333] _Tholoma_ = Dome-building; [Greek: tholôma].

[334] Alternating, [Greek: metallassôn].

[335] _Staurotholus_ = Cupolas cross-wise disposed; [Greek: stauros],
[Greek: tholos].

[336] _Staurotholonium_ = Small shell with cupolas cross-wise disposed;
[Greek: stauros], [Greek: tholônion].

[337] _Tholocubus_ = Cube with six cupolas on its sides; [Greek: tholos],
[Greek: kubos].

[338] _Tholonium_ = Shell with cupolas; [Greek: tholônion].

[339] _Cubotholus_ = Shell with cupolas disposed on six cube-sides; [Greek:
kubos], [Greek: tholos].

[340] _Cubotholonium_ = Small shell with cupolas disposed on the six sides
of a cube; [Greek: kubos], [Greek: tholônion].

[341] _Zonarium_ = Small girdle; [Greek: zônarion].

[342] _Zoniscus_ = Elegant girdle; [Greek: zôniskos].

[343] _Zonidium_ = Little girdle; [Greek: zônidion].

[344] _Spirema_ = Convolution, turning; [Greek: speirêma].

[345] _Lithelius_ = Stony sun; [Greek: lithos], [Greek: hêlios].

[346] _Larcospira_ = Spiral basket; [Greek: larkos], [Greek: speira].

[347] _Pylospira_ = Spiral shell with internal gates; [Greek: pylê],
[Greek: speira].

[348] _Tholospira_ = Shell with spiral domes; [Greek: tholos], [Greek:
speira].

[349] _Spironium_ = Shell with spiral structure; [Greek: speirônion].

[350] _Streblonia_ = Screw-shell; [Greek: streblônion].

[351] _Streblacantha_ = Screw-shell with spines; [Greek: streblê], [Greek:
akantha].

[352] _Streblopyle_ = Screw-shell with gates; [Greek: streblê], [Greek:
pylê].

[353] _Phorticium_ = Small vessel; [Greek: phortikion].

[354] _Spongophortis_ = Spongy vessel; [Greek: spongos], [Greek: phortis].

[355] _Soreuma_ = [Greek: sôreuma], heap.

[356] _Sorolarcus_ = Basket heap; [Greek: sôros], [Greek: larkos].

[357] _Actinelius_ = Radiant sun; [Greek: aktis], [Greek: hêlios].

[358] _Astrolophus_ = Star-like bunch; [Greek: astron], [Greek: lophos].

[359] _Actinastrum_ = Radiant star; [Greek: aktis], [Greek: astron].

[360] _Litholophus_ = Stony brush; [Greek: lithos], [Greek: lophos].

[361] _Chiastolus_ = With crossed arms; [Greek: chiastos], [Greek: ôlos].

[362] _Acanthochiasma_ = Spine-cross; [Greek: akantha], [Greek: chiasma].

[363] _Acanthometron_ = Spine proportion; [Greek: akantha], [Greek:
metron].

[364] _Zygacantha_ = Spines opposite in pairs; [Greek: zyga], [Greek:
akantha].

[365] _Acanthonia_ = Spiny article; [Greek: akantha], [Greek: ônia].

[366] _Lithophyllium_ = With stony leaves; [Greek: lithos], [Greek:
phyllion].

[367] _Phractacantha_ = Spines enclosed by a hedge; [Greek: phraktos],
[Greek: akantha].

[368] _Doracantha_ = Spear-like spine; [Greek: dory], [Greek: akantha].

[369] _Astrolonche_ = Star-spear; [Greek: astron], [Greek: lonchê].

[370] _Xiphacantha_ = Sword spine; [Greek: xiphos], [Greek: akantha].

[371] _Stauracantha_ = Cruciate spine; [Greek: stauros], [Greek: akantha].

[372] _Phatnacantha_ = Spine with fretwork; [Greek: phatnê], [Greek:
akantha].

[373] _Pristacantha_ = Saw-spine; [Greek: pristos], [Greek: akantha].

[374] _Acanthostaurus_ = Spine-cross; [Greek: akantha], [Greek: stauros].

[375] _Belonostaurus_ = Needle cross; [Greek: belonê], [Greek: stauros].

[376] _Lonchostaurus_ = Spear-cross; [Greek: lonchê], [Greek: stauros].

[377] _Zygostaurus_ = Pair-cross; [Greek: zygon], [Greek: stauros].

[378] _Quadrilonche_ = Square-spear.

[379] _Xiphoptera_ = Sword-wing; [Greek: xiphos], [Greek: pteron].

[380] _Lithoptera_ = Stone wing; [Greek: lithos], [Greek: pteron].

[381] _Amphilonche_ = With two opposed spears; [Greek: amphi], [Greek:
lonchê].

[382] _Amphibelone_ = With two opposed needles; [Greek: amphi], [Greek:
belonê].

[383] _Acantholonche_ = Spiny spear; [Greek: akantha], [Greek: lonchê].

[384] _Sphærocapsa_ = Spherical capsule; [Greek: sphaira], [Greek: kapsa].

[385] _Astrocapsa_ = Star-capsule; [Greek: astron], [Greek: kapsa].

[386] _Porocapsa_ = Porous capsule; [Greek: poros], [Greek: kapsa].

[387] _Cannocapsa_ = Tubular capsule; [Greek: kanna], [Greek: kapsa].

[388] _Cenocapsa_ = Hollow Capsule; [Greek: kenos], [Greek: kapsa].

[389] _Phractaspis_ = Hedging shield; [Greek: phraktos], [Greek: aspis].

[390] _Pleuraspis_ = Shield formed by ribs; [Greek: pleura], [Greek:
aspis].

[391] _Dorataspis_ = Spear on the shield; [Greek: dory], [Greek: aspis].

[392] _Diporaspis_ = Shield with two pores; [Greek: diporos], [Greek:
aspis].

[393] _Orophaspis_ = Roof shield; [Greek: orophos], [Greek: aspis].

[394] _Ceriaspis_ = Dimply shield; [Greek: kêrion], [Greek: aspis].

[395] _Hystrichaspis_ = Porcupine-shield; [Greek: hystrix], [Greek: aspis].

[396] _Coscinaspis_ = Sieve-shield; [Greek: koskinon], [Greek: aspis].

[397] _Acontaspis_ = Shield with spears; [Greek: akontion], [Greek: aspis].

[398] _Stauraspis_ = Cross-shield; [Greek: stauros], [Greek: aspis].

[399] _Echinaspis_ = Urchin with shields; [Greek: echinos], [Greek: aspis].

[400] _Zonaspis_ = Shell with a girdle of shield; [Greek: zônê], [Greek:
aspis].

[401] _Dodecaspis_ = Shell with twelve shields; [Greek: dôdeka], [Greek:
aspis].

[402] _Tessaraspis_ = Shield with four pores; [Greek: tessara], [Greek:
aspis].

[403] _Lychnaspis_ = Lantern-shield; [Greek: lychnos], [Greek: aspis].

[404] _Icosaspis_ = Shell with twenty shields; [Greek: eikosi], [Greek:
aspis].

[405] _Hylaspis_ = Forest shield; [Greek: hylê], [Greek: aspis].

[406] _Phractopelta_ = Hedging shield; [Greek: phraktos], [Greek: peltê].

[407] _Pantopelta_ = Shell everywhere with shields; [Greek: pantê], [Greek:
peltê].

[408] _Octopelta_ = Shell with eight shields; [Greek: oktô], [Greek:
peltê].

[409] _Dorypelta_ = Spear with a light shield; [Greek: dory], [Greek:
peltê].

[410] _Stauropelta_ = Cross-shield; [Greek: stauros], [Greek: peltê].

[411] _Thoracaspis_ = Cuirass-shield; [Greek: thôrax], [Greek: aspis].

[412] _Belonaspis_ = Arrow-shield; [Greek: belonê], [Greek: aspis].

[413] _Dictyaspis_ = Net-shield; [Greek: diktyon], [Greek: aspis].

[414] _Coleaspis_ = Shield with sheaths; [Greek: koleos], [Greek: aspis].

[415] _Phatnaspis_ = Panel-shield or fretwork; [Greek: phatnê], [Greek:
aspis].

[416] _Hexalaspis_ = Shield with six wings; [Greek: hexalos], [Greek:
aspis].

[417] _Hexaconus_ = Shell with six cones; [Greek: hexa], [Greek: kônos].

[418] _Hexonaspis_ = Shield with six auricles; [Greek: hexonos], [Greek:
aspis].

[419] _Hexacolpus_ = Shell with six bosoms; [Greek: hexa], [Greek: kolpos].

[420] _Diploconus_ = Double cone; [Greek: diploos], [Greek: kônos].

[421] _Diplocolpus_ = Double bosom; [Greek: diploos], [Greek: kolpos].

* * * * *

Corrections made to printed text

Page iii, § 7: 'Radiolaria' corrected from 'Radioloria'.

Page xiv, § 29: 'Octodendron' corrected from 'Octodendrum'.

Page xvii, § 36: 'allopolar' corrected from 'alloplar'.

Page xxii, § 47: 'rhombic' corrected from 'rombic'.

Page xxvi, § 54B: 'Capsule' corrected from 'Caspule'.

Page xxxiii, § 63: 'Thalassosphærida' corrected from 'Thallassosphærida'.

Page xxxv, § 66.5: 'Nassellaria' corrected from 'Nasellaria'.

Page xlvii, § 77A: 'Mediterranean' corrected from 'Meditteranean'.

Page lxxvi, § 112: 'Phæodinida' corrected from 'Phoeodinida';
'Cannorrhaphida' from 'Cannorhaphida'.

Page lxxxvi, § 131: Heliosphæra referred to Pl. 28, figs. 1-3 but these are
not of this genus.

Page xciv, § 142B: 'Sphærozoea' corrected from 'Sphæozoea'.

Page xcix § 151: Reference to '§ 213' corrected from '§ 273'.

Page cx, § 166: 'interrupted' corrected from 'interruped'.

Page clxxiii, § 249: 'Cannorrhaphida' corrected from 'Cannorrhapida'.

Page clxxix, § 252: 'Karl' (Brandt) corrected from 'Karlt'.

Page clxxxii, § 252: 'Kenntniss' corrected from 'Kentniss'.

Page 29, Thalassosphærida, definition: 'Cannorrhaphida' corrected from
'Cannorhaphida'.

Page 36, Thalassoplancta, definition: 'Cannorrhaphis' corrected from
'Cannorhaphis'.

Page 43, Sphærozoum armatum: 'fig. 9' corrected from 'figs. 1, 9'.

Page 85, Cromyosphæra antarctica refers to Pl. 30, fig. 8 but that does not
show this species

Page 87, Genus 26: 'Liosphærida' corrected from 'Liospærida'.

Page 99, Buccinosphæra invaginata, crystals: '0.008' corrected from
'0.088'.

Ib., Buccinosphæra tubaria, pores: '0.008 to 0.012' corrected from '0.008
to 0.002' (cf. "three to four times as broad as the bars").

Page 130, Xiphostylus alca: 'Pl. 13' corrected from 'Pl. 14'.

Page 132, Saturnalis annularis, major axis: '0.27 to 0.3' corrected from
'0.27 to 0.03'.

Page 137, Stylosphæra jugata, pores: '0.005 to 0.02' corrected from '0.005
to 0.002'.

Page 141, Sphærostylus ophidium, pores: '0.005 to 0.008' corrected from
'0.05 to 0.008'.

Page 201, Hexacromyida, definition: 'concentric' corrected from
'concentic'.

Page 203, Cubosphæra concentrica, dimensions (A) and (B): '0.02' and '0.04'
corrected from '0.2', '0.4' (cf. the given radial proportions).

Page 222, Coscinomma: 'n. gen.' corrected from 'n. sp.'.

Page 224, Cladococcinus: 'Subgenus' corrected from 'Sugenus'.

Page 241, Heliosoma hastatum, outer pores: '0.006' corrected from '0.06'.

Ib., Heliosoma indicum, outer pores: '0.004 to 0.01' corrected from '0.004
to 0.1' (cf. "twice to five times as broad as the bars").

Page 253, Actinomma japonicum, middle: '0.1' corrected from '0.01'.

Page 260, Pityomma piniferum, middle: '0.05' corrected from '0.5'; inner
'0.025' from '0.25'.

Ib., Pityomma drymodes, middle: '0.1' corrected from '0.01'.

Page 261, Cromyomma perplexum, diameter (B): '0.13' corrected from '0.013'.

Page 271, Spongiomma spathillatum: 'Octodendron' corrected from
'Octodendrom'.

Page 291, Cenellipsis ehrenbergii, shorter axis: '0.08 to 0.1' corrected
from '0.8 to 0.1'.

Page 307, synopsis, 141: 'Cromyatractus' corrected from 'Cromyotractus'.

Page 308, Druppula cocos, medullary shell: '0.03 and 0.035' corrected from
'0.03 and 0.35'.

Page 309, Druppula phoenix, minor axis: '0.08' corrected from '0.8'.

Page 310, Druppula elliptica: 'irregularly' corrected from 'irreguarlly'.

Page 314, Prunulum triplex, minor axis: '0.065' corrected from '0.65'.

Page 324, Druppatractus hippocampus: 'Druppatractus' corrected from
'Druppactractus'.

Page 372, Cyphinidium apicatum, pores: '0.005 to 0.01' corrected from
'0.005 to 0.001'.

Page 380, Panartus fusiformis, breadth: '0.08 to 0.1' corrected from '0.08
to 0.01'.

Ib., Panartus profundissimus, pores: '0.015' corrected from '0.15' (cf.
"six to eight times as broad as the bars").

Page 405, ref. to Saturnalis: 'Pl. 13, fig. 16' corrected from 'fig. 6'.

Page 418, Trochodiscus solaris: reference to 'Heliodiscus solaster'
corrected from 'solaris' per main text entry.

Page 468, Coccocyclia heliantha: 'Coccocyclia' corrected from 'Coccocylia'.

Page 469, Diplactura: reference to 'Spongolena' corrected from
'Spongolene'.

Page 485, synopsis, 236: 'Tessarastrum' corrected from 'Tesserastrum'.

Page 508, Staurodictya ocellata: 'Stylodictya' corrected from 'Stylodicta'.

Page 513, Stylodictya dujardinii: 'Stylodictya' corrected from
'Stylodicyta'.

Page 538: 'Chitonastrum dicranoides' corrected from 'Chitonastrum
dicranodes' (per Index).

Page 544, Hagiastrum christi: 'Tessarastrum' corrected from 'Tesserastrum'.

Page 548, subgenus 2: 'Tessarastromma' corrected from 'Tessarostromma'.

Page 557, Pentinastrum, definition: 'Histiastrum' corrected from
'Histriastrum'.

Page 561, last paragraph: 'Archidiscus hexoniscus' corrected from
'Archidiscus hexonicus'.

Page 564, Triolena trispinosa, diameter: '0.55' corrected from '0.55'.

Page 577, Spongodiscus favus, outer meshes: '0.008 to 0.012' corrected from
'0.08 to 0.012'.

Page 580, Spongolonche conostyla refers to Pl. 48, fig. 7 which does not
show this species.

Page 593, Dictyocoryne trigona: (basal) 'breadth' corrected from 'breath'.

Page 606, first paragraph: 'Actilarcus' corrected from 'Actolarcus'.

Page 606, Cenolarcus: 'Actilarcus' corrected from 'Actolarcus'.

Page 608, Larcarium amphistylum, breadth: '0.12' corrected from '0.012'.

Page 612, Larcidium polyacanthum, medullary shell: '0.03 to 0.09' corrected
from '0.3 to 0.09'.

Page 625, Larnacoma quadruplex, breadth (A): '0.016' corrected from '0.16'.

Ib., Larnacoma hexagonium: 'inner cortical and outer medullary' corrected
from 'outer cortical and inner medullary'.

Page 632, synopsis: 'Amphipylonium' corrected from 'Amhipylonium'.

Page 654, Octopyle decastyle, breadth: '0.032' corrected from '0.32'.

Page 657, Amphipylonium spinosissimum, length of the inner cortical shell:
'0.16' corrected from '0.016'.

Page 657, Amphipylonium tetraceros: 'breadth' corrected from 'breath'.

Page 660, Tholonida: (we distinguish in this) 'family' corrected from
'subfamily'.

Page 665, Tholodes, definition: 'Tholartus' corrected from 'Thalartus'.

Page 666, Amphitholus: 'lentelliptical' corrected from 'lentelleptical'. So
also p. 666, Amphitholonium octostylium.

Page 677, pores: '0.004 to 0.008' corrected from '0.04 to 0.008', cf. "once
to three times as broad as the bars".

Page 709, Phorticium pylonium, breadth: '0.045' corrected from '0.45'.

Page 742: 'Acanthometron elasticum' corrected from 'Acathometron'.

Page 748, Zygacantha semicompressa, habitat (author): 'Car' corrected from
'Czar'.

Page 765, Phatnacantha: 'Tessaraspida', 'Tessaraspis' corrected from
'Tesseraspida', 'Tesseraspis'.

Page 786, Amphilonche messanensis, length of the minor spines: '0.05 to
0.09' corrected from '0.5 to 0.09'.

Page 826, Coscinaspis rhacopora, diameter: '0.15' corrected from '1.15'.

Page 827, Coscinaspis orthopora: 'Phatnaspis lacunaria' corrected from
'Phatnaspis lacumaria'.

Page 834, Dodecaspis tricinata, diameter: '0.016' corrected from '0.16'.

Page 844, Icosaspis elegans, smaller pores: '0.001 to 0.003' corrected from
'0.01 to 0.003'.

Page 852, Phractopelta: 'apophyses' corrected from 'aphophyses'.

Page 867, Coleaspis occulta: 'Coleaspis' corrected from 'Coleapsis'.

*** End of this LibraryBlog Digital Book "Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia) - Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII" ***

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